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Ver. 19.2
April 17th, 2024

210.0000 Family Zygophyllaceae R.Br.; M. Flinders, Voy. Terra Austral. 2: 545 (1814), nom. cons.
26 genera, ca. 280 spp.
Species listed in database: 282; hybrids: 1
= Agialidaceae Wettst.; Handb. Syst. Bot., ed. 2, 2: 620 (1911)
= Balanitaceae M.Roem.; Fam. Nat. Syn. Monogr. 1: 26 (1846), nom. cons.
= Tribulaceae Trautv.; Estestv. Istor. Gub. Kievsk. Uchebn. Okr. Bot. Sist.: 28 (1853)
210.5005 Genus Zygophyllum L.; Sp. Pl. 1: 385 (1753) s. str.
Distribution: E-Mediterranean (1), mostly W-Asia to China, few Africa (?). A few African species still need transfer into Tetraena or Roepera.
Species listed in database: 45
= Agrophyllum Neck.; Elem. 2: 227 (1790)
= Fabago Mill.; Gard. Dict. Abridg. Ed. 4 (1754)
= Halimiphyllum (Engl.) Boriss.; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 18: 144 (1957)
= Sarcozygium Bunge; Linnaea 17: 7. t. 1 (1843)
species :
Zygophyllum atriplicoides Fisch. & C.A.Mey.; Ind. Sem. Horti Petrop. 1: 41 (1835)
subspecies :
subsp. atriplicoides
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Armenia; Iraq (NW-Iraq, W-Iraq: Desert); Iran (EC-Iran, E-Iran, NE-Iran: Mts., S-Iran, W-Iran); Afghanistan (Badakshan, Bamyan, Badghys, Farah, Herat, Samangan, Zabul); Syria (C-Syrian Desert, Jazira, NW-Syria); Pakistan
32 KGZ TKM TZK UZB 33 ARM 34 AFG IRN IRQ SYR 40 PAK
Synonyms: hidden
= Halimiphyllum atriplicoides (Fisch. & C.A.Mey.) Boriss.; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 18: 150 (1957)
= Halimiphyllum megacarpum (Boriss.) Boriss.; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 18: 152 (1957)
= Zygophyllum atriplicoides var. megacarpum (Boriss.) Akhiani; Iranian J. Bot. 5(1): 42 (1991)
= Zygophyllum megacarpum Boriss.; Komarov, Fl. USSR 14: 730 (1949)
subspecies :
subsp. eurypterum (Boiss. & Buhse) Popov; Byull. Sredne-Aziatsk. Gosud. Univ. 12: 119 (1926)
Distribution: Turkmenistan; Iraq (NE-Iraq); Iran (E-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Balkh, Bamyan, Farah, Ghorat, Helmand, Herat, Parwan, Samangan, Zabul); Lebanon; Pakistan (Baluchistan, Quetta)
32 TKM 34 AFG IRN IRQ LEB 40 PAK
Synonyms: hidden
= Halimiphyllum eurypterum (Boiss. & Buhse) Boriss.; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 18: 146 (1957)
= Zygophyllum eurypterum Boiss. & Buhse; Nouv. Mém. Soc. Imp. Naturalistes Moscou 12: 49 (1860)
species :
Zygophyllum balchaschense Boriss.; Komarov, Fl. USSR 14: 725 (1949)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum betpakdalense Golosk. & Semiotr.; Bull. Acad. Sci. Kazakk SSR, Bot. & Soil Sci. Ser., Fasc. (7) 74 (1960)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum borissovae Beier & Thulin; Pl. Syst. Evol. 240(1-4): 33 (2003), nom. nov.
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Zygophyllum microcarpum Boriss.; Komarov, Fl. USSR 14: 724 (1949)
species :
Zygophyllum brachypterum Kar. & Kir.; Bull. Soc. Imp. Naturalistes Moscou 397 (1841)
Distribution: Kazakhstan; C-Iran; China (W-Gansu, Xinjiang); Mongolia
32 KAZ 34 IRN 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum fabago subsp. brachypterum (Kar. & Kir.) Popov; (1925)
species :
Zygophyllum cuspidatum Boriss.; Komarov, Fl. USSR 14: 726 (1949)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum darvasicum Boriss.; Komarov, Fl. USSR 14: 729 (1949)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Halimiphyllum darvasicum (Boriss.) Boriss.; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 18: 150 (1957)
species :
Zygophyllum dregeanum Sond.; Harv. & Sond. Fl. Cap. 1: 365 (1860)
Distribution: South Africa (N-Cape Prov.)
27 SAF
Remarks: belongs in another genus
Synonyms: hidden
= Zygophyllum densiflorum Schinz; Verh. Bot. Ver. Brandenb. 31: 181 (1896)
species :
Zygophyllum eichwaldii C.A.Mey.; K. E. I. von Eichwald, Pl. Nov. Casp.-Cauc.: 15 (1831)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
Synonyms: hidden
= Zygophyllum coccineum Lepech.; Tagebuch Reise Russ. 1: 310 (1774)
species :
Zygophyllum fabago L.; Sp. Pl. 385 (1753)
Name: Bohnen-Jochblatt (DE), Syrian beancaper (EN)
Distribution: Germany [I]; Spain; Baleares; France [I]; Sardinia [I]; Romania; E-European Russia; Ukraine; Crimea; Egypt (Eastern Desert); Kazakhstan; Turkmenistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, S-Anatolia, SE-Anatolia); Iraq (NW-Iraq, SE-Iraq); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Badghys, Bamyan, Faryab, Ghazni, Herat, Kabul, Parwan, Qunduz, Samangan); Lebanon (Antilebanon); Syria (C-Syrian Desert, Jazira, NW-Syria, W-Syrian Mountains); Israel (N-Israel); Sinai peninsula (N-Sinai); Jordania (W-Jordania); China (NW-Gansu, W-Nei Mongol, Qinghai, Xinjiang); Pakistan (Baluchistan, Quetta); USA [I] (California [I], Colorado [I], Idaho [I], Kansas [I], Montana [I], New Mexico [I], Nevada [I], New York [I], Pennsylvania [I], Texas [I], Washington State [I])
11 ger 12 BAL fra sar SPA 13 ROM 14 KRY RUS UKR 20 EGY 32 KAZ TKM 33 ARM AZE GEO NCS 34 AFG IRN IRQ ISR JOR LEB SIN SYR TUR 36 CHI 40 PAK 73 usa 74 usa 75 usa 76 usa 77 usa
Synonyms: hidden
= Fabago alata Moench; Suppl. Meth. (Moench) 90 (1802)
= Fabago major Sweet; Brit. Fl. Gard. 6: t. 226 (1834)
= Zygophyllum fabagineum St.-Lag.; Ann. Soc. Bot. Lyon 7: 137 (1880)
= Zygophyllum fabago subsp. dolichocarpum Popov ex Hadidi; Fl. Iran. 98: 27 (1972)
= Zygophyllum fabago subsp. orientale Boriss. ex Hadidi; Fl. Iran. 98: 27 (1972)
= Zygophyllum micronatum Opiz; Naturalientausch 6: 42 (1824)
= Zygophyllum typicum (Popov) C.Regel; Candollea 16: 232 (1958)
species :
Zygophyllum fabagoides Popov; Bull. Univ. As. Centr. 11: 113 (1925)
Distribution: Kazakhstan; China (Gansu, Xinjiang)
32 KAZ 36 CHI
species :
Zygophyllum furcatum C.A.Mey.; Ledeb. Fl. Alt. 2: 106 (1830)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum gobicum Maxim.; Enum. Fl. Mongol. 125 (1889)
Distribution: Kazakhstan; China (N-Gansu, W-Nei Mongol, Qinghai, E-Xinjiang); Mongolia
32 KAZ 36 CHI 37 MON
species :
Zygophyllum gontscharovii Boriss.; Komarov, Fl. USSR 14: 729 (1949)
Distribution: Turkmenistan; Tajikistan; Iran; Afghanistan (Bamyan, Farah, Samangan); Pakistan
32 TKM TZK 34 AFG IRN 40 PAK
Synonyms: hidden
= Halimiphyllum gontscharovii (Boriss.) Boriss.; Not. Syst. Herb. Inst. Bot. Acad. Sci. URSS 18: 148 (1957)
= Zygophyllum eurypterum subsp. gontscharovii (Boriss.) Hadidi; Fl. Iran. 98: 24 (1972)
species :
Zygophyllum heterocladum Rech.f. & Patzak; K. Danske Vid. Selsk., Biol. Skrift. 13: No. 4 (Symb. Afghan. v.) 15 (1963)
Distribution: Afghanistan (Badakshan)
34 AFG
Synonyms: hidden
= Halimiphyllum heterocladum (Rech.f. & Patzak) Adylov; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 20: 6 (1982)
species :
Zygophyllum iliense Popov; Bull. Univ. As. Centr. 12: 112 (1926)
Distribution: Kazakhstan; China (Gansu, W-Nei Mongol, Xinjiang)
32 KAZ 36 CHI
species :
Zygophyllum jaxarticum Popov; Fl. Uzbekistan. 4: 58 (1959)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; China (Xinjiang)
32 KAZ TKM TZK UZB 36 CHI
Synonyms: hidden
= Zygophyllum dielsianum (Popov) Popov; Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 3, 75 (1928) non Schltr.
= Zygophyllum miniatum subsp. dielsianum Popov
species :
Zygophyllum kansuense Y.X.Liou; Acta Phytotax. Sin. 18(4): 484 (1980)
Distribution: China (Gansu)
36 CHI
species :
Zygophyllum karatavicum Boriss.; Komarov, Fl. USSR 14: 726 (1949)
Distribution: Kazakhstan (Karatau)
32 KAZ
species :
Zygophyllum kaschgaricum Boriss.; Komarov, Fl. USSR 14: 728 (1949)
Distribution: ?Kazakhstan; China (Xinjiang); Mongolia
36 CHI 37 MON
Synonyms: hidden
= Sarcozygium kaschgaricum (Boriss.) Y.X.Liou; Fl. Reipubl. Popul. Sin. 43(1): 142 (1998)
species :
Zygophyllum kegense Boriss.; Komarov, Fl. USSR 14: 727 (1949)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Zygophyllum kopalense Boriss.; Komarov, Fl. USSR 14: 726 (1949)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum lehmannianum Bunge; Arb. Naturf. Ver. Riga 1: II. 202 (1847)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan
32 KAZ TKM TZK UZB
Synonyms: hidden
= Zygophyllum karelini Fisch. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou (1839) 149, nomen
species :
Zygophyllum loczyi Kanitz; Növenyt. Gyujtesek Eredm. Grof Szechenyi Bela Keletazsiai Utjabol 13 (1891)
Distribution: ?Kazakhstan; China (Gansu, W-Nei Mongol, Qinghai, Xinjiang)
36 CHI
species :
Zygophyllum macropodum Boriss.; Komarov, Fl. USSR 14: 161, 724 (1949)
Distribution: Kazakhstan; China (Xinjiang)
32 KAZ 36 CHI
species :
Zygophyllum melongena Bunge; Ledeb. Fl. Alt. 2: 104 (1830)
Distribution: Siberia (Altai); Mongolia
30 SIB 37 MON
species :
Zygophyllum miniatum Cham. & Schltdl.; Linnaea 5: 49 (1830)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Afghanistan (Badghys, Bamyan, Kabul)
32 KAZ KGZ TKM TZK UZB 34 AFG
variant :
var. englerianum Popov; (1925)
Distribution: Turkmenistan; Uzbekistan (Samarkand, Aralo-Caspia); Afghanistan
32 TKM UZB 34 AFG
Synonyms: hidden
= Zygophyllum englerianum (Popov) Popov; Fl. Uzbekistan. 4: 57 (1959)
species :
Zygophyllum mucronatum Maxim.; Bull. Acad. Petersb. 27: 438 (1881)
Distribution: China (Gansu, W-Nei Mongol, Ningxia, Qinghai, Xinjiang); Mongolia
36 CHI 37 MON
species :
Zygophyllum neglectum Grubov; Opred. Sosud. Rast. Mongolii (Key Vasc. Pl. Mongolia): 176 (1982)
Distribution: Mongolia
37 MON
species :
Zygophyllum obliquum Popov; Bull. Univ. As. Centr. 11: 113 (1925)
Distribution: Kazakhstan; Kyrgyzstan; Tajikistan; Afghanistan (Wakhan); China (Gansu, Xinjiang)
32 KAZ KGZ TZK 34 AFG 36 CHI
species :
Zygophyllum ovigerum Fisch. & C.A.Mey. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou (1839) 142, nomen, Boiss. Fl. Orient. 1: 914
Distribution: SE-European Russia; W-Kazakhstan; Turkmenistan; Uzbekistan
14 RUS 32 KAZ TKM UZB
species :
Zygophyllum oxianum Boriss.; Komarov, Fl. USSR 14: 723 (1949)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; Iran (NE-Iran: Mts., N-Iran); Afghanistan (Faryab, Samangan)
32 KAZ TKM TZK UZB 34 AFG IRN
Synonyms: hidden
= Zygophyllum fabago var. oxianum (Boriss.) Kitam.
species :
Zygophyllum oxycarpum Popov; Bull. Univ. As. Centr. 12: 112 (1926), et in Bull. Univ. As. Centr. Suppl.,12
Distribution: Kazakhstan; China (Xinjiang)
32 KAZ 36 CHI
Synonyms: hidden
= Zygophyllum budunense Semiotr.; Not. Syst. Herb. Inst. Bot. Acad. Sci. Kasachst. 3: 19 (1965)
species :
Zygophyllum pamiricum Grubov; Novosti Sist. Vyssh. Rast. 31: 175 (1998)
Distribution: China (Xinjiang)
36 CHI
species :
Zygophyllum pinnatum Cham. & Schltdl.; Linnaea 5: 48 (1830)
Distribution: C-European Russia; E-European Russia (incl. S-Urals); Siberia (Altai, Krasnoyarsk); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Iran (N-Iran); China (Xinjiang); Mongolia
14 RUS 30 SIB 32 KAZ KGZ TKM TZK UZB 34 IRN 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum chakassicum (Peschkova) A.L.Ebel; Botanica Pacifica 12(2): 129 (2023)
= Zygophyllum macropterum C.A.Mey.; Ledebour, Fl. Altaic. 2: 102 (1830)
= Zygophyllum pinnatum subsp. chakassicum Peschkova; Fl. Sibir. 10: 32 (1996)
= Zygophyllum sieversianum Stephan ex Ledeb.; Fl. Ross. (Ledeb.) 1(3): 484 (1843)
species :
Zygophyllum potaninii Maxim.; Bull. Acad. Petersb. 27: 488 (1881)
Distribution: Kazakhstan; China (Gansu, W-Nei Mongol, Xinjiang); Mongolia
32 KAZ 36 CHI 37 MON
species :
Zygophyllum pterocarpum Bunge; Ledeb. Fl. Alt. 2: 103 (1830)
Distribution: Siberia (Altai, Tuva); Kazakhstan; China (Gansu, W-Nei Mongol, Xinjiang); Mongolia
30 SIB 32 KAZ 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum altaicum Stephan ex Ledeb.; Fl. Ross. 1: 484 (1843)
= Zygophyllum pterocarpum subsp. tuvinicum Peschkova; Fl. Sibir. 10: 33 (1996)
= Zygophyllum pterocarpum var. microcarpum Y.X.Liou; Acta Phytotax. Sin. 18(4): 485 (1980)
= Zygophyllum tuvinicum (Peschkova) A.L.Ebel; Botanica Pacifica 12(2): 129 (2023)
species :
Zygophyllum rosowii Bunge; Linnaea 17: 5 (1843)
Distribution: Kazakhstan; Kyrgyzstan; Tajikistan; China (Gansu, Xinjiang); Mongolia
32 KAZ KGZ TZK 36 CHI 37 MON
variant :
var. latifolium (Schrenk) Popov; Bjull. Sredne-Aziatsk. Gosud. Univ. 11: 118 (1925)
Distribution: Kazakhstan; China (Gansu, Nei Mongol, Xinjiang); Mongolia
32 KAZ 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum latifolium Schrenk; Bull. Phys.-Math. Acad. Petersb. 2: 198 (1844)
species :
Zygophyllum sinkiangense Y.X.Liou; Acta Phytotax. Sin. 18: 484 (1980)
Distribution: China (Xinjiang)
36 CHI
species :
Zygophyllum stenopterum Schrenk; Bull. Phys.-Math. Acad. Petersb. 3: 308 (1845)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum subtrijugum C.A.Mey.; Ledeb. Fl. Alt. 2: 105 (1830)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum taldykurganicum Boriss.; Komarov, Fl. USSR 14: 727 (1949)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum trialatum Blatt. & Hallb.; Journ. Ind. Bot. 1: 90 (1919)
Distribution: Pakistan (Baluchistan)
40 PAK
Remarks: provisionally accepted name
species :
Zygophyllum turcomanicum Fisch. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou (1839) 149, nomen, Boiss. Fl. Orient. 1: 914
Distribution: Kazakhstan; Turkmenistan; Uzbekistan
32 KAZ TKM UZB
species :
Zygophyllum xanthoxylum (Bunge) Maxim.; Fl. Tangut. 103 (1889)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang, Qinghai, Nei Mongol, NC-China); Mongolia
32 KGZ TZK UZB 36 CHI 37 MON
Synonyms: hidden
= Sarcozygium xanthoxylum Bunge; Linnaea 17: 8 (1843)
= Zygophyllum xanthoxylum (Bunge) Engl.; Engl. & Prantl, Natürl. Pflanzenfam. 3: 4 (1890) 81, isonym
= Zygophyllum zanthoxylum ´Engl.´ ex Dippel; Handb. Laubholzk. 2: 358 (1893)
= Zygophyllum ferganense (Drobow) Boriss.; Komarov, Fl. USSR 14: 184 (1949)
= Zygophyllum xanthoxylon var. ferganense Drobow
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A complete, synonymic checklist of the Higher Plants of the World


Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life