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043.0000 Family Cystopteridaceae (Payer) Shmakov; Turczaninowia 4: 60 (2001)
3-4 genera, ca. 35 spp. Family status confirmed by Rothfels et al. (2012)
Species listed in database: 39; hybrids: 11
= Cystopteridinae Payer; Bot. Cryptog. 198 (1850) (Cystopterideae)
= Cystopterioideae Ching & Z.R.Wang; Acta Phytotax. Sin. 35: 34 (1997)
043.0002 Genus Acystopteris Nakai; Bot. Mag. (Tokyo) 47: 180 (1933)
Distribution: Himalaya to Japan and Malesia. Valid genus separate from Cystopteris fide Fraser-Jenkins et al. (2018)
Species listed in database: 3
= Cystopteris subgen.Acystopteris (Nakai) Blasdell; Mem. Torrey Bot. Club 21(4): 49 (1963)
species :
Acystopteris japonica (Luerss.) Nakai; Bot. Mag. (Tokyo) 47: 180 (1933)
Distribution: China (Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Yunnan, Zhejiang); Japan
36 CHI 38 JAP
Remarks: not in Taiwan
Synonyms: hidden
= Acystopteris abbreviata W.M.Chu; Acta Phytotax. Sin. 21: 221 (1983)
= Acystopteris angustifolia S.F.Wu; J. Wuhan Bot. Res. 13(1): 24-26, f. 1 (1995)
= Acystopteris attenuata Ching; Acta Phytotax. Sin. 9: 68 (1964)
= Acystopteris hunanensis Z.R.Wang & S.F.Wu; Acta Phytotax. Sin. 28: 165 (1990)
= Acystopteris jinfoshanensis Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 11-12, f. 13 (1984)
= Acystopteris longipes Ching; Acta Phytotax. Sin. 9: 68 (1964)
= Acystopteris membranacea Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 10-11, f. 12 (1984)
= Acystopteris pachyphylla Ching; Acta Phytotax. Sin. 9: 66 (1964)
= Acystopteris petiolata Ching; Acta Phytotax. Sin. 9: 67 (1964)
= Cystopteris japonica Luerss.; Bot. Jahrb. Syst. 4: 363 (1883)
species :
Acystopteris taiwaniana (Tagawa) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977)
Distribution: Japan; Taiwan
38 JAP TAI
Synonyms: hidden
= Acystopteris japonica var. taiwaniana (Tagawa) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 13: 51 (1976)
= Cystopteris japonica var. taiwaniana Tagawa; Acta Phytotax. Geobot. 4(2): 57 (1935)
species :
Acystopteris tenuisecta (Blume) Tagawa; Acta Phytotax. Geobot. 7: 73 (1938)
Distribution: China (Yunnan, Sichuan, Guangxi); Tibet; S-Japan (incl. Yakushima Isl.); Taiwan; Nepal; Bhutan; India (Sikkim, Arunachal Pradesh, Meghalaya, Nagaland, Uttarakhand, West Bengal); Myanmar (Kachin, etc.); Thailand; Vietnam; peninsular Malaysia (Cameron Highlands, Pahang, Perak, Selangor); Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; Moluccas (Seram); New Guinea (Papua New Guinea)
36 CHI TIB 38 JAP NNS TAI 40 BHU IND NEP 41 MYA THA VIE 42 BOR JAW MLY MOL NWG PHI SUL
Synonyms: hidden
= Alsophila tenuisecta Blume ex T.Moore; Index Fil. [T. Moore] 68 (1857)
= Aspidium tenuisectum Blume; Enum. Pl. Javae 2: 170 (1828)
= Asplenium tenuisectum (Blume) Hook.; Sp. Fil. [W.J.Hooker] 3: 222 (1860)
= Athyrium tenuisectum (Blume) T.Moore; Index Fil. [T. Moore] 188 (1860)
= Cornopteris tenuisecta (Blume) Tardieu; Amer. Fern J. 48: 32 (1958)
= Ctenitis tenerifrons (Christ) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 125 (1947)
= Cystopteris formosana Hayata; Icon. Pl. Formosan. 4: 143, f. 83 (1914)
= Cystopteris setosa (Bedd.) Bedd.; Ferns Brit. India, t. 312 (1869)
= Cystopteris tenuisecta (Blume) Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 241 (1864)
= Davallia setosa (Bedd.) Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 2: 468 (1874), non Sm. (1808)
= Dryopteris dennstaedtioides Copel.; Philipp. J. Sci. 56: 473, t. 3 (1935)
= Dryopteris tenerifrons (Christ) C.Chr.; Index Filic., Suppl. Tert. 99 (1934)
= Hypolepis tenerifrons Christ; Philipp. J. Sci. 3: 274 (1908)
= Lastrea setosa Bedd.; Ferns Brit. India, t. 262 (1868) (non C. Presl 1848, nec T. Moore (1858))
= Thelypteris tenerifrons (Christ) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 254 (1941)
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.02 (Feb. 1st, 2026)

Covering 358,774 accepted species,
8,847 hybrids, 48,003 accepted infraspecific taxa and 1,006,442 synonyms,
in total 1,422,066* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

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