045.0000
Family
Woodsiaceae (A.Gray) Herter; Revista Sudamer. Bot. 9: 14 (1949) s. str.
2 genera, ca. 50 spp. Systematics see Lu et al. (2020).
Species listed in database: 52; hybrids: 5
= Protowoodsioideae Shmakov; Turczaninowia 4(1-2): 65 (2001), Subfamily
= Woodsieae A.Gray; Man. Bot. [A.Eaton], ed. 2: 588 (1856)
= Woodsiinae Hook.; Sp. Fil. [W.J.Hooker] 1: 58 (1844)
= Woodsioideae Fomin; Fl. Caucas. Crit. 3(1): 2, 5 (1911)
045.0003
Genus
Physematium Kaulf.; Flora 12: 341 (1829)
Distribution: North America (subgen. Woodsiopsis), New World and southern Africa/Madagascar (subgen. Physematium), C-Asia to E-Asia (subgen. Cheilanthopsis)
Species listed in database: 24; hybrids: 2
= Cheilanthopsis Hieron.; Notizbl. Bot. Gart. Berlin-Dahlem 7: 406 (1920)
= Hymenocystis C.A.Mey.; Verz. Pfl. Casp. Meer [C.A. von Meyer] 229 (1831)
= Physematium subgen.Cheilanthopsis (Hieron.) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
= Physematium subgen.Woodsiopsis (Shmakov) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
= Protowoodsia Ching; Lingnan Sci. J. 21: 36 (1945)
= Woodsia subgen.Cheilanthopsis (Hieron.) X.C.Zhang & R.Wei; PLoS ONE 13(2): e0136318 (13) (2015)
= Woodsiopsis Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Physematium angolense
(Schelpe) Shmakov; Turczaninowia 18(2): 12 (2015)
Distribution: Angola; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal)
26 ANG 27 SAF
Synonyms: hidden
= Woodsia angolensis Schelpe; Garcia de Orta, Ser. Bot. 3(1): 53 (1976)
species :
Physematium burgessianum
(Gerrard ex Hook. & Baker) Shmakov; Turczaninowia 18(2): 12 (2015)
Distribution: Zimbabwe; South Africa (Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho; Madagascar
26 ZIM 27 LES SAF 29 MDG
Synonyms: hidden
= Diacalpe burgessiana (Gerrard ex Hook. & Baker) Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
= Woodsia burgessiana Gerrard ex Hook. & Baker; Syn. Fil. [W.J.Hooker & Baker] 48 (1866)
= Woodsia montevidensis var. burgessiana (Gerrard ex Hook. & Baker) Schelpe; J. S. African Bot. 35: 138 (1969)
species :
Physematium canescens
(Kunze) Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
Distribution: Mexico (Ciudad de Mexico, Mexico State, Oaxaca, Puebla, San Luis Potosi)
79 MEX
Synonyms: hidden
= Cheilanthes canescens (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 25, no. 18 (1859)
= Cheilanthes canescens Kunze; Linnaea 18: 143 (1839)
= Woodsia canescens (Kunze) Mett.; Ann. Sci. Nat., Bot., Sér. 5, 2: 249 (1864)
species :
Physematium cochisense
(Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
Name: Cochise cliff fern
Distribution: USA (SE-Arizona, SW-New Mexico); Mexico (Chihuahua, Sonora, ?Baja California, Sur, Durango, Sinaloa)
76 USA 77 USA 79 MEX
Conservation Status: IUCN R
Synonyms: hidden
= Woodsia cochisensis Windham; Contr. Univ. Michigan Herb. 19: 54 (1993)
= Woodsiopsis cochisensis (Windham) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Physematium cystopteroides
(Windham & Mickel) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
Distribution: Mexico (Chihuahua, Durango, Sinaloa, Sonora)
79 MEX
Synonyms: hidden
= Woodsia cystopteroides Windham & Mickel; Mem. New York Bot. Gard. 88 [Pterid. Mexico]: 688 (2004)
= Woodsiopsis cystopteroides (Windham & Mickel) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Physematium elongatum
(Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
Distribution: China (NW-Yunnan); Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, West Bengal, Darjeeling, Arunachal Pradesh); Myanmar
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: subgen. Cheilanthopsis
Synonyms: hidden
= Cheilanthopsis elongata (Hook.) Copel.; Univ. Calif. Publ. Bot. 12: 395 (1931)
= Woodsia elongata Hook.; Sp. Fil. [W.J.Hooker] 1: 62, t. 21 c (1844)
species :
Physematium fragile
(Trevir.) Kunze; Analecta Pteridogr. 42 (1837)
Distribution: Northern Caucasus; Georgia [Caucasus] (Abkhazia, Samegrelo, Adzharia, Imereti, Shiga Kartli, Mtiuleti, Tush-Pshav-Khevsureti); Azerbaijan
33 AZE GEO NCS
Synonyms: hidden
= Cystopteris caucasica (C.A.Mey.) Fée; Mém. Foug. 5 (Gen. Filic.) 299 (1852)
= Diacalpe fragilis (Trevir.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1875)
= Dicksonia fragilis Trevir.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 7: 155, t. 3, f. 18-19 (1816)
= Hymenocystis caucasica C.A.Mey.; Verz. Pfl. Casp. Meer [C.A. von Meyer] 229 (1831)
= Hymenocystis fragilis (Trevir.) Askerov; Izv. Akad. Nauk Azerbaidzhansk. S.S.R., Ser. Biol. Med. Nauk 1986(3): 52 (1986)
= Woodsia caucasica (C.A.Mey.) J.Sm.; J. Bot. [Hooker] 4: 191 (1841)
= Woodsia fragilis (Trevir.) T.Moore; Index Fil. [T. Moore] 283 (1861), nom. illeg.
species :
Physematium hieronymi
Ponce & Arana; Willdenowia 55(1): 34 (2025)
Distribution: Argentina (Catamarca, Jujuy, Salta, Tucumán)
85 ARG
Remarks: subgen. Physematium. To be expected in Bolivia
species :
Physematium indusiosum
(Christ) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
Distribution: China (NW-Yunnan, SW-Sichuan); ?Bhutan
36 CHI
Remarks: subgen. Cheilanthopsis. Not in India
Synonyms: hidden
= Cheilanthes straminea Brause; Hedwigia 54: 205, t. 4 H (1914)
= Cheilanthopsis indusiosa (Christ) Ching; Sinensia 8: 154 (1932) (indusiora)
= Cheilanthopsis straminea (Brause) Hieron. ex Copel.; Univ. Calif. Publ. Bot. 16: 95 (1929)
= Woodsia indusiosa Christ; Notul. Syst. (Paris) 1: 44 (1909)
species :
Physematium jujuiense
(Copel.) Ponce & Arana; Willdenowia 55(1): 38 (2025)
Distribution: Peru (Cuzco); Bolivia (Cochabamba); Argentina (Catamarca, Jujuy, La Rioja, San Juan, Salta, Tucumán)
83 BOL PER 85 ARG
Remarks: subgen. Physematium
Synonyms: hidden
= Woodsia jujuiensis Copel.; Univ. Calif. Publ. Bot. 19: 297, t. 50 (1941)
= Woodsia montevidensis var. fuscipes Hieron.; Hedwigia 46: 322 (1907)
species :
Physematium kangdingense
(H.S.Kung, Li Bing Zhang & X.S.Guo) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
Distribution: China (Sichuan)
36 CHI
Remarks: subgen. Cheilanthopsis
Synonyms: hidden
= Cheilanthopsis kangdingensis (H.S.Kung, Li Bing Zhang & X.S.Guo) Shmakov; Pteridol. New Millennium 64 (2003)
= Woodsia kangdingensis H.S.Kung, Li Bing Zhang & X.S.Guo; Acta Bot. Yunnan. 17(4): 421 (1995)
species :
Physematium manchuriense
(Hook.) Nakai; Bot. Mag. (Tokyo) 39: 176 (1925)
Distribution: Russian Far East (Khabarovsk, Primorye); China (Anhui, Guizhou, Hebei, Heilongjiang, Henan, Jiangxi, Jilin, Liaoning, Shandong, Sichuan, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI 38 JAP KOR NKO
Remarks: subgen. Cheilanthopsis
Synonyms: hidden
= Diacalpe manchuriensis (Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1876)
= Protowoodsia manchuriensis (Hook.) Ching; Lingnan Sci. J. 21: 37 (1945)
= Woodsia insularis Hance; Ann. Sci. Nat., Bot., Sér. 4, 15: 228 (1861)
= Woodsia manchuriensis Hook.; Sec. Cent. Ferns, t. 98 (1861)
species :
Physematium mexicanum
(Fée) comb.ined.
Distribution: Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chihuahua, Coahuila, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Jalisco, Mexico State, Nuevo Leon, Queretaro, San Luis Potosi, Sinaloa, Sonora, Veracruz, Zacatecas)
79 MEX
Synonyms: hidden
= Woodsia mexicana Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 66, t. 26, f. 3 (1857)
= Woodsia oregana f. pusilla (E.Fourn.) B.Boivin; Bull. Soc. Bot. France 113: 408 (1967)
= Woodsia oregana var. mexicana (Fée) B.Boivin; Bull. Soc. Bot. France 113: 408 (1967)
= Woodsia pusilla E.Fourn.; Bull. Soc. Bot. France 27: 329 (1880)
= Woodsia pusilla var. mexicana (Fée) T.M.C.Taylor; Amer. Fern J. 37: 86 (1947)
= Woodsiopsis mexicana (Fée) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Physematium molle
Kaulf.; Flora 1829(1): 341 (1829)
Name: Mexican Cliff Fern
Distribution: Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala; El Salvador; Honduras; Nicaragua
79 MEX 80 ELS GUA HON NIC
Remarks: subgen. Physematium
Synonyms: hidden
= Diacalpe guatemalensis (Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1875)
= Lastrea bulbosa C.Presl; Tent. Pterid. 75 (1836)
= Woodsia fragilis Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 5, 1: 280 (1849) (non T. Moore (1857))
= Woodsia guatemalensis Hook.; Sp. Fil. [W.J.Hooker] 1: 60, t. 21 (1844)
= Woodsia mollis (Kaulf.) J.Sm.; J. Bot. [Hooker] 4: 191 (1841)
species :
Physematium montevidense
(Spreng.) Shmakov; Turczaninowia 18(2): 12 (2015)
Distribution: Brazil (Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Uruguay (Lavalleja, Maldonado, Minas, Rocha, Treinta y Tres); Argentina (Buenos Aires, Cordoba, La Rioja, Mendoza, Rio Negro, San Juan, San Luis, Tucuman)
84 BRA 85 ARG URU
Remarks: subgen. Physematium. Confirmed by Ponce et al. 2025 only for S-Brazil, Uruguay and Argentina. The formerly recorded distribution in the northern part of South America as well as the synonymy need furthern checking
Synonyms: hidden
= Cheilanthes andina Griseb.; Abh. Königl. Ges. Wiss. Göttingen 24: 342 (1879) (unconfirmed synonym)
= Cheilanthes crenata Kunze; Linnaea 9: 84 (1834) (unconfirmed synonym)
= Cystopteris vestita C.Presl; Tent. Pterid. 93 (1836), nom. nud. (unconfirmed synonym)
= Dicksonia montevidensis Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 122 (1827)
= Physematium euporolepis Trevis.; Nuovo Giorn. Bot. Ital. 7: 155 (1875) (unconfirmed synonym)
= Physematium incisum (Gillies ex Hook. & Grev.) C.Presl; Tent. Pterid. 66 (1836)
= Polypodium flabellatum Klotzsch; Linnaea 20: 382 (1847) (unconfirmed synonym)
= Woodsia crenata (Kunze) Hieron.; Bot. Jahrb. Syst. 34: 440 (1904) (unconfirmed synonym)
= Woodsia crenata var. pallidipes Hieron.; Bot. Jahrb. Syst. 34: 440 (1904) (unconfirmed synonym)
= Woodsia euporolepis (Trevis.) Trevis.; Rendiconti Reale Ist. Lombardo Sci. 2(9): 812 (1876) (unconfirmed synonym)
= Woodsia incisa Gillies ex Hook. & Grev.; Icon. Filic. 2: t. 191 (1830)
= Woodsia montevidensis (Spreng.) Hieron.; Bot. Jahrb. Syst. 22: 363 (1896)
= Woodsia obtusa var. andina Hook.; Syn. Fil. [W.J.Hooker & Baker] 48 (1866) (unconfirmed synonym)
= Woodsia obtusa var. montevidensis (Spreng.) Hicken
species :
Physematium neomexicanum
(Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
Name: New Mexican Cliff Fern (EN)
Distribution: USA (Arizona, New Mexico, S-Colorado, W-Texas, C-Oklahoma, S-Dakota); Mexico (Coahuila, Nuevo Leon, Puebla, Tamaulipas, Zacatecas)
73 USA 74 USA 76 USA 77 USA 79 MEX
Synonyms: hidden
= Woodsia neomexicana Windham; Contr. Univ. Michigan Herb. 19: 52 (1993)
= Woodsiopsis neomexicana (Windham) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Physematium obtusum
(Spreng.) Hook.; Fl. Bor.-Amer. [Hooker] 2: 259 (1840)
subspecies :
subsp. obtusum
Name: Blunt-lobed Cliff Fern
Distribution: Canada (Ontario, Québec); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, Nebraska, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee, Texas, Virginia, Vermont, Wisconsin, West Virginia)
72 CDN 74 USA 75 USA 77 USA 78 USA
Remarks: subgen. Physematium
Synonyms: hidden
= Alsophila perriniana Spreng.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 10: 232 (1821)
= Aspidium obtusum (Spreng.) Sw.; Syn. Fil. (Kiliae) [O.Swartz] 420 (1806)
= Aspidium pennsylvanicum F.Dietr.; Nachtr. Vollst. Lex. Gärtn. 9: 374 (1838), nom. superfl.
= Cystopteris albescens Link; Fil. Spec. 47 (1841)
= Cystopteris obtusa (Spreng.) C.Presl; Tent. Pterid. 93 (1836)
= Cystopteris perriniana (Spreng.) Link; Hort. Berol. [Link] 2: 131 (1833)
= Physematium perrinianum (Spreng.) C.Presl; Tent. Pterid. 66 (1836)
= Physematium vestitum (C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 66 (1851)
= Polypodium obtusum Spreng.; Anleit. Kenntn. Gew. 3: 93 (1804)
= Woodsia obtusa (Spreng.) Torr.; Rep. Bot. Dep. Surv. New York, no. 50, Assembly: 195 (1840)
= Woodsia obtusa f. darlingtonii (Stowell) M.Broun; Index No. Amer. Ferns: 166 (1938)
= Woodsia obtusa var. angusta Peck; Working List N. Amer. Pterid. 39 (1901)
= Woodsia obtusa var. darlingtonii Stowell; J. Trenton Nat. Hist. Soc. 1: 25 (1887)
= Woodsia perriniana (Spreng.) Hook. & Grev.; Icon. Filic. t. 68 (1828)
= Woodsiopsis obtusa (Spreng.) Shmakov; Turczaninowia 18(2): 13 (2015)
subspecies :
subsp. occidentale
(Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
Name: Western cliff fern
Distribution: USA (Arkansas, Kansas, Oklahoma, Texas)
74 USA 77 USA 78 USA
Synonyms: hidden
= Woodsia obtusa subsp. occidentalis Windham; Contr. Univ. Michigan Herb. 19: 56 (1993)
= Woodsiopsis obtusa subsp. occidentalis (Windham) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Physematium oreganum
(D.C.Eaton) Trevis.; Nuovo Giorn. Bot. Ital. 7: 162 (1875)
subspecies :
subsp. oreganum
Name: Oregon cliff fern
Distribution: Canada (Alberta, British Columbia, Saskatchewan); USA (California, Idaho, Montana, Nevada, Oregon, Utah, Washington State, Wyoming)
71 CDN 73 USA 76 USA
Remarks: subgen. Physematium
Synonyms: hidden
= Woodsia oregana D.C.Eaton; Canad. Naturalist & Quart. J. Sci., Ser. 2: 90 (1865)
= Woodsiopsis oregana (D.C.Eaton) Shmakov; Turczaninowia 18(2): 13 (2015)
subspecies :
subsp. cathcartianum
(B.L.Rob.) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
Name: Oregon cliff fern
Distribution: Canada (Manitoba, Ontario, Québec, Saskatchewan); USA (Arizona, California, Colorado, Iowa, Idaho, Kansas, Michigan, Minnesota, Montana, North Dakota, Nebraska, New Mexico, Nevada, New York, Oklahoma, South Dakota, Utah, Wisconsin, Wyoming)
71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA
Synonyms: hidden
= Woodsia cathcartiana B.L.Rob.; Rhodora 10: 30 (1908)
= Woodsia oregana f. cathcartiana (B.L.Rob.) B.Boivin; Bull. Soc. Bot. France 113: 408 (1966)
= Woodsia oregana f. glandulosa T.M.C.Taylor; Amer. Fern J. 37: 85 (1947)
= Woodsia oregana subsp. cathcartiana (B.L.Rob.) Windham; Contr. Univ. Michigan Herb. 19: 58 (1993)
= Woodsia oregana var. cathcartiana (B.L.Rob.) C.V.Morton; Amer. Fern J. 40: 222 (1950)
= Woodsia pusilla var. cathcartiana (B.L.Rob.) T.M.C.Taylor; Amer. Fern J. 37: 86 (1947)
= Woodsia scopulina f. cathcartiana (B.L.Rob.) Clute; Our Ferns 133 (1901)
= Woodsia scopulina var. cathcartiana (B.L.Rob.) Clute; Fern Bull. 15: 127 (1908)
= Woodsiopsis oregana subsp. cathcartiana (B.L.Rob.) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Physematium pallidum
(Copel.) Ponce & Arana; Willdenowia 55(1): 45 (2025)
Distribution: Argentina (Catamarca, Jujuy, La Rioja, San Juan, Salta, Tucumán); Bolivia (Chuquisaca, Potosí); Chile (Antofagasta, Arica y Parinacota, Tarapacá)
83 BOL 85 ARG CLE
Remarks: subgen. Physematium
Synonyms: hidden
= Woodsia pallida Copel.; Univ. Calif. Publ. Bot. 19: 2917, t. 51 (1941)
species :
Physematium peruvianum
(Hook.) Ponce & Arana; Willdenowia 55(1): 36 (2025)
Distribution: Ecuador; Peru
83 ECU PER
Remarks: subgen. Physematium
Synonyms: hidden
= Diacalpe peruviana (Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1875)
= Woodsia peruviana Hook.; Sp. Fil. [W.J.Hooker] 1: 61, t. 21 (1844)
species :
Physematium phillipsii
(Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
Name: Philipps´s Cliff Fern
Distribution: USA (S-Arizona, S-New Mexico, W-Texas); Mexico (Chihuahua, Coahuila, Durango, Jalisco, Mexico State, Sonora)
76 USA 77 USA 79 MEX
Synonyms: hidden
= Woodsia phillipsii Windham; Contr. Univ. Michigan Herb. 19: 52 (1993)
= Woodsiopsis phillipsii (Windham) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Physematium plummerae
(Lemmon) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
Name: Plummer´s Cliff Fern
Distribution: USA (Arizona, California, Colorado, New Mexico, Oklahoma, Texas); Mexico (Baja California Norte, Baja California Sur, Chihuahua, Jalisco, Sonora)
73 USA 74 USA 76 USA 77 USA 79 MEX
Synonyms: hidden
= Woodsia obtusa var. glandulosa D.C.Eaton & Faxon; Bull. Torrey Bot. Club 9: 50 (1882)
= Woodsia obtusa var. plummerae (Lemmon) Maxon; Proc. U.S. Natl. Mus. 23: 644 (1901)
= Woodsia plummerae Lemmon; Bot. Gaz. 7: 6 (1882)
= Woodsia pusilla var. glandulosa (D.C.Eaton & Faxon) T.M.C.Taylor; Amer. Fern J. 37: 86 (1947)
= Woodsiopsis plummerae (Lemmon) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Physematium pubescens
(Spreng.) Bevis; Hort. Brit. [Loudon] ed. 3, 656 (1839)
Distribution: Jamaica
81 JAM
Synonyms: hidden
= Woodsia pubescens Spreng.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 10: 233, t. 16, f. 5-7 (1821)
species :
Physematium scopulinum
Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
subspecies :
subsp. scopulinum
Name: Rocky Mountain woodsia
Distribution: Alaska; Canada (Alberta, British Columbia, Manitoba, Saskatchewan, Yukon); USA (California, Colorado, Idaho, Montana, Nevada, Oregon, Utah, Washington State, Wyoming)
70 ASK CDN 71 CDN 73 USA 76 USA
Synonyms: hidden
= Woodsia obtusa f. nana (Lemmon ex Gilbert) Clute; Fern Bull. 15: 126 (1908)
= Woodsia obtusa var. lyallii Hook.; Syn. Fil. [W.J.Hooker & Baker], ed. 1: 48 (1866)
= Woodsia obtusa var. nana Lemmon ex Gilbert; Working List N. Amer. Pterid. 21, 40 (1901)
= Woodsia oregana var. lyallii (Hook.) B.Boivin; Bull. Soc. Bot. France 113: 409 (1966)
= Woodsia scopulina D.C.Eaton; Canad. Naturalist & Quart. J. Sci., Ser. 2: 91 (1865)
= Woodsia scopulina f. nana (Lemmon) M.Broun; Index No. Amer. Ferns: 167 (1938)
= Woodsiopsis scopulina (D.C.Eaton) Shmakov; Turczaninowia 18(2): 13 (2015)
subspecies :
subsp. appalachianum
(T.M.C.Taylor) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
Name: Appalachian cliff fern
Distribution: USA (Arkansas, Georgia, Kentucky, North Carolina, Oklahoma, Tennessee, Virginia, West Virginia)
74 USA 75 USA 78 USA
Synonyms: hidden
= Woodsia appalachiana T.M.C.Taylor; Amer. Fern J. 37: 88 (1947)
= Woodsia oregana var. appalachiana (T.M.C.Taylor) B.Boivin; Bull. Soc. Bot. France 113: 409 (1967)
= Woodsia scopulina subsp. appalachiana (T.M.C.Taylor) Windham; Contr. Univ. Michigan Herb. 19: 58 (1993)
= Woodsia scopulina var. appalachiana (T.M.C.Taylor) C.V.Morton; Amer. Fern J. 40: 224 (1950)
= Woodsiopsis appalachiana (T.M.C.Taylor) Shmakov; Turczaninowia 18(2): 12 (2015)
subspecies :
subsp. laurentianum
(Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
Name: Laurent´s cliff fern
Distribution: Canada (Alberta, British Columbia, Ontario, Québec); USA (Arizona, California, Colorado, Idaho, Minnesota, Montana, Nevada, Oregon, South Dakota, Utah, Washington State, Wyoming)
71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA
Synonyms: hidden
= Woodsia scopulina subsp. laurentiana Windham; Contr. Univ. Michigan Herb. 19: 59 (1993)
= Woodsiopsis scopulina subsp. laurentiana (Windham) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Physematium ×kansanum
(R.E.Brooks) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
Distribution: USA (Kansas)
74 USA
Remarks: Woodsia obtusa x oregana var. oregana
Synonyms: hidden
= Woodsia ×kansana R.E.Brooks; Amer. Fern J. 72(3): 81 (1982)
species :
Physematium ×maxonii
(R.M.Tryon) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
Distribution: Canada (Ontario); USA (Minnesota)
72 CDN 75 USA
Remarks: Woodsia oregana var. cathcartiana x Woodsia scopulina
Synonyms: hidden
= Woodsia ×maxonii R.M.Tryon; Amer. Fern J. 38: 169 (1949)
