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044.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
5-34 genera (depending on genus concept), 1100-1200 spp. New genus concept adopted here from Fawcett & Smith (2021), based on molecular results.
Species listed in database: 1180; hybrids: 39
044.2006 Genus Steiropteris (C.Chr.) Pic.Serm.; Webbia 28: 449 (1973)
Distribution: Neotropics. Status, position and separation from Thelypteris see Salino et al. (2015)
Species listed in database: 28
= Glaphyropteris Fée; Crypt. Vasc. Brésil 2: 40 (1873)
= Dryopteris subgen.Glaphyropteris (Fée) C.Chr.; Biol. Arbejder Tilegnede Eug. Warming 80 (1911)
= Thelypteris subgen.Glaphyropteris (Fée) Alston; J. Wash. Acad. Sci. 48: 234 (1958)
= Thelypteris sect.Glaphyropteris (Fée) C.V.Morton; Amer. Fern J. 51: 31 (1961)
= Thelypteris subgen.Steiropteris (C.Chr.) K.Iwats.; Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 31: 31 (1964)
= Glaphyropteris (Fée) C.Presl ex Fée; Crypt. Vasc. Brésil 2: 40 (1873)
= Glaphyropteris (Fée) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 5: 6: 344 (1848), nomen
= Phegopteris subgen.Glaphyropteris Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 58 (1866)
= Steiropteris subgen.Glaphyropteris (Fée) A.R.Sm.; PhytoKeys 57: 44 (2015)
species :
Steiropteris alstonii Salino & A.R.Sm.; Phytotaxa 340(2): 176 (2018)
Distribution: Colombia (Nariño)
82 CLM
species :
Steiropteris buchtienii (A.R.Sm.) Salino & T.E.Almeida; PhytoKeys 57: 44 (2015)
Distribution: +Bolivia (+La Paz)
83 BOL
Synonyms: hidden
= Thelypteris buchtienii A.R.Sm.; Univ. Calif. Publ. Bot. 76: 10 (1980)
species :
Steiropteris clypeolutata (Desv.) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Nevis; St. Kitts; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada
81 DOM GDL GRE MAR MTS STK STL STV
Synonyms: hidden
= Aspidium asperulum Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 80, t. 23, f. 1 (1866)
= Aspidium capitainei Fée & L´Herm.; Mém. Foug. 11 (Hist. Foug. Antil.): 80, t. 22 f.-3 (1866)
= Aspidium clypeolutatum (Desv.) Mett.; Nomencl. Gefäßkrypt. 47 (1883)
= Aspidium conterminum var. asperulum (Fée) Duss; Div. Foug. Antill. Franç. 100 (1903)
= Aspidium lherminieri Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(2) [Pheg. u. Asp.]: 369 (-370) (1858)
= Aspidium trichophorum Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 81, t. 23, f. 2 (1866)
= Cyclosorus clypeolutatus (Desv.) Christenh.; Bot. J. Linn. Soc. 161(3): 247 (2009)
= Dryopteris clypeolutata (Desv.) Weath.; Contr. Gray Herb. 114: 28 (1936)
= Dryopteris decumbens C.Chr.; Index Filic. 261 (1905)
= Dryopteris lherminieri (Kunze ex Mett.) C.Chr.; Index Filic. 275 (1905)
= Dryopteris trichophora (Fée) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Lastrea lherminieri T.Moore; Index Fil. [T.Moore] 5: 95 (1858) nom. nud.
= Nephrodium clypeolutatum Desv.; Mém. Soc. Linn. Paris 6(2): 258 (1827)
= Nephrodium trichophorum (Fée) Baker; Syn. Fil. [W.J.Hooker & Baker] 265 (1867)
= Thelypteris clypeolutata (Desv.) Proctor; Rhodora 61: 306 (1960)
variant :
var. holmei (Baker) Salino & T.E.Almeida; PhytoKeys 57: 45 (2015)
Distribution: St. Kitts; Montserrat; Dominica; Martinique
81 DOM MAR MTS STK
Synonyms: hidden
= Dryopteris holmei (Baker) C.Chr.; Index Filic. 271 (1905)
= Nephrodium holmei Baker; Ann. Bot. (Oxford) 4: 317 (1891)
= Thelypteris clypeolutata var. holmei (Baker) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 11 (1980)
species :
Steiropteris comosa (C.V.Morton) Salino & T.E.Almeida; PhytoKeys 57: 45 (2015)
Distribution: Peru; Brazil (Acre)
83 PER 84 BRA
Synonyms: hidden
= Cyclosorus comosus (C.V.Morton) Mazumdar & R.Mukhop.; Bionature 33(1): 16 (2013)
= Dryopteris comosa C.V.Morton; J. Wash. Acad. Sci. 28: 528 (1938)
= Thelypteris comosa (C.V.Morton) C.V.Morton; Amer. Fern J. 51: 38 (1961)
species :
Steiropteris connexa (Kuhn ex Baker) comb.ined.
Distribution: French Guiana; ?Brazil
82 FRG
Remarks: generic placement tentative, not mentioned in Fawcett & Smith 2021. Probably a Steiropteris fide Hoess (in litt.)
Synonyms: hidden
= Aspidium connexum (Kuhn ex Baker) Salomon; Nomencl. Gefäßkrypt. 47 (1883)
= Cyclosorus martinii (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 246 (1941)
= Dryopteris connexa (Kuhn ex Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris martinii C.Chr.; Index Filic. 276 (1905)
= Nephrodium connexum Kuhn ex Baker; Fl. Bras. [Martius] 1(2): 489 (1870)
= Thelypteris connexa (Kuhn ex Baker) C.F.Reed; Phytologia 17: 268 (1968)
species :
Steiropteris decussata (L.) A.R.Sm.; PhytoKeys 57: 45 (2015)
Distribution: Guatemala; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Saba; Nevis; St. Kitts; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Trinidad; Colombia (Antioquia, Caldas, Caquetá, Cauca, Chocó, Meta, Nariño, Putumayo, Valle); Venezuela (Aragua, Bolívar, Carabobo, Falcon, Miranda, Sucre, Tachira, Trujillo); Guyana; ?Surinam; French Guiana; Ecuador; Peru; Bolivia (Cochabamba, La Paz); Brazil (Acre, Bahia, Minas Gerais, Pernambuco, Paraná, Santa Catarina, São Paulo)
80 COS GUA NIC PAN 81 CUB DOM GDL GRE HIS JAM MAR MTS PUE SAB STK STL STV TRT 82 CLM FRG GUY VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Aspidium decussatum (L.) Christ; Denkschr. Kaiserl. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 79: 16 (1908)
= Cyclosorus decussatus (L.) Christenh.; Bot. J. Linn. Soc. 161(3): 248 (2009)
= Cyclosorus tatei (Maxon & C.V.Morton) Mazumdar & R.Mukhop.; Bionature 33(1): 32 (2013)
= Dryopteris decussata (L.) Urb.; Symb. Antill. [Urban] 4: 19 (1903)
= Dryopteris percivalii (Jenman) C.Chr.; Index Filic. 284 (1905), nom. nud.
= Dryopteris tatei Maxon & C.V.Morton ex C.V.Morton; J. Wash. Acad. Sci. 28: 529 (1938)
= Glaphyropteris decussata (L.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 5: 344 (1848)
= Glaphyropteris decussata (L.) Fée; Crypt. Vasc. Brésil 2: 40 (1873), isonym
= Gymnogramma decussata (L.) Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 44 (1857)
= Gymnogramma microcarpon Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 43, t. 20, f. 5 (1857)
= Lastrea decussata (L.) C.Presl; Tent. Pterid. 75 (1836)
= Nephrodium decussatum (L.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 171 (1899)
= Phegopteris decussata (L.) Mett.; Fil. Hort. Bot. Lips. 83, t. 17, f. 8 (1856)
= Polypodium decussatum L.; Sp. Pl. [Linnaeus] 2: 1093 (1753)
= Polypodium grammicum Spreng.; Neue Entdeck. Pflanzenk. [Sprengel] 3: 6 (1822)
= Polypodium percivalii Jenman ex Baker; Ann. Bot. (Oxford) 5: 56 (1891), nom. nud.
= Thelypteris decussata (L.) Proctor; Bull. Inst. Jamaica, Sci. Ser. 6: 59 (1953)
= Thelypteris tatei (Maxon & C.V.Morton) C.V.Morton; Amer. Fern J. 51: 38 (1961)
variant :
var. brasiliensis (C.Chr.) Salino & T.E.Almeida; PhytoKeys 57: 45 (2015)
Distribution: Bolivia (La Paz); Brazil (Minas Gerais, Pernambuco, Paraná, Santa Catarina, São Paulo)
83 BOL 84 BRA
Synonyms: hidden
= Dryopteris brasiliensis (C.Chr.) C.V.Morton; J. Wash. Acad. Sci. 28: 529 (1938)
= Dryopteris decussata var. brasiliensis C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 7, 10: 161 (1913)
= Thelypteris brasiliensis (C.Chr.) C.V.Morton; Amer. Fern J. 51: 38 (1961)
= Thelypteris decussata var. brasiliensis (C.Chr.) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 15 (1980)
variant :
var. costaricensis (A.R.Sm.) Salino & T.E.Almeida; PhytoKeys 57: 45 (2015)
Distribution: Belize; Honduras; Costa Rica; Panama
80 BLZ COS HON PAN
Synonyms: hidden
= Thelypteris decussata var. costaricensis A.R.Sm.; Univ. Calif. Publ. Bot. 76: 16 (1980)
variant :
var. mapiriensis (Rosenst.) Salino & T.E.Almeida; PhytoKeys 57: 45 (2015)
Distribution: Peru; Bolivia (Chuquisaca, Cochabamba, La Paz)
83 BOL PER
Synonyms: hidden
= Dryopteris mapiriensis Rosenst.; Repert. Spec. Nov. Regni Veg. 6: 313 (1908)
= Lastrea mapiriensis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 139 (1947)
= Thelypteris decussata var. mapiriensis (Rosenst.) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 16 (1980)
= Thelypteris mapiriensis (Rosenst.) Alston; J. Wash. Acad. Sci. 48: 234 (1958)
variant :
var. velutina (Sodiro) Salino & T.E.Almeida; PhytoKeys 57: 45 (2015)
Distribution: Colombia (Cauca, Chocó, Nariño, Putumayo, Valle); Ecuador; Peru; Bolivia (Cochabamba); ?Brazil
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Dryopteris decussata f. velutina (Sodiro) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 7, 10(2): 160 (1913)
= Polypodium velutinum Sodiro; Recens. Crypt. Vasc. Quit. 59 (1883)
= Thelypteris decussata f. velutina (Sodiro) C.F.Reed; Phytologia 17: 271 (1968)
= Thelypteris decussata var. velutina (Sodiro) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 17 (1980)
species :
Steiropteris deltoidea (Sw.) Pic.Serm.; Webbia 28: 451 (1973)
Name: Deltoid maiden fern
Distribution: Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Virgin Isl. (St. Thomas); Guadeloupe; Martinique
81 CUB GDL HIS JAM MAR PUE VIR
Synonyms: hidden
= Aspidium deltoideum (Sw.) Sw.; J. Bot. [Schrader] 1800(2): 34 (1804)
= Cyclosorus deltoideus (Sw.) Mazumdar & R.Mukhop.; Bionature 33(1): 18 (2013)
= Dryopteris deltoidea (Sw.) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Lastrea deltoidea (Sw.) T.Moore; Index Fil. [T. Moore] 89 (1858)
= Nephrodium deltoideum (Sw.) Desv.; Mém. Soc. Linn. Paris 6(2): 259 (1827)
= Polypodium appendiculatum Poir.; Encycl. [J.Lamarck et al.] 5: 533 (1804)
= Polypodium deltoideum Sw.; Prodr. [Swartz] 133 (1788)
= Thelypteris deltoidea (Sw.) Proctor; Bull. Inst. Jamaica, Sci. Ser. 5: 59 (1953)
species :
Steiropteris fendleri (D.C.Eaton) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Venezuela (Aragua, Carabobo, Monagas, Sucre)
82 VEN
Synonyms: hidden
= Aspidium fendleri D.C.Eaton; Mem. Amer. Acad. Arts, n. s. 8: 210 (1860)
= Cyclosorus fendleri (D.C.Eaton) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 244 (1941)
= Dryopteris fendleri (D.C.Eaton) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Goniopteris fendleri (D.C.Eaton) Alston; J. Wash. Acad. Sci. 48: 232 (1958)
= Nephrodium fendleri Hook.; Sp. Fil. [W.J.Hooker] 4: 82 (1862)
= Thelypteris fendleri (D.C.Eaton) C.F.Reed; Phytologia 17: 276 (1968)
species :
Steiropteris gardneriana (Baker) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Colombia (Cauca, Magdalena); Venezuela (Amazonas); Ecuador; Galapagos; Peru; Bolivia (Cochabamba, La Paz); Brazil (Amazonas, Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
82 CLM VEN 83 BOL ECU GAL PER 84 BRA
Synonyms: hidden
= Cyclosorus gardnerianus (Baker) Mazumdar & R.Mukhop.; Bionature 33(1): 20 (2013)
= Dryopteris densiloba C.Chr.; Index Filic. 261 (1905) (nom. illeg.)
= Dryopteris gardneriana (Kunze) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris supralineata Rosenst.; Repert. Spec. Nov. Regni Veg. 8: 277 (1910)
= Lastrea densiloba (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 138 (1947)
= Nephrodium gardnerianum Baker; Fl. Bras. [Martius] 1(2): 474 (1870)
= Thelypteris gardneriana (Baker) C.F.Reed; Phytologia 17: 278 (1968)
species :
Steiropteris glabra A.R.Sm. & M.Kessler; Phytotaxa 331(1): 30 (2017)
Distribution: Bolivia (La Paz)
83 BOL
Remarks: known only from type
species :
Steiropteris glandulosa (Desv.) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Guatemala; Honduras; Costa Rica; Panama; Nevis; St. Kitts; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Tobago; Trinidad; Colombia (Antioquia, Chocó, Tolima, Valle); Venezuela (Venezuela Distrito Federal, Falcon, Miranda, Nueva Esparta, Sucre); Guyana; Surinam; French Guiana; Ecuador; Galapagos
80 COS GUA HON PAN 81 DOM GDL GRE MAR MTS STK STL STV TRT 82 CLM FRG GUY SUR VEN 83 ECU GAL
Remarks: subgenus Steiropteris
Synonyms: hidden
= Dryopteris dejecta (Jenman) C.Chr.; Index Filic. 261 (1905)
= Dryopteris glandulosa (Desv.) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 7, 10: 171 (1913)
= Goniopteris abbreviata (Sieber) C.Presl; Tent. Pterid. 183 (1836)
= Goniopteris rostrata Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 64, t. 17, f. 3 (1866)
= Lastrea glandulosa (Desv.) Brade; Bradea 1: 222 (1972)
= Nephrodium dejectum Jenman; Gard. Chron., Ser. 3, 18: 640 (1895)
= Nephrodium grayi Jenman; Ferns Brit. W. Ind. 235 (1908)
= Phegopteris abbreviata (Sieber) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): 307 [Pheg. u. Asp. 23, no. 45] (1868)
= Phegopteris plumieri J.Sm.; Bot. Voy. Herald [Seemann] 6: 228 (1854)
= Polypodium abbreviatum Sieber; Exsicc. (Fl. Mixta) no. 168 (1811)
= Polypodium glandulosum Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 317 (1811)
= Thelypteris dejecta (Jenman) C.F.Reed; Phytologia 17: 271 (1968)
= Thelypteris glandulosa (Desv.) Proctor; Rhodora 61: 306 (1960)
variant :
var. brachyodus (Kunze) Salino & T.E.Almeida; PhytoKeys 57: 46 (2015)
Distribution: Mexico (Chiapas); Guatemala; Belize; Honduras; Nicaragua; Costa Rica; Colombia; Ecuador; Galapagos; Peru
79 MEX 80 BLZ COS GUA HON NIC 82 CLM 83 ECU GAL PER
Synonyms: hidden
= Aspidium brachyodus (Kunze) Griseb.; Fl. Brit. W. I. [Grisebach] 693 (1864)
= Aspidium brachyodus var. auriculatum Kuhn; Bot. Jahrb. Syst. 24(1): 120 (1897)
= Cyclosorus brachyodus (Kunze) Farw.; Amer. Midl. Naturalist 12: 258 (1931)
= Dryopteris brachyodus (Kunze) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris glandulosa var. brachyodus (Kunze) C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 7, 10(2): 171 (1913)
= Lastrea brachyodus (Kunze) Brade; Bradea 1: 221 (1972)
= Nephrodium brachyodus (Kunze) Hook.; Sp. Fil. [W.J.Hooker] 4: 83 (1862)
= Phegopteris brachyodus (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 2(2): 307 [Pheg. u. Asp. 21, no. 38] (1858)
= Polypodium brachyodus Kunze; Linnaea 9: 48 (1834)
= Thelypteris brachyodus (Kunze) Ching; Bull. Fan Mem. Inst. Biol., Bot. 6: 286 (1936)
= Thelypteris glandulosa var. brachyodus (Kunze) A.R.Sm.; Phytologia 34(3): 233 (1976)
variant :
var. longipilosa (A.R.Sm.) Salino & T.E.Almeida; PhytoKeys 57: 46 (2015)
Distribution: Costa Rica; Panama; Colombia; Ecuador
80 COS PAN 82 CLM 83 ECU
Synonyms: hidden
= Thelypteris glandulosa var. longipilosa A.R.Sm.; Univ. Calif. Publ. Bot. 76: 22 (1980)
species :
Steiropteris hatschbachii (A.R.Sm.) Salino & T.E.Almeida; PhytoKeys 57: 46 (2015)
Distribution: Brazil (Minas Gerais, Paraná, Rio de Janeiro, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Thelypteris hatschbachii A.R.Sm.; Univ. Calif. Publ. Bot. 76: 22 (1980)
species :
Steiropteris hottensis (C.Chr.) Salino & T.E.Almeida; PhytoKeys 57: 46 (2015)
Distribution: Hispaniola (Haiti)
81 HIS
Synonyms: hidden
= Cyclosorus hottensis (C.Chr.) Mazumdar & R.Mukhop.; Bionature 33(1): 22 (2013)
= Thelypteris hottensis (C.Chr.) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 23 (1980)
= Dryopteris hottensis C.Chr.; Kungl. Svenska Vetenskapsakad. Handl., Ser. 3, 16(2): 26, t. 2(4-6) (1937)
species :
Steiropteris insignis (Mett.) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Colombia; Ecuador
82 CLM 83 ECU
Synonyms: hidden
= Amauropelta insignis (Mett.) Salino & T.E.Almeida; PhytoKeys 57: 24 (2015)
= Aspidium insigne Mett.; Ann. Sci. Nat., Bot., Sér. 5, 2: 247 (1864)
= Cyclosorus insignis (Mett.) Alston; Bull. Jard. Bot. État Bruxelles 27: 57 (1957)
= Dryopteris insignis (Mett.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris lata Hieron.; Hedwigia 46: 327 (1906)
= Nephrodium insigne (Mett.) Baker; Syn. Fil. [W.J.Hooker & Baker] 262 (1867)
= Thelypteris insignis (Mett.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 252 (1941)
species :
Steiropteris leprieurii (Hook.) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Panama; Colombia (Antioquia, Cauca, Chocó, Cundinamarca, Huila, Meta, Putumayo, Valle); Venezuela (Amazonas, Bolívar, Sucre); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (La Paz); Brazil (Amazonas, Brasilia Distrito Federal, Goiás, Minas Gerais, Mato Grosso, Pernambuco, Roraima, São Paulo); Paraguay (Canindeyu)
80 PAN 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA 85 PAR
Synonyms: hidden
= Aspidium leprieurii (Hook.) Christ; Bull. Herb. Boissier, Sér. 2, 6: 161 (1906)
= Cyclosorus leprieurii (Hook.) Mazumdar & R.Mukhop.; Bionature 33(1): 24 (2013)
= Dryopteris leprieurii (Hook.) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Dryopteris leprieurii var. minor Hieron.; Hedwigia 46: 328 (1907)
= Dryopteris leprieurii var. palmensis C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 7, 10(2): 169 (1913)
= Lastrea leprieurii (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 139 (1947)
= Nephrodium leprieurii Hook.; Sp. Fil. [W.J.Hooker] 4: 106 (1862)
= Thelypteris leprieurii (Hook.) R.M.Tryon; Rhodora 69: 6 (1967)
variant :
var. costalis (Baker) A.R.Sm.; PhytoKeys 57: 46 (2015)
Distribution: Venezuela (Aragua)
82 VEN
Synonyms: hidden
= Cyclosorus densisorus (C.Chr.) Mazumdar & R.Mukhop.; Bionature 33(1): 18 (2013)
= Dryopteris costalis (Baker) Kuntze; Revis. Gen. Pl. 2: 812 (1891)
= Dryopteris densisora C.Chr.; Index Filic. 261 (1905), nom. nov. illeg. for Nephrodium costale
= Lastrea densisora (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 138 (1947)
= Nephrodium costale Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 2: 495 (1874)
= Steiropteris densisora (C.Chr.) Pic.Serm.; Webbia 28: 451 (1973)
= Thelypteris densisora (C.Chr.) C.F.Reed; Phytologia 17: 271 (1968)
= Thelypteris leprieurii var. costalis (Baker) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 25 (1980)
variant :
var. glandifera (A.R.Sm) A.R.Sm.; PhytoKeys 57: 46 (2015)
Distribution: Nicaragua; Costa Rica; Panama; Colombia; Peru; Bolivia (Beni, Cochabamba, La Paz); Brazil (Mato Grosso, Goiás, São Paulo)
80 COS NIC PAN 82 CLM 83 BOL PER 84 BRA
Synonyms: hidden
= Thelypteris leprieurii var. glandifera A.R.Sm.; Univ. Calif. Publ. Bot. 76: 25 (1980)
variant :
var. incana (Christ) A.R.Sm.; PhytoKeys 57: 46 (2015)
Distribution: Venezuela (Sucre); Ecuador; Peru; Brazil (Amazonas)
82 VEN 83 ECU PER 84 BRA
Synonyms: hidden
= Aspidium incanum Christ; Hedwigia 44: 367 (1905)
= Cyclosorus incanus (Christ) Mazumdar & R.Mukhop.; Bionature 33(1): 22 (2013)
= Dryopteris incana (Christ) C.Chr.; Index Filic. 272 (1905)
= Steiropteris incana (Christ) Pic.Serm.; Webbia 28: 451 (1973)
= Thelypteris leprieurii var. incana (Christ) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 26 (1980)
variant :
var. subcostalis (A.R.Sm.) A.R.Sm.; PhytoKeys 57: 46 (2015)
Distribution: Costa Rica; Panama
80 COS PAN
Synonyms: hidden
= Thelypteris leprieurii var. subcostalis A.R.Sm.; Univ. Calif. Publ. Bot. 76: 26 (1980)
species :
Steiropteris lonchodes (D.C.Eaton) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Aspidium lonchodes D.C.Eaton; Mem. Amer. Acad. Arts, n. s. 8: 210 (1860)
= Cyclosorus lonchodes (D.C.Eaton) Mazumdar & R.Mukhop.; Bionature 33(1): 24 (2013)
= Dryopteris lonchodes (D.C.Eaton) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium lonchodes (D.C.Eaton) Hook.; Sp. Fil. [W.J.Hooker] 4: 99 (1862)
= Phegopteris lonchodes (D.C.Eaton) Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 52 (1866)
= Thelypteris lonchodes (D.C.Eaton) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 252 (1941)
species :
Steiropteris mexiae (C.Chr. ex Copel.) Salino & T.E.Almeida; PhytoKeys 57: 47 (2015)
Distribution: Brazil (Minas Gerais, Pernambuco, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Dryopteris mexiae C.Chr. ex Copel.; Univ. Calif. Publ. Bot. 17: 32, t. 6 (1932)
= Lastrea mexiae (C.Chr. ex Copel.) Copel.; Gen. Fil. [Copeland] 139 (1941)
= Thelypteris mexiae (C.Chr. ex Copel.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 252 (1941)
species :
Steiropteris nana (A.Rojas) A.Rojas; Phytoneuron 2024-89: 4 (2024)
Distribution: Isla del Coco
80 COC
Remarks: status uncertain, may be a diminutive specimen of Steiropteris gardneriana
Synonyms: hidden
= Thelypteris nana A.Rojas; Brenesia 75-76: 9 (2011)
species :
Steiropteris parva (A.R.Sm. & M.Kessler) Salino & T.E.Almeida; PhytoKeys 57: 47 (2015)
Distribution: Bolivia (Cochabamba, La Paz, Santa Cruz)
83 BOL
Synonyms: hidden
= Thelypteris parva A.R.Sm. & M.Kessler; Brittonia 60(1): 57 (2008)
species :
Steiropteris pennellii (A.R.Sm.) Salino & T.E.Almeida; PhytoKeys 57: 47 (2015)
Distribution: Colombia (Tolima)
82 CLM
Synonyms: hidden
= Cyclosorus pennellii (A.R.Sm.) Mazumdar & R.Mukhop.; Bionature 33(1): 28 (2013)
= Thelypteris pennellii A.R.Sm.; Univ. Calif. Publ. Bot. 76: 28 (1980)
species :
Steiropteris polyphlebia (C.Chr.) Salino & T.E.Almeida; PhytoKeys 57: 47 (2015)
Distribution: Costa Rica; Colombia (Antioquia, Chocó, Nariño, Valle); Ecuador
80 COS 82 CLM 83 ECU
Synonyms: hidden
= Cyclosorus polyphlebius (C.Chr.) Mazumdar & R.Mukhop.; Bionature 33(1): 28 (2013)
= Dryopteris polyphlebia C.Chr.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 7, 10: 161 (1913)
= Thelypteris polyphlebia (C.Chr.) C.V.Morton; Amer. Fern J. 51: 38 (1961)
species :
Steiropteris polypodioides (Raddi) Salino & T.E.Almeida; PhytoKeys 57: 47 (2015)
Distribution: Brazil (Bahia, Espírito Santo, Minas Gerais, Pernambuco, Rio de Janeiro, São Paulo)
84 BRA
Synonyms: hidden
= Ceterach polypodioides Raddi; Opusc. Sci. 3: 284 (1819)
= Dryopteris polypodioides (Raddi) C.Chr.; Index Filic. 285 (1905), non Knowlt. (1898), fossil name.
= Grammitis polypodioides (Raddi) C.Presl; Tent. Pterid. 209 (1836)
= Grammitis severa C.Presl; Tent. Pterid. 209 (1836)
= Gymnogramma aspidioides Desv.; Mém. Soc. Linn. Paris 6(2): 214 (1827)
= Gymnogramma polypodioides (Raddi) Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 40 (1827)
= Gymnogramma raddiana Link; Hort. Berol. [Link] 2: 50 (1833), nom. nov.
= Leptogramma polypodioides (Raddi) J.Sm.; J. Bot. [Hooker] 4: 52 (1841)
= Phegopteris polypodioides (Raddi) Mett.; Fil. Hort. Bot. Lips. 82 (1856) (non Fée (1850))
= Polystichum gymnogrammoides Gaudich.; Voy. Uranie [Freycinet], Bot. 324 (1827)
= Thelypteris polypodioides (Raddi) C.F.Reed; Phytologia 17: 305 (1968)
species :
Steiropteris praetervisa (Kuhn) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Venezuela (Aragua)
82 VEN
Synonyms: hidden
= Aspidium praetervisum Kuhn; Linnaea 36: 111 (1869)
= Cyclosorus praetervisus (Kuhn) Vareschi; Fl. Venez. 1: 443 (1969)
= Dryopteris praetervisa (Kuhn) Kuntze; Revis. Gen. Pl. 2: 813 (1891)
= Nephrodium praetervisum (Kuhn) Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 2: 495 (1874)
= Thelypteris praetervisa (Kuhn) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 29 (1980)
species :
Steiropteris seemannii (J.Sm.) Salino & T.E.Almeida; PhytoKeys 57: 47 (2015)
Distribution: Costa Rica; Colombia (Chocó, Nariño, Valle); Ecuador
80 COS 82 CLM 83 ECU
Synonyms: hidden
= Cyclosorus seemannii (J.Sm.) Mazumdar & R.Mukhop.; Bionature 33(1): 30 (2013)
= Phegopteris seemannii J.Sm.; Bot. Voy. Herald [Seemann] 228 (1854)
= Thelypteris seemannii (J.Sm.) A.R.Sm.; Univ. Calif. Publ. Bot. 76: 30 (1980)
species :
Steiropteris setulosa (A.R.Sm.) A.R.Sm. & S.E.Fawc.; Sida, Bot. Misc. 59: 86 (2021)
Distribution: ?Panama; Colombia (Chocó, Valle)
82 CLM
Synonyms: hidden
= Cyclosorus setulosus (A.R.Sm.) Mazumdar & R.Mukhop.; Bionature 33(1): 30 (2013)
= Thelypteris setulosa A.R.Sm.; Univ. Calif. Publ. Bot. 76: 30 (1980)
species :
Steiropteris valdepilosa (Baker) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Costa Rica; Panama; Colombia (Cauca, Chocó, Valle); Ecuador; Peru; Brazil (Acre)
80 COS PAN 82 CLM 83 ECU PER 84 BRA
Synonyms: hidden
= Cyclosorus valdepilosus (Baker) Mazumdar & R.Mukhop.; Bionature 33(1): 33 (2013)
= Dryopteris valdepilosa (Baker) C.Chr.; Index Filic. 299 (1905)
= Nephrodium valdepilosum Baker; J. Bot. 19: 204 (1881)
= Thelypteris valdepilosa (Baker) C.F.Reed; Phytologia 17: 323 (1968)
species :
Steiropteris villosa (Link) Salino & T.E.Almeida; PhytoKeys 57: 47 (2015)
Distribution: Brazil (Bahia, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Dryopteris dasyphylla C.Chr.; Index Filic. 260 (1905)
= Grammitis villosa (Link) C.Presl; Tent. Pterid. 209 (1836)
= Gymnogramma villosa Link; Hort. Berol. [Link] 2: 51 (1833)
= Lastrea dasyphylla (C.Chr.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 138 (1947)
= Leptogramma villosa (Link) J.Sm.; J. Bot. [Hooker] 4: 52 (1841)
= Phegopteris villosa (Link) Mett.; Fil. Hort. Bot. Lips. 82 (1856)
= Thelypteris villosa (Link) C.F.Reed; Phytologia 17: 323 (1968)
species :
Steiropteris wrightii (Mett. ex D.C.Eaton) Pic.Serm.; Webbia 28: 451 (1973)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Aspidium wrightii Mett. ex D.C.Eaton; Mem. Amer. Acad. Arts, n. s. 8: 210 (1860)
= Cyclosorus wrightii (Mett. ex D.C.Eaton) Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 250 (1941)
= Dryopteris wrightii (Mett. ex D.C.Eaton) Kuntze; Revis. Gen. Pl. 2: 814 (1891)
= Nephrodium wrightii (Mett. ex D.C.Eaton) Hook.; Sp. Fil. [W.J.Hooker] 4: 64, t. 239 (1862)
= Thelypteris wrightii (Mett. ex D.C.Eaton) C.F.Reed; Phytologia 17: 324 (1968)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.11 (Nov. 11th, 2025)

Covering 357,753 accepted species,
8,806 hybrids, 48,075 accepted infraspecific taxa and 1,004,944 synonyms,
in total 1,419,578* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life