030.0000
Family
Pteridaceae E.D.M.Kirchn.; Schul-Bot. 109 (1831)
48 genera, ca. 1350 spp. The family has been split extensively in smaller subgroups, but Smith et al. (2006) and PPG-I argue for keeping them together.
Species listed in database: 1405; hybrids: 68
= Acrostichaceae Mett. ex A.B.Frank; Syn. Pflanzenk., ed. 2, 3: 1458 (1877), nom. rej.
= Actiniopteridaceae Pic.Serm.; Webbia 17: 5 (1962)
= Adiantaceae Newman; Hist. Brit. Ferns 5 (1840) nom. cons. over Parkeriaceae (1825)
= Anopteraceae Doweld; Tent. Syst. Pl. Vasc. (Tracheophyta) 51 (2001)
= Antrophyaceae (Link) Ching; Acta Phytotax. Sin. 16: 11 (1978)
= Ceratopteridaceae Underw.; Native Ferns, ed. 6: 65, 78 (1900)
= Cheilanthaceae B.K.Nayar; Taxon 19: 233 (1970)
= Cryptogrammaceae Pic.Serm.; Webbia 17: 299 (1963)
= Hemionitidaceae Pic.Serm.; Webbia 21: 487 (1966)
= Negripteridaceae Pic.Serm.; Nuovo Giorn. Bot. Ital., n. s. 53: 160 (1946)
= Parkeriaceae Hook.; Exot. Fl. 2: t. 147 (1825), nom. rej. in favor of Adiantaceae
= Platyzomataceae Nakai; Bull. Natl. Sci. Mus., Tokyo, 29: 4 (1950)
= Pteridaceae Reichenb.; Handb. Nat. Pfl.-Syst. 138 (1837)
= Sinopteridaceae Koidz.; Acta Phytotax. Geobot. 3: 50 (1934)
= Taenitidaceae (C.Presl) Pic.Serm.; Webbia 29: 1 (1974)
= Vittariaceae Ching; Sunyatsenia 5: 232 (1940)
030.3011
Genus
Pellaea Link; Fil. Spec. 48, 59 (1841), nom. cons., s. l.
Distribution: (s. str.) SW-USA, Mexico, to Galapagos and Haiti, (Paraceterach) Australia, some Pacific, 1 Malesia, (Sericopteris) Himalayas. The separation of Paraceterach and Sericopteris was rejected by PPG-II.
Species listed in database: 41; hybrids: 3
= Bakeropteris Kuntze; Revis. Gen. Pl. 2: 807 (1891)
= Paraceterach (F.Muell.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 75 (1947)
= Platyloma J.Sm.; J. Bot. [Hooker] 4: 160 (1841)
= Sericopteris Jing Zhao, X.M.Zhou & Z.R.He; Phytotaxa 693(2): 167 (2025)
= Synochlamys Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 35, t. 20, f. 4 (1857)
species :
Pellaea ambigua
(Fée) Baker; Syn. Fil. [W.J.Hooker & Baker] 147 (1867)
Distribution: Colombia (Sa. Nevada de Santa Marta)
82 CLM
Synonyms: hidden
= Allosorus ambiguus (Fée) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Cheilanthes ambigua (Fée) Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 49, no. 78 (1859) (non A. Rich (1832) nec Brack. (1854))
= Hemionitis ambigua (Fée) Christenh.; Global Fl. 4: 9 (2018)
= Synochlamys ambigua Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 35, t. 20, f. 4 (1857)
species :
Pellaea andromedifolia
(Kaulf.) Fée; Mém. Foug. 5 (Gen. Filic.) 129 (1852)
Name: Coffee cliffbrake
Distribution: USA (S-Oregon, California); Mexico (Baja California Norte, Baja California Sur, Isla Cedros)
73 USA 76 USA 79 MEX
Synonyms: hidden
= Allosorus andromedifolius Kaulf. ex Kuntze; Linnaea 9: 56 (1834)
= Cassebeera andromedifolia Kaulf. ex Farw.; Amer. Midl. Naturalist 12: 280 (1931)
= Cassebeera andromedifolia var. gracilis (R.W.Summers) Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Cassebeera andromedifolia var. pubescens (D.C.Eaton) Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Cassebeera andromedifolia var. rubens (D.C.Eaton) Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Hemionitis andromedifolia (Kaulf.) Christenh.; Global Fl. 4: 9 (2018)
= Notholaena andromedifolia (Kaulf.) Keyserl.; Polyp. Herb. Bunge 29 (1873)
= Pellaea andromedifolia f. gracilis (R.W.Summers) M.Broun; Index No. Amer. Ferns: 129 (1938)
= Pellaea andromedifolia f. pubescens (D.C.Eaton) M.Broun; Index No. Amer. Ferns: 129 (1938)
= Pellaea andromedifolia var. gracilis R.W.Summers ex Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Pellaea andromedifolia var. pubescens D.C.Eaton; Ferns N. Amer. 1: 205 (1878)
= Pellaea andromedifolia var. rubens D.C.Eaton; Ferns N. Amer. 1: 205 (1878)
= Pellaea rafaelensis Moxley; Amer. Fern J. 6: 107, t. 8 (1915)
= Platyloma andromedifolium (Kaulf.) J.Sm.; J. Bot. [Hooker] 4: 160 (1841)
= Pteris andromedifolia Kaulf.; Enum. Filic. [Kaulfuss] 188 (1824)
species :
Pellaea angolensis
Schelpe; Garcia de Orta, Ser. Bot. 3(1): 53 (1976)
Distribution: Angola
26 ANG
Remarks: of uncertain taxonomic position, unsampled in phylogenetic studies
Synonyms: hidden
= Hemionitis angolensis (Schelpe) Christenh.; Global Fl. 4: 9 (2018)
species :
Pellaea atropurpurea
(L.) Link; Fil. Spec. 59 (1841)
Name: Purple cliffbrake
Distribution: Canada (Ontario, Québec); USA (Alabama, Arkansas, Arizona, Colorado, Connecticut, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Michigan, Minnesota, Missouri, Mississippi, North Carolina, Nebraska, New Jersey, New Mexico, Nevada, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia, Vermont, Wisconsin, West Virginia, Wyoming); Mexico (Chiapas, Chihuahua, Coahuila, Colima, Hidalgo, Jalisco, Nuevo Leon, Oaxaca, Puebla, Sonora, Tamaulipas, Veracruz); Guatemala
72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA 79 MEX 80 GUA
Synonyms: hidden
= Allosorus atropurpureus (L.) Kunze; Tent. Pterid. 153 (1836)
= Asplenium atropurpureum (L.) Bernh.; Neues J. Bot. 1(2): 10 (1808)
= Cassebeera atropurpurea (L.) Kaulf. ex Farw.; Amer. Midl. Naturalist 12: 281 (1930)
= Cassebeera atropurpurea var. minima Eggert ex Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Hemionitis atropurpurea (L.) Christenh.; Global Fl. 4: 10 (2018)
= Notholaena atropurpurea (L.) Keyserl.; Polyp. Herb. Bunge 30 (1873)
= Pellaea atropurpurea f. bifida Hahne; Fern Bull. 12: 117 (1904)
= Pellaea atropurpurea f. furcata Hahne; Fern Bull. 12: 117 (1904)
= Pellaea atropurpurea var. cristata Trel.; Rep. (Annual) Missouri Bot. Gard. 12: 77, pl. 34 (1901)
= Pellaea atropurpurea var. latifolia Pickett; Amer. Fern J. 4: 101 (1914)
= Pellaea atropurpurea var. minima Eggert ex Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Pellaea atropurpurea var. mucronata (Fée) E.Fourn.; Mexic. Pl. 1: 119 (1872), nom. nud.
= Pellaea atropurpurea var. nana (Richardson ex Franklin) Hook.; Sp. Fil. [W.J.Hooker] 2: 139 (1858)
= Pellaea atropurpurea var. punctata Gilbert; Working List N. Amer. Pterid. 18 (1901)
= Pellaea atropurpurea var. venosa Gilbert; Working List N. Amer. Pterid. 18 (1901)
= Pellaea mucronata Fée; Mém. Foug. 9 (Cat. Foug. Mex.): 8 (1857), nom. nud.
= Pellaea spiculata (Schkuhr) Fée; Mém. Foug. 5 (Gen. Filic.) 129 (1852)
= Platyloma atropurpureum (L.) J.Sm.; J. Bot. [Hooker] 4: 160 (1841)
= Pteris atropurpurea L.; Sp. Pl. [Linnaeus] 2: 1076 (1753)
= Pteris spiculata Schkuhr; Deutschl. Krypt. Gew. 1: 92, t. 100 (1809)
species :
Pellaea bipinnata
(Christ) G.M.Zhang & Y.Yin; Bull. Bot. Res., Harbin 44(4): 499 (2024)
Distribution: China (Gansu, Shaanxi, Sichuan, Yunnan); Tibet; Nepal; India (Arunachal Pradesh)
36 CHI TIB 40 IND NEP
Synonyms: hidden
= Gymnopteris bipinnata Christ; Notul. Syst. (Paris) 1: 55 (1909), non Notholaena bipinnata Liebm. 1849, nec al.
= Hemionitis bipinnata (Christ) Mickel; Brittonia 31(3): 411 (1979)
= Notholaena dipinnata Fraser-Jenk.; Fern Gaz. 18(5): 217 (2009), nom. nov.
= Notholaena dipinnula Fraser-Jenk.; Annot. Checkl. Ind. Pterid. 1: 250 (2016), nom. superfl.
= Paraceterach bipinnata (Christ) R.M.Tryon; Amer. Fern J. 76(4): 186 (1986)
= Paragymnopteris bipinnata (Christ) K.H.Shing; Indian Fern J. 10(1-2): 230 (1993 publ. 1994)
= Sericopteris bipinnata (Christ) Jing Zhao & X.M.Zhou; Phytotaxa 693(2): 169 (2025)
species :
Pellaea borealisinensis
(Kitag.) comb.ined.
Distribution: China (Gansu, Hebei, Henan, W-Hubei, Nei Mongol, Sichuan, Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Uttarakhand)
36 CHI TIB 40 BHU IND NEP
Remarks: Pellaea auriculata as combination would be homonym
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Gymnogramma vestita var. auriculata Franch.; Nouv. Arch. Mus. Hist. Nat., Sér. 2, 10: 123 (1887)
= Gymnopteris bipinnata var. auriculata (Franch.) Ching; Lingnan Sci. J. 15(3): 398 (1936)
= Gymnopteris borealisinensis Kitag.; Rep. Exped. Manchoukuo, Sect. IV, 2 (Contr. Cogn. Fl. Manshuricae): 83, t. 12 (1935)
= Gymnopteris vestita var. auriculata (Franch.) Ching; Ill. Chin. Pl. Pterid. 85 (1957)
= Notholaena borealisinensis (Kitag.) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 183 (1997), comb. nov.
= Paragymnopteris bipinnata var. auriculata (Franch.) K.H.Shing; Indian Fern J. 10(1-2): 230 (1993 publ. 1994)
= Sericopteris auriculata (Franch.) Jing Zhao & X.M.Zhou; Phytotaxa 693(2): 169 (2025)
species :
Pellaea brachyptera
(T.Moore) Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 2: 477 (1874)
Name: Sierra cliffbrake
Distribution: USA (S-Oregon, N-California, C-Washington)
73 USA 76 USA
Synonyms: hidden
= Allosorus bellus (T.Moore) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Allosorus brachypterus (T.Moore) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Hemionitis brachyptera (T.Moore) Christenh.; Global Fl. 4: 10 (2018)
= Pellaea bella (T.Moore) Baker; Syn. Fil. [W.J.Hooker & Baker], ed. 2: 477 (1874)
= Pellaea ornithopus var. brachyptera (T.Moore) D.C.Eaton; Rep. U.S. Geogr. Surv., Wheeler 6(Botany): 322 (1878)
= Platyloma bellum T.Moore; Gard. Chron. 1873: 213 (1873)
= Platyloma brachypterum T.Moore; Gard. Chron. 1873: 141 (1873)
species :
Pellaea bridgesii
Hook.; Sp. Fil. [W.J.Hooker] 2: 238 (1858)
Name: Bridges´ cliffbrake
Distribution: W-USA (C-California, W-Nevada, W-Idaho, Oregon)
73 USA 76 USA
Synonyms: hidden
= Allosorus bridgesii (Hook.) Luerss.; Flora 290 (1876)
= Platyloma bridgesii (Hook.) J.Sm.; Hist. Fil. 165 (1875)
= Hemionitis bridgesii (Hook.) Christenh.; Global Fl. 4: 10 (2018)
species :
Pellaea calidirupium
Brownsey & Lovis; New Zealand J. Bot. 28(3): 197 (1990)
Distribution: Australia (S-Queensland, New South Wales, Australian Capital Territory, Victoria, Tasmania); New Zealand (North Isl., South Isl.)
50 AUS 51 NZL
Synonyms: hidden
= Hemionitis calidirupium (Brownsey & Lovis) Christenh.; Global Fl. 4: 11 (2018)
= Paraceterach calidirupium (Brownsey & Lovis) Li Bing Zhang, X.M.Zhou & Jing Zhao; Phytotaxa 731(3): 229 (2025)
species :
Pellaea falcata
(R.Br.) Fée; Mém. Foug. 5 (Gen. Filic.) 129 (1852)
Name: Sickle fern (EN)
Distribution: S-India (Tamil Nadu, Kerala); Sri Lanka; peninsular Malaysia; Sulawesi; Lesser Sunda Isl. (Lombok); New Guinea (Papua New Guinea); Australia (Australian Capital Territory, Queensland, Victoria, New South Wales, Tasmania); Lord Howe Isl.; New Caledonia
40 IND SRL 42 LSI MLY NWG SUL 50 AUS LHI 60 NWC
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Allosorus falcatus (R.Br.) Kunze; Linnaea 23: 219 (1850)
= Allosorus integricuspis (E.Fourn.) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Hemionitis pellaea Christenh.; Global Fl. 4: 19 (2018), nom. nov. for Pteris falcata R. Br.
= Paraceterach falcata (R.Br.) Li Bing Zhang, X.M.Zhou & Jing Zhao; Phytotaxa 731(3): 229 (2025)
= Pellaea falcata var. denticulata Bonap.; Notes Ptéridol. 5: 126 (1917)
= Pellaea falcata var. setosa Hook.; Sp. Fil. [W.J.Hooker] 2: 136, t. 111B (1858)
= Pellaea integricuspis E.Fourn.; Ann. Sci. Nat., Bot., Sér. 5, 18: 324 (1873)
= Pellaea seticaulis (Hook.) S.R.Ghosh; J. Econ. Taxon. Bot. 7(3): 681 (1985 publ. 1986) (fide Fraser-Jenkins 1997)
= Platyloma falcatum (R.Br.) J.Sm.; J. Bot. [Hooker] 4: 160 (1841)
= Platyloma falcatum var. caudatum E.J.Lowe; Ferns 3: 84, sub t. XXX-B (1857)
= Platyloma falcatum var. setosum Bedd.; Ferns S. India 7, t. 22 (1863)
= Pteris alternifolia Wall.; Numer. List [Wallich] no. 2182 (1830), nomen
= Pteris falcata R.Br.; Prodr. Fl. Nov. Holland. 154 (1810)
= Pteris falcata var. deltoidea F.Muell.; Fragm. [Mueller] 5: 188 (1865)
= Pteris seticaulis Hook.; Hooker´s Icon. Pl. 3: t. 209 (1839)
species :
Pellaea furcata
M.A.Hern. & O.G.Martínez; Kew Bull. 76(3): 471 (2021)
Distribution: Argentina (Jujuy, Salta)
85 ARG
species :
Pellaea gastonyi
Windham; Contr. Univ. Michigan Herb. 19: 40 (1993)
Name: Gastony´s cliffbrake
Distribution: Canada (Alberta, British Columbia, Saskatchewan); USA (Missouri, South Dakota, Wyoming)
71 CDN 73 USA 74 USA 75 USA
Synonyms: hidden
= Hemionitis gastonyi (Windham) Christenh.; Global Fl. 4: 14 (2018)
species :
Pellaea glabella
Mett.; Linnaea 36: 87 (1869)
subspecies :
subsp. glabella
Name: Smooth cliffbrake
Distribution: Canada (Ontario, Québec); USA (Arkansas, Connecticut, Iowa, Illinois, Indiana, Kansas, Kentucky, Maryland, Michigan, Minnesota, Missouri, Nebraska, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Tennessee, Texas, Virginia, Vermont, Wisconsin, West Virginia)
72 CDN 74 USA 75 USA 77 USA 78 USA
Synonyms: hidden
= Cassebeera atropurpurea var. glabella (Mett.) Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Hemionitis glabella (Mett.) Christenh.; Global Fl. 4: 14 (2018)
= Pellaea atropurpurea f. glabella (Mett.) Clute; Our Ferns 109 (1901)
= Pellaea atropurpurea unranked glabella (Mett.) F.C.Gates; Trans. Kansas Acad. Sci. 42: 135 (1940)
= Pellaea atropurpurea var. bushii Mack.; Man. Fl. Jackson County 5 (1902)
= Pellaea atropurpurea var. glabella (Mett.) F.C.Gates; Trans. Kansas Acad. Sci. 42: 135 (1940)
subspecies :
subsp. missouriensis
(Gastony) Windham; Contr. Univ. Michigan Herb. 19: 39 (1993)
Name: Missouri cliffbrake
Distribution: USA (C-Missouri)
75 USA
Synonyms: hidden
= Pellaea glabella var. missouriensis Gastony; Amer. Fern J. 78(2): 64 (1988)
subspecies :
subsp. occidentalis
(E.Nelson) Windham; Contr. Univ. Michigan Herb. 19: 39 (1993)
Name: Western dwarf cliffbrake
Distribution: Canada (Alberta, Manitoba, Northern Territories, Saskatchewan); USA (Montana, North Dakota, South Dakota, Utah, Wyoming)
70 CDN 71 CDN 73 USA 74 USA 76 USA
Synonyms: hidden
= Pellaea atropurpurea var. nana (Richardson) Hook.; Sp. Fil. [W.J.Hooker] 2: 139 (1858)
= Pellaea atropurpurea unranked occidentalis E.E.Nelson; Fern Bull. 7: 30 (1899)
= Pellaea glabella var. nana (Richardson) Cody; Canad. Field-Naturalist 77: 109 (1963)
= Pellaea glabella var. occidentalis (E.Nelson) Butters; Amer. Fern J. 7: 82 (1917)
= Pellaea occidentalis (E.Nelson) Rydb.; Mem. New York Bot. Gard. 1: 466 (1900)
= Pellaea pumila Rydb.; Mem. New York Bot. Gard. 1: 4 (1900)
= Pteris gracilis var. nana Richardson; Narr. Journey Polar Sea [Franklin] 39 (1823)
subspecies :
subsp. simplex
(Butters) Á.Löve & D.Löve; Taxon 26: 325 (1977)
Name: Smooth cliffbrake
Distribution: Canada (Alberta, British Columbia); USA (Arizona, Colorado, Idaho, Montana, New Mexico, Nevada, Utah, Washington State, Wyoming)
71 CDN 73 USA 76 USA 77 USA
Synonyms: hidden
= Pellaea atropurpurea var. simplex (Butters) C.V.Morton; Leafl. W. Bot. 16: 156 (1951)
= Pellaea glabella var. simplex Butters; Amer. Fern J. 7: 84 (1917)
= Pellaea occidentalis subsp. simplex (Butters) Gastony; Amer. Fern J. 78(2): 64 (1988)
= Pellaea suksdorfiana Butters; Amer. Fern J. 11: 40, 75-82 (1921)
species :
Pellaea himalaica
(Fraser-Jenk.) G.M.Zhang & Y.Yin; Bull. Bot. Res., Harbin 44(4): 499 (2024)
Name: Mouse-ear fern (EN)
Distribution: China (W-Guizhou, Hebei, Shanxi, Sichuan, Yunnan); Tibet; Taiwan; Pakistan (Swat, Hazara); Nepal; Bhutan; India (Arunachal Pradesh, Himachal Pradesh, Manipur, Sikkim, Uttarakhand); Myanmar; N-Thailand; Philippines
36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA THA 42 PHI
Synonyms: hidden
= Grammitis vestita Wall.; Numer. List [Wallich] no. 12 (1829), nomen
= Grammitis wightiana Wall.; Numer. List [Wallich] no. 2161 (1830), nom. inval.
= Gymnogramma vestita var. bipinnata Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16(199-201): 130 (1906)
= Gymnogramma vestita Wall. ex C.Presl; Tent. Pterid. 218 (1836)
= Gymnogramma vestita Wall. ex Hook.; Hooker´s Icon. Pl. 2: t. 115 (1837)
= Gymnopteris vestita (Wall. ex Hook.) Underw.; Bull. Torrey Bot. Club 29: 627 (1902)
= Hemionitis himalaica (Fraser-Jenk.) Christenh.; Global Fl. 4: 15 (2018), nom. superfl.
= Hemionitis vestita (Wall. ex Hook.) J.Sm.; Hist. Fil. 151 (1875)
= Neurogramma vestita (Wall. ex Hook.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 262 (1899)
= Notholaena himalaica Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 183 (1997), nom. nov.
= Paraceterach vestita (Wall. ex Hook.) R.M.Tryon; Amer. Fern J. 76(4): 186 (1986)
= Paragymnopteris vestita (Wall. ex C.Presl) K.H.Shing; Indian Fern J. 10(1-2): 230 (1993 publ. 1994)
= Sericopteris vestita (Hook.) Jing Zhao & X.M.Zhou; Phytotaxa 693(2): 169 (2025)
= Syngramma vestita (Wall. ex Hook.) T.Moore; Index Fil. [T. Moore] 60 (1857)
species :
Pellaea intermedia
Mett.; Linnaea 38: 84 (1869)
Name: Intermediate cliffbrake
Distribution: USA (W-Texas, S-New Mexico, Arizona); Mexico (Chihuahua, Coahuila, Nuevo Leon, Sonora, Tamaulipas, Zacatecas)
76 USA 77 USA 79 MEX
Synonyms: hidden
= Cassebeera intermedia (Mett.) Kaulf.; Amer. Midl. Naturalist 12: 281 (1931)
= Hemionitis horokana Christenh.; Global Fl. 4: 15 (2018), nom. nov.
= Pellaea flexuosa var. sagittata E.Fourn.; Mexic. Pl. 1: 141 (1872)
= Pellaea intermedia f. pubescens (Mett.) M.Broun; Index No. Amer. Ferns: 132 (1938)
= Pellaea intermedia var. pubescens Mett.; Linnaea 36: 85 (1869)
species :
Pellaea kitchingii
Baker; J. Bot. 18: 327 (1880)
Distribution: Madagascar
29 MDG
Remarks: of uncertain taxonomic position, unsampled in phylogenetic studies
Synonyms: hidden
= Allosorus kitchingii (Baker) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Doryopteris kitchingii (Baker) Bonap. ex C.Chr.; Index Filic., Suppl. (1906-1912) 16 (1913)
= Doryopteris latiloba C.Chr.; Arch. Bot. Bull. Mens. 2: 213 (1928)
= Hemionitis kitchingii (Baker) Christenh.; Global Fl. 4: 16 (2018)
= Hemionitis latiloba (C.Chr.) Christenh.; Global Fl. 4: 16 (2018)
species :
Pellaea lyngholmii
Windham; Contr. Univ. Michigan Herb. 19: 40 (1993)
Name: Lyngholm´s cliffbrake
Distribution: USA (C-Arizona)
76 USA
Conservation Status: IUCN V
Synonyms: hidden
= Hemionitis lyngholmii (Windham) Christenh.; Global Fl. 4: 17 (2018)
species :
Pellaea mucronata
(D.C.Eaton) D.C.Eaton; Rep. U.S. Mex. Bound., Bot. [Emory] 2(1): 233 (1859)
subspecies :
subsp. mucronata
Name: Bird´s Foot cliffbrake
Distribution: USA (Arizona, California, Nevada, Oregon); Mexico (Baja California Norte, Sonora); Isla Guadalupe
73 USA 76 USA 79 GLP MEX
Synonyms: hidden
= Allosorus mucronatus D.C.Eaton; Amer. J. Sci. Arts, Ser. 2, 22: 138 (1856)
= Allosorus ornithopus (Hook.) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Cassebeera ternifolia var. longimucronata (Hook.) Farw.; Amer. Midl. Naturalist 12: 282 (1931)
= Cassebeera ternifolia var. mucronata (D.C.Eaton) Farw.; Amer. Midl. Naturalist 12: 282 (1931)
= Cassebeera ternifolia var. ornithopus (Hook.) Farw.; Amer. Midl. Naturalist 12: 282 (1931)
= Cassebeera ternifolia var. stipitata Farw.; Amer. Midl. Naturalist 12: 282 (1931)
= Cheilanthes mucronata (D.C.Eaton) T.Moore; Index Fil. [T. Moore] 45, 248 (1861)
= Cheilanthes ornithopus (Hook.) T.Moore; Index Fil. [T. Moore] 250 (1861)
= Hemionitis mucronata (D.C.Eaton) Christenh.; Global Fl. 4: 18 (2018)
= Pellaea longimucronata Hook.; Sp. Fil. [W.J.Hooker] 2: 143, t. 115 A (1858)
= Pellaea longimucronata var. minor Hook.; Sp. Fil. [W.J.Hooker] 2: 143 (1858)
= Pellaea mucronata f. stipitata (Farw.) M.Broun; Index No. Amer. Ferns: 132 (1938)
= Pellaea ornithopus Hook.; Sp. Fil. [W.J.Hooker] 2: 143, t. 116 A (1858) (non Pellaea ornithopus Mett. ex Hook. & Baker)
= Pellaea wrightiana var. longimucronata (Hook.) Davenp.; Cat. Davenport Herb., Suppl. 46 (1883)
subspecies :
subsp. californica
(Lemmon) Windham; Contr. Univ. Michigan Herb. 19: 42 (1993)
Distribution: W-USA (C-California, S-California, Oregon)
73 USA 76 USA
Synonyms: hidden
= Cheilanthes horridula f. compacta (Clute) M.Broun; Index No. Amer. Ferns: 49 (1938)
= Pellaea aspera f. compacta (Davenp.) Clute; Fern Bull. 12: 44 (1904)
= Pellaea compacta (Davenp.) Maxon; Proc. Biol. Soc. Washington 80: 183 (1917)
= Pellaea mucronata var. californica (Lemmon) Munz & Johnst.; Amer. Fern J. 12: 106 (1922)
= Pellaea wrightiana var. californica Lemmon; Ferns Pacif. Coast 10 (1882)
= Pellaea wrightiana var. compacta Davenp.; Cat. Davenport Herb., Suppl. 46 (1883)
species :
Pellaea muelleri
(Hook.) A.R.Field; Austral. Syst. Bot. 33(1): 49 (2020)
Distribution: Australia (Queensland)
50 AUS
Synonyms: hidden
= Grammitis muelleri (Hook.) F.Muell.; Fragm. [Mueller] 5: 138 (1866)
= Gymnogramma muelleri Hook.; Sp. Fil. [W.J.Hooker] 6: 143, t. 295 (1864)
= Gymnopteris muelleri (Hook.) Underw.; Bull. Torrey Bot. Club 29: 627 (1902)
= Hemionitis hynreckii Christenh.; Global Fl. 4: 15 (2018), nom. nov.
= Hemionitis muelleri (Hook.) J.Sm.; Hist. Fil. 151 (1875)
= Neurogramma muelleri (Hook.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 262 (1899)
= Notholaena muelleri (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 75 (1947), isonym
= Notholaena muelleri (Hook.) Fraser-Jenk.; Fern Gaz. 18(5): 218 (2009), isonym
= Notholaena muelleri (Hook.) Mottet; Dict. Prat. Hort. 3: 484 (1895-1896)
= Paraceterach muelleri (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 75 (1947)
species :
Pellaea myrtillifolia
Mett.; Linnaea 36: 85 (1869)
Name: Coca, hierba coca, lendo del cerro [Chile]
Distribution: Chile (Atacama, Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins)
85 CLE
Conservation Status: IUCN R
Synonyms: hidden
= Allosorus myrtillifolius (Mett.) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Hemionitis myrtillifolia (Mett.) Christenh.; Global Fl. 4: 18 (2018)
= Pellaea andromedifolia Gay; Fl. Chil. [Gay] 6: 493 (1854), nom. illeg.
species :
Pellaea nana
(Hook.) Bostock; Fl. Australia 48: 708 (1998)
Distribution: Australia (Australian Capital Territory, Queensland, New South Wales, Victoria)
50 AUS
Synonyms: hidden
= Hemionitis nana (Hook.) Christenh.; Global Fl. 4: 18 (2018)
= Paraceterach nana (Hook.) Li Bing Zhang, X.M.Zhou & Jing Zhao; Phytotaxa 731(3): 229 (2025)
= Pellaea falcata var. nana Hook.; Sp. Fil. [W.J.Hooker] 2: 136 (1858)
= Platyloma falcatum var. nanum (Hook.) F.M.Bailey; Handb. Queensl. Ferns 36 (1874)
= Pteris falcata var. nana (Hook.) F.M.Bailey; Fern World Australia 45 (1881)
species :
Pellaea notabilis
Maxon; Contr. U.S. Natl. Herb. 10: 500 (1908)
Distribution: Mexico (Nuevo Leon, Tamaulipas)
79 MEX
Synonyms: hidden
= Hemionitis notabilis (Maxon) Christenh.; Global Fl. 4: 18 (2018)
species :
Pellaea oaxacana
Mickel & Beitel; Mem. New York Bot. Gard. 46 [Pterid. Fl. Oaxaca]: 271 (1988), pro hybr.
Distribution: Mexico (Colima, Jalisco, México State, Nayarit, Oaxaca); Guatemala
79 MEX 80 GUA
species :
Pellaea ovata
(Desv.) Weath.; Contr. Gray Herb. 114: 34 (1936)
Name: Ovate-leaved cliffbrake
Distribution: USA (S-Texas); Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México State, Michoacán, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tlaxcala, Veracruz, Zacatecas); Guatemala; Honduras; Nicaragua; Costa Rica; Hispaniola (Dominican Republic, Haiti); Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, Nariño, Putumayo, Santander, Valle); Venezuela (Aragua, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Tachira, Trujillo); Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Tarija, Santa Cruz); Brazil (São Paulo); N-Argentina (Catamarca, Jujuy, La Rioja, Salta, Tucuman)
77 USA 79 MEX 80 COS GUA HON NIC 81 HIS 82 CLM VEN 83 BOL ECU PER 84 BRA 85 ARG
Remarks: the Dominican Republic distribution might be a separate species
Synonyms: hidden
= Allosorus flexuosus (Kaulf. ex Schltdl. & Cham.) Kaulf.; Linnaea 13: 136 (1839)
= Hemionitis ovata (Desv.) Christenh.; Global Fl. 4: 18 (2018)
= Pellaea cordata var. flexuosa (Kaulf. ex Schltdl. & Cham.) Baker; Syn. Fil. [W.J.Hooker & Baker] 153 (1867)
= Pellaea flexuosa (Kaulf. ex Schltdl. & Cham.) Link; Fil. Spec. 60 (1841)
= Platyloma flexuosum (Kaulf. ex Schltdl. & Cham.) J.Sm.; Bot. Mag. 72 (Compan.): 21 (1846)
= Pteris flexuosa Kaulf. ex Schltdl. & Cham.; Linnaea 5: 614 (1830)
= Pteris ovata Desv.; Mém. Soc. Linn. Paris 6(2): 301 (1827)
species :
Pellaea paradoxa
(R.Br.) Hook.; Sp. Fil. [W.J.Hooker] 2: 135, t. 111 A (1858)
Distribution: Australia (E-Queensland, New South Wales); Lord Howe Isl.
50 AUS LHI
Synonyms: hidden
= Adiantum paradoxum R.Br.; Prodr. Fl. Nov. Holland. 155 (1810)
= Allosorus falcatus var. paradoxus (R.Br.) Luerss.; J. Mus. Godeffroy 3: (1873)
= Allosorus paradoxus (R.Br.) Kunze; Linnaea 23: 219 (1850)
= Hemionitis paradoxa (R.Br.) Christenh.; Global Fl. 4: 19 (2018)
= Paraceterach paradoxa (R.Br.) Li Bing Zhang, X.M.Zhou & Jing Zhao; Phytotaxa 731(3): 229 (2025)
= Pellaea cordata Fée; Mém. Foug. 5 (Gen. Filic.) 130 (1852)
= Pellaea paradoxa var. trichophora Domin; Biblioth. Bot. 20: 130 (1915)
= Platyloma brownii J.Sm.; J. Bot. [Hooker] 4: 160 (1841)
= Pteris cordata Sieber; Exsicc. (Fl. Mixta) no. 269 (1827), non Cav. 1802
= Pteris falcata var. paradoxa (R.Br.) F.M.Bailey; Syn. Queensl. Fl. 699 (1883)
= Pteris paradoxa (R.Br.) Desv.; Mém. Soc. Linn. Paris 6(2): 294 (1827)
species :
Pellaea pringlei
Davenp.; Gard. & Forest 4: 555, t. 88 (1891)
Distribution: Mexico (Colima, Guerrero, Jalisco, México State, Michoacán, Morelos, Nayarit, Sinaloa)
79 MEX
Synonyms: hidden
= Allosorus pringlei (Davenp.) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Hemionitis hysbaldii Christenh.; Global Fl. 4: 15 (2018), nom. nov.
= Windhamia pringlei (Davenp.) Li Bing Zhang, X.M.Zhou & Jing Zhao; Phytotaxa 731(3): 231 (2025)
species :
Pellaea prolifera
Schelpe; Bull. Jard. Bot. Natl. Belg. 38: 193, f. 1 (1968)
Distribution: Congo [Brazzaville]
23 CON
Remarks: of uncertain taxonomic position, unsampled in phylogenetic studies
Synonyms: hidden
= Hemionitis hykrionii Christenh.; Global Fl. 4: 15 (2018), nom. nov.
species :
Pellaea reynoldsii
(F.Muell.) A.R.Field; Austral. Syst. Bot. 33(1): 49 (2020)
Distribution: Australia (Western Australia, Northern Territory, South Australia)
50 AUS
Synonyms: hidden
= Grammitis reynoldsii (F.Muell.) Benth.; Fl. Austral. [Bentham] 7: 776 (1878)
= Gymnogramma reynoldsii (F.Muell.) J.M.Black; Fl. S. Austral. [J.M.Black] 1: 40 (1922)
= Gymnopteris reynoldsii (F.Muell.) C.Chr.; Index Filic. 341 (1905)
= Hemionitis reynoldsii (F.Muell.) Christenh.; Global Fl. 4: 20 (2018)
= Neurogramma reynoldsii (F.Muell.) Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 16: 262 (1899)
= Notholaena reynoldsii F.Muell.; Fragm. [Mueller] 8: 175 (1874)
= Paraceterach reynoldsii (F.Muell.) Tindale; Fl. S. Austral. [J.M.Black], ed. 3, 1: 53 (1978)
species :
Pellaea ribae
A.Mend. & Windham; Acta Bot. Mex. 57: 15 (2001)
Distribution: Mexico (San Luis Potosi)
79 MEX
species :
Pellaea rotundifolia
(G.Forst.) Hook.; Sp. Fil. [W.J.Hooker] 2: 136 (1858)
Name: New Zealand Cliff brake
Distribution: Norfolk Isl.; New Zealand (Three Kings Isl., North Isl., South Isl.); Kermadec Isl.; Chatham Isl.; Marshall Isl.. [I] (Majuro [I])
50 NFK 51 CTM KER NZL
Synonyms: hidden
= Allosorus rotundifolius (G.Forst.) Kunze; Linnaea 23: 219 (1850)
= Hemionitis rotundifolia (G.Forst.) Christenh.; Global Fl. 4: 20 (2018)
= Paraceterach rotundifolia (G.Forst.) Li Bing Zhang, X.M.Zhou & Jing Zhao; Phytotaxa 731(3): 229 (2025)
= Pellaea rotundifolia var. oblongifolia (G.Forst.) Hook.; Sp. Fil. [W.J.Hooker] 2: 136 (1858)
= Platyloma rotundifolium (G.Forst.) J.Sm.; J. Bot. [Hooker] 4: 160 (1841)
= Pteris falcata var. rotundifolia (G.Forst.) F.M.Bailey; Fern World Australia 45 (1881)
= Pteris rotundifolia G.Forst.; Fl. Ins. Austr. 79 (1786)
species :
Pellaea rufa
A.F.Tryon; Ann. Missouri Bot. Gard. 42: 101, t. 7 (1955)
Distribution: South Africa (W-Cape Prov.)
27 SAF
Remarks: of uncertain taxonomic position, unsampled in phylogenetic studies
Conservation Status: IUCN R
Synonyms: hidden
= Hemionitis yikka Christenh.; Global Fl. 4: 23 (2018), nom. nov.
species :
Pellaea sagittata
(Cav.) Link; Fil. Spec. 60 (1841)
Distribution: ?USA (?Texas); Mexico (Aguascalientes, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México State, Michoacán, Morelos, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sonora, Tlaxcala, Veracruz, Zacatecas); Guatemala; Costa Rica; Colombia (Cauca, Cundinamarca, Nariño, Santander, Valle); Ecuador; Peru; Bolivia (Chuquisaca, La Paz); Argentina (Salta)
79 MEX 80 COS GUA 82 CLM 83 BOL ECU PER 85 ARG
Synonyms: hidden
= Allosorus sagittatus (Cav.) C.Presl; Tent. Pterid. 151 (1836)
= Hemionitis emperatricella Christenh.; Global Fl. 4: 13 (2018), nom. nov. for Pteris sagittata Cav.
= Platyloma sagittatum (Cav.) J.Sm.; J. Bot. [Hooker] 4: 160 (1841)
= Pteris sagittata Cav.; Descr. Pl. [Cavanilles] 267 (1801)
= Windhamia sagittata (Cav.) Li Bing Zhang, X.M.Zhou & Jing Zhao; Phytotaxa 731(3): 231 (2025)
species :
Pellaea sargentii
(Christ) G.M.Zhang & Y.Yin; Bull. Bot. Res., Harbin 44(4): 499 (2024)
Distribution: China (W-Sichuan, NW-Yunnan); Tibet
36 CHI TIB
Synonyms: hidden
= Gymnopteris sargentii Christ; Bot. Gaz. 51: 355 (1911)
= Hemionitis sargentii (Christ) Christenh.; Global Fl. 4: 20 (2018)
= Notholaena sargentii (Christ) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 184 (1997)
= Paraceterach sargentii (Christ) R.M.Tryon; Amer. Fern J. 76(4): 186 (1986)
= Paragymnopteris sargentii (Christ) K.H.Shing; Indian Fern J. 10(1-2): 230 (1993 publ. 1994)
= Sericopteris sargentii (Christ) Jing Zhao & X.M.Zhou; Phytotaxa 693(2): 169 (2025)
species :
Pellaea striata
(Desv.) C.Chr.; Dansk Bot. Ark. 7: 117, t. 44, f. 12-15 (1932)
Distribution: Madagascar
29 MDG
Remarks: of uncertain taxonomic position, unsampled in phylogenetic studies
Synonyms: hidden
= Doryopteris madagascariensis Tardieu; Fl. Madagasc., Fam. 5, 1: 148 (1958) (synonymy fide Rakontondrainibe in sched. holotype)
= Hemionitis striata (Desv.) Christenh.; Global Fl. 4: 21 (2018)
= Hemionitis tenrecorum Christenh.; Global Fl. 4: 22 (2018), nom. nov. for Doryopteris madagascariensis Tardieu
= Pellaea sulcata Bonap.; Notes Ptéridol. 6: 63 (1917)
= Pteris striata Desv.; Mém. Soc. Linn. Paris 6(2): 300 (1827)
species :
Pellaea ternifolia
(Cav.) Link; Fil. Spec. 59 (1841)
subspecies :
subsp. ternifolia
Name: Ternate-leaved cliffbrake, Trans-Pecos cliffbrake (EN); Qusupi, qusupe, yuquelahuén [Chile]
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.); USA (SW-Texas: Big Bend NP); Mexico (Chiapas, Baja California, Sonora, Chihuahua, Durango, Sinaloa, Nayarit, Zacatecas, Aguascalientes, Guanajuato, Jalisco, Michoacán, Guerrero, México State, Queretaro, Morelos, Tlaxcala, Oaxaca, Puebla, Veracruz, Hidalgo, San Luis Potosi, Coahuila); Guatemala; Honduras; Nicaragua; Costa Rica; Hispaniola (Dominican Republic, Haiti); Colombia (Boyacá, Cundinamarca, Magdalena, Nariño, Santander, Valle); Venezuela (Merida); Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Oruro, Potosí, Santa Cruz, Tarija); Argentina (Buenos Aires, Catamarca, Chaco, Cordoba, Jujuy, La Pampa, La Rioja, Mendoza, Salta, Santiago del Estero, San Juan, San Luis, Tucuman); Chile (Arica y Parinacota, Tarapacá, Antofagasta, Valparaíso, O´Higgins, Biobío, Araucanía, Los Rios)
63 HAW 77 USA 79 MEX 80 COS GUA HON NIC 81 HIS 82 CLM VEN 83 BOL ECU PER 85 ARG CLE
Synonyms: hidden
= Adiantum ternatum Sessé & Moc.; Naturaleza (Mexico City), Ser. 2, 1(App.): 182 (1890), nom. illeg.
= Allosorus subverticillatus C.Presl; Tent. Pterid. 153 (1836)
= Allosorus ternifolius (Cav.) Kunze ex Klotzsch; Linnaea 20: 339 (1847)
= Cassebeera ternifolia (Cav.) Kaulf. ex Farw.; Amer. Midl. Naturalist 12: 281 (1931)
= Cheilanthes ternifolia (Cav.) T.Moore; Index Fil. [T. Moore] 255 (1861)
= Cheilanthes ternifolia var. lanuginosa (Fée) T.Moore; Index Fil. [T. Moore] 255 (1861)
= Cheilanthes weddelliana T.Moore; Index Fil. [T. Moore] 257 (1861)
= Hemionitis ternifolia (Cav.) Christenh.; Global Fl. 4: 22 (2018)
= Notholaena ternifolia (Cav.) Keyserl.; Polyp. Herb. Bunge 29 (1873)
= Pellaea lanuginosa Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 115 (1857)
= Pellaea ternifolia subsp. weddelliana (Fée) Arana; Fl. Vasc. Argent. 2: 315 (2016)
= Pellaea ternifolia var. petiolulata C.Chr.; Ark. Bot. 20A(7): 18 (1926)
= Pellaea weddelliana Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 74 (1857)
= Platyloma ternifolium (Cav.) J.Sm.; Bot. Mag. 72 (Compan.): 21 (1846)
= Pteris peruviana Poir.; Encycl. [J.Lamarck et al.] 5: 718 (1804)
= Pteris subverticillata Sw.; Syn. Fil. (Kiliae) [O.Swartz] 103 (1806)
= Pteris ternifolia Cav.; Descr. Pl. [Cavanilles] 266 (1801)
= Pteris triphylla Bertero; Mercurio Chileno 16: 748 (1829), nom. nud.
subspecies :
subsp. arizonica
Windham; Contr. Univ. Michigan Herb. 19: 42 (1993)
Distribution: USA (South Carolina, SE-Arizona, SW-Texas: Big Bend NP); Mexico (Chihuahua, Guerrero, Guanajuato, Jalisco, México State, Morelos, Nayarit, Oaxaca, Puebla, Sinaloa, San Luis Potosi, Sonora, Veracruz)
76 USA 77 USA 78 USA 79 MEX
subspecies :
subsp. brandegeei
(C.C.Hall) Mickel; Mem. New York Bot. Gard. 88 [Pterid. Mexico]: 451 (2004)
Distribution: Mexico (Baja California Sur)
79 MEX
Synonyms: hidden
= Pellaea brandegeei C.C.Hall; Amer. Fern J. 37: 111, t. 6 (1947)
species :
Pellaea timorensis
Alderw.; Bull. Dept. Agric. Indes Neerl. 18: 11 (1908)
Distribution: Thailand (SE-Thailand); Cambodia; Vietnam; Lesser Sunda Isl. (Timor)
41 CBD THA VIE 42 LSI
Remarks: of uncertain taxonomic position, unsampled in phylogenetic studies
Synonyms: hidden
= Hemionitis timorensis (Alderw.) Christenh.; Global Fl. 4: 22 (2018)
species :
Pellaea tripinnata
Baker; J. Linn. Soc., Bot. 25: 350 (1889)
Distribution: Madagascar
29 MDG
Remarks: of uncertain taxonomic position, unsampled in phylogenetic studies
Synonyms: hidden
= Allosorus tripinnatus (Baker) Kuntze; Revis. Gen. Pl. 2: 806 (1891)
= Hemionitis tripinnata (Baker) Christenh.; Global Fl. 4: 22 (2018)
= Pellaea ovalifolia Bonap.; Notes Ptéridol. 5: 62 (1917)
species :
Pellaea truncata
Goodd.; Muhlenbergia 8: 94 (1912)
Name: Spiny cliffbrake
Distribution: SW-USA (S-Nevada, SE-California, S-Utah, Arizona, S-New Mexico, W-Texas, C-Colorado); Mexico (Baja California Norte, Baja California Sur, Sonora)
73 USA 76 USA 77 USA 79 MEX
Synonyms: hidden
= Hemionitis truncata (Goodd.) Christenh.; Global Fl. 4: 22 (2018)
species :
Pellaea villosa
(Windham) Windham & Yatsk.; Novon 13: 361 (2003)
Distribution: USA (W-Texas: Davis Mts., Arizona); Mexico (Chihuahua, Coahuila, Guanajuato, Nuevo Leon, Durango, Hidalgo, México State, Michoacán, Puebla, San Luis Potosi, Sonora)
76 USA 77 USA 79 MEX
Synonyms: hidden
= Pellaea ternifolia subsp. villosa Windham; Contr. Univ. Michigan Herb. 19: 43 (1993)
species :
Pellaea wrightiana
Hook.; Sp. Fil. [W.J.Hooker] 2: 142 (1858)
Name: Wright´s cliffbrake
Distribution: USA (Arizona, Arkansas, Colorado, North Carolina, New Mexico, Oklahoma, South Carolina, Texas, Utah); Mexico (Baja California Norte, Baja California Sur, Sonora, Chihuahua, Coahuila)
73 USA 74 USA 76 USA 77 USA 78 USA 79 MEX
Synonyms: hidden
= Cheilanthes mucronata var. wrightiana (Hook.) T.Moore; Index Fil. [T. Moore] 248 (1861)
= Hemionitis wrightiana (Hook.) Christenh.; Global Fl. 4: 23 (2018)
= Pellaea ternifolia var. wrightiana (Hook.) A.F.Tryon; Ann. Missouri Bot. Gard. 44: 153 (1957)
species :
Pellaea zygophylla
(Riddell) P.J.Alexander; J. Semiarid Environm. 2: 2 (2021)
Distribution: USA (Texas); Mexico (Coahuila, Morelos, Nuevo Leon, San Luis Potosí, Tamaulipas)
77 USA 79 MEX
Remarks: a diploid entity formerly included in Pellaea ovata
Synonyms: hidden
= Pteris zygophylla Riddell; New Orleans Med. Surg. J. 9: 616 (1853)
species :
Pellaea ×glaciogena
W.H.Wagner; Madroño 30(2): 70 (1983)
Distribution: USA (C-California: Sa. Nevada)
76 USA
Remarks: Pellaea bridgesii x Pellaea mucronata
species :
Pellaea ×gooddingii
Windham; Amer. J. Bot. 109(5): 845 (2022)
Distribution: USA (Arizona)
76 USA
Remarks: Pellaea wightiana x ternifolia subsp. arizonica
species :
Pellaea ×wagneri
Windham; Amer. J. Bot. 109(5): 845 (2022)
Distribution: USA (Arizona)
76 USA
Remarks: Pellaea truncata x wightiana
