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481.0000 Family Boraginaceae Juss.; Gen. Pl. [Jussieu] 128 (1789), nom. cons., s. str.
118 genera, ca. 2000 spp.
Species listed in database: 2062; hybrids: 48
= Anchusaceae Vest; Anleit. Stud. Bot. 274, 302 (1818)
= Cerinthaceae Bercht. & J.Presl; Přir. Rostlin 244 (1820)
= Cynoglossaceae Döll; Rhein. Fl. [Döll] 406 (1843)
= Echiaceae Raf.; Fl. Tellur. 2: 61 (1837)
= Onosmaceae Martinov; Tekhno-Bot. Slovar 437 (1820)
481.3825 Genus Pectocarya DC. ex Meisn.; Pl. Vasc. Gen. [Meisner] 79 (1840)
Distribution: Pacific coastal America from British Columbia to Chile
Species listed in database: 12
= Gruvelia A.DC.; Prodr. [A.P. de Candolle] 10: 119 (1846)
= Ktenospermum Lehm.; Index Sem. (Hamburg) 7 (1837)
= Ctenospermum Post & Kuntze; Lex. Gen. Phan. [Post & Kuntze] 152 (1903), nomen
species :
Pectocarya anisocarpa Veno; Aliso 31(1): 3 (2013)
Distribution: USA (California, Arizona, Utah); Mexico (Baja California Norte)
76 USA 79 MEX
species :
Pectocarya anomala I.M.Johnst.; Contr. Gray Herb. 70: 35 (1924)
Distribution: Peru; Chile (Arica y Parinacota, Tarapacá)
83 PER 85 CLE
species :
Pectocarya boliviana (I.M.Johnst.) I.M.Johnst.; Contr. Gray Herb. 78: 118 (1927)
Distribution: Bolivia (La Paz, Oruro, Potosí); Argentina (Jujuy, Salta)
83 BOL 85 ARG
Synonyms: hidden
= Pectocarya gracilis var. boliviana I.M.Johnst.; Contr. Gray Herb. 70: 37 (1924)
species :
Pectocarya dimorpha I.M.Johnst.; Contr. Gray Herb. 78: 115 (1927)
Distribution: Chile (Antofagasta, Atacama, Coquimbo)
85 CLE
Synonyms: hidden
= Pectocarya gracilis var. dimorpha I.M.Johnst.; Contr. Gray Herb. 70: 37 (1924)
species :
Pectocarya heterocarpa (I.M.Johnst.) I.M.Johnst.; J. Arnold Arbor. 20: 399 (1939)
Name: Chuckwalla combseed
Distribution: USA (Arizona, California, New Mexico, Nevada, Texas, Utah); Mexico (Baja California Norte, Sonora)
76 USA 77 USA 79 MEX
Synonyms: hidden
= Pectocarya penicillata var. heterocarpa I.M.Johnst.; Contr. Gray Herb. 70: 37 (1924)
species :
Pectocarya linearis (Ruiz & Pav.) DC.; Prodr. [A.P. de Candolle] 10: 120 (1846)
subspecies :
subsp. linearis
Distribution: Germany [I]; Ecuador; Peru; Argentina (Buenos Aires, Chubut, Mendoza, Neuquén, Rio Negro, Santa Cruz, Tierra del Fuego); Chile (Antofagasta, Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Biobío, Aisén)
11 ger 83 ECU PER 85 ARG CLE
Synonyms: hidden
= Cynoglossum lateriflorum Bertero ex A.DC.; Prodr. [A.P. de Candolle] 10: 120 (1846)
= Cynoglossum lateriflorum Lam.; Encycl. [J.Lamarck et al.] 2: 239 (1786)
= Cynoglossum lineare Ruiz & Pav.; Fl. Peruv. [Ruiz & Pavon] 2: 6 (1799)
= Cynoglossum pilosum Ruiz & Pav.; Fl. Peruv. [Ruiz & Pavon] 2: 6 (1799)
= Echinospermum gracile Lehm.; Pl. Asperif. Nucif. 129 (1818)
= Ktenospermum linifolium Lehm.; Index Sem. (Hamburg) 1837: 5, 7 (1837)
= Mattia lateriflorum (Lam.) G.Don; Gen. Hist. 4: 310 (1837)
= Mattia pilosa (Roem. & Schult.) G.Don; Gen. Hist. 4: 310 (1837)
= Omphalodes lateriflora (Lam.) J.F.Macbr.; Proc. Amer. Acad. Arts 51: 513 (1916)
= Pectocarya chilensis DC.; Prodr. [A.P. de Candolle] 10: 120 (1846)
= Pectocarya gracilis (Ruiz & Pav.) I.M.Johnst.; Contr. Gray Herb. 70: 36 (1924)
= Pectocarya lateriflora (Lam.) DC.; Prodr. [A.P. de Candolle] 10: 120 (1846)
= Pectocarya linearis var. penicillata M.E.Jones; Proc. Calif. Acad. Sci., Ser. 2, 5: 709 (1895)
= Rindera lateriflora Roem. & Schult.; Syst. Veg., ed. 15bis [Roemer & Schultes] 4: 762 (1819)
= Rindera pilosa Roem. & Schult.; Syst. Veg., ed. 15bis [Roemer & Schultes] 4: 762 (1819)
= Rochelia gracilis Roem. & Schult.; Syst. Veg., ed. 15bis [Roemer & Schultes] 4: 111 (1819)
subspecies :
subsp. ferocula (I.M.Johnst.) Thorne; Aliso 9: 191 (1978)
Name: Sagebrush combseed
Distribution: USA (California); Mexico (Baja California Norte, Baja California Sur); Isla Guadalupe
76 USA 79 GLP MEX
Synonyms: hidden
= Pectocarya linearis var. ferocula I.M.Johnst.; Contr. Arnold Arbor. 3: 95 (1932)
species :
Pectocarya penicillata (Hook. & Arn.) A.DC.; Prodr. [A.P. de Candolle] 10: 120 (1846)
Name: Sleeping combseed
Distribution: Canada (British Columbia); USA (Arizona, California, Idaho, Nevada, Oregon, Washington State, Wyoming); Mexico (Baja California Norte)
71 CDN 73 USA 76 USA 79 MEX
Synonyms: hidden
= Ctenospermum penicillatum (Hook. & Arn.) J.Adams; Bull. Dept. Agric. Domin. Canada, n. s. 78: 63 (1927)
= Cynoglossum nuttallii Spreng.; Syst. Veg., ed. 16 [Sprengel] 1: 566 (1824)
= Cynoglossum penicillatum Hook. & Arn.; Bot. Beechey Voy. [Hooker & Arnott] 371 (1839)
= Pectocarya lateriflora var. nuttallii (Spreng.) Brand; Pflanzenreich [Engler] IV.252, Heft 78 (Borag.-Cynogloss.): 95 (1921)
= Pectocarya linearis var. penicillata (Hook. & Arn.) M.E.Jones; Proc. Calif. Acad. Sci., Ser. 2, 5: 709 (1895)
= Pectocarya miser A.Nelson; Bot. Gaz. 37: 278 (1904)
species :
Pectocarya peninsularis I.M.Johnst.; J. Arnold Arbor. 20: 401 (1939)
Name: Peninsular pectocarya
Distribution: USA (California); Mexico (Baja California Norte, Baja California Sur)
76 USA 79 MEX
species :
Pectocarya platycarpa (Munz & I.M.Johnst.) Munz & I.M.Johnst.; Contr. Gray Herb. 81: 81 (1928)
Name: Broadfruit combseed
Distribution: USA (Arizona, California, New Mexico, Nevada, Texas, Utah); Mexico (Baja California Norte, Sonora)
76 USA 77 USA 79 MEX
Synonyms: hidden
= Pectocarya gracilis var. platycarpa Munz & I.M.Johnst.; Contr. Gray Herb. 70: 36 (1924)
= Pectocarya linearis var. platycarpa (Munz & I.M.Johnst.) Cronquist; Vasc. Pl. Pacif. N. W. [C.L.Hitchcock et al.] 4: 235 (1959)
species :
Pectocarya pusilla (A.DC.) A.Gray; Proc. Amer. Acad. Arts 12: 81 (1877)
Name: Little combseed
Distribution: USA (California, Oregon, Washington State); Chile (Valparaíso, O´Higgins)
73 USA 76 USA 85 CLE
Synonyms: hidden
= Gruvelia pusilla A.DC.; Prodr. [A.P. de Candolle] 10: 119 (1846)
= Pectocarya chilensis var. californica Torr.; Pacif. Railr. Rep. 4(5, no. 4), Bot. [Whipple] 124 (1857)
= Pectocarya pusilla var. flagellaris Brand; Pflanzenreich [Engler] IV.252, Heft 78 (Borag.-Cynogloss.): 96 (1921)
species :
Pectocarya recurvata I.M.Johnst.; Contr. Arnold Arbor. 3: 97 (1932)
Name: Curvenut combseed
Distribution: USA (Arizona, California, New Mexico, Nevada, Utah); Mexico (Baja California Norte, Sonora); Isla Guadalupe
76 USA 77 USA 79 GLP MEX
species :
Pectocarya setosa A.Gray; Proc. Amer. Acad. Arts 12: 81 (1877)
Name: Moth combseed
Distribution: USA (Arizona, California, Idaho, New Mexico, Nevada, Oregon, Utah, Washington State); Mexico (Baja California Norte)
73 USA 76 USA 77 USA 79 MEX
Synonyms: hidden
= Gruvelia setosa (A.Gray) Rydb.; Bull. Torrey Bot. Club 40: 479 (1913)
= Pectocarya setosa var. aptera I.M.Johnst.; Contr. Gray Herb. 70: 38 (1924)
= Pectocarya setosa var. holoptera I.M.Johnst.; Contr. Gray Herb. 70: 38 (1924)
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life