034.0000
Family
Dryopteridaceae Herter; Revista Sudamer. Bot. 9: 15 (1949), nom. cons.
26 genera, ca. 2100 spp. Latest phylogeny: Zuo et al. (2025)
Species listed in database: 2183; hybrids: 202
= Aspidiaceae Burnett; Outlines Bot. [Burnett] 324, 1156 (1835), nom. illeg.
= Aspidiaceae Mett. ex A.B.Frank; Syn. Pflanzenk., ed. 2, 3: 1469 (1877), nom. illeg.
= Bolbitidaceae (Pic.Serm.) Ching; Acta Phytotax. Sin. 16: 15 (1978)
= Elaphoglossaceae Pic.Serm.; Webbia 23: 209 (1968)
= Filicaceae Juss.; Gen. Pl. [Jussieu] 14 (1789), as ´Filices´, nom. illeg., rej.
= Peranemataceae (C.Presl) Ching; Sunyatsenia 5: 246 (1940), nom. rej.
034.6005
Genus
Polybotrya Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 99 (1810) s. str.
Distribution: Neotropics, mostly Andes and SE-Brazil
Species listed in database: 39
= Soromanes Fée; Mém. Foug. 2 (Hist. Acrostich.): 16, 82 (1845)
= Amphisoria Trevis.; Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 2, 2: 168 (1851)
= Botryothallus Klotzsch; Bot. Zeitung (Berlin) 104 (1846)
= Eotoneura Fée; Mém. Foug. 2 (Hist. Acrostich.): 75 (1845)
= Granulina Bory ex Fée; Mém. Foug. 5 (Gen. Filic.) 48 (1852)
species :
Polybotrya aequatoriana
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 94 (1987)
Distribution: Ecuador; Bolivia (La Paz)
83 BOL ECU
species :
Polybotrya alata
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34: 92 (1987)
Distribution: Panama
80 PAN
species :
Polybotrya alfredii
Brade; Bradea 1: 12 (1969)
Distribution: Nicaragua; Costa Rica; Panama; Colombia (Antioquia); Ecuador; Peru; Bolivia (La Paz)
80 COS NIC PAN 82 CLM 83 BOL ECU PER
Synonyms: hidden
= Polybotrya alfredii f. carpinterae Brade; Bradea 1(1): 13 (1969)
species :
Polybotrya altescandens
C.Chr.; Index Filic. 7 (1905), 503 (1906)
Distribution: Colombia (Antioquia, Boyacá); Ecuador; Peru
82 CLM 83 ECU PER
Synonyms: hidden
= Acrostichum chrysolepis Sodiro; Crypt. Vasc. Quit. 485 (1893) (non Fée (1869))
species :
Polybotrya andina
C.Chr.; Index Filic. 11 (1905), 504 (1906)
Distribution: Ecuador; Peru
83 ECU PER
Synonyms: hidden
= Acrostichum insigne Baker; J. Bot. 25: 167 (1887) (non Fée (1872))
species :
Polybotrya appressa
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 94 (1987)
Distribution: Ecuador; Peru
83 ECU PER
species :
Polybotrya attenuata
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 74 (1987)
Distribution: Colombia (Antioquia, Nariño); Ecuador; Bolivia (Cochabamba, La Paz)
82 CLM 83 BOL ECU
species :
Polybotrya aureisquama
A.Rojas; Lankesteriana 7(3): 558 (2007)
Distribution: Costa Rica
80 COS
species :
Polybotrya bipinnata
A.Rojas; Brenesia 75-76: 9 (2011)
Distribution: Isla del Coco
80 COC
species :
Polybotrya botryoides
(Baker) C.Chr.; Index Filic. 6 (1905)
Distribution: Colombia (Antioquia); Bolivia (Cochabamba, La Paz)
82 CLM 83 BOL
Synonyms: hidden
= Acrostichum botryoides Baker; J. Bot. 19: 206 (1881)
species :
Polybotrya canaliculata
Klotzsch; Linnaea 20: 429 (1847)
Distribution: Venezuela (Aragua, Falcon, Yaracuy)
82 VEN
Synonyms: hidden
= Acrostichum canaliculatum (Klotzsch) Hook.; Sp. Fil. [W.J.Hooker] 6: 247 (1864)
species :
Polybotrya caudata
Kunze; Linnaea 9: 23 (1834)
Distribution: Mexico (Oaxaca, Chiapas); Guatemala; Belize; Honduras; Nicaragua; Costa Rica; Panama; Trinidad; Colombia (Amazonas, Antioquia, Bolívar, Caldas, Caquetá, Cauca, Chocó, Cundinamarca, Nariño, Santander, Valle, Vaupés); Venezuela (Amazonas, Aragua, Bolívar, Delta Amacuro, Miranda, Portuguesa, Sucre, Tachira); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Beni, Cochabamba, La Paz, Pando, Santa Cruz); Brazil (Acre, Amazonas, Goiás, Maranhão, Mato Grosso do Sul, Mato Grosso, Pará, Roraima)
79 MEX 80 BLZ COS GUA HON NIC PAN 81 TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Acrostichum caudatum (Kunze) Hook.; Sp. Fil. [W.J.Hooker] 6: 244 (1864)
= Acrostichum osmundaceum var. caudatum (Kunze) Krug; Bot. Jahrb. Syst. 24(1): 142 (1897)
= Amphisoria caudata (Kunze) Trevis.; (1851)
= Olfersia caudata (Kunze) Kunze; Linnaea 21: 206 (1848)
= Polybotrya acuminata var. villosa Christ; Prim. Fl. Costaric. 3(1): 9, n9 268 (1901)
= Polybotrya costaricensis Brade; Bradea 1: 11, t. 1, f. 1 (1969)
= Polybotrya villosula Christ; Bull. Herb. Boissier, Sér. 2, 6: 168 (1906)
= Psomiocarpa caudata (Kunze) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 162 (1851)
species :
Polybotrya crassirhizoma
Lellinger; Amer. Fern J. 62: 49 (1972)
Distribution: Colombia (Amazonas); Venezuela (Tachira); Ecuador; Peru; Bolivia (Cochabamba); Brazil (Acre)
82 CLM VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Polybotrya macbridei Lellinger; Amer. Fern J. 62: 51 (1972)
species :
Polybotrya cylindrica
Kaulf.; Enum. Filic. [Kaulfuss] 56 (1824)
Distribution: Brazil (Alagoas, Bahia, Espírito Santo, Pernambuco, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Acrostichum cylindricum (Kaulf.) Hook.; Sp. Fil. [W.J.Hooker] 6: 246 (1864)
= Acrostichum incisum (Link) Endl.; Cat. Horti Vindob. 1: 1 (1842)
= Acrostichum incisum (Link) Hook.; Sp. Fil. [W.J.Hooker] 5: 245 (1864), isonym
= Polybotrya cylindrica var. frondosa (Fée) Brade; (1971)
= Polybotrya frondosa Fée; Crypt. Vasc. Brésil 1: 15 (1869)
= Polybotrya incisa Link; Hort. Berol. [Link] 2: 135 (1833)
= Polybotrya osmundacea var. cylindrica (Kaulf.) Luetzelb.; Estud. Bot. Nordéste 3: 245 (1926)
= Polybotrya osmundacea var. frondosa (Fée) Luetzelb.; Estud. Bot. Nordéste 3: 245 (1926)
= Polybotrya osmundacea var. incisa (Link) Luetzelb.; Estud. Bot. Nordéste 3: 245 (1926)
= Psomiocarpa incisa (Link) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 162 (1851)
species :
Polybotrya espiritosantensis
Brade; Rodriguésia 10(21): 28, t. 3,4(1) (1948)
Distribution: Brazil (Espírito Santo)
84 BRA
species :
Polybotrya fractiserialis
(Baker) J.Sm.; Hist. Fil. 133 (1875)
Distribution: Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Beni, Cochabamba, La Paz, Santa Cruz)
82 FRG GUY SUR 83 BOL ECU PER
Synonyms: hidden
= Acrostichum fractiseriale Baker; Syn. Fil. [W.J.Hooker & Baker] 414 (1868)
= Acrostichum plumbicaule Baker; Syn. Fil. [W.J.Hooker & Baker] 413 (1868)
= Polybotrya plumbicaulis (Baker) J.Sm.; Hist. Fil. 133 (1875)
species :
Polybotrya glandulosa
Mett.; Linnaea 36: 65 (1869)
Distribution: Venezuela (Amazonas); NE-Peru; Brazil (Amazonas)
82 VEN 83 PER 84 BRA
Synonyms: hidden
= Polybotrya subelliptica Lellinger; Amer. Fern J. 62: 56 (1972)
species :
Polybotrya gomezii
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34: 99 (1987)
Distribution: Nicaragua; Costa Rica
80 COS NIC
species :
Polybotrya goyazensis
Brade; Bradea 1: 24, f. 1 (1969)
Distribution: Brazil (Brasilia Distrito Federal, Goiás, Mato Grosso do Sul, Mato Grosso, Pará, São Paulo); Paraguay (Amambay)
84 BRA 85 PAR
Synonyms: hidden
= Polybotrya macedoi Brade; Bradea 1: 24 (1969)
species :
Polybotrya gracilis
Brade; Bradea 1: 14, t. 1, f. 3 &, t. 3 (1969)
Distribution: Costa Rica
80 COS
species :
Polybotrya hickeyi
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 88 (1987)
Distribution: Colombia (Boyacá, Cauca, Cundinamarca, Norte de Santander); Peru; Bolivia (Cochabamba)
82 CLM 83 BOL PER
species :
Polybotrya insularis
A.Rojas; Lankesteriana 7(3): 560 (2007)
Distribution: Isla del Coco
80 COC
species :
Polybotrya latisquamosa
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 108 (1987)
Distribution: Colombia (Meta)
82 CLM
species :
Polybotrya lechleriana
Mett.; Fil. Lechl. 1: 4, t. 1, f. 1-5 (1856)
Distribution: Colombia (Cauca, Chocó, Nariño); Guyana (Mt. Roraima); Ecuador; Peru; Bolivia (Cochabamba, La Paz)
82 CLM GUY 83 BOL ECU PER
Synonyms: hidden
= Acrostichum lechlerianum (Mett.) Hook.; Sp. Fil. [W.J.Hooker] 6: 246 (1864) (non Mett. (1856))
species :
Polybotrya lourteigiana
Lellinger; Proc. Biol. Soc. Washington 89(61): 723 (1977)
Distribution: Colombia (Antioquia, Chocó, Nariño, Valle); Ecuador
82 CLM 83 ECU
species :
Polybotrya matosii
Canestraro & Labiak; Brittonia 67(3): 200 (2015)
Distribution: Brazil (Bahia)
84 BRA
species :
Polybotrya osmundacea
Humb. & Bonpl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 99 (1810)
Distribution: Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti); Guadeloupe; Grenada; Trinidad; Colombia (Amazonas, Antioquia, Boyacá, Cauca, Chocó, Cundinamarca, Magdalena, Meta, Putumayo, Risaralda, Santander, Valle); Venezuela (Amazonas, Anzoategui, Apure, Aragua, Barinas, Venezuela Distrito Federal, Falcon, Lara, Miranda, Monagas, Nueva Esparta, Sucre, Tachira, Yaracuy, Zulia); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Cochabamba, La Paz); Brazil (Roraima, Pará, Amazonas)
80 COS GUA HON NIC PAN 81 CUB GDL GRE HIS JAM TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Acrostichum osmundaceum Hook.; Sp. Fil. [W.J.Hooker] 6: 246 (1864)
= Polybotrya aristeguietae Brade; Bradea 1: 19, t. 1 (1969)
= Polybotrya cyathifolia Fée & L´Herm. ex Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 6, t. 2 (1866)
= Polybotrya trapezoides Link; Fil. Spec. 164 (1841)
= Polybotrya vareschii Brade; Bradea 1: 20, t. 2 (1969)
species :
Polybotrya pilosa
Brade; Bradea 1: 27, t. 1, f. 4 (1969)
Distribution: Brazil (Rio de Janeiro)
84 BRA
species :
Polybotrya pittieri
Lellinger; Proc. Biol. Soc. Washington 89(61): 725 (1977)
Distribution: Colombia (Cauca); Ecuador
82 CLM 83 ECU
species :
Polybotrya polybotryoides
(Baker) Christ; Bull. Herb. Boissier, Sér. 2, 1: 70 (1901)
Distribution: Mexico (Chiapas, Oaxaca, Veracruz); Guatemala; Belize; Honduras; Nicaragua; Costa Rica; Panama; Colombia (Amazonas, Antioquia, Chocó, Nariño, Norte de Santander, Putumayo, Valle); Ecuador; Peru
79 MEX 80 BLZ COS GUA HON NIC PAN 82 CLM 83 ECU PER
Synonyms: hidden
= Acrostichum juglandifolium Baker; J. Bot. 19: 207 (1881) (non Kaulf. (1824))
= Acrostichum polybotryoides Baker; J. Bot. 19: 207 (1881)
= Polybotrya aucuparia Christ; Bull. Herb. Boissier, Sér. 2, 6: 166 (1906)
= Polybotrya juglandifolia (Baker) Christ; Bull. Herb. Boissier, Sér. 2, 4: 965 (1904)
= Polybotrya juglandifolia var. lobata Christ; Bull. Herb. Boissier, Sér. 2, 6: 168 (1906)
= Polybotrya kalbreyeri C.Chr.; Index Filic. 12 (1905), 504 (1906)
species :
Polybotrya pubens
Mart. ex Kunze; Linnaea 9: 22 (1834)
Distribution: Colombia (Amazonas, Caquetá, Putumayo); Ecuador; Peru; Bolivia (Cochabamba, La Paz); Brazil (Acre, Amazonas, Pará)
82 CLM 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Acrostichum caudatum var. pubens (Mart.) Hook. ex Baker; Fl. Bras. [Martius] 1(2): 586 (1870)
= Acrostichum pubens (Mart. ex Kunze) Baker; Syn. Fil. [W.J.Hooker & Baker] 414 (1868)
= Dryopteris guentheri Rosenst.; Repert. Spec. Nov. Regni Veg. 25: 59 (1928)
= Lastrea guentheri (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 139 (1947)
= Polybotrya crespiana Bosco; Nuovo Giorn. Bot. Ital., n. s. 46: 145, t. 9(1) (1938)
= Polybotrya decorata Lellinger; Amer. Fern J. 62: 54 (1972)
= Thelypteris guentheri (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 251 (1941)
species :
Polybotrya puberulenta
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 90 (1987)
Distribution: Ecuador; ?Peru; Bolivia (Cochabamba, La Paz)
83 BOL ECU
species :
Polybotrya semipinnata
Fée; Crypt. Vasc. Brésil 1: 16 (1869)
Distribution: Brazil (Bahia, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Acrostichum acuminatum (Kaulf. ex Link) Hook.; Sp. Fil. [W.J.Hooker] 5: 245 (1864)
= Aspidium scandens Raddi; Pl. Bras. Nov. Gen. 1: 34, t. 49 (1825)
= Polybotrya acuminata Kaulf. ex Link; Hort. Berol. [Link] 2: 135 (1833)
= Polybotrya scandens (Raddi) Christ; Bull. Herb. Boissier, Sér. 2, 4: 965 (1904)
= Psomiocarpa acuminata (Kaulf. ex Link) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 162 (1851)
species :
Polybotrya serratifolia
(Fée) Klotzsch; Linnaea 20: 430 (1847)
Distribution: Trinidad; Colombia (Antioquia, Meta); Venezuela (Aragua, Venezuela Distrito Federal, Falcon, Lara, Merida, Monagas, Portuguesa, Trujillo, Yaracuy)
81 TRT 82 CLM VEN
Synonyms: hidden
= Acrostichum hartii Baker; J. Bot. 26: 371 (1888)
= Polybotrya crassa C.V.Morton; Fieldiana, Bot. 28: 13 (1951)
= Polybotrya hartii (Baker) C.Chr.; Index Filic. 11 (1905), 504 (1906)
= Soromanes integrifolium Fée; Mém. Foug. 2 (Hist. Acrostich.): 82, t. 42 (1845)
= Soromanes serratifolium Fée; Mém. Foug. 2 (Hist. Acrostich.): 82 (1845)
species :
Polybotrya sessilisora
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 108 (1987)
Distribution: Colombia (Caquetá, Vaupés); Venezuela (Bolívar); ?W-Guyana; Brazil (Amazonas)
82 CLM VEN 84 BRA
species :
Polybotrya sorbifolia
Mett.; Linnaea 36: 64 (1869)
Distribution: Costa Rica; Colombia (Boyacá, Casanare, Cundinamarca, Meta, Santander); Venezuela (Amazonas, Aragua, Bolívar, Portuguesa, Yaracuy); Brazil (Alagoas, Goiás, Maranhão, Mato Grosso, Pará, Pernambuco, Roraima)
80 COS 82 CLM VEN 84 BRA
Remarks: to be expected in the Guianas
Conservation Status: IUCN R
Synonyms: hidden
= Acrostichum caenopteris Kunze; Sp. Fil. [W.J.Hooker] 6: 266 (1864)
= Acrostichum caenopteris var. salicifolium Hook.; Sp. Fil. [W.J.Hooker] 5: 257 (1864)
= Botryothallus kunzei Klotzsch; Bot. Zeitung (Berlin) 104 (1846)
= Polybotrya coenopteris (Kunze) Christ; Farnkr. Erde 44 (1897)
= Polybotrya kunzei (Klotzsch) Ettingsh.; Denkschr. Kaiserl. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 22: 66 (1864)
= Polybotrya salicifolia Lellinger; Amer. Fern J. 62: 54 (1972)
= Soromanes caenopteris (Kunze) Christ; Bull. Herb. Boissier, Sér. 2, 3: 613 (1903)
= Soromanes dentatum Fée; Mém. Foug. 2 (Hist. Acrostich.): t. 43 (1845)
species :
Polybotrya speciosa
Schott; Gen. Fil. [Schott] t. 7 (1834)
Distribution: Brazil (Bahia, Espírito Santo, Minas Gerais, Rio de Janeiro, São Paulo)
84 BRA
Synonyms: hidden
= Polybotrya litoralis Brade; Bradea 1: t. 1, f. 2 (1969)
= Polybotrya osmundacea var. crispo-paleacea Rosenst.; Repert. Spec. Nov. Regni Veg. 21: 349 (1925)
= Polybotrya rosenstockiana Brade; Bradea 1: 27. t, 1, f. 3 (1969)
= Polybotrya tomentosa Brade; Arch. Inst. Biol. (São Paulo) 1: 224, f. 2, t. 2, 3 (1935)
species :
Polybotrya stolzei
R.C.Moran; Bull. Illinois Nat. Hist. Surv. 34(1): 76 (1987)
Distribution: Colombia (Antioquia, Boyacá, Chocó, Nariño, Valle); Ecuador
82 CLM 83 ECU
species :
Polybotrya suberecta
(Baker) C.Chr.; Index Filic. 19 (1905)
Distribution: Colombia (Antioquia, Boyacá, Chocó, Cundinamarca, Magdalena, Nariño, Santander); Ecuador; Peru
82 CLM 83 ECU PER
Synonyms: hidden
= Acrostichum hackelianum Sodiro; Crypt. Vasc. Quit. 491 (1893)
= Acrostichum suberectum Baker; J. Bot. 19: 207 (1881)
