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041.0000 Family Polypodiaceae J. & C.Presl; Delic. Prag. 159 (1822)
67 genera, ca. 1700 spp. Phylogeny: Wei & Zhang 2022
Species listed in database: 1750; hybrids: 50
= Platyceriaceae Ching; Acta Phytotax. Sin. 16(3): 18 (1978)
= Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
= Grammitidaceae Newman; Hist. Brit. Ferns 7 (1840) (Grammitaceae auct., missp.)
= Gymnogrammitidaceae Ching; Acta Phytotax. Sin. 11: 12 (1966)
= Loxogrammaceae Ching ex Pic.Serm.; Webbia 29: 11 (1975)
= Platyceriaceae Ching; Sunyatsenia 5: ? (1940)
= Pleurosoriopsidaceae Sa.Kurata & Ikebe ex Ching; Acta Phytotax. Sin. 16(4): 17 (1978)
041.3402 Genus Leptochilus Kaulf.; Enum. Filic. [Kaulfuss] 147 (1824)
Distribution: E- and tropical Asia, 3 to Queensland. Phylogeny and species circumscription: Zhang et al. (2024)
Species listed in database: 54; hybrids: 2
= Dendroglossa C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 149 (1851)
= Gymnopteris C.Presl; Tent. Pterid. 244 (1836)
= Paraleptochilus Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 198 (1947)
= Colysis C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 146 (1851) (fide Fraser-Jenkins 2008 and others)
= Selliguea Blume; (1826) (nec Bory (1824))
= Nistarika B.K.Nayar, Madhus. & Molly; Fern Gaz. 13: 33 (1985)
= Kontumia S.K.Wu & P.K.Lôc; Novon 15(1): 245 (2005) (fide PPG-I)
species :
Leptochilus axillaris (Cav.) Kaulf.; Enum. Filic. [Kaulfuss] 147, t. 1, f. 10 (1824)
Distribution: China (Guizhou, Yunnan); India (Maharashtra, Assam State, S-India, Bengal); Bangladesh; Nicobar Isl.; N-Myanmar; Thailand; Vietnam; peninsular Malaysia (Perak); Sumatra; Borneo (Sabah); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok); Moluccas; New Guinea (Papua New Guinea)
36 CHI 40 BAN IND 41 MYA NCB THA VIE 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM
Remarks: Colysis clade, Leptochilus subclade
Synonyms: hidden
= Acrostichum axillare Cav.; Anales Hist. Nat. 1: 101 (1799)
= Colysis axillaris (Cav.) Rajkumar & Manickam; Indian Fern J. 18(1-2): 125 (2001)
= Colysis bahupunctika (B.K.Nayar, Madhus. & Molly) Rajkumar & Manickam; Indian Fern J. 18(1-2): 125 (2001) (fide Fraser-Jenkins 2008)
= Gymnogramma axillaris (Cav.) Ces.; Atti Reale Accad. Sci. Napoli 7(8): 29 (1876)
= Gymnopteris axillaris (Cav.) C.Presl; Tent. Pterid. 244, t. 11, f. 4-5 (1836)
= Gymnopteris variabilis var. axillaris (Cav.) Bedd.; Ferns Brit. India, t. 271 (1968)
= Leptochilus bahupunctikus (B.K.Nayar, Madhus. & Molly) B.K.Nayar, Madhus. & Molly; Fern Fl. S. India 59 (1998)
= Leptochilus platyphyllus Copel.; Philipp. J. Sci. 37: 340, t. 2 (1928)
= Lomaria serpens Wall.; Numer. List [Wallich] no. 32 (1829), nomen
= Nistarika bahupunctika B.K.Nayar, Madhus. & Molly; Fern Gaz. 13(1): 33 (1985) (fide Fraser-Jenkins 2008)
= Polypodium hymenodes Wall.; Numer. List [Wallich] no. 283 (1829), nomen
= Polypodium leptochiloides Kuhn; Forschungsr. Gazelle, Bot. Alg. 4, Farne 10 (1889)
species :
Leptochilus bolikhamsaiensis Liang Zhang, Thepkaysone & Z.Zhou; Phytotaxa 659(3): 217 (2024)
Distribution: Laos
41 LAO
Remarks: related to L. pentaphyllus fide molecular data
species :
Leptochilus bolsteri (Copel.) Parris; Fern Gaz. 20(7): 305 (2018)
Distribution: Borneo (Sarawak); Philippines (Mindanao)
42 BOR PHI
Remarks: often included in L. macrophyllus
Synonyms: hidden
= Colysis bolsteri (Copel.) Copel.; Fern Fl. Philipp. 3: 489 (1960)
= Pleopeltis bolsteri (Copel.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 11 (1909)
= Polypodium bolsteri Copel.; Philipp. J. Sci. 1, Suppl. 4: 257, t. 4 A (1906)
species :
Leptochilus brevipes Liang Zhang, Z.L.Liang & Zhi Y.Yu; Phytotaxa 634(2): 144 (2024)
Distribution: China (Yunnan)
36 CHI
Remarks: Colysis clade, Yunnan subclade
species :
Leptochilus cantoniensis (Baker) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 343 (1933)
Distribution: China (Guangdong, Hainan); Vietnam
36 CHI 41 VIE
Remarks: ellipticus clade, cantoniensis subclade
Synonyms: hidden
= Campium cantoniense (Baker) Ching; Sinensia 1: 53 (1930)
= Campium dilatatum Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Christopteris cantoniensis (Baker) Christ; J. Bot. [Morot] 21: 273 (1908)
= Dendroglossa cantoniensis (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 199 (1947)
= Drymoglossum cordatum Christ; Notul. Syst. (Paris) 1: 375 (1911)
= Gymnogramma cantoniensis (Baker) Baker; Hooker´s Icon. Pl. t. 1685 (1887)
= Leptochilus cordatus (Christ) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 343 (1933)
= Myuropteris cordata (Christ) C.Chr.; Dansk Bot. Ark. 6(3): 73, t. 9, f. 1-2, t. 10, f. 3 (1929)
= Polypodium cantoniense (Baker) Baker; J. Bot. 17: 304 (1879)
species :
Leptochilus chilangensis (V.N.Tu) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 172 (2018)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Colysis chilangensis V.N.Tu; Bot. Zhurn. (Moscow & Leningrad) 65(4): 582 (1980)
species :
Leptochilus chingii Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018), nom. nov.
Distribution: China
36 CHI
Remarks: ellipticus clade, henryi subclade
Synonyms: hidden
= Colysis bonii Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
species :
Leptochilus daklakensis Liang Zhang, X.M.Zhou, T.T.Luong & Li Bing Zhang; Phytotaxa 659(3): 219 (2024)
Distribution: Vietnam
41 VIE
Remarks: ellipticus clade, henryi subclade
species :
Leptochilus decurrens Blume; Enum. Pl. Javae 2: 206 (1828)
Distribution: China (Guangxi, Guizhou, Hainan, Hongkong, Yunnan); Japan; Ogasawara Isl. (Hahajima); Taiwan; Nepal; Bhutan; India (Sikkim, E-India, C-India, Cachar, S-India, Arunachal Pradesh, Sikkim, Darjeeling); Sri Lanka; Myanmar; Thailand; Laos; ?Cambodia; Vietnam; peninsular Malaysia (Perak, Pahang); Sumatra; Borneo (Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Bali, Lombok); Moluccas; Christmas Isl. [Austr.]; New Guinea (Papua New Guinea)
36 CHI 38 JAP TAI 40 BHU IND NEP SRL 41 LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI SUL SUM XMS
Remarks: Colysis clade, Paraleptochilus subclade. Leptochilus lanceolatus is included here by Flora of China but a separate Indian species fide Fraser-Jenkins (2008). Inclusion of L. metallicus, L. thwaitesianus and L. wallii fide Handbook of Ceylon (2006)
Synonyms: hidden
= Acrostichum decurrens Wall.; Numer. List [Wallich] no. 18 (1829), nomen
= Acrostichum decurrens Wall. ex Thwaites; Enum. Pl. Zeyl. [Thwaites] 381 (1864)
= Acrostichum feejanum Ettingsh.; Denkschr. Kaiserl. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 22: 54 (1864)
= Acrostichum listeri Baker; J. Linn. Soc., Bot. 26: 361 (1889)
= Acrostichum rivulare Wall.; Numer. List [Wallich] no. 2165 (1829), nomen
= Acrostichum variabile Hook.; Sp. Fil. [W.J.Hooker] 5: 277 (1864)
= Acrostichum wallii Baker; J. Bot. 10: 146 (1872)
= Anapausia decurrens (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 186 (1851)
= Campium decurrens (Blume) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium metallicum (Bedd.) Copel.; Philipp. J. Sci. 37: 347 (1928)
= Campium wallii (Baker) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Colysis decurrens (Blume) Panigrahi; Abstr. & Souv. Nation. Symp. Curr. Trends Pterid. Palayamkottai 13 (1991)
= Colysis diversifolia W.M.Chu; Acta Bot. Yunnan. 1(2): 93 (1979)
= Colysis metallica (Bedd.) Rajkumar & Manickam; Indian Fern J. 18(1-2): 126 (2001)
= Colysis ×beddomei Manickam & Irudayaraj; Taxon 46: 267 (1997) (fide Kandel & Fraser-Jenkins 2020)
= Dendroglossa metallica (Bedd.) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 100 (1974)
= Dendroglossa wallii (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 200 (1947)
= Gymnogramma decurrens (Blume) Hook.; Sp. Fil. [W.J.Hooker] 6: 161 (1864)
= Gymnopteris decurrens (Blume) T.Moore; Index Fil. [T. Moore] 29 (1857)
= Gymnopteris dichotomophlebia Hayata; Icon. Pl. Formosan. 4: 201, f. 136 (1914)
= Gymnopteris feei f. anomala Bedd.; Ferns Brit. India, t. 274 (1868)
= Gymnopteris feei T.Moore; Index Fil. [T. Moore] 30 (1857)
= Gymnopteris metallica Bedd.; Ferns Brit. India Suppl. 26, t. 390 (1876)
= Gymnopteris obtusifolia C.Presl; Tent. Pterid. 244 (1836)
= Gymnopteris variabilis (Hook.) Bedd.; Ferns Brit. India, t. 272 (1868)
= Gymnopteris wallichiana C.Presl; Tent. Pterid. 244 (1836), nom. nud.
= Gymnopteris wallii (Baker) Bedd.; Ferns Brit. India Suppl. 26, t. 389 (1876)
= Leptochilus dichotomophlebia Hayata; Icon. Pl. Formosan. 4: 202 (1914), pro syn.
= Leptochilus listeri (Baker) C.Chr.; Index Filic. 12 (1905), 386 (1906)
= Leptochilus metallicus (Bedd.) C.Chr.; Index Filic. 386 (1906)
= Leptochilus obtusifolius (C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 185 (1851)
= Leptochilus trifidus Alderw.; Bull. Dépt. Agric. Indes Néerl. 18: 26 (1908) (probably a hybrid L. decurrens x macrophyllus)
= Leptochilus wallii (Baker) C.Chr.; Index Filic. 20 (1905), 388 (1906)
= Leptochilus ×beddomei (Manickam & Irudayaraj) X.C.Zhang & Noot.; Fl. China 2-3: 834 (2013)
= Paraleptochilus decurrens (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 198 (1947)
= Paraleptochilus metallicus (Bedd.) Á.Löve & D.Löve; Taxon 26(2-3): 324 (1977)
= Selliguea decurrens (Blume) C.Presl; Tent. Pterid. 216 (1836)
species :
Leptochilus digitatus (Baker) Noot.; Blumea 42: 282 (1997)
Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Colysis digitata (Baker) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 328 (1933)
= Colysis digitata f. annamensis (Christ) Ching
= Colysis digitata f. cadieri (Christ) Ching
= Colysis digitata f. laciniata Ching
= Colysis tridactylis Fée; Mém. Foug. 5 (Gen. Filic.) 176 (1852), nomen
= Colysis triphylla Ching; Acta Phytotax. Sin. 8: 170, t. 23(31) (1959)
= Grammitis finlaysoniana Wall.; Numer. List [Wallich] no. 248 (1829), nomen
= Gymnogramma digitata Baker; J. Bot. 28: 267 (1890)
= Gymnogramma finlaysoniana (Wall.) Baker; Ann. Bot. (Oxford) 6: 486 (1891)
= Phymatodes digitata (Baker) Ching; Contr. Inst. Bot. Natl. Acad. Peiping 2: 77 (1933)
= Polypodium ampelideum Christ; J. Bot. [Morot] 19: 78 (1905)
= Polypodium annamense Christ; J. Bot. [Morot] 19: 77 (1905)
= Polypodium cadieri Christ; J. Bot. [Morot] 19: 76 (1905)
= Polypodium digitatum (Baker) C.Chr.; Index Filic. 522 (1906)
= Polypodium podopterum Christ; J. Bot. [Morot] 19: 125 (1905)
= Selliguea finlaysoniana Wall. ex T.Moore; Index Fil. [T. Moore] 64 (1857), nomen
species :
Leptochilus dissimilialatus (Bonap.) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018)
Distribution: Vietnam; etc.
41 VIE
Remarks: frequently included in L. ellipticus s. l.
Synonyms: hidden
= Colysis dissimilialata (Bonap.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Polypodium dissimilialatum Bonap.; Notes Ptéridol. 14: 155 (1924)
species :
Leptochilus dolichophyllus H.H.Fu & H.J.Wei; Ecol. Evol. 15(10)-e72293: 9 (2025)
Distribution: China (Fujian)
36 CHI
species :
Leptochilus dolichopterus (Copel.) Fraser-Jenk. & Amoroso; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008)
Distribution: Philippines
42 PHI
Synonyms: hidden
= Pleopeltis dolichoptera (Copel.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 9 (1909)
= Polypodium dolichopterum Copel.; Philipp. J. Sci. 1, Suppl. 2: 162 (1906)
species :
Leptochilus ellipticus (Thunb.) Noot.; Blumea 42: 283 (1997)
Distribution: China (Anhui, Guangxi, Guizhou, Hainan, Hongkong, Hunan, Jiangsu, Jiangxi, Yunnan, Zhejiang); Tibet; South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; Nepal; Bhutan; NE-India (Bengal, Arunachal Pradesh, Manipur, Meghalaya, Sikkim, Darjeeling, Himachal Pradesh); Myanmar; Thailand; Laos; Vietnam; Philippines (Luzon, Leyte, Mindanao, Samar)
36 CHI TIB 38 JAP KOR NNS TAI 40 BHU IND NEP 41 LAO MYA THA VIE 42 PHI
Remarks: Fraser-Jenkins et al. (2015) accepts no varieties, but distinguishes between this taxon and L. pothifolius
Synonyms: hidden
= Colysis boisii (Christ) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Colysis decurrens (Wall. ex Hook. & Grev.) Nakaike; New Fl. Jap. Pterid. 13 (1991) [fide Zimmer, in litt.]
= Colysis elegans Sa.Kurata; J. Geobot. 12: 41 (1963)
= Colysis elliptica (Thunb.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 333 (1933)
= Colysis elliptica var. longipes (Ching) L.Shi & X.C.Zhang; Acta Phytotax. Sin. 37(1): 76 (1999)
= Colysis flavescens (Ching) Nakaike, S.Matsumoto & V.L.Gurung; Cryptog. Himalayas 2 C. & E. 189 (1990)
= Colysis flavescens (Ching) S.R.Ghosh; Pterid. Fl. E. India 497 (2004), isonym
= Colysis leptophylla H.Itô; Bot. Mag. (Tokyo) 53: 68 (1939)
= Colysis longipes Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933), nom. illeg., non Polypodium longipes auct.
= Colysis pothifolia f. bipinnatifida H.Itô; J. Jap. Bot. 11: 89 (1935)
= Colysis pothifolia var. membranacea Nakai; Bull. Natl. Sci. Mus., Tokyo, 27: 23 (1949)
= Colysis ×kiusiana Sa.Kurata; J. Geobot. 10: 37 (1961)
= Grammitis decurrens Wall. ex Hook. & Grev.; Icon. Filic. tab. 6 (1827) [fide Zimmer, in litt.]
= Gymnogramma elliptica (Thunb.) Hook. & Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 2: 389 (1874) (pro parte)
= Gymnogramma elliptica C.B.Clarke; Trans. Linn. Soc. London, Bot., Ser. 2, 1: 570 (1880) (pro parte)
= Gymnogramma elliptica var. furcans Tutcher; J. Linn. Soc., Bot. 37: 69 (1904)
= Gymnogramma pothifolia (Buch.-Ham. ex D.Don) Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 39 (1827)
= Leptochilus elegans (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan 35 (2013)
= Leptochilus ellipticus var. longipes (Ching) Noot.; Fl. China 2-3: 837 (2013)
= Leptochilus ×kiusianus (Sa.Kurata) Nakaike; Checkl. Lycophytes Ferns Japan 35 (2013) (L. ellipticus ×
= Leptochilus longipes (Ching) X.C.Zhang; Lycophytes Ferns China 655 (2012)
= Leptochilus neopothifolius Nakaike; Checkl. Lycophytes Ferns Japan 35 (2013)
= Pleopeltis elliptica (Thunb.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 12 (1909)
= Polypodium boisii Christ; J. Bot. [Morot] 19: 76 (1905)
= Polypodium chunii C.Chr.; Index Filic., Suppl. Tert. 146 (1934)
= Polypodium ellipticum f. breve Y.C.Wu; Bull. Dept. Biol. Sun Yatsen Univ. 3: 322 (1932)
= Polypodium ellipticum Thunb.; Syst. Veg., ed. 14 [J.A.Murray] 935 (1784)
= Polypodium faurianum Nakai; J. Coll. Sci. Imp. Univ. Tokyo 31: 416 (1911)
= Polypodium flavescens Ching; Bull. Fan Mem. Inst. Biol., Bot. 2: 22, t. 8 (1931) (fide Fraser-Jenkins 2008)
= Polypodium mediosorum Ching; Bull. Fan Mem. Inst. Biol., Bot. 2: 19, t. 4 (1931)
= Polypodium neoellipticum Koidz.; Bot. Mag. (Tokyo) 43: 388 (1929)
= Selliguea coraiensis Christ; Repert. Spec. Nov. Regni Veg. 5: 11 (1908) (non Polypodium coraiense Christ (1908))
= Selliguea elliptica (C.B.Clarke) Bedd.; Ferns Brit. India, t. 150 (1867) (pro parte)
= Selliguea elliptica (Thunb.) Bedd.; Ferns Brit. India, t. 150 (1867) (pro parte)
= Selliguea elliptica var. flagellaris Christ; Bull. Herb. Boissier 7: 6 (1899) (non Polypodium flagellare Christ (1896))
= Selliguea pentaphylla (Baker) Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 248 (1906), not Baker (1891)
species :
Leptochilus evrardii (Tardieu) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018)
Distribution: Vietnam
41 VIE
Remarks: Colysis clade, Laos subclade
Synonyms: hidden
= Colysis evrardii Tardieu; Bull. Soc. Bot. France 87: 372, t. 2(3-4) (1941)
species :
Leptochilus flexilobus (Christ) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: China (Chongqing, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan); Taiwan; Vietnam
36 CHI 38 TAI 41 VIE
Remarks: frequently included in L. ellipticus s. lat.
Synonyms: hidden
= Colysis elliptica var. flexiloba (Christ) L.Shi & X.C.Zhang; Acta Phytotax. Sin. 37: 74 (1999)
= Colysis flexiloba (Christ) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Colysis flexiloba var. undulatocrenata (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 331 (1933)
= Colysis flexiloba var. undulatorepanda (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4(10): 330 (1933)
= Leptochilus ellipticus var. flexilobus (Christ) X.C.Zhang; Lycophytes Ferns China 652 (2012)
= Pleopeltis flexiloba (Christ) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Polypodium ellipticum var. undulatocrenatum C.Chr.; Bull. Acad. Int. Géogr. Bot. 107 (1904)
= Polypodium ellipticum var. undulatorepandum C.Chr.; Bull. Acad. Int. Geogr. Bot. 22: 141 (1913)
= Polypodium flexilobum Christ; Bull. Acad. Int. Géogr. Bot. 107 (1904)
= Polypodium flexilobum var. undulatocrenatum Ching; Bull. Fan Mem. Inst. Biol., Bot. 2(2): 20 (1931), nom. nud.
= Selliguea flexiloba Christ; Bull. Herb. Boissier, Sér. 2, 6: 992 (1906)
species :
Leptochilus gracilis Z.L.Liang, Liang Zhang & Li Bing Zhang; Phytotaxa 468(1): 117 (2020)
Distribution: China (Yunnan)
36 CHI
species :
Leptochilus hemionitideus (C.Presl) Noot.; Blumea 42: 285 (1997)
Distribution: China (Anhui, Fujian, Guizhou, Guangdong, Guangxi, Hainan, Jiangxi, Sichuan, Yunnan); Tibet; S-Japan; Ryukyu Isl.; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam, Karnataka, Kerala, Meghalaya, Mizoram, Nagaland, Odisha, Sikkim, Darjeeling, Tamil Nadu, Tripura, Uttarakhand, West Bengal); Bangladesh; Myanmar (Kachin, Chin, etc.); Thailand; Laos; Vietnam; Philippines
36 CHI TIB 38 JAP NNS TAI 40 BAN BHU IND NEP 41 LAO MYA THA VIE 42 PHI
Remarks: Colysis clade, Colysis subclade
Synonyms: hidden
= Colysis hemionitidea (C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 147 (1851)
= Colysis hemionitidea (C.Presl) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933), isonym
= Drynaria hemionitidea (C.Presl) J.Sm.; J. Bot. [Hooker] 4: 61 (1841), nomen
= Gymnogramma plantaginea Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 39 (1827), nom. nov. rej. prop. for Hemionitis plantaginea D. Don 1825, non Sm. 1793
= Hemionitis plantaginea D.Don; Prodr. Fl. Nepal. (1825), nomen
= Leptochilus decurrens subsp. hemionitideus (C.Presl) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 63 (2008)
= Microsorum ensato-sessilifrons (Hayata) H.Itô; J. Jap. Bot. 11: 96 (1935)
= Microsorum hemionitideum (C.Presl) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)
= Pleopeltis hemionitidea (C.Presl) T.Moore; Index Fil. [T. Moore] 436 (1862)
= Polypodium ensato-sessilifrons Hayata; Icon. Pl. Formosan. 6: 312, f. 126 (1915)
= Polypodium hemionitideum (C.Presl) Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2: 112 (1856), et Ann. Mus. Bot. Lugduno-Batavi 4(2): 215 (1866)
= Selliguea hemionitidea C.Presl; Tent. Pterid. 216 (1836), nomen
species :
Leptochilus henryi (Baker) X.C.Zhang; Lycophytes Ferns China 654 (2012)
Distribution: China (Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Taiwan; Bangladesh; Myanmar (Kachin); Vietnam
36 CHI 38 TAI 40 BAN 41 MYA VIE
Remarks: ellipticus clade, henryi subclade
Synonyms: hidden
= Colysis henryi (Baker) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Colysis lioui Ching; Fl. Fujianica 1: 602 (1982)
= Gymnogramma henryi Baker; J. Bot. 25: 171 (1887)
= Leptochilus chittagongensis Fraser-Jenk. & Gias; Taxon. Revis. Indian Subcontinental Pteridophytes 67 (2008) (fide Fraser-Jenkins et al. 2019)
= Polypodium henryi (Baker) C.Chr.; Bull. Herb. Boissier 6: 873 (1898)
= Polypodium mon-changense C.Chr.; Index Filic., Suppl. (1906-1912) 60 (1913)
= Selliguea cochlearis Christ; Bull. Acad. Int. Géogr. Bot. 142 (1907) (non Polypodium cochleare Brause (1912))
= Selliguea henryi (Baker) Christ; Bull. Herb. Boissier 6: 870 (1898)
species :
Leptochilus heterophyllus (S.K.Wu & L.K.Phan) Christenh.; Ann. Bot. (Oxford), n. s. 113(4): 590 (2014)
Distribution: Vietnam
41 VIE
Remarks: ellipticus clade, Kontumia subclade
Synonyms: hidden
= Kontumia heterophylla S.K.Wu & L.K.Phan; Novon 15(1): 245 (2005)
= Leptochilus heterophyllus (S.K.Wu & L.K.Phan) R.Kr.Singh & V.K.Rawat; J. Biodivers. Conservation 9(2): 74 (2025), isonym
species :
Leptochilus kachinensis Liang Zhang & Li Bing Zhang; Phytotaxa 659(3): 220 (2024)
Distribution: Myanmar (Kachin)
41 MYA
Remarks: Colysis clade, Colysis subclade
species :
Leptochilus khammouanensis Liang Zhang, Thepkaysone & Z.Zhou; Phytotaxa 659(3): 221 (2024)
Distribution: C-Laos
Error
Remarks: pteropus clade
species :
Leptochilus lanceolatus Fée; Mém. Foug. 2 (Hist. Acrostich.): 87, t. 47, f. 1 (1844)
Distribution: NE-India; ?Sri Lanka; Bangladesh
40 BAN IND
Remarks: valid species fide Fraser-Jenkins (2008)
Synonyms: hidden
= Acrostichum lanceolatum (Fée) Hook.; Sp. Fil. [W.J.Hooker] 6: 276 (1864) (non L. (1753))
= Acrostichum variabile var. laciniatum Hook.; Sp. Fil. [W.J.Hooker] 5: 255 (1864)
= Campium laciniatum (Hook.) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium lanceolatum (Fée) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium zeylanicum (Fée) Copel.; Philipp. J. Sci. 37: 352 (1928)
= Colysis laciniata (Hook.) Panigrahi; Pl. Sci. Res. (India) 20: 20 (1998)
= Colysis lanceolata (Fée) Panigrahi; Pl. Sci. Res. (India) 20 (Abstr. 23rd Ann. Conf. Khurda): 35 (1998)
= Dendroglossa lanceolata (Fée) Fée; Mém. Foug. 5 (Gen. Filic.) 81 (1852), nom. illeg.
= Dendroglossa zeylanica (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 199 (1947)
= Gymnopteris feei f. triloba Bedd.; Ferns Brit. India, t. 273 (1868)
= Gymnopteris feei var. pinnatifida Bedd.; Ferns S. India, t. 211 (1864)
= Gymnopteris feei var. triloba Bedd.; Ferns Brit. India, t. 273 (1864)
= Gymnopteris lanceolata (Fée) Bedd.; Ferns Brit. India Suppl. 26. (1876)
= Gymnopteris variabilis var. lanceolata Bedd.; Ferns Brit. India 429 (1883)
= Leptochilus decurrens f. lanceolatus (Fée) Sledge; Ann. Mag. Nat. Hist., Ser. 12, 7: 869 (1956)
= Leptochilus laciniatus (Hook.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 344 (1933)
= Leptochilus lanceolatus var. simplex Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 344 (1933)
= Leptochilus zeylanicus Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 8, t. 31, f. 1 (1865) (fide Fraser-Jenkins 2008)
= Paraleptochilus decurrens var. lanceolata (Fée) R.D.Dixit; Indian Fern J. 6(1-2): 148 (1989)
= Pleopeltis feei (T.Moore) Alderw.; Malayan Ferns Fern Allies 405 (1917), nom. illeg., non (Bory) Alderw. 1909
species :
Leptochilus leveillei (Ching) X.C.Zhang; Lycophytes Ferns China 654 (2012)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou)
36 CHI
Synonyms: hidden
= Colysis leveillei (Christ) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Colysis leveillei f. angusta (C.Chr.) Ching; Bull. Dept. Biol. Sun Yatsen Univ. 3: ? (1933)
= Colysis leveillei f. major (C.Chr.) Ching; Bull. Dept. Biol. Sun Yatsen Univ. 3: ? (1933)
= Leptochilus leveillei (Christ) X.C.Zhang & Noot.; Fl. China 2-3: 835 (2013), isonym
= Polypodium leveillei (Christ) C.Chr.; Index Filic., Suppl. (1906-1912) 60 (1913)
= Polypodium leveillei f. angusta C.Chr.; Bull. Dept. Biol. Sun Yatsen Univ. 3: ? (1932)
= Polypodium leveillei f. majus C.Chr.; Bull. Dept. Biol. Sun Yatsen Univ. 3: ? (1932)
= Selliguea leveillei Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 236 (1906)
species :
Leptochilus locii Liang Zhang, N.T.Lu & Li Bing Zhang; Phytotaxa 659(3): 222 (2024)
Distribution: C-Vietnam (Quang Binh)
41 VIE
Remarks: ellipticus clade, henryi subclade
species :
Leptochilus longissimus (Blume) L.Y.Kuo; Taiwania 64(4): 378 (2019)
Distribution: China (Hainan, Yunnan, Hongkong); Japan; Ryukyu Isl.; Taiwan; India (Kerala, Assam); Bangladesh; Myanmar; Thailand; Vietnam; peninsular Malaysia (Perak, Selangor, Terengganu); Singapore; Sumatra; Borneo (Sabah); N-Sulawesi; Philippines; Java; Moluccas (Seram, Talaud); Bismarck Arch. (Admiralty Isl.); Palau (Babeldaob)
36 CHI 38 JAP NNS TAI 40 BAN IND 41 MYA THA VIE 42 BOR JAW MLY MOL PHI SNG SUL SUM 43 BIS 62 PLU
Remarks: traditionally often treated as Microsorum rubidum, which still might be the valid name
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012). Endangered (Singapore)
Synonyms: hidden
= Colysis longissima (Blume) J.Sm.; Hist. Fil. 101 (1875)
= Drynaria rubida J.Sm.; J. Bot. [Hooker] 3: 397 (1841), nom. nud.
= Microsorum rubidum (Kunze) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 197 (1947)
= Phymatodes longissima (Blume) J.Sm.; Cult. Ferns 10 (1857)
= Phymatodes suisha-stagnalis (Hayata) H.Itô; J. Jap. Bot. 11: 98 (1935)
= Phymatosorus longissimus (Blume) Pic.Serm.; Webbia 28: 459 (1973)
= Phymatosorus longissimus var. sordidum (Copel.) Nakai
= Phymatosorus suisha-stagnalis (Hayata) Pic.Serm.; Webbia 28: 460 (1973)
= Pleopeltis longissima (Blume) Bedd.; Ferns Brit. India Suppl. 388 (1876), isonym
= Pleopeltis longissima (Blume) Blume; Enum. Pl. Javae 2: 127 (1828)
= Pleopeltis longissima (Blume) T.Moore; Index Fil. [T. Moore] 78 (1857), isonym
= Polypodium longissimum Blume; Enum. Pl. Javae 2: 127 (1828)
= Polypodium rubidum Kunze; Bot. Zeitung (Berlin) 117 (1848) (fide Fraser-Jenkins 1997)
= Polypodium suisha-stagnale Hayata; Icon. Pl. Formosan. 6: 160 (1916), 156 (1919)
species :
Leptochilus luangprabangensis Liang Zhang, Thepkaysone & Z.Zhou; Phytotaxa 659(3): 224 (2024)
Distribution: Laos (Luang Prabang Prov.)
41 LAO
Remarks: ellipticus clade, henryi subclade
species :
Leptochilus macrophyllus (Blume) Noot.; Blumea 42: 289 (1997)
Distribution: Nicobar Isl.; Myanmar (Kachin, etc.); peninsular Thailand; peninsular Malaysia; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Lombok, Bali); Moluccas (Seram, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Britain, Admiralty Isl., New Ireland); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Malaita, Nggatokae, Ranongga, New Georgia, Pavuvu, Rendova, Vangunu, Vella Lavella)
41 MYA NCB THA 42 BOR JAW LSI MLY MOL NWG PHI PLW SUL SUM 43 BIS SOL
Remarks: macrophyllus clade
Synonyms: hidden
= Colysis acuminata Holttum; Rev. Fl. Malaya 2: 162, f. 73 (1955), nom. superfl.
= Colysis acuminata var. angustata Holttum; Rev. Fl. Malaya, ed. 2, 2: 163 (1966), nom. inval.
= Colysis acuminata var. fluminalis (Ridl.) Hennipman; Rheophytes World [Steenis] 157 (1981)
= Colysis campyloneuroides (Baker) Parris; Sandakania 50(4): 264 (1997), et in Malayan Nat. J. 9: 81 (1997)
= Colysis hosei (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 199 (1947)
= Colysis loxogrammoides (Copel.) M.G.Price; Contr. Univ. Michigan Herb. 16: 193 (1987)
= Colysis macrophylla (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 147 (1851)
= Colysis polysora (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 199 (1947)
= Colysis regularis (Mett.) Ching; Sunyatsenia 5: 261 (1940)
= Colysis spuria (Mett.) Ching; Sunyatsenia 6: 261 (1940)
= Grammitis macrophylla Blume; Enum. Pl. Javae 2: 119 (1828)
= Gymnogramma acuminata Baker; J. Bot. 26: 326 (1888) (non Kaulf. (1824), nec Polypodium acuminatum Don (1825))
= Gymnogramma macrophylla (Blume) Hook.; Sp. Fil. [W.J.Hooker] 5: 159 (1864)
= Gymnogramma regularis (Mett.) Baker; Syn. Fil. [W.J.Hooker & Baker] 388 (1868)
= Gymnogramma spuria (Mett.) Baker; Syn. Fil. [W.J.Hooker & Baker] 388 (1868)
= Pleopeltis campyloneuroides (Baker) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 7 (1909)
= Pleopeltis hosei (C.Chr.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 12 (1909)
= Pleopeltis interrupta (C.Chr.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 11 (1909)
= Pleopeltis linealifolia (Rosenst.) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Pleopeltis loxogrammoides (Copel.) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Pleopeltis macrophylla (Blume) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 12 (1909)
= Pleopeltis pseudoloxogramma Alderw.; Bull. Jard. Bot. Buitenzorg, Sér. 3, 6: 218 (1922)
= Pleopeltis regularis (Mett.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 11 (1909)
= Pleopeltis selligueoides Alderw.; Bull. Jard. Bot. Buitenzorg, Sér. 2, 23: 18 (1916) (non Baker (1874))
= Pleopeltis spuria (Mett.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 12 (1909)
= Polypodium campyloneuroides Baker; J. Linn. Soc., Bot. 20: 229 (1883)
= Polypodium hosei C.Chr.; Index Filic. 334 (1905), 534 (1906)
= Polypodium interruptum C.Chr.; Index Filic. 333 (1905)
= Polypodium linealifolium Rosenst.; Nova Guinea 8: 728 (1912)
= Polypodium loxogrammoides Copel.; Philipp. J. Sci. 7: 65 (1912)
= Polypodium macrophyllum (Blume) Reinw. ex Mett.; Fil. Hort. Bot. Lips. 37, t. 25, f. 22-23 (1856)
= Polypodium polysorum Brause; Bot. Jahrb. Syst. 56: 203 (1920)
= Polypodium pseudoloxogramma (Alderw.) C.Chr.; Index Filic., Suppl. Tert. 156 (1934)
= Polypodium regulare Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 225 (1866)
= Polypodium spathulifolium Blume; Enum. Pl. Javae 2: 134 (1828)
= Polypodium spurium Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 226 (1866)
= Selliguea campyloneuroides (Baker) Bedd.; Suppl. Ferns Brit. Ind. 101 (1892)
= Selliguea macrophylla (Blume) Blume; Enum. Pl. Javae 2: addend. (1828)
= Selliguea membranacea var. fluminalis Ridl.; J. Malayan Branch Roy. Asiat. Soc. 4: 89 (1926)
= Selliguea polysora (Brause) Ching; Sunyatsenia 6: 260 (1940)
= Selliguea pseudoloxogramma (Alderw.) Ching; Sunyatsenia 5: 260 (1940)
= Syngramma campyloneuroides (Baker) Ridl.; J. Malayan Branch Roy. Asiat. Soc. 4: 105 (1926)
species :
Leptochilus mengsongensis M.X.Zhao; Phytotaxa 317(2): 144 (2017)
Distribution: China (Yunnan)
36 CHI
species :
Leptochilus minor Fée; Mém. Foug. 2 (Hist. Acrostich.): 87, t. 25 (1845)
Distribution: NE-India (Meghalaya); Myanmar (Kachin, etc.); Thailand; Vietnam; peninsular Malaysia; Sumatra; Borneo (Sabah); Sulawesi; Philippines (Luzon)
40 IND 41 MYA THA VIE 42 BOR MLY PHI SUL SUM
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Acrostichum minus (Fée) Mett.; Fil. Hort. Bot. Lips. 20 (1856)
= Campium minus (Fée) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Campium minutulum (Fée) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Colysis minor (Fée) M.G.Price; Kalikasan 3: 176 (1974)
= Colysis minutula (Fée) Panigrahi; Pl. Sci. Res. (India) 20 (Abstr. 23rd Ann. Conf. Khurda): 35 (1998)
= Dendroglossa minor (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 199 (1947)
= Dendroglossa minutula (Fée) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 199 (1947) (fide Fraser-Jenkins 2008)
= Dendroglossa normalis (J.Sm.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 149 (1851)
= Gymnopteris minor (Fée) Hook.; Sec. Cent. Ferns, t. 78 (1861)
= Gymnopteris normalis J.Sm.; J. Bot. [Hooker] 3: 403 (1841), nomen
= Leptochilus minutulus Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 8, t. 31, f. 2 (1865) (fide Fraser-Jenkins 2008)
= Leptochilus normalis (J.Sm.) Copel.; Philipp. J. Sci. 8: 31 (1908)
= Leptochilus rizalianus Christ; Bull. Herb. Boissier, Sér. 2, 6: 1004 (1906)
species :
Leptochilus morsei (Ching) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 2: 443 (2018)
Distribution: China; India
36 CHI 40 IND
Remarks: separate species from L. ellipticus fide Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Colysis morsei (Ching) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Polypodium morsei Ching; Bull. Fan Mem. Inst. Biol., Bot. 2: 17, t. 1 (1931)
species :
Leptochilus multilobus Liang Zhang & X.M.Zhou; Phytotaxa 659(3): 225 (2024)
Distribution: SE-Tibet (Medog)
36 TIB
Remarks: Colysis clade
species :
Leptochilus neolongipes Liang Zhang, X.M.Zhou, T.T.Luong & Li Bing Zhang; Phytotaxa 659(3): 227 (2024)
Distribution: S-Vietnam
41 VIE
Remarks: ellipticus clade, Khanhhoa subclade
species :
Leptochilus oblongus Li Bing Zhang, Liang Zhang & N.T.Lu; Phytotaxa 234(2): 195 (2015)
Distribution: N-Vietnam
41 VIE
Remarks: ellipticus clade, oblongus subclade
species :
Leptochilus ornithopus T.Fujiw. & B.H.Quang; Phytotaxa 584(3): 153 (2023)
Distribution: Vietnam
41 VIE
Remarks: ellipticus clade, Khanhhoa subclade
species :
Leptochilus ovatifolius Zhe Zhang, S.W.Yao & Yi Huang; J. S. W. Forest. Univ. 43(5): 185 (2023)
Distribution: China (Guangdong)
36 CHI
species :
Leptochilus ovatus Copel.; Philipp. J. Sci. 9: 229 (1914)
Distribution: Thailand; Laos; Vietnam
41 LAO THA VIE
Remarks: Colysis clade; separate from L. pedunculatus fide molecular data (Zhang et al. 2024). Some synonyms listed there might belong here.
Synonyms: hidden
= Bolbitis ovata (Copel.) Ching; Index Filic., Suppl. Tert. 49 (1934)
= Campium ovatum (Copel.) Copel.; Philipp. J. Sci. 37: 341-402 (1928)
= Paraleptochilus ovatus (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 198 (1947)
species :
Leptochilus pedunculatus (Hook. & Grev.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008)
Distribution: China (Yunnan, Guangxi, Hainan); Nepal; Bhutan; NE-India (Assam, Meghalaya, Arunachal Pradesh, Sikkim, Darjeeling); Sri Lanka; Bangladesh; Myanmar (Kachin, etc.); Thailand (widespread); Laos; Vietnam; peninsular Malaysia (Langkawi Isl., Perak, Selangor, Kelantan, Pahang); Sumatra; Borneo (Sabah); Philippines; Java
36 CHI 40 BAN BHU IND NEP SRL 41 LAO MYA THA VIE 42 BOR JAW MLY PHI SUM
Remarks: Colysis clade. L. ovatus is a separate species fide Zhang et al. 2024
Synonyms: hidden
= Ceterach pedunculatum Hook. & Grev.; Icon. Filic. t. 5 (1827)
= Colysis hemitoma (Hance) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Colysis hemitoma f. integra Ching ex S.H.Fu
= Colysis intermedia Ching & Chu H.Wang; Acta Phytotax. Sin. 8: 170, t. 23(30) (1959)
= Colysis macrophylla var. pedunculata (Hook. & Grev.) S.Linds.; Edinburgh J. Bot. 66(2): 356 (2009)
= Colysis marginata (Meyen ex C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 148 (1851)
= Colysis membranacea (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 147 (1851)
= Colysis pedunculata (Hook. & Grev.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 321 (1933)
= Colysis selliguea (Mett.) Ching; Sunyatsenia 6: 261 (1940)
= Grammitis hamiltoniana Wall.; Numer. List [Wallich] no. 9 (1829), nomen
= Grammitis membranacea Blume; Enum. Pl. Javae 2: 118 (1828) (non Polypodium membranaceum Don (1825))
= Grammitis pedunculata (Hook. & Grev.) Blume; Enum. Pl. Javae 2: 118 (1828)
= Gymnogramma hamiltoniana Wall. ex Hook.; Sp. Fil. [W.J.Hooker] 6: 161 (1864)
= Gymnogramma membranacea (Blume) Hook.; Sp. Fil. [W.J.Hooker] 5: 159 (1864)
= Leptochilus macrophyllus var. pedunculatus (Hook. & Grev.) Noot.; Blumea 42: 290 (1997)
= Leptochilus ×hemitomus (Hance) Noot.; Blumea 42: 293 (1997)
= Pleopeltis pedunculata (Hook. & Grev.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 12 (1909)
= Pleopeltis selliguea (Mett.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 11 (1909)
= Polypodium bonii Christ; Bull. Fan Mem. Inst. Biol., Bot. 4: 322 (1933)
= Polypodium hamiltonianum (Wall. ex Hook.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 316 (1899)
= Polypodium hemitomum Hance; J. Bot. 21: 269 (1883)
= Polypodium pedunculatum (Hook. & Grev.) Mett.; Nomencl. Gefäßkrypt. 312 (1883)
= Polypodium selliguea Mett.; Abh. Senckenberg. Naturf. Ges. 2(1) [Polypodium]: 111 (1856)
= Scolopendrium lanceolatum Roxb.; Calcutta J. Nat. HIst. 4: 501 (1844)
= Selliguea aliena Brack.; U.S. Expl. Exped. 16 (Filic.): 58 (1854)
= Selliguea hamiltoniana (Wall. ex Hook.) Bedd.; Ferns Brit. India, t. 239 (1867)
= Selliguea hamiltonii (Wall. ex Hook.) C.Presl; Tent. Pterid. 216 (1836)
= Selliguea hookeri C.Presl; Tent. Pterid. 216 (1836)
= Selliguea marginata Meyen ex C.Presl; Tent. Pterid. 216 (1836)
= Selliguea membranacea (Blume) Blume; Enum. Pl. Javae 2: addend. (1828)
= Selliguea pedunculata (Hook. & Grev.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 146 (1851)
= Selliguea selliguea (Mett.) Christ; Bull. Herb. Boissier, Sér. 2, 6: 992 (1906)
species :
Leptochilus pentaphyllus (Baker) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: China (Guangdong, Guangxi, Guizhou, Yunnan); Tibet; Laos
36 CHI TIB 41 LAO
Remarks: pentaphyllus clade
Synonyms: hidden
= Colysis elliptica var. pentaphylla (Baker) L.Shi & X.C.Zhang; Acta Phytotax. Sin. 37(1): 77 (1999)
= Colysis longisora (Baker) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 331 (1933) (fide Fraser-Jenkins 2008)
= Colysis pentaphylla (Baker) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 332 (1932)
= Gymnogramma longisora Baker; J. Bot. 28: 267 (1890) (fide Fraser-Jenkins 2008)
= Gymnogramma pentaphylla Baker; Bull. Misc. Inform. Kew 1898: 233 (1898) (non Polypodium pentaphyllum Baker (1891))
= Leptochilus ellipticus var. pentaphyllus (Baker) X.C.Zhang & Noot.; Fl. China 2-3: 837 (2013)
= Polypodium ellipticum var. pentaphyllum (Baker) C.Chr.
= Polypodium longisorum (Baker) C.Chr.; Index Filic. 541 (1906) (fide Fraser-Jenkins 2008)
species :
Leptochilus phanerophlebius Liang Zhang, Z.L.Liang & Zhi Y.Yu; Phytotaxa 716(2): 132 (2025)
Distribution: China (SE-Yunnan)
36 CHI
species :
Leptochilus poilanei (C.Chr. & Tardieu) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: Vietnam; etc.
41 VIE
Synonyms: hidden
= Colysis poilanei C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 201 (1939)
species :
Leptochilus pothifolius (Buch.-Ham. ex D.Don) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 60 (2008)
Distribution: China (Chongqing, Fujian, Guangxi, Guizhou, Hainan, Hongkong, Hunan, Jiangxi, Yunnan, Zhejiang); Japan; Ryukyu Isl.; Ogasawara Isl. (Chichijima, Hahajima); ?Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Meghalaya); Myanmar; Thailand; Laos; Cambodia; Vietnam; Philippines
36 CHI 38 JAP NNS 40 BHU IND NEP 41 CBD LAO MYA THA VIE 42 PHI
Remarks: included sometimes as variety (or completely) in L. ellipticus, but accepted as full species by Fraser-Jenkins et al. (2008, 2015)
Synonyms: hidden
= Colysis elliptica f. simplex Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 335 (1933)
= Colysis elliptica var. furcans (Tutcher) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 335 (1933)
= Colysis elliptica var. pothifolia (Buch.-Ham. ex D.Don) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 334 (1933)
= Colysis latiloba (Ching) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 330 (1933)
= Colysis pothifolia (Buch.-Ham. ex D.Don) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 148 (1851)
= Colysis pothifolia (Buch.-Ham. ex D.Don) H.Itô; J. Jap. Bot. 11: 89 (1935), isonym
= Colysis pothifolia f. furcans (Ching) S.R.Ghosh; Pterid. Fl. E. India 499 (2004)
= Colysis pothifolia f. simplex (Ching) S.R.Ghosh; Pterid. Fl. E. India 500 (2004)
= Hemionitis pothifolia Buch.-Ham. ex D.Don; Prodr. Fl. Nepal. 13 (1825)
= Leptochilus ellipticus var. pothifolius (Buch.-Ham. ex D.Don) X.C.Zhang; Lycophytes Ferns China 653 (2012)
= Polypodium ellipticum var. furcans (Tutcher) Ching; Bull. Fan Mem. Inst. Biol., Bot. 2(2): 21 (1931)
= Polypodium ellipticum var. pothifolium (Buch.-Ham. ex D.Don) Makino; Bot. Mag. (Tokyo) 23: 72 (1909)
= Polypodium latilobum Ching; Bull. Fan Mem. Inst. Biol., Bot. 2: 21, t. 7 (1931)
= Polypodium pothifolium (Buch.-Ham. ex D.Don) Mett.; Fil. Hort. Bot. Lips. 130, t. 25, f. 21 (1856)
= Polypodium pothifolium var. furcans (Tutcher) Ching; Bull. Fan Mem. Inst. Biol., Bot. 2(2): 21 (1931)
= Selliguea pothifolia (Buch.-Ham. ex D.Don) J.Sm.; J. Bot. [Hooker] 3: 399 (1841)
species :
Leptochilus pteropus (Blume) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008)
subspecies :
subsp. pteropus
Distribution: China (Fujian, Guangxi, Guizhou, Hainan, Hongkong, Hunan, Jiangxi, Yunnan); Ryukyu Isl. (Iriomote Isl., Ishigakijima); Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tripura, West Bengal); Bangladesh; Andaman Isl.; Myanmar; Thailand (widespread); Laos; Vietnam; peninsular Malaysia (Langkawi Isl., Perak, Kuala Lumpur, Terengganu, Pahang); Sumatra; Borneo (Sabah, Sarawak); Philippines; Java; Lesser Sunda Isl. (Lombok); Moluccas (Halmaheira); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.)
36 CHI 38 NNS TAI 40 BAN BHU IND NEP 41 AND LAO MYA THA VIE 42 BOR JAW LSI MLY MOL NWG PHI SUM 43 BIS
Remarks: pteropus clade
Synonyms: hidden
= Colysis pteropus (Blume) Bosman; Leiden Bot. Ser. 14: 112 (1991)
= Colysis tridactylum (Wall. ex Hook. & Grev.) J.Sm.; Hist. Fil. 101 (1875)
= Drynaria dubia J.Sm.; J. Bot. [Hooker] 3: 397 (1841)
= Drynaria tridactyla (Wall. ex Hook. & Grev.) Fée; Mém. Foug. 5 (Gen. Filic.) 271 (1852)
= Kaulinia pteropus (Blume) B.K.Nayar; Taxon 13: 67 (1964)
= Lepisorus pteropus (Blume) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 17 (1933)
= Lepisorus pteropus (Blume) R.D.Dixit; Census Indian Pteridophytes [Dixit] 42 (1984), isonym
= Microsorum paucijugum (Alderw.) K.Iwats. & M.Kato; Acta Phytotax. Geobot. 32(1-4): 123 (1981)
= Microsorum pteropus (Blume) Copel.; Univ. Calif. Publ. Bot. 16: 111-116 (1929)
= Phymatodes tridactyla C.Presl; Tent. Pterid. 196 (1836), nom. nud.
= Pleopeltis aquatica (Christ) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Pleopeltis paucijuga (Alderw.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 8 (1909)
= Pleopeltis pteropus (Blume) Bedd.; Ferns Brit. India 359 (1883), non T. Moore (1857)
= Pleopeltis pteropus (Blume) T.Moore; Index Fil. [T. Moore] 78 (1857)
= Pleopeltis tridactyla (Wall. ex Hook. & Grev.) T.Moore; Index Fil. [T. Moore] 78 (1857)
= Polypodium aquaticum Christ; Nova Guinea 8: 153 (1909)
= Polypodium paucijugum Alderw.; Bull. Dépt. Agric. Indes Néerl. 18: 24 (1908)
= Polypodium pteropus Blume; Enum. Pl. Javae 2: ed. 3 (1828)
= Polypodium tridactylum Wall. ex Hook. & Grev.; Icon. Filic. 2: t. 209 (1831)
subspecies :
subsp. minor (Bedd.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 62 (2008), comb. inval., et Indian J. Forest. 42(4): 311 (2019)
Distribution: S-India (Karnataka, Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Colysis zosteriformis (Wall. ex Mett.) J.Sm.; Hist. Fil. 100 (1875)
= Kaulinia pteropus var. minor (Bedd.) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 87 (1974)
= Kaulinia zosteriformis (Wall. ex Mett.) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 87 (1974)
= Lepisorus pteropus var. minor (Bedd.) N.P.Balakr.; Bull. Bot. Surv. India 22 (1980)
= Lepisorus pteropus var. minor (Bedd.) R.D.Dixit; Census Indian Pteridophytes [Dixit] 42 (1984) (as (Blume) Ching´), isonym
= Microsorum brassii Copel.; J. Arnold Arbor. 10: 181 (1929)
= Microsorum pteropus f. minus (Bedd.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 312 (1933)
= Microsorum pteropus var. minus (Bedd.) C.Chr. & Tardieu; Notul. Syst. (Paris) 8: 194 (1939)
= Microsorum pteropus var. minus (Bedd.) S.R.Ghosh; Pterid. Fl. E. India 565 (2004), isonym
= Microsorum pteropus var. zosteriforme (Wall. ex Mett.) S.Kaur & Subh.Chandra; Nomencl. Guide to Beddome´s Ferns S. India & Brit. India: 49 (1987)
= Microsorum zosteriforme (Wall. ex Mett.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 295-311 (1933)
= Phymatodes zosteriformis (Wall. ex Mett.) C.Presl; Tent. Pterid. 196 (1836)
= Pleopeltis pteropus var. minor Bedd.; Ferns Brit. India 361 (1833), et Ferns S. India, t. 179 (1865)
= Pleopeltis pteropus var. zosteriformis (Wall. ex Mett.) Bedd.; Ferns Brit. India 362 (1883)
= Pleopeltis raapii (Alderw.) Alderw.; Bull. Dépt. Agric. Indes Néerl. 27: 6 (1909)
= Pleopeltis zosteriformis (Wall. ex Mett.) Bedd.; Ferns Brit. India, t. 123 (1866)
= Polypodium brassii (Copel.) C.Chr.; Index Filic., Suppl. Tert. 145 (1934)
= Polypodium micropteris Baker; Bull. Misc. Inform. Kew 1906: 14 (1906) (non Index Filic. 545)
= Polypodium pteropus var. minor (Bedd.) Y.C.Wu, K.Wong & Pong; Bull. Dept. Biol. Sun Yatsen Univ. 3: 298 (1932)
= Polypodium raapii Alderw.; Bull. Dépt. Agric. Indes Néerl. 18: 23, t. 4 (1908)
= Polypodium udum Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 20: 140 (1910)
= Polypodium zosteriforme Wall.; Numer. List [Wallich] no. 280 (1829), nomen
= Polypodium zosteriforme Wall. ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(1) [Polypodium]: 86 (1856)
species :
Leptochilus sanjiangensis (H.G.Zhou & Hua Li) Liang Zhang & Li Bing Zhang; Phytotaxa 659(3): 233 (2024)
Distribution: China (Guangxi)
36 CHI
Synonyms: hidden
= Colysis sanjiangensis H.G.Zhou & Hua Li; Acta Bot. Yunnan. 15(3): 254 (1993)
= Colysis elliptica var. sanjiangensis (H.G.Zhou & Hua Li) R.H.Jiang; Fl. Guangxi 6: 406 (2013)
species :
Leptochilus sarawakensis Li Bing Zhang & Liang Zhang; Phytotaxa 379(3): 269 (2018), nom. nov.
Distribution: Borneo (Sabah, Sarawak); Sulawesi; Philippines
42 BOR PHI SUL
Remarks: frequently included in L. pedunculatus
Synonyms: hidden
= Colysis fluviatilis (Lauterb.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Leptochilus fluviatilis (Lauterb.) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 173 (2018), nom. illeg., non (Hook.) C. Chr.
= Leptochilus macrophyllus var. fluviatilis (Lauterb.) Noot.; Blumea 42: 289 (1997)
= Pleopeltis fluviatilis (Lauterb.) Alderw.; Malayan Ferns Fern Allies, Suppl. 383-406 (1917)
= Polypodium fluviatile Lauterb.; Bot. Jahrb. Syst. 44: 507 (1910)
species :
Leptochilus saxicola (H.G.Zhou & Hua Li) Liang Zhang & Li Bing Zhang; Phytotaxa 374(2): 174 (2018)
Distribution: China (Yunnan, etc.)
36 CHI
Remarks: frequently included in L. pedunculatus
Synonyms: hidden
= Colysis saxicola H.G.Zhou & Hua Li; Acta Bot. Yunnan. 15(3): 253 (1993)
species :
Leptochilus scandens H.J.Wei & Yi Huang; Phytotaxa 618(1): 80 (2023)
Distribution: China (Guangdong)
36 CHI
Remarks: ellipticus clade, Taiwan subclade
species :
Leptochilus sinovietnamica Liang Zhang, N.T.Lu & Li Bing Zhang; Phytotaxa 659(3): 228 (2024)
Distribution: China (Yunnan); C-Vietnam; N-Vietnam
36 CHI 41 VIE
Remarks: Vietnam clade
species :
Leptochilus thwaitesianus Fée; Mém. Foug. 10 (Icon. Esp. Nouv.): 7 (1865)
Distribution: S-India
40 IND
Remarks: according to Fraser-Jenkins et al. 2019 a valid tetraploid species intermediate between L. lanceolatus and L. decurrens
Synonyms: hidden
= Colysis thwaitesianus (Fée) Rajkumar & Arunach.; Amer. Fern J. 88(4): 183 (1999)
species :
Leptochilus tridigitatus H.J.Wei; Phytotaxa 650(2): 150 (2024)
Distribution: China (Guangdong, Hongkong)
36 CHI
species :
Leptochilus vietnamensis Liang Zhang, N.T.Lu & Li Bing Zhang; Phytotaxa 659(3): 230 (2024)
Distribution: N-Vietnam
41 VIE
Remarks: ellipticus clade, henryi subclade
species :
Leptochilus wrightii (Hook.) X.C.Zhang; Lycophytes Ferns China 656 (2012)
Distribution: China (Fujian, Guangxi, Hongkong, Jiangxi, Yunnan); Japan; Ryukyu Isl.; Taiwan; Vietnam
36 CHI 38 JAP NNS TAI 41 VIE
Synonyms: hidden
= Colysis hokouensis Ching; Bull. Fan Mem. Inst. Biol., Bot., Ser. 2, 1: 284 (1949)
= Colysis longifrons Tagawa; J. Jap. Bot. 14: 110 (1938)
= Colysis megalolepis Tagawa; J. Jap. Bot. 12: 492 (1936)
= Colysis subsessilifolia Ching; Wuyi Sci. J. 1: 11 (1981)
= Colysis wrightii (Hook.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Colysis wrightii f. laciniata Sa.Kurata
= Gymnogramma wrightii Hook.; Sp. Fil. [W.J.Hooker] 5: 160 (1864)
= Leptochilus macrophyllus var. wrightii (Hook. & Baker) Noot.; Blumea 42(2): 291 (1997)
= Polypodium kusukusense Hayata; Icon. Pl. Formosan. 6: 319, f. 131 (1915)
= Polypodium wrightii (Hook.) Mett. ex Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 316 (1899)
= Selliguea wrightii (Hook.) J.Sm.; Hist. Fil. 102 (1875)
species :
Leptochilus wui (Ching) Liang Zhang & Li Bing Zhang; Phytotaxa 659(3): 233 (2024)
Distribution: China (Guangxi)
36 CHI
Synonyms: hidden
= Colysis wui Ching; Bull. Fan Mem. Inst. Biol., Bot. 4: 313-331 (1933)
= Polypodium wui (Ching) C.Chr.; Bull. Dept. Biol. Sun Yatsen Univ. 6: 17 (1933)
species :
Leptochilus wusugongii Liang Zhang & Li Bing Zhang; Phytotaxa 659(3): 231 (2024)
Distribution: Laos (Luang Prabang Prov.)
41 LAO
species :
Leptochilus ×nepalensis Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 64 (2008)
Distribution: Nepal
40 NEP
Remarks: Leptochilus pothifolius x decurrens
species :
Leptochilus ×shintenensis (Hayata) Nakaike; Checkl. Lycophytes Ferns Japan 35 (2013)
Distribution: South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Taiwan; Vietnam
38 JAP KOR NNS TAI 41 VIE
Remarks: Leptochilus ellipticus x L. wrightii
Synonyms: hidden
= Colysis elliptica var. simplicifrons (Christ) Masam.; Sci. Rep. Kanazawa Univ., Biol. 2: 102 (1953)
= Colysis wrightii var. lacerata Nakai
= Colysis ×shintenensis (Hayata) H.Itô; J. Jap. Bot. 11: 90 (1935)
= Colysis ×simplicifrons (Christ) Tagawa; J. Jap. Bot. 25: 114 (1950)
= Leptochilus ×shintenensis (Hayata) X.C.Zhang & Noot.; Fl. China 2-3: 835 (2013), isonym
= Leptochilus ×simplicifrons (Christ) Nakaike; Checkl. Lycophytes Ferns Japan 36 (2013)
= Polypodium ellipticum var. simplicifrons Christ; Bull. Herb. Boissier, Sér. 2, 2: 832 (1902)
= Polypodium ×shintenense Hayata; Icon. Pl. Formosan. 8: 154, f. 85-86 (1919)
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.02 (Feb. 1st, 2026)

Covering 358,774 accepted species,
8,847 hybrids, 48,003 accepted infraspecific taxa and 1,006,442 synonyms,
in total 1,422,066* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

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With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

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