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049.0000 Family Onocleaceae Pic.Serm.; Webbia 24: 708 (1970)
1-4 genera, 5-6 spp.
Species listed in database: 6
= Onocleeae Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 29 (1866)
= Onocleinae Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: I, 4: 159 (1899)
049.0004 Genus Pentarhizidium Hayata; Bot. Mag. (Tokyo) 41: 715 (1927)
Distribution: E-Asia to Indian subcontinent
Species listed in database: 2
= Pteretis Raf.; Amer. Monthly Mag. & Crit. Rev. 2: 268 (1818)
= Pterilis Raf.; Amer. Monthly Mag. & Crit. Rev. 4: 195 (1819)
species :
Pentarhizidium intermedium (C.Chr.) Hayata; Bot. Mag. (Tokyo) 42: 316 (1928)
Distribution: China (Guizhou, Hebei, Hubei, Shanxi, Sichuan, Yunnan); Nepal; N-India (Uttarakhand, Sikkim, Arunachal Pradesh)
36 CHI 40 IND NEP
Remarks: valid species fide Fraser-Jenkins 2008 and 2018, not a hybrid
Synonyms: hidden
= Matteuccia intermedia C.Chr.; Bot. Gaz. 66: 337 (1913)
= Onoclea intermedia (C.Chr.) M.Kato; Fern Gaz. 14(2): 71 (1991)
= Pteretis intermedia (C.Chr.) Ching; Lingnan Sci. J. 21: 36 (1945)
species :
Pentarhizidium orientale (Hook.) Hayata; Bot. Mag. (Tokyo) 42: 345 (1928)
Distribution: Russian Far East (Kuril Isl., Primorye, Sakhalin); China (Anhui, Chongqing, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shanxi, Sichuan, Zhejiang); Tibet; North Korea; South Korea; Japan; Taiwan; Bhutan; NE-India (Arunachal Pradesh, Meghalaya); ?Myanmar
31 RFE 36 CHI TIB 38 JAP KOR NKO TAI 40 BHU IND
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris peregrina C.Chr.; Index Filic. 284 (1905)
= Matteuccia cavaleriana (Christ) C.Chr.; Index Filic. 420 (1906)
= Matteuccia japonica (Hayata) C.Chr.; Index Filic., Suppl. Tert. 127 (1934)
= Matteuccia orientalis (Hook.) Trevis.; Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 3, 14: 586 (1869)
= Matteuccia orientalis f. monstra Ching & K.H.Shing; J. Arnold Arbor. 64(1): 25 (1983)
= Nephrodium regulare Baker; J. Bot. 13: 200 (1875) (non Desv. (1827))
= Onoclea orientalis (Hook.) Hook.; Sp. Fil. [W.J.Hooker] 4: 161 (1862)
= Pentarhizidium japonicum Hayata; Bot. Mag. (Tokyo) 42: 345 (1928)
= Pteretis japonica (Hayata) Ching; Lingnan Sci. J. 21: 36 (1945)
= Pteretis orientalis (Hook.) Ching; Lingnan Sci. J. 21: 36 (1945)
= Pterinodes orientalis (Hook.) Kuntze; Revis. Gen. Pl. 2: 820 (1891)
= Struthiopteris cavaleriana Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 13: 118 (1904)
= Struthiopteris orientalis Hook.; Sec. Cent. Ferns 4 (1861)
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A complete, synonymic checklist of the Higher Plants of the World


Version 26.05 (May 20th, 2026)

Covering 359,646 accepted species,
8,890 hybrids, 47,679 accepted infraspecific taxa and 1,012,517 synonyms,
in total 1,428,732* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life