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338.0000 Family Phyllanthaceae Martinov; Tekhno-Bot. Slovar 369 (1820), nom. cons.
58 genera, ca. 2200 spp.
Species listed in database: 2113; hybrids: 5
= Antidesmataceae Loudon; Hort. Brit. [Loudon] 534 (1830)
= Aporosaceae Lindl. ex Planch.; Ann. Sci. Nat., Bot., Sér. 4, 2: 265 (1854)
= Bischofiaceae Airy Shaw; Kew Bull. 18: 252 (1965)
= Hymenocardiaceae Airy Shaw; Kew Bull. 18: 261 (1965)
= Porantheraceae Hurus.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 6: 224 (1954)
= Scepaceae Lindl.; Intr. Nat. Syst. Bot., ed. 2: 171 (1836)
= Stilaginaceae C.Agardh; Aphor. Bot. 199 (1824)
= Uapacaceae Airy Shaw; Kew Bull. 18: 270 (1965)
338.1301 Genus Uapaca Baill.; Étude Euphorb. 595 (1858)
Distribution: Tropical Africa (many), Madagascar (8)
Species listed in database: 25
= Canariastrum Engl.; Bot. Jahrb. Syst. 26: 364 (1899)
= Gymnocarpus Thouars ex Baill.; Étude Euphorb. 595 (1858)
species :
Uapaca acuminata (Hutch.) Pax & K.Hoffm.; Pflanzenreich [Engler] IV.147.XV, Heft 81 (Euph.-Phyllantheae): 308 (1922)
Distribution: Nigeria; Cameroon; Equatorial Guinea; Gabon
22 NGA 23 CMN EQG GAB
Synonyms: hidden
= Uapaca heudelotii var. acuminata Hutch.; Fl. Trop. Afr. [Oliver et al.] 6(1): 636 (1912)
species :
Uapaca ambanjensis Leandri; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 8: 255 (1957)
Distribution: N-Madagascar
29 MDG
species :
Uapaca amplifolia Denis; Arch. Bot. Bull. Mens. 1: 224 (1927)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Uapaca rivularis Denis; Arch. Bot. Bull. Mens. 1: 225 (1927)
species :
Uapaca bojeri Baill.; Adansonia 11: 176 (1874)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Uapaca clusiacea Baker; J. Linn. Soc., Bot. 18: 278 (1881)
= Chorizotheca macrophylla Heckel; Ann. Mus. Colon. Marseille, Sér. 2, 8: 215 (1910)
species :
Uapaca densifolia Baker; J. Linn. Soc., Bot. 20: 252 (1884)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Uapaca angustifolia Denis; Arch. Bot. Bull. Mens. 1: 227 (1927)
= Uapaca densifolia f. orientalis Leandri; Fl. Madagasc., Fam. 111: 174 (1958), no type indicated.
= Uapaca densifolia var. angustifolia (Denis) Leandri; Fl. Madagasc., Fam. 111: 176 (1958)
= Uapaca densifolia var. humbertii Leandri; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 8: 257 (1957)
= Uapaca densifolia var. perrieri (Denis) Leandri; Fl. Madagasc., Fam. 111: 176 (1958)
= Uapaca densifolia var. sambiranensis (Denis) Leandri; Fl. Madagasc., Fam. 111: 176 (1958)
= Uapaca myricifolia Baker; J. Linn. Soc., Bot. 21: 440 (1885)
= Uapaca perrieri Denis; Arch. Bot. Bull. Mens. 1: 228 (1927)
= Uapaca sambiranensis Denis; Arch. Bot. Bull. Mens. 1: 226 (1927)
species :
Uapaca ferruginea Baill.; Étude Euphorb. 596 (1858)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Uapaca clusioides Baker; J. Linn. Soc., Bot. 21: 441 (1885)
= Uapaca ferruginea var. carnotiana Leandri; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 8: 257 (1957)
species :
Uapaca guineensis Müll.Arg.; Flora 47: 517 (1864)
Distribution: Guinea-Bissau; Guinea; Sierra Leone; Liberia; SW-Mali; SW-Burkina Faso; Ivory Coast; Ghana; Togo; Benin; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; Equatorial Guinea; Gabon; Congo [Brazzaville]; Central African Republic; D. R. Congo [Zaire]; Tanzania; Angola
22 BEN BKN GHA GNB GUI IVO LBR MLI NGA SIE TOG 23 BIO CAF CMN CON EQG GAB PRI SAO ZAI 25 TAN 26 ANG
Synonyms: hidden
= Antidesma guineensis G.Don ex Hook.; Niger Fl. [W.J.Hooker] 515 (1849) (nomen)
= Uapaca bingervillensis Beille; Mém. Soc. Bot. France 8: 66 (1908)
= Uapaca gabonensis Pierre ex Pax & K.Hoffm.; Pflanzenreich [Engler] IV.147.XV, Heft 81 (Euph.-Phyllantheae): 306 (1922)
= Uapaca seretii De Wild.; Ann. Mus. Congo Belge, Bot., Sér. 5, 2: 274 (1908)
species :
Uapaca heudelotii Baill.; Adansonia 1: 81 (1860 publ. 1861)
Distribution: Guinea-Bissau; Guinea; Sierra Leone; Liberia; SW-Mali; S-Burkina Faso; Ivory Coast; Ghana; Togo; Benin; Nigeria; Cameroon; Equatorial Guinea; Gabon; Congo [Brazzaville]; Central African Republic; D. R. Congo [Zaire]; Angola
22 BEN BKN GHA GNB GUI IVO LBR MLI NGA SIE TOG 23 CAF CMN CON EQG GAB ZAI 26 ANG
Synonyms: hidden
= Uapaca bossenge De Wild.; Ann. Mus. Congo Belge, Bot., Sér. 5, 2: 271 (1908)
= Uapaca marquesii Pax; Bot. Jahrb. Syst. 23: 522 (1897)
= Uapaca prominenticarinata De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 47: 37 (1927)
species :
Uapaca kirkiana Müll.Arg.; Flora 47: 517 (1864)
Distribution: D. R. Congo [Zaire]; Burundi; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique
23 BUR ZAI 25 TAN 26 ANG MLW MOZ ZAM ZIM
Synonyms: hidden
= Uapaca albida De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 57 (1933)
= Uapaca goetzei Pax; Bot. Jahrb. Syst. 28: 418 (1900)
= Uapaca greenwayi Suess.; Trans. Rhodesia Sci. Assoc. 43: 11 (1951)
= Uapaca homblei De Wild.; Ann. Soc. Sci. Bruxelles 45: 309 (1926)
= Uapaca kirkiana var. goetzei Pax; Bot. Jahrb. Syst. 34: 370 (1904)
= Uapaca kirkiana var. sessilifolia P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 891 (1949)
= Uapaca neomasuku De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 52: 208 (1932)
variant :
var. benguelensis (Müll.Arg.) Meerts; Kew Bull. 74(3)-38: 14 (2019)
Distribution: D. R. Congo [Zaire]; Angola; Zambia
23 ZAI 26 ANG ZAM
Synonyms: hidden
= Uapaca angolensis Hutch. ex Pax & K.Hoffm.; Pflanzenreich [Engler] IV.147.XV, Heft 81 (Euph.-Phyllantheae): 301 (1922), pro syn.
= Uapaca benguelensis Müll.Arg.; J. Bot. 2: 332 (1864)
= Uapaca benguelensis var. pedunculata P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 890 (1949)
variant :
var. gossweileri (Hutch.) Meerts; Kew Bull. 74(3)-38: 12 (2019)
Distribution: D. R. Congo [Zaire]; Angola
23 ZAI 26 ANG
Synonyms: hidden
= Uapaca gossweileri Hutch.; Bull. Misc. Inform. Kew 1912: 101 (1912), et in Fl. Trop. Afr. [Oliver et al.] 6(1): 635 (1912)
= Uapaca kirkiana var. kwangoensis P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 891 (1949)
species :
Uapaca lissopyrena Radcl.-Sm.; Kew Bull. 48(3): 612 (1993)
Distribution: Zambia; Malawi; Zimbabwe; Mozambique
26 MLW MOZ ZAM ZIM
species :
Uapaca littoralis Denis; Arch. Bot. Bull. Mens. 1: 228 (1927)
Distribution: E-Madagascar
29 MDG
Synonyms: hidden
= Uapaca betamponensis Leandri; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 8: 255 (1957)
= Uapaca mangorensis Leandri; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 8: 258 (1957)
= Uapaca silvestris Leandri; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 8: 259 (1957)
species :
Uapaca louvelii Denis; Arch. Bot. Bull. Mens. 1: 230 (1927)
Distribution: E-Madagascar
29 MDG
Synonyms: hidden
= Uapaca louvelii var. paralia Leandri; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 8: 258 (1957)
species :
Uapaca mole Pax; Bot. Jahrb. Syst. 19: 79 (1894)
Distribution: Guinea; Liberia; Ivory Coast; Ghana; Benin; Nigeria; Cameroon; Gabon; Congo [Brazzaville]; D. R. Congo [Zaire]; Uganda; Tanzania
22 BEN GHA GUI IVO LBR NGA 23 CMN CON GAB ZAI 25 TAN UGA
Synonyms: hidden
= Uapaca angustipyrena De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 52: 205 (1932)
= Uapaca brevipedunculata De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 145 (1933)
= Uapaca casteelsii De Wild.; Compt.-Rend. Hebd. Séances Mém. Soc. Biol. 96: 700 (1927)
= Uapaca ealaensis De Wild.; Ann. Mus. Congo Belge, Bot., Sér. 5, 2: 272 (1908)
= Uapaca ferrarii De Wild.; Compt.-Rend. Hebd. Séances Mém. Soc. Biol. 96: 118 (1927)
= Uapaca kibuati De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 309 (1933)
= Uapaca laurentii De Wild.; Ann. Mus. Congo Belge, Bot., Sér. 5, 2: 272 (1908)
= Uapaca lebrunii De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 146 (1933)
= Uapaca macrostipulata De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 47: 36 (1927)
= Uapaca multinervata De Wild.; Compt.-Rend. Hebd. Séances Mém. Soc. Biol. 96: 118 (1927)
= Uapaca paludosa Aubrév. & Leandri; Bull. Soc. Bot. France 82: 50 (1935)
= Uapaca samfii De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 310 (1933)
= Uapaca stipularis Pax & K.Hoffm.; Pflanzenreich [Engler] IV.147.XV, Heft 81 (Euph.-Phyllantheae): 303 (1922)
= Uapaca vanderystii De Wild.; Compt.-Rend. Hebd. Séances Mém. Soc. Biol. 96: 701 (1927)
= Uapaca verruculosa De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 148 (1933)
species :
Uapaca niangadoumae Breteler; Pl. Ecol. Evol. 145(1): 129 (2012)
Distribution: Gabon
23 GAB
species :
Uapaca nitida Müll.Arg.; Flora 47: 517 (1864)
Distribution: D. R. Congo [Zaire]; Burundi; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique
23 BUR ZAI 25 KEN TAN 26 ANG MLW MOZ ZAM ZIM
Synonyms: hidden
= Uapaca microphylla Pax; Bot. Jahrb. Syst. 23: 523 (1897)
= Uapaca microphylla var. hendrickxii De Wild.; Études Fl. Bas- Moyen-Congo 2: 273 (1908)
= Uapaca nitida f. aucta P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 888 (1949)
= Uapaca nitida f. bianoensis P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 889 (1949)
= Uapaca nitida f. latiuscula P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 888 (1949)
= Uapaca nitida f. scalarinervosa P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 889 (1949)
= Uapaca nitida var. mulengo P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 889 (1949)
= Uapaca nitida var. nsambi P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 888 (1949)
= Uapaca nitida var. sokolobe P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 889 (1949)
= Uapaca nitida var. suffrutescens P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 889 (1949)
= Uapaca similis Pax & K.Hoffm.; Pflanzenreich [Engler] IV.147.XV, Heft 81 (Euph.-Phyllantheae): 305 (1922)
variant :
var. longifolia (P.A.Duvign.) Radcl.-Sm.; Kew Bull. 48(3): 616 (1993)
Distribution: D. R. Congo [Zaire]; Malawi; Zimbabwe; Mozambique
23 ZAI 26 MLW MOZ ZIM
Synonyms: hidden
= Uapaca nitida f. longifolia P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 890 (1949)
species :
Uapaca pilosa Hutch.; Bull. Misc. Inform. Kew 1912: 102 (1912), et in Fl. Trop. Afr. [Oliver et al.] 6(1): 635 (1912)
Distribution: Cameroon; D. R. Congo [Zaire]; Tanzania; Zambia; Malawi
23 CMN ZAI 25 TAN 26 MLW ZAM
Synonyms: hidden
= Uapaca masuku De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 45: 311 (1926)
= Uapaca pilosa f. hirsuta P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 891 (1949), nom. illeg.
variant :
var. petiolata P.A.Duvign.; (1949)
Distribution: D. R. Congo [Zaire]; Tanzania; Zambia; Malawi
23 ZAI 25 TAN 26 MLW ZAM
Synonyms: hidden
= Uapaca macrocephala Pax & K.Hoffm.; Pflanzenreich [Engler] IV.147.XV, Heft 81 (Euph.-Phyllantheae): 305 (1922)
= Uapaca pilosa f. subglabra P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 891 (1949)
= Uapaca sapinii De Wild.; Proc. Int. Congr. Pl. Sci. Ithaca 1420 (1929), in clavi.
= Uapaca sapinii De Wild. ex De Wild.; Mém. Inst. Roy. Colon. Belge, Sect. Sci. Nat. (8vo) 4(5): 174 (1936), in adnot., latine.
species :
Uapaca pynaertii De Wild.; Ann. Mus. Congo Belge, Bot., Sér. 5, 2: 274 (1908)
Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Ghana; Nigeria; Cameroon; ?Congo [Brazzaville]; Gabon; D. R. Congo [Zaire]
22 GHA GUI IVO LBR NGA SIE 23 CMN GAB ZAI
Synonyms: hidden
= Uapaca corbisieri De Wild.; Compt.-Rend. Hebd. Séances Mém. Soc. Biol. 96: 118 (1927)
= Uapaca esculenta A.Chev. ex Aubrév. & Leandri; Bull. Soc. Bot. France 82: 52 (1935)
= Uapaca goossensii De Wild.; Compt.-Rend. Hebd. Séances Mém. Soc. Biol. 96: 118 (1927)
= Uapaca nymphaeantha Pax & K.Hoffm.; Pflanzenreich [Engler] IV.147.XV, Heft 81 (Euph.-Phyllantheae): 306 (1922)
species :
Uapaca robynsii De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 60 (1933)
Distribution: D. R. Congo [Zaire]; Zambia; Malawi
23 ZAI 26 MLW ZAM
species :
Uapaca rufopilosa (De Wild.) P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 890 (1949)
Distribution: D. R. Congo [Zaire]; Angola; Zambia
23 ZAI 26 ANG ZAM
Synonyms: hidden
= Uapaca nitida var. rufopilosa De Wild.; Mém. Inst. Roy. Colon. Belge, Sect. Sci. Nat. (8vo) 4(5): 161 (1936)
species :
Uapaca sansibarica Pax; Bot. Jahrb. Syst. 34: 370 (1904)
Distribution: D. R. Congo [Zaire]; Burundi; South Sudan; Uganda; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique
23 BUR ZAI 24 SSU 25 TAN UGA 26 ANG MLW MOZ ZAM ZIM
Synonyms: hidden
= Uapaca sansibarica var. cuneata Pax; Bot. Jahrb. Syst. 34: 370 (1904)
species :
Uapaca staudtii Pax; Bot. Jahrb. Syst. 23: 522 (1897)
Distribution: Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Equatorial Guinea; Gabon
22 NGA 23 BIO CMN EQG GAB
species :
Uapaca teusczii Pax; Bot. Jahrb. Syst. 19: 79 (1894)
Distribution: D. R. Congo [Zaire]; Angola; Zambia
23 ZAI 26 ANG ZAM
Synonyms: hidden
= Uapaca benguelensis f. glabra P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 890 (1949)
= Uapaca dubia De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 307 (1933)
= Uapaca kirkiana var. dubia (De Wild.) P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 891 (1949)
= Uapaca munamensis De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 52: 206 (1932)
form :
f. pilosa (P.A.Duvign.) Meerts; Kew Bull. 74(3)-38: 8 (2019)
Distribution: D. R. Congo [Zaire]
23 ZAI
Synonyms: hidden
= Uapaca benguelensis f. pilosa P.A.Duvign.; Bull. Séances Inst. Roy. Colon. Belge 20: 890 (1949)
= Uapaca katentaniensis De Wild.; Ann. Soc. Sci. Bruxelles, Sér. B, 53: 58 (1933)
species :
Uapaca thouarsii Baill.; Étude Euphorb. 598 (1858)
Distribution: E-Madagascar
29 MDG
species :
Uapaca togoensis Pax; Bot. Jahrb. Syst. 34: 371 (1904)
Distribution: Guinea-Bissau; Guinea; Sierra Leone; SW-Mali; Burkina Faso; Ivory Coast; Ghana; Togo; Benin; Nigeria; Cameroon; Congo [Brazzaville]; Central African Republic; SW-Chad; Malawi
22 BEN BKN GHA GNB GUI IVO MLI NGA SIE TOG 23 CAF CMN CON 24 CHA 26 MLW
Synonyms: hidden
= Uapaca chevalieri Beille; Mém. Soc. Bot. France 8: 68 (1908)
= Uapaca guignardii A.Chev. ex Beille; Mém. Soc. Bot. France 8: 66 (1908)
= Uapaca guignardii var. sudanica Beille; Mém. Soc. Bot. France 8: 67 (1908)
= Uapaca guineensis var. sudanica (Beille) Hutch.; Fl. Trop. Afr. [Oliver et al.] 6(1): 641 (1912)
= Uapaca perrotii Beille; Mém. Soc. Bot. France 8: 67 (1908)
= Uapaca somon Aubrév. & Leandri; Bull. Soc. Bot. France 82: 50 (1935)
species :
Uapaca vanhouttei De Wild.; Ann. Mus. Congo Belge, Bot., Sér. 5, 2: 275 (1908)
Distribution: S-Nigeria; Cameroon; Gabon; Congo [Brazzaville]; D. R. Congo [Zaire]
22 NGA 23 CMN CON GAB ZAI
Synonyms: hidden
= Uapaca brieyi De Wild.; Bull. Jard. Bot. État Bruxelles 4: 362 (1914)
= Uapaca letestuana A.Chev.; Vég. Utiles Afrique Trop. Franç. 9: 304 (1917), A. Chev. in Mém. Soc. Bot. France 8: 297 (1917)
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A complete, synonymic checklist of the Higher Plants of the World


Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life