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503.0000 Family Plantaginaceae Juss.; Gen. Pl. [Jussieu] 89 (1789), nom. cons.
93 genera, ca. 2050 spp.
Species listed in database: 2097; hybrids: 91
= Antirrhinaceae Pers.; Syn. Pl. [Persoon] 2: 154, 640 (1807)
= Aragoaceae D.Don; Edinburgh New Philos. J. 19: 109, 113 (1835)
= Callitrichaceae Link; Enum. Hort. Berol. Alt. 1: 7 (1821), nom. cons.
= Chelonaceae Martinov; Tekhno-Bot. Slovar 124 (1820)
= Digitalidaceae Martinov; Tekhno-Bot. Slovar 202 (1820)
= Ellisiophyllaceae Honda; Daikou Nippon Shokubutsu Bunraigaku [Syst. Pl. Japon.]: 372 (1930)
= Erinaceae Pfeiff.; Nomencl. Bot. [Pfeiffer] 1: 1236 (1874)
= Globulariaceae DC.; Fl. Franç. [de Candolle & Lamarck], éd. 3, 3: 427 (1805), nom. cons.
= Gratiolaceae Martinov; Tekhno-Bot. Slovar 294 (1820)
= Hippuridaceae Vest; Anleit. Stud. Bot. 265, 278 (1818), nom. cons.
= Linariaceae Bercht. & J.Presl; Přir. Rostlin 243 (1820)
= Littorellaceae Gray; Nat. Arr. Brit. Pl. 2: 290, 294 (1822)
= Oxycladaceae Schnizl.; Iconogr. Fam. Regn. Veg. 2: ad t. 151xx (1857-1870)
= Psylliaceae Horan.; Prim. Lin. Syst. Nat. 69 (1834)
= Sibthorpiaceae D.Don; Edinburgh New Philos. J. 19: 114 (1835)
= Trapellaceae Honda & Sakis.; Daikou Nippon Shokubutsu Bunraigaku [Syst. Pl. Japon.]: 378 (1930)
= Veronicaceae Cassel; Lehrb. Nat. Pflanzenord. 366 (1817)
503.1206 Genus Sophronanthe Benth.; Intr. Nat. Syst. Bot., ed. 2: 445 (1836)
Distribution: North America. Often included in Gratiola but a sister group fide Estes 2008
Species listed in database: 2
= Tragiola Small & Pennell; Man. S.E. Fl. [Small] 1192 (1933)
species :
Sophronanthe hispida Benth.; Intr. Nat. Syst. Bot., ed. 2: 445 (1836)
Name: Rough hedgehyssop (EN)
Distribution: USA (Alabama, Florida, Georgia, Mississippi)
78 USA
Synonyms: hidden
= Gratiola subulata Baldwin ex Benth.; Prodr. [A.P. de Candolle] 10: 405 (1846)
= Gratiola hispida Pollard; Bull. Torrey Bot. Club 24: 157 (1897)
species :
Sophronanthe pilosa (Michx.) Small; Fl. S.E. U.S. [Small] 1067 (1903)
Name: Shaggy hedgehyssop (EN)
Distribution: USA (Alabama, Arkansas, District of Columbia, Delaware, Florida, Georgia, Kentucky, Louisiana, Maryland, Mississippi, North Carolina, New Jersey, Oklahoma, South Carolina, Tennessee, Texas, Virginia)
74 USA 75 USA 77 USA 78 USA
Synonyms: hidden
= Gratiola peruviana Walter; Fl. Carol. [Walter] 62 (1788) non L.
= Gratiola pilosa var. epilis Pennell; Proc. Acad. Nat. Sci. Philadelphia 71: 239 (1920)
= Gratiola pilosa Michx.; Fl. Bor.-Amer. [Michaux] 1: 7 (1803)
= Tragiola pilosa (Michx.) Small & Pennell; Man. S.E. Fl. [Small] 1192 (1933)
= Tragiola pilosa subsp. epilis (Small & Pennell) Small & Pennell; Monogr. Acad. Nat. Sci. Philadelphia 1: 99 (1935)
= Tragiola pilosa var. epilis Small & Pennell; Man. S.E. Fl. [Small] 1192 (1933)
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A complete, synonymic checklist of the Higher Plants of the World


Version 26.03 (March 30th, 2026)

Covering 359,184 accepted species,
8,882 hybrids, 47,737 accepted infraspecific taxa and 1,007,635 synonyms,
in total 1,423,438* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life