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492.0000 Family Rubiaceae Juss.; Gen. Pl. [Jussieu] 196 (1789), nom. cons.
589 genera, ca. 14.200 spp. mostly tropics, comparably few temperate. Phylogeny: Razafimandimbison & Rydin (2024)
Species listed in database: 14370; hybrids: 47
= Aparinaceae Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 2: 3, 38 (1813-1829)
= Asperulaceae Cham. ex Spenn.; Handb. Angew. Bot. 2: 493 (1835)
= Catesbaeaceae Martinov; Tekhno-Bot. Slovar 111 (1820)
= Cephalanthaceae Raf.; Ann. Gén. Sci. Phys. 6: 86 (1820)
= Cinchonaceae Batsch; Tab. Affin. Regni Veg. 234 (1802)
= Coffeaceae Batsch; Tab. Affin. Regni Veg. 233 (1802)
= Coutareaceae Martinov; Tekhno-Bot. Slovar 168 (1820)
= Dialypetalanthaceae Rizzini & Occhioni; Lilloa 17: 253 (1948), nom. cons.
= Galiaceae Lindl.; Intr. Nat. Syst. Bot., ed. 2: 249 (1836)
= Gardeniaceae Dumort.; Anal. Fam. Pl. 29, 32 (1829)
= Guettardaceae Batsch; Tab. Affin. Regni Veg. 235 (1802)
= Hameliaceae Mart.; Consp. Regn. Veg. [Martius] 31 (1835)
= Hedyotidaceae Dumort.; Commentat. Bot. [Dumortier] 57 (1822)
= Henriqueziaceae Bremek.; Acta Bot. Neerl. 6 (Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 141): 371 (1957)
= Houstoniaceae Raf.; Good Book 21 (1840)
= Hydrophylacaceae Martinov; Tekhno-Bot. Slovar 318 (1820)
= Lippayaceae Meisn.; Pl. Vasc. Gen. [Meisner], Tab. Diagn. 156, Comm. 112 (1838)
= Lygodisodeaceae Bartl.; Ord. Nat. Pl. 123, 207 (1830)
= Naucleaceae Wernham; New Phytol. 11: 225 (1912)
= Nonateliaceae Martinov; Tekhno-Bot. Slovar 420 (1820)
= Operculariaceae Juss. ex Perleb; Vers. Arzneikr. Pfl. 207 (1818)
= Pagamaeaceae Martinov; Tekhno-Bot. Slovar 447 (1820)
= Psychotriaceae F.Rudolphi; Syst. Orb. Veg. 49 (1830)
= Randiaceae Martinov; Tekhno-Bot. Slovar 534 (1820)
= Sabiceaceae Martinov; Tekhno-Bot. Slovar 555 (1820)
= Spermacoceaceae Bercht. & J.Presl; Přir. Rostlin 256 (1820)
= Theligonaceae Dumort.; Anal. Fam. Pl. 15, 17 (1829), nom. cons.
492.8610 Genus Atractocarpus Schltr. & K.Krause; Bot. Jahrb. Syst. 40(Beibl. 92): 43 (1908)
Distribution: Australia, Pacific, many in New Caledonia
Species listed in database: 40
= Franciella Guillaumin; Bull. Mus. Natl. Hist. Nat. 28: 197 (1922)
= Neofranciella Guillaumin; Bull. Mus. Natl. Hist. Nat. 31: 481 (1925)
= Sukunia A.C.Sm.; Bull. Bernice P. Bishop Mus. 141: 136 (1936)
= Sulitia Merr.; Philipp. J. Sci. 29: 494 (1926)
= Trukia Kaneh.; Bot. Mag. (Tokyo) 49: 278 (1935)
species :
Atractocarpus aragoensis Guillaumin; Arch. Bot. Mém. 3(5): 11 (1930)
Distribution: New Caledonia
60 NWC
species :
Atractocarpus artensis (Montrouz.) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Gardenia artensis Montrouz.; Mém. Acad. Roy. Sci. Lyon, Sect. Sci., Sér. 2, 10: 215 (1860)
= Gardenia edulis Vieill.; Ann. Sci. Nat., Bot., Sér. 4, 16: 65 (1862), nom. illeg.
= Gynochthodes artensis (Montrouz.) Razafim. & B.Bremer; Adansonia, Sér. 3, 33(2): 287 (2011)
= Morinda artensis Montrouz.; Mém. Acad. Roy. Sci. Lyon, Sect. Sci., Sér. 2, 10: 219 (1860)
= Randia artensis (Montrouz.) Guillaumin; Arch. Bot. Mém. 3(5): 8 (1930)
species :
Atractocarpus baladicus (Montrouz. ex Guillaumin & Beauvis.) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Atractocarpus sessilifolius Guillaumin; Arch. Bot. Mém. 3(5): 13 (1930)
= Gardenia baladica Montrouz. ex Guillaumin & Beauvis.; Ann. Soc. Bot. Lyon 38: 95 (1913 publ. 1914)
= Gardenia edulis Montrouz.; Mém. Acad. Roy. Sci. Lyon, Sect. Sci., Sér. 2, 10: 216 (1860), nom. illeg.
= Randia baladica (Montrouz. ex Guillaumin & Beauvis.) Guillaumin; Arch. Bot. Mém. 3(5): 8 (1930)
species :
Atractocarpus benthamianus (F.Muell.) Puttock; Austral. Syst. Bot. 12(2): 277 (1999)
subspecies :
subsp. benthamianus
Distribution: Australia (Queensland, New South Wales)
50 AUS
Synonyms: hidden
= Gardenia benthamiana F.Muell.; Fragm. [Mueller] 9: 180 (1875), pro syn.
= Gardenia chartacea var. latifolia F.Muell. ex Benth.; Fl. Austral. [Bentham] 3: 410 (1867)
subspecies :
subsp. glaber Puttock; Austral. Syst. Bot. 12(2): 281 (1999)
Distribution: Australia (New South Wales, Queensland)
50 AUS
species :
Atractocarpus brandzeanus (Baill.) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Randia brandzeana Baill.; Adansonia 12: 244 (1879)
species :
Atractocarpus carolinensis (Valeton) Puttock; Austral. Syst. Bot. 12(2): 300 (1999)
Distribution: Fed. States of Micronesia (Yap, Truk, Moen, Dublon, Tol, Fefan, Udot, Uman)
62 MIC
Synonyms: hidden
= Randia carolinensis Valeton; Bot. Jahrb. Syst. 63: 301 (1930)
= Rhopalobrachium megacarpum (Kaneh.) Kaneh.; Bot. Mag. (Tokyo) 46: 674 (1932)
= Timonius megacarpus Kaneh.; Trop. Woods, No. 29: 5, nomen, et in Bot. Mag., Tokyo,1932 46: 494 (1932), descr.
= Trukia carolinensis (Valeton) Kaneh. & Hatus.; Bot. Mag. (Tokyo) 50: 606 (1936)
= Trukia megacarpa (Kaneh.) Kaneh.; Bot. Mag. (Tokyo) 49: 279 (1935)
species :
Atractocarpus chartaceus (F.Muell.) Puttock; Austral. Syst. Bot. 12(2): 282 (1999)
Distribution: Australia (Queensland, New South Wales)
50 AUS
Synonyms: hidden
= Gardenia chartacea F.Muell.; Essay Pl. Burdekin 12 (1860)
= Randia chartacea (F.Muell.) F.Muell.; Fragm. [Mueller] 9: 180 (1875)
species :
Atractocarpus colnettianus (Guillaumin) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Gardenia colnettiana Guillaumin; Mém. Mus. Natl. Hist. Nat., Sér. B, Bot., n. s. 8: 93 (1957)
species :
Atractocarpus confertus (Guillaumin) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Gardenia conferta Guillaumin; Arch. Bot. Mém. 3(5): 5 (1930)
= Genipa conferta Baill. ex Guillaumin; Arch. Bot. Mém. 3(5): 5 (1930)
species :
Atractocarpus crosbyi (Burkill) Puttock; Austral. Syst. Bot. 12(2): 301 (1999)
Distribution: Tonga (Vava´u)
60 TON
Synonyms: hidden
= Porterandia crosbyi (Burkill) A.C.Sm. & S.P.Darwin; Fl. Vit. Nova 4: 190 (1988)
= Randia crosbyi Burkill; J. Linn. Soc., Bot. 35: 41 (1890)
species :
Atractocarpus cucumicarpus S.Moore; J. Linn. Soc., Bot. 45: 330 (1921)
Distribution: New Caledonia
60 NWC
species :
Atractocarpus decorus (Valeton) Puttock; Austral. Syst. Bot. 12(2): 301 (1999)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Randia decora Valeton; Bot. Jahrb. Syst. 60: 90, et in Nova Guinea 14: 266 (1925)
species :
Atractocarpus deplanchei (Vieill. ex Guillaumin) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Gardenia deplanchei Vieill. ex Guillaumin; Arch. Bot. Mém. 3(5): 6 (1930)
= Genipa deplanchei Baill. ex Guillaumin; Arch. Bot. Mém. 3(5): 6 (1930)
species :
Atractocarpus fitzalanii (F.Muell.) Puttock; Austral. Syst. Bot. 12(2): 285 (1999)
Distribution: Australia (Queensland)
50 AUS
Synonyms: hidden
= Atractocarpus fitzalanii subsp. tenuipes Puttock; Austral. Syst. Bot. 12(2): 288 (1999)
= Gardenia fitzalanii F.Muell.; Essay Pl. Burdekin 12 (1860)
= Randia fitzalanii (F.Muell.) F.Muell. ex Benth.; Fl. Austral. [Bentham] 3: 411 (1867)
= Trukia fitzalanii (F.Muell.) Fosberg; Phytologia 62: 175 (1987)
species :
Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis.; Ann. Soc. Bot. Lyon 38: 96 (1913 publ. 1914), et Sp. Montrouz. 22 (1914) (reprint)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Atractocarpus balansanus Guillaumin; Arch. Bot. Mém. 3(5): 12 (1930)
= Atractocarpus bracteatus Schltr. & K.Krause; Bot. Jahrb. Syst. 40(Beibl. 92): 43 (1908)
= Atractocarpus heterophyllus Guillaumin & Beauvis.; Ann. Soc. Bot. Lyon 38: 22 (1913 publ. 1914)
= Gardenia fusiformis Baill. ex Heckel; Ann. Mus. Colon. Marseille, Sér. 2, 10: 256 (1912)
= Gardenia heterophylla Montrouz.; Mém. Acad. Roy. Sci. Lyon, Sect. Sci., Sér. 2, 10: 216 (1860)
= Genipa balansana Baill. ex Guillaumin; Arch. Bot. Mém. 3(5): 12 (1930)
= Genipa xylopioides Baill. ex Guillaumin; Arch. Bot. Mém. 3(5): 12 (1930)
species :
Atractocarpus hirtus (F.Muell.) Puttock; Austral. Syst. Bot. 12(2): 289 (1999)
Distribution: Australia (Northern Territory, Queensland)
50 AUS
Synonyms: hidden
= Gardenia hirta F.Muell.; Fragm. [Mueller] 7: 46 (1869)
= Randia hirta (F.Muell.) F.Muell.; Syst. Census Austral. Pl. 1: 74 (1882)
= Rothmannia hirta (F.Muell.) Fagerl.; Ark. Bot. 30A(7): 39 (1943)
species :
Atractocarpus longipes (A.C.Sm.) Puttock; Austral. Syst. Bot. 12(2): 302 (1999)
Distribution: Fiji
60 FIJ
Synonyms: hidden
= Sukunia longipes A.C.Sm.; Pacific Sci. 23: 392 (1969)
species :
Atractocarpus longistipitatus Guillaumin; Arch. Bot. Mém. 3(5): 12 (1930)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Genipa longistipitata Baill. ex Guillaumin; Arch. Bot. Mém. 3(5): 12 (1930)
species :
Atractocarpus macarthurii (F.Muell.) Puttock; Austral. Syst. Bot. 12(2): 303 (1999)
Distribution: New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Solomon Isl.
42 NWG 43 BIS SOL
Synonyms: hidden
= Gardenia dryadum S.Moore; J. Bot. 65: 246 (1927)
= Randia albituba Valeton; Bot. Jahrb. Syst. 60: 92 (1925)
= Randia dryadum (S.Moore) Merr. & L.M.Perry; J. Arnold Arbor. 25: 201 (1944)
= Randia macarthurii F.Muell.; Descr. Notes Papuan Pl. 1: 68 (1876)
= Trukia dryadum (S.Moore) Fosberg; Phytologia 62(3): 175 (1987)
= Trukia macarthurii (F.Muell.) Fosberg; Phytologia 62(3): 176 (1987)
species :
Atractocarpus merikin (F.M.Bailey) Puttock; Austral. Syst. Bot. 12(2): 292 (1999)
Distribution: Australia (Queensland)
50 AUS
Synonyms: hidden
= Gardenia merikin F.M.Bailey; Queensl. Fl. 6: 2005 (1902)
species :
Atractocarpus mollis (Schltr.) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Gardenia mollis Schltr.; Bot. Jahrb. Syst. 39: 256 (1906)
= Genipa conica Baill. ex Guillaumin; Arch. Bot. Mém. 3(5): 6 (1930)
species :
Atractocarpus ngoyensis (Schltr.) Mouly; Pl. Ecol. Evol. 154(1): 117 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Gardenia ngoyensis Schltr.; Bot. Jahrb. Syst. 39: 257 (1906)
= Randia ngoyensis (Schltr.) Hutch. ex S.Moore; J. Linn. Soc., Bot. 45: 328 (1921)
species :
Atractocarpus nigricans (Schltr.) Mouly; Pl. Ecol. Evol. 154(1): 118 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Randia nigricans Schltr.; Bot. Jahrb. Syst. 39(2): 258 (1906)
species :
Atractocarpus oblongus S.Moore; J. Linn. Soc., Bot. 45: 331 (1921)
Distribution: New Caledonia
60 NWC
species :
Atractocarpus obscurinervius (Merr.) Puttock; Austral. Syst. Bot. 12(2): 304 (1999)
Distribution: Philippines (Luzon, Mindanao, Catanduanes)
42 PHI
Synonyms: hidden
= Gardenia obscurinervia Merr.; Philipp. J. Sci., C 11: 32 (1916)
= Sulitia longiflora Merr.; Philipp. J. Sci. 29: (1926)
= Sulitia obscurinervia (Merr.) Ridsdale; Blumea 25(1): 301 (1979)
species :
Atractocarpus pancherianus (Guillaumin) Mouly; Pl. Ecol. Evol. 154(1): 118 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Randia pancheriana Guillaumin; Arch. Bot. Mém. 3(5): 10 (1930)
species :
Atractocarpus pentagonioides (Seem.) Puttock; Austral. Syst. Bot. 12(2): 305 (1999)
Distribution: Fiji
60 FIJ
Synonyms: hidden
= Sukunia pentagonioides (Seem.) A.C.Sm.; Bull. Bernice P. Bishop Mus. 141, 137 (1936)
= Gardenia pentagonioides Seem.; Fl. Vit. [Seemann] 122 (1866)
species :
Atractocarpus platixylon (Vieill. ex Pancher & Sebert) Guillaumin; Arch. Bot. Mém. 3(5): 13 (1930)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Gardenia platixylon Vieill. ex Pancher & Sebert; Not. Bois Nouv. Caléd. 183 (1874)
species :
Atractocarpus pseudoterminalis (Guillaumin) Mouly; Pl. Ecol. Evol. 154(1): 118 (2021)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Randia pseudoterminalis Guillaumin; Mém. Mus. Natl. Hist. Nat., Sér. B, Bot., n. s. 8: 96 (1957)
species :
Atractocarpus pterocarpon (Guillaumin) Puttock; Austral. Syst. Bot. 12(2): 305 (1999)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Franciella pterocarpon Guillaumin; Bull. Mus. Natl. Hist. Nat. 28: 197 (1922)
= Neofranciella pterocarpon Guillaumin; Bull. Mus. Natl. Hist. Nat. 31: 481 (1925)
species :
Atractocarpus rotundifolius Guillaumin; Mém. Mus. Natl. Hist. Nat., Sér. B, Bot., n. s. 8: 95 (1957)
Distribution: New Caledonia
60 NWC
species :
Atractocarpus sessilis (F.Muell.) Puttock; Austral. Syst. Bot. 12(2): 294 (1999)
Distribution: New Guinea (Papua New Guinea); Australia (Queensland)
42 NWG 50 AUS
Synonyms: hidden
= Gardenia sessilis F.Muell.; Fragm. [Mueller] 7: 47 (1869), pro syn.
= Randia sessilis F.Muell.; Fragm. [Mueller] 7: 47 (1869)
species :
Atractocarpus sezitat (Guillaumin) Mouly; Pl. Ecol. Evol. 154(1): 118 (2021)
Distribution: New Caledonia; Vanuatu
60 NWC VAN
Synonyms: hidden
= Genipa sezitat Baill. ex Guillaumin; Bull. Mus. Natl. Hist. Nat. 25: 646 (1919)
= Randia sezitat Guillaumin; Arch. Bot. Mém. 3(5): 9 (1930)
species :
Atractocarpus simulans Guillaumin; Bull. Mus. Natl. Hist. Nat., Sér. 2, 5: 247 (1933)
Distribution: New Caledonia
60 NWC
species :
Atractocarpus stipularis (F.Muell.) Puttock ex P.S.Green; Kew Bull. 45(2): 246 (1990)
Distribution: Lord Howe Isl.
50 LHI
Synonyms: hidden
= Gardenia stipulosa F.Muell.; Fragm. [Mueller] 7: 47 (1869)
= Randia stipularis F.Muell.; Descr. Notes Papuan Pl. 69 (1890)
= Randia stipulosa C.Moore & F.Muell.; Fragm. [Mueller] 7: 47 (1869), nom. superfl.
species :
Atractocarpus tahitiensis (Nadeaud) Puttock; Austral. Syst. Bot. 12(2): 306 (1999)
Distribution: Society Isl. (Tahiti)
61 SCI
Synonyms: hidden
= Randia tahitiensis Nadeaud; Énum. Pl. Tahiti 54 (1873)
= Trukia tahitensis (Nadeaud) Fosberg; Phytologia 62(3): 174 (1987)
species :
Atractocarpus teamotuaitaui Mouly & Butaud; Adansonia, Sér. 3, 45(10): 158 (2023)
Distribution: Society Isl. (Tahiti)
61 SCI
species :
Atractocarpus tenuiflorus (A.C.Sm.) Puttock; Austral. Syst. Bot. 12(2): 306 (1999)
Distribution: Fiji
60 FIJ
Synonyms: hidden
= Porterandia tenuiflora (A.C.Sm.) A.C.Sm. & S.P.Darwin; Fl. Vit. Nova 4: 192 (1988)
= Randia tenuiflora A.C.Sm.; Bull. Bernice P. Bishop Mus. 141, 135 (1936)
species :
Atractocarpus vaginatus Guillaumin; Arch. Bot. Mém. 3(5): 14 (1930)
Distribution: New Caledonia
60 NWC
species :
Atractocarpus versteegii (Valeton) Puttock; Austral. Syst. Bot. 12(2): 307 (1999)
Distribution: New Guinea (Irian Jaya)
42 NWG
Synonyms: hidden
= Randia versteegii Valeton; Nova Guinea 8: 466 (1911). 8: 466 (1911)
= Randia eymai Steenis; Bull. Jard. Bot. Buitenzorg, Sér. 3, 17: 399 (1948)
= Randia klossii (Wernham) Valeton; Nova Guinea 14: 267 (1925), nec Ridl. (1920)
= Gardenia klossii Wernham; Trans. Linn. Soc. London, Bot., Ser. 2, 9: 72 (1916)
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life