388.0000 Family Caryophyllaceae Juss.; Gen. Pl.: 299 (1789), nom. cons.

102 genera, ca. 3250 spp. Higher systematics: Xue et al. (2023)

Species listed in database: 3335; hybrids: 59

388.0401 Genus Spergularia (Pers.) J.Presl & C.Presl; Fl. tech.: 94 (1819) (nom. conserv.)

Distribution: cosmopolitan, some halophytes. Borders between this genus and Spergula very unclear.

Species listed in database: 60; hybrids: 2

species :

Spergularia aberrans I.M.Johnst.; Contrib. Gray Herb. n. s. 85: 147 (1929)

Distribution: Chile (Antofagasta)

85 CLE

species :

Spergularia andina Rohrb.; Linnaea 37: 234 (1871 publ. 1873)

Distribution: Ecuador; Peru; Bolivia (Cochabamba, La Paz, Oruro, Potosí, Tarija)

83 BOL ECU PER

species :

Spergularia arbuscula (Gay) I.M.Johnst.; Contrib. Gray Herb. n. s. 85: 40 (1929)

Distribution: Chile (Arica y Parinacota, Tarapacá, Antofagasta, Atacama, Coquimbo)

85 CLE

Synonyms: hidden

= Arenaria lignosa Phil.; Fl. Atacam. 10. (1860)

= Arenaria teretifolia Phil.; Fl. Atacam. 10. (1860)

= Paronychia arbuscula Gay; Fl. Chil. 2: 520 (1847)

= Spergularia fruticosa Phil.; Anales Univ. Chile 81: 763 (1892)

= Spergularia lignosa (Phil.) Rohrb.; Linnaea 37: 244 (1872)

= Spergularia teretifolia (Phil.) Phil.

= Tissa fruticosa (Phil.) Reiche; Anales Univ. Chile 91: 347 (1895)

= Tissa lignosa (Phil.) Reiche; Anales Univ. Chile 91: 346 (1895)

= Tissa teretifolia (Phil.) Reiche; Anales Univ. Chile 91: 346 (1895)

species :

Spergularia atrosperma R.Rossbach; Rhodora 42: 80 (1940)

Name: Blackseed Sandspurrey

Distribution: USA (California, Nevada)

76 USA

species :

Spergularia azorica (Kindb.) Lebel; Mem. Soc. Sci. Nat. Cherb. 14: 47 (1869)

Distribution: Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.)

21 AZO

Synonyms: hidden

= Lepigonum azoricum Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) in Ind. 30. t. 11, f. 15 (1863)

species :

Spergularia bocconei (Scheele) Graebn.; Asch. & Graebn., Syn. Mitteleur. Fl. 5(I): 849 (1919)

Name: Griechische Schuppenmiere (DE); Greek Sea-spurrey (EN)

Distribution: England [I]; Netherlands [I]; Germany [I]; Portugal; Spain; Baleares; France; Channel Isl. [I]; Corsica; Sardinia; Italy; Sicily; Croatia; European Turkey; Greece (coastal widespread, incl. all islands); Crete (incl. islets); Morocco; Algeria; Tunisia; Libya; Egypt (Desert Oases, NE-Egypt, Nile Delta, NW-coastal Egypt); Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl., Ilhas Desertas, Porto Santo Isl.); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); South Africa [I]; East Aegaean Isl. (widespread, incl. Rhodos); Turkey (NW-Anatolia: Bithynia, SSW-Anatolia, W-Anatolia, WN-Anatolia); Iran (N-Iran, S-Iran, W-Iran); Cyprus (C-Mountains, E-Cyprus, N-Cyprus, W-Cyprus); Lebanon (C-Lebanon, coastal W-Lebanon); Syria (coastal W-Syria, C-Syrian Desert, Jazira); Israel (N-Israel); Sinai peninsula (N-Sinai); Saudi Arabia (Hejaz, Nejd Desert); Bahrain; United Arab Emirates; Oman (Mascat & Oman); New Zealand [I]; USA [I] (California [I], Oregon [I]); Uruguay [I]; Argentina [I]; Chile [I] (Coquimbo [I], Valparaíso [I], Metropolitana de Santiago [I], Ñuble [I], Biobío [I], Araucanía [I], Los Rios [I], Magallanes [I]); Desventuradas Isl. [I]; Juan Fdz. Isl. [I]

10 grb 11 ger net 12 BAL cis COR FRA POR SAR SPA 13 CRO GRC ITA KRI SIC TUE 20 ALG EGY LBY MOR TUN 21 AZO CNY MDR 27 saf 34 CYP EAI IRN ISR LEB SIN SYR TUR 35 BAR OMA SAU UAE 51 nzl 73 usa 76 usa 85 arg cle dsv jnf uru

Synonyms: hidden

= Alsine atheniensis (Asch.) Druce; Proc. Linn. Soc. 1906-1907, 77 (1907)

= Alsine bocconei Scheele; Flora (Regensburg) 26: 43 (1843)

= Alsine luteola (Greene) House; Amer. Midl. Naturalist 7: 134 (1921)

= Alsine marina Webb & Berthel.; Hist. Nat. Iles Canaries 3(2

= Alsine maritima proles atheniensis (Heldr. & Sartori) Samp.; Apêndice Lista Esp. Herb. Portug.: 6 (1914)

= Alsine maritima var. pallens (Samp.) Samp.; Fl. Portugal, ed. 2: 344 (1947), nom. illeg.

= Arenaria bocconei (Scheele) Soleirol ex Steud.; Nomencl. Bot., ed. 2, 1: 123 (1840)

= Corion atheniense (Asch.) Merino; Fl. Galic. 1: 519 (1905)

= Corion atheniense var. salinarium (Samp.) Merino; Fl. Galicia 1: 519 (1905)

= Lepigonum campestre (L.) Kindb.; Monogr. Gen. Lepigon. 35 (1863)

= Spergula atheniensis (Asch.) Murb.; Acta Univ. Lund. 32(12): 45 (1896)

= Spergula bocconei (Scheele) Pedersen; Fl. Patagonica 4a: 256 (1984)

= Spergula jallui Maire; Bull. Soc. Hist. Nat. Afrique N. 27: 214 (1936)

= Spergula rubra subsp. atheniensis (Asch.) Maire; Jahand. & Maire, Cat. Pl. Maroc: 210 (1932)

= Spergula rubra var. decipiens (Sarato ex Burnat) Maire & Weiller; Fl. Afrique N. 9: 123 (1963)

= Spergula rubra var. jallui (Maire) Maire; Cat. Pl. Maroc 4: 982 (1941)

= Spergula rubra var. marcelii Sennen ex Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 171 (1932)

= Spergula salina var. decipiens (Sarato ex Burnat) Maire; Bull. Soc. Hist. Nat. Afrique N. 26: 190 (1935)

= Spergularia atheniensis Asch.; Schweinf., Beitr. Fl. Aethiopiens: 207 (1867)

= Spergularia atheniensis proles salinaria (Samp.) Samp.; Ann. Sci. Nat. (Oporto) 10: 22 (1906)

= Spergularia atheniensis var. decipiens Sarato ex Burnat; Fl. Alpes Marit. 1: 272 (1892)

= Spergularia atheniensis var. salinaria Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 160 (1904)

= Spergularia bourgeaui Lebel; Mem. Soc. Sci. Nat. Cherbourg 14: 42 (1869)

= Spergularia campestris (Kindb.) Willk.; Willk. & Lange, Prodr. Fl. Hispan. 3: 165 (1874), nom. illeg., non Asch.

= Spergularia diandra subsp. campestris (Kindb.) Nyman; Consp. Fl. Eur.: 123 (1878)

= Spergularia marcelii Sennen; Diagn. Nouv.: 105, 184 (1936)

= Spergularia modesta proles atheniensis (Heldr. & Sartori) Samp.; Bol. Soc. Brot. 24: 27 (1909), nom. illeg.

= Spergularia modesta Samp.; Bol. Soc. Brot. 24: 27 (1908)

= Spergularia modesta var. pallens Samp.; Man. Fl. Port.: 285 (1911)

= Spergularia modesta var. pallescens Samp.; Ann. Sci. Acad. Polytecn. Porto 5: 50 (1909)

= Spergularia modesta var. salinaria (Samp.) Samp.; Ann. Sci. Acad. Polytecn. Porto 5: 50 (1909)

= Spergularia rubra subsp. atheniensis (Asch.) Rouy & Foucaud; Fl. France 3: 310 (1896)

= Spergularia rubra subsp. marcelii Sennen ex Maire

= Spergularia rubra var. atheniensis Heldr. & Sartori; Exsicc. (Herb. Graec. Norm.) 1856: no. 590 (1856)

= Spergularia rubra var. campestris (Kindb.) Batt.; Batt. & Trab., Fl. Algérie, Dicot.: 161 (1888), nom. illeg.

= Tissa campestris (Kindb.) Prantl; Engl. & Prantl, Natürl. Pflanzenfam. 3: 1b. 85 (1889)

= Tissa luteola Greene; Pittonia 5: 114 (1903)

species :

Spergularia canadensis (Pers.) G.Don; Gen. Syst. 1: 426 (1831)

subspecies :

subsp. canadensis

Name: Canadian Sandspurrey

Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Nunavut, Ontario, Prince Edward Isl., Quebec); St. Pierre and Miquelon; USA (Connecticut, Maine, Massachusetts, New Hampshire, New York, Rhode Island)

70 CDN 72 CDN 75 USA

Synonyms: hidden

= Alsine canadensis (Pers.) Samp.; Herb. Portug.: 79 (1913)

= Arenaria canadensis Pers.; Syn. 1: 504 (1805)

= Melargyra canadensis (Pers.) Raf.; Fl. Tellur. 3: 81 (1837)

= Spergula canadensis (Pers.) D.Dietr.; Syn. Pl. 2: 1598 (1840)

= Tissa canadensis (Pers.) Britton; Mem. Torr. Bot. Club, v. 152 (1894)

subspecies :

subsp. occidentalis (R.Rossbach) A.Haines; Stantec Bot. Notes (2010)

Name: Western Sandspurrey

Distribution: Alaska; Canada (British Columbia, Saskatchewan); USA (California, Oregon, Washington State)

70 ASK 71 CDN 73 USA 76 USA

Synonyms: hidden

= Spergularia canadensis var. occidentalis R.Rossbach; Rhodora 42: 116 (1940)

species :

Spergularia capillacea (Kindb. & Lange) Willk.; Willk. & Lange, Prodr. Fl. Hispan. 3: 163 (1874)

Distribution: Portugal; Spain

12 POR SPA

Synonyms: hidden

= Alsine radicans var. capillacea (Kindb.) Samp.; Lista Esp. Herb. Port.: 79 (1913)

= Corion radicans proles capillaceum (Kindb.) Merino; Fl. Galicia 1: 518 (1905)

= Lepigonum capillaceum Kindb. & Lange; Bot. Notiser 1857: 46 (1857)

= Spergula capillacea (Kindb. & Lange) G.López; Lagascalia 30: 16 (2010)

= Spergularia capillacea var. glandulosa Cout.; Fl. Portugal: 205 (1913), nom. illeg.

= Spergularia capillacea var. glandulosa Rouy; Ill. Pl. Eur. 1: 2 (1895)

= Spergularia radicans proles capillacea (Kindb.) Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 158 (1904)

= Spergularia radicans var. capillacea (Kindb.) Samp.; Man. Fl. Port.: 285 (1911)

= Spergularia rubra subsp. capillacea (Kindb.) Rivas Mart.; Publ. Inst. Biol. Aplicada 42: 122 (1967)

species :

Spergularia collina I.M.Johnst.; Contrib. Gray Herb. n. s. 81: 89 (1928)

Distribution: SW-Peru

83 PER

species :

Spergularia colombiana Rossbach; Rhodora 42: 180 (1940)

Distribution: Colombia (Cundinamarca)

82 CLM

species :

Spergularia confertiflora Steud.; Flora 39: 425 (1856)

Distribution: Desventuradas Isl.; Juan Fdz. Isl.

85 DSV JNF

Synonyms: hidden

= Arenaria rubra var. polyphylla Phil.

= Spergularia confertiflora var. glaberrima Skottsb.; Nat. Hist. Juan Fernandez, reimpr. 2: 123 (1921)

= Spergularia confertiflora var. polyphylla (Phil.) Skottsb.; Nat. Hist. Juan Fernandez (Botany) 2: 123 (1921)

= Spergularia polyphylla (Phil.) Rohrb.; Linnaea 37: 232 (1872)

= Tissa polyphylla (Phil.) Reiche; Anales Univ. Chile 91: 345 (1895)

species :

Spergularia congestifolia I.M.Johnst.; Contrib. Gray Herb. n. s. 81: 90 (1928)

Distribution: SW-Peru

83 PER

species :

Spergularia cremnophila I.M.Johnst.; Contrib. Gray Herb. n. s. 85: 41 (1929)

Distribution: Chile (Antofagasta, Atacama)

85 CLE

species :

Spergularia denticulata (Phil.) Phil.; Anales Univ. Chile 81: 769 (1892)

Distribution: Chile (Antofagasta, Atacama)

85 CLE

Synonyms: hidden

= Arenaria denticulata Phil.; Fl. Atacam. 10 (1860)

= Tissa denticulata (Phil.) Reiche; Anales Univ. Chile 91: 347 (1895)

species :

Spergularia depauperata (Naudin) Rohrb.; Linnaea 37: 231 (1872)

Distribution: Argentina (Neuquen, Rio Negro, San Juan); Chile (Valparaíso, Metropolitana de Santiago, Ñuble, Araucanía, Los Lagos)

85 ARG CLE

Synonyms: hidden

= Arenaria depauperata Naudin; Gay, Fl. Chil. 1: 270 (1846)

= Lepigonum depauperatum (Naudin) Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 25. (1863)

= Lepigonum grandiflorum Kindb.; Monogr. Gen. Lepigon. 34 (1863)

= Spergula depauperata (Naudin) Pedersen; Fl. Patagonica (Colecc. Cient. 8), 4a: 258 (1984)

= Spergularia brevinodis Kunze ex Kindb.; Monogr. Gen. Lepigon.: 34 (1863)

= Spergularia depauperata (Naudin) Phil.; Anales Univ. Chil. 82: 765 (1893), isonym

= Spergularia depauperata var. tenella (Phil.) Hauman & Irigoyen; Anales Mus. Nac. Hist. Nat. Buenos Aires 32: 191 (1923)

= Spergularia grandiflora Poepp. ex Kindb.; Monogr. Gen. Lepigon. 34. (1863)

= Spergularia rengifoi Phil.; Anales Univ. Chile 81: 769 (1892)

= Spergularia tenella Phil.; Anales Univ. Chil. 82: 766 (1893)

= Tissa depauperata (Naudin) Reiche; Anales Univ. Chile 91: 347 (1895)

= Tissa depauperata var. tenella (Phil.) Reiche; Anales Univ. Chile 91: 348 (1895)

species :

Spergularia diandra (Guss.) Heldr.; Herb. Graec. Norm. no. 492. no. 1124 (1855)

Name: Zweimännige Schuppenmiere (DE); Two-stamen Sandspurrey (EN)

Distribution: Germany [I]; Portugal; Spain; Baleares; France; Corsica; Sardinia; Italy; Sicily; Isole Pelagie (Lampedusa); Malta; Albania; European Turkey; Greece (Attiki, Skiros, E-Peloponnisos, widespread Kiklades); Crete (incl. islets); E-European Russia; Morocco; Algeria; Tunisia; Libya; Egypt (Desert Oases, Eastern Desert, Great Southwestern Desert, NE-Egypt, Nile Delta, NW-coastal Egypt, SE-Egypt); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria); Siberia (Altai, W-Siberia); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Georgia [Caucasus]; Armenia; Azerbaijan (incl. Nachichevan); East Aegaean Isl. (Lipsi, Chalki, Rhodos); Turkey (S-Anatolia); Iraq (NW-Iraq, SE-Iraq: Mesopotamia, S-Iraq, W-Iraq: Desert); Iran (EC-Iran, NE-Iran: Mts., N-Iran, S-Iran, W-Iran); Afghanistan (Badghys, Farah, Helmand, Herat, Kandahar, Laghman, Qunduz, Samangan, Zabul); Cyprus (C-Mountains, E-Cyprus, N-Cyprus, W-Cyprus); Syria (C-Syrian Desert, Jazira, NW-Syria); Israel (C-Israel, coastal W-Israel, Rift Valley, N-Israel, N-Negev Desert, Judean Desert, Negev Desert); Sinai peninsula (C-Sinai, N-Sinai, S-Sinai); Jordania (S-Jordania); Saudi Arabia (C-Saudi Arabia, Dahana, NE-Saudi Arabia, Hejaz, Asir); Kuwait; Bahrain; Qatar; United Arab Emirates; Oman (Mascat & Oman); China (Gansu, Ningxia, Qinghai, Xinjiang); Pakistan (Baluchistan, Waziristan, N. W. Frontier Prov., Peshawar, Pakistani Punjab, Rawalpindi); Canada [I] (Alberta [I], British Columbia [I], Saskatchewan [I]); USA [I] (Idaho [I], Massachusetts [I], Oregon [I], Washington [I]); Argentina [I]

11 ger 12 BAL COR FRA POR SAR SPA 13 ALB GRC ITA KRI MLT PEL SIC TUE 14 RUS 20 ALG EGY LBY MOR TUN 21 CNY 30 SIB 32 KAZ KGZ TKM TZK UZB 33 ARM AZE GEO 34 AFG CYP EAI IRN IRQ ISR JOR SIN SYR TUR 35 BAR KUW OMA QAT SAU UAE 36 CHI 40 PAK 71 cdn 73 usa 75 usa 85 arg

Synonyms: hidden

= Alsine bracteata (B.L.Rob.) House; Amer. Midl. Naturalist 7: 134 (1921)

= Alsine diandra (Guss.) House; Am. Midl. Nat. 7: 133 (1921)

= Alsine diandra Guss.; Fl. Sicul. Syn. 1: 501 (1843)

= Alsine rubra C.A.Mey.; Verz. Pfl. Casp. Meer. (C. A. Mey.). 217 (1831) non (L.) Crantz

= Arenaria diandra Guss.; Fl. Sicul. Prodr. 1: 515 (1827)

= Arenaria salsuginea Bunge ex Ledeb.; Fl. Altaic. [Ledeb.]. 2: 163 (1830)

= Buda diandra (Guss.) Kuntze; Revis. Gen. Pl. 50 (1891)

= Lepigonum diandrum (Guss.) Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 35 (1863), isonym

= Lepigonum diandrum (Guss.) Nyman; Syll. 250

= Lepigonum diandrum var. viscidum Regel; Index Seminum (LE, Petropolitanus) 1856: 35 (1856)

= Lepigonum melanospermum Lange ex Kindb.; Monogr. Gen. Lepigon.: 42 (1863)

= Lepigonum salsugineum (Bunge) Fisch. & C.A.Mey.; Index Sem. Hort. Petrop. 1: 10 (1835)

= Lepigonum salsugineum Kindb.; Monogr. Gen. Lepigon.: 42 (1863)

= Lepigonum salsugineum var. leiospermum Bunge; Index Seminum (TU, Dorpatensis) 1843: viii (1843)

= Spergula diandra (Guss.) Murb.; Contr. Fl. Tunisie [Murbeck] 1, Acta Univ. Lund 33(12): 44 (1897)

= Spergula diandra var. leiosperma (Bunge) Maire; Bull. Soc. Hist. Nat. Afrique N. 30: 268 (1939)

= Spergula diandra var. maura Pau & Sennen

= Spergula dillenii Lebel; Mem. Soc. Sci. Nat. Cherbourg 14: 43 (1869)

= Spergularia adenophora Vved.; Bot. Mat. Gerb. Bot. Inst. Uzb. Fil. Akad. Nauk SSSR 3: 4 (1941)

= Spergularia alleluia Sennen; Diagn. Nouv.: 210 (1936)

= Spergularia amurensis Pomel; Nouv. Mat. Fl. Atl. 205 (1875)

= Spergularia balearica Sennen; Bol. Soc. Iber. Ci. Nat. 25: 219 (1927)

= Spergularia bracteata (B.L.Rob.) A.Nelson & J.F.Macbr.; Bot. Gaz. 61: 30 (1916)

= Spergularia degenii Sennen; Diagn. Nouv.: 210 (1936)

= Spergularia diandra (Guss.) Boiss.; Fl. Orient. 1: 733 (1867), isonym

= Spergularia diandra var. leiosperma (Bunge) Asch. & Schweinf.; Ill. Fl. Egypte: 47 (1887)

= Spergularia nematopoda Vved.; Bot. Mater. Gerb. Bot. Inst. Uzbekistansk. Fil. Akad. Nauk SSSR 3: 5 (1941)

= Spergularia patens Hochst. ex Boiss.; Fl. Orient. [Boiss. ] 1: 733 (1867)

= Spergularia rubra subsp. diandra (Guss.) Arcang.; Comp. Fl. Ital.: 107 (1882)

= Spergularia salsuginea (Bunge) Fenzl; Ledeb., Fl. Ross. 2: 166 (1843)

= Spergularia salsuginea var. bracteata B.L.Rob.; Syn. Fl. N. Amer. 1, pt. 1: 251 (1897)

= Spergularia salsuginea var. tenuissima Rouy & Foucaud; Fl. France 3: 308 (1896)

= Tissa bracteata (B.L.Rob.) Small; Fl. S.E. U.S.: 418 (1903)

= Tissa diandra (Guss.) Brandegee; Proc. Calif. Acad. Ser. II. 2: 131 (1889)

= Tissa diandra subsp. bracteata (B.L.Rob.) Piper; Contr. U.S. Natl. Herb. 11: 264 (1906)

species :

Spergularia diandroides L.G.Adams; Austral. Syst. Bot. 21(4): 266 (2008)

Distribution: Australia (South Australia, Northern Territory, New South Wales, Queensland, Western Australia)

50 AUS

species :

Spergularia doumerguei Foucaud & Monnier; Nat. Monspel., Ser. Bot., Fasc. 19, 101 (1968)

Distribution: Morocco; Algeria

20 ALG MOR

species :

Spergularia echinosperma (Celak.) Asch. & Graebn.; Ber. Deutsch. Bot. Ges. 11: 516 (1893)

subspecies :

subsp. echinosperma

Name: Bristleseed Sandspurrey (EN), Igelsamige Schuppenmiere (DE)

Distribution: Germany (Brandenburg); Poland; Czech Republic; Slovakia; France; Morocco; USA [I] (Alabama [I], Georgia [I], Louisiana [I], Texas [I])

11 CZE GER POL SLO 12 FRA 20 MOR 77 usa 78 usa

Synonyms: hidden

= Spergula echinosperma (Celak.) E.H.L.Krause; Sturm, Fl. Deutschland, ed. 2, 5: 21 (1901)

= Spergula rubra subsp. radiata Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 171 (1932)

= Spergularia campestris subsp. echinosperma (Celak.) Hegi; 422 (1911)

= Spergularia echinosperma (Hayata) S.S.Ying; New Taxa New Names 2: 40 (2022), isonym

= Spergularia rubra subsp. echinosperma Celak.; Arch. Naturw. Land. Boehm. (Prodr. Fl. Böhmen) 4: 867 (1881)

subspecies :

subsp. albensis Kúr, Amarell, Jage & Stech; Phytotaxa 305(3) : 157 (2017)

Name: Braunsamige Schuppenmiere (DE)

Distribution: Germany (Brandenburg, Mecklenburg-Vorpommern, Niedersachsen, Schleswig-Holstein, Sachsen, Sachsen-Anhalt); N-Czech Republic

11 CZE GER

species :

Spergularia embergeri Monnier; Nat. Monspel., Ser. Bot., Fasc. 19, 97 (1968)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Spergularia marginata var. melilensis Sennen & Mauricio; [nom. nud.]

= Spergularia marginata var. valerianii Sennen & Mauricio; [nom. nud.]

= Spergularia media var. latifolia Pau; Font Quer, Exsicc. (Iter Marocc.) 1929: no. 135 (1930)

species :

Spergularia fasciculata Phil.; Anal. Mus. nac. Chile (1891) 6

Name: Alhucema [Chile]

Distribution: Peru; Bolivia (La Paz); Chile (Arica y Parinacota, Tarapacá, Antofagasta)

83 BOL PER 85 CLE

Synonyms: hidden

= Spergularia laciniata Baehni & J.F.Macbr.; Publ. Field Mus. Nat. Hist., Chicago, Bot. Ser. 13: 11. 631 (1937)

= Spergularia stuebelii (Hieron.) I.M.Johnst.; Contrib. Gray Herb. n. s. 81: 90 (1928)

= Tissa fasciculata (Phil.) Hieron.; Bot. Jahrb. Syst. 21: 308 (1895)

= Tissa stuebelii Hieron.; Bot. Jahrb. Syst. 21: 308 (1895)

species :

Spergularia fimbriata Boiss. & Reut.; Boiss., Diagn. Pl. Orient. ser. 2, 1: 94 (1854)

Distribution: S-Portugal; S-Spain (Cadiz); Gibraltar; S-Morocco; Western Sahara; Canary Isl. (Fuerteventura, Gran Canaria, Tenerife)

12 GIB POR SPA 20 MOR WSA 21 CNY

Synonyms: hidden

= Alsine fimbriata (Boiss. & Reut.) Pau; Mem. Soc. Esp. Hist. Nat. 12: 292 (1924)

= Arenaria fimbriata Salzm. ex Kindb.; Monogr. Gen. Lepigon.: 22 (1863), nom. inval.

= Lepigonum fimbriatum (Boiss. & Reut.) Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 21 (1863)

= Spergula fimbriata (Boiss. & Reut.) Murb.; Acta Univ. Lund. ser. 2, 18(3): 33 (1922), isonym

= Spergula fimbriata (Boiss. & Reut.) Pit. & Proust; Iles Canaries [Pit. & Proust]: 124 (1909)

= Spergula fimbriata f. albiflora (H.Lindb.) Maire; Jahand. et al., Cat. Pl. Maroc 3: 876 (1934)

= Spergula fimbriata f. tenuis (Ball) Maire; Fl. Afrique N. 9: 99 (1963)

= Spergula fimbriata subvar. glabrifolia (H.Lindb.) Maire; Jahand. et al., Cat. Pl. Maroc 3: 876 (1934)

= Spergula fimbriata subvar. tenuis (Ball) Maire; Fl. Afrique N. 9: 99 (1963)

= Spergula fimbriata var. condensata Ball; Spicil. Marocc. [Ball]: 368 (1877)

= Spergula fimbriata var. hebephylla Maire & Weiller; Fl. Afrique N. 9: 99 (1963)

= Spergula fimbriata var. pulvinata Sauvage; Bull. Soc. Sci. Nat. Maroc 25-27: 365 (1949)

= Spergula fimbriata var. tenuis Ball; Spicil. Marocc. [Ball]: 369 (1877)

= Spergularia fimbriata f. albiflora H.Lindb.; Acta Soc. Sci. Fenn., Ser. B, Opera Biol. 1(2): 49 (1932)

= Spergularia fimbriata subsp. condensata (Ball) H.Lindb.; Itin. Medit., Acta Soc. Sci. Fenn., Nov. Ser. B, 1(2): 49 (1932)

= Spergularia fimbriata var. condensata Ball; Spicil. Fl. Marocc. [Ball]: 368 (1877)

= Spergularia fimbriata var. glabrifolia H.Lindb.; Acta Soc. Sci. Fenn., Ser. B, Opera Biol. 1(2): 49 (1932)

= Spergularia fimbriata var. tenuis Ball; Spicil. Fl. Marocc. [Ball]: 369 (1877)

= Spergularia rubra var. fimbriata (Boiss. & Reut.) Knoche; Fl. Baléar. 1: 458 (1921)

variant :

var. interclusa Svent.; Addit. Fl. Canar.: 7 (1960)

Distribution: Canary Isl. (Lanzarote)

21 CNY

species :

Spergularia flaccida (Madden) I.M.Turner; Willdenowia 47(3): 214 (2017)

Name: Trügerischer Spörgel (DE)

Distribution: Germany [I]; SE-Spain; Sicily; Morocco; Algeria (incl. Hoggar); Tunisia; Libya; Egypt (Eastern Desert, Great Southwestern Desert, NE-Egypt, NW-coastal Egypt, SE-Egypt); Madeira (Madeira Isl., Ilhas Desertas, Porto Santo Isl.); Islas Selvagens; Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro); Sudan; NE-Somalia; Soqotra; Iraq (SE-Iraq: Mesopotamia, S-Iraq); Iran (S-Iran, W-Iran); Afghanistan; Syria (C-Syrian Desert); Israel (C-Israel, coastal W-Israel, Rift Valley, Judean Desert, Negev Desert); Sinai peninsula (C-Sinai, S-Sinai); Jordania (S-Jordania); Saudi Arabia (C-Saudi Arabia, Dahana, NE-Saudi Arabia, N-Saudi Arabia, Nafud Desert, Hejaz, Asir, Nejd Desert); Kuwait; Bahrain; Qatar; United Arab Emirates; Oman (Mascat & Oman); Yemen (W-Yemen); Pakistan (Karachi, Baluchistan, Peshawar, Pakistani Punjab); India (widespread)

11 ger 12 SPA 13 SIC 20 ALG EGY LBY MOR TUN 21 CNY MDR SEL 24 SOC SOM SUD 34 AFG IRN IRQ ISR JOR SIN SYR 35 BAR KUW OMA QAT SAU UAE YEM 40 IND PAK

Synonyms: hidden

= Arenaria flaccida Roxb.; Hort. Beng. 34 (1814)

= Lepigonum eximium Kindb.; Monogr. Gen. Lepigon. 32 (1863)

= Lepigonum fallax (Lowe) Lowe; Fl. Salvag. Tent. 9 (1869)

= Spergula fallax (Lowe) E.H.L.Krause; Sturm, Fl. Deutschland ed. 2, 5-19 (1901)

= Spergula flaccida (Madden) Asch.; Verh. Bot. Vereins Prov. Brandenburg 30: XLIII. 1889 (nom. illeg.)

= Spergula flaccida f. purpurascens Maire & Sauvage; Bull. Soc. Hist. Nat. Afrique N. 32: 204 (1941)

= Spergula indica Royle; Ill. Bot. Himal. Mts. 78 (1834), nom. inval.

= Spergula pentandra subsp. flaccida (Madden) Jafri; Fl. Karachi 124 (1966)

= Spergula pentandra var. intermedia Boiss.; Diagn. Pl. Orient., ser. 2, 1: 93 (1853)

= Spergularia fallax Lowe; Hooker´s J. Bot. Kew Gard. Misc. 8: 289 (1856)

= Stipularia fallax Roxb. ex Edgew.; J. Proc. Linn. Soc., Bot. 6: 185 (1862)

= Stipularia flaccida Madden; J. Asiat. Soc. Bengal 18: 621 (1849)

species :

Spergularia floribunda (Gay) Rohrb.; Linnaea 37: 230 (1871 publ. 1873)

Distribution: Chile (Atacama, Coquimbo)

85 CLE

Synonyms: hidden

= Arenaria floribunda Gay; Fl. Chil. 1: 269 (1846)

= Lepigonum floribundum (Gay) Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 5 (1863)

= Spergularia coquimbensis Phil.; Anales Univ. Chile 81: 764 (1892)

= Tissa coquimbensis (Phil.) Reiche; Anales Univ. Chile 91: 346 (1895)

= Tissa floribunda (Gay) Reiche; Anales Univ. Chile 91: 345 (1895)

species :

Spergularia glandulosa (Jacq.) Heynh.; Alph. Aufz. Gew.: 689 (1847)

Distribution: South Africa (W-Cape Prov.)

27 SAF

Remarks: provisionally accepted name

Synonyms: hidden

= Arenaria glandulosa Jacq.; Pl. Hort. Schoenbr. 3: 56 (1798)

= Buda glandulosa (Jacq.) Desv.; J. Bot. Agric. 3: 226 (1816)

= Lepigonum glandulosum (Jacq.) Fisch. & C.A.Mey.; Index Seminum (LE, Petropolitanus) 1: 10 (1835)

= Lepigonum marginatum var. glandulosum (Jacq.) Sond.; Fl. Cap. (Harvey) 1: 135 (1860)

species :

Spergularia hanoverensis E.Simon ex M.Á.Alonso, M.B.Crespo, Mart.-Azorín & Mucina; Plants [Basel] 12(13)-2481: 7 (2023)

Distribution: South Africa (N-Cape Prov., W-Cape Prov.)

27 SAF

species :

Spergularia heldreichii Foucaud; Note sur le Spergularia rubra var. pinguis Fenzl, 5 (1903)

Name: Heldreichs Schuppenmiere (DE); Heldreich´s Sea-spurrey (EN)

Distribution: Portugal; Spain; Gibraltar; Baleares; France; Corsica; Italy; Morocco; ?Algeria; Tunisia; East Aegaean Isl. (Rhodos)

12 BAL COR FRA GIB POR SPA 13 ITA 20 MOR TUN 34 EAI

Synonyms: hidden

= Spergula heldreichii (Foucaud) G.López; Lagascalia 30: 17 (2010)

= Spergularia heldreichii E.Simon sec. & P.Monnier; Bull. Soc. Bot. France 105: 263 (1958)

= Spergularia insularis Foucaud; Ann. Soc. Sci. Nat. Charente-Infér. 32: 211 (1899)

= Spergularia rubra subsp. heldreichii (E.Simon sec. & P.Monnier) O.Bolòs & Vigo; Butll. Inst Catalana Hist. Nat. 38: 87 (1974)

species :

Spergularia hohuanshanensis S.S.Ying; New Taxa New Names 2: 34 (2022)

Distribution: Taiwan

38 TAI

Remarks: provisionally accepted name

species :

Spergularia kurkae F.Dvorák; Scripta Fac. Sci. Nat. Univ. Purkyn. Brunensis, Biol. 19(9): 320 (1989), pro hybr.

Distribution: Germany (Brandenburg, Niedersachsen, Sachsen, Sachsen-Anhalt); Czech Republic; Slovakia

11 CZE GER SLO

Remarks: derived from Spergularia echinosperma x rubra

Synonyms: hidden

= Spergula ×kurkae (F.Dvorák) B.Bock; Bull. Soc. Bot. Centre-Ouest 43: 218 (2012)

species :

Spergularia macrorrhiza (Req. ex Loisel.) Heynh.; Alph. Aufz. Gew. 689 (1847)

Distribution: Corsica; Sardinia; Italy

12 COR SAR 13 ITA

Synonyms: hidden

= Arenaria macrorrhiza Req. ex Loisel.; Nouv. Not. 22

= Lepigonum macrorhizum Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 22 (1863)

= Spergula macrorrhiza (Req. ex Loisel.) B.Bock & J.-M.Tison; Bull. Soc. Bot. Centre-Ouest 43: 212 (2012)

species :

Spergularia macrotheca (Hornem. ex Cham. & Schltdl.) Heynh.; Nom. 2: 689 (1846)

Name: Sticky Sandspurrey

Distribution: Canada (British Columbia); USA (California, Oregon, Washington); Mexico (Baja California Norte); Isla Guadalupe

71 CDN 73 USA 76 USA 79 GLP MEX

Synonyms: hidden

= Alsine greenei House; Amer. Midl. Naturalist 7: 134 (1921)

= Alsine macrotheca (Hornem. ex Cham. & Schltdl.) House; Am. Midl. Nat. 7: 134 (1921)

= Alsine valida (Greene) House; Amer. Midl. Naturalist 7: 134 (1921)

= Arenaria macrotheca Hornem. ex Cham. & Schltdl.; Linnaea 1: 53 (1826)

= Buda macrotheca (Hornem. ex Cham. & Schltdl.) Kuntze; Revis. Gen. Pl. 50 (1891)

= Lepigonum chilense Fisch. & C.A.Mey.; Index Seminum (LE, Petropolitanus) 3: 14 (1837)

= Lepigonum macrothecum (Hornem. ex Cham. & Schltdl.) Fisch. & C.A.Mey.; Index Sem. Hort. Petrop. 3: 14 (1835)

= Spergularia chilensis (Fisch. & C.A.Mey.) A.Braun & C.D.Bouché; Index Seminum (B, Berolinensis) 1862: 10 (1862)

= Spergularia macrotheca (Hornem. ex Cham. & Schltdl.) B.L.Rob.; Proc. Amer. Acad. Arts 29: 312 (1894), isonym

= Spergularia macrotheca var. scariosa (Britton) B.L.Rob.; Proc. Amer. Acad. Arts 29: 313 (1894)

= Spergularia macrotheca var. talinum (Greene) Jeps.; Fl. Calif. 1: 493 (1915)

= Tissa macrotheca (Hornem. ex Cham. & Schltdl.) Brandegee; Proc. Calif. Acad. Ser. II. 2: 131 (1889)

= Tissa macrotheca var. scariosa Britton; Bull. Torrey Bot. Club 16: 129 (1889)

= Tissa pallida Greene ex Britton; Bull. Torrey Bot. Club 16: 129 (1889)

= Tissa talinum Greene; Erythea 1: 106 (1893)

= Tissa valida Greene; Erythea 1: 107 (1893)

variant :

var. leucantha (Greene) B.L.Rob.; Proc. Amer. Acad. Arts. 29: 313 (1894)

Distribution: USA (California)

76 USA

Synonyms: hidden

= Alsine leucantha (Greene) House; Am. Midl. Nat. 7: 134 (1921)

= Spergularia macrotheca var. glabra (Davidson) Munz; Man. S. Calif. Bot.: 164 (1935)

= Tissa leucantha Greene; Pittonia 1: 301 (1889)

= Tissa leucantha var. glabra Davidson; Bull. S. Calif. Acad. Sci. 25: 84 (1926)

= Tissa macrotheca var. leucantha (Greene) Jeps.; Fl. W. Calif.: 170 (1901)

variant :

var. longistyla R.Rossbach; Rhodora. 42: 78, plate 589 (1940)

Distribution: USA (California)

76 USA

species :

Spergularia manicata (Skottsb.) Kool & Thulin; Taxon 66(3): 620 (2017)

Distribution: Desventuradas Isl. (San Ambrosio Isl.)

85 DSV

Synonyms: hidden

= Paronychia manicata Skottsb.; Acta Hort. Gotoburg. 17: 50 (1947)

= Sanctambrosia manicata Skottsb. ex Kuschel; Arkiv Bot., Stockh., andra ser. 4: 418 (1962), in adnot., Skottsb. in Arkiv Bot., Stockh., andra ser. 4: 469 (1963)

species :

Spergularia marina (L.) Besser; Enum. Pl.: 97 (1822)

Name: Salz-Schuppenmiere (DE); Salt Sandspurrey (EN)

Distribution: Iceland; England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Germany (Brandenburg, Baden-Württemberg, Bayern, Hessen, Hessen [I], Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Gibraltar; Baleares; France; Channel Isl.; Corsica; Sardinia; Italy; Sicily; Isole Pelagie (Lampedusa); Malta; Slovenia; Croatia; Montenegro; Serbia; Albania; Romania; Bulgaria; European Turkey; Greece (coastal widespread, incl. almost all islands); Crete (incl. islets); Estonia; Latvia; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Morocco; Western Sahara; Algeria; Tunisia; Libya; Egypt (Great Southwestern Desert, Nile Delta, Nile Valley, NW-coastal Egypt); Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Corvo Isl.); Madeira (Madeira Isl., Porto Santo Isl.); Canary Isl. (Gran Canaria, Tenerife, Lanzarote, Fuerteventura); Mauritania; N-Chad (Tibesti); Siberia (Altai, Buryatia, Chita, Irkutsk, W-Siberia, Yakutia); Russian Far East (Khabarovsk, Kuril Isl., Primorye, Sakhalin); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; East Aegaean Isl. (widespread, incl. Rhodos); Turkey (N-Anatolia, NW-Anatolia: Bithynia, S-Anatolia, SSW-Anatolia, SW-Anatolia, W-Anatolia); Iraq (NE-Iraq, NW-Iraq, SE-Iraq: Mesopotamia, S-Iraq, W-Iraq: Desert); Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Faryab, Ghazni, Ghorat, Helmand, Kabul, Qunduz, Takhar, Zabul); Cyprus (C-Mountains, E-Cyprus, N-Cyprus, W-Cyprus); Lebanon (Antilebanon, coastal W-Lebanon); Syria (C-Syrian Desert, Jazira, NW-Syria); Israel (C-Israel, coastal W-Israel, Rift Valley); Sinai peninsula (C-Sinai, N-Sinai); Jordania (S-Jordania, W-Jordania); Saudi Arabia (NE-Saudi Arabia, Asir); Kuwait; Bahrain; United Arab Emirates; Yemen (W-Yemen); China (Gansu, Hebei, Heilongjiang, Henan, Jiangsu, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan, Xinjiang, Yunnan, Zhejiang); Mongolia; South Korea; Japan (Hokkaido, Honshu, Kyushu); Australia [I] (Western Australia [I], South Australia [I], New South Wales [I], Victoria [I], Tasmania [I]); New Zealand [I]; Hawaii [I] (Kure Isl. [I], Midway Isl. [I], French Frigate Shoals [I], Kauai [I], Oahu [I], Molokai [I], Maui [I]); Canada [I] (Alberta [I], British Columbia [I], Manitoba [I], New Brunswick [I], N. W. T. [I], Nova Scotia [I], Ontario [I], Prince Edward Isl. [I], Quebec [I], Saskatchewan [I]); St. Pierre and Miquelon [I]; USA [I] (Alabama [I], Arizona [I], California [I], Colorado [I], Connecticut [I], Delaware [I], District of Columbia [I], Florida [I], Georgia [I], Idaho [I], Illinois [I], Iowa [I], Kentucky [I], Louisiana [I], Maine [I], Maryland [I], Massachusetts [I], Michigan [I], Mississippi [I], Missouri [I], Montana [I], Nebraska [I], Nevada [I], New Hampshire [I], New Jersey [I], New Mexico [I], New York [I], North Carolina [I], North Dakota [I], Ohio [I], Oklahoma [I], Oregon [I], Pennsylvania [I], Rhode Island [I], South Carolina [I], Texas [I], Utah [I], Virginia [I], Washington [I], Wisconsin [I], Wyoming [I]); Mexico [I]; Isla Guadalupe [I]; Colombia [I]; Brazil [I]; Uruguay [I]; Argentina [I]; Chile [I] (Coquimbo [I], Valparaíso [I], Metropolitana de Santiago [I], Biobío [I], Los Lagos [I], Aisén [I], Magallanes [I]); Falkland Isl. [I]

10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN NET POL SLO 12 BAL CIS COR FRA GIB POR SAR SPA 13 ALB BUL CRO GRC ITA KRI MLT MNT PEL ROM SIC SLV SRB TUE 14 BLR EST KRY LAT MLD RUS UKR 20 ALG EGY LBY MOR TUN WSA 21 AZO CNY MDR 22 MTN 24 CHA 30 SIB 31 RFE 32 KAZ KGZ TKM TZK UZB 33 ARM AZE GEO NCS 34 AFG CYP EAI IRN IRQ ISR JOR LEB SIN SYR TUR 35 BAR KUW SAU UAE YEM 36 CHI 37 MON 38 JAP KOR 50 aus 51 nzl 63 haw 70 cdn 71 cdn 72 cdn 73 usa 74 usa 75 usa 76 usa 77 usa 78 usa 79 glp mex 82 clm 84 bra 85 arg cle uru 91 fal

Remarks: validity of name S. marina vs. S. salina highly disputed

Synonyms: hidden

= Alsine atheniensis proles marina (L.) Samp.; Lista Esp. Herb. Port.: 79 (1913)

= Alsine atheniensis proles urbica (Leffler) Samp.; Lista Esp. Herb. Port.: 19 (1913)

= Alsine marginata C.A.Mey.; Verz. Pfl. Casp. Meer 217 (1831), nom. illeg.

= Alsine marina (L.) Mert. & W.D.J.Koch; Röhl., Deutschl. Fl., ed. 3, 2: 293 (1826)

= Alsine marina var. succosior Mert. & W.D.J.Koch; Röhl. Deutsch. Fl. 2: 294 (1831)

= Alsine maritima proles urbica (Leffler) Samp.; Lista Esp. Herb. Port., Ap. 2: 6 (1914)

= Alsine salina (J.Presl & C.Presl) Heldmann; Oberhess. Fl. (1837)

= Alsine salina (J.Presl & C.Presl) J.Groves; Bab. Man. Brit. Bot. ed. 9: 68 (1904)

= Alsine sparsiflora (Greene) House; Amer. Midl. Naturalist 7: 134 (1921)

= Alsine tenuis (Greene) House; Amer. Midl. Naturalist 7: 133 (1921)

= Alsinella media Hornem.; Nomencl. Fl. Danic. 33 (1827)

= Arenaria halophila (T.Marsson) Bunge; Ledeb., Fl. Altaica 2: 163 (1830)

= Arenaria heterosperma Guss.; Suppl. Fl. Sicul. Prodr.: 129 (1832)

= Arenaria marina (L.) All.; Fl. Pedem. 2: 114 (1785)

= Arenaria marina (L.) Weber; Suppl. Fl. Holsat. 7 (1787), nom. illeg.

= Arenaria maritima (L.) Steud.; Nomencl. Bot. ed. 2, 1: 125 (1840), nom. illeg.

= Arenaria media DC.; Prodr. 1: 104 (1824), nom. illeg.

= Arenaria miquelonensis Bach.Pyl. ex Lebel; Bull. Soc. Bot. France 15: 58 (1868), nom. inval.

= Arenaria rubra subsp. marina (L.) Ehrh.; Hannover Mag. 18: 222 (1780)

= Arenaria rubra var. marina L.; Sp. Pl. 423 (1753)

= Arenaria salina (J.Presl & C.Presl) Ser.; DC., Prodr. 1: 401 (1824)

= Buda marina (L.) Dumort.; Fl. Belg.: 110 (1827)

= Buda marina var. minor S.Watson; A. Gray, Manual, ed. 6: 90 (1890)

= Buda media var. marina (Griseb.) Kuntze; Revis. Gen. Pl. 1: 50 (1891)

= Corion halophilum (T.Marsson) Merino; Fl. Galic. 1: 514 (1905)

= Corion halophilum var. dillenii (Lebel) Merino; Fl. Galic. 1: 515 (1905)

= Corion halophilum var. leiospermum (Kindb.) Merino; Fl. Galic. 1: 515 (1905)

= Corion halophilum var. urbicum (Leffler) Merino; Fl. Galicia 1: 515 (1905)

= Corion marinum (L.) N.E.Br.; J. E. Sm., Engl. Bot., ed. 3, Suppl. 48 (1891)

= Corion marinum var. leiospermum (Kindb.) N.E.Br.; J. E. Sm., Engl. Bot., ed. 3, Suppl.: 48 (1891)

= Holosteum sperguloides Lehm.; Nov. Stirp. Pug. 1: 21 (1828)

= Lepigonum caninum Leffler; Oesterr. Bot. Z. 19: 103 (1869), nom. illeg.

= Lepigonum caninum var. leiospermum (Kindb.) Leffl.; 104 (1869)

= Lepigonum caninum var. salinum (J.Presl & C.Presl) Leffl.; 103 (1869)

= Lepigonum leiospermum Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 23 (1863)

= Lepigonum marinum (L.) Wahlenb.; Fl. Gothob. 1: 45 (1820)

= Lepigonum marinum (Pall.) Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 18 (1863), isonym

= Lepigonum marinum (Pall.) Wahlenb.; Fl. Gothob. 45 (1820)

= Lepigonum medium Fr.; Novit. Fl. Suec. Mant. 3: 33 (1842)

= Lepigonum neglectum Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 6 (1863)

= Lepigonum salinum (J.Presl & C.Presl) G.Don; Oesterr. Bot. Zeitschr. 10: 103 (1869)

= Lepigonum tenue Greene; Pittonia 1: 63 (1887)

= Lepigonum trachyspermum subsp. murale Kindb.; Monogr. Gen. Lepigon. 31 (1863)

= Spergula canina subsp. sperguloides (Lehm.) Ahlfv.; Neuman, Sverig. Fl. 555 (1901)

= Spergula coreana H.Lév.; Repert. Spec. Nov. Regni Veg. 11: 495 (1913)

= Spergula diandra var. australis Lebel; Mem. Soc. Sci. Nat. Cherbourg 14: 44 (1869)

= Spergula dillenii var. australis Lebel

= Spergula heterosperma (Guss.) E.A.Durand & Barratte; Fl. Libyc. Prodr.: 41 (1910)

= Spergula longicaulis Pomel; Nouv. Mat. Fl. Atlant.1: 203 (1874)

= Spergula longicaulis var. sperguloides (Lehm.) Maire; Cat. Pl. Maroc 4: 982 (1941)

= Spergula marina (L.) Bartl. & H.L.Wendl.; Beitr. Bot. 2: 64 (1825)

= Spergula salina (J.Presl & C.Presl) D.Dietr.; Syn. Pl. 1: 1598 (1840)

= Spergula salina subvar. halophila (Bunge) Maire & Weiller; Fl. Afrique N. 9: 112 (1963)

= Spergula salina var. australis (Lebel) Maire; Cat. Pl. Maroc 2: 210 (1932)

= Spergula salina var. faurei Maire; Bull. Soc. Hist. Nat. Afrique N. 31: 100 (1940)

= Spergula salina var. halophila (Bunge) Maire; Bull. Soc, Hist. Nat. Afrique N. 26: 190 (1935)

= Spergula salina var. leiosperma (Kindb.) Maire & Weiller; Fl. Afrique N. 9: 112 (1963)

= Spergula salina var. longicaulis (Pomel) Maire & Weiller; Fl. Afrique N. 9: 113 (1963)

= Spergula salina var. sperguloides (Lehm.) Maure; Jahandiez et al., Cat. Pl. Maroc 4: 982 (1941)

= Spergula salina var. stenopetala (A.Chev.) Maire & Weiller; Fl. Afrique N. 9: 113 (1963)

= Spergula salina var. urbicum (Leffler) Gürke; Cat. Pl. Maroc 2: 210 (1932)

= Spergula salina var. urbicum (Leffler) Maire; Jahand. et al., Cat. Pl. Maroc 2: 210 (1932)

= Spergula seminulifera (Hy) Maire; Jahand. & Maire, Cat. Pl. Maroc: 211 (1932)

= Spergularia alata Wiegand; Rhodora 22: 15 (1920)

= Spergularia bofillii Sennen; Bol. Soc. Iber. Ci. Nat. 25: 217 (1926 publ. 1927)

= Spergularia bungeana Tzvelev; Novosti Sist. Vyssh. Rast. 32: 29 (2000), nom. nov. for S. halophila T. Marsson

= Spergularia caballeroi Sennen & Mauricio; Cat. Fl. Rif. Orient. 22 (1933), nom. nud.

= Spergularia campestris var. pinguis (Fenzl) P.Fourn.; Quatre Fl. France: 294 (1936)

= Spergularia canina var. salina (J.Presl & C.Presl) F.Aresch.; 265 (1881)

= Spergularia dillenii Lebel; Mém. Soc. Sci. Nat. Cherbourg 14: 43 (1868)

= Spergularia dillenii proles salina (J.Presl & C.Presl) Rouy & Foucaud; Fl. France 3: 304 (1896)

= Spergularia dillenii subsp. urbica (Leffler) Rouy & Foucaud; Fl. France [Rouy & Foucaud] 3: 304 (1896)

= Spergularia dillenii var. australis Lebel; Mém. Soc. Sci. Nat. Cherbourg 14: 44 (1868)

= Spergularia dillenii var. confusa Rouy & Foucaud; Fl. France 3: 304 (1896)

= Spergularia dillenii var. confusa Rouy; Fl. France [Rouy & Foucaud] 3: 304 (1896)

= Spergularia dillenii var. pedicellaris Rouy & Foucaud; Fl. France [Rouy & Foucaud] 3: 304 (1896)

= Spergularia dillenii var. perennis Rouy & Foucaud; Fl. France [Rouy & Foucaud] 3: 303 (1896)

= Spergularia fauriei (Nakai) Nakai; Bull. Nat. Sci. Mus., Tokyo, No. 31, 38 (1952), nom. inval.

= Spergularia halophila T.Marsson; Fl. Neu-Vorpommern, 77 (1869), nom. inval.

= Spergularia halophila var. dillenii (Lebel) Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 164 (1904)

= Spergularia halophila var. marina (L.) T.Marsson; Fl. Neu-Vorpommern 77 (1869), nom. illeg.

= Spergularia halophila var. urbica (Lebel) Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 164 (1904)

= Spergularia heterosperma (Guss.) Lebel; Mém. Soc. Sci. Nat. Cherbourg 14: 45 (1868)

= Spergularia leiosperma (Kindb.) F.Schmidt; Reisen Amurl. 131 (1868)

= Spergularia longicaulis Pomel; Bull. Soc. Sci. Phys. Algérie 11: 203 (1874)

= Spergularia marina (L.) Griseb.; Spic. Fl. Rumel. 1: 213 (1843)

= Spergularia marina f. longicaulis (Pomel) Batt.; Fl. Algerie [Batt. & Trab.] Dicot.: 161 (1888)

= Spergularia marina subsp. neglecta (Kindb.) Hook.f.; 63 (1870)

= Spergularia marina var. asiatica (H.Hara) H.Hara; J. Jap. Bot. 17: 24 (1941)

= Spergularia marina var. cleistogama Y.X.Ma & D.L.Cui; Pl. Diversity Resources 34: 155 (2012)

= Spergularia marina var. densiflora Hosny ex Hosni; Taeckholmia 11: 48 (1988)

= Spergularia marina var. heterosperma (Fenzl) Neilr.; Fl. Nied.-Oesterr. 782 (1859)

= Spergularia marina var. leiosperma (Kindb.) Gürke

= Spergularia marina var. melanocaulos (Merino) Merinio; Contr. Fl. Galicia 117 (1897)

= Spergularia marina var. neglecta (Kindb.) Hook.f.; 63 (1870)

= Spergularia marina var. salina (J.Presl & C.Presl) Hook.f.; 63 (1870)

= Spergularia marina var. simonii O.Deg. & I.Deg.; Fl. Hawaiiensis 119: s. p. (1962)

= Spergularia marina var. stenopetala A.Chev.; Bull. Herb. Boiss. , ser. 2, 5: 440 (1905)

= Spergularia marina var. tenuis (Greene) R.Rossbach; Rhodora 42: 138 (1940)

= Spergularia marina var. urbica (Leffler) Cout.; Fl. Portugal 205 (1913)

= Spergularia media var. heterosperma Gren. & Godr.; Fl. France Corse 1: 276 (1848)

= Spergularia miquelonensis Lebel; Bull. Soc. Bot. France 15: 58 (1868), nom. illeg.

= Spergularia modesta proles marina (L.) Samp.; Bol. Soc. Brot. 24: 27 (1909)

= Spergularia modesta proles urbica (Leffler) Samp.; Bol. Soc. Brot. 24: 27 (1909), nom. illeg.

= Spergularia neglecta (Kindb.) Syme; J. E. Sm., Engl. Bot. ed. 3, 2: 129 (1864) non Weihe

= Spergularia neglecta var. salina (J.Presl & C.Presl) Syme; 130 (1864)

= Spergularia pinguis (Fenzl) Tzvelev; Novosti Sist. Vyssh. Rast. 32: 29 (2000), nom. illeg.

= Spergularia pinguis Rouy & Foucaud; Fl. France 3: 307 (1896)

= Spergularia rodrigoi Sennen & Mauricio; Diagn. Nouv. Pl. Esp. Maroc 183 (1936)

= Spergularia rubra subsp. salina (J.Presl & C.Presl) Celak.; Oesterr. Bot. Z. 20: 48 (1870)

= Spergularia rubra var. dillenii (Lebel) Knoche; Fl. Balear. 1: 458 (1921)

= Spergularia rubra var. marina (L.) A.Gray; Manual 64 (1848)

= Spergularia rubra var. marina (L.) Fiori & Paol.; A. Fiori et al., Fl. Anal. Italia 1: 139 (1898)

= Spergularia rubra var. pinguis Fenzl; Ledeb., Fl. Ross. 2: 167 (1843)

= Spergularia rubra var. sperguloides (Lehm.) Ball; Spicil. Fl. Marocc. [Ball]: 369 (1877)

= Spergularia salina f. glabriuscula Zapal.; 183 (1911)

= Spergularia salina J.Presl & C.Presl; Fl. Cech.: 95 (1819)

= Spergularia salina var. asiatica H.Hara; J. Jap. Bot. 13: 171 (1937)

= Spergularia salina var. australis (Lebel) Gürke; K. Richter, Pl. Eur. 2: 197 (1899)

= Spergularia salina var. drohobycensis Zapal.; 183 (1911)

= Spergularia salina var. involucrata (B.L.Rob.) Jeps.; Fl. W. Calif. ed. 2: 156 (1911)

= Spergularia salina var. leiosperma (Kindb.) Lindm.; 246 (1918)

= Spergularia salina var. minor (S.Watson) B.L.Rob.; Proc. Amer. Acad. Arts 29: 312 (1894)

= Spergularia salina var. minor Döll; 621 (1843)

= Spergularia salina var. sordida (Greene) Jeps.; Fl. W. Calif., ed. 2: 156 (1911)

= Spergularia salina var. tenuior Rupr.; Fl. Ingrica: 193 (1860)

= Spergularia salina var. tenuis (Greene) Jeps.; Fl. W. Calif., ed. 2: 156 (1911)

= Spergularia salina var. urbica (Leffler) Gürke; K. Richter, Pl. Eur. 2: 196 (1899)

= Spergularia seminulifera Hy; Revis. Gen. Pl. Bot. 25: 316 (1913)

= Spergularia sparsiflora (Greene) A.Nelson; Coult. & Nels., Man. Bot. Rocky Mt., ed. 2: 187 (1909)

= Spergularia sperguloides (Lehm.) Heynh.; Nom. Bot. Hört.: 690 (1841)

= Spergularia tenuis (Greene) B.L.Rob.; Proc. Amer. Acad. Arts 29: 311 (1894)

= Spergularia tenuis var. involucrata B.L.Rob.; A. Gray, Syn. Fl. N. Amer. 1(1): 251 (1897)

= Spergularia uliginosa Pomel; Nouv. Mat. Fl. Atl. 204 (1875)

= Spergularia urbica Nyman; Consp. Fl. Eur. 122 (1878)

= Stipularia marina (L.) Haw.; Syn. Pl. Succ. 104 (1812)

= Tissa coreana (H.Lév.) Nakai; Bot. Mag., Tokyo 48: 775 (1934)

= Tissa fauriei Nakai; Bot. Mag., Tokyo 27: 129 (1913), nomen

= Tissa leiosperma (Kindb.) A.Heller; Muhlenbergia 6: 84 (1910)

= Tissa marina (L.) Britton; Bull. Torrey Bot. Club 16: 126 (1889)

= Tissa marina var. minor (S.Watson) Britton; Man. Fl. N. States 403 (1901)

= Tissa salina (J.Presl & C.Presl) A.Heller; Muhlenbergia 6: 96 (1910), isonym

= Tissa salina (J.Presl & C.Presl) Britton; Bull. Torrey Bot. Club 16: 127 (1889)

= Tissa salina var. involucrata (B.L.Rob.) Jeps.; Fl. W. Calif. 170 (1901)

= Tissa salina var. sanfordii Greene; Fl. Francisc. 129 (1891)

= Tissa salina var. sordida Greene; Fl. Francisc. 129 (1891)

= Tissa salina var. tenuis (Greene) Jeps.; Fl. W. Calif. 170 (1901)

= Tissa sparsiflora Greene; Erythea 3: 46 (1895)

= Tissa tenuis (Greene) Greene; Bull. Torrey Bot. Club 16: 128 (1889)

species :

Spergularia masafuerana Skottsb.; Nat. Hist. Juan Fernandez & Easter Island 2: 778 (1953)

Distribution: Juan Fdz. Isl.

85 JNF

species :

Spergularia media (L.) C.Presl; Fl. Sicul. 161 (1826) nomen, et ex Griseb. Spicil. Fl. Rumel. 1: 213

subspecies :

subsp. media

Name: Flügelsamige Schuppenmiere (DE); Greater Sea-spurrey (EN)

Distribution: England; Ireland; Denmark; Norway; Sweden; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Gibraltar; Baleares; France; Corsica; Sardinia; Italy; Sicily; Malta; Slovenia; Croatia; Montenegro; Serbia; North Macedonia; Romania; Bulgaria; European Turkey; Greece (rare coastal, Kiklades, Thasos, Samothraki, Limnos, Kithira); Crete; C-European Russia; E-European Russia; Moldova; Ukraine; Crimea; Morocco; Algeria; Egypt (Desert Oases, Great Southwestern Desert, NW-coastal Egypt); Azores (Pico); Madeira (Madeira Isl.); Canary Isl. (Gomera, Tenerife, Gran Canaria, Lanzarote, Fuerteventura); Angola [I]; Namibia [I]; South Africa [I] (widespread); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; East Aegaean Isl. (Lesvos, Samos, Rhodos); Turkey (Inner Anatolia, N-Anatolia, NE-Anatolia, W-Anatolia, WN-Anatolia); Iran (EC-Iran, E-Iran, N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Baghlan, Faryab, Kabul, Kandahar, Kunar / Nuristan, Laghman, Nangarhar, Samangan, Takhar); Afghanistan (Baghlan, Helmand, Herat, Kabul, Logar, Qunduz); Lebanon (coastal W-Lebanon); Syria (C-Syrian Desert); Israel (coastal W-Israel, Rift Valley, N-Israel, Negev Desert); Sinai peninsula (N-Sinai); Jordania (S-Jordania); China (Nei Mongol, Xinjiang); Mongolia; ?Ryukyu Isl.; peninsular Malaysia [I] (Perak [I], Pahang [I]); Australia [I] (South Australia [I], Victoria [I], Tasmania [I]); Lord Howe Isl. [I]; Norfolk Isl. [I]; New Zealand [I]; Canada [I] (Ontario [I], Quebec [I]); USA [I] (California [I], Colorado [I], Illinois [I], Indiana [I], Massachusetts [I], Michigan [I], Montana [I], New Mexico [I], New York [I], North Dakota [I], Ohio [I], Oregon [I], Pennsylvania [I], Utah [I], Wyoming [I]); Mexico [I]; Chile [I] (Antofagasta [I], Coquimbo [I], Valparaíso [I], Metropolitana de Santiago [I], O´Higgins [I], Maule [I], Ñuble [I], Biobío [I], Araucanía [I], Los Rios [I], Los Lagos [I], Aisén [I], Magallanes [I]); Desventuradas Isl. [I]; Macquarie Isl. [I]; Gough Isl. [I]; Tristan da Cunha [I]

10 DEN GRB IRE NOR SWE 11 AUT CZE HUN POL SLO 12 BAL COR FRA GIB POR SAR SPA 13 BUL CRO GRC ITA KRI MLT MNT NMA ROM SIC SLV SRB TUE 14 KRY MLD RUS UKR 20 ALG EGY MOR 21 AZO CNY MDR 26 ang 27 nam saf 32 KAZ KGZ TKM TZK UZB 33 ARM AZE GEO NCS 34 AFG EAI IRN ISR JOR LEB SIN SYR TUR 36 CHI 37 MON 42 mly 50 aus lhi nfk 51 nzl 72 cdn 73 usa 74 usa 75 usa 76 usa 77 usa 79 mex 85 cle dsv 91 maq tdc

Synonyms: hidden

= Alsine marginata (DC.) Fenzl; Verbreit. Alsin. tab. ad p. 18 (1833)

= Alsine marginata (DC.) Rchb.; Fl. Germ. Excurs. 2: 566 (1832), nom. illeg.

= Alsine maritima (All.) Samp.; Herb. Port., Ap. 2: 6 (1914)

= Alsine media (L.) Druce; Ann. Scott. Nat. Hist. 1906: 221 (1906), nom. illeg.

= Alsine rubra var. media (L.) Wahlenb.; Fl. Suecica 1: 281 (1824)

= Alsinella marina Hartm.; Handb. Skand. Fl. 175 (1820)

= Alsinella media (L.) Druce; Ann. Scott. Nat. Hist. 1906: 221 (1906), nom. illeg.

= Alsinella media (L.) Hornem.; Nom. 33 (1827)

= Arenaria litoralis Phil.; Linnaea 28: 673 (1856), nom. illeg.

= Arenaria macrorrhiza Guirão ex Willk. & Lange; Prodr. Fl. Hispan. 3: 166 (1874)

= Arenaria marginata DC.; Fl. Franc. [DC. & Lam.], ed. 3, 5: 793 (1805), nom. illeg.

= Arenaria marina All.; Fl. Pedem. 2: 114 (1785)

= Arenaria marina var. media (L.) Fr.; Novit. Fl. Svec. 49 (1817)

= Arenaria maritima All.; Auct. Syn. Meth. Stirp. Hort. Regii Taur.: 35 (1773)

= Arenaria media Hook.f.; Handb. N. Zeal. Fl. 1: 25 (1864), nom. illeg.

= Arenaria media L.; Sp. Pl. (ed. 2) 606 (1762)

= Arenaria pubescens Steud.; Nom. ed. I. 66 (1821)

= Arenaria spergula Dufour; Ann. Gen. Sci. Phys. 7: 293 (1820)

= Buda media (L.) Desv.; J. Bot. Agric. 3: 226 (1816)

= Buda media (L.) Dumort.; Fl. Belg. 110 (1827), isonym

= Cerastium medium (L.) Crantz; Inst. 2: 401 (1766)

= Corion halophilum var. marginatum (C.A.Mey.) Merino; Fl. Galicia 1: 515 (1905)

= Corion medium (L.) N.E.Br.; Bot. Exch. Club Brit. Isles, Report for 1892 359 (1893)

= Lepigonum australe F.Muell. ex Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) in Ind. (1863)

= Lepigonum canadense Fisch. & C.A.Mey.; Index Seminum (LE, Petropolitanus) 3: 14 (1837)

= Lepigonum capense Fisch. & C.A.Mey.; Index Seminum (LE, Petropolitanus) 1: 10 (1835)

= Lepigonum dubium Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 18. t. 1, f. 6 (1863)

= Lepigonum halophilum Fisch. & C.A.Mey. ex Avé-Lall.; Index Seminum (LE, Petropolitanus) 10: 22 (1845)

= Lepigonum heterospermum Guss.; Enum. Pl. Inarim.: 51 (1855)

= Lepigonum heterospermum Schur; Enum. Pl. Transsilv.: 126 (1866), nom. illeg.

= Lepigonum heterospermum var. capense (Haw.) Kindb.; Monogr. Gen. Lepigon.: 29 (1863)

= Lepigonum marginatum (DC.) W.D.J.Koch; Syn. Fl. Germ. ed. II. 121 (1843)

= Lepigonum marginatum var. glabra L.

= Lepigonum marginatum var. glandulosa Fenzl

= Lepigonum marinum var. apterum E.S.Marshall; J. Bot. 39: 267 (1901)

= Lepigonum medium (L.) Wahlenb.; Fl. Gothob. 45 (1820)

= Lepigonum medium var. grandiflorum Regel; Index Seminum (LE, Petropolitanus) 1856: 33 (1856)

= Lepigonum medium Wahlenb.; Fl. Gothob. 1: 45 (1820)

= Lepigonum pallidum (Salisb.) Britten; J. Bot. 54: 61 (1916)

= Melargyra media (L.) Raf.; Fl. Tellur. 3: 81 (1836)

= Spergula filipensis D.Dietr.; Syn. Plant. 2: 1598 (1840)

= Spergula marginata (DC.) Murb.; Contr. Fl. Tunisie [Murbeck] 1, Acta Univ. Lund 33(12): 45 (1897)

= Spergula marginata var. glandulosissima Faure & Maire; Bull. Soc. Hist. Nat. Afrique N. 27: 214 (1936)

= Spergula marginata var. kralikii Kralik ex Maire; Bull. Soc. Hist. Nat. Afrique N. 30: 268 (1939)

= Spergula maritima (All.) Pedersen; Fl. Patagonica (Collecc. Cient. 8), 4a: 267 (1984), isonym

= Spergula maritima Hill; Brit. Herb.: 185 (1756), opus utique oppr.

= Spergula media (L.) Bartl. & H.L.Wendl.; Beitr. Bot. 2: 64 (1825)

= Spergula pallida Salisb.; Prodr. Stirp. Chap. Allerton: 298 (1796)

= Spergula rubra var. media (L.) Fiori; Fiori et al., Fl. Anal. Italia 1: 339 (1898), nom. illeg.

= Spergularia branlyi Sennen; Bol. Soc. Ibér. Ci. Nat. 25: 217 (1926 publ. 1927)

= Spergularia capensis Heynh.; Alph. Aufz. Gew.: 689 (1847)

= Spergularia halophila var. marginata (DC.) Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 165 (1904)

= Spergularia halophila var. media (L.) T.Marsson; Fl. Neu-Vorpommern.: 77 (1869), nom. illeg.

= Spergularia marginata (DC.) Kitt.; Taschenb. ed. II. 1003 (1844)

= Spergularia marginata subsp. attalii Sennen; Diagn. Nouv. 1928-1935: 93 (1936)

= Spergularia marginata var. attalii Sennen; Diagn. Nouv.: 93 (1936)

= Spergularia marginata var. crassa Hy ex Pit.; Expl. Sci. Maroc [Pit.] 1: 17 (1913)

= Spergularia marginata var. grandiflora Pit.; Iles Canaries [Pit. & Proust]: 123 (1909)

= Spergularia marginata var. illerdensis Sennen; Diagn. Nouv.: 138 (1936)

= Spergularia marginata var. monticola Debeaux & É.Rev. ex Foucaud; Bull. Acad. Int. Géogr. Bot. 15(185-186): 65 (1905)

= Spergularia marginata var. pedicellaris Pit.; Iles Canaries [Pit. & Proust]: 123 (1909)

= Spergularia maritima (All.) Chiov.; Ann. Bot. (Roma) 10: 22 (1912)

= Spergularia maritima (All.) Druce; Rep. Bot. Exch. Cl. Brit. Isles 3: 438 (1913), isonym

= Spergularia media f. diemenica Gand.; Bull. Soc. Bot. France 47: 306 (1901)

= Spergularia media proles nobreana (Samp.) Samp.; Bol. Soc. Brot. 24: 28 (1909)

= Spergularia media subsp. marginata (DC.) Arcang.; Comp. Fl. Ital.: 107 (1882)

= Spergularia media var. aptera (E.S.Marshall) P.D.Sell; Fl. Gr. Brit. Ireland 1: 686 (2018)

= Spergularia media var. heterocarpa Merino; Contr. Fl. Galicia, Supl. 1: 29 (1898)

= Spergularia media var. marginata (DC.) Gren. & Godr.; Fl. France Corse 1: 276 (1848)

= Spergularia media var. nobreana (Samp.) Samp.; Man. Fl. Port.: 285 (1911)

= Spergularia nicaeensis var. hispanica Pau; Anales Soc. Esp. Hist. Nat. 1897: 122 (1897), nom. illeg.

= Spergularia pallida G.Don; Gen. Hist. 1: 425 (1831)

= Spergularia racemosa G.Don; Gen. Hist. 1: 426 (1831)

= Spergularia rubra subsp. marginata (DC.) Celak.; Oesterr. Bot. Z. 20: 48 (1870)

= Spergularia rubra subsp. media (L.) Bonnier & Layens; Tabl. Syn. Pl. Vasc. France 53 (1894)

= Spergularia rubra var. marginata (DC.) Knoche; Fl. Baléar. 1: 458 (1921)

= Spergularia rubra var. media (L.) Fiori & Paol.; Fl. Anal. Italia 1: 139 (1898)

= Spergularia salina subsp. marginata Celak.; Prodr. Fl. Böhmen: 491 (1875)

= Spergularia spergula G.Don; Gen. Syst. 1: 426 (1831)

= Stipularia media (L.) Haw.; Syn. Pl. Succ. 103 (1812)

= Stipularia pubescens Haw.; Syn. Pl. Succ. 103 (1812)

= Tissa marginata (DC.) A.Heller; Muhlenbergia 6: 96 (1910)

= Tissa media (L.) Reiche; Anales Univ. Chile 91: 349 (1895)

subspecies :

subsp. angustata (Clavaud) Kerguélen & Lambinon; Nouv. Fl. Belg., Luxemb., N. France, ed. 4: 988 (1992)

Name: Atlantische Flügelsamige Schuppenmiere (DE)

Distribution: Netherlands; Belgium; Germany (Baden-Württemberg [I], Hessen [I], Mecklenburg-Vorpommern, Niedersachsen, +Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Sachsen-Anhalt, Thüringen); Portugal; Spain; France; Estonia (Saaremaa Isl.); ?Morocco

11 BGM GER NET 12 FRA POR SPA 14 EST

Synonyms: hidden

= Alsine canadensis var. angustata (Clavaud) Samp.; Lista Esp. Herb. Port. 79 (1913)

= Lepigonum caninum var. urbicum Leffler; Oesterr. Bot. Z. 19: 103 (1869), nom. illeg.

= Lepigonum fasciculare E.J.Lönnr.; Obs. Crit. Pl. Suec.: 13 (1854)

= Lepigonum marinum var. fasciculare (Lönnr.) Leffl.; 104 (1869), nom. illeg.

= Lepigonum rubrum var. glabrum Wahlberg; 46 (1820)

= Spergula fascicularis (Lönnr.) Johanss.; Bot. Notiser 1910: 235 (1910)

= Spergula fascicularis var. glabra Johanss.; Bot. Notiser 1910: 225 (1910)

= Spergula marginata subsp. fascicularis (Lönnr.) Ahlfv.; Neuman, 555 (1901)

= Spergularia angustata (Clavaud) Tzvelev; Novosti Sist. Vyssh. Rast. 32: 27 (2000)

= Spergularia fascicularis (Lönnr.) Nyman; Consp. 122 (1878)

= Spergularia fascicularis Nyman; Consp. Fl. Eur.: 122 (1878)

= Spergularia halophila f. macrantha T.Marsson; 77 (1869)

= Spergularia halophila f. micrantha T.Marsson; 77 (1869)

= Spergularia marginata subsp. angustata (Clavaud) P.Monnier; Candollea 30: 152 (1975)

= Spergularia marginata var. angustata Clavaud; Actes Soc. Linn. Bordeaux 35: 401 (1882)

= Spergularia maritima subsp. angustata (Clavaud) Greuter & Burdet; Willdenowia 12(2): 190 (1982)

= Spergularia salina f. urbica (Leffl.) Lindm.; 246 (1918)

= Spergularia urbica (Leffler) Nyman; Consp. Fl. Eur.: 122 (1878)

subspecies :

subsp. intermedia (Maire) Lambinon & Dobignard; J. Bot. Soc. Bot. France 20: 42 (2002)

Distribution: Morocco; Western Sahara; W-Mauritania

20 MOR WSA 22 MTN

Synonyms: hidden

= Spergula marginata var. intermedia Maire; Bull. Soc. Hist. Nat. Afrique N. 26: 191 (1935)

= Spergularia marginata var. intermedia (Maire) P.Monnier; Candollea 30: 152 (1975)

= Spergularia maritima subsp. intermedia (Maire) Greuter & Burdet; Willdenowia 12: 190 (1982)

subspecies :

subsp. occidentalis (P.Monnier) Lambinon & Dobignard; J. Bot. Soc. Bot. France 20: 42 (2002)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Spergularia maritima subsp. occidentalis P.Monnier; Willdenowia 13: 282 (1984)

subspecies :

subsp. sauvagei (P.Monnier) Lambinon & Dobignard; J. Bot. Soc. Bot. France 20: 43 (2002)

Distribution: Morocco; Western Sahara; Algeria; Tunisia; ?Libya

20 ALG MOR TUN WSA

Synonyms: hidden

= Spergula battandieri E.A.Durand & Barratte; Fl. Libyc. Prodr.: 42 (1910)

= Spergula gamostyla (Pomel) Maire; Fl. Afrique N. 9: 104 (1963)

= Spergula marginata var. battandieri (E.A.Durand & Barratte) Maire; Bull. Soc. Hist. Nat. Afrique N. 24: 205 (1933)

= Spergula marginata var. reverchonii (Foucaud ex Batt.) Maire; Jahand. et al., Cat. Pl. Maroc 4: 982 (1932)

= Spergula marginata var. robertii (Foucaud ex Batt.) Maire; Jahand. et al., Cat. Pl. Maroc 4: 983 (1932)

= Spergularia battandieri (E.A.Durand & Barratte) Foucaud ex Pamp.; Pl. Tripol.: 84 (1914)

= Spergularia gamostyla Pomel; Nouv. Mat. Fl. Atlant. 1: 202 (1874)

= Spergularia marginata subsp. chevallieri (Hy) Pott.-Alap.; Fl. Tunisie 1: 103 (1979) (nom. inval.)

= Spergularia marginata subsp. sauvagei P.Monnier; Candollea 30: 152 (1975) (nom. inval.)

= Spergularia maritima subsp. sauvagei P.Monnier; Willdenowia 13: 282 (1983 publ. 1984)

= Spergularia media f. heterosperma Batt.; Fl. Algerie [Batt. & Trab.] Dicot.: 161 (1888)

= Spergula media subsp. sauvagei (P.Monnier) Dobignard; Evaxiana 2: 172 (2016), isonym

= Spergularia media var. battandieri (E.A.Durand & Barratte) Batt.; Fl. Algerie [Batt. & Trab.] Suppl. Phan.: 29 (1910)

= Spergularia media var. reverchonii Foucaud ex Batt.; Fl. Algèrie [Batt. & Trab.], Suppl. Phan.: 29 (1910)

= Spergularia media var. robertii Foucaud ex Batt.; Fl. Algerie (Batt.), Suppl. Phan.: 29 (1910)

subspecies :

subsp. tunetana (Maire) Lambinon & Dobignard; J. Bot. Soc. Bot. France 20: 43 (2002)

Distribution: Sicily; Algeria; Tunisia; Libya

13 SIC 20 ALG LBY TUN

Synonyms: hidden

= Spergula tunetana Maire; Bull. Soc. Hist. Nat. Afrique N. 32: 205 (1941)

= Spergula tunetana var. chevallieri (Hy) Maire & Weiller; [nom. inval.]

= Spergula tunetana var. mixta Maire; Bull. Soc. Hist. Nat. Afrique N. 32: 205 (1941)

= Spergula tunetana var. pseudopycnorrhiza Maire; Bull. Soc. Hist. Nat. Afrique N. 32: 205 (1941)

= Spergularia chevallieri Hy; Revis. Gen. Pl. Bot. 25: 312 (1913), nomen

= Spergularia marginata subsp. tunetana (Maire) Monnier; Candollea 30: 152 (1975)

= Spergularia maritima subsp. tunetana (Maire) Greuter & Burdet; Willdenowia 12(2): 191 (1982)

= Spergularia tunetana (Maire) Jalas; Ann. Bot. Fenn. 20: 110 (1983)

species :

Spergularia melanocaulos Merino; Aigunas Pl. Rar. de La Guardia (Pontevedra): 26 (1895)

Distribution: W-Portugal; W-Spain

12 POR SPA

Synonyms: hidden

= Alsine purpurea var. crassipes (Samp.) Samp.; Lista Esp. Herb. Port.: 79 (1913)

= Alsine rupicola var. australis (Samp.) Samp.; Lista Esp. Herb. Port.: 79 (1913)

= Corion rupicola var. australe (Samp.) Merino; Fl. Galicia 1: 516 (1905)

= Spergula australis (Samp.) G.López; Lagascalia 30: 15 (2010)

= Spergularia australis (Samp.) P.Monnier & Ratter; Tutin & al., Fl. Eur. 1: 155 (1964)

= Spergularia australis (Samp.) Ratter; Bot. J. Linn. Soc., 109(3): 322 (1992), isonym

= Spergularia colorata proles australis (Samp.) Samp.; Ann. Sci. Acad. Polytecn. Porto 5: 51 (1909), nom. illeg.

= Spergularia colorata proles crassipes (Samp.) Samp.; Bol. Soc. Brot. 24: 27 (1909), nom. illeg.

= Spergularia colorata var. australis (Samp.) Samp.; Man. Fl. Port.: 284 (1911)

= Spergularia colorata var. crassipes (Samp.) Samp.; Man. Fl. Port.: 284 (1911), nom. illeg.

= Spergularia purpurea var. crassipes Samp.; Ann. Sci. Nat. (Oporto) 10: 23 (1906)

= Spergularia rupicola proles australis (Samp.) Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 161 (1904)

= Spergularia rupicola subsp. australis (Samp.) Malag.; Sin. Fl. Iber.: 285 (1975)

= Spergularia rupicola var. australis Samp.; Ann. Sci. Nat. (Oporto) 6: 71 (1899)

= Spergularia rupicola var. crassipes (Samp.) Cout.; Fl. Portugal: 204 (1913)

species :

Spergularia mexicana Hemsl.; Diagn. Pl. Nov. 22 (1879)

Distribution: Mexico (Aguascalientes, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacan, Queretaro, San Luis Potosi, Tlaxcala, Veracruz, Zacatecas)

79 MEX

Synonyms: hidden

= Alsine mexicana (Hemsl.) House; Am. Midl. Nat. 7: 134 (1921)

= Tissa mexicana (Hemsl.) Britton; Bull. Torrey Bot. Club 16: 129 (1889)

species :

Spergularia microsperma (Kindb.) Vved.; Fl. Uzbekistan. 2: 373 (1953)

subspecies :

subsp. microsperma

Distribution: Kazakhstan; Uzbekistan; Tajikistan

32 KAZ TZK UZB

Synonyms: hidden

= Lepigonum microspermum Kindb.; Monogr. Gen. Lepigon.: 26 (1863)

= Spergula microsperma (Kindb.) Murb.; Acta Univ. Lund. 32(12): 44 (1896)

= Spergularia diandra var. microspermoides (Vved.) Täckh. & Boulos; Publ. Cairo Univ. Herb. 5: 15 (1972 publ. 1974)

= Spergularia glaucophylla Vved.; Bot. Mater. Gerb. Bot. Inst. Uzbekistansk. Fil. Akad. Nauk SSSR 3: 5 (1941)

= Spergularia microspermoides Vved.; Bot. Mater. Gerb. Bot. Inst. Uzbekistansk. Fil. Akad. Nauk SSSR 3: 5 (1941)

subspecies :

subsp. fontenellei (Maire) Greuter & Burdet; Willdenowia 12: 191 (1982)

Distribution: S-Algeria (Hoggar); N-Chad (Tibesti)

20 ALG 24 CHA

Synonyms: hidden

= Spergula fontenellei Maire; Bull. Soc. Hist. Nat. Afrique N. 20: 174 (1929)

= Spergularia fontenellei (Maire) Ozenda; Fl. Sahara: 213 (1958) (comb. inval.)

= Spergularia tibestica Quézel & Monnier; Bull. Soc. Hist. Nat. Afr. Nord 49: 284 (1959)

subspecies :

subsp. oreophila (Litard. & Maire) P.Monnier; Naturalia Monspel., Ser. Bot. 19: 105 (1968)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Spergula rubra subsp. oreophila (Litard. & Maire) Maire; Jahand. & Maire, Cat. Pl. Maroc: 209 (1932)

= Spergularia rubra subsp. oreophila Litard. & Maire; Mém. Soc. Sci. Nat. Maroc 4: 9 (1924)

species :

Spergularia munbyana Pomel; Bull. Soc. Sci. Phys. Algérie 11: 203 (1874) ex E. A. Durand & Barratte, Fl. Libya. Prodr. 42 (1910)

Distribution: ?Morocco; Algeria; Tunisia; Libya

20 ALG LBY TUN

Synonyms: hidden

= Spergula marginata var. munbyana (Pomel) Maire; Cat. Pl. Maroc 2: 210 (1932)

= Spergula munbyana (Pomel) E.A.Durand & Barratte; Fl. Libyc. Prodr.: 42 (1910)

= Spergularia marginata var. munbyana (Pomel) Batt.; Fl. Algerie [Batt. & Trab.] Suppl. Phan.: 30 (1910)

= Spergularia media var. munbyana (Pomel) Batt.; Fl. Algerie [Batt. & Trab.], Suppl. Phan.: 30 (1910)

species :

Spergularia namaquensis Schltr. ex M.Á.Alonso, M.B.Crespo, Mart.-Azorín & Mucina; Pl. Syst. Evol. 307(Art. 24): 24, figs. 1-6 (2021)

Distribution: Namibia

27 NAM

species :

Spergularia nesophila L.G.Adams; Austral. Syst. Bot. 21(4): 269 (2008)

Distribution: Australia (Western Australia)

50 AUS

species :

Spergularia nicaeensis Burnat; Fl. Alpes Marit. 1: 269 (1892)

Distribution: Spain; France; Corsica; Sardinia; Italy; ?Algeria

12 COR FRA SAR SPA 13 ITA

Synonyms: hidden

= Spergula nicaeensis (Sarato ex Burnat) G.López; Lagascalia 30: 17 (2010)

= Spergula rubra subsp. nicaeensis (Burnat) Maire; Jahand. & Maire, Cat. Pl. Maroc: 210 (1932)

= Spergularia rubra subsp. nicaeensis (Sarato ex Burnat) Briq.; Prodr. Fl. Corse 1: 490 (1910)

species :

Spergularia pazensis (Rusby) Rossbach; Rhodora 42: 205 (1940)

Distribution: Bolivia (Cochabamba, La Paz, Santa Cruz, Tarija)

83 BOL

Synonyms: hidden

= Tissa pazensis Rusby; Bull. N. Y. Bot. Gard. 6: 503 (1910)

species :

Spergularia pissisi (Phil.) I.M.Johnst.; Rev. Chil. Hist. Nat. 33: 26 (1929)

Distribution: Argentina (San Juan); Chile (Atacama, Coquimbo)

85 ARG CLE

Synonyms: hidden

= Arenaria pissisi Phil.; Linnaea 33: 20 (1864 publ. 1865)

= Spergula pissisi (Phil.) Volponi; R. Kiesling, Fl. San Juan 1: 176 (1994)

= Tissa pissisi (Phil.) Reiche; Fl. Chile 1: 200 (1895)

species :

Spergularia pitardiana Hy ex Pit.; Explor. Sci. Maroc, Bot. 17 (1912)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Spergula pitardiana (Hy ex Pit.) Maire; Bull. Soc. Hist. Nat. Afr. Nord 20: 17 (1929)

= Spergula pitardiana var. villosa (Emb. & Maire) Maire; Mem. Soc. Sci. Nat. Maroc 21-22: 27 (1930)

= Spergularia pitardiana var. villosa Emb. & Maire; Mem. Soc. Sci. Nat. Maroc 22: 27 (1929)

species :

Spergularia purpurea (Pers.) G.Don; Gen. Hist. 1: 425 (1831)

Name: Purple Sandspurrey

Distribution: Portugal; Spain; ?Italy; Morocco; USA [I] (Massachusetts [I])

12 POR SPA 20 MOR 75 usa

Synonyms: hidden

= Alsine purpurea (Pers.) Heynh.; Nom. Bot. Hort.: 38 (1840)

= Alsine purpurea var. indurata (Samp.) Samp.; Lista Esp. Herb. Port.: 79 (1913)

= Alsine purpurea var. longipes (Lange) Samp.; Lista Esp. Herb. Port.: 79 (1913)

= Arenaria purpurea Pers.; Syn. Pl. 1: 504 (1805)

= Corion purpureum (Pers.) Pau; Bol. Soc. Arag. 4: 311 (1905)

= Corion purpureum var. longipes (Lange) Merino; Fl. Galicia 1: 517 (1905)

= Lepigonum firmum Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 33 (1863)

= Lepigonum longipes Murb.; Lunds Univ. Arsskrift, n. s. 18: No. 3, 33 (1922)

= Lepigonum purpureum Kindb.; Monogr. Gen. Lepigon.: 33 (1863)

= Lepigonum rubrum var. longipes Lange; Pug. Pl. Hispan. 4: 101 (1865)

= Melargyra purpurea Raf.; Fl. Tellur. 3: 81 (1837)

= Spergula longipes (Lange) Murb.; Contr. Fl. Maroc [Murbeck]1, Acta Univ. Lund 18(3): 33 (1922)

= Spergula purpurea (Pers.) D.Dietr.; Syn. Pl. 2: 1598 (1840)

= Spergula rubra subsp. longipes (Lange) Maire; Jahand. & Maire, Cat. Pl. Maroc: 210 (1932)

= Spergula rubra var. rouyana Cout.; Fl. Portugal: 205 (1912)

= Spergularia colorata proles indurata (Samp.) Samp.; Bol. Soc. Brot. 24: 26 (1909)

= Spergularia colorata proles purpurea (Pers.) Samp.; Bol. Soc. Brot. 24: 26 (1909), nom. illeg.

= Spergularia colorata Samp.; Bol. Soc. Brot. 24: 26 (1908)

= Spergularia colorata var. indurata (Samp.) Samp.; Man. Fl. Port.: 284 (1911)

= Spergularia colorata var. longipes (Lange) Samp.; Bol. Soc. Brot. 24: 26 (1909)

= Spergularia langei Foucaud; Note sur le Spergularia rubra var. pinguis Fenzl: 5 (1903)

= Spergularia longipes (Lange) Rouy; Bull. Herb. Boiss. 3: 224 (1895)

= Spergularia longipes var. langeana Cout.; Fl. Portugal: 205 (1913), nom. illeg.

= Spergularia longipes var. rouyana Cout.; Fl. Portugal: 205 (1913)

= Spergularia purpurea proles indurata Samp.; Ann. Sci. Nat. (Oporto) 10: 22 (1906)

= Spergularia purpurea var. indurata Samp.; Ann. Sci. Nat. (Oporto) 10: 22 (1906)

= Spergularia purpurea var. longipes (Lange) Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 155 (1904)

= Spergularia rubra subsp. longipes (Lange) Briq.; Prodr. Fl. Corse 1: 490 (1910)

species :

Spergularia pycnantha Rossbach; Rhodora 42: 170 (1940)

Distribution: Chile (Atacama, Coquimbo, Valparaíso, Metropolitana de Santiago)

85 CLE

species :

Spergularia pycnorrhiza Foucaud; ex Batt., Fl. Alger., Suppl. Phan. 29 (1910), nomen subnudum, et ex Maire, Bull. Soc. Hist. Nat. Afrique N. 26: 190 (1935), cum descr. ampl.

Distribution: Algeria

20 ALG

Synonyms: hidden

= Spergula marginata var. pycnorrhiza (Foucaud ex Batt.) Maire; Bull. Soc. Hist. Nat. Afrique N. 24: 205 (1933)

= Spergula pycnorrhiza (Foucaud ex Batt.) Maire; Bull. Soc. Hist. Nat. Afrique N. 26: 190 (1935)

= Spergularia media f. macrorhiza Batt.; Fl. Algèrie [Batt. & Trab.] Dicot.: 162 (1888)

species :

Spergularia quartzicola M.Á.Alonso, M.B.Crespo, Mart.-Azorín & Mucina; Pl. Biosystems 156(2): 507 (2021)

Distribution: South Africa (N-Cape Prov., W-Cape Prov.)

27 SAF

species :

Spergularia rubra (L.) J.Presl & C.Presl; Fl. Cech.: 94 (1819)

Name: Rote Schuppenmiere, Roter Spärkling (DE), Red Sandspurrey (EN)

Distribution: England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; Baleares; France; +Monaco; Channel Isl.; Corsica; Sardinia; Italy; Sicily; Pantelleria; Isole Pelagie (Lampedusa); Malta; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (widespread mainland, rare Kiklades, Lefkada); Crete; Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Algeria; Libya; Egypt (Great Southwestern Desert, Nile Delta, Nile Valley, NW-coastal Egypt); Sudan; Ethiopia; Eritrea; South Africa [I]; Siberia (Altai [I], Buryatia, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); Kazakhstan; Turkmenistan; Uzbekistan [I]; Kyrgyzstan [I]; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; East Aegaean Isl. (Lesvos, Chios, Ikaria, Samos, Nisyros); Turkey (Inner Anatolia, N-Anatolia, NW-Anatolia: Bithynia, SW-Anatolia, SSW-Anatolia, W-Anatolia); Iran (N-Iran); Israel (C-Israel, coastal W-Israel, Rift Valley, N-Israel); Sinai peninsula (N-Sinai, S-Sinai); Jordania (S-Jordania); Saudi Arabia (C-Saudi Arabia, Nejd Desert); China (Xinjiang); South Korea [I]; Japan (Hokkaido); Pakistan (Sind, Baluchistan, Quetta, N. W. Frontier Prov., Swat, Pakistani Punjab, Rawalpindi); India (Jammu & Kashmir, Haryana, Uttar Pradesh); Australia [I] (Western Australia [I], Northern Territory [I], South Australia [I], Queensland [I], New South Wales [I], Victoria [I], Tasmania [I]); New Zealand [I]; Chatham Isl. [I]; Alaska [I]; Canada [I] (British Columbia [I], New Brunswick [I], Newfoundland [I], Nova Scotia [I], Ontario [I], Prince Edward Isl. [I], Quebec [I], Yukon [I]); St. Pierre and Miquelon [I]; USA [I] (California [I], Colorado [I], Connecticut [I], Delaware [I], District of Columbia [I], Idaho [I], Illinois [I], Indiana [I], Kentucky [I], Maine [I], Maryland [I], Massachusetts [I], Michigan [I], Minnesota [I], Montana [I], Nevada [I], New Hampshire [I], New Jersey [I], New Mexico [I], New York [I], Ohio [I], Oregon [I], Pennsylvania [I], Rhode Island [I], Utah [I], Vermont [I], Virginia [I], Washington [I], Wisconsin [I], Wyoming [I]); Chile [I] (Coquimbo [I], Metropolitana de Santiago [I], O´Higgins [I], Maule [I], Ñuble [I], Biobío [I], Araucanía [I]); Juan Fdz. Isl. [I]

10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO BAL CIS COR FRA MCO POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS KRI MLT MNT NMA PEL PTL ROM SIC SLV SRB TUE 14 BLR EST KRY LAT LIT MLD RUS UKR 20 ALG EGY LBY 24 ERI ETH SUD 27 saf 30 SIB 31 RFE 32 KAZ kgz TKM uzb 33 ARM AZE GEO NCS 34 EAI IRN ISR JOR SIN TUR 35 SAU 36 CHI 38 JAP kor 40 IND PAK 50 aus 51 ctm nzl 70 ask cdn 71 cdn 72 cdn 73 usa 74 usa 75 usa 76 usa 77 usa 78 usa 85 cle jnf

Synonyms: hidden

= Alsine alpina Willk.; Bot. Zeitung (Berlin) 5: 238 (1847)

= Alsine radicans (C.Presl) Guss.; Fl. Sicul. Syn. 1: 502 (1843)

= Alsine rubra (L.) Crantz; Inst. Rei Herb. 2: 407 (1766)

= Alsine rubra var. glabrata Kabath; 105 (1846)

= Alsinella rubra (L.) Hartm.; Handb. Skand. Fl. 175 (1820)

= Arenaria campestris (L.) All.; Fl. Pedem. 2: 114 (1785)

= Arenaria campestris H.Mart.; 80 (1817), nom. illeg.

= Arenaria campestris var. urbana H.Mart.; 80 (1817), nom. illeg.

= Arenaria membranacea Gaterau; Descr. Pl. Montaub. 83 (1789)

= Arenaria radicans (C.Presl) Spreng.; Syst. Veg. ed. 16, 2: 400 (1825)

= Arenaria rubra L.; Sp. Pl.: 423 (1753)

= Arenaria rubra subsp. campestris (L.) Ehrh.; Hannover. Mag. 18: 222 (1780)

= Arenaria rubra var. campestris L.; Sp. Pl. 423 (1753), nom. inval.

= Arenaria rubra var. canadensis Eaton; Man. Bot., ed. 2: 143 (1818)

= Arenaria rubra var. pubera Lej. & Courtois; Comp. Fl. Belg. 2: 96 (1831), nom. superfl.

= Buda borealis S.Watson; A. Gray, Man. Bot. N. Am. Stat. ed. 6: 90 (1890)

= Buda campestris (L.) Kuntze; Revis. Gen. Pl. 3(2): 13 (1898)

= Buda rubra (L.) Desv.; J. de Bot. 3: (No. 5) 226 (1818)

= Buda rubra (L.) Dumort.; Fl. Belg. 110 (1827), isonym

= Corion radicans var. campestre (L.) Merino; Fl. Galicia 1: 518 (1905)

= Corion rubrum (L.) N.E.Br.; Syme, Engl. Bot. ed. 3, Suppl. 48 (1891)

= Fasciculus ruber (L.) Dulac; Fl. Hautes-Pyr. 245 (1867)

= Lepigonum anceps Bartl.; Lehm. Pl. Preiss. 1: 244 (1845)

= Lepigonum anceps var. glandulosum Kindb.; Symb. Syn. Gen. Lepigon.: 15 (1856), nom. nud.

= Lepigonum brevifolium Bartl.; Lehm. Pl. Preiss. 1: 243 (1845)

= Lepigonum brevifolium var. anceps (Bartl.) Kindb.; Monogr. Gen. Lepigon.: 36 (1863)

= Lepigonum brevifolium var. laxiflorum (Bartl.) Kindb.; Monogr. Gen. Lepigon.: 36 (1863)

= Lepigonum diandrum Kindb.; Symb. Syn. Gen. Lepigon.: 35 (1856), nom. illeg.

= Lepigonum largiflorum F.Muell. ex Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 29 (1863)

= Lepigonum laxiflorum Bartl.; Lehm. Pl. Preiss. 1: 244 (1845)

= Lepigonum radicans (C.Presl) Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 10 (1863)

= Lepigonum rubrum (L.) Wahlenb.; Fl. Gothob. 45 (1820)

= Lepigonum rubrum var. gramineum H.Post; Bot. Notiser 1844: 135 (1844)

= Lepigonum rubrum var. perennans Kindb.; Monogr. Gen. Lepigon. 41 (1858)

= Lepigonum rubrum var. radicans (C.Presl) Leffner; Bot. Notiser 1865: 97 (1865)

= Lepigonum sperguloides Fisch. & C.A.Mey.; Index Sem. Hort. Petrop. 1: 10 (1835)

= Melargyra rosea Raf.; Fl. Tellur. 3: 81 (1836)

= Melargyra rubra (L.) Raf.; Fl. Tellur. 3: 81 (1836)

= Spergula campestris (L.) Murb.; Lunds Univ. Arsshrift, n. s. 18: No. 3, 33 (1922)

= Spergula maxima Dufour ex Steud.; Nomencl. Bot., ed. 2, 2: 617 (1841), nom. inval.

= Spergula radicans (C.Presl) D.Dietr.; Syn. Pl. 2: 1598 (1840)

= Spergula rubra (L.) Bartl.; Linnaea 7: 625 (1832)

= Spergula rubra (L.) D.Dietr.; Syn. Pl. 2: 1598 (1840), isonym

= Spergula rubra subsp. campestris (L.) Maire; Jahand. & Maire, Cat. Pl. Maroc: 210 (1932)

= Spergula rubra subsp. radicans (J.Presl & C.Presl) O.Schwarz; Mitt. Thüring. Bot. Ges., n. s. 1: 99 (1949)

= Spergularia alpina Willk.; Flora 34: 706 (1851)

= Spergularia anceps (Bartl.) Walp.; Ann. Bot. Syst. 1: 83 (1848)

= Spergularia arenosa Foucaud ex Merino; Mém. Real Soc. Esp. Hist. Nat- 2: 496 (1904) (nom. inval.)

= Spergularia borealis (S.Watson) B.L.Rob.; Proc. Amer. Acad. Arts 39: 312 (1894)

= Spergularia brevifolia (Bartl.) Walp.; Ann. Bot. Syst. 1: 83 (1848)

= Spergularia campestris (L.) Asch.; Bot. Zeitung (Berlin) 17: 292 (1859), et Fl. Brandenburg 1: 94 (1860), nom. illeg.

= Spergularia campestris f. pubescens Hauman; Anales Soc. Ci. Argent. 89: 208 (1920), nom. nud.

= Spergularia campestris var. alpina (Boiss.) Gürke; K. Richter, Pl. Eur. 2: 194 (1899)

= Spergularia campestris var. longipes (Lange) P.Fourn.; Quatre FL. France 294 (1936)

= Spergularia campestris var. radicans (C.Presl) Gürke; K. Richter, Pl. Eur. 2: 194 (1899)

= Spergularia catalaunica P.Monnier; Guin. & R. Vilm., Fl. France: 262 (1973) (nom. inval.)

= Spergularia laxiflora (Bartl.) Walp.; Ann. Bot. Syst. 1: 83 (1848)

= Spergularia longipes Nyman; Consp. 123 (1878)

= Spergularia lycia Monnier & Quézel; Candollea 25: 359 (1970)

= Spergularia polyphylla Rohrb.; Linnaea 37: 232 (1871 publ. 1873)

= Spergularia radicans C.Presl; J. Presl & C. Presl, Delic. Prag.: 64 (1822)

= Spergularia radicans var. campestris (L.) Samp.; Ann. Sci. Acad. Polytecn. Porto 27: 157 (1904), nom. illeg.

= Spergularia rubra proles polysperma Sennen; Bol. Soc. Ibér. Ci. Nat. 25: 218 (1926)

= Spergularia rubra subsp. alpina (Willk.) Losa & Rivas Goday; Arch. Inst. Aclim. Cons. Super. Invest. Ci. 13: 133 (1974), no basionym ref.

= Spergularia rubra subsp. campestris (L.) Celak.; Prodr. Fl. Böhmen 867 (1881)

= Spergularia rubra subsp. campestris (L.) Rouy & Foucaud; Fl. France 3: 309 (1896)

= Spergularia rubra subsp. radicans (J.Presl & C.Presl) Nyman; Consp. Fl. Eur. 123 (1878)

= Spergularia rubra subsp. radicans (J.Presl & C.Presl) O.Schwarz; 99 (1949), isonym

= Spergularia rubra var. amurensis (Pomel) Batt.; Fl. Algerie [Batt. & Trab.] Dicot.: 161 (1888)

= Spergularia rubra var. campestris (L.) Fenzl; Ledeb., Fl. Ross. 2: 167 (1843)

= Spergularia rubra var. campestris (Willk.) Batt.; Fl. Algerie [Batt. & Trab.] Dicot.: 161 (1888)

= Spergularia rubra var. campestris A.Gray; Manual, ed. 5: 95 (1867), nom. illeg.

= Spergularia rubra var. cuatrecasasii Sennen; Diagn. Nouv. 280 (1936)

= Spergularia rubra var. elongata (Fenzl) Hook.f.; Fl. Tasman. 1: 41 (1855)

= Spergularia rubra var. glabrata (Kabath) Fiek; 67 (1881)

= Spergularia rubra var. longipes Lange; Willk. & Lange, Prodr. Fl. Hispan. 3: 164 (1874)

= Spergularia rubra var. parviflora Zapal.; 182 (1911)

= Spergularia rubra var. perennans B.L.Rob.; A. Gray, Syn. Fl. N. Amer. 1(1): 250 (1897)

= Spergularia rubra var. stipularis Boiss.; Fl. Orient. 1: 732 (1867)

= Spergularia rupestris Fenzl; Endl. et al., Enum. Pl. Hügel 9 (1837), nom. illeg.

= Spergularia rupestris var. elongata Fenzl; Endl. et al., Enum. Pl. Hügel 9 (1837), nom. illeg.

= Spergularia sperguloides (Fisch. & C.A.Mey.) Hausskn.; Mitth. Thür. Bot. Ver. N. Folge 5: 58 (1893)

= Stipularia rubra (L.) Haw.; Syn. Pl. Succ. 103 (1812)

= Tissa rubra (L.) Brandegee; Proc. Calif. Acad. Ser. II. 2: 131 (1889)

= Tissa rubra (L.) Britton; Bull. Torrey Bot. Club 16: 127 (1889)

= Tissa rubra var. perennans (Kindb.) Greene; Pittonia 2: 229 (1892)

species :

Spergularia rupicola Lebel; Mém. Soc. Sci. Nat. Cherbourg 7: 274 (1860)

Name: Felsen-Spärkling (DE), Cliff Sand Spurrey (EN)

Distribution: England; Ireland; Portugal; Spain; France; Channel Isl.

10 GRB IRE 12 CIS FRA POR SPA

Synonyms: hidden

= Alsine rupestris (Lebel ex Kindb.) Druce; Ann. Scott. Nat. Hist., 221 (1906)

= Alsine rupicola (Lebel) Hiern; J. Bot. 37: 318 (1899)

= Alsine rupicola var. rupicoloides (Samp.) Samp.; Fl. Portugal, ed. 2: 343 (1847)

= Buda rupestris (Lebel) F.Hanb.; Lond. Cat. Pl. ed. 9: 12 (1895)

= Corion rupestre (Lebel) N.E.Br.; Syme, Engl. Bot. ed. 3, Suppl. 49 (1891)

= Corion rupicola (Lebel) Merino; Merino, Fl. Galic. 1: 516 (1905)

= Lepigonum rupestre Lebel ex Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 29 (1863)

= Spergula lebeliana Maire; Bull. Soc. Hist. Nat. Afrique N. 26: 191 (1935), nom. superfl.

= Spergula rupicola (Lebel) G.López; Lagascalia 30: 15 (2010)

= Spergularia colorata proles rupicola (Lebel ex Le Jol.) Samp.; Man. Fl. Port.: 284 (1911)

= Spergularia colorata proles rupicoloides Samp.; Bol. Soc. Brot. 24: 26 (1909)

= Spergularia colorata var. rupicoloides (Samp.) Samp.; Man. Fl. Port. 284 (1911)

= Spergularia lebeliana Rouy; Bull. Herb. Boiss. 3: 223 (1895) (nom. illeg. superfl.)

= Spergularia macrorrhiza var. rupestris (Lebel) Tanfani; Parlatore, Fl. Ital. 9: 623 (1892)

= Spergularia marina subsp. rupestris (Lebel) Hook.f.; Student. Fl. Brit. Isl. 63 (1870)

= Spergularia rubra subsp. rupicola (Lebel) Bonnier; Fl. Ill. France 2: 64 (1913)

= Spergularia rupestris Lebel ex Willk. & Lange; Prodr. Fl. Hispan. 3: 166 (1874), nom. illeg.

= Spergularia rupicola var. guimaraesii Foucaud ex Cout.; Fl. Portugal 294 (1913)

species :

Spergularia segetalis (L.) G.Don; Gen. Hist. 1: 425 (1831)

Name: Saat-Schuppenmiere, Getreidemiere (DE)

Distribution: Netherlands; Belgium; Germany (+Brandenburg, +Baden-Württemberg, +Bayern, +Hessen, Mecklenburg-Vorpommern [I], +Niedersachsen, ++Nordrhein-Westfalen, +Rheinland-Pfalz, +Sachsen-Anhalt, +Thüringen); Switzerland; Poland; Portugal; Spain; France; Italy; E-European Russia; Morocco; Kazakhstan; Mongolia

11 BGM GER NET POL SWI 12 FRA POR SPA 13 ITA 14 RUS 20 MOR 32 KAZ 37 MON

Remarks: often included in genus Spergula. Citations for C-Asia, Mongolia and European Russia are somewhat doubtful

Conservation Status: highly threatened in the whole area and almost extinct

Synonyms: hidden

= Alsine segetalis L.; Sp. Pl.: 272 (1753)

= Alsine segetum Christm.; Pflanzensyst. 6: 223 (1780)

= Alsine unilateralis Moench; Methodus 228 (1794), nom. illeg.

= Arenaria segetalis (L.) Lam.; Fl. Franç. 3: 43 (1779)

= Arenaria unilateralis (Moench) Steud.; Nom. ed. II. 1: 127 (1840)

= Delia segetalis (L.) Dumort.; Fl. Belg.: 110 (1827)

= Lepigonum segetale (L.) Bartl.; Index Sem. Hort. Acad. Gött. 1839: 5 (1839), isonym

= Lepigonum segetale (L.) Bluff, Nees & Schauer; Comp. Fl. German., ed. alt. 1(2): 92 (1837)

= Lepigonum segetale (L.) W.D.J.Koch; Syn. Fl. Germ. ed. II. 120 (1843), isonym

= Segetella vulgaris Desv.; J. de Bot. 3: (No. 5) 225 (1816)

= Spergula segetalis (L.) Jess.; 292 (1879), isonym

= Spergula segetalis (L.) Vill.; Hist. Pl. Dauph. 3: 657 (1789)

= Spergularia exilis Fenzl; Ledeb., Fl. Ross. 2: 166 (1843)

= Spergularia semidecandra Kitt.; Taschenb. ed. II. 1003 (1844)

= Tissa segetalis (L.) Warb.; Pflanzenw. 1: 576 (1913)

species :

Spergularia sezer-zenginii Yild.; Ot Sist. Bot. Dergisi 4(2): 4 (1997 publ. 1998)

Distribution: Turkey

34 TUR

species :

Spergularia spruceana Rossbach; Rhodora 42: 180 (1940)

Distribution: Ecuador

83 ECU

species :

Spergularia stenocarpa (Phil.) I.M.Johnst.; Contrib. Gray Herb. n. s. 85: 41 (1929)

Distribution: Chile (Arica y Parinacota, Tarapacá, Antofagasta, Atacama)

85 CLE

Synonyms: hidden

= Arenaria stenocarpa Phil.; Fl. Atacam. 10 (1860)

= Spergularia borchersii Phil.; Anales Univ. Chile 81: 769 (1892)

= Spergularia larranagae Phil.; Anales Univ. Chile 81: 767 (1892)

= Tissa borchersii (Phil.) Reiche; Anales Univ. Chile 91: 347 (1895)

= Tissa larranagae (Phil.) Reiche; Anales Univ. Chile 91: 350 (1895)

= Tissa stenocarpa (Phil.) Reiche; Anales Univ. Chile 91: 349 (1895)

species :

Spergularia syvaschica Tzvelev; Novosti Sist. Vyssh. Rast. 32: 28 (2000)

Distribution: Crimea

14 KRY

species :

Spergularia tangerina P.Monnier; Feddes Repert. 69: 50 (1964)

Distribution: Portugal; Spain; France; Corsica; Morocco; Algeria

12 COR FRA POR SPA 20 ALG MOR

Synonyms: hidden

= Spergula tangerina (P.Monnier) G.López; Lagascalia 30: 16 (2010)

= Spergularia marina f. microcarpa Batt.; Fl. Algèrie [Batt. & Trab.] Dicot.: 161 (1888)

= Spergularia salina subsp. microcarpa (Batt.) P.Monnier; Quézel & Santa, Nouv. Fl. Algerie 1: 324 (1962)

species :

Spergularia tasmanica (Kindb.) L.G.Adams; Austral. Syst. Bot. 21(4): 258 (2008)

Distribution: Australia (Tasmania); New Zealand

50 AUS 51 NZL

Synonyms: hidden

= Lepigonum tasmanicum Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 31 (1863)

species :

Spergularia tenuifolia Pomel; Bull. Soc. Sci. Phys. Algérie 11: 205 (1874)

Distribution: Morocco; Algeria

20 ALG MOR

Synonyms: hidden

= Spergula rubra subsp. tenuifolia (Pomel) Maire; Fl. Afrique N. 9: 120 (1963)

= Spergula tenuifolia (Pomel) Maire; Bull. Soc. Hist. Nat. Afr. Nord 20: 174 (1929), et 26: 191 (1935)

= Spergularia diandra var. tenuifolia (Pomel) Batt.; Fl. Algerie [Batt. & Trab.] Dicot.: 160 (1888)

species :

Spergularia villosa (Pers.) Cambess.; A. St.-Hil. Fl. Bras., Mer. 2: 178 (1830)

Name: Hairy Sandspurrey

Distribution: USA [I] (California [I], Oregon [I]); Mexico [I] (Baja California [I]); Peru; Uruguay (Montevideo); Argentina (Buenos Aires, Catamarca, Chubut, Cordoba, Entre Rios, Jujuy, La Rioja, Rio Negro, Santa Cruz, Santa Fe, San Juan, San Luis); Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Biobío, Los Lagos)

73 usa 76 usa 79 mex 83 PER 85 ARG CLE URU

Synonyms: hidden

= Arenaria alba var. villosa Larrañaga; Escritos Damaso Antonio Larrañaga 2: 154 (1923)

= Arenaria berteroana Phil.; Linnaea 28: 673 (1856)

= Buda rupestris (Cambess.) F.Hanb.; H. C. Watson & auct. suc. (eds.), London Cat. Brit. Pl., ed. 9: 12 (1895)

= Lepigonum arenarium Kindb.; Monogr. Lepig. (Syn. Framst. Lepig.) 13 (1863)

= Lepigonum glandulosum Liebm.; Index Seminum (C, Hauniensi) 1853: 21 (1853), nom. illeg.

= Lepigonum liebmannianum Lange; Index Sem. Hort. Haun. App. 2 (1858)

= Lepigonum trachyspermum Kindb.; Monogr. Gen. Lepigon.: 31 (1863)

= Lepigonum villosum (Pers.) Fisch. & C.A.Mey.; Index Seminum (LE, Petropolitanus) 4: 15 (1838)

= Spergula villosa Pers.; Syn. 1: 522 (1805)

= Spergularia aprica Phil.; Anales Univ. Chile 81: 766 (1892)

= Spergularia liebmanniana (Lange) Rohrb.; Linnaea 37: 241 (1872)

= Spergularia remotiflora Steud.; Flora 39: 424 (1856)

= Spergularia rupestris Cambess.; A. St.-Hil. Fl. Bras., Mer. 2: 176 (1829)

= Spergularia villosa var. berteroana (Phil.) Rohrb.; Fl. Bras. (Mart.) 14, pt. 2: 269 (1872)

= Spergularia villosa var. rupestris (Cambess.) Rohrb.; Mart., Fl. Bras. 14(2): 269 (1872)

= Tissa aprica (Phil.) Reiche; Anales Univ. Chile 91: 349 (1895)

= Tissa glandulosa Reiche; Anales Univ. Chile 91: 344 (1895), nom. superfl.

= Tissa media var. berteroana (Phil.) Reiche; Anales Univ. Chile 91: 349 (1895)

= Tissa villosa (Pers.) Britton; Bull. Torr. Bot. Club. 16: 62, 129 (1889)

species :

Spergularia ×hybrida Hausskn.; Mitth. Thüring. Bot. Vereins, n.f. 5: 59 (1893)

Distribution: Europe

Remarks: S. bocconei x S. diandra

species :

Spergularia ×pseudoazorica (Reyn.) P.Fourn.; Quatre Fl. France: 295 (1936)

Distribution: Europe

Remarks: S. marina x S. media

Synonyms: hidden

= Spergularia marginata var. pseudoazorica Reyn.; Bull. Soc. Bot. France 1916: 255 (1916)

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A complete, synonymic checklist of the Higher Plants of the World

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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

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How to Cite:

Search

+ Family

Filter

Order

Family (40)

Genus (104)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species, 8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms, in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]