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044.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
3-34 genera (depending on genus concept), 1100-1200 spp. The genus concepts were wildly fluctuating and disputed, from the inclusion of everything into one monster genus Thelypteris to the split of over 30 genera. We use the new genus concept by Fawcett & Smith (2021), which is based on molecular results and adopts a compromise approachl
Species listed in database: 1181; hybrids: 39
044.2010 Genus Leptogramma J.Sm.; J. Bot. [Hooker] 4: 51 (1841)
Distribution: Mostly Himalayas (S-China) to Japan and Malesia, 1 South Africa, 1 Europe, 2 North America
Species listed in database: 29
= Thelypteris subgen.Leptogramma (J.Sm.) C.F.Reed; Phytologia 17: 254 (1968)
= Craspedosorus Ching & W.M.Chu; Acta Phytotax. Sin. 16(4): 24, t. 1 (1978)
species :
Leptogramma amabilis Tagawa; Acta Phytotax. Geobot. 7: 76 (1938)
Distribution: Ryukyu Isl.
38 NNS
Conservation Status: NT in Japan
Synonyms: hidden
= Leptogramma totta var. amabilis (Tagawa) H.Itô; (1938)
= Stegnogramma amabilis (Tagawa) Nakaike; Checkl. Lycophytes Ferns Japan 15 (2013)
= Stegnogramma gymnocarpa subsp. amabilis (Tagawa) K.Iwats.; Acta Phytotax. Geobot. 7: 76 (1938)
= Thelypteris gymnocarpa subsp. amabilis (Tagawa) C.V.Morton; Amer. Fern J. 56: 179 (1966)
species :
Leptogramma burksiorum (J.E.Watkins & Farrar) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 178 (2019)
Name: Alabama maiden fern
Distribution: USA (Alabama)
78 USA
Conservation Status: IUCN V
Synonyms: hidden
= Cyclosorus burksiorum (J.E.Watkins & Farrar) Mazumdar & R.Mukhop.; Bionature 33(1): 16 (2013)
= Leptogramma pilosa var. alabamensis (Crawford) Wherry; S. Fern Guide 346 (1964)
= Stegnogramma burksiorum (J.E.Watkins & Farrar) Weakley; J. Bot. Res. Inst. Texas 5(2): 451 (2011)
= Stegnogramma pilosa var. alabamensis (Crawford) K.Iwats.; Amer. Fern J. 54: 142 (1964)
= Thelypteris burksiorum J.E.Watkins & Farrar; Amer. Fern J. 92(2): 172 (2002), nom. nov.
= Thelypteris pilosa var. alabamensis Crawford; Amer. Fern J. 41: 19 (1951)
species :
Leptogramma celebica Ching; Sinensia 7: 99, t. 5 (1936) (non Thelypteris celebica (Baker) C. F. Reed)
Distribution: SW-Sulawesi
42 SUL
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Stegnogramma celebica (Ching) Holttum; Fl. Males., Ser. 2, Pterid. 543 (1981)
= Stegnogramma gymnocarpa subsp. celebica K.Iwats.; Acta Phytotax. Geobot. 19: 123 (1963)
= Thelypteris bunnemeyeri C.F.Reed; Phytologia 17: 265 (1968), nom. nov.
= Thelypteris rivendellensis Christenh.; Global Fl. 4: 37 (2018), nom. nov. superfl. for Leptogramma celebica Ching
species :
Leptogramma centrochinensis Ching ex Y.X.Lin; Fl. Reipubl. Popularis Sin. 4(1): 323 (1999)
Distribution: China (W-Hubei)
36 CHI
Synonyms: hidden
= Stegnogramma centrochinensis (Ching ex Y.X.Lin) H.F.Wang, X.Y.Dong & K.P.Ma; Checkl. Vasc. Pl. E. Asia 1: 59 (2025)
= Thelypteris centrochinensis (Ching ex Y.X.Lin) Christenh.; Global Fl. 4: 28 (2018)
species :
Leptogramma crenata (Holttum) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 50 (2021)
Distribution: Sulawesi
42 SUL
Synonyms: hidden
= Stegnogramma crenata Holttum; Blumea 33(2): 404 (1998)
= Thelypteris legolas Christenh.; Global Fl. 4: 33 (2018), nom. nov.
species :
Leptogramma cyrtomioides (C.Chr.) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 177 (2019)
Distribution: China (Guizhou, Sichuan, Hunan, Yunnan)
36 CHI
Synonyms: hidden
= Dryopteris stegnogramme var. cyrtomioides C.Chr.; Acta Horti Gothob. 1: 56 (1924)
= Thelypteris cyrtomioides (C.Chr.) C.F.Reed; Phytologia 17: 465 (1969)
= Stegnogramma cyrtomioides (C.Chr.) Ching; Sinensia 7: 95, t. 3 (1936)
= Stenogramma petiolulata Ching; (1976), nom. nud.
species :
Leptogramma dissitifolia (Holttum) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 178 (2019)
Distribution: Philippines (Luzon)
42 PHI
Synonyms: hidden
= Stegnogramma dissitifolia Holttum; Fl. Males., Ser. 2, Pterid. 543 (1981)
= Thelypteris dissitifolia (Holttum) Christenh.; Global Fl. 4: 29 (2018)
species :
Leptogramma gymnocarpa (Copel.) Ching; Acta Phytotax. Sin. 8: 318 (1963)
Distribution: ?Sulawesi; Philippines (Mindanao)
42 PHI
Synonyms: hidden
= Dryopteris gymnocarpa Copel.; Leafl. Philipp. Bot. 3: 807 (1910)
= Lastrea gymnocarpa (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 139 (1947)
= Phegopteris gymnocarpa (Copel.) Alderw.; Malayan Ferns Fern Allies 313 (1917)
= Stegnogramma gymnocarpa (Copel.) K.Iwats.; Acta Phytotax. Geobot. 19: 122 (1963)
= Thelypteris gymnocarpa (Copel.) C.V.Morton; Amer. Fern J. 56: 179 (1966)
species :
Leptogramma himalaica Ching; Sinensia 7: 100, t. 6 (1936)
Distribution: China (Yunnan); India (Himachal Pradesh, ?Tamil Nadu)
36 CHI 40 IND
Remarks: separate from L. mollissima fide Kuo et al. 2019
Synonyms: hidden
= Leptogramma yunnanensis Ching; Icon. Filic. Sin. 5: t. 228 (1958)
= Stegnogramma himalaica (Ching) K.Iwats.; Acta Phytotax. Geobot. 19: 122 (1963)
= Thelypteris himalaica (Ching) C.F.Reed; Phytologia 17: 283 (1968)
= Thelypteris pozoi subsp. himalaica (Ching) C.V.Morton; Amer. Fern J. 56(4): 179 (1966)
species :
Leptogramma huishuiensis Ching ex Y.X.Lin; Fl. Reipubl. Popularis Sin. 4(1): 323 (1999)
Distribution: China (Guizhou)
36 CHI
Synonyms: hidden
= Leptogramma pinfaensis Ching; (1976), nom. nud.
= Stegnogramma huishuiensis (Ching ex Y.X.Lin) H.F.Wang, X.Y.Dong & K.P.Ma; Checkl. Vasc. Pl. E. Asia 1: 59 (2025)
= Thelypteris huishuiensis (Ching ex Y.X.Lin) Christenh.; Global Fl. 4: 32 (2018)
species :
Leptogramma intermedia Ching ex Y.H.Chang & L.Y.Kuo; Cladistics 36: 179 (2019)
Distribution: China (Zhejiang, Fujian, Guizhou)
36 CHI
Synonyms: hidden
= Leptogramma intermedia Ching; Sporae Pterid. Sin. 286 (1976) nom. nud.
= Leptogramma intermedia Ching ex Y.X.Lin; Fl. Zhejiang 1: 154 (1993), nom. nud.
= Stegnogramma intermedia (Ching ex Y.H.Chang & L.Y.Kuo) H.F.Wang, X.Y.Dong & K.P.Ma; Checkl. Vasc. Pl. E. Asia 2: 2303 (2025)
species :
Leptogramma jinfoshanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 17 (1984)
Distribution: China (Chongqing)
36 CHI
Remarks: valid species fide Kuo et al. 2019
Synonyms: hidden
= Thelypteris jinfoshanensis (Ching & Z.Y.Liu) Christenh.; Global Fl. 4: 32 (2018)
species :
Leptogramma jinyunensis Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 177 (2019), nom. nov.
Distribution: China (Chongqing)
36 CHI
Remarks: valid species fide Kuo et al. 2019
Synonyms: hidden
= Stegnogramma diplazioides Ching ex Y.X.Lin; Fl. Reipubl. Popularis Sin. 4(1): 288 (1999), nom. illeg.
species :
Leptogramma latipinna (Ching ex Y.X.Lin) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 177 (2019)
Distribution: China (Yunnan)
36 CHI
Remarks: valid species fide Kuo et al. 2019
Synonyms: hidden
= Stegnogramma latipinna Ching ex Y.X.Lin; Fl. Reipubl. Popularis Sin. 4(1): 350 (1999)
= Stegnogramma latipinna K.H.Shing ex Ching; Vasc. Pl. Hengduan Mount. 1: 99 (1993), et Acta Phytotax. Sin. 31(6): 571 (1993), without latin descr. or type
= Thelypteris samwisei Christenh.; Global Fl. 4: 37 (2018), nom. nov.
species :
Leptogramma leptogrammoides (K.Iwats.) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 178 (2019)
Distribution: China (Yunnan); ?Tibet; India (Sikkim)
36 CHI 40 IND
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Thelypteris kingii C.F.Reed; Phytologia 17: 465 (1969) (nom. nov.)
= Stegnogramma leptogrammoides K.Iwats.; Acta Phytotax. Geobot. 19: 119 (1963)]
species :
Leptogramma liui Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 178 (2019), nom. nov.
Distribution: China (Chongqing, Sichuan, Yunnan)
36 CHI
Synonyms: hidden
= Stegnogramma jinfoshanensis Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 3(4): 13 (1983)
species :
Leptogramma mollissima (Fisch. ex Kunze) Ching; Sinensia 7: 102, t. 9 (1936)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; South Korea (incl. Cheju Isl.); Japan; Ryukyu Isl.; Bonin Isl. (Chichijima, Hahajima); Volcano Isl. (Minami-Iwojima); Taiwan (Lan Yu Isl.); Nepal; Bhutan; India (Jammu & Kashmir, Arunachal Pradesh, Himachal Pradesh, Sikkim, Tamil Nadu, Uttarakhand)
36 CHI TIB 38 JAP KOR KZN NNS OGA TAI 40 BHU IND NEP
Remarks: will need further splitting in the future because the type of L. mollissima is a tetraploid of southern India fide Kuo et al. 2019. Eastern Asian material is diploid.
Synonyms: hidden
= Gymnogramma mollissima Fisch. ex Kunze; Linnaea 23: 255, 310 (1850), nomen
= Gymnogramma totta var. mollissima (Fisch. ex Kunze) Kunze; Linnaea 24: 249 (1851)
= Lastrea mollissima (Fisch. ex Kunze) Akasawa; Bull. Kochi Women´s Univ., Ser. Nat. Sci. 8: 34 (1960)
= Lastrea mollissima (Fisch. ex Kunze) Okuyama; Col. Ill. Wild. Pl. Japan 6 (Index): 19 (1962), isonym
= Lastrea mollissima f. cristata (Nakai) Okuyama; Col. Ill. Wild. Pl. Japan 6 (Index): 19 (1962)
= Leptogramma mollissima f. cristata (Nakai) Sa.Kurata; Coll. & Cult. Ferns & Fern Allies 306, 341 (1961)
= Leptogramma mollissima f. pilosissima (H.Itô) Nakaike; New Fl. Jap. Pterid. 841 (1992)
= Leptogramma mollissima f. yakusimensis (H.Itô) Sa.Kurata; Coll. & Cult. Ferns & Fern Allies 306, 341 (1961)
= Leptogramma mollissima f. takaoyamensis Hayashi; Bull. Gov. Forest Exp. Sta. 125: 77, t. 7 (1960)
= Leptogramma mollissima var. pilosissima H.Itô; J. Geobot. 7: 41 (1958)
= Leptogramma mollissima var. yakusimensis (H.Itô) Nakaike; New Fl. Jap. Pterid. 841 (1992)
= Leptogramma pozoi f. pilosissima (H.Itô) Nakaike; Enum. Pterid. Jap., Filic. 284 (1975)
= Leptogramma pozoi subsp. mollissima (Kunze) Nakaike; Enum. Pterid. Jap., Filic. 283 (1975)
= Leptogramma pozoi var. yakusimensis (H.Itô) Nakaike; Enum. Pterid. Jap., Filic. 284 (1975)
= Leptogramma totta f. cristata Nakai; Bot. Mag. (Tokyo) 45: 105 (1931)
= Leptogramma totta f. pilosissima (H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 165 (1939)
= Leptogramma totta var. pilosissima H.Itô; Bot. Mag. (Tokyo) 49: 434 (1935)
= Leptogramma totta var. yakusimensis H.Itô; Bot. Mag. (Tokyo) 49: 434 (1935)
= Stegnogramma mollissima (Fisch. ex Kunze) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 237 (1997)
= Stegnogramma mollissima (Kunze) Nakaike; Checkl. Lycophytes Ferns Japan 15 (2013), isonym
= Stegnogramma mollissima f. pilosissima (H.Itô) Nakaike; Checkl. Lycophytes Ferns Japan 16 (2013)
= Stegnogramma mollissima var. truncata Kholia; Ferns Fern-Allies Sikkim 2: 196 (2014)
= Stegnogramma mollissima var. yakusimensis (H.Itô) Nakaike; Checkl. Lycophytes Ferns Japan 16 (2013)
= Stegnogramma pozoi f. pilosissima (H.Itô) Sugim.; Keys Herb. Pl. Jap. 323, 407 (1966)
= Stegnogramma pozoi f. yakusimensis (H.Itô) Sugim.; Keys Herb. Pl. Jap. 322, 407 (1966)
= Stegnogramma pozoi subsp. mollissima (Fisch. ex Kunze) K.Iwats.; Acta Phytotax. Geobot. 19: 125 (1963)
= Stegnogramma pozoi var. mollissima (Fisch. ex Kunze) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 8(1): 50 (1981)
= Thelypteris mollissima (Fisch. ex Kunze) Thapa; Pterid. Nepal 95 (2002)
= Thelypteris mollissima var. truncata Kholia; Ferns Fern-Allies Sikkim 2: 196 (2014)
= Thelypteris pozoi subsp. mollissima (Fisch. ex Kunze) C.V.Morton; Amer. Fern J. 56(4): 178 (1966)
species :
Leptogramma petiolata Ching; Acta Phytotax. Sin. 8: 319 (1963)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Stegnogramma pozoi var. petiolata (Ching) Sledge; Bull. Brit. Mus. (Nat. Hist.), Bot. 8(1): 49 (1981)
= Thelypteris stipitata Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 196 (2008), nom. nov. for Leptogramma petiolata Ching
species :
Leptogramma pilosa (M.Martens & Galeotti) Underw.; Bull. Torrey Bot. Club 29: 626 (1902)
Name: Streak-sorus Fern
Distribution: Mexico (Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala; Honduras; Nicaragua; Panama
79 MEX 80 GUA HON NIC PAN
Synonyms: hidden
= Dryopteris pilosa (M.Martens & Galeotti) C.Chr.; Index Filic. 284 (1905)
= Dryopteris pilosa var. procurrens (Fée) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 7, 10(2): 197 (1913), comb. illeg.
= Dryopteris pseudototta Christ; Bull. Herb. Boissier, Sér. 2, 7: 415 (1907)
= Gymnogramma pilosa M.Martens & Galeotti; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15(5): 27 (1842)
= Gymnogramma pilosa var. major E.Fourn.; Mexic. Pl. 1: 73 (1872)
= Gymnogramma procurrens Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 78 (1857)
= Gymnogramma totta var. procurrens (Fée) Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 2: 515 (1874)
= Lastrea pilosa (M.Martens & Galeotti) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 139 (1947)
= Leptogramma pilosa var. major (E.Fourn.) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 180 (2019)
= Phegopteris pilosa (M.Martens & Galeotti) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 18, no. 30 (1858)
= Phegopteris procurrens (Fée) Mett. ex Salomon; Nomencl. Gefäßkrypt. 277 (1883)
= Stegnogramma pilosa (M.Martens & Galeotti) K.Iwats.; Amer. Fern J. 54: 141 (1964)
= Stegnogramma pilosa var. major (E.Fourn.) K.Iwats.; Amer. Fern J. 54: 141 (1964)
= Thelypteris pilosa (M.Martens & Galeotti) Crawford; Amer. Fern J. 41: 16, t. 3 (1951)
= Thelypteris pilosa var. major (E.Fourn.) Crawford; Amer. Fern J. 41: 19 (1951)
species :
Leptogramma pozoi (Lag.) Heywood; Feddes Repert. Spec. Nov. Regni Veg. 64: 19 (1961)
Distribution: Spain; France; Azores (Sao Miguel Isl., Terceira, Pico, Faial, Flores Isl.); Madeira (Madeira Isl.)
12 FRA SPA 21 AZO MDR
Synonyms: hidden
= Acrostichum pilosiusculum Wikstr.; Kongl. Vetensk. Acad. Handl. 439 (1826)
= Acrostichum pilosum Buch; Phys. Beschr. Canar. Ins. [Buch] 189 (1825), nomen
= Ceterach pozoi (Lag.) A.Braun ex Milde; Bot. Zeitung (Berlin) 24: 310 (1866)
= Cyclosorus pozoi (Lag.) C.M.Kuo; Taiwania 47(2): 172 (2002)
= Gymnogramma lowei Hook. & Grev.; Icon. Filic. t. 89 (1829)
= Gymnogramma pozoi (Lag.) Desv.; Mém. Soc. Linn. Paris 6(2): 216 (1827)
= Hemionitis pozoi Lag.; Gen. Sp. Pl. [Lagasca] 33 (1816)
= Lastrea pozoi (Lag.) Ohwi; J. Jap. Bot. 41: 54 (1966)
= Leptogramma lowei (Hook. & Grev.) J.Sm.; J. Bot. [Hooker] 4: 52 (1841)
= Leptogramma pilosiuscula (Wikstr.) Alston; Bol. Soc. Brot., Ser. 2, 30: 17 (1956)
= Pleurosorus pozoi (Lag.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 245 (1899), isonym
= Pleurosorus pozoi (Lag.) Trevis.; Atti Soc. Ital. Sci. Nat. 17: 256 (1875)
= Polypodium eliasii Sennen & Pau; Bull. Acad. Int. Géogr. Bot., Sér. 3, 20: 94 (1910)
= Stegnogramma pozoi (Lag.) K.Iwats.; Acta Phytotax. Geobot. 29: 124 (1963)
= Thelypteris pozoi (Lag.) C.V.Morton; Bull. Soc. Bot. France 106: 234 (1959)
species :
Leptogramma scallanii (Christ) Ching; Sinensia 7: 101, t. 7 (1936)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); NE-India (Meghalaya, Arunachal Pradesh, ?Manipur); Vietnam
36 CHI 40 IND 41 VIE
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium scallanii Christ; Bull. Soc. Bot. Ital. 1901: 296 (1901)
= Cyclosorus scallanii (Christ) Mazumdar & R.Mukhop.; Bionature 33(1): 30 (2013)
= Dryopteris scallanii (Christ) C.Chr.; Index Filic. 291 (1905)
= Stegnogramma scallanii (Christ) K.Iwats.; Acta Phytotax. Geobot. 19: 124 (1963)
= Thelypteris scallanii (Christ) C.V.Morton; Amer. Fern J. 56: 179 (1966)
species :
Leptogramma sinensis (Ching & W.M.Chu) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 178 (2019)
Distribution: China (NE-Yunnan)
36 CHI
Synonyms: hidden
= Craspedosorus sinensis Ching & W.M.Chu; Acta Phytotax. Sin. 16(4): 34, t. 1 (1978)
= Stegnogramma sinensis (Ching & W.M.Chu) L.J.He & X.C.Zhang; Lycophytes Ferns China 32 (2012)
= Thelypteris sinensis (Ching & W.M.Chu) Christenh.; Global Fl. 4: 38 (2018)
species :
Leptogramma sinica Ching ex Y.X.Lin; Fl. Reipubl. Popularis Sin. 4(1): 323 (1999)
Distribution: China (Hunan, Guizhou)
36 CHI
Synonyms: hidden
= Cyclosorus sinicus (Ching ex Y.X.Lin) Mazumdar & R.Mukhop.; Bionature 33(1): 30 (2013)
= Stegnogramma sinica (Ching ex Y.X.Lin) H.F.Wang, X.Y.Dong & K.P.Ma; Checkl. Vasc. Pl. E. Asia 1: 60 (2025)
= Thelypteris sinica (Ching ex Y.X.Lin) Christenh.; Global Fl. 4: 38 (2018)
species :
Leptogramma stegnogrammopsis (C.F.Reed) Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 3: 355 (2020), et Indian J. Forest. 43(1): 77 (2021)
Distribution: China (NW-Yunnan); Tibet; India (Arunachal Pradesh, Meghalaya, Sikkim, Uttarakhand)
36 CHI TIB 40 IND
Remarks: to be expected in Nepal
Synonyms: hidden
= Dryopteris stenogramme var. asplenioides C.Chr.; Index Filic. 294 (1905), et Acta Horti Gothob. 1: 56 (1924) (non Thelypteris asplenioides (Sw.) Proctor)
= Leptogramma chandrae (Fraser-Jenk.) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 177 (2019)
= Stegnogramma asplenioides (C.Chr.) J.Sm. ex Ching; Sinensia 7: 94, t. 2 (1936), non Sphaerostephanos asplenioides J. Sm. ex Hook., Gen. Fil. [Hooker] t. 24 (1839), nec Stegnogramma asplenioides (J. Sm. ex Hook.) Fée. 1852, nec Thelypteris asplenioides (Sw.) Proctor. 1953
= Thelypteris chandrae Fraser-Jenk.; Taiwania 53(2): 173-209 (2008), nom. nov. superfl. for Dryopteris stegnogramme var. asplenioides C. Chr.
= Thelypteris dasypoda C.V.Morton; Contr. U.S. Natl. Herb. 38: 239 (1973), nom. nov. superfl. for Dryopteris stegnogramme var. asplenioides C. Chr.
= Thelypteris fangornica Christenh.; Global Fl. 4: 30 (2018), nom. nov. superfl.
= Thelypteris stegnogrammopsis C.F.Reed; Phytologia 17: 466 (1969), nom. nov.
species :
Leptogramma subcalcarata (Alderw.) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 181 (2019)
Distribution: Sumatra
42 SUM
Conservation Status: Only known from 2 collections
Synonyms: hidden
= Phegopteris totta var. subcalcarata Alderw.; Malayan Ferns Fern Allies 515 (1917)
= Stegnogramma subcalcarata (Alderw.) Holttum; Fl. Males., Ser. 2, Pterid. 1(5): 543 (1982)
= Thelypteris subcalcarata (Alderw.) Christenh.; Global Fl. 4: 38 (2018)
species :
Leptogramma totta (Schltdl.) J.Sm.; J. Bot. [Hooker] 4: 52 (1841)
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; D.R.Congo [Zaire]; Rwanda; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Zambia; Malawi; Zimbabwe; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Lesotho; Eswatini [Swaziland]; Madagascar; Comores
23 BIO CMN RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 MLW ZAM ZIM 27 LES SAF SWZ 29 COM MDG
Synonyms: hidden
= Aspidium africanum (Desv.) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] ed. 2, 1: 28, 153 (1912)
= Aspidium totta (Schltdl.) Engl.; Hochgebirgsfl. Afrika [Engler] 99 (1892)
= Dryopteris africana (Desv.) C.Chr.; Index Filic. 251 (1905)
= Dryopteris totta (Schltdl.) Masam.; Mem. Fac. Agric. Taihoku Imp. Univ. 11 (Bot. no. 4): 58 (1934)
= Grammitis totta (Schltdl.) C.Presl; Tent. Pterid. 209, t. 9, f. 4 (1836)
= Gymnogramma totta (Willd.) Schltdl.; Adumbr. Pl. 15 (1825)
= Lastrea africana (Desv.) Ching; Contr. Biol. Lab. Sci. Soc. China, Bot. Ser. 9: 37 (1933), nom. superfl.
= Lastrea africana (Desv.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 138 (1947), isonym
= Lastrea totta (Schltdl.) Ohwi; Bull. Natl. Sci. Mus., Tokyo, 3: 98 (1956)
= Leptogramma africana (Desv.) Nakai ex T.Mori; Enum. Pl. Corea 13 (1922), nom. superfl.
= Nephrodium totta (Schltdl.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 170 (1899), as ´tottum´
= Phegopteris totta (Schltdl.) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 302 (1858)
= Polypodium africanum Desv.; Mém. Soc. Linn. Paris 6(2): 239 (1827), nom. nov. for Polypodium tottum Willd., non Thunb. (1800)
= Stegnogramma totta (Schltdl.) Dhir; Biblioth. Pteridol. 2: 70 (1981), without basionym ref.
= Thelypteris totta (Schltdl.) B.K.Nayar & Kaur; Companion Handb. Ferns Brit. India [Nayar & Kaur] 90 (1974) (non (Thunb.) Schelpe)
species :
Leptogramma tottoides Hayata ex H.Itô; Bot. Mag. (Tokyo) 49: 434, f. 7(13) (1935)
Distribution: China (Chongqing, Fujian, Guizhou, Hunan, Jiangxi, Zhejiang); Taiwan
36 CHI 38 TAI
Synonyms: hidden
= Cyclosorus tottoides (Hayata ex H.Itô) C.M.Kuo; Taiwania 47(2): 172 (2002)
= Leptogramma caudata Ching; Sinensia 7: 98, t. 4 (1936)
= Leptogramma totta var. tottoides (Hayata ex H.Itô) H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 166 (1938)
= Stegnogramma tottoides (Hayata ex H.Itô) K.Iwats.; Acta Phytotax. Geobot. 19: 121 (1963)
= Thelypteris caudata (Ching) C.F.Reed; Phytologia 17: 266 (1968)
= Thelypteris tottoides (Hayata ex H.Itô) C.M.Kuo; Taiwania 30: 60 (1985), isonym
= Thelypteris tottoides (Hayata ex H.Itô) C.V.Morton; Amer. Fern J. 56: 179 (1966)
species :
Leptogramma xingwenensis (Ching ex Y.X.Lin) Y.H.Chang & L.Y.Kuo; Cladistics 36(2): 178 (2019)
Distribution: China (Sichuan)
36 CHI
Synonyms: hidden
= Stegnogramma xingwenensis Ching ex Y.X.Lin; Fl. Reipubl. Popularis Sin. 4(1): 350 (1999)
= Thelypteris xingwenensis (Ching ex Y.X.Lin) Christenh.; Global Fl. 4: 40 (2018)
species :
Leptogramma yahanensis Ching ex Y.X.Lin; Fl. Reipubl. Popularis Sin. 4(1): 324 (1999)
Distribution: China (SW-Sichuan)
36 CHI
Synonyms: hidden
= Stegnogramma yahanensis (Ching ex Y.X.Lin) H.F.Wang, X.Y.Dong & K.P.Ma; Checkl. Vasc. Pl. E. Asia 1: 60 (2025)
= Thelypteris yahanensis (Ching ex Y.X.Lin) Christenh.; Global Fl. 4: 40 (2018)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.12 (Dec. 21st, 2025)

Covering 358,226 accepted species,
8,847 hybrids, 48,003 accepted infraspecific taxa and 1,006,142 synonyms,
in total 1,421,218* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life