492.0000
Family
Rubiaceae Juss.; Gen. Pl. [Jussieu] 196 (1789), nom. cons.
589 genera, ca. 14.200 spp. mostly tropics, comparably few temperate. Phylogeny: Razafimandimbison & Rydin (2024)
Species listed in database: 14370; hybrids: 47
= Aparinaceae Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 2: 3, 38 (1813-1829)
= Asperulaceae Cham. ex Spenn.; Handb. Angew. Bot. 2: 493 (1835)
= Catesbaeaceae Martinov; Tekhno-Bot. Slovar 111 (1820)
= Cephalanthaceae Raf.; Ann. Gén. Sci. Phys. 6: 86 (1820)
= Cinchonaceae Batsch; Tab. Affin. Regni Veg. 234 (1802)
= Coffeaceae Batsch; Tab. Affin. Regni Veg. 233 (1802)
= Coutareaceae Martinov; Tekhno-Bot. Slovar 168 (1820)
= Dialypetalanthaceae Rizzini & Occhioni; Lilloa 17: 253 (1948), nom. cons.
= Galiaceae Lindl.; Intr. Nat. Syst. Bot., ed. 2: 249 (1836)
= Gardeniaceae Dumort.; Anal. Fam. Pl. 29, 32 (1829)
= Guettardaceae Batsch; Tab. Affin. Regni Veg. 235 (1802)
= Hameliaceae Mart.; Consp. Regn. Veg. [Martius] 31 (1835)
= Hedyotidaceae Dumort.; Commentat. Bot. [Dumortier] 57 (1822)
= Henriqueziaceae Bremek.; Acta Bot. Neerl. 6 (Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 141): 371 (1957)
= Houstoniaceae Raf.; Good Book 21 (1840)
= Hydrophylacaceae Martinov; Tekhno-Bot. Slovar 318 (1820)
= Lippayaceae Meisn.; Pl. Vasc. Gen. [Meisner], Tab. Diagn. 156, Comm. 112 (1838)
= Lygodisodeaceae Bartl.; Ord. Nat. Pl. 123, 207 (1830)
= Naucleaceae Wernham; New Phytol. 11: 225 (1912)
= Nonateliaceae Martinov; Tekhno-Bot. Slovar 420 (1820)
= Operculariaceae Juss. ex Perleb; Vers. Arzneikr. Pfl. 207 (1818)
= Pagamaeaceae Martinov; Tekhno-Bot. Slovar 447 (1820)
= Psychotriaceae F.Rudolphi; Syst. Orb. Veg. 49 (1830)
= Randiaceae Martinov; Tekhno-Bot. Slovar 534 (1820)
= Sabiceaceae Martinov; Tekhno-Bot. Slovar 555 (1820)
= Spermacoceaceae Bercht. & J.Presl; Přir. Rostlin 256 (1820)
= Theligonaceae Dumort.; Anal. Fam. Pl. 15, 17 (1829), nom. cons.
492.3016
Genus
Hedyotis L.; Sp. Pl. [Linnaeus] 1: 101 (1753) s. str.
Distribution: Old World tropics, S-China, W-Pacific. Former Hedyotideae.
Species listed in database: 201
= Allaeophania Thwaites; Enum. Pl. Zeyl. [Thwaites] 147 (1859)
= Anistelma Raf.; Autik. Bot. 69 (1840)
= Diplophragma (Wight & Arn.) Meisn.; Pl. Vasc. Gen. [Meisner] 160 (1839)
= Macrandria (Wight & Arn.) Meisn.; Pl. Vasc. Gen. [Meisner] 5(Tab. Diagn.): 160 (1838)
= Metabolos Blume; Bijdr. Fl. Ned. Ind. 990 (1826)
= Metabolus A.Rich.; Mém. Rubiac. 157 (1830), orth. var.
= Pleiocraterium Bremek.; Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 56: 440 (1939)
= Sclerococcus Bartl.; Ord. Nat. Pl. 210 (1830), nom. inval.
= Symphyllarion Gagnep.; Bull. Soc. Bot. France 95: 32 (1948)
species :
Hedyotis acutangula
Champ. ex Benth.; Hooker´s J. Bot. Kew Gard. Misc. 4: 171 (1852)
Distribution: China (Guangdong, Hainan, Fujian, Hongkong); Andaman Isl.; Myanmar (Taninthayi); N-Thailand; Laos; Cambodia; Vietnam
36 CHI 41 AND CBD LAO MYA THA VIE
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis helferi Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 53 (1880)
= Hedyotis lawsoniae (DC.) Benth. ex Hook.; London J. Bot. 1: 486 (1842), nom. illeg.
= Oldenlandia acutangula (Champ. ex Benth.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Oldenlandia helferi (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Wendlandia lawsoniae DC.; Prodr. [A.P. de Candolle] 4: 413 (1830)
species :
Hedyotis albonervia
Bedd.; Icon. Pl. Ind. Or. [Beddome] 1 (1868)
Distribution: India (Tamil Nadu, Kerala)
40 IND
Synonyms: hidden
= Oldenlandia albonervia (Bedd.) Gamble; Fl. Madras 599 (1921)
species :
Hedyotis articularis
R.Br. ex G.Don; Gen. Hist. 3: 527 (1834), descr.
subspecies :
subsp. articularis
Distribution: India (Karnataka, Kerala, Tamil Nadu)
40 IND
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia articularis (R.Br. ex G.Don) Gamble; Fl. Madras 597 (1921)
subspecies :
subsp. santapaui
(Shetty & Vivek.) Deb & Ratna Dutta; Taxon 32(2): 284 (1983)
Distribution: India (Kerala)
40 IND
Synonyms: hidden
= Hedyotis santapaui Shetty & Vivek.; Bull. Bot. Surv. India 11(3-4): 447 (1969 publ. 1972)
species :
Hedyotis assimilis
Tutcher; Rep. Bot. Forest. Dept. Hongkong 32 (1914 publ. 1915)
Distribution: China (Guangdong)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia assimilis (Tutcher) Chun; Sunyatsenia 1: 310 (1934)
species :
Hedyotis atropurpurea
Merr.; Philipp. J. Sci. 17: 426 (1920 publ. 1921)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis austroguangxiensis
M.D.Yuan & R.J.Wang; PhytoKeys 263: 142 (2025)
Distribution: China (Guangxi)
36 CHI
species :
Hedyotis austrosinica
L.Wu & L.H.Yang; Nordic J. Bot. 36(4)-e01499: 2 (2018)
Distribution: China (Guangdong, Guangxi)
36 CHI
species :
Hedyotis bahaii
J.F.Maxwell; Nat. Hist. Bull. Siam Soc. 50(1): 3 (2002)
Distribution: E-Thailand; Laos; ?Cambodia
41 LAO THA
species :
Hedyotis bambusetorum
Merr.; Philipp. J. Sci. 20: 469 (1922)
Distribution: Palawan
42 PLW
species :
Hedyotis baotingensis
W.C.Ko; J. S. China Agric. Univ. 16(4): 43 (1995)
Distribution: China (Hainan)
36 CHI
Synonyms: hidden
= Hedyotis dinganensis Qing L.Wang, L.X.Yuan & Z.N.Wang; Phytotaxa 399: 44 (2019)
species :
Hedyotis barberi
(Gamble) A.N.Henry & Subraman.; Proc. Indian Acad. Sci., B 76(1): 29 (1972)
Distribution: India (Tamil Nadu)
40 IND
Synonyms: hidden
= Oldenlandia barberi Gamble; Bull. Misc. Inform. Kew 1920: 68 (1920)
species :
Hedyotis beddomei
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 52 (1880)
Distribution: India (Kerala)
40 IND
Synonyms: hidden
= Hedyotis capitata Bedd.; Icon. Pl. Ind. Or. [Beddome] 44 (1873), nom. illeg.
= Oldenlandia beddomei (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis benguetensis
(Elmer) Elmer; Leafl. Philipp. Bot. 3: 976 (1911)
Distribution: Sulawesi; Philippines (Luzon)
42 PHI SUL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia benguetensis Elmer; Leafl. Philipp. Bot. 1: 18 (1906)
= Hedyotis capitata Korth.; Ned. Kruidk. Arch. 2(2): 159 (1851), nom. illeg.
species :
Hedyotis bourdillonii
(Gamble) R.S.Rao & Hemadri; Indian Forester 99(6): 378 (1973)
Distribution: India (Kerala)
40 IND
Synonyms: hidden
= Hedyotis bourdillonii (Gamble) R.S.Rao & Hemadri ex N.C.Nair, V.J.Nair & R.Ansari; Bull. Bot. Surv. India 22(1-4): 205 (1983)
= Hedyotis silent-valleyensis Vajr., Rathakr. & Bhargavan; J. Bombay Nat. Hist. Soc. 80(2): 402 (1983 publ. 1984)
= Oldenlandia bourdillonii Gamble; Bull. Misc. Inform. Kew 1919: 404 (1919)
species :
Hedyotis brachyantha
Merr.; Philipp. J. Sci. 17: 427 (1920 publ. 1921)
Distribution: Philippines (Luzon)
42 PHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis bracteosa
Hance; J. Bot. 23: 323 (1885)
Distribution: China (Guangdong, Hongkong)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia bracteosa (Hance) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis buxifolia
Bedd.; Madras J. Lit. Sci., Ser. 1: 49 (1864)
Distribution: S-India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Oldenlandia buxifolia (Bedd.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis cagayanensis
Merr.; Philipp. J. Sci., C 5: 231 (1910)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis camarinensis
Merr.; Philipp. J. Sci. 20: 468 (1922)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis cantoniensis
F.C.How ex W.C.Ko; J. S. China Agric. Univ. 16(4): 42 (1995)
Distribution: China (Guangdong)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis cardiophylla
Quisumb. & Merr.; Philipp. J. Sci. 37: 202 (1928)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis catanduanensis
Merr.; Philipp. J. Sci. 17 (1920 publ. 1921)
Distribution: Philippines (Catanduanes)
42 PHI
species :
Hedyotis cathayana
W.C.Ko; Fl. Hainan. 3: 579 (1974)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis caudata
Merr.; Philipp. J. Sci., C 8: 33 (1913)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis caudatifolia
Merr. & F.P.Metcalf; J. Arnold Arbor. 23: 228 (1942)
Distribution: China (Fujian, Guangdong, Guangxi, Hunan, Jiangxi, Zhejiang)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis hui Diels; Notizbl. Bot. Gart. Berlin-Dahlem 9: 1032 (1926)
species :
Hedyotis cephalophora
(Craib) M.D.Yuan & R.J.Wang; J. Trop. Subtrop. Bot. 30(3): 383 (2022)
Distribution: Thailand
41 THA
Synonyms: hidden
= Hedyotis uncinella var. cephalophora Craib; Fl. Siam 2: 50 (1932)
species :
Hedyotis ceylanica
N.Wikstr. & Neupane; Taxon 62(2): 370 (2013), nom. nov.
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Allaeophania arnottii Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 48 (1880)
= Allaeophania decipiens Thwaites; Enum. Pl. Zeyl. [Thwaites] 147 (1859)
= Allaeophania decipiens var. arnottii (Hook.f.) Trimen; (1894)
= Allaeophania decipiens var. flavescens Thwaites; Enum. Pl. Zeyl. [Thwaites] 147 (1859)
= Hedyotis nodulosa Thwaites; Enum. Pl. Zeyl. [Thwaites] 143, partim (1859)
= Metabolos decipiens (Thwaites) Ridsdale; Blumea 41: 460 (1996)
= Metabolos rugosus f. javanicus Hochr.; Candollea 5: 279 (1934)
= Metabolos rugosus f. zeylanicus Hochr.; Candollea 5: 279 (1934)
= Metabolos rugosus var. decipiens (Thwaites) Hochr.; Candollea 5: 280 (1934)
= Metabolos rugosus var. flavescens (Thwaites) Hochr.; Candollea 5: 279 (1934)
species :
Hedyotis cheniana
R.J.Wang; Novon 18(2): 264 (2008)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis cinereoviridis
Thwaites; Enum. Pl. Zeyl. [Thwaites] 419 (1859)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Oldenlandia cinereoviridis (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis communis
W.C.Ko; Fl. Hainan. 3: 579 (1974)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis consanguinea
Hance; Ann. Sci. Nat., Bot., Sér. 4, 18: 221 (1862)
Distribution: China (Guangdong, Fujian, Hongkong, Macao, Zhejiang, Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia consanguinea (Hance) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis coprosmoides
Trimen; Syst. Cat. Fl. Pl. Ceylon 42 (1885), et Handb. Fl. Ceylon 2: 3(16 (1894)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis corallicola
Fosberg; Kew Bull. 33(1): 136 (1978)
Distribution: Aldabra
29 ALD
Remarks: status uncertain
species :
Hedyotis cornifolia
Kaneh.; Trans. Nat. Hist. Soc. Taiwan 25: 5 (1935)
Distribution: Palau (Babeldaob)
62 PLU
species :
Hedyotis cryptantha
Dunn; Bull. Misc. Inform. Kew 1912: 367 (1912)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia cryptantha (Dunn) Chun; Sunyatsenia 1: 311 (1934)
species :
Hedyotis cushingiae
Fosberg; Allertonia 6(3): 221 (1991)
Distribution: Fed. States of Micronesia
62 MIC
species :
Hedyotis cyanescens
Thwaites; Enum. Pl. Zeyl. [Thwaites] 143 (1859)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Oldenlandia cyanescens (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis dawsoniana
Craib; Bull. Misc. Inform. Kew 1931: 218 (1931)
Distribution: Thailand
41 THA
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis decora
E.T.Geddes; Bull. Misc. Inform. Kew 1931: 219 (1931)
Distribution: peninsular Thailand
41 THA
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis dendroides
Alston; Handb. Fl. Ceylon 6: Suppl., 147 (1931)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis lessertiana var. major Thwaites; Enum. Pl. Zeyl. [Thwaites] 141 (1859)
species :
Hedyotis densa
Craib; Bull. Misc. Inform. Kew 1931: 219 (1931)
Distribution: peninsular Thailand
41 THA
species :
Hedyotis devicolamensis
Deb & Ratna Dutta; J. Bombay Nat. Hist. Soc. 82(3): 619 (1985 publ. 1986)
Distribution: S-India (Tamil Nadu, Kerala)
40 IND
species :
Hedyotis diffusissima
Merr.; Philipp. J. Sci. 17: 429 (1920 publ. 1921)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis divaricata
(Valeton) Hosok.; Trans. Nat. Hist. Soc. Taiwan 24: 204 (1934)
Distribution: Palau (Koror)
62 PLU
Synonyms: hidden
= Hedyotis laciniata var. takamatsui Fosberg; Occas. Pap. Bernice P. Bishop Mus. 15: 215 (1940)
= Hedyotis plurifurcata Hosok.; Trans. Nat. Hist. Soc. Taiwan 24: 203 (1934)
= Oldenlandia divaricata Valeton; Bot. Jahrb. Syst. 63: 292 (1930)
variant :
var. capillothyrsa
Fosberg; Allertonia 6(3): 224 (1991)
Distribution: Fed. States of Micronesia
62 MIC
species :
Hedyotis edanoii
Quisumb. & Merr.; Philipp. J. Sci. 37: 203 (1928)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis effusa
Hance; J. Bot. 17: 11 (1879)
Distribution: China (Guangdong, Guangxi)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia effusa (Hance) Kuntze; Revis. Gen. Pl. 1: 292 (1891), nom. illeg.
species :
Hedyotis elegans
Wall. ex Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 46(2): 133 (1877)
Distribution: Myanmar; N- & C-Thailand
41 MYA THA
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia elegans (Wall. ex Kurz) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Hedyotis sessilifolia E.T.Geddes; Bull. Misc. Inform. Kew 1928: 241 (1928)
species :
Hedyotis equisetiformis
Wernham; J. Nat. Hist. Soc. Siam 4: 134 (1921)
Distribution: Thailand; Vietnam
41 THA VIE
species :
Hedyotis eualata
(Gamble) A.N.Henry & Subraman.; Proc. Indian Acad. Sci., B 76(1): 28 (1972)
Distribution: India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Oldenlandia eualata Gamble; Bull. Misc. Inform. Kew 1919: 404 (1919)
variant :
var. agastyamalayana
A.N.Henry & Subraman.; Proc. Indian Acad. Sci., B 76(1): 28 (1972)
Distribution: India (Tamil Nadu)
40 IND
species :
Hedyotis eucapitata
Merr.; Philipp. J. Sci., C 2: 303 (1907)
Distribution: Philippines (Mindoro)
42 PHI
species :
Hedyotis evania
Thwaites; Enum. Pl. Zeyl. [Thwaites] 140, 419 (1859)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Oldenlandia evania (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis exserta
Merr.; Lingnan Sci. J. 13: 72 (1934)
Distribution: China (Hainan); Vietnam
36 CHI 41 VIE
species :
Hedyotis fissistipula
Merr.; J. Malayan Branch Roy. Asiat. Soc. 1: 38 (1923)
Distribution: Borneo
42 BOR
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis flavescens
Thwaites; Enum. Pl. Zeyl. [Thwaites] 141 (1859)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis lessertiana var. flavescens (Thwaites) Thwaites; (1864)
species :
Hedyotis flexuosa
Ridl.; J. Straits Branch Roy. Asiat. Soc. 35: 15 (1901)
Distribution: peninsular Malaysia (Kedah, Melaka, Johor)
42 MLY
species :
Hedyotis fruticosa
L.; Sp. Pl. [Linnaeus] 1: 101 (1753)
Distribution: S-India (Kerala); Sri Lanka; Bangladesh; Myanmar; Laos
40 BAN IND SRL 41 LAO MYA
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis corymbosa Spreng.; Pl. Min. Cogn. Pug. 2: 27 (1815), nom. illeg.
= Oldenlandia fruticosa (L.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis fruticulosa
(Volkens) Merr.; Philipp. J. Sci. 15: 544 (1919)
Distribution: Palau (Babeldaob, Ulebsechel, Yap); Guam
62 GUM PLU
Synonyms: hidden
= Oldenlandia fruticulosa Volkens; Bot. Jahrb. Syst. 31: 475 (1901)
variant :
var. atroglomerata
Fosberg; Allertonia 6(3): 228 (1991)
Distribution: Fed. States of Micronesia
62 MIC
variant :
var. yapensis
(Kaneh.) Fosberg; Allertonia 6(3): 228 (1991)
Distribution: Fed. States of Micronesia (Yap)
62 MIC
Synonyms: hidden
= Hedyotis yapensis Kaneh.; Trans. Nat. Hist. Soc. Taiwan 25: 5 (1935)
species :
Hedyotis fumata
Alston; Handb. Fl. Ceylon 6: Suppl., 148 (1931)
Distribution: Sri Lanka
40 SRL
species :
Hedyotis gamblei
A.N.Henry & Subr.; Proc. Indian Acad. Sci., B 76: 26 (1972)
Distribution: S-India (Tamil Nadu)
40 IND
species :
Hedyotis gardneri
Thwaites; Enum. Pl. Zeyl. [Thwaites] 142 (1859)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia gardneri (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis garrettii
Craib; Bull. Misc. Inform. Kew 1931: 220 (1931)
Distribution: N- & NE-Thailand
41 THA
species :
Hedyotis gartmorensis
Ridsdale; Blumea 41: 455 (1996)
Distribution: Sri Lanka
40 SRL
species :
Hedyotis gentianifolia
(Bremek.) N.Wikstr. & Neupane; Taxon 62(2): 370 (2013)
Distribution: Sumatera
42 SUM
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Pleiocraterium gentianifolium Bremek.; Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 56: 441 (1939), et in Recueil Trav. Bot. Néerl. 36: 441 (1939 publ. 1940)
species :
Hedyotis globiceps
Ridl.; Fl. Malay Penins. 2: 47 (1923)
Distribution: peninsular Malaysia (Langkawi Isl.)
42 MLY
species :
Hedyotis griffithii
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 58 (1880)
Distribution: Bhutan
40 BHU
Synonyms: hidden
= Oldenlandia griffithii (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis hainanensis
(Chun) W.C.Ko; Fl. Hainan. 3: 580 (1974)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia hainanensis Chun; Sunyatsenia 1: 311 (1934)
= Oldenlandia oligantha Chun; Sunyatsenia 1: 313 (1934)
species :
Hedyotis hamiguitanensis
Santor, D.D.B.Santiago & Alejandro; Webbia 76(2): 196 (2021)
Distribution: Philippines
42 PHI
species :
Hedyotis havilandii
King; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 72: 164 (1903)
Distribution: Vietnam; peninsular Thailand; peninsular Malaysia (Penang, Perak, Pahang)
41 THA VIE 42 MLY
Synonyms: hidden
= Exallage havilandii (King) Bremek.; Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2) (Afr. Sp. Oldenlandia): 142 (1952)
= Hedyotis betongensis Geddes; Bull. Misc. Inform. Kew 1931: 216 (1931)
= Oldenlandia havilandii (King) Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 133 (1922)
species :
Hedyotis hirsutissima
Bedd.; Madras J. Lit. Sci., Ser. 3, 1: 49 (1864)
Distribution: +S-India (+Tamil Nadu)
40 IND
Synonyms: hidden
= Oldenlandia hirsutissima (Bedd.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis howii
G.B.Jiang & R.J.Wang; Nordic J. Bot. 39(6)-e02934: 4 (2021)
Distribution: China (Hainan)
36 CHI
species :
Hedyotis humilis
Merr.; Philipp. J. Sci., C 8: 33 (1913)
Distribution: Philippines (Camiguin, Mindanao)
42 PHI
species :
Hedyotis inamoena
Thwaites; Enum. Pl. Zeyl. [Thwaites] 143, partim. (1859)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Oldenlandia inamoena (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis indirae
K.M.P.Kumar & Aiswarya; Webbia 74(2): 276 (2019)
Distribution: India (Kerala)
40 IND
species :
Hedyotis interrupta
G.B.Jiang & R.J.Wang; Nordic J. Bot. 39(6)-e02934: 5 (2021)
Distribution: China (Hainan)
36 CHI
species :
Hedyotis jinghongensis
M.D.Yuan & R.J.Wang; PhytoKeys 263: 137 (2025)
Distribution: China (Yunnan)
36 CHI
species :
Hedyotis kamputensis
(Pit.) Wangwasit & Chantar.; Thai Forest Bull., Bot. 30: 69 (2002)
Distribution: SE-Thailand; Cambodia; Vietnam
41 CBD THA VIE
Synonyms: hidden
= Hedyotis diversifolia Geddes; Bull. Misc. Inform. Kew 1931: 220 (1931)
= Oldenlandia kamputensis Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 117 (1922)
species :
Hedyotis konhanungensis
B.H.Quang, T.A.Le, K.S.Nguyen & Neupane; PhytoKeys 221: 79 (2023)
Distribution: Vietnam
41 VIE
species :
Hedyotis korrorensis
(Valeton) Hosok.; Trans. Nat. Hist. Soc. Taiwan 24: 204 (1934)
Distribution: Fed. States of Micronesia; Palau (Babeldaob, Koror)
62 MIC PLU
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis aimiriikensis Kaneh.; Trans. Nat. Hist. Soc. Taiwan 25: 4 (1935)
= Hedyotis korrorensis var. hosokawana Fosberg; Allertonia 6: 230 (1991)
= Hedyotis korrorensis var. mollis (Valeton) Hosok.; Trans. Nat. Hist. Soc. Taiwan 24: 132 (1934)
= Hedyotis korrorensis var. subglomerata Fosberg; Allertonia 6: 231 (1991)
= Oldenlandia korrorensis Valeton; Bot. Jahrb. Syst. 63: 294 (1930)
= Oldenlandia korrorensis var. mollis Valeton; Bot. Jahrb. Syst. 63: 295 (1930)
species :
Hedyotis kottangathattiensis
M.B.Viswan. & Manik.; Edinburgh J. Bot. 65(3): 388 (2008)
Distribution: India (Andhra Pradesh, Gujarat, Karnataka, Madhya Pradesh, Maharashtra, Odisha, Rajasthan, Tamil Nadu)
40 IND
species :
Hedyotis kurzii
Merr.; Pap. Michigan Acad. Sci. 19: 195 (1933 publ. 1934)
Distribution: Nicobar Isl. (Central Nicobar Isl.); Myanmar (Taninthayi); Thailand; Vietnam; peninsular Malaysia; Borneo
41 MYA NCB THA VIE 42 BOR MLY
Synonyms: hidden
= Hedyotis galioides Wall.; Numer. List [Wallich] no. 866 (1828), nomen
= Hedyotis nicobarensis W.H.Lewis; Ann. Missouri Bot. Gard. 53: 109 (1966), nom. nov. superfl.
= Hedyotis wallichii Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 45: 136 (1876), nom. illeg.
= Oldenlandia rosea Ridl.; J. Straits Branch Roy. Asiat. Soc. 59: 110 (1911)
= Oldenlandia wallichii Craib; Bull. Misc. Inform. Kew 1911: 388 (1911)
species :
Hedyotis laciniata
Kaneh.; Trans. Nat. Hist. Soc. Taiwan 25: 6 (1935)
Distribution: Northern Marianas (Alamagan, Anatahan, Sarigan, Saipan, Rota); Guam
62 GUM MRN
Synonyms: hidden
= Hedyotis alamaganensis Hosok.; J. Soc. Trop. Agric. 6: 669, nomen, et in Trans. Nat. Hist. Soc. Formosa, 1935 25: 37 (1934), descr.
= Hedyotis sariganensis Hosok.; J. Soc. Trop. Agric. 6: 669, nomen, et in Trans. Nat. Hist. Soc. Formosa, 1935 25: 36 (1934), descr.
species :
Hedyotis lallanii
R.Kr.Singh; J. Biodivers. Conservation 8(4): 64 (2024), nom. nov.
Distribution: Philippines
42 PHI
Synonyms: hidden
= Hedyotis microphylla Merr.; Philipp. J. Sci. 1(Suppl.): 236 (1906), nom. illeg.
species :
Hedyotis lancea
Thunb. ex Maxim.; Bull. Acad. Imp. Sci. St.-Pétersbourg, Sér. 3, 29: 161 (1883)
Distribution: China (Guangdong, Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis lancelimba var. epedunculosa Merr.; Mitt. Inst. Allg. Bot. Hamburg 7: 273 (1937)
= Hedyotis lancelimba var. scaberulifolia Merr.; Mitt. Inst. Allg. Bot. Hamburg 7: 273 (1937)
= Oldenlandia lancea (Thunb. ex Maxim.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis laotica
(Pit.) Wangwasit & Chantar.; Phytotaxa 206(1): 51 (2015)
Distribution: N- & NE-Thailand; Laos
41 LAO THA
Synonyms: hidden
= Oldenlandia laotica Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 123 (1922)
species :
Hedyotis lawsoniae
Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 407 (1834)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia lawsoniae (Wight & Arn.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis leschenaultiana
DC.; Prodr. [A.P. de Candolle] 4: 422 (1830)
Distribution: India (Tamil Nadu)
40 IND
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis anamalayana (Gamble) R.S.Rao & Hemadri; Indian Forester 99(6): 378 (1973)
= Hedyotis sisaparensis Gage; J. Proc. Asiat. Soc. Bengal 1: 244 (1906)
= Knoxia wightiana Schltdl. ex Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 51 (1880), non Wall. ex Wight & Arn. 1834
= Oldenlandia anamalayana Gamble; Bull. Misc. Inform. Kew 1920: 67 (1920)
= Oldenlandia sisaparensis (Gage) Gamble; Fl. Madras 599 (1921)
variant :
var. wynaadensis
(Gamble) Deb & Ratna Dutta; Taxon 34(2): 296 (1985)
Distribution: India (Tamil Nadu)
40 IND
Synonyms: hidden
= Hedyotis wynaadensis (Gamble) R.S.Rao & Hemadri; Indian Forester 99(6): 379 (1973)
= Oldenlandia wynaadensis Gamble; Bull. Misc. Inform. Kew 1919: 405 (1919)
species :
Hedyotis lessertiana
Arn.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18(1): 339 (1836)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis confertiflora (Thwaites) Alston; Handb. Fl. Ceylon 6: Suppl., 147 (1931)
= Hedyotis lessertiana var. confertiflora Thwaites; Enum. Pl. Zeyl. [Thwaites] 141 (1859)
= Hedyotis lessertiana var. pilosa Thwaites; Enum. Pl. Zeyl. [Thwaites] 141 (1859)
= Hedyotis neolessertiana Ridsdale; Blumea 41: 456 (1996), nom. nov. superfl.
= Oldenlandia lessertiana (Arn.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis leuserensis
N.Wikstr. & Neupane; Taxon 62(2): 370 (2013), nom. nov.
Distribution: Sumatera
42 SUM
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Pleiocraterium sumatranum Bremek.; Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 56: 440 (1939), et in Recueil Trav. Bot. Néerl. 36: 440 (1939 publ. 1940)
species :
Hedyotis linearifolia
(Pit.) M.D.Yuan & R.J.Wang; J. Trop. Subtrop. Bot. 30(3): 385 (2022)
Distribution: Laos
41 LAO
Synonyms: hidden
= Oldenlandia multiglomerulata var. linearifolia Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 131 (1922)
species :
Hedyotis loganioides
Benth.; Fl. Hongk. 149 (1861)
Distribution: China (Hongkong, Guangdong)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis bodinieri H.Lév.; Repert. Spec. Nov. Regni Veg. 11: 64 (1912)
= Oldenlandia bodinieri (H.Lév.) Chun; Sunyatsenia 1: 310 (1934)
= Oldenlandia loganioides (Benth.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis longiexserta
Merr. & F.P.Metcalf; J. Arnold Arbor. 23: 229 (1942)
Distribution: China (Guangxi)
36 CHI
species :
Hedyotis longipedunculata
Merr.; Philipp. J. Sci. 17: 430 (1920 publ. 1921)
Distribution: Philippines (Catanduanes)
42 PHI
species :
Hedyotis longipetala
Merr.; J. Arnold Arbor. 8: 18 (1927)
Distribution: China (Guangdong, Fujian)
36 CHI
Synonyms: hidden
= Oldenlandia longipetala (Merr.) Chun; Sunyatsenia 1: 312 (1934)
species :
Hedyotis longiramulis
Y.D.Xu & R.J.Wang; PhytoKeys 230: 276 (2023)
Distribution: China (Guangdong)
36 CHI
species :
Hedyotis luzoniensis
Merr.; Philipp. J. Sci., C 12: 160 (1917)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis lychnidifolia
Craib; Bull. Misc. Inform. Kew 1931: 276 (1931)
Distribution: peninsular Thailand
41 THA
Remarks: known only from type locality
species :
Hedyotis macgregorii
Merr.; Philipp. J. Sci., C 5: 160 (1910)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis macraei
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 50 (1880)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Oldenlandia macraei (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis macrophylla
Wall. ex Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 408 (1834)
Distribution: Laos; Cambodia; Vietnam; NE- & peninsular Thailand; peninsular Malaysia (Ulu Kerling, Perak); Jawa
41 CBD LAO THA VIE 42 JAW MLY
Synonyms: hidden
= Exallage kunstleri (King) Bremek.; Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2) (Afr. Sp. Oldenlandia): 142 (1952)
= Exallage latifolia (Blume) Bremek.; Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2) (Afr. Sp. Oldenlandia): 142 (1952)
= Hedyotis fraterna (Pierre ex Pit.) P.H.Hô; Câyco Vietnam (Illustr. Fl. Vietnam) 3(1): 138 (1993)
= Hedyotis kunstleri King; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 72: 162 (1904)
= Hedyotis latifolia (Blume) Reinw. ex de Vriese; Pl. Ind. Bat. Orient. 159 (1856), nom. superfl.
= Hedyotis latifolia var. ovata Miq.; Fl. Ned. Ind. 2: 178 (1857)
= Hedyotis reinwardtii Bakh.f.; Blumea 12: 63 (1963)
= Metabolos latifolius Blume; Bijdr. Fl. Ned. Ind. 991 (1826)
= Oldenlandia fraterna Pierre ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 128 (1922)
= Oldenlandia latifolia (Blume) Kuntze; Revis. Gen. Pl. 1: 292 (1891), nom. superfl.
species :
Hedyotis macrostegia
Stapf; Trans. Linn. Soc. London, Bot., Ser. 2, 4: 170 (1894)
Distribution: Borneo
42 BOR
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis maingayi
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 49 (1880)
Distribution: peninsular Malaysia (Gunung Ledang, Melaka)
42 MLY
Synonyms: hidden
= Oldenlandia maingayi (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis manasensis
D.Baro, Bawri & Borthakur; Natl. Acad. Sci. Lett. [1] (2022)
Distribution: NE-India (Assam)
40 IND
species :
Hedyotis marginata
Thwaites ex Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 53 (1880), pro syn.
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis lessertiana var. marginata Thwaites ex Trimen; (1894)
= Hedyotis marginata (Thwaites ex Trimen) Alston; Handb. Fl. Ceylon 6: 147 (1931)
species :
Hedyotis matthewii
Dunn; J. Bot. 47: 376 (1909)
Distribution: China (Guangdong, Guangxi, Fujian, Jiangxi, Hunan)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis acuminatissima Merr.; Philipp. J. Sci., C 13: 159 (1918)
= Hedyotis mellii Tutcher; Rep. Bot. Forest. Dept. Hongkong 32 (1914 publ. 1915)
= Hedyotis wulsinii Merr.; J. Arnold Arbor. 8: 17 (1927)
= Oldenlandia matthewii (Dunn) Chun; Sunyatsenia 1: 312 (1934)
= Oldenlandia mellii (Tutcher) Chun; Sunyatsenia 1: 313 (1934)
= Oldenlandia speciosa Hand.-Mazz.; Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 58: 93 (1921)
= Oldenlandia wulsinii (Merr.) Chun; Sunyatsenia 1: 314 (1934)
species :
Hedyotis megalantha
Merr.; Philipp. J. Sci., C 9: 143 (1914)
Distribution: Guam
62 GUM
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia megalantha (Merr.) Valeton; Bot. Jahrb. Syst. 63: 298 (1930)
species :
Hedyotis membranacea
Thwaites; Enum. Pl. Zeyl. [Thwaites] 143 (1859)
Distribution: S-India (Kerala); Sri Lanka
40 IND SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis asperifolia Alston; Handb. Fl. Ceylon 6: Suppl., 148 (1931)
= Oldenlandia membranacea (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis merrillii
R.Kr.Singh; J. Biodivers. Conservation 8(4): 64 (2024), nom. nov.
Distribution: Philippines
42 PHI
Remarks: provisionally accepted name, status unconfirmed
Synonyms: hidden
= Hedyotis laxiflora Merr.; Philipp. J. Sci. 17: 430 (1920 publ. 1921), nom. illeg.
species :
Hedyotis mindorensis
Quisumb.; Philipp. J. Sci. 41: 361, t. 25 (1930)
Distribution: Philippines (Mindoro)
42 PHI
Remarks: known only from type collection
species :
Hedyotis minutopuberula
Merr. & F.P.Metcalf; J. Arnold Arbor. 23: 229 (1942)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis montana
Merr.; Philipp. J. Sci., C 2: 304 (1907)
Distribution: Philippines (Mindoro)
42 PHI
species :
Hedyotis nairii
Murug. & V.Balas.; Rheedea 17(1-2): 25 (2007)
Distribution: India (Tamil Nadu)
40 IND
species :
Hedyotis nankunshanensis
R.J.Wang & S.J.Deng; Nordic J. Bot. 30(3): 302 (2012)
Distribution: China (Guangdong)
36 CHI
species :
Hedyotis nanlingensis
R.J.Wang; Phytotaxa 206(1): 37 (2015)
Distribution: China (Guangdong)
36 CHI
species :
Hedyotis nigrescens
Merr.; J. Arnold Arbor. 23: 193 (1942)
Distribution: Vietnam
41 VIE
species :
Hedyotis nodiflora
Wall. ex G.Don; Gen. Hist. 3: 526 (1834)
Distribution: S-China; Myanmar; N- & E-Thailand; Laos; Cambodia; Vietnam
36 CHI 41 CBD LAO MYA THA VIE
species :
Hedyotis nodulosa
Arn.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18(1): 340 (1836)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis axillaris Gardner ex Thwaites; Enum. Pl. Zeyl. [Thwaites] 143 (1859)
= Hedyotis nodulosa var. walkeri (Arn.) Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 55 (1880)
= Hedyotis walkeri Arn.; Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 18: 339 (1836)
= Oldenlandia nodulosa (Arn.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis novoguineensis
Merr. & L.M.Perry; J. Arnold Arbor. 26: 4 (1945)
Distribution: New Guinea (Papua New Guinea); Australia (Queensland)
42 NWG 50 AUS
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis nutans
(Valeton) P.Royen; Alpine Fl. New Guinea 4: 2718 (1983)
Distribution: New Guinea (Irian Jaya)
42 NWG
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia nutans Valeton; Fl. Arfak Mts. [Gibbs] 180 (1917)
species :
Hedyotis obscura
Thwaites; Enum. Pl. Zeyl. [Thwaites] 141 (1859)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia obscura (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis ovalis
(Korth.) Walp.; Ann. Bot. Syst. [Walpers] 2: 769 (1852)
Distribution: Borneo
42 BOR
Synonyms: hidden
= Diplophragma ovalis Korth.; Ned. Kruidk. Arch. 2(2): 150 (1857)
species :
Hedyotis ovata
Thunb. ex Maxim.; Bull. Acad. Imp. Sci. St.-Pétersbourg 29: 161 (1883)
Distribution: China (Hainan); Vietnam
36 CHI 41 VIE
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia ovata (Thunb.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis papafranciscoi
Alejandro; Antoninus J. 1: 75, figs. 1-2 (2015)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis paridifolia
Dunn; Bull. Misc. Inform. Kew 1912: 366 (1912)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia paridiflora (Dunn) Chun; Sunyatsenia 1: 313 (1934)
species :
Hedyotis patens
Ridl.; J. Linn. Soc., Bot. 38: 311 (1908)
Distribution: peninsular Malaysia (Gunung Tahan, Pahang)
42 MLY
species :
Hedyotis phanerophlebia
Merr.; Philipp. J. Sci., C 8: 34 (1913)
Distribution: Philippines (Mindanao)
42 PHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis philippensis
(Willd. ex Spreng.) Merr. ex C.B.Rob.; Philipp. J. Sci., C 6: 222 (1911)
Distribution: Philippines
42 PHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Exallage philippensis (Willd. ex Spreng.) Bremek.; Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2): 142 (1952)
= Spermacoce philippensis Willd. ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 1: 401 (1825)
species :
Hedyotis pilosissima
Merr.; Philipp. J. Sci., C 5: 232 (1910)
Distribution: Philippines (Luzon, Catanduanes, Panay, Samar, Mindanao)
42 PHI
species :
Hedyotis pinaster
Ridl.; Bull. Misc. Inform. Kew 1940: 611 (1940)
Distribution: Borneo
42 BOR
species :
Hedyotis plantaginifolia
Arn.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18(1): 340 (1836)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Hedyotis plantaginea Walp.; Ann. Bot. Syst. [Walpers] 1: 376 (1848)
= Hedyotis verticillaris subsp. plantaginifolia (Arn.) Deb & Ratna Dutta; Rev. Ind. Hedyotis 88 (1985)
= Pleiocraterium plantaginifolium (Arn.) Bremek.; Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 56: 443 (1939), et in Recueil Trav. Bot. Néerl. 36: 443 (1939 publ. 1940)
species :
Hedyotis ponapensis
(Valeton) Kaneh.; Fl. Micron. 463 (1933)
Distribution: Fed. States of Micronesia (Truk, Moen, Pohnpei)
62 MIC
Synonyms: hidden
= Hedyotis ponapensis var. robusta Hosok.; J. Jap. Bot. 13: 616 (1937)
= Oldenlandia ponapensis Valeton; Bot. Jahrb. Syst. 63: 296 (1930)
species :
Hedyotis prostrata
Blume; Cat. Gew. Buitenzorg [Blume] 40 (1823)
Distribution: China (Hainan); Nicobar Isl.; Myanmar (Taninthayi); Thailand; Cambodia; Vietnam; peninsular Malaysia; Borneo; Philippines; Jawa
36 CHI 41 CBD MYA NCB THA VIE 42 BOR JAW MLY PHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Exallage congesta (R.Br. ex G.Don) Bremek.; Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2) (Afr. Sp. Oldenlandia): 142 (1952)
= Hedyotis argentea Wall. ex G.Don; Gen. Hist. 3: 526 (1834)
= Hedyotis congesta R.Br. ex G.Don; Gen. Hist. 3: 526 (1834)
= Hedyotis congesta var. longifolia (Valeton) Merr. & L.M.Perry; J. Arnold Arbor. 25: 4 (1945)
= Hedyotis congesta var. nicobarica King; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 72(2): 161 (1903)
= Hedyotis laevigata (DC.) Miq.; Fl. Ned. Ind. 2: 178 (1857)
= Hedyotis quocensis Pierre ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 134 (1922), pro syn.
= Hedyotis subcarnosa Wall. ex Craib; Fl. Siam 2: 48 (1932)
= Metabolos laevigatus DC.; Prodr. [A.P. de Candolle] 4: 436 (1830)
= Oldenlandia congesta (R.Br. ex G.Don) Kuntze; Revis. Gen. Pl. 1: 292 (1891), not Baker (1877)
= Oldenlandia laevigata (DC.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Oldenlandia prostrata (Blume) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Oldenlandia quocensis Pierre ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 134 (1922)
= Oldenlandia rigida var. longifolia Valeton; Nova Guinea 8: 438 (1911)
species :
Hedyotis protrusa
Stapf; Trans. Linn. Soc. London, Bot., Ser. 2, 4: 169 (1894)
Distribution: Borneo
42 BOR
species :
Hedyotis pruinosa
Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 408 (1834)
Distribution: Pakistan (Hazara, Pakistani Punjab, Rawalpindi, Mirpur); S-India (Kerala); Thailand; Laos; Cambodia
40 IND PAK 41 CBD LAO THA
Synonyms: hidden
= Hedyotis corymbiformis E.T.Geddes; Bull. Misc. Inform. Kew 1931: 217 (1931)
= Hedyotis corymbosa Herb.Madr. ex Wall.; Numer. List [Wallich] no. 6202 (1832)
= Hedyotis lentiginosa Bedd.; Icon. Pl. Ind. Or. [Beddome] 1: 2, t. 6 (1868)
= Oldenlandia corymbosa Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 408 (1834), nom. inval., non L.
= Oldenlandia pruinosa (Wight & Arn.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Oldenlandia pruinosa var. massiei Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 116 (1922)
species :
Hedyotis puberulifolia
Y.D.Xu & R.J.Wang; Phytotaxa 502(2): 161 (2021)
Distribution: China (Guangdong)
36 CHI
species :
Hedyotis pubescens
(Valeton) Merr. & L.M.Perry; J. Arnold Arbor. 26: 3 (1945)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Exallage pubescens (Valeton) Bremek.; Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 48(2) (Afr. Sp. Oldenlandia): 142 (1952)
= Oldenlandia pubescens Valeton; Nova Guinea 8: 439 (1911)
species :
Hedyotis pubicaulis
M.D.Yuan & R.J.Wang; Taiwania 67(1): 142 (2022)
Distribution: China (Guangxi, Yunnan)
36 CHI
species :
Hedyotis pubirachis
Y.D.Xu & R.J.Wang; Nordic J. Bot. 39(12)-e03264: 3 (2021)
Distribution: China (Guangdong)
36 CHI
species :
Hedyotis puffii
Wangwasit & Chantar.; Phytotaxa 206(1): 49 (2015)
Distribution: peninsular Thailand
41 THA
species :
Hedyotis pulchella
Stapf; Trans. Linn. Soc. London, Bot., Ser. 2, 4: 169 (1894)
Distribution: Borneo
42 BOR
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis pulchella var. magnistipula Wernham; J. Linn. Soc., Bot. 42: 89 (1914)
species :
Hedyotis pulcherrima
Dunn; Bull. Misc. Inform. Kew 1912: Add., Ser. 10: 127 (1912)
Distribution: China (Guangdong)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia pulcherrima (Dunn) Chun; Sunyatsenia 1: 314 (1934)
species :
Hedyotis punicea
Craib; Bull. Misc. Inform. Kew 1931: 277 (1931)
Distribution: NE-Thailand; Laos
41 LAO THA
species :
Hedyotis purpurascens
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 50 (1880)
Distribution: India (Tamil Nadu, Kerala)
40 IND
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis purpurascens var. pallida (Gamble) M.Mohanan; J. Econ. Taxon. Bot. 6(2): 480 (1985)
= Hedyotis purpurea Bedd.; Icon. Pl. Ind. Or. [Beddome] 1 (1868), nom. illeg.
= Oldenlandia purpurascens (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Oldenlandia purpurascens var. pallida Gamble; Fl. Madras 597 (1921)
species :
Hedyotis quinquenervia
Thwaites; Enum. Pl. Zeyl. [Thwaites] 141 (1859)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia quinquenervia (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis rajasekaranii
Karupp. & Ravich.; Edinburgh J. Bot. 71(1): 59 (2014)
Distribution: India (Tamil Nadu)
40 IND
species :
Hedyotis ramarowii
(Gamble) R.S.Rao & Hemadri; Indian Forester 99(6): 378 (1973)
Distribution: India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Oldenlandia ramarowii Gamble; Bull. Misc. Inform. Kew 1919: 405 (1919)
= Hedyotis ramarowii var. kannikattica Sundaresan, Manickam, Jothi & Murugan; Indian J. Forest. 25(1-2): 102 (2002)
species :
Hedyotis recurvata
Jiji, Jeomol, Manudev & Sunojk.; Phytotaxa 583(3): 287 (2023)
Distribution: S-India (Tamil Nadu)
40 IND
species :
Hedyotis resupinata
Ridl.; Bull. Misc. Inform. Kew 1926: 67 (1926)
Distribution: Sumatera (Mentawai Isl.)
42 SUM
species :
Hedyotis rhinophylla
Thwaites ex Trimen; J. Bot. 23: 208 (1885)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis rigida
(Blume) Walp.; Ann. Bot. Syst. [Walpers] 2: 772 (1852)
Distribution: China (Guangdong, Guangxi, Hainan, Yunnan); NE-India (Assam, Meghalaya, Nagaland, Tripura); Bangladesh; Andaman Isl. (Little Andaman Isl.); Nicobar Isl. (Car Nicobar Isl., North Nicobar Isl., Central Nicobar Isl., Great Nicobar Isl., Little Nicobar Isl.); Myanmar (Chin, Kayin, Mandalay, Taninthayi); Thailand; Cambodia; Vietnam; peninsular Malaysia; Singapore; Sumatera; Borneo; Sulawesi; Palawan (incl. Busuanga, Culion); Philippines (Luzon, Polillo, Mindoro, Sibuyan, Panay, Negros, Camiguin, Basilan, Mindanao); Jawa; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Australia (Queensland); Palau (Babeldaob, Kasioru)
36 CHI 40 BAN IND 41 AND CBD MYA NCB THA VIE 42 BOR JAW MLY MOL NWG PHI PLW SNG SUL SUM 50 AUS 62 PLU
Remarks: Hedyotis s. str. confirmed. See also Exallage cristata which frequently has been confused with this taxon. Distribution data might possibly be mixed.
Conservation Status: vulnerable (Singapore)
Synonyms: hidden
= Exallage rigida (Blume) Bremek.; Verh. Kon. Ned. Akad. Wetensch, Afd. Natuurk., Sect. 2, 48(2): 142 (1952)
= Hedyotis asperrima (Merr.) Merr.; Enum. Philipp. Fl. Pl. 3: 495 (1923)
= Hedyotis costata var. lutescens (Kaneh.) Fosberg; Smithsonian Contr. Bot. 45: 28 (1980)
= Hedyotis gitingensis Elmer; Leafl. Philipp. Bot. 3: 979 (1911)
= Hedyotis leucocarpa Elmer; Leafl. Philipp. Bot. 2: 524 (1908)
= Hedyotis lutescens Kaneh.; Trans. Nat. Hist. Soc. Taiwan 25: 4 (1935)
= Hedyotis philippensis var. asperrima Merr.; Philipp. J. Sci., C 8: 38 (1913)
= Hedyotis prostrata (Blume) Korth.; Ned. Kruidk. Arch. 2(2): 160 (1851), nom. illeg.
= Hedyotis pulgarensis Elmer; Leafl. Philipp. Bot. 4: 1333 (1912)
= Hedyotis rigida (Blume) Miq.; Fl. Ned. Ind. 2: 181 (1857)
= Hedyotis vestita var. lutescens (Kaneh.) Fosberg; Allertonia 6(3): 244 (1991)
= Metabolos prostratus Blume; Bijdr. Fl. Ned. Ind. 991 (1826)
= Metabolos rigidus Blume; Bijdr. Fl. Ned. Ind. 992 (1826)
= Oldenlandia prostrata (Blume) Koord.; Exkurs.-Fl. Java 3: 241 (1912), nom. illeg.
= Oldenlandia rigida (Blume) Kuntze; Revis. Gen. Pl. 1: 292 (1891), nom. illeg.
= Sclerococcus laevigatus Bartl. ex DC.; Prodr. [A.P. de Candolle] 4: 436 (1830), nom. inval.
= Sclerococcus lineatus Bartl. ex DC.; Prodr. [A.P. de Candolle] 4: 435 (1830)
= Scleromitrion rigidum (Blume) Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 46: 136 (1877)
species :
Hedyotis rivalis
Ridl.; J. Fed. Malay States Mus. 6: 153 (1915)
Distribution: peninsular Malaysia (Pahang, Johor)
42 MLY
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis rosmarinifolia
(Pit.) Craib; Fl. Siam 2: 47 (1932)
Distribution: Cambodia; Vietnam; NE-Thailand
41 CBD THA VIE
Synonyms: hidden
= Oldenlandia rosmarinifolia Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 127 (1922)
= Hedyotis valerianelloides (Pit.) P.H.Hô; Câyco Vietnam (Illustr. Fl. Vietnam) 3(1): 137 (1993)
= Oldenlandia valerianelloides Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 124 (1922)
species :
Hedyotis rugosa
(Blume) Korth.; Ned. Kruidk. Arch. 2(2): 160 (1851)
Distribution: Sumatera; Jawa
42 JAW SUM
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Allaeophania rugosa (Blume) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 124 (1891)
= Metabolos rugosus Blume; Bijdr. Fl. Ned. Ind. 991 (1826)
= Metabolos rugosus f. glabrescens Hochr.; Candollea 5: 278 (1934), nom. inval.
= Metabolos rugosus f. pubescens Hochr.; Candollea 5: 279 (1934)
= Oldenlandia rugosa (Blume) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Scleromitrion rugosum (Korth.) Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 46(2): 137 (1877)
species :
Hedyotis sachetiana
Fosberg; Allertonia 6(3): 235 (1991)
Distribution: Fed. States of Micronesia
62 MIC
species :
Hedyotis scaberrima
Merr.; Philipp. J. Sci. 17: 432 (1920 publ. 1921)
Distribution: Philippines (Panay)
42 PHI
species :
Hedyotis scaberula
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 50 (1880)
Distribution: Myanmar (Mon)
41 MYA
species :
Hedyotis scabridifolia
Kaneh.; Trans. Nat. Hist. Soc. Taiwan 25: 7 (1935)
Distribution: Northern Marianas (Pagan, Saipan)
62 MRN
Synonyms: hidden
= Hedyotis saipanensis Hosok.; J. Soc. Trop. Agric. 6: 669, nomen, et in Trans. Nat. Hist. Soc. Formosa, 1935 25: 37 (1934), descr.
variant :
var. stonei
Fosberg; Allertonia 6(3): 237 (1991)
Distribution: Northern Marianas
62 MRN
species :
Hedyotis schlechteri
Merr. & L.M.Perry; J. Arnold Arbor. 26: 1 (1945)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis schlechteri var. acuminata (Valeton) Merr. & L.M.Perry; J. Arnold Arbor. 26: 1 (1945)
= Oldenlandia schlechteri Valeton; Bot. Jahrb. Syst. 60: 11 (1925), nom. illeg., non Schinz
= Oldenlandia schlechteri var. acuminata Valeton; Bot. Jahrb. Syst. 60: 12 (1925)
species :
Hedyotis scoparia
(Pit.) P.H.Hô; Câyco Vietnam (Illustr. Fl. Vietnam) 3(1): 132 (1993)
Distribution: Myanmar; Thailand; Laos; Cambodia; Vietnam
41 CBD LAO MYA THA VIE
Synonyms: hidden
= Hedyotis cytisoides Craib; Bull. Misc. Inform. Kew 1931: 217 (1931)
= Hedyotis impedita Craib; Bull. Misc. Inform. Kew 1931: 275 (1931)
= Hedyotis praecox (Pit.) P.H.Hô; Câyco Vietnam (Illustr. Fl. Vietnam) 3(1): 131 (1993)
= Oldenlandia praecox Pierre ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 121 (1922)
= Oldenlandia scoparia Pierre ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 121 (1922)
species :
Hedyotis shenzhenensis
Tao Chen; Edinburgh J. Bot. 64(3): 331 (2007)
Distribution: China (Guangdong)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis shiuyingiae Tao Chen; Harvard Pap. Bot. 13: 283 (2008)
species :
Hedyotis shettyi
K.Ravik. & V.Lakshm.; Rheedea 9(1): 69 (1999)
Distribution: India (Tamil Nadu)
40 IND
Remarks: provisionally accepted name
species :
Hedyotis shoolamudi
Sunil, Naveen Kum. & K.M.P.Kumar; Phytotaxa 438(2): 159 (2020)
Distribution: India (Kerala)
40 IND
species :
Hedyotis sibuyanensis
Elmer; Leafl. Philipp. Bot. 3: 977 (1911)
Distribution: Philippines (Sibuyan)
42 PHI
species :
Hedyotis similis
E.T.Geddes; Bull. Misc. Inform. Kew 1928: 241 (1928)
Distribution: Thailand; peninsular Malaysia
41 THA 42 MLY
species :
Hedyotis simplex
Merr.; Philipp. J. Sci. 17: 433 (1920 publ. 1921)
Distribution: Philippines (Catanduanes, Biliran, Mindanao)
42 PHI
species :
Hedyotis simplicissima
(Lour.) Merr.; Trans. Amer. Philos. Soc., n. s. 24(2): 11, 362 (1935)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Petesia simplicissima Lour.; Fl. Cochinch. 77 (1790)
species :
Hedyotis sithiravaraiensis
Muruganand., Devanath., S.Ravik. & D.Naras.; J. Asia-Pacific Biodivers. 13: 750, figs. 1-3 (2020)
Distribution: India (Tamil Nadu)
40 IND
species :
Hedyotis srilankensis
Deb & Ratna Dutta; Taxon 32(2): 285 (1983), nom. nov.
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Hedyotis cymosa Thwaites; Enum. Pl. Zeyl. [Thwaites] 142 (1859)
= Oldenlandia cymosa (Thwaites) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis stelligera
Ridl.; J. Straits Branch Roy. Asiat. Soc. 59: 109 (1911)
Distribution: Thailand; peninsular Malaysia (once from Besut, Terengganu)
41 THA 42 MLY
species :
Hedyotis stylosa
R.Br. ex Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 407 (1834)
Distribution: India (Tamil Nadu)
40 IND
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia stylosa (R.Br. ex Wight & Arn.) Kuntze; Revis. Gen. Pl. 1: 293 (1891)
species :
Hedyotis suborthogona
Hosok.; Trans. Nat. Hist. Soc. Taiwan 28: 66 (1938)
Distribution: Palau (Babeldaob)
62 PLU
species :
Hedyotis subvelutina
Elmer; Leafl. Philipp. Bot. 3: 978 (1911)
Distribution: Philippines (Sibuyan)
42 PHI
species :
Hedyotis subvenosa
Merr.; Philipp. J. Sci. 17: 432 (1920, publ. 1921)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis subverticillata
Alston; Handb. Fl. Ceylon 6: Suppl., 148 (1931)
Distribution: Sri Lanka
40 SRL
species :
Hedyotis swertioides
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 51 (1880)
Distribution: S-India (Kerala, Tamil Nadu)
40 IND
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia swertiodes (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis symphyllarionoides
Ridsdale; Blumea 24(2): 363 (1978), nom. nov.
Distribution: Laos; Cambodia; Vietnam
41 CBD LAO VIE
Synonyms: hidden
= Symphyllarion herbaceum Gagnep.; Bull. Soc. Bot. France 95: 32 (1948)
species :
Hedyotis taishanensis
G.T.Wang & R.J.Wang; Phytotaxa 367(1): 39 (2018)
Distribution: China (Guangdong)
36 CHI
species :
Hedyotis tavoyensis
N.P.Balakr.; J. Bombay Nat. Hist. Soc. 71(1): 168 (1974)
Distribution: Myanmar (Chin, Taninthayi)
41 MYA
species :
Hedyotis tenuipes
Hemsl. ex Forbes & Hemsl.; J. Linn. Soc., Bot. 23: 375 (1888)
Distribution: China (Guangdong, Fujian)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia tenuipes (Hemsl.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis terminaliflora
Merr. & Chun; Sunyatsenia 2: 326 (1935)
Distribution: China (Hainan)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis ternata
(Pit.) P.H.Hô; Câyco Vietnam (Illustr. Fl. Vietnam) 3(1): 133 (1993)
Distribution: E-Thailand; Laos; Cambodia; Vietnam
41 CBD LAO THA VIE
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia ternata Pierre ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 122 (1922)
= Hedyotis ternata (Pierre ex Pit.) J.F.Maxwell; Nat. Hist. Bull. Siam Soc. 50(1): 4 (2002), isonym
species :
Hedyotis tetrangularis
(Korth.) Walp.; Ann. Bot. Syst. [Walpers] 2: 769 (1852)
Distribution: China (Guangdong, Guangxi); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Sumatera; Borneo
36 CHI 41 CBD LAO THA VIE 42 BOR MLY SUM
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Diplophragma tetrangularis Korth.; Ned. Kruidk. Arch. 2(2): 149 (1851)
= Hedyotis parryi Hance; Ann. Sci. Nat., Bot., Sér. 4, 18: 221 (1862)
= Hedyotis quadrangularis Miq.; Fl. Ned. Ind. 2: 180 (1857)
= Hedyotis subdivaricata Drake ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 124 (1922)
= Oldenlandia parryi (Hance) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Oldenlandia quadrangularis (Miq.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
= Oldenlandia subdivaricata Drake; J. Bot. [Morot] 9: 211 (1895)
= Oldenlandia tetrangularis (Korth.) Merr.; Pap. Michigan Acad. Sci. 23: 193 (1937 publ. 1938)
species :
Hedyotis thwaitesii
Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 54 (1880)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis macrophylla Thwaites; Enum. Pl. Zeyl. [Thwaites] 142 (1859), nom. illeg.
= Oldenlandia thwaitesii (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 293 (1891)
species :
Hedyotis tomentosa
(Valeton) Hosok.; Trans. Nat. Hist. Soc. Taiwan 24: 204 (1934)
Distribution: Palau (Babeldaob, Koror, Ngarakabesang)
62 PLU
Synonyms: hidden
= Oldenlandia tomentosa Valeton; Bot. Jahrb. Syst. 63: 292 (1930)
species :
Hedyotis tonggulingensis
G.B.Jiang & R.J.Wang; Phytotaxa 406(3): 193 (2019)
Distribution: China (Hainan)
36 CHI
species :
Hedyotis travancorica
Bedd.; Icon. Pl. Ind. Or. [Beddome] 1: 2 (1868)
Distribution: S-India (Tamil Nadu)
40 IND
Synonyms: hidden
= Oldenlandia travancorica (Bedd.) Kuntze; Revis. Gen. Pl. 1: 293 (1891)
species :
Hedyotis trichoneura
Alston; Handb. Fl. Ceylon 6: Suppl., 146 (1931)
Distribution: Sri Lanka
40 SRL
species :
Hedyotis tridentata
Ridsdale; Blumea 41: 456 (1996)
Distribution: Sri Lanka
40 SRL
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis trimenii
Deb & Ratna Dutta; Taxon 34(2): 297 (1985)
Distribution: India (Tamil Nadu); Sri Lanka
40 IND SRL
Remarks: Hedyotis s. str. confirmed
variant :
var. orbicularifolia
Ridsdale; Blumea 41(2): 4,57 (1996)
Distribution: Sri Lanka
40 SRL
species :
Hedyotis trisecta
Elmer; Leafl. Philipp. Bot. 9: 3249 (1934)
Distribution: Philippines (Luzon)
42 PHI
species :
Hedyotis tuyamae
Hosok.; Trans. Nat. Hist. Soc. Taiwan 28: 67 (1938)
Distribution: Palau (Babeldaob)
62 PLU
species :
Hedyotis uncinella
Hook. & Arn.; Bot. Beechey Voy. [Hooker & Arnott] 192 (1833)
Distribution: China (Guangdong, Hainan, Hunan, Guizhou, Hongkong); Taiwan; NE-India (Meghalaya, Sikkim); Bangladesh; Myanmar (Mandalay); Thailand; Laos; Cambodia; Vietnam
36 CHI 38 TAI 40 BAN IND 41 CBD LAO MYA THA VIE
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Hedyotis borrerioides Champ. ex Benth.; Hooker´s J. Bot. Kew Gard. Misc. 4: 171 (1852)
= Hedyotis cephalophora R.Br. ex Wall.; Numer. List [Wallich] no. 842 (1829), nomen
= Hedyotis kuraruensis Hayata; Icon. Pl. Formosan. 9: 53 (1920)
= Oldenlandia uncinella (Hook. & Arn.) Kuntze; Revis. Gen. Pl. 1: 293 (1891)
variant :
var. mekongensis
(Pierre ex Pit.) W.C.Chen; Fl. Yunnan. 15: 39 (2003)
Distribution: China (Yunnan); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Oldenlandia uncinella var. mekongensis Pierre ex Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 126 (1922)
species :
Hedyotis vachellii
Hook. & Arn.; Bot. Beechey Voy. [Hooker & Arnott] 194 (1833)
Distribution: China (Hongkong)
36 CHI
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia vachelii (Hook. & Arn.) Kuntze; Revis. Gen. Pl. 1: 292 (1891)
species :
Hedyotis valetoniana
Merr. & L.M.Perry; J. Arnold Arbor. 26: 2 (1945)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia coprosmoidea Valeton ex H.J.Lam; Natuurk. Tijdschr. Ned.-Indie 89: 72, 101, 133, 134 (1929), nomen
species :
Hedyotis verticillaris
Wall. ex Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 409 (1834)
Distribution: India (Arunachal Pradesh, Assam, Bihar, Himachal Pradesh, Meghalaya, Mizoram Nagaland, Sikkim, Tripura, Uttar Pradesh, West Bengal)
40 IND
Remarks: Hedyotis s. str. confirmed
Synonyms: hidden
= Oldenlandia verticillaris (Wall. ex Wight & Arn.) Kuntze; Revis. Gen. Pl. 1: 293 (1891)
= Pleiocraterium verticillare (Wight & Arn.) Bremek.; Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 56: 440 (1939), et in Recueil Trav. Bot. Néerl. 36: 440 (1939 publ. 1940)
species :
Hedyotis wangii
R.J.Wang; Acta Phytotax. Sin. 41(1): 85 (2003)
Distribution: China (Yunnan)
36 CHI
species :
Hedyotis whiteheadii
Merr.; Philipp. J. Sci., C 2: 303 (1907)
Distribution: Philippines (Mindoro)
42 PHI
species :
Hedyotis wuzhishanensis
R.J.Wang; Novon 18(2): 266 (2008)
Distribution: China (Hainan)
36 CHI
species :
Hedyotis xinyiensis
X.Guo & R.J.Wang; Ann. Bot. Fenn. 48(5): 443 (2011)
Distribution: China (Guangdong)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis yangchunensis
W.C.Ko & G.C.Zhang; J. S. China Agric. Univ. 16(4): 45 (1995)
Distribution: China (Guangdong)
36 CHI
Remarks: Hedyotis s. str. confirmed
species :
Hedyotis yazhouensis
F.W.Xing & R.J.Wang; Acta Phytotax. Sin. 41(1): 87 (2003)
Distribution: China (Hainan)
36 CHI
species :
Hedyotis zhihaoana
Huan C.Wang & Xiao Lan Liu; Phytotaxa 461(3): 168 (2020)
Distribution: China (Yunnan)
36 CHI
