001.0000
Family
Lycopodiaceae P.Beauv. ex Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 4: 293 (1802)
17 genera, ca. 470 spp. Phylogeny: Chen et al. (2021).
Species listed in database: 476; hybrids: 20
= Huperziaceae Rothm.; Feddes Repert. 66: 236 (1962), nom. nov.
= Urostachyaceae Rothm.; Feddes Repert. 54: 58 (1944) (nom. illeg.)
= Phylloglossaceae Kunze; Bot. Zeitung (Berlin) 1: 722 (1843)
= Lycopodiellaceae Val.N.Tikhom.; Novosti Sist. Vyssh. Rast. 49: 149 (2018)
001.4003
Genus
Huperzia Bernh.; J. Bot. (Schrader) 1800(2): 126 (1801) ex Schrank & Mart. (1829)
Distribution: Probably only 25 spp. temperate, alpine and arctic, tropical Asian mountains. Many remaining species still need transfer into other split genera.
Species listed in database: 62; hybrids: 8
= Mirmau Adans.; Fam. Pl. 2: 491 (1763) (p. p., nom. illegit.)
= Plananthus P.Beauv. ex Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 3: 476, 4: 310 (1802) (nom. superfl.)
= Selago Boehm.; Ludwig, Def. Gen. Pl., ed. Boehmer 484 (1760) (non L. 1753, nec Browne 1756, nec Hill (1757) nom. illeg.)
= Selago Hill; Brit. Herb. 533 (1757) (non L. 1753, nec Browne 1756, nom. inval.)
= Selago Schur; Enum. pl. Transsilv. 825. 1866 (non L. 1753, nec Browne 1756, nec Hill (1757))
= Urostachys Herter; Beih. Bot. Centralbl. 39(2): 249 (1922) (nom. superfl.)
species :
Huperzia acicularis
Björk; Ann. Bot. Fenn. 57(4-6): 274 (2020)
Distribution: England; Norway; N-European Russia; Canada (British Columbia)
10 GRB NOR 14 RUS 71 CDN
Remarks: provisionally accepted name, validity highly disputed
species :
Huperzia appalachiana
Beitel & Mickel; Amer. Fern J. 82: 45 (1992)
Name: Appalachian clubmoss
Distribution: Greenland; Canada (Labrador, New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec); St. Pierre et Miquelon; USA (Georgia, Massachusetts, Maine, Michigan, Minnesota, North Carolina, New Hampshire, New York, South Carolina, Tennessee, Virginia, Vermont, Wisconsin)
70 GNL 72 CDN SPM 74 USA 75 USA 78 USA
Remarks: most probably a synonym of H. (selago ssp.) appressa fide Fraser-Jenkins et al. (2017)
species :
Huperzia archboldiana
(Nessel) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Remarks: selago agg.
Synonyms: hidden
= Urostachys archboldianus Nessel; Repert. Spec. Nov. Regni Veg. 39: 61, t. 189a (1935)
= Lycopodium archboldianum (Nessel) Alston; J. Bot. 77: 290 (1939)
species :
Huperzia arunachalensis
(D.D.Pant & P.S.Pandey) Fraser-Jenk.; Ferns and Fern-allies of Nepal 1: 45 (2015) (published in January)
Distribution: S-China; Tibet; Nepal; Bhutan; NE-India (Arunachal Pradesh, West Bengal, Sikkim)
36 CHI TIB 40 BHU IND NEP
Remarks: selago agg.
Synonyms: hidden
= Huperzia mazumdariana C.Das; Int. J. Adv. Res. Innov. Ideas Educ. 3(3): 3112 (2017), nom. nov. superfl. illeg. for Lycopodium arunachalense D. D. Pant & P. S. Pandey
= Lycopodium arunachalense D.D.Pant & P.S.Pandey; Phyta Monogr. 3: 11-13, t. 3A (1985)
= Lycopodium sahnii D.D.Pant & P.S.Pandey; Phyta Monogr. 3: 16, t. 10A (1985)
species :
Huperzia asiatica
(Ching) N.Shrestha & X.C.Zhang; Phytotaxa 219(3): 249 (2015)
Distribution: China (Jilin: Changbai-san); North Korea (Baekdu-san)
36 CHI 38 NKO
Synonyms: hidden
= Huperzia asiatica (Ching) B.Y.Sun & J.Lim; Korean J. Pl. Taxon. 45(4): 302 (2015), isonym
= Huperzia lucidula var. asiatica Ching; Acta Bot. Yunnan. 3(3): 296 (1981)
species :
Huperzia australiana
(Herter) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
Distribution: Borneo (Sabah); Sulawesi; Australia (SE-New South Wales, E-Victoria, Tasmania); New Zealand (North Isl., South Isl., Stewart Isl.); Chatham Isl.; Auckland Isl.; Macquarie Isl.; Campbell Isl.
42 BOR SUL 50 AUS 51 AUC CTM NZL 91 CMP MAQ
Remarks: selago agg.
Synonyms: hidden
= Lycopodium australianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 42 (1909)
= Urostachys australianus (Herter) Herter; Bot. Jahrb. Syst. 43: 42 (1909) ex Nessel, Bärlappgew. 49 (1939)
species :
Huperzia beccarii
(Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
Distribution: Sumatra; Borneo (Sabah)
42 BOR SUM
Remarks: selago agg.
Synonyms: hidden
= Lycopodium beccarii Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 29 (1912)
= Urostachys beccarii (Alderw.) Herter ex Nessel; Bärlappgew. 44 (1939)
species :
Huperzia beiteliana
Mickel; Brittonia 44(3): 312, f. 1A-E (1992)
Distribution: Mexico (San Luis Potosi, Oaxaca)
79 MEX
Remarks: selago agg.
species :
Huperzia bucahwangensis
Ching; Acta Bot. Yunnan. 3(3): 301 (1981)
Distribution: Tibet; India (Arunachal Pradesh); Myanmar (Kachin, etc.)
36 TIB 40 IND 41 MYA
Remarks: selago agg.
Synonyms: hidden
= Huperzia arunachalensis Mazumdar & R.Mukhop.; Bionature 34(2): 33 (2014 publ. 2015), nom. illeg.
= Huperzia kingdonwardiana Fraser-Jenk.; Indian J. Forest. 38(4): 335 (2015)
species :
Huperzia campestris
(Alderw.) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: selago agg.
Synonyms: hidden
= Lycopodium campestre Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 3 (1917)
= Urostachys campestris (Alderw.) Herter ex Nessel; Bärlappgew. 45 (1939)
species :
Huperzia catharinae
(Christ) Holub; Folia Geobot. Phytotax. 20: 71 (1985)
Distribution: Brazil (Santa Catarina)
84 BRA
Synonyms: hidden
= Lycopodium catharinae Christ; Schwacke, Pl. Nov. Mineiras 2: 39 (1900)
= Urostachys catharinae (Christ) Herter ex Nessel & Hoehne; Arch. Bot. São Paulo 1: 387 (1927)
species :
Huperzia ceylanica
(Spring) Rothm.; Feddes Repert. 54: 59 (1951)
Distribution: China (W-Sichuan, W-Yunnan); India (Tamil Nadu, Arunachal Pradesh, Kerala, Tripura, West Bengal, Meghalaya); Sri Lanka; Sumatra
36 CHI 40 IND SRL 42 SUM
Remarks: selago agg.
Conservation Status: IUCN NT (Fraser-Jenkins 2012)
Synonyms: hidden
= Huperzia vorwerkii (Nessel) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
= Lycopodium ceylanicum Spring; Bull. Acad. Roy. Sci. Bruxelles 8(2): 514 (1841)
= Urostachys ceylanicus (Spring) Herter ex Nessel; Bärlappgew. 52 (1939)
= Urostachys vorwerkii Nessel; Repert. Spec. Nov. Regni Veg. 36(12-15): 178 (1934)
species :
Huperzia chinensis
(Christ) Ching; Acta Bot. Yunnan. 3(3): 304 (1981), et Fl. Xizangica 1: 9 (1983)
Distribution: Siberia (E-Siberia); Russian Far East (Amur, Ussuri); China (Hubei, Hunan, Shaanxi, Sichuan); ?Vietnam
30 SIB 31 RFE 36 CHI
Remarks: selago agg.
Synonyms: hidden
= Huperzia selago subsp. chinensis (Christ) Á.Löve & D.Löve
= Huperzia takingensis Ching; Acta Bot. Yunnan. 3(3): 300-301 (1981)
= Lycopodium chinense Christ; Nuov. Giorn. Bot. Ital. n. s. 4(1): 1011, t. 3, f. 4 (1897)
= Lycopodium selago subsp. chinense (Christ) Hultén; Fl. Alaska (Hulten) 26 (1968), nom. inval.
= Lycopodium selago var. chinense (Christ) Ohwi; Fl. Jap. Pterid. 24 (1965) (nom. illeg., non Lycopodium selago subsp. chinense (Christ) Hultén)
= Urostachys chinense (Christ) Herter ex Nessel; Bärlappgew. 27, Abb. 4, f. 2 (1939)
species :
Huperzia chishuiensis
X.Y.Wang & P.S.Wang; Bull. Bot. Res., Harbin 16(4): 417, f. 1-3 (1996)
Distribution: China (N-Guizhou)
36 CHI
species :
Huperzia continentalis
Testo, A.Haines & A.V.Gilman; Syst. Bot. 41(4): 894 (2016)
Distribution: Norway; Russian Far East (Magadan); Alaska; Greenland; Canada (Alberta, British Columbia, Manitoba, Northwest Territories, Newfoundland, Nunavut, Ontario, Québec, Saskatchewan, Yukon); USA (Washington State, Colorado, Montana, Wyoming)
10 NOR 31 RFE 70 ASK CDN GNL 71 CDN 72 CDN 73 USA
species :
Huperzia crispata
(Ching) Ching; Acta Bot. Yunnan. 3(3): 293 (1981)
Distribution: China (Chongqing, Guizhou, Jiangxi, W-Hubei, W-Hunan, Sichuan, NE-Yunnan)
36 CHI
Remarks: selago agg.
Synonyms: hidden
= Lycopodium crispatum Ching; Acta Phytotax. Sin. 18(2): 236, f. 1: 3 (1980)
species :
Huperzia delavayi
(Christ & Herter) Ching; Acta Bot. Yunnan. 3: 303 (1981)
Distribution: China; Tibet; Nepal; N-Myanmar (Kachin, etc.)
36 CHI TIB 40 NEP 41 MYA
Remarks: selago agg.; a valid species separate from H. ceylanica fide Fraser-Jenkins et al. 2018
Synonyms: hidden
= Lycopodium delavayi Christ & Herter; Herter, Bot. Jahrb. Syst. 43(1, Beibl. 98): 41 (1909)
= Urostachys delavayi (Christ & Herter) Herter ex Nessel; Bärlappgew. 31, Abb. 4, f. 9 (1939)
species :
Huperzia emeiensis
(Ching & H.S.Kung) Ching & H.S.Kung; Acta Bot. Yunnan. 3(3): 299-300 (1981)
Distribution: China (Chongqing, N-Guizhou, W-Hubei, Hunan, Sichuan, NE-Yunnan); Taiwan
36 CHI 38 TAI
Remarks: selago agg.
Synonyms: hidden
= Lycopodium emeiense Ching & H.S.Kung ex H.S.Kung; Acta Phytotax. Sin. 18(2): 235, f. 2 (1980)
species :
Huperzia erosa
Beitel & W.H.Wagner; Contr. Univ. Michigan Herb. 22: 140 (1999), nom. nov.
Distribution: Hawaii (Kauai, Oahu, Lanai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Huperzia serrata var. dentata (Hillebr.) Kartesz & Gandhi; Phytologia 70(3): 201 (1991)
= Lycopodium serratum var. dentatum Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 642 (-643) (1888)
species :
Huperzia erubescens
(Brack.) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Remarks: selago agg.
Synonyms: hidden
= Lycopodium erubescens Brack.; U.S. Expl. Exped., Filic. 16: 320, atlas, t. 45, f. 1 (1854)
= Urostachys erubescens (Brack.) Herter ex Nessel; Bärlappgew. 61 (1939)
species :
Huperzia europaea
Björk; Ann. Bot. Fenn. 57(4-6): 276 (2020)
Distribution: Iceland; Great Britain; Norway; Sweden; Finland; ?Germany; Poland; Czech Republic; France; Italy; Romania; Estonia; N-European Russia; Ukraine
10 FIN GRB ICE NOR SWE 11 CZE POL 12 FRA 13 ITA ROM 14 EST RUS UKR
Remarks: provisionally accepted name, validity highly disputed
species :
Huperzia everettii
(Herter) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
Distribution: S-Sulawesi
42 SUL
Remarks: selago agg.
Synonyms: hidden
= Lycopodium everettii Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Urostachys everettii (Herter) Herter ex Nessel; Bärlappgew. 66 (1939)
species :
Huperzia fuegiana
(Roiv.) Holub; Folia Geobot. Phytotax. 20: 72 (1985)
Distribution: Argentina (Santa Cruz Prov., Chubut, Tierra del Fuego); Chile (Aisen, Magallanes); Falkland Isl.
85 ARG CLE 91 FAL
Remarks: selago agg.
Conservation Status: IUCN R. Endangered in Falkland Islands
Synonyms: hidden
= Lycopodium fuegianum Roiv.; Ann. Bot. Soc. Zool.-Bot. Vanamo 6(8): 13-15, f. 4-5 (1936)
= Urostachys fuegianus (Roiv.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 62 (1949)
= Urostachys selago var. aschersonianus Herter ex Nessel; Bärlappgew. 40 (1939) (nom. illeg.), et Rev. Sudam. Bot. 6: 157 (1940)
= Urostachys selago var. hessei Herter ex Nessel; Bärlappgew. 40 (1939) (nom. illeg.), et Rev. Sudam. Bot. 6: 157 (1940)
species :
Huperzia gedeana
(Alderw.) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
Distribution: Java
42 JAW
Remarks: selago agg.
Synonyms: hidden
= Lycopodium gedeanum Alderw.; Malayan Ferns Fern Allies Suppl 1(Corr.): 65 (1917) (Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 43, t. 8 (1918)
= Urostachys gedeanus (Alderw.) Herter ex Nessel; Bärlappgew. 45 (1939)
species :
Huperzia goliathensis
(Alderw.) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
Distribution: peninsular Malaysia (Pahang); Borneo (Sabah)
42 BOR MLY
Remarks: selago agg.
Synonyms: hidden
= Lycopodium goliathense Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 7: 29 (1912)
= Urostachys goliathensis (Alderw.) Herter ex Nessel; Bärlappgew. 44 (1939)
species :
Huperzia haleakalae
(Brack.) Holub; Folia Geobot. Phytotax. 20: 73 (1985)
Name: Pacific clubmoss
Distribution: +Hawaii (+Maui)
63 HAW
Remarks: selago agg.
Synonyms: hidden
= Lycopodium haleakalae Brack.; U.S. Expl. Exped., Filic. 16: 321 (1854)
= Lycopodium selago var. haleakalae (Brack.) Warb.; Monsunia 1: 98 (1900)
= Urostachys haleakalae (Brack.) Herter ex Nessel; Bärlappgew. 30 (1939)
species :
Huperzia herteriana
(Kümmerle) T.Sen & U.Sen; Fern Gaz. 11(6): 415-417, f. 1, k-r (1978)
Distribution: China (Chongqing, Guizhou, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Sikkim, West Bengal, Manipur, Darjeeling, Arunachal Pradesh); Myanmar (Kachin, Chin, etc.); Thailand; ?Java
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA THA
Remarks: selago agg.
Synonyms: hidden
= Huperzia kamaensis Ching & S.K.Wu; Acta Bot. Yunnan. 3(3): 300 (1981)
= Huperzia maerhkangensis Ching; Acta Bot. Yunnan. 3. 295 (1981)
= Huperzia multidichotoma Ching; Acta Bot. Yunnan. 3: 295 (1981)
= Huperzia obscure-denticulata Ching; Acta Bot. Yunnan. 3: 295 (1981)
= Huperzia thwaitesii (Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
= Lycopodium herterianum Kümmerle; Mag. Bot. Lapok 26: 99 (1927 publ. 1928)
= Lycopodium sikkimense Herter; Bot. Jahrb. Syst. 43(1, Beibl. 98): 42-43 (1909), nom. illeg., non Lycopodium sikkimense Müll. Hal. (1861)
= Urostachys herterianus (Kümmerle) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 64 (1949)
= Urostachys sikkimensis (Herter) Herter ex Nessel; Bärlappgew. 52 (1939)
= Urostachys thwaitesii Herter ex Nessel; Bärlappgew. 52 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 158, t. 8, f. 14 (1940)
species :
Huperzia javanica
(Sw.) C.Y.Yang; Bull. Acad. Military Med. Sci. 13: 368 (1989)
Distribution: China; Tibet; South Korea (incl. Cheju Isl.); Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, West Bengal); Sri Lanka; Myanmar (Kachin, etc.); Thailand; Cambodia; Vietnam; peninsular Malaysia (Pahang, Perak); Sumatra; Borneo (Sabah, Mt. Kinabalu); Sulawesi; Sulawesi; ?Philippines; Philippines; Lesser Sunda Isl. (Lombok); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal); Australia (NE-Queensland); New Caledonia; Vanuatu; ?Fiji (?Viti Levu); [Western] Samoa (Savai´i); Hawaii (Kauai, Oahu, Molokai, Lanai, Hawaii Isl.); Mexico (Hidalgo, Oaxaca); Hispaniola (Haiti)
36 CHI TIB 38 KOR TAI 40 BHU IND NEP SRL 41 CBD MYA THA VIE 42 BOR LSI MLY MOL NWG PHI SUL SUM 43 SOL 50 AUS 60 NWC SAM VAN 63 HAW 79 MEX 81 HIS
Remarks: inclusion of var. longipetiolata fide Fraser-Jenkins et al. (2018). Used to be included in H. serrata. All records of "serrata" outside of East Asia belong here, but identity of the American and Australian taxa not confirmed (Fraser-Jenkins in litt.)
Synonyms: hidden
= Huperzia indica S.R.Ghosh; Pterid. Fl. E. India: 51 (2004), herbarium of type not stated.
= Huperzia javanica (Sw.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 10 (2008), isonym
= Huperzia longipetiolata (Spring) Y.C.Yang; Chin. Tradit. Herbal Drugs 16(7): 32 (1982), nom. superfl.
= Huperzia rubricaulis (Alderw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Huperzia serrata f. longipetiolata (Spring) Ching; Acta Bot. Yunnan. 3(3): 294 (1981)
= Huperzia serrata subsp. javanica (Sw.) Satou; Hikobia 12(3): 268 (1997)
= Huperzia serrata var. longipetiolata (Spring) H.M.Chang; Taiwania 46(4): 378 (2001)
= Huperzia serrata var. longipetiolata (Spring) S.R.Ghosh; Pterid. Fl. E. India 55 (57) (2004), isonym
= Lycopodium javanicum Sw.; J. Bot. (Schrader) 1800(2): 114 (1801) (Syn. Fil. 175, 399 (1806))
= Lycopodium khasianum D.D.Pant & P.S.Pandey; Phyta Monogr. 3: 16, t. 6B (1985)
= Lycopodium rubricaule Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 24: 3 (1917)
= Lycopodium sargassifolium Liebm.; Overs. Kongel. Danske Vid. Selsk. Forh. Medlemmers Arbeider 1847: 41 (1847)
= Lycopodium serratum var. javanicum (Sw.) Makino; Bot. Mag. 12(131): 13 (1898)
= Lycopodium serratum var. longipetiolatum Spring; Mém. Acad. Roy. Sci. Belgique 24: 18 (1849)
= Lycopodium serratum var. minus D.D.Pant & P.S.Pandey; Phyta Monogr. 3: 19 (1985) (tentative synonym)
= Urostachys javanicus (Sw.) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 66 (1949)
= Urostachys rubricaulis (Alderw.) Herter ex Nessel; Bärlappgew. 44 (1939)
= Urostachys serratus var. japonico-neotropicus Herter ex Nessel; Bärlappgew. 59 (1939), without Latin diagnosis, et Revista Sudamer. Bot. 6: 159, tab. 8, fig. 19 (1940)
species :
Huperzia jejuensis
B.Y.Sun & J.Lim; Korean J. Pl. Taxon. 45(1): 19 (2015)
Distribution: South Korea (Cheju Isl.)
38 KOR
species :
Huperzia kangdingensis
(Ching) Ching; Acta Bot. Yunnan. 3(3): 294 (1981)
Distribution: China (W-Sichuan, NE-Yunnan)
36 CHI
Remarks: selago agg.
Synonyms: hidden
= Huperzia tahkuanensis Ching; Acta Bot. Yunnan. 3(3): 297 (1981)
= Lycopodium kangdingense Ching; Acta Phytotax. Sin. 18(2): 236-237, f. 1: 4, t. 6, f. 3 (1980)
species :
Huperzia kunmingensis
Ching; Acta Bot. Yunnan. 3: 297 (1981)
Distribution: China (Guangdong, Guangxi, W-Guizhou, Yunnan)
36 CHI
Remarks: selago agg.
species :
Huperzia laipoensis
Ching; Acta Bot. Yunnan. 3: 300 (1981)
Distribution: China (S-Sichuan)
36 CHI
Remarks: selago agg.
species :
Huperzia lajouensis
Ching; Acta Bot. Yunnan. 3(3): 305 (1981)
Distribution: China (Himalayas); Tibet; Nepal; India (Sikkim); N-Myanmar
36 CHI TIB 40 IND NEP 41 MYA
Remarks: partially treated as ´Huperzia dixitiana´ in earlier works. H. cavei is maintained as separate in Shalimov et al. 2017 but Fraser-Jenkins (in litt.) believes it to be a synonym here.
Synonyms: hidden
= Huperzia cavei Fraser-Jenk. & Kholia; Ferns and Fern-Allies of Nepal 1: 46 (2015)
species :
Huperzia leishanensis
X.Y.Wang; Bull. Bot. Res., Harbin 14(4): 355-356, f. 1-3 (1994)
Distribution: China (Guizhou, Sichuan, Yunnan)
36 CHI
species :
Huperzia liangshanica
(H.S.Kung) Ching & H.S.Kung; Acta Bot. Yunnan. 3(3): 296 (1981)
Distribution: China (Sichuan, ?Yunnan); Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Remarks: selago agg.
Synonyms: hidden
= Lycopodium liangshanicum H.S.Kung; Acta Bot. Yunnan. 2(1): 100-101, f. 1-3 (1980)
species :
Huperzia lucidula
(Michx.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
Name: Shining Clubmoss
Distribution: Canada (Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec); St. Pierre et Miquelon; USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, North Carolina, New Hampshire, New Jersey, New Mexico, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)
71 CDN 72 CDN SPM 74 USA 75 USA 77 USA 78 USA
Remarks: selago agg.
Synonyms: hidden
= Huperzia lucidula var. tryonii (Mohlenbr.) Mohlenbr.; Illustr. Fl. Illinois Ferns (ed. 2) 180 (1999)
= Huperzia selago subsp. lucidula (Michx.) Á.Löve & D.Löve; Univ. Colorado Stud. 17: 5 (1965)
= Lycopodium lucidulum Michx.; Fl. bor. Amer. 2: 284 (1803)
= Lycopodium lucidulum var. tryonii Mohlenbr.; Ill. Fl. Illinois, Grasses: Bromus-Paspalum 26, fig. 61e (1967)
= Lycopodium reflexum (P.Beauv.) Sw.; Syn. Fil. 175 (1806) (non Lam. (1792))
= Lycopodium selago var. lucidulum (Michx.) Clute; Fern Bull. 11: 47 (1903)
= Urostachys lucidulus (Michx.) Herter ex Nessel; Bärlappgew. 49 (1939)
= Urostachys lucidulus var. reclinatus (Dup) Nessel; Bärlappgew. 50 (1939)
species :
Huperzia medogensis
Ching & Y.X.Lin; Acta Phytotax. Sin. 22(3): 193 (1984)
Distribution: Tibet
36 TIB
species :
Huperzia meghalaica
Fraser-Jenk.; Indian J. Forest. 38(4): 336 (2015)
Distribution: China (Yunnan); India (Meghalaya, Nagaland, Mizoram); Myanmar
36 CHI 40 IND 41 MYA
species :
Huperzia miniata
(Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
Distribution: Sulawesi; Philippines; Java; New Guinea (Irian Jaya, Papua New Guinea)
42 JAW NWG PHI SUL
Remarks: selago agg.
Synonyms: hidden
= Lycopodium miniatum Spring; Bull. Acad. Roy. Soc. Bruxelles 8: 513 (1841)
= Lycopodium selago var. miniatum (Spring) B.C.Tan & Tolentino; Philipp. J. Sci. 116(4): 437-438 (1987)
= Urostachys miniatus (Spring) Herter ex Nessel; Bärlappgew. 42 (1939)
species :
Huperzia minima
(Herter) Holub; Folia Geobot. Phytotax. 20: 74 (1985)
Distribution: Philippines
42 PHI
Remarks: selago agg.
Synonyms: hidden
= Urostachys minimus Herter; Bot. Arch. 3: 13 (1923)
species :
Huperzia miyoshiana
(Makino) Ching; Acta Bot. Yunnan. 3(3): 303, 304 (1981)
Name: Chinese clubmoss
Distribution: Russian Far East (Kamchatka, Kuril Isl., Primorye, Sakhalin); China (Heilongjiang, Jilin, Liaoning); North Korea; South Korea (incl. Cheju Isl.); Japan; Alaska; Canada (British Columbia, Newfoundland, Ontario); USA (Oregon, Washington State)
31 RFE 36 CHI 38 JAP KOR NKO 70 ASK 71 CDN 72 CDN 73 USA
Synonyms: hidden
= Huperzia chinensis subsp. miyoshiana (Makino) Tzvelev; Bot. Zhurn. (Moscow & Leningrad) 84(1): 84 (1999)
= Huperzia miyoshiana var. coreana (Hayata) Ching; Acta Bot. Yunnan. 3(3): 304 b(1981)
= Huperzia selago var. miyoshiana (Makino) Roy L.Taylor & MacBryde; Canad. J. Bot. 56: 185 (1978)
= Lycopodium chinense Tagawa; Colenso Ill. Jap. Pterid. 8, pl. 1, f. 3 (1959)
= Lycopodium miyoshianum Makino; Bot. Mag. (Tokyo) 12: 36 (1898)
= Lycopodium miyoshianum var. coreanum Hayata; Icon. Pl. Formosan. 5: 255 (1915)
= Lycopodium selago subsp. miyoshianum (Makino) Calder & Roy L.Taylor; Canad. J. Bot. 43: 1388 (1965)
= Lycopodium selago var. chinense Ohwi; Fl. Jap. 24 (1965)
= Lycopodium selago var. miyoshianum (Makino) Makino; Bot. Mag. 16(189): 199 (1902)
= Lycopodium tenuifolium Herter; Bot. Jahrb. Syst. 43(1, Beibl. 98): 41 (1909)
= Urostachys miyoshianus (Makino) Herter ex Nessel; Bärlappgew. 28, Abb. 4, f. 4 (1939)
= Urostachys miyoshianus var. coreanus (Hayata) Herter ex Nessel; Bärlappgew. 28, Abb. 4, f. 5 (1939)
species :
Huperzia muscicola
W.M.Chu; Fl. Yunnanica 20: 718 (5-6) (2006)
Distribution: China (Yunnan)
36 CHI
species :
Huperzia myriophyllifolia
(Hayata) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
Distribution: Taiwan
38 TAI
Remarks: extremely rare. Status as full species see Wan et al. 2023
Synonyms: hidden
= Huperzia serrata var. myriophyllifolia (Hayata) H.M.Chang; Taiwania 46(4): 380 (2001)
= Lycopodium serratum var. myriophyllifolium Hayata; Icon. Pl. Formosan. 4: 133 (1914)
= Urostachys myriophyllifolius (Hayata) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 20: 71 (1949)
species :
Huperzia nanchuanensis
(Ching & H.S.Kung) Ching & H.S.Kung; Acta Bot. Yunnan. 3(3): 302 (1981)
Distribution: China (Chongqing, N-Guizhou, W-Hubei, Hunan, NE-Yunnan)
36 CHI
Remarks: selago agg.
Synonyms: hidden
= Huperzia hupehensis Ching; Acta Bot. Yunnan. 3: 301 (1981)
= Huperzia nanchuanense (Ching & H.S.Kung) S.R.Ghosh; Pterid. Fl. E. India 50 (2004), isonym
= Lycopodium nanchuanense Ching & H.S.Kung ex H.S.Kung; Acta Phytotax. Sin. 18(2): 235 (1980)
species :
Huperzia nanlingensis
Shrestha, F.W.Xing, X.P.Qi, Y.H.Yan & X.C.Zhang; Phytotaxa 175(5): 73-79 (2014)
Distribution: China (Guangdong, Hunan); Vietnam
36 CHI 41 VIE
species :
Huperzia occidentalis
(Clute) Kartesz & Gandhi; Phytologia 70: 201 (1991)
Name: Western clubmoss
Distribution: Alaska; Canada (Alberta, British Columbia, SW-Yukon); USA (Idaho, Montana, Oregon, Washington State)
70 ASK CDN 71 CDN 73 USA
Synonyms: hidden
= Huperzia occidentalis (Clute) Beitel; Amer. Fern J. 82(2): 46 (1992) (nom. illeg.)
= Lycopodium lucidulum f. occidentale Clute; Fern Bull. 11: 13 (1903)
= Lycopodium lucidulum var. occidentale (Clute) L.R.Wilson; Rhodora 34: 170 (1932)
= Lycopodium selago var. occidentale (Clute) B.Boivin; Le Naturaliste Canadien 93: 359 (1966)
species :
Huperzia porophila
(F.E.Lloyd & Underw.) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
Name: Rock clubmoss
Distribution: Canada (Ontario); USA (Alabama, Georgia, Iowa, Illinois, Indiana, Kentucky, Maryland, Minnesota, Missouri, North Carolina, Ohio, Pennsylvania, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)
72 CDN 74 USA 75 USA 78 USA
Remarks: selago agg.
Synonyms: hidden
= Huperzia selago var. porophila (F.E.Lloyd & Underw.) Á.Löve & D.Löve; Univ. Colorado Stud. Ser. Biol. Ser. no. 17: 5 (1965)
= Lycopodium lucidulum f. porophilum (F.E.Lloyd & Underw.) Clute; Fern Bull. 16: 22 (1908)
= Lycopodium lucidulum var. porophilum (F.E.Lloyd & Underw.) Clute; Fern-Allies 262 (1905)
= Lycopodium porophilum F.E.Lloyd & Underw.; Bull. Torrey Bot. Club 27(4): 150 (1900)
= Lycopodium selago var. porophila (F.E.Lloyd & Underw.) Clute; Fern Bull. 11: 47 (1903), sphalm. ´porophyllum´
= Urostachys lucidulus var. porophilus (F.E.Lloyd & Underw.) Nessel; Bärlappgew. 50 (1939)
= Urostachys porophilus (F.E.Lloyd & Underw.) Herter; Philipp. J. Sci. 22: 179 (1923)
species :
Huperzia quasipolytrichoides
(Hayata) Ching; Acta Bot. Yunnan. 3(3): 299 (1981)
Distribution: China (Anhui, Hunan); Taiwan
36 CHI 38 TAI
Remarks: selago agg.
Synonyms: hidden
= Huperzia reflexointegra (Hayata) Holub; Folia Geobot. Phytotax. 20: 76 (1985)
= Huperzia whangshanensis Ching & P.C.Chiu; Acta Bot. Yunnan. 3(3): 299 (1981)
= Lycopodium quasipolytrichoides Hayata; Icon. Pl. Formosan. 5: 252-254, f. 89 (1915)
= Lycopodium reflexointegrum Hayata; Icon. Pl. Formosan. 5: 254-255, f. 90 (1915)
= Urostachys quasipolytrichoides (Hayata) Herter ex Nessel; Bärlappgew. 64-65, Abb. 8, f. 37 (1939)
= Urostachys reflexointeger (Hayata) Herter ex Nessel; Bärlappgew. 64 (1939)
variant :
var. rectifolia
(J.F.Cheng) H.S.Kung & Li Bing Zhang; Acta Phytotax. Sin. 36(6): 528 (1998)
Distribution: China (Jiangxi)
36 CHI
Synonyms: hidden
= Huperzia whangshanensis var. rectifolia J.F.Cheng; Fl. Jiangxi 1: 27, 505, f. 12 (1993)
species :
Huperzia rubicaulis
S.K.Wu & X.Cheng; Acta Phytotax. Sin. 23(5): 400, f. 1 (1985)
Distribution: China (NW-Yunnan)
36 CHI
Remarks: since "rubicaulis" was accepted as original spelling, although in error for "rubricaulis", there is no need for a replacement name.
Synonyms: hidden
= Huperzia wusugongii Li Bing Zhang, X.G.Xu & X.M.Zhou; Phytotaxa 296(1): 98 (2017), nom. nov. superfl.
species :
Huperzia saururoides
(Bory) Rothm.; Feddes Repert. 54: 59 (1944)
Distribution: Argentina (Santa Cruz, Tierra del Fuego); Falkland Isl.
85 ARG 91 FAL
Remarks: selago agg.
Synonyms: hidden
= Lycopodium selago var. saururoides Bory; Fl. Iles Malouin. 26 (1825)
= Lycopodium selago var. saururoides Bory & d´Urv.; Mém. Soc. Linn. Paris 4: 597, pl. 4 (1825 publ. 1826), nom. inval., et ex Baker, Handb. Fern-Allies 9 (1887)
= Lycopodium taxifolium var. saururoides (Bory) Christ ex Nessel; Bärlappgew. 171 (1939)
= Urostachys saururoides (Bory) Herter ex Nessel; Bärlappgew. 30 (1939)
species :
Huperzia selago
(L.) Bernh.; J. Bot. (Schrader) 2: 121-35 (1801) ex Schrank & Mart., Hort. Monac. 3 (1829)
subspecies :
subsp. selago
Name: Tannen-Teufelsklaue (DE), Northern Firmoss, Fir Clubmoss (EN)
Distribution: Iceland; Jan Mayen; Faeroe Isl.; England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, +Berlin, Baden-Württemberg, Bayern, Hessen, +Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, +Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Italy; Malta; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Estonia; Latvia; Lithuania; Belarus; European Russia (N-European Russia, C-European Russia, W-European Russia, E-European Russia); Ukraine; Moldova; Madagascar; La Réunion; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Kamchatka, Khabarovsk, Kuril Isl., Primorye, Sakhalin); Kazakhstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia); China (Jilin, Xinjiang); Mongolia; North Korea; South Korea; Japan (incl. Yakushima Isl.); Australia (Victoria, Tasmania); Alaska; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); St. Pierre et Miquelon; USA (Connecticut, Massachusetts, Maine, Michigan, Minnesota, New Hampshire, New York, Vermont, Wisconsin)
10 DEN FIN FOR GRB ICE IRE JMY NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO FRA SPA 13 ALB BIH BUL CRO ITA KOS MLT MNT NMA ROM SLV SRB 14 BLR EST LAT LIT MLD RUS UKR 29 MDG REU 30 SIB 31 RFE 32 KAZ 33 ARM AZE GEO NCS 34 TUR 36 CHI 37 MON 38 JAP KOR NKO NNS 50 AUS 70 ASK CDN 71 CDN 72 CDN SPM 74 USA 75 USA
Remarks: selago agg.
Synonyms: hidden
= Huperzia apolinari-mariae (Nessel) Holub; Folia Geobot. Phytotax. 20: 70 (1985)
= Huperzia selago f. imbricata (Neilr.) Dostál; Folia Mus. Rerum Nat. Bohemia Occid., Bot. 21 (1984), nom. inval.
= Huperzia selago f. sarmatica (Wolosz.) Dostál; Folia Mus. Rerum Nat. Bohemia Occid., Bot. 21 (1984), nom. inval.
= Huperzia selago f. squarrosa (Bodino) Dostál; Folia Mus. Rerum Nat. Bohemia Occid., Bot. 21 (1984), nom. inval.
= Huperzia selago var. densa Trevis.; Atti Soc. Ital. Sci. Nat. 17(3): 248 (1875)
= Huperzia selago var. patens (P.Beauv.) Trevis.; Atti Soc. Ital. Sci. Nat. 17(3): 249 (1875)
= Lycopodium abietiforme Gray; Nat. Arr. Brit. Pl. 2: 22 (1821 publ. 1822)
= Lycopodium densum Lam.; Fl. franc. 1 (Meth. anal.) (33) (1779)
= Lycopodium disseminatum Dulac; Fl. Hautes-Pyrénées: 37 (1867), nom. illeg.
= Lycopodium selago f. alpinum (G.Lawson) M.Broun; Index No. Amer. Ferns: 112 (1938)
= Lycopodium selago f. patens (P.Beauv.) Clute; Fern Bull. 17: 20 (1909)
= Lycopodium selago f. sylvaticum (G.Lawson) M.Broun; Index No. Amer. Ferns 112 (1938)
= Lycopodium selago L.; Sp. Pl. 2: 1102 (1753)
= Lycopodium selago subsp. patens (P.Beauv.) Calder & Roy L.Taylor; Canad. J. Bot. 43: 1388 (1965)
= Lycopodium selago var. densum (Lam.) Wallr.; Fl. Crypt. Germ. 32 (1831)
= Lycopodium selago var. laxum Desv.; Mém. Soc. Linn. Paris 6, pt. 2: 180 (1827)
= Lycopodium selago var. patens (P.Beauv.) Desv.; Mém. Soc. Linn. Paris 6, pt. 2: 180 (1827)
= Lycopodium selago var. vulgatum Syme & N.E.Br.; (1886), nom. inval.
= Mirmau selago (L.) H.P.Fuchs; Verh. Naturf. Ges. Basel 66: 43-44 (1955)
= Plananthus fastigiatus Opiz; Kratos 1(4): 19 (1819), et Böhm. Gew. 114 (1823)
= Plananthus patens P.Beauv.; Prodr. Aethéogam. 100, 111 (1805)
= Plananthus selago (L.) P.Beauv.; Prod. Aetheogam. 112 (1805)
= Selago selago (L.) Börner; (1912), nom. inval.
= Selago vulgaris Schur; Enum. pl. Transsilv. 825 (1866)
= Urostachys apolinari-mariae Nessel; Repert. Spec. Nov. Regni Veg. 36: 177, t. 170 (1934)
= Urostachys selago (L.) Herter; Bot. Centralbl. 39(2): 249 (1922) (nom. illegit.)
= Urostachys selago var. groenlandicus T.Wulff ex Nessel; Bärlappgew. 39 (1939)
= Urostachys selago var. incurvus (Beutelsp.) Herter ex Nessel; Bärlappgew. 38 (1939)
subspecies :
subsp. appressa
(La Pylaie ex Desv.) D.Löve; Bot. Notiser 109: 164 (1956)
Distribution: Svalbard; Scotland; Norway; N-European Russia; Siberia (Buryatia, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Sakhalin); Northern Caucasus; Georgia [Caucasus]; China (Jilin, Shaanxi, Sichuan, Yunnan); Tibet; Japan; Taiwan; Pakistan (Gilgit); Nepal; India (Sikkim, Uttarakhand, Jammu & Kashmir); Myanmar; Greenland; Canada (Labrador, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Saskatchewan, Yukon); USA (Michigan)
10 GRB NOR SVA 14 RUS 30 SIB 31 RFE 33 GEO NCS 36 CHI TIB 38 JAP TAI 40 IND NEP PAK 41 MYA 70 CDN GNL 71 CDN 72 CDN 75 USA
Remarks: treated by Fraser-Jenkins et al. (2015 et in litt.) and Flora of Great Britain as subspecies of H. selago.
Synonyms: hidden
= Huperzia appressa (La Pylaie ex Desv.) Á.Löve & D.Löve; Bot. Notis. 114: 34 (1961)
= Huperzia arctica (Tolm.) Sipliv.; Novosti Sist. Vyssh. Rast. 10: 347 (1973)
= Huperzia dixitiana P.Mondal & R.K.Ghosh; Fern Gaz. (U. K.) 15(2): 72 (1995) (fide Fraser-Jenkins 1997, 2015, 2017)
= Huperzia petrovii Sipliv.; Novosti Sist. Vyssh. Rast. 10: 346 (1973)
= Huperzia selago f. recurva (Kit. ex Willd.) Dostál; 20 (1984), nom. inval.
= Huperzia selago f. reducta Christ; Boll. Soc. Bot. Ital. 1898: 184 (1898)
= Huperzia selago f. reductum-angustinum Christ; Boll. Soc. Bot. Ital. 1898: 184 (1898)
= Huperzia selago subsp. arctica (Tolm.) Á.Löve & D.Löve; Bot. Notis. (Lund) 114: 35 (1961)
= Huperzia selago var. appressa (La Pylaie ex Desv.) Ching; Acta Bot. Yunnan. 3(3): 303 (1981)
= Huperzia selago var. appressa (La Pylaie ex Desv.) Kukkonen; Hämet-Ahti & Kukkonen 209 (1984), isonym
= Huperzia selago var. sulcata Ching; Acta Bot. Yunnan. 3(3): 303 (1981) (tentative synonym)
= Lycopodium appressum (La Pylaie ex Desv.) Petrov; Fl. Jakutiae 37. (p. p. excl. basionymo & pl., non arct., non F. E. Lloyd & Underw (1930))
= Lycopodium arcticum (Tolm.) Sipliv.; Novost. Sis. Vyssh. Rast. 10: 347 (1973)
= Lycopodium recurvum Kit. ex Willd.; Sp. Pl. ed. 4. 5: 50 (1810)
= Lycopodium selago f. alpestre Berlin; Öfvers. Kongl. Vetensk.-Akad. Förh. 41(7): 83 (1884)
= Lycopodium selago f. appressum (La Pylaie ex Desv.) Gelert; Ostenf., Fl. Arct.: 12 (1902)
= Lycopodium selago f. appressum (Desv.) Zapal.; 86 (1905)
= Lycopodium selago f. recurvum (Kit. ex Willd.) Zapal.; 86 (1905)
= Lycopodium selago subsp. appressum (La Pylaie ex Desv.) Hultén; Ark. Bot. (n. s.) 7(1): 7 (1968) p. p.
= Lycopodium selago subsp. arcticum Grossh. ex Tolm.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk SSSR 20: 39, f. 1-2 (1960)
= Lycopodium selago var. appressum La Pylaie ex Desv.; Mém. Soc. Linn. Paris 6: 180 (1827)
= Lycopodium selago var. recurvum (Kit. ex Willd.) Desv.; Mém. Soc. Linn. Paris 6: 180 (1827)
= Lycopodium selago var. recurvum (Kit. ex Willd.) Syme & N.E.Br.; 12 (1886) nom. illeg.
= Plananthus recurvus (Kit. ex Willd.) Opiz; Kratos 1(4): 19 (1819)
= Urostachys selago f. alpestris Berlin ex Nessel; Bärlappgew. 39 (1939)
= Urostachys selago f. reductus (Christ) Herter; Bärlappgew. 37 (1939)
= Urostachys selago f. reductus-angustinus (Christ) Herter ex Nessel; Bärlappgew. 37 (1939)
= Urostachys selago var. appressus (La Pylaie ex Desv.) Herter ex Nessel; Bärlappgew. 38 (1939)
species :
Huperzia serrata
(Thunb.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
Distribution: Russian Far East (Khabarovsk, Kuril Isl., Primorye, Sakhalin); China (Fujian, Guangdong, Guangxi, Guizhou, Hebei, Heilongjiang, Hubei, Hunan, Jiangxi, Jilin, Liaoning, Shaanxi, Sichuan, Yunnan); Tibet; North Korea; South Korea; Japan; Ryukyu Isl.
31 RFE 36 CHI TIB 38 JAP KOR NKO NNS
Remarks: selago agg. Restricted to E-Asia, the rest of the records belong to H. javanica (Fraser-Jenkins in litt.)
Synonyms: hidden
= Huperzia selago var. serrata (Thunb.) Á.Löve & D.Löve; Univ. Colorado Stud. 17: 5 (1965)
= Huperzia serrata (Thunb.) Rothm.; Feddes Repert. 54(1): 59 (1944), isonym
= Huperzia serrata f. intermedia (Nakai) Ching; Acta Bot. Yunnan. 3(3): 294 (1981)
= Huperzia serrata var. intermedia (Nakai) Satou; Hikobia 12(3): 268 (1997)
= Huperzia serrata var. paucidentata (Sugim.) Satou; Hikobia 12(3): 268 (1997)
= Lycopodium serratum f. intermedium Nakai; Bot. Mag. 39(463): 196-197 (1925)
= Lycopodium serratum Thunb.; Syst. Veg. ed. 14 944 (1784), also Thunb., Fl. Jap. 341, t. 38 (1784)
= Lycopodium serratum var. thunbergii Makino; Bot. Mag. (Tokyo) 12: 12 (1898), nom. superfl.
= Plananthus serratus (Thunb.) P.Beauv.; Prodr. Aethéogam. 100, 111 (1805)
= Urostachys sachalinensis (Nessel) Herter; Est. Bot. Reg. Uruguaya 20 [Index Lycopod.]: 80 (1949)
= Urostachys serratus (Thunb.) Herter; Bot. Arch. 3: 13 (1923)
= Urostachys serratus var. sachalinensis Nessel; Repert. Spec. Nov. Regni Veg. 36: 179, t. 171 (1934)
species :
Huperzia shresthae
Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 407 (2018)
Distribution: China (Zhejiang, Fujian, Guangxi); India (West Bengal); Myanmar (Kachin)
36 CHI 40 IND 41 MYA
Remarks: selago agg.
species :
Huperzia somae
(Hayata) Ching; Acta Bot. Yunnan. 3: 301 (1981) (somai auct.)
Distribution: Japan; Taiwan; Philippines (N-Luzon)
38 JAP TAI 42 PHI
Remarks: selago agg.
Synonyms: hidden
= Lycopodium chinense var. somae (Hayata) Masam.; Mem. Fac. Sci. Agric. Taihoku Univ. II (Bot. no. 4): 114 (1934)
= Lycopodium selago var. somae (Hayata) Masam.; Short Fl. Formos. 32 (1936)
= Lycopodium somae Hayata; Icon. Pl. Formosan. 5: 255, f. 91 (1915)
= Urostachys somae (Hayata) Herter ex Nessel; Bärlappgew. 54 (1939)
species :
Huperzia sprengeri
(Nessel) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distribution: Society Isl. (Tahiti)
61 SCI
Remarks: selago agg.
Synonyms: hidden
= Urostachys sprengeri Nessel; Repert. Spec. Nov. Regni Veg. 36: 179, t. 171 (1934)
species :
Huperzia suberecta
(Lowe) Tardieu; Adansonia 10: 20 (1970)
Distribution: Azores (Sao Miguel Isl., Terceira, Sao Jorge, Pico, Faial, Flores Isl.); Madeira (Madeira Isl.)
21 AZO MDR
Remarks: bona sp. fide Fernandez-Prieto et al. 2008
Synonyms: hidden
= Huperzia selago subsp. suberecta (Lowe) Franco & Vasc.; Bol. Soc. Brot. Ser. 2, 41: 23 (1967)
= Lycopodium selago subsp. suberectum (Lowe) Romariz; Rev. Fac. Cienc. Lisboa ser. 2, C. 3: 3: 71, t. 1, f. 3 (1953)
= Lycopodium selago var. suberectum (Lowe) C.Chr.; Dansk Bot. Ark. 7: 188 (1932)
= Lycopodium suberectum Lowe; Trans. Cambridge Philos. Soc. 4(1): 9 (1830)
= Urostachys suberectus (Lowe) Herter ex Nessel; Bärlappgew. 41 (1939)
species :
Huperzia subintegra
(Hillebr.) Beitel & W.H.Wagner; Contr. Univ. Michigan Herb. 22: 140 (1999)
Name: Kauai clubmoss
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Lycopodium serratum var. subintegrum Hillebr.; Fl. Hawaiian Isl. [Hillebr.] 643 (1888)
species :
Huperzia sumatrana
(Alderw.) Holub; Folia Geobot. Phytotax. 20: 77 (1985)
Distribution: Sumatra; New Guinea (Irian Jaya, Papua New Guinea)
42 NWG SUM
Remarks: selago agg.
Synonyms: hidden
= Lycopodium sumatranum Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 44 (1918)
= Urostachys sumatranus (Alderw.) Herter ex Nessel; Bärlappgew. 66 (1939)
species :
Huperzia sutchueniana
(Herter) Ching; Acta Bot. Yunnan. 3: 297 (1981)
Distribution: China (Anhui, Chongqing, Guangdong, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Zhejiang)
36 CHI
Remarks: selago agg.
Synonyms: hidden
= Huperzia minimadenta J.F.Cheng; Bull. Bot. Res. North-East. Forest. Inst. 7(3): 113 (1987)
= Lycopodium serratum var. alpestre Christ; Bull. Acad. Int. Geogr. Bot. 16(199-201): 141 (1906)
= Lycopodium sutchuenianum Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Urostachys sutchuenianus (Herter) Herter ex Nessel; Bärlappgew. 53 (1939)
species :
Huperzia tibetica
(Ching) Ching; Acta Bot. Yunnan. 3: 304 (1981), et Fl. Xizangica 1: 7 (1983)
Distribution: China (NW-Yunnan)
36 CHI
Remarks: selago agg.
Synonyms: hidden
= Lycopodium tibeticum Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 17 (1940)
species :
Huperzia yakusimensis
(Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distribution: Japan (Yakushima Isl.)
38 JAP NNS
Remarks: selago agg.; possibly a synonym of H. somae
Synonyms: hidden
= Lycopodium yakusimense Herter ex Nessel; Bärlappgew. 65 (1939) (nom. inval.), Revista Sudamer. Bot. 6: 160, t. 8, f. 27 (1940)
species :
Huperzia zollingeri
(Herter) Holub; Folia Geobot. Phytotax. 20: 78 (1985)
Distribution: Sumatra; Java; New Guinea (Irian Jaya, Papua New Guinea)
42 JAW NWG SUM
Remarks: selago agg.
Synonyms: hidden
= Huperzia monticola (Alderw.) Holub; Folia Geobot. Phytotax. 20: 75 (1985)
= Lycopodium monticola Alderw.; Malayan Ferns Fern Allies Suppl 1(Corr.): 65 (1917) (Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 43, t. 7 (1918))
= Lycopodium zollingeri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 48 (1909)
= Urostachys monticola (Alderw.) Herter ex Nessel; Bärlappgew. 120 (1939)
= Urostachys zollingeri (Herter) Herter ex Nessel; Bärlappgew. 120 (1939)
species :
Huperzia ×bartleyi
(Cusick) Holub; Folia Geobot. Phytotax. 26(1): 92 (1991)
Distribution: USA (Arkansas, Iowa, Illinois, Kentucky, Missouri, North Carolina, New York, Ohio, South Carolina, Wisconsin)
74 USA 75 USA 78 USA
Remarks: selago agg., Huperzia lucidula x porophila
Synonyms: hidden
= Lycopodium ×bartleyi Cusick; Amer. Fern J. 77(3): 100 (1987)
species :
Huperzia ×buttersii
(Abbe) Kartesz & Gandhi; Phytologia 70: 201 (1991)
Distribution: Canada (Newfoundland, Ontario); USA (Maine, Michigan, Minnesota, New Hampshire, Ohio, Vermont, Wisconsin)
72 CDN 74 USA 75 USA
Remarks: selago agg., Huperzia lucidula x selago
Synonyms: hidden
= Lycopodium ×buttersii Abbe; Rhodora 55(651): 91-92 (1953)
species :
Huperzia ×carlquistii
Beitel & W.H.Wagner; Contr. Univ. Michigan Herb. 22: 142 (1999)
Distribution: Hawaii (Maui)
63 HAW
Remarks: Huperzia erubescens x Huperzia subintegra
species :
Huperzia ×gillettii
Beitel & W.H.Wagner; Contr. Univ. Michigan Herb. 22: 142 (1999)
Distribution: Hawaii (Maui, Kauai, Oahu, Molokai, Lanai, Hawaii Isl.)
63 HAW
Remarks: Huperzia serrata x Huperzia subintegra
species :
Huperzia ×josephbeitelii
A.Haines; Fam. Huperziac. Lycopodiac. New England ed. 2: 100 (2003)
Distribution: Canada (Labrador, Newfoundland, Nova Scotia, Ontario, Québec); St. Pierre et Miquelon
72 CDN SPM
Remarks: Huperzia selago x appressa
Synonyms: hidden
= Huperzia ×beitelii A.Haines; Fam. Huperziac. Lycopodiac. New England 86 (-87) (2003), non B. Øllg. (1990)
species :
Huperzia ×medeirosii
Beitel & W.H.Wagner; Contr. Univ. Michigan Herb. 22: 142 (1999)
Distribution: Hawaii (Maui)
63 HAW
Remarks: Huperzia haleakalae x Huperzia subintegra
species :
Huperzia ×protoporophila
A.Haines; Fam. Huperziac. Lycopodiac. New England 88 (2003)
Distribution: USA (Maine)
75 USA
Remarks: Huperzia lucidula x appressa
species :
Huperzia ×sulcinervia
(Spring) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 248 (1875)
Distribution: Hawaii (Oahu, Lanai, Hawaii Isl.)
63 HAW
Remarks: Huperzia erubescens x Huperzia serrata
Synonyms: hidden
= Huperzia sulcinervia (Spring) Rothm.; Feddes Repert. 54: 59 (1951)
= Lycopodium sulcinervium Spring; Bull. Acad. Roy. Soc. Bruxelles 8: 514 (1841)
= Lycopodium ×helleri Herter; Bot. Jahrb. Syst. 43: Beibl. 98: 43 (1909)
= Huperzia ×helleri (Herter) Kartesz & Gandhi; Phytologia 70(3): 201 (1991)
= Urostachys helleri (Herter) Herter ex Nessel; Bärlappgew. 53 (1939)
= Urostachys sulcinervius (Spring) Herter ex Nessel; Bärlappgew. 54 (1939)
