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060.0000 Family Zamiaceae Rchb.; Handb. Nat. Pfl.-Syst. 139 (1837)
9 genera, ca. 230 spp.
Species listed in database: 260; hybrids: 1
= Boweniaceae D.W.Stev.; Amer. J. Bot. 68: 1114 (1981)
= Bowenieae J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 63 (1932)
= Bowenioideae Pilg.; Nat. Pflanzenfam. [Engler & Prantl], 2. Aufl., 13: 73 (1926)
= Ceratozamieae J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 64 (1932)
= Dioaceae Doweld; Tent. Syst. Pl. Vasc.: xv. (2001)
= Diooneae J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 64 (1932)
= Dioonoideae Pilg.; Nat. Pflanzenfam. [Engler & Prantl], 2. Aufl., 13: 73 (1926)
= Encephalartaceae Schimp. & Schenk; Handb. Palaeontol., Palaeophyt. 2: 215 (1880)
= Encephalarteae Miq.; Prodr. Syst. Cycad. 5 (1861)
= Encephalartinae A.DC.; Prodr. [A.P. de Candolle] 16(2): 530 (1868) (Encephalarteae)
= Encephalartoideae D.W.Stev.; Brittonia 44: 221 (1992)
= Macrocycadinae D.W.Stev.; Brittonia 44: 222 (1992)
= Macrozamieae J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 64 (1932)
= Macrozamiinae D.W.Stev.; Brittonia 44: 222 (1992)
= Macrozamioideae J.H.Schaffn.; Ohio Naturalist 13: 106 (1913) (Macrozamiatae)
= Microcycadaceae Tarbaeva; Anat.-Morf. Str. Sem. Cycad. 19 (1991)
= Microcycadeae J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 64 (1932)
= Microcycadinae W.D.Stev.; Brittonia 44: 222 (1992)
= Microcycadoideae Tarbaeva; Byull. Glavn. Bot. Sada 173: 111, 112 (1996) (Microcycaideae)
= Stangeriaceae Schimp. & Schenk; Handb. Palaeontol., Palaeophyt. 216 (1880)
= Stangerieae Miq.; Prodr. Syst. Cycad. 5 (1861)
= Stangeriinae A.DC.; Prodr. [A.P. de Candolle] 16(2): 529 (1868) (Stangerieae)
= Stangerioideae Pilger; Nat. Pflanzenfam. [Engler & Prantl], 2. Aufl., 13: 73 (1926)
= Zamieae Miq.; Prodr. Syst. Cycad. 5 (1861)
= Zamiinae Benth. & Hook.f.; Gen. Pl. [Bentham & Hooker f.] 3: 444 (1880) (Zamieae)
= Zamiineae D.W.Stev.; Brittonia 44: 222 (1992)
= Zamioideae Potonié; Lehrb. Pfl.-Palaeont. 3: 275, 276 (1898) (Zamieae)
= Zamioideae J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 84 (1932)
= Zamioideae Pilg.; Nat. Pflanzenfam. [Engler & Prantl], 2. Aufl., 13: 73 (1926)
060.0007 Genus Ceratozamia Brongn.; Ann. Sci. Nat., Bot., Sér. 3, 5: 7, t. 1 (1846)
Distribution: Mexico to Belize. Monograph: Martinez-Dominguez et al. (2022)
Species listed in database: 43
= Dipsacozamia Lehm.; Veg. Kingd. 225 (1847)
= Eriozamia J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 131 (1932), syn. (sine descr)
species :
Ceratozamia alba Pérez-Farr., Gut.Ortega & Vovides; Phytotaxa 666(4): 265 (2024)
Distribution: Mexico (Chiapas)
79 MEX
species :
Ceratozamia alvarezii Pérez-Farr., Vovides & Iglesias; Novon 9, 410 (1999)
Distribution: Mexico (Chiapas)
79 MEX
Conservation Status: Endangered (known only from a small area)
species :
Ceratozamia aurantiaca Pérez-Farr., Gut.Ortega, J.L.Haynes & Vovides; Taxonomy 1(3): 249 (2021)
Distribution: Mexico (Oaxaca)
79 MEX
Synonyms: hidden
= Ceratozamia martinezii Mart.-Domínguez, Nic.-Mor. & D.W.Stev.; Nordic J. Bot. 2022(1)-e03348: 2 (2021)
species :
Ceratozamia becerrae Pérez-Farr., Vovides & Schutzman; Bot. J. Linn. Soc. 146(1): 124 (123-128), figs.
Distribution: Mexico (Tabasco, Chiapas)
79 MEX
species :
Ceratozamia brevifrons Miq.; Tijdschr. Wis- Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 1: 41 (1848)
Distribution: Mexico (Veracruz)
79 MEX
species :
Ceratozamia chamberlainii Mart.-Domínguez, Nic.-Mor. & D.W.Stev.; Phytotaxa 317(1): 22 (2017)
Distribution: Mexico (San Luis Potosi, Queretaro, Hidalgo)
79 MEX
species :
Ceratozamia chimalapensis Pérez-Farr. & Vovides; Bot. J. Linn. Soc. 157(2): 169 (-175), figs. 1-4)
Distribution: Mexico (NE-Oaxaca)
79 MEX
species :
Ceratozamia chinantlensis Pérez-Farr., Ram.-Oviedo & Gut.-Ortega; Taxonomy 4(4): 738 (2024)
Distribution: Mexico (Oaxaca)
79 MEX
species :
Ceratozamia decumbens Vovides, Avendaño, Pérez-Farr. & Gonz.-Astorga; Novon 18(1): 110 (2008)
Distribution: Mexico (C-Veracruz)
79 MEX
species :
Ceratozamia delucana Vázq.Torres, A.Moretti & Carv.-Hern.; Delpinoa 50-51: 129 (2013)
Distribution: Mexico (Veracruz, Puebla)
79 MEX
species :
Ceratozamia euryphyllidia Vázq.Torres, Sabato & D.W.Stev.; Brittonia 38(1): 17 (1986)
Distribution: SC-Mexico (Veracruz, Oaxaca)
79 MEX
Remarks: miqueliana agg.
Conservation Status: Critically endangered
species :
Ceratozamia fuscoviridis W.Bull; Cat. New Beautiful Rare Pl. [W.Bull] 154: 4 (1879)
Distribution: Mexico (Hidalgo, Veracruz)
79 MEX
Synonyms: hidden
= Ceratozamia mexicana f. fuscoviridis (W.Bull) J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 132 (1932)
species :
Ceratozamia gigantea Mart.-Domínguez, Nic.-Mor., D.W.Stev. & Gonz.-Aguilar; Kew Bull. 79: 546 (2024)
Distribution: Mexico (Tabasco)
79 MEX
species :
Ceratozamia guatemalensis Pérez-Farr., Gut.Ortega & M.L.Quezada; Phytotaxa 668(1): 71 (2024)
Distribution: Guatemala
80 GUA
species :
Ceratozamia hildae G.P.Landry & M.C.Wilson; Brittonia 31(3): 422 (1979)
Distribution: Mexico (Queretaro, San Luis Potosi)
79 MEX
Conservation Status: Critically endangered
species :
Ceratozamia hondurensis J.L.Haynes, Whitelock, Schutzman & R.S.Adams; Cycad Newslett. 31(2-3): 18 (16-21), figs., map
Distribution: Honduras
80 HON
species :
Ceratozamia huastecorum Avendaño, Vovides & Cast.-Campos; Bot. J. Linn. Soc. 141(3): 395 (2003)
Distribution: Mexico (Veracruz)
79 MEX
species :
Ceratozamia kuesteriana Regel; Bull. Soc. Imp. Naturalistes Moscou 30(1): 187 (1857)
Distribution: Mexico (Tamaulipas)
79 MEX
Conservation Status: Endangered (IUCN V)
Synonyms: hidden
= Ceratozamia angustifolia Lindl.; Ill. Hort. 28: 32 (1881), nom. nud.
= Ceratozamia eriolepis J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 132 (1932)
species :
Ceratozamia latifolia Miq.; Tijdschr. Wis- Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 1(4): 206 (1848)
Name: Costilla de leon, palma imperial, piña del monte (ES), Mexican horncone (EN)
Distribution: Mexico (San Luis Potosi)
79 MEX
Conservation Status: Rare (IUCN V)
Synonyms: hidden
= Ceratozamia karsteniana Dyer; Biol. Centr.-Amer., Bot. 3: 192 (1884)
= Ceratozamia mexicana var. latifolia (Miq.) J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 131 (1932)
= Ceratozamia microstrobila Vovides & J.D.Rees; Madroño 30(1): 39 (1983)
= Eriozamia latifolia J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 131 (1932), nomen
= Zamia latifolia Karw. ex J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 131 (1932), nomen
species :
Ceratozamia leptoceras Mart.-Domínguez, Nic.-Mor., D.W.Stev. & Lorea-Hern.; PhytoKeys 156: 13, figs. 1, 3-8 (2020)
Distribution: Mexico (Guerrero)
79 MEX
species :
Ceratozamia matudae Lundell; Lloydia 2: 75 (1939)
Distribution: Mexico (SW-Chiapas)
79 MEX
Conservation Status: Rare (IUCN R)
species :
Ceratozamia mexicana Brongn.; Ann. Sci. Nat., Bot., Sér. 3, 5: 8 (1846)
Distribution: Mexico (Veracruz)
79 MEX
Synonyms: hidden
= Ceratozamia ensiformis J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 130 (1932), pro syn.
= Ceratozamia fusca J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 132 (1932), pro syn.
= Ceratozamia intermedia Miq.; Tijdschr. Wis- Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 1: 40 (1848)
= Ceratozamia longifolia Miq.; Tijdschr. Wis- Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 1: 40 (1848)
= Ceratozamia longifolia var. minor Miq.; Tijdschr. Wis- Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 2: 290 (1849)
= Ceratozamia longipinnata J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 130 (1932), pro syn.
= Ceratozamia mexicana var. longifolia (Miq.) Dyer; Biol. Centr.-Amer., Bot. 3: 193 (1884)
= Ceratozamia ottonis J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 130 (1932), pro syn.
= Ceratozamia purpurea Matte; Rech. App. Lib. Lign. Cycad.: 125 (1904)
= Dipsacozamia mexicana (Brongn.) Liebm. ex Dyer; Biol. Centr.-Amer., Bot. 3: 192 (1884)
= Macrozamia breviformis J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 130 (1932), pro syn.
= Zamia amplifrons J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 130, 142 (1932), pro syn.
= Zamia atropurpurea J.Parm.; Cat. Coll. Pl. Rar. 7 (1853)
= Zamia atropurpurea J.Parm. ex Miq.; Prodr. Syst. Cycad. 23 (1861)
= Zamia macrophylla J.Parm. ex Miq.; Prodr. Syst. Cycad., 23 (1861), nomen, in syn.
= Zamia mexicana (Brongn.) G.Don; Encycl. Pl. [Loudon], new ed., Suppl. 2: 1526 (1855)
species :
Ceratozamia miqueliana H.Wendl.; Index Palm. 68 (1854)
Distribution: Mexico (Veracruz, Chiapas, Tabasco)
79 MEX
Remarks: miqueliana agg.
Conservation Status: Rare
Synonyms: hidden
= Ceratozamia ghiesbrechtii Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 4: 299 (1876)
= Ceratozamia mexicana var. miqueliana (H.Wendl.) J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 131 (1932)
species :
Ceratozamia mirandae Vovides, Pérez-Farr. & Iglesias; Bot. J. Linn. Soc. 137(1): 81 (2001)
Distribution: Mexico (Chiapas)
79 MEX
species :
Ceratozamia mixeorum Chemnick, T.J.Greg. & Salas-Mor.; Phytologia 83: 47-52 (1997 publ. 1998)
Distribution: Mexico (Oaxaca)
79 MEX
Conservation Status: Endangered
species :
Ceratozamia morettii Vázq.Torres & Vovides; Novon 8, 87-90 (1998)
Distribution: Mexico (Veracruz)
79 MEX
Conservation Status: Endangered
species :
Ceratozamia norstogii D.W.Stev.; Brittonia 34(2): 181 (1982)
Distribution: Mexico (Chiapas, Oaxaca)
79 MEX
Conservation Status: Endangered
species :
Ceratozamia oliversacksii D.W.Stev., Mart.-Domínguez & Nic.-Mor.; Kew Bull. 77: 212 (2022)
Distribution: Mexico (Oaxaca)
79 MEX
species :
Ceratozamia osbornei D.W.Stev., Mart.-Domínguez & Nic.-Mor.; PhytoKeys 208: 68, figs. 5-6, 22 (2022)
Distribution: Belize (Toledo)
80 BLZ
species :
Ceratozamia reesii Vovides, Pérez-Farr. & Gut.Ortega; Phytotaxa 575(3): 241 (2022)
Distribution: Mexico (San Luis Potosí)
79 MEX
species :
Ceratozamia robusta Miq.; Tijdschr. Wis- Natuurk. Wetensch. Eerste Kl. Kon. Ned. Inst. Wetensch. 1: 42 (1848)
Distribution: Mexico (Chiapas); Guatemala
79 MEX 80 GUA
Synonyms: hidden
= Ceratozamia mexicana var. robusta (Miq.) Dyer; Biol. Centr.-Amer., Bot. 3: 193 (1884)
species :
Ceratozamia rosea Pérez-Farr., Gut.Ortega & Vovides; Phytotaxa 595(1): 79 (2023)
Distribution: Mexico (Chiapas)
79 MEX
species :
Ceratozamia sabatoi Vovides, Vázq.Torres, Schutzman & Iglesias; Novon 3(4): 502 (1993)
Distribution: Mexico (Hidalgo, Queretaro)
79 MEX
species :
Ceratozamia sancheziae Pérez-Farr., Gut.Ortega & Vovides; Phytotaxa 500(3): 209 (2021)
Distribution: Mexico (Chiapas)
79 MEX
species :
Ceratozamia santillanii Pérez-Farr. & Vovides; Syst. Biodivers. 7: 433. 2009
Distribution: Mexico (Chiapas)
79 MEX
Remarks: miqueliana agg.
species :
Ceratozamia schiblii Pérez-Farr. & Gut.Ortega; Taxonomy 2(3): 330 (2022)
Distribution: Mexico (Oaxaca)
79 MEX
species :
Ceratozamia subroseophylla Mart.-Domínguez & Nic.-Mor.; Phytotaxa 268(1): 35 (2016)
Distribution: Mexico (SE-Veracruz, Tabasco)
79 MEX
Synonyms: hidden
= Ceratozamia dominguezii Pérez-Farr. & Gut.Ortega; Taxonomy 1(4): 353 (2021)
species :
Ceratozamia tenuis (Dyer) D.W.Stev. & Vovides; Bot. Sci. 94(2): 425 (2016)
Distribution: Mexico (Veracruz)
79 MEX
Synonyms: hidden
= Ceratozamia mexicana f. tenuis (Dyer) J.Schust.; Pflanzenr. [Engler] IV.1, Heft 99 (Cycad.): 132 (1932)
= Ceratozamia mexicana var. tenuis Dyer; Biol. Centr.-Amer., Bot. 3: 193 (1884)
species :
Ceratozamia totonacorum Mart.-Domínguez & Nic.-Mor.; Brittonia 69(4): 518 (2017)
Distribution: Mexico (Hidalgo, Puebla, Veracruz)
79 MEX
species :
Ceratozamia vovidesii Pérez-Farr. & Iglesias; Bot. J. Linn. Soc. 153(4): 394 (393-400), figs. 1-6 (2007)
Distribution: Mexico (Chiapas); Guatemala
79 MEX 80 GUA
species :
Ceratozamia whitelockiana Chemnick & T.J.Greg.; Phytologia 79(1): 51 (1995 publ. 1996)
Distribution: Mexico (Oaxaca)
79 MEX
Conservation Status: Endangered
species :
Ceratozamia zaragozae Medellín; Brittonia 15: 175 (1963)
Distribution: Mexico (San Luis Potosi)
79 MEX
Conservation Status: Endangered
species :
Ceratozamia zoquorum Pérez-Farr., Vovides & Iglesias; Bot. J. Linn. Soc. 137(1): 77 (2001)
Distribution: Mexico (Chiapas)
79 MEX
Remarks: miqueliana agg.
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.05 (May 20th, 2026)

Covering 359,646 accepted species,
8,890 hybrids, 47,679 accepted infraspecific taxa and 1,012,517 synonyms,
in total 1,428,732* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life