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519.0000 Family Lamiaceae Martinov; Tekhno-Bot. Slovar 355 (1820), nom. cons., nom. alt.
Latest phylogeny: Zhao et al. 2021
Species listed in database: 7866; hybrids: 382
= Aegiphilaceae Raf.; Sylva Tellur. 161 (1838)
= Ajugaceae Döll; Rhein. Fl. [Döll] 375 (1843)
= Chloanthaceae Hutch.; Fam. Fl. Pl., Dicot., ed. 2: 396 (1959)
= Glechomaceae Martinov; Tekhno-Bot. Slovar 288 (1820)
= Labiatae Juss.; Gen. Pl. [Jussieu] 110 (1789), nom. cons.
= Melissaceae Bercht. & J.Presl; Přir. Rostlin 245 (1820)
= Melittidaceae Martinov; Tekhno-Bot. Slovar 390 (1820)
= Menthaceae Burnett; Outlines Bot. [Burnett] 969, 1095, 1106 (1835)
= Monardaceae Döll; Fl. Baden [Döll] 2: 661 (1858)
= Nepetaceae Bercht. & J.Presl; Přir. Rostlin 245 (1820)
= Salazariaceae F.A.Barkley; Phytologia 32: 304 (1975)
= Salviaceae Bercht. & J.Presl; Přir. Rostlin 245 (1820)
= Saturejaceae Döll; Fl. Baden [Döll] 2: 664. med (1858)
= Scutellariaceae Döll; Rhein. Fl. [Döll] 373 (1843)
= Siphonanthaceae Raf.; Fl. Tellur. 4: 87 (1838)
= Stachydaceae Döll; Rhein. Fl. [Döll] 366 (1843)
= Symphoremataceae Wight; Icon. Pl. Ind. Orient. [Wight] 4(3): 13 (1849)
= Viticaceae Juss.; Gen. Pl. [Jussieu] 106 (1789)
519.2263 Genus Hypenia (Mart. ex Benth.) Harley; Bot. J. Linn. Soc. 98: 91 (1988)
Distribution: Tropical S-America, mostly Brazil
Species listed in database: 29
species :
Hypenia aristulata (Epling) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis aristulata Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 231 (1936)
species :
Hypenia brachystachys (Pohl ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: WC-Brazil (Goiás, Brasilia Distrito Federal); SE-Brazil
84 BRA
Synonyms: hidden
= Hyptis brachystachys Pohl ex Benth.; Labiat. Gen. Spec. [Bentham] 137 (1833)
= Mesosphaerum brachystachyum (Pohl ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia calycina (Pohl ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis calycina Pohl ex Benth.; Labiat. Gen. Spec. [Bentham] 140 (1833)
= Mesosphaerum calycinum (Pohl ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia concinna (Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: N-Brazil (Tocantins)
84 BRA
Synonyms: hidden
= Hyptis concinna Benth.; Prodr. [A.P. de Candolle] 12: 136 (1848)
= Mesosphaerum concinnum (Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia crispata (Pohl ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis crispata Pohl ex Benth.; Labiat. Gen. Spec. [Bentham] 139 (1833)
= Mesosphaerum crispatum (Pohl ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia densiflora (Pohl ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: N-Brazil (Tocantins); WC-Brazil (Goiás, Brasilia Distrito Federal); SE-Brazil
84 BRA
Synonyms: hidden
= Hyptis densiflora Pohl ex Benth.; Labiat. Gen. Spec. [Bentham] 187 (1833)
= Hyptis densiflora var. dolichodon Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 228 (1936)
= Hyptis melochioides A.St.-Hil. ex Benth.; Labiat. Gen. Spec. [Bentham] 137 (1833)
= Mesosphaerum densiflorum (Pohl ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
= Mesosphaerum melochioides (A.St.-Hil. ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia durifolia (Epling) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hypenia sclerophylla (Epling) Harley & J.F.B.Pastore; Phytotaxa 58: 24 (2012)
= Hyptis durifolia Epling; Revista Mus. La Plata, Secc. Bot. 7: 235 (1949)
= Hyptis sclerophylla Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 234 (1936)
species :
Hypenia filicifolia Antar, Faria & A.Soares; Nordic J. Bot. 2025(12)-e04917: 7 (2025)
Distribution: Brazil (Brasília Distrito Federal)
84 BRA
species :
Hypenia gardneriana (Benth.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis gardneriana (Benth.) Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 233 (1936)
= Hyptis glauca var. gardneriana Benth.; Prodr. [A.P. de Candolle] 12: 140 (1848)
species :
Hypenia glauca (A.St.-Hil. ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: S-Brazil (Paraná); Paraguay (Amambay, Canindeyu)
84 BRA 85 PAR
Synonyms: hidden
= Hyptis glauca A.St.-Hil. ex Benth.; Labiat. Gen. Spec. [Bentham] 141 (1833)
= Hyptis glauca var. amplifrons Briq.; Bull. Herb. Boissier, Sér. 2, 7: 613 (1907)
= Mesosphaerum glaucum (A.St.-Hil. ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia hypoleuca Zavatin & Harley; Phytotaxa 682(1): 82 (2025)
Distribution: Brazil (Minas Gerais)
84 BRA
species :
Hypenia inelegans (Epling) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis inelegans Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 226 (1936)
species :
Hypenia irregularis (Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: NE-Brazil (Bahia); WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis irregularis Benth.; Prodr. [A.P. de Candolle] 12: 137 (1848)
= Mesosphaerum irregulare (Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia kalunga A.Soares, Harley & Antar; Brittonia 77(2): 208 (2025)
Distribution: WC-Brazil (Goiás)
84 BRA
species :
Hypenia machrisae (Epling) Harley & J.F.B.Pastore; Syst. Bot. 46: 1088 (2021)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Eriope machrisae (Epling) Harley; Bot. J. Linn. Soc. 98(2): 93 (1988)
= Hyptis machrisae Epling; Contr. Sci. Nat. Hist. Mus. Los Angeles County 6: 1 (1957)
species :
Hypenia macrantha (A.St.-Hil. ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: SE-Brazil (Minas Gerais)
84 BRA
Synonyms: hidden
= Hyptis macrantha A.St.-Hil. ex Benth.; Labiat. Gen. Spec. [Bentham] 140 (1833)
= Mesosphaerum macranthum (A.St.-Hil. ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia macrosiphon (Briq.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: Bolivia (Santa Cruz); N-Brazil (Rondônia); WC-Brazil (Mato Grosso, Goiás)
83 BOL 84 BRA
Synonyms: hidden
= Hyptis lindmaniana Briq.; Ark. Bot. 2(10): 4 (1904)
= Hyptis macrosiphon Briq.; Bull. Herb. Boissier 4: 785 (1896)
= Hyptis mattogrossensis Pilg.; Bot. Jahrb. Syst. 30: 191 (1901)
= Mesosphaerum lindmanianum Briq.; Ark. Bot. 2(10): 4 (1904)
= Mesosphaerum macrosiphon Briq.; Bull. Herb. Boissier 4: 785 (1896)
species :
Hypenia marifolia (Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis marifolia Benth.; Prodr. [A.P. de Candolle] 12: 186 (1848)
= Mesosphaerum marifolium (Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia micrantha (Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: WC-Brazil (Mato Grosso)
84 BRA
Synonyms: hidden
= Eriope micrantha Benth.; Prodr. [A.P. de Candolle] 12: 141 (1848)
= Hyptis effusa S.Moore; Trans. Linn. Soc. London, Bot., Ser. 2, 4: 441 (1894)
species :
Hypenia paniculata (Benth.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: SE-Brazil (Minas Gerais)
84 BRA
Synonyms: hidden
= Hyptis paniculata Benth.; Labiat. Gen. Spec. [Bentham] 139 (1833)
= Mesosphaerum paniculatum (Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
species :
Hypenia paradisi (Harley) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis paradisi Harley; Kew Bull. 29(1): 126 (1974)
species :
Hypenia pauliana (Epling) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: SE-Brazil (São Paulo)
84 BRA
Synonyms: hidden
= Hyptis pauliana Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 232 (1936)
species :
Hypenia perplexa (Epling) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: SE-Brazil (Minas Gerais)
84 BRA
Synonyms: hidden
= Hyptis perplexa Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 231 (1936)
species :
Hypenia pruinosa (Pohl ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis pruinosa Pohl ex Benth.; Labiat. Gen. Spec. [Bentham] 137 (1833)
= Mesosphaerum pruinosum (Pohl ex Benth.) Kuntze; Revis. Gen. Pl. 2: 527 (1891)
species :
Hypenia reticulata (Mart. ex Benth.) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: Bolivia (Santa Cruz); N-Brazil (Tocantins); NE-Brazil; WC-Brazil (Goiás, Brasilia Distrito Federal); SE-Brazil (Minas Gerais, São Paulo); S-Brazil (Paraná)
83 BOL 84 BRA
Synonyms: hidden
= Hyptis laxiflora Mart. ex Benth.; Labiat. Gen. Spec. [Bentham] 139 (1833)
= Hyptis longiflora Pohl ex Benth.; Labiat. Gen. Spec. [Bentham] 140 (1833)
= Hyptis reticulata Mart. ex Benth.; Labiat. Gen. Spec. [Bentham] 141 (1833)
= Hyptis reticulata var. longiflora Mart. ex Benth.; Labiat. Gen. Spec. [Bentham] 141 (1833)
= Mesosphaerum laxiflorum (Mart. ex Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
= Mesosphaerum longiflorum (Pohl ex Benth.) Kuntze; Revis. Gen. Pl. 2: 525 (1891)
species :
Hypenia salzmannii (Benth.) Harley; Bot. J. Linn. Soc. 98(2): 91 (1988)
Distribution: Venezuela (Anzoátegui, Bolívar, Monagas); Guyana; NE-Brazil (Piauí, Ceará, Paraíba, Pernambuco, Bahia, Sergipe); SE-Brazil (Minas Gerais)
82 GUY VEN 84 BRA
Synonyms: hidden
= Hyptis salzmannii Benth.; Labiat. Gen. Spec. [Bentham] 138 (1833)
= Hyptis salzmannii var. filipes Benth.; Prodr. [A.P. de Candolle] 12: 137 (1848)
= Hyptis salzmannii var. tomentosa Benth.; Labiat. Gen. Spec. [Bentham] 138 (1833)
= Mesosphaerum salzmannii (Benth.) Kuntze; Revis. Gen. Pl. 2: 527 (1891)
species :
Hypenia simplex (A.St.-Hil. ex Benth.) Harley & J.F.B.Pastore; Phytotaxa 58: 24 (2012)
Distribution: N-Brazil (Tocantins); WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Eriope simplex (A.St.-Hil. ex Benth.) Harley; Bot. J. Linn. Soc. 98: 93 (1988)
= Hyptis campanulata Benth.; Prodr. [A.P. de Candolle] 12: 137 (1848)
= Hyptis simplex A.St.-Hil. ex Benth.; Labiat. Gen. Spec. [Bentham] 138 (1833)
= Mesosphaerum campanulatum (Benth.) Kuntze; Revis. Gen. Pl. 2: 526 (1891)
= Mesosphaerum simplex (A.St.-Hil. ex Benth.) Kuntze; Revis. Gen. Pl. 2: 527 (1891)
species :
Hypenia subrosea (Harley) Harley; Bot. J. Linn. Soc. 98(2): 92 (1988)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Hyptis subrosea Harley; Kew Bull. 29(1): 128 (1974)
species :
Hypenia violacea Mart.Gord. & S.Valencia; Brittonia 61(1): 67 (2009)
Distribution: Mexico (Guerrero, Oaxaca)
79 MEX
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A complete, synonymic checklist of the Higher Plants of the World


Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life