492.0000
Family
Rubiaceae Juss.; Gen. Pl. [Jussieu] 196 (1789), nom. cons.
589 genera, ca. 14.200 spp. mostly tropics, comparably few temperate. Phylogeny: Razafimandimbison & Rydin (2024)
Species listed in database: 14370; hybrids: 47
= Aparinaceae Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 2: 3, 38 (1813-1829)
= Asperulaceae Cham. ex Spenn.; Handb. Angew. Bot. 2: 493 (1835)
= Catesbaeaceae Martinov; Tekhno-Bot. Slovar 111 (1820)
= Cephalanthaceae Raf.; Ann. Gén. Sci. Phys. 6: 86 (1820)
= Cinchonaceae Batsch; Tab. Affin. Regni Veg. 234 (1802)
= Coffeaceae Batsch; Tab. Affin. Regni Veg. 233 (1802)
= Coutareaceae Martinov; Tekhno-Bot. Slovar 168 (1820)
= Dialypetalanthaceae Rizzini & Occhioni; Lilloa 17: 253 (1948), nom. cons.
= Galiaceae Lindl.; Intr. Nat. Syst. Bot., ed. 2: 249 (1836)
= Gardeniaceae Dumort.; Anal. Fam. Pl. 29, 32 (1829)
= Guettardaceae Batsch; Tab. Affin. Regni Veg. 235 (1802)
= Hameliaceae Mart.; Consp. Regn. Veg. [Martius] 31 (1835)
= Hedyotidaceae Dumort.; Commentat. Bot. [Dumortier] 57 (1822)
= Henriqueziaceae Bremek.; Acta Bot. Neerl. 6 (Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 141): 371 (1957)
= Houstoniaceae Raf.; Good Book 21 (1840)
= Hydrophylacaceae Martinov; Tekhno-Bot. Slovar 318 (1820)
= Lippayaceae Meisn.; Pl. Vasc. Gen. [Meisner], Tab. Diagn. 156, Comm. 112 (1838)
= Lygodisodeaceae Bartl.; Ord. Nat. Pl. 123, 207 (1830)
= Naucleaceae Wernham; New Phytol. 11: 225 (1912)
= Nonateliaceae Martinov; Tekhno-Bot. Slovar 420 (1820)
= Operculariaceae Juss. ex Perleb; Vers. Arzneikr. Pfl. 207 (1818)
= Pagamaeaceae Martinov; Tekhno-Bot. Slovar 447 (1820)
= Psychotriaceae F.Rudolphi; Syst. Orb. Veg. 49 (1830)
= Randiaceae Martinov; Tekhno-Bot. Slovar 534 (1820)
= Sabiceaceae Martinov; Tekhno-Bot. Slovar 555 (1820)
= Spermacoceaceae Bercht. & J.Presl; Přir. Rostlin 256 (1820)
= Theligonaceae Dumort.; Anal. Fam. Pl. 15, 17 (1829), nom. cons.
492.5609
Genus
Timonius Rumph. ex DC.; Prodr. [A.P. de Candolle] 4: 461 (1830), nom. cons.
Distribution: Indian Ocean through tropical Asia to Pacific, mostly Malesia (very many in Borneo)
Species listed in database: 265
= Abbottia F.Muell.; Fragm. [Mueller] 9: 181 (1875)
= Burneya Cham. & Schltdl.; Linnaea 4: 189 (1829), nom. rej.
= Erithalis G.Forst.; Fl. Ins. Austr. 17 (1786), nom. illeg.
= Eupyrena Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 422 (1834)
= Helospora Jacq.; Trans. Linn. Soc. London 14: 127 (1824), nom. rej.
= Polyphragmon Desf.; Mém. Mus. Hist. Nat. 6: 5 (1820), nom. rej.
= Porocarpus Gaertn.; Fruct. Sem. Pl. 2: 473 (1791), nom. rej.
species :
Timonius abanii
Jun H.Chen; Phytotaxa 181(3): 139 (2014)
Distribution: Borneo (Sabah, Sarawak); Brunei
42 BOR BRN
species :
Timonius affinis
A.Gray; Proc. Amer. Acad. Arts 4: 36 (1860)
Distribution: Solomon Isl.; Santa Cruz Isl.; Vanuatu; Fiji
43 SOL 60 FIJ SCZ VAN
variant :
var. samoensis
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 28 (1994)
Distribution: [Western] Samoa (Savai´i, ´Upolu)
60 SAM
variant :
var. sapotifolius
(A.Gray) Fosberg; Sargentia 1: 121 (1942)
Distribution: Fiji
60 FIJ
Synonyms: hidden
= Timonius sapotifolius A.Gray; Proc. Amer. Acad. Arts 4: 35 (1860)
species :
Timonius akunensis
S.P.Darwin; Candollea 65(2): 222 (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius albus
Volkens; Bot. Jahrb. Syst. 31: 475 (1901)
Distribution: Fed. States of Micronesia (Yap)
62 MIC
species :
Timonius alejandroanus
J.G.Chavez, Meve & Liede; Nordic J. Bot. 38(7)-e02730: 2 (2020)
Distribution: Philippines
42 PHI
species :
Timonius alius
S.P.Darwin; Brittonia 62(2): 127 (2010)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius amboinicus
(Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891)
Distribution: Moluccas (Ambon)
42 MOL
Synonyms: hidden
= Polyphragmon amboinicum Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 242 (1869)
species :
Timonius amungwiwanensis
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 29 (1994)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius anodon
(Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891)
Distribution: Moluccas (Ambon)
42 MOL
Remarks: probably in Guettardella
Synonyms: hidden
= Antirhea anodon (Miq.) Chaw; Tulane Stud. Zool. & Bot. 28(2): 60 (1992)
= Polyphragmon anodon Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 242 (1869)
species :
Timonius appendiculatus
Merr.; Philipp. J. Sci. 4: 327 (1909)
Distribution: Philippines (Luzon, Mindoro)
42 PHI
species :
Timonius arabii
P.K.Hoo & S.Y.Wong; Nordic J. Bot. 2025(6)-e04595: 2 (2025)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius arboreus
Elmer; Leafl. Philipp. Bot. 1: 72 (1906)
Distribution: Taiwan (Lan Yu Isl., Lutao Isl.); Palawan; Philippines (Luzon, Mindoro, Sibuyan, Ticao, Panay, Guimaras, Negros, Leyte, Mindanao)
38 TAI 42 PHI PLW
Synonyms: hidden
= Timonius gamillii Elmer; Leafl. Philipp. Bot. 4: 1359 (1912)
species :
Timonius argenteus
Valeton; Bot. Jahrb. Syst. 61: 35 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius ariffinii
Junhao Chen & K.M.Wong; Timonius Borneo 74 (2023)
Distribution: Brunei
42 BRN
species :
Timonius avenis
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 46 (1909)
Distribution: Borneo (E-Kalimantan); Sulawesi; Moluccas; New Guinea (Irian Jaya, Papua New Guinea)
42 BOR MOL NWG SUL
variant :
var. magnifructus
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 36 (1994)
Distribution: New Guinea (Irian Jaya)
42 NWG
variant :
var. pubipetalus
Valeton; Bot. Jahrb. Syst. 61: 37 (1927)
Distribution: New Guinea
42 NWG
Synonyms: hidden
= Timonius pubipetalus (Valeton) Merr. & L.M.Perry; J. Arnold Arbor. 26: 236 (1945)
variant :
var. vogelkopensis
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 37 (1994)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius azlanii
Junhao Chen & K.M.Wong; Timonius Borneo 78 (2023)
Distribution: Borneo (Sarawak, Sabah); Brunei
42 BOR BRN
species :
Timonius balansanus
(Baill.) Schltr.; Bot. Jahrb. Syst. 39: 257 (1906)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Guettarda balansana Baill.; Adansonia 12: 238 (1879)
species :
Timonius baziae
Junhao Chen & K.M.Wong; Timonius Borneo 63 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius beamanii
K.M.Wong & Jun H.Chen; Phytotaxa 181(3): 141 (2014)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius belensis
Merr. & L.M.Perry; J. Arnold Arbor. 26: 236 (1945)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius benomensis
P.K.Hoo & Sugumaran; Sandakania 27: 64 (2025)
Distribution: peninsular Malaysia
42 MLY
species :
Timonius berhamanii
Junhao Chen & K.M.Wong; Timonius Borneo 87 (2023)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius billitonensis
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 59 (1909)
Distribution: Sumatera (Belitung Isl.); Borneo (Sarawak, Kalimantan); Brunei
42 BOR BRN SUM
species :
Timonius bismarckensis
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 40 (1994)
Distribution: Bismarck Arch.
43 BIS
species :
Timonius bogoriensis
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 43 (1909)
Distribution: ?Malesia
Error
species :
Timonius borneensis
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 48 (1909)
Distribution: Sumatera; Borneo (Kalimantan, Sabah, Sarawak); Brunei; Sulawesi
42 BOR BRN SUL SUM
species :
Timonius bosaviensis
S.P.Darwin; Brittonia 62(2): 128 (2010)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius bougainvillensis
Merr. & L.M.Perry; J. Arnold Arbor. 26: 249 (1945)
Distribution: Bougainville
43 SOL
species :
Timonius boycei
Junhao Chen & K.M.Wong; Timonius Borneo 102 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius bracteatus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 246 (1945)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius bracteoides
S.P.Darwin; Candollea 65(2): 224 (-225, fig.2) (2010)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius branderhorstii
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 33 (1909)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Timonius strumarius Merr. & L.M.Perry; J. Arnold Arbor. 26: 248 (1945)
species :
Timonius bubuensis
P.K.Hoo & Sugumaran; Sandakania 27: 66 (2025)
Distribution: peninsular Malaysia
42 MLY
species :
Timonius budaensis
Junhao Chen & K.M.Wong; Timonius Borneo 107 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius bullatus
K.M.Wong & Jun H.Chen; Phytotaxa 181(3): 143 (2014)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius buwaldae
Junhao Chen & K.M.Wong; Timonius Borneo 112 (2023)
Distribution: Borneo (C-Kalimantan)
42 BOR
species :
Timonius calcicola
Junhao Chen & K.M.Wong; Timonius Borneo 114 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius carii
S.P.Darwin; Candollea 65(2): 226 (-227, fig.3) (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius carstensensis
Wernham; Trans. Linn. Soc. London, Bot., Ser. 2, 9: 73 (1916)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius caudatifolius
Elmer; Leafl. Philipp. Bot. 5: 1898 (1913)
Distribution: N-Sulawesi; Philippines (Mindanao)
42 PHI SUL
species :
Timonius caudatus
Valeton; Bot. Jahrb. Syst. 61: 38 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius celebicus
Koord.; Natuurk. Tijdschr. Ned.-Indië 63: 87 (1897)
Distribution: Sulawesi
42 SUL
species :
Timonius chaianthus
Junhao Chen & K.M.Wong; Timonius Borneo 117 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius chanii
Junhao Chen & K.M.Wong; Timonius Borneo 120 (2023)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius cheechiewii
Junhao Chen & K.M.Wong; Timonius Borneo 123 (2023)
Distribution: Borneo (Sarawak, E-Kalimantan)
42 BOR
species :
Timonius chinanthus
Junhao Chen & K.M.Wong; Timonius Borneo 127 (2023)
Distribution: Borneo (W-Kalimantan)
42 BOR
species :
Timonius christinae
Junhao Chen & K.M.Wong; Timonius Borneo 131 (2023)
Distribution: Borneo (Sarawak: Hose Mts.)
42 BOR
species :
Timonius clathratus
S.P.Darwin; Candollea 65(2): 227 (-229, fig.4) (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius clementis
Merr.; J. Straits Branch Roy. Asiat. Soc. 77: 244 (1917)
Distribution: Borneo (Sabah: Mt. Kinabalu)
42 BOR
species :
Timonius collinus
Junhao Chen & K.M.Wong; Timonius Borneo 136 (2023)
Distribution: Borneo (Sarawak: Kelabit Highlands)
42 BOR
species :
Timonius confertiflorus
Merr.; Philipp. J. Sci., C 10: 114 (1915)
Distribution: Philippines (Samar, Mindanao)
42 PHI
species :
Timonius coodei
Junhao Chen & K.M.Wong; Timonius Borneo 140 (2023)
Distribution: Borneo (C-Kalimantan, Sarawak); Brunei
42 BOR BRN
species :
Timonius corneri
K.M.Wong; Kew Bull. 43(3): 500 (1988)
Distribution: Thailand; peninsular Malaysia (Kelantan, Terengganu)
41 THA 42 MLY
variant :
var. penangianus
(Ridl.) K.M.Wong; Kew Bull. 43(3): 502 (1988)
Distribution: Thailand; peninsular Malaysia (Kedah, Penang, Perak, Pahang)
41 THA 42 MLY
Synonyms: hidden
= Timonius wallichianus var. penangianus Ridl.; Fl. Malay Penins. 2: 116 (1923)
species :
Timonius corymbosus
Valeton; Bot. Jahrb. Syst. 63: 308 (1930)
Distribution: Palau (Babeldaob, Koror, Peleliu, Kaiguru, Ulebsechel)
62 PLU
Synonyms: hidden
= Timonius korrensis Kaneh.; Bot. Mag. (Tokyo) 45: 351 (1931)
variant :
var. takamatsui
Fosberg & Sachet; Micronesica 20(1-2): 159 (1987)
Distribution: Palau
62 PLU
species :
Timonius creber
S.P.Darwin; Brittonia 62(2): 130 (2010)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius cryptophlebus
S.Moore; Proc. Roy. Soc. Queensland 34: 55 (1922)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius cuneatus
Warb.; Bot. Jahrb. Syst. 13: 433 (1891)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius cyathicalyx
S.P.Darwin; Brittonia 62(2): 131 (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius decipiens
Merr. & L.M.Perry; J. Arnold Arbor. 26: 246 (1945)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius denigricans
P.K.Hoo; Phytotaxa 599(3): 178 (2023)
Distribution: peninsular Malaysia
42 MLY
species :
Timonius densiflorus
Valeton; Bot. Jahrb. Syst. 61: 39 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius devogelii
Junhao Chen & K.M.Wong; Timonius Borneo 145 (2023)
Distribution: Borneo (Sarawak: Gunung Serapi)
42 BOR
species :
Timonius diffusus
Ridl.; J. Fed. Malay States Mus. 4: 32 (1909)
Distribution: peninsular Malaysia (Pahang, Selangor)
42 MLY
species :
Timonius dolichophyllus
(Merr. & L.M.Perry) S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 47 (1994)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Timonius trichanthus var. dolichophyllus Merr. & L.M.Perry; J. Arnold Arbor. 26: 239 (1945)
species :
Timonius dumagat
J.G.Chavez; Nordic J. Bot. 38(7)-e02730: 3 (2020)
Distribution: Philippines
42 PHI
species :
Timonius elegans
S.P.Darwin; Syst. Bot. 22(1): 93 (1997)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius elisabethiae
Junhao Chen & K.M.Wong; Timonius Borneo 147 (2023)
Distribution: Borneo (Sarawak: Kuching)
42 BOR
species :
Timonius enderianus
Warb.; Bot. Jahrb. Syst. 13: 434 (1891)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius endertii
Junhao Chen & K.M.Wong; Timonius Borneo 149 (2023)
Distribution: Borneo (Kalimantan: Kutai)
42 BOR
species :
Timonius epiphyticus
Elmer; Leafl. Philipp. Bot. 3: 1011 (1911)
Distribution: Philippines (Mindanao, Basilan)
42 PHI
species :
Timonius eranthus
Junhao Chen & K.M.Wong; Timonius Borneo 152 (2023)
Distribution: Borneo (Kalimantan, Sabah)
42 BOR
species :
Timonius eremiticus
J.G.Chavez & Banag; Blumea 65(2): 105 (2020)
Distribution: Philippines
42 PHI
species :
Timonius esherianus
W.W.Sm.; Notes Roy. Bot. Gard. Edinburgh 8: 327 (1915)
Distribution: Borneo (Sarawak); Brunei
42 BOR BRN
species :
Timonius eximius
(Baill.) Schltr.; Bot. Jahrb. Syst. 39: 257 (1906)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Guettarda eximia Baill.; Adansonia 12: 241 (1879)
species :
Timonius ferrugineus
Merr.; Philipp. J. Sci., C 10: 115 (1915)
Distribution: Borneo; Palawan
42 BOR PLW
species :
Timonius filipes
Wernham; Trans. Linn. Soc. London, Bot., Ser. 2, 9: 73 (1916)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius finlaysonianus
(Wall. ex G.Don) Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 127 (1880)
Distribution: Nicobar Isl. (Great Nicobar Isl.); peninsular Malaysia (all coasts); Singapore; Sumatera; Borneo; Philippines; Jawa
41 NCB 42 BOR JAW MLY PHI SNG SUM
Conservation Status: critically endangered (Singapore)
Synonyms: hidden
= Guettarda finlaysoniana Wall. ex G.Don; Gen. Hist. 3: 552 (1834), descr.
= Polyphragmon compressicaule Miq.; Fl. Ned. Ind. 2: 235 (1857)
= Polyphragmon compressicaule var. floribunda Miq.; Fl. Ned. Ind. 2: 235 (1857)
= Timonius compressicaulis (Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891)
= Timonius jambosella var. finlaysoniana (Wall. ex G.Don) King & Gamble; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 54 (1904)
species :
Timonius flavescens
(Jack) Baker; Fl. Mauritius [J.G.Baker] 144 (1877)
Distribution: Seychelles; Sri Lanka; Andaman Isl. (South Andaman Isl.); Nicobar Isl. (Car Nicobar Isl., North Nicobar Isl., Central Nicobar Isl., Great Nicobar Isl., Little Nicobar Isl.); Myanmar; Thailand; Vietnam; peninsular Malaysia (throughout); Singapore; Sumatera; Borneo (Kalimantan, Sabah, Sarawak); Brunei; New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl.
29 SEY 40 SRL 41 AND MYA NCB THA VIE 42 BOR BRN MLY NWG SNG SUM 43 SOL
Conservation Status: critically endangered (Singapore)
Synonyms: hidden
= Antirhea borneensis Valeton; Beih. Bot. Centralbl. 34(2): 43 (1916)
= Bobea glabra (Wight & Arn.) Korth.; Ned. Kruidk. Arch. 2(2): 211 (1851)
= Bobea inaequisepala Miq.; Fl. Ned. Ind., Eerste Bijv. 1: 545 (1860)
= Bobea sericantha f. glabrior Miq.; Fl. Ned. Ind., Eerste Bijv. 1: 545 (1860)
= Bobea sericantha Miq.; Fl. Ned. Ind. 2: 355 (1857)
= Eupyrena glabra Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 423 (1834)
= Guettarda brunonis Wall. ex G.Don; Gen. Hist. 3: 552 (1834)
= Guettarda missionis Wall. ex G.Don; Gen. Hist. 3: 550 (1834)
= Guettarda peduncularis Wall. ex G.Don; Gen. Hist. 3: 551 (1834)
= Guettarda pervilleana Baill.; Adansonia 12: 319 (1879)
= Helospora flavescens Jacq.; Trans. Linn. Soc. London 14: 127, t. 4, f. 3 (1824)
= Polyphragmon flavescens (Jacq.) Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 45(2): 134 (1876)
= Polyphragmon flavescens var. macrocarpum Kurz; J. Asiat. Soc. Bengal 46(2): 154 (1877)
= Polyphragmon mutabile (Korth.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 240 (1869)
= Polyphragmon sericanthum (Miq.) Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 242 (1869)
= Timonius flavescens var. brevipetiolatus Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 35 (1909)
= Timonius flavescens var. grandiflorus Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 35 (1909)
= Timonius jambosella Thwaites; Enum. Pl. Zeyl. [Thwaites] 153 (1859)
= Timonius koenigii Blume ex Korth.; Ned. Kruidk. Arch. 2(2): 211 (1851), nom. inval.
= Timonius peduncularis (Wall. ex G.Don) Ridl.; Fl. Malay Penins. 2: 113 (1923)
= Timonius sechellensis Summerh.; Bull. Misc. Inform. Kew 1928: 391 (1928)
= Timonius sericanthus (Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891)
= Timonius sericanthus var. inaequisepalus (Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891)
species :
Timonius fuscus
(Korth.) Walp.; Ann. Bot. Syst. [Walpers] 2: 765 (1852)
Distribution: Sumatera
42 SUM
Synonyms: hidden
= Bobea fusca Korth.; Ned. Kruidk. Arch. 2: 213 (1851)
= Timonius fuscus Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891), et Bull. Dep. Agric. Indes Neerl. 26: 60 (1909)
species :
Timonius gammillii
Elmer; Leafl. Philipp. Bot. 3: 1013 (1911)
Distribution: Philippines (Sibuyan)
42 PHI
species :
Timonius ganesanii
Junhao Chen & K.M.Wong; Timonius Borneo 170 (2023)
Distribution: Borneo (C-Sarawak)
42 BOR
species :
Timonius glabrior
(Valeton) K.M.Wong; Kew Bull. 43(3): 505 (1988)
Distribution: peninsular Malaysia (Perak)
42 MLY
Synonyms: hidden
= Timonius wallichianus var. glabrior Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 37 (1909)
species :
Timonius gracilipes
Merr.; Philipp. J. Sci., C 8: 60 (1913)
Distribution: Philippines (Cebu)
42 PHI
species :
Timonius grandifolius
Valeton; Bot. Jahrb. Syst. 61: 41 (1927)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius hallieri
Junhao Chen & K.M.Wong; Timonius Borneo 174 (2023)
Distribution: Borneo (W-Kalimantan)
42 BOR
species :
Timonius hentyi
S.P.Darwin; Syst. Bot. 22(1): 92 (1997)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
species :
Timonius heptamerus
Wernham; Trans. Linn. Soc. London, Bot., Ser. 2, 9: 73 (1916)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius hughtanii
Junhao Chen & K.M.Wong; Nat. Singapore Suppl.1(e2022071): 34 (2022)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius hydrangeifolius
(Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891)
Distribution: Sumatera
42 SUM
Synonyms: hidden
= Polyphragmon hydrangeifolium Miq.; Fl. Ned. Ind. 2: 235 (1857)
species :
Timonius imitans
Merr. & L.M.Perry; J. Arnold Arbor. 26: 245 (1945)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius insularis
S.P.Darwin; Brittonia 62(2): 133 (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius involucratus
Merr.; J. Straits Branch Roy. Asiat. Soc. 77: 241 (1917)
Distribution: Borneo (SW-Sarawak, W-Kalimantan); Brunei
42 BOR BRN
species :
Timonius jangarunii
Junhao Chen & K.M.Wong; Timonius Borneo 189 (2023)
Distribution: Brunei
42 BRN
species :
Timonius jiewhoei
Junhao Chen & K.M.Wong; Timonius Borneo 192 (2023)
Distribution: Borneo (Sarawak); Brunei
42 BOR BRN
species :
Timonius jobiensis
Wernham; J. Bot. 56: 131 (1918)
Distribution: New Guinea (Japen Isl.)
42 NWG
species :
Timonius joffreanthus
Junhao Chen & K.M.Wong; Timonius Borneo 198 (2023)
Distribution: Brunei
42 BRN
species :
Timonius juliasangiae
Junhao Chen & K.M.Wong; Timonius Borneo 199 (2023)
Distribution: Borneo (E-Sabah)
42 BOR
species :
Timonius kalimantanensis
Junhao Chen & K.M.Wong; Timonius Borneo 203 (2023)
Distribution: Borneo (C-Kalimantan, W-Kalimantan)
42 BOR
species :
Timonius kaniensis
Valeton; Bot. Jahrb. Syst. 61: 42 (1927)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius keningauensis
Junhao Chen & K.M.Wong; Timonius Borneo 208 (2023)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius keranjiensis
Junhao Chen & K.M.Wong; Timonius Borneo 210 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius kessleri
Junhao Chen & K.M.Wong; Timonius Borneo 213 (2023)
Distribution: Borneo (E-Kalimantan)
42 BOR
species :
Timonius khoonmengii
P.K.Hoo & Sugumaran; Sandakania 27: 76 (2025)
Distribution: peninsular Malaysia
42 MLY
species :
Timonius kiahiae
Junhao Chen & K.M.Wong; Timonius Borneo 217 (2023)
Distribution: Brunei
42 BRN
species :
Timonius kinabaluensis
Jun H.Chen; Phytotaxa 181(3): 144 (2014)
Distribution: Borneo (Sabah: Mt. Kinabalu)
42 BOR
species :
Timonius klossii
Wernham; Trans. Linn. Soc. London, Bot., Ser. 2, 9: 74 (1916)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius koikokoensis
Wernham; J. Bot. 56: 130 (1918)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius koordersii
Valeton; Icon. Bogor. [Boerlage] t. 297 (1909)
Distribution: Sulawesi
42 SUL
species :
Timonius korthalsii
Junhao Chen & K.M.Wong; Timonius Borneo 221 (2023)
Distribution: Borneo (S-Kalimantan)
42 BOR
species :
Timonius kostermansii
S.P.Darwin; Syst. Bot. 22(1): 95 (1997)
Distribution: Moluccas (Morotai)
42 MOL
species :
Timonius kuteiensis
Junhao Chen & K.M.Wong; Timonius Borneo 224 (2023)
Distribution: Borneo (E-Kalimantan)
42 BOR
species :
Timonius laevigatus
Valeton; Bot. Jahrb. Syst. 61: 43 (1927)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius lagenifer
S.P.Darwin; Allertonia 7(1): 33 (1993), as ´lageniferus´
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius lamii
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 49 (1994)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius lanceolatus
Merr.; Philipp. J. Sci. 17: 318 (1920 publ. 1921)
Distribution: Philippines (Dinagat, Mindanao)
42 PHI
species :
Timonius lasianthoides
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 48 (1909)
Distribution: Borneo (SW-Sabah, Sarawak, W-Kalimantan, E-Kalimantan); Brunei
42 BOR BRN
species :
Timonius latifolius
K.Schum. & Lauterb.; Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 568 (1900)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius laugerioides
Wernham; J. Bot. 56: 131 (1918)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius lawalawaensis
Junhao Chen & K.M.Wong; Timonius Borneo 235 (2023)
Distribution: Borneo (E-Sabah)
42 BOR
species :
Timonius laxus
King & Gamble; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 56 (1904)
Distribution: peninsular Malaysia (only known from type, Ulu Bubong, Perak)
42 MLY
species :
Timonius ledermannii
Valeton; Bot. Jahrb. Syst. 63: 305 (1930)
Distribution: Fed. States of Micronesia (Truk, Tol, Pohnpei)
62 MIC
species :
Timonius leopoldii
Jun H.Chen & K.M.Wong; Phytotaxa 181(3): 145 (2014)
Distribution: Borneo (Sabah: Mt. Kinabalu)
42 BOR
species :
Timonius leptocalyx
S.P.Darwin; Candollea 65(2): 230 (-231, fig.5) (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius longifolius
Valeton; Bot. Jahrb. Syst. 61: 44 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius longistipulus
Merr.; Philipp. J. Sci., C 8: 59 (1913)
Distribution: Philippines (Leyte)
42 PHI
species :
Timonius longitubus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 241 (1945)
Distribution: Solomon Isl. (Bougainville, Guadalcanal)
43 SOL
variant :
var. pubescens
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 54 (1994)
Distribution: Bismarck Arch.
43 BIS
species :
Timonius louiseanthus
Junhao Chen & K.M.Wong; Timonius Borneo 239 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius macrophyllus
(Teijsm. & Binn.) Valeton ex K.Heyne; Nutt. Pl. Ned.-Ind. 4: 200 (1917)
Distribution: Moluccas (Ambon)
42 MOL
Synonyms: hidden
= Greenea latifolia Teijsm. & Binn.; Cat. Hort. Bot. Bogor. [Teijsmann & Binnendijk] 116 (1866)
= Greenea macrophylla Teijsm. & Binn.; Natuurk. Tijdschr. Ned.-Indië 29: 248 (1867)
species :
Timonius malawaliensis
Junhao Chen & K.M.Wong; Timonius Borneo 243 (2023)
Distribution: Borneo (Sabah: Malawali Isl.)
42 BOR
species :
Timonius mapuluensis
Junhao Chen & K.M.Wong; Timonius Borneo 246 (2023)
Distribution: Borneo (E-Kalimantan)
42 BOR
species :
Timonius matangensis
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 58 (1909)
Distribution: Borneo (SW-Sarawak: Mt. Matang)
42 BOR
species :
Timonius megalocalyx
P.K.Hoo; Phytotaxa 599(3): 175 (2023)
Distribution: peninsular Malaysia; Singapore
42 MLY SNG
species :
Timonius melanophloeus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 251 (1945)
Distribution: Solomon Isl. (Ysabel)
43 SOL
species :
Timonius meliganensis
Junhao Chen & K.M.Wong; Timonius Borneo 254 (2023)
Distribution: Borneo (Sabah: Meligan FR)
42 BOR
species :
Timonius meridionalis
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 55 (1994)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius merokensis
Wernham; J. Bot. 56: 130 (1918)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius middletonii
Junhao Chen & K.M.Wong; Timonius Borneo 258 (2023)
Distribution: Borneo (SW-Sarawak: Mt. Matang)
42 BOR
species :
Timonius minahassae
Koord.; Natuurk. Tijdschr. Ned.-Indië 63: 87 (1904)
Distribution: Sulawesi; Moluccas
42 MOL SUL
Synonyms: hidden
= Timonius stipulosus Valeton; Icon. Bogor. [Boerlage] 3: 275 (1909), nom. superfl.
species :
Timonius minutifolius
Valeton; Bot. Jahrb. Syst. 61: 45 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius modestus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 237 (1945)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Timonius mohamedii
Junhao Chen & K.M.Wong; Timonius Borneo 260 (2023)
Distribution: Borneo (Sarawak: Batang Ai NP)
42 BOR
species :
Timonius mollis
Valeton; Bot. Jahrb. Syst. 63: 308 (1930)
Distribution: Palau (Babeldaob, Ngarakabesang, Malakal)
62 PLU
variant :
var. submollis
Fosberg & Sachet; Micronesica 20(1-2): 161 (1987)
Distribution: Palau
62 PLU
variant :
var. villosissimus
(Kaneh.) Fosberg & Sachet; Micronesica 20(1-2): 161 (1987)
Distribution: Palau
62 PLU
Synonyms: hidden
= Timonius villosissimus Kaneh.; Bot. Mag. (Tokyo) 48: 923 (1934)
species :
Timonius moluccanus
S.P.Darwin; Candollea 65(2): 233 (fig.6) (2010)
Distribution: Moluccas (Morotai)
42 MOL
species :
Timonius monkoboensis
Junhao Chen & K.M.Wong; Timonius Borneo 263 (2023)
Distribution: Borneo (Sabah: Gunung Monkobo)
42 BOR
species :
Timonius montanus
Ridl.; J. Linn. Soc., Bot. 38: 312 (1908)
Distribution: peninsular Malaysia (Gunung Lawit, Terengganu, Gunung Tahan & Gunung Tapis, Pahang); Sumatera
42 MLY SUM
species :
Timonius morobensis
S.P.Darwin; Syst. Bot. 22(1): 94 (1997)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius multinervis
Valeton; Bot. Jahrb. Syst. 61: 45 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius murudensis
Junhao Chen & K.M.Wong; Timonius Borneo 265 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius mutabilis
(Korth.) Walp.; Ann. Bot. Syst. [Walpers] 2: 765 (1852)
Distribution: Borneo (Kalimantan)
42 BOR
Synonyms: hidden
= Bobea mutabilis Korth.; Ned. Kruidk. Arch. 2(2): 212 (1851)
species :
Timonius neobritannicus
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 59 (1994)
Distribution: New Guinea (Papua New Guinea); Bismarck Arch.
42 NWG 43 BIS
species :
Timonius neocaledonicus
Schltr. & K.Krause; Bot. Jahrb. Syst. 40(Beibl. 92): 37 (1908)
Distribution: New Caledonia
60 NWC
species :
Timonius niahensis
Junhao Chen & K.M.Wong; Timonius Borneo 273 (2023)
Distribution: Borneo (NE-Sarawak)
42 BOR
species :
Timonius nitens
Merr. & L.M.Perry; J. Arnold Arbor. 26: 243 (1945)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius nitidus
(Bartl. ex DC.) F.Vill.; Fl. Filip. [Blanco], ed. 3, Nov. App. 109 (1880)
Distribution: Guam
62 GUM
Synonyms: hidden
= Petesia nitida Bartl. ex DC.; Prodr. [A.P. de Candolle] 4: 395 (1830)
species :
Timonius noli-tangere
J.G.Chavez, Alejandro & Meve; Bot. Lett. 167(3): 365 (2020)
Distribution: Philippines
42 PHI
species :
Timonius novoguineensis
Warb.; Bot. Jahrb. Syst. 13: 434 (1891)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Timonius bammleri K.Schum.; Notizbl. Königl. Bot. Gart. Berlin 1: 56 (1895)
species :
Timonius nudiceps
Valeton; Bot. Jahrb. Syst. 61: 47 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius nymannii
Valeton; Bot. Jahrb. Syst. 61: 48 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius oblanceolatus
Valeton; Bot. Jahrb. Syst. 61: 48 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius oblongus
Valeton; Bot. Jahrb. Syst. 61: 49 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius obovatifolius
Junhao Chen & K.M.Wong; Timonius Borneo 276 (2023)
Distribution: Borneo (C-Kalimantan)
42 BOR
species :
Timonius obovatus
Elmer; Leafl. Philipp. Bot. 1: 36 (1906)
Distribution: Philippines (Luzon, Sibuyan)
42 PHI
species :
Timonius octonervius
Merr.; J. Arnold Arbor. 18: 132 (1937)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Timonius ferrugineus Valeton; Bot. Jahrb. Syst. 61: 40 (1927), non Merr. (1915)
species :
Timonius oktediensis
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 60 (1994)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius ophioliticus
Jun H.Chen; Phytotaxa 181(3): 146 (2014)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius oreophilus
Ridl.; J. Straits Branch Roy. Asiat. Soc. 61: 22 (1912)
Distribution: peninsular Malaysia (Pahang, Selangor)
42 MLY
species :
Timonius ovalis
(Korth.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 132 (1891)
Distribution: Borneo (Kalimantan)
42 BOR
Synonyms: hidden
= Polyphragmon ovale Korth.; Ned. Kruidk. Arch. 2(2): 216 (1851)
species :
Timonius ovalistipulus
Valeton; Bot. Jahrb. Syst. 61: 49 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius oxyphyllus
Gand.; Bull. Soc. Bot. France 65: 34 (1918)
Distribution: peninsular Malaysia
42 MLY
species :
Timonius pachyphyllus
Merr.; Philipp. J. Sci. 26: 494 (1925)
Distribution: Philippines (Luzon)
42 PHI
species :
Timonius paiawensis
S.P.Darwin; J. Arnold Arbor. 64(4): 617 (1983)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius palawanensis
Elmer; Leafl. Philipp. Bot. 4: 1360 (1912)
Distribution: Borneo (Sabah, Sarawak, Kalimantan); Brunei; Palawan
42 BOR BRN PLW
Synonyms: hidden
= Timonius heterophyllus Merr.; J. Straits Branch Roy. Asiat. Soc. 77: 243 (1917)
= Timonius villamilii Merr.; J. Straits Branch Roy. Asiat. Soc. 77: 240 (1917)
species :
Timonius paludosus
Junhao Chen & K.M.Wong; Timonius Borneo 298 (2023)
Distribution: Borneo (Kalimantan)
42 BOR
species :
Timonius panayensis
Merr.; Philipp. J. Sci. 17: 317 (192t) (1920)
Distribution: Philippines (Panay)
42 PHI
species :
Timonius pannosus
Jun H.Chen; Phytotaxa 181(3): 146 (2014)
Distribution: Borneo (Sabah: Mt. Kinabalu)
42 BOR
species :
Timonius papuanus
Merr.; J. Arnold Arbor. 18: 132 (1937)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Timonius involucratus Valeton; Bot. Jahrb. Syst. 61: 41 (1927), nom. illeg.
species :
Timonius pasohensis
P.K.Hoo; J. Trop. Forest Sci. 36(3): 329 (2024)
Distribution: peninsular Malaysia
42 MLY
species :
Timonius paulleongii
Junhao Chen & K.M.Wong; Timonius Borneo 304 (2023)
Distribution: Borneo (SW-Sarawak)
42 BOR
species :
Timonius pearceae
Junhao Chen & K.M.Wong; Timonius Borneo 307 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius pedunculatus
(Thunb.) Ormerod; Harvard Pap. Bot. 29(1): 139 (2024)
Distribution: Japan
38 JAP
Synonyms: hidden
= Laurus pedunculata Thunb.; Syst. Veg., ed. 14 [J.A.Murray] 383 (1784)
= Cinnamomum japonicum Siebold; Verh. Batav. Genootsch. Kunst. 12: 23 (1830), nom. illeg.
species :
Timonius pereirae
Junhao Chen & K.M.Wong; Timonius Borneo 311 (2023)
Distribution: Borneo (Sabah, E-Kalimantan)
42 BOR
species :
Timonius philippinensis
Merr.; Bull. Bur. Forest. Philipp. Islands 1: 54 (1903)
Distribution: Philippines (Mindoro, Samar, Mindanao, Dinagat, Bucas Grande)
42 PHI
species :
Timonius piusii
Junhao Chen & K.M.Wong; Timonius Borneo 315 (2023)
Distribution: Borneo (Sabah: Trus Madi FR)
42 BOR
species :
Timonius platycarpus
Montrouz.; Mém. Acad. Roy. Sci. Lyon, Sect. Sci., Sér. 2, 10: 221 (1860)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Guettarda platycarpa (Montrouz.) Guillaumin; Arch. Bot. Mém. 3(5): 17 (1930)
species :
Timonius politus
Junhao Chen & K.M.Wong; Timonius Borneo 316 (2023)
Distribution: Borneo (NE-Sarawak)
42 BOR
species :
Timonius polyneurus
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 59 (1909)
Distribution: Borneo (W-Kalimantan)
42 BOR
species :
Timonius ponapensis
Valeton; Bot. Jahrb. Syst. 63: 306 (1930)
Distribution: Fed. States of Micronesia (Pohnpei)
62 MIC
species :
Timonius postaranthus
Junhao Chen & K.M.Wong; Timonius Borneo 321 (2023)
Distribution: Borneo (Sabah, Sarawak); Brunei
42 BOR BRN
species :
Timonius pseudaffinis
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 63 (1994)
Distribution: Solomon Isl. (South Solomons)
43 SOL
species :
Timonius pseudoarboreus
J.G.Chavez; Nordic J. Bot. 38(7)-e02730: 5 (2020)
Distribution: Philippines
42 PHI
species :
Timonius pseudocapitatus
(Scheff.) F.Muell.; Descr. Notes Papuan Pl. 1: 26 (1876)
Distribution: Moluccas
42 MOL
Synonyms: hidden
= Polyphragmon pseudocapitatum Scheff.; Ann. Jard. Bot. Buitenzorg 1: 29 (1876)
= Timonius pseudocapitatus (Scheff.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 132 (1891), isonym
species :
Timonius puayyokii
Junhao Chen & K.M.Wong; Timonius Borneo 325 (2023)
Distribution: Borneo (C-Sarawak)
42 BOR
species :
Timonius pubistipulus
S.P.Darwin; J. Arnold Arbor. 64(4): 612 (1983)
Distribution: New Guinea (Papua New Guinea)
42 NWG
variant :
var. pubescens
S.P.Darwin; J. Arnold Arbor. 64(4): 614 (1983)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius puffii
Junhao Chen & K.M.Wong; Timonius Borneo 329 (2023)
Distribution: Brunei
42 BRN
species :
Timonius pulgarensis
Elmer; Leafl. Philipp. Bot. 4: 1360 (1912)
Distribution: Palawan
42 PLW
species :
Timonius pullenii
S.P.Darwin; Brittonia 62(2): 134 (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius pulposus
C.T.White; J. Arnold Arbor. 31: 115 (1950)
Distribution: New Guinea (Papua New Guinea); Solomon Isl.
42 NWG 43 SOL
species :
Timonius quinqueflorus
Merr.; Philipp. J. Sci., C 10: 116 (1915)
Distribution: Philippines (Luzon)
42 PHI
species :
Timonius rarissimus
P.K.Hoo & Sugumaran; Sandakania 27: 83 (2025)
Distribution: peninsular Malaysia
42 MLY
species :
Timonius reflexus
S.P.Darwin; Candollea 65(2): 234 (-235, fig.7) (2010)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius repertus
S.P.Darwin; Candollea 65(2): 235 (-236, fig.8) (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius ridhanthus
Junhao Chen & K.M.Wong; Timonius Borneo 333 (2023)
Distribution: Borneo (E-Kalimantan)
42 BOR
species :
Timonius ridleyi
Merr.; J. Arnold Arbor. 18: 132 (1937), nom. nov.
Distribution: peninsular Malaysia
42 MLY
Synonyms: hidden
= Timonius hirsutus Ridl.; J. Straits Branch Roy. Asiat. Soc. 79: 81 (1918), non T. hirsutus Merr. 79: 81 (1918), non T. hirsutus Merr.
species :
Timonius ridsdalei
J.G.Chavez; Nordic J. Bot. 38(7)-e02730: 6 (2020)
Distribution: Philippines
42 PHI
species :
Timonius rigidus
(Miq.) F.Muell.; Descr. Notes Papuan Pl. 1: 26 (1876)
Distribution: Lesser Sunda Isl. (Timor)
42 LSI
Synonyms: hidden
= Timonius rigidus Boerl. ex Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 57 (1909)
= Polyphragmon rigidum Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 243 (1869)
species :
Timonius rimiae
Junhao Chen & K.M.Wong; Timonius Borneo 335 (2023)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius riswanii
Junhao Chen & K.M.Wong; Timonius Borneo 338 (2023)
Distribution: Borneo (W-Kalimantan, C-Kalimantan)
42 BOR
species :
Timonius rivularis
Merr. & L.M.Perry; J. Arnold Arbor. 26: 251 (1945)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius roemeri
Valeton; Nova Guinea 8: 476 (1911)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius rosselensis
S.P.Darwin; Candollea 65(2): 236 (-237, fig.9) (2010)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius rufescens
(Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 133 (1891)
Distribution: Moluccas (Ternate); New Guinea (Irian Jaya, Papua New Guinea)
42 MOL NWG
Synonyms: hidden
= Polyphragmon rufescens Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 241 (1869)
species :
Timonius rufus
Valeton; Bot. Jahrb. Syst. 61: 50 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius runiae
Junhao Chen & K.M.Wong; Timonius Borneo 340 (2023)
Distribution: Borneo (Sarawak, Sabah)
42 BOR
species :
Timonius salicifolius
Valeton; Icon. Bogor. [Boerlage] t. 298 (1909)
Distribution: Borneo (Kalimantan)
42 BOR
species :
Timonius salsedoi
Fosberg & Sachet; Micronesica 20(1-2): 162 (1987)
Distribution: Fed. States of Micronesia
62 MIC
species :
Timonius sampitensis
Junhao Chen & K.M.Wong; Timonius Borneo 349 (2023)
Distribution: Borneo (C-Kalimantan)
42 BOR
species :
Timonius scaber
Valeton; Bot. Jahrb. Syst. 61: 50 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius scabriflorus
(Valeton) Merr. & L.M.Perry; J. Arnold Arbor. 26: 238 (1945)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Timonius avenis var. scabriflorus Valeton; Bot. Jahrb. Syst. 61: 37 (1927)
species :
Timonius schefferi
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 49 (1909)
Distribution: Moluccas (Gebe Isl.)
42 MOL
Synonyms: hidden
= Polyphragmon stipulosum Scheff. ex Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 49 (1909)
species :
Timonius schumannii
Koord. ex Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 47 (1909)
Distribution: Sulawesi
42 SUL
species :
Timonius secundiflorus
S.P.Darwin; Syst. Bot. Monogr. 12 (Syst. Timonius subgen. Abbottia): 69 (1994)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
Synonyms: hidden
= Timonius avenis f. triflora Valeton; Bot. Jahrb. Syst. 61: 38 (1927)
species :
Timonius seowchoenanthus
Junhao Chen & K.M.Wong; Timonius Borneo 352 (2023)
Distribution: Borneo (Sarawak, Kalimantan)
42 BOR
species :
Timonius sepilokensis
Junhao Chen & K.M.Wong; Timonius Borneo 357 (2023)
Distribution: Borneo (E-Sabah)
42 BOR
species :
Timonius sepulutensis
Junhao Chen & K.M.Wong; Timonius Borneo 359 (2023)
Distribution: Borneo (Sabah: Sepulut FR)
42 BOR
species :
Timonius serenae
Junhao Chen & K.M.Wong; Timonius Borneo 361 (2023)
Distribution: Borneo (Kalimantan, SW-Sarawak)
42 BOR
species :
Timonius sessilis
(Scheff.) Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 51 (1909)
Distribution: Moluccas
42 MOL
Synonyms: hidden
= Polyphragmon sessile Scheff.; Ann. Jard. Bot. Buitenzorg 1: 67 (1876), nomen
species :
Timonius silamensis
Junhao Chen & K.M.Wong; Timonius Borneo 364 (2023)
Distribution: Borneo (E-Sabah)
42 BOR
species :
Timonius singularis
(F.Muell.) L.S.Sm.; Proc. Roy. Soc. Queensland 68: 50 (1957)
Distribution: Australia (Queensland)
50 AUS
Synonyms: hidden
= Abbottia singularis F.Muell.; Fragm. [Mueller] 9: 181 (1875)
species :
Timonius sinyengiae
Junhao Chen & K.M.Wong; Timonius Borneo 366 (2023)
Distribution: Borneo (Sarawak)
42 BOR
species :
Timonius solomonensis
Merr. & L.M.Perry; J. Arnold Arbor. 26: 242 (1945)
Distribution: Solomon Isl. (Ysabel, San Christobal)
43 SOL
species :
Timonius spes-vitarum
J.G.Chavez; Nordic J. Bot. 38(7)-e02730: 8 (2020)
Distribution: Philippines
42 PHI
species :
Timonius splendens
(Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 132 (1891)
Distribution: Sumatera
42 SUM
Synonyms: hidden
= Polyphragmon splendens Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 241 (1869)
species :
Timonius stenolobus
Jun H.Chen & K.M.Wong; Phytotaxa 181(3): 147 (2014)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius stevendarwinii
J.G.Chavez; Nordic J. Bot. 38(7)-e02730: 10 (2020)
Distribution: Philippines
42 PHI
species :
Timonius subauritus
Valeton; Bot. Jahrb. Syst. 63: 307 (1930)
Distribution: Palau (Babeldaob, Koror, Ngarakabesang)
62 PLU
variant :
var. strigosus
Fosberg & Sachet; Micronesica 20(1-2): 163 (1987)
Distribution: Palau
62 PLU
species :
Timonius subavenis
(Valeton) S.P.Darwin; J. Arnold Arbor. 64(4): 616 (1983)
Distribution: New Guinea (Irian Jaya)
42 NWG
Synonyms: hidden
= Timonius scaber var. subavenis Valeton; Bot. Jahrb. Syst. 61: 51 (1927)
species :
Timonius subcoriaceus
Valeton; Bot. Jahrb. Syst. 61: 52 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
variant :
var. sessilis
S.P.Darwin; Syst. Bot. 22(1): 97 (1997)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius subsessilis
Valeton; Bull. Dépt. Agric. Indes Néerl. 10: 64 (1907)
Distribution: Sulawesi; New Guinea (Irian Jaya)
42 NWG SUL
species :
Timonius sulitii
Merr. & Quisumb. ex J.G.Chavez & Tandang; Bot. Lett. 167(3): 368 (2020)
Distribution: Philippines
42 PHI
species :
Timonius sunsuronensis
Junhao Chen & K.M.Wong; Timonius Borneo 375 (2023)
Distribution: Borneo (Sabah: Tambunan)
42 BOR
species :
Timonius suzanae
Junhao Chen & K.M.Wong; Timonius Borneo 377 (2023)
Distribution: Borneo (E-Sabah)
42 BOR
species :
Timonius sylvestris
S.Moore; J. Bot. 65: 247 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius tambuyukonensis
Jun H.Chen; Phytotaxa 181(3): 148 (2014)
Distribution: Borneo (Sabah: Mt. Kinabalu)
42 BOR
species :
Timonius ternifolius
(Bartl. ex DC.) Fern.-Vill.; Fl. Filip. [Blanco], ed. 3, Nov. App. 109 (1880)
Distribution: Palawan (incl. Busuanga, Culion); Philippines (Camiguin, Luzon, Babuyan Isl., Panay)
42 PHI PLW
Synonyms: hidden
= Petesia ternifolia Bartl. ex DC.; Prodr. [A.P. de Candolle] 4: 395 (1830)
species :
Timonius teysmannii
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 38 (1909)
Distribution: Sulawesi
42 SUL
species :
Timonius thomasii
Junhao Chen & K.M.Wong; Timonius Borneo 384 (2023)
Distribution: Borneo (W-Kalimantan)
42 BOR
species :
Timonius timon
(Spreng.) Merr.; J. Arnold Arbor. 18: 131 (1937)
Distribution: peninsular Malaysia (Perak, Pahang, Selangor, Negeri Sembilan, Melaka, Johor); Sulawesi; Jawa; Lesser Sunda Isl.; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Solomon Isl.; Australia (Western Australia, Northern Territory, Queensland)
42 JAW LSI MLY MOL NWG SUL 43 BIS SOL 50 AUS
Synonyms: hidden
= Axanthes timoriensis DC.; Prodr. [A.P. de Candolle] 4: 441 (1830)
= Erithalis polygama var. timonius Willd.; Sp. Pl., ed. 4 [Willdenow] 1: 997 (1798)
= Erithalis timon Spreng.; Pl. Min. Cogn. Pug. 1: 18 (1813)
= Guettarda polygama (Oken) Baill.; Hist. Pl. [Baillon] 7: 377 (1880)
= Guettarda polyphragmoides F.Muell.; Fragm. [Mueller] 2: 134 (1861)
= Nelitris timon (Spreng.) Britten; J. Bot. 39: 68 (1901)
= Polyphragmon sericeum Desf.; Mém. Mus. Hist. Nat. 6: 6, t. 2 (1820)
= Polyphragmon spiciflorum Zipp. ex Span.; Linnaea 15: 317 (1841)
= Timonius brassii S.Moore; J. Bot. 65: 265 (1927)
= Timonius pleiomera Lauterb. & K.Schum.; Notizbl. Königl. Bot. Gart. Berlin 2: 152 (1898)
= Timonius polygamus Oken; Allg. Naturgesch. 3(2): 875 (1841)
= Timonius sericeus (Desf.) K.Schum.; Fl. Kais. Wilh. Land [Schumann & Hollrung] 131 (1889)
= Timonius sericeus var. pleiomera Valeton; Bot. Jahrb. Syst. 61: 51 (1927)
variant :
var. buloloensis
S.P.Darwin; Allertonia 7(1): 26 (1993)
Distribution: New Guinea (Papua New Guinea)
42 NWG
variant :
var. grandiflorus
(K.Schum.) S.P.Darwin; Allertonia 7(1): 25 (1993)
Distribution: New Guinea
42 NWG
Synonyms: hidden
= Timonius sericeus var. grandiflorus Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 53 (1909), nom. inval.
= Timonius sericeus var. grandiflora K.Schum.; Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 568 (1900)
variant :
var. reticulatus
S.P.Darwin; Allertonia 7(1): 28 (1993)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
variant :
var. whiteanus
(S.Moore) S.P.Darwin; Allertonia 7(1): 25 (1993)
Distribution: New Guinea
42 NWG
Synonyms: hidden
= Timonius whiteanus S.Moore; J. Bot. 65: 246 (1927)
species :
Timonius tjiasmantoi
Junhao Chen & K.M.Wong; Timonius Borneo 387 (2023)
Distribution: Borneo (C-Kalimantan)
42 BOR
species :
Timonius tomentosus
(Valeton) S.P.Darwin; Allertonia 7(1): 29 (1993)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Timonius sericeus var. tomentosa Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 53 (1909)
species :
Timonius trichanthus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 238 (1945)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius trichocaulon
(Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 2(1): 132 (1891)
Distribution: Moluccas
42 MOL
Synonyms: hidden
= Polyphragmon trichocaulon Miq.; Ann. Mus. Bot. Lugduno-Batavi 4: 241 (1869)
species :
Timonius trichocladus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 239 (1945)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius trichophorus
Merr.; Philipp. J. Sci., C 9: 388 (1914)
Distribution: Philippines (Leyte, Samar, Mindanao, Bucas Grande)
42 PHI
Synonyms: hidden
= Greenea hirsuta Elmer; Leafl. Philipp. Bot. 5: 1857 (1913)
= Timonius hirsutus (Elmer) Merr.; Philipp. J. Sci. 17: 480 (1920 publ. 1921), nom. illeg.
species :
Timonius uniflorus
(C.F.Gaertn.) Govaerts; Bot. J. Linn. Soc. 157(1): 120 (2008)
Distribution: Solomon Isl.; New Caledonia; Vanuatu; Tonga (Vava´u); Niue; Fiji; Wallis & Futuna (Alofi Isl.); Cook Isl. (widespread, incl. Rarotonga); Society Isl. (Mopelia Atoll, Tahiti, Bora Bora, Tetiaroa Atoll, Tupai); Tuamotu Arch.; Tuamotu Arch. (Rangiroa Atoll, Takapoto Atoll, Makatea Isl., Niau, Takaroa, Tikei, Arutua, Manihi, Hiti, Takaroa, Makemo, Katiu, Aratika, Taiaro, Kaukura); Austral Isl. (Maria Atoll, Rimatara); Pitcairn Isl.; Henderson Isl.; Palau [I] (Peleliu [I], Angaur [I]); Marshall Isl.
43 SOL 60 FIJ NUE NWC TON VAN WAL 61 AUI COO HEN PIT SCI TUA 62 MRS plu
Synonyms: hidden
= Bobea forsteri (DC.) Korth.; Ned. Kruidk. Arch. 2(2): 211 (1851), nom. superfl.
= Bobea gaertneri Korth.; Ned. Kruidk. Arch. 2(2): 211 (1851), nom. illeg.
= Burneya forsteri Cham. & Schltdl.; Linnaea 4: 189 (1829), nom. illeg.
= Erithalis cymosa G.Forst. ex Spreng.; Pl. Min. Cogn. Pug. 1: 17 (1813)
= Erithalis polygama G.Forst.; Fl. Ins. Austr. 17 (1786)
= Erithalis uniflora Banks ex C.F.Gaertn.; Fruct. Sem. Pl. 3: 93 (1805)
= Polyphragmon minus A.Rich. ex DC.; Prodr. [A.P. de Candolle] 4: 445 (1830), A. Rich., Mém. Soc. Hist. Nat. Paris 5: 231 (1834)
= Timonius forsteri DC.; Prodr. [A.P. de Candolle] 4: 461 (1830), nom. superfl.
= Timonius polygamus (G.Forst.) C.B.Rob.; Proc. Amer. Acad. Arts 45: 408 (1910), nom. illeg.
species :
Timonius vaccinioides
Wernham ex Ridl.; Hooker´s Icon. Pl. 31: t. 3054 (1916)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius valetonii
Elmer; Leafl. Philipp. Bot. 3: 1012 (1911)
Distribution: Philippines (Sibuyan, Mindanao)
42 PHI
species :
Timonius vanderentii
Junhao Chen & K.M.Wong; Timonius Borneo 392 (2023)
Distribution: Borneo (Sabah)
42 BOR
species :
Timonius versteegii
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 39 (1909)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius villosus
Valeton; Bot. Jahrb. Syst. 61: 52 (1927)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius virgatus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 240 (1945)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius wallichianus
Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 36 (1909)
Distribution: peninsular Malaysia; Singapore; Sumatera; Borneo (Kalimantan)
42 BOR MLY SNG SUM
Synonyms: hidden
= Bobea wallichiana Korth.; Ned. Kruidk. Arch. 2(2): 211 (1851), nomen
= Gardenia trichoclada Miq.; Fl. Ned. Ind. 2: 231 (1857)
= Timonius rumphii DC.; Prodr. [A.P. de Candolle] 4: 461 (1830), nom. illeg.
= Timonius wallichianus var. glabrior Valeton; Bull. Dépt. Agric. Indes Néerl. 26: 37 (1909)
species :
Timonius wollastonii
Wernham; Trans. Linn. Soc. London, Bot., Ser. 2, 9: 74 (1916)
Distribution: New Guinea (Irian Jaya)
42 NWG
species :
Timonius wrayi
King & Gamble; J. Asiat. Soc. Bengal 73: 54 (1904)
Distribution: peninsular Malaysia (Terengganu, Perak, Pahang, Johor); +Singapore
42 MLY SNG
species :
Timonius xanthocarpus
Merr. & L.M.Perry; J. Arnold Arbor. 26: 247 (1945)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Timonius zuckianus
S.P.Darwin; J. Arnold Arbor. 64(4): 611 (1983)
Distribution: New Guinea (d´Entrecasteaux Isl., Louisiade Arch.)
42 NWG
