045.0000
Family
Woodsiaceae (A.Gray) Herter; Revista Sudamer. Bot. 9: 14 (1949) s. str.
1-2 genera, ca. 50 spp. Systematics see Lu et al. (2020).
Species listed in database: 57; hybrids: 5
= Protowoodsioideae Shmakov; Turczaninowia 4(1-2): 65 (2001), Subfamily
= Woodsieae A.Gray; Man. Bot. [A.Eaton], ed. 2: 588 (1818)
= Woodsiinae Hook.; Sp. Fil. [W.J.Hooker] 1: 58 (1844)
= Woodsioideae Fomin; Fl. Caucas. Crit. 3(1): 2, 5 (1911)
045.0001
Genus
Woodsia R.Br.; Prodr. Fl. Nov. Holland. 158 (1810)
Distribution: North-temperate holarctic, few China and Taiwan (subgen. Eriosorus). There are significant delimitation problems in Siberia. It has been proposed to split the genus in two (Woodsia and Physematium) with a hybrid genus Woodsimatium between them, but PPG-II is not following this split.
Species listed in database: 57; hybrids: 5
= Acrolysis Nakai; Bot. Mag. (Tokyo) 89: 176 (1925)
= Cheilanthopsis Hieron.; Notizbl. Bot. Gart. Berlin-Dahlem 7: 406 (1920)
= Eriosoriopsis (Kitag.) Ching & S.H.Wu; Fern Fam. Gen. China 402 (1991)
= Hymenocystis C.A.Mey.; Verz. Pfl. Casp. Meer [C.A. von Meyer] 229 (1831)
= Petrinia Hook.; Sp. Fil. [W.J.Hooker] 1: 62 (1844)
= Physematium Kaulf.; Flora 12: 341 (1829)
= Preslia Opiz ex Milde; Fil. Eur. 164 (1867)
= Protowoodsia Ching; Lingnan Sci. J. 21: 36 (1945)
= Trichocyclus Dulac; Fl. Hautes-Pyrenées 31 (1867) (nom. superfl.)
= Woodsia subgen.Eriosorus (Ching) Shmakov; Pteridol. New Millennium 59 (2003)
= Woodsiopsis Shmakov; Turczaninowia 18(2): 12 (2015)
= xWoodsimatium Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
Search for: Woodsia R.Br.
Nothing found!
species :
Woodsia acuminata
(Fomin) Sipliv.; Novosti Sist. Vyssh. Rast. 11: 327 (1974)
Distribution: Siberia (SW-Siberia, Altay-Yenisei region, Tunguska-Lena region, Baikal region); Russian Far East (Upper Amur, Russian Manchuria/Primorye)
30 SIB 31 RFE
Remarks: ilvensis agg.; often included in W. ilvensis
Synonyms: hidden
= Woodsia ilvensis var. acuminata Fomin; Visn. Kyyivs´koho Bot. Sadu 3: 3, t. 1 (1925)
species :
Woodsia alpina
(Bolton) Gray; Nat. Arr. Brit. Pl. 2: 17 (1821)
Name: Alpen-Wimperfarn (DE), Alpine Cliff Fern (EN)
Distribution: Iceland; England (N-Wales, S-Scotland); Norway; Sweden; Finland; Germany (Bayern); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Italy; Slovenia; Romania; C-European Russia; N-European Russia; W-European Russia; Ukraine; Siberia (Sverdlovsk); Kazakhstan; Kyrgyzstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (NE-Anatolia); NW-Iran; Tibet; Japan; Pakistan (Kurram, Swat, Hazara); N-India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand); Alaska; Greenland; Canada (British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); USA (Maine, Michigan, Minnesota, New York, Vermont)
10 FIN GRB ICE NOR SWE 11 AUT CZE GER HUN LIE POL SLO SWI 12 ADO FRA SPA 13 ITA ROM SLV 14 RUS UKR 30 SIB 32 KAZ KGZ 33 ARM AZE GEO NCS 34 IRN TUR 36 TIB 38 JAP 40 IND PAK 70 ASK CDN GNL 71 CDN 72 CDN 75 USA
Remarks: alpina agg.
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Acrostichum alpinum Bolton; Fil. Brit. 76, t. 42 (1790)
= Acrostichum hyperboreum Lilj.; Utkast Sv. Fl. 307 (1792)
= Ceterach alpinum (Bolton) DC.; Fl. Franç. [de Candolle & Lamarck], éd. 3, 567 (1805)
= Ceterach hyperboreum (Lilj.) Clairv.; Man. Herbor. Suisse 301 (1811)
= Polypodium alpinum (Bolton) With.; Bot. Arr. Veg. Gr. Brit., ed. 2, 3(2): 58 (1792), nom. illeg.
= Polypodium arvonicum With.; Bot. Arr. Veg. Gr. Brit., ed. 3, 3: 774 (1796)
= Polypodium hyperboreum (Lilj.) Sw.; J. Bot. [Schrader] 1800(2): 27 (1801)
= Trichocyclus hyperboreus (Lilj.) Dulac; Fl. Hautes-Pyrenées 31 (1867)
= Woodsia alpina subsp. bellii (G.Lawson) Á.Löve & D.Löve; Taxon 26(2-3): 326 (1977)
= Woodsia alpina Tausch; Flora 1839(2): 480 (1839), non (Bolton) Gray
= Woodsia alpina var. bellii (G.Lawson) G.Lawson; Canad. Naturalist Geol., n. s. 1: 4 (1864)
= Woodsia bellii (G.Lawson) A.E.Porsild; Rhodora 47: 147 (1945)
= Woodsia glabella var. bellii G.Lawson; Edinburgh New Philos. J., Ser. 2, 19: 281 (1864)
= Woodsia himalaica Ching & S.K.Wu; Fl. Xizang. 1: 189 (1983) (fide Fraser-Jenkins 1997)
= Woodsia hyperborea (Lilj.) R.Br.; Trans. Linn. Soc. London 11: 173 (1815)
= Woodsia hyperborea var. alpina (Bolton) Jess.; Deut. Excurs.-Fl. 620 (1879)
= Woodsia hyperborea var. arvonica (With.) Milde; Fil. Eur. 161 (1867)
= Woodsia ilvensis subsp. alpina (Bolton) Asch.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 1: 46 (1896)
= Woodsia ilvensis subsp. arvonica (With.) Milde; Sporenpfl. 74 (1865)
= Woodsia ilvensis subsp. hyperborea (Lilj.) Hartm.; Sv. Norsk Excurs.-Fl. 152 (1846)
= Woodsia ilvensis var. alpina (Bolton) Bab.; Man. Brit. Bot. [Babington], ed. 4: 420 (1847)
= Woodsia ilvensis var. alpina (Bolton) Watt.; Canad. Naturalist & Quart. J. Sci., Ser. 3: 160 (1866), isonym
= Woodsia pubescens Opiz; Kratos 2(1): 4 (1820), et Bohem. Phan. Crypt. Gew. (1833)
species :
Woodsia andersonii
(Bedd.) Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 45 (1905)
Distribution: China (Gansu, Shaanxi, Qinghai, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; India (Uttarakhand)
36 CHI TIB 38 TAI 40 BHU IND NEP
Remarks: subgen. Woodsia s. str.
Conservation Status: IUCN Ex/E (India)
Synonyms: hidden
= Gymnogramma andersonii Bedd.; Ferns Brit. India, t. 190 (1867)
= Eriosoriopsis andersonii (Bedd.) Shmakov; Turczaninowia 18(2): 15 (2015)
species :
Woodsia angolensis
Schelpe; Garcia de Orta, Ser. Bot. 3(1): 53 (1976)
Distribution: Angola; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal)
26 ANG 27 SAF
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium angolense (Schelpe) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Woodsia asiatica
Kiselev & Shmakov; Obzor Vidov. Sem. Woodsiaceae Evraz. 40 (1995)
Distribution: Siberia (SW-Siberia, Altay-Yenisei region, Baikal region, N-Yakutia); Russian Far East (Upper Amur, Russian Manchuria/Primorye, Kamchatka, Okhotsk / Magadan, Chukotka)
30 SIB 31 RFE
Remarks: ilvensis agg.
species :
Woodsia asplenioides
Rupr.; Dist. Crypt. Vasc. Ross. 55 (1845)
Distribution: Siberia (Tunguska-Lena region, Baikal region)
30 SIB
Remarks: glabella agg.
Synonyms: hidden
= Woodsia glabella subsp. asplenioides (Rupr.) Stepanov; Sibirsk. Biol. Zhurn. 1993(1): 47 (1993)
species :
Woodsia burgessiana
Gerrard ex Hook. & Baker; Syn. Fil. [W.J.Hooker & Baker] 48 (1866)
Distribution: Zimbabwe; South Africa (Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho; Madagascar
26 ZIM 27 LES SAF 29 MDG
Remarks: subgen. Physematium
Synonyms: hidden
= Diacalpe burgessiana (Gerrard ex Hook. & Baker) Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
= Physematium burgessianum (Gerrard ex Hook. & Baker) Shmakov; Turczaninowia 18(2): 12 (2015)
= Woodsia montevidensis var. burgessiana (Gerrard ex Hook. & Baker) Schelpe; J. S. African Bot. 35: 138 (1969)
species :
Woodsia calcarea
(Fomin) Shmakov; Obzor Vidov. Sem. Woodsiaceae Evraz. 29 (1995)
Distribution: Siberia (SW-Siberia, Altay-Yenisei region, Baikal region); Russian Far East (Upper Amur); Kazakhstan; Mongolia
30 SIB 31 RFE 32 KAZ 37 MON
Remarks: ilvensis agg.
Synonyms: hidden
= Woodsia ilvensis var. calcarea Fomin; Fl. Sibir. Orient. Extremi 5: 21 (1930)
species :
Woodsia canescens
(Kunze) Mett.; Ann. Sci. Nat., Bot., Sér. 5, 2: 249 (1864)
Distribution: Mexico (Ciudad de Mexico, México State, Oaxaca, Puebla, San Luis Potosi)
79 MEX
Synonyms: hidden
= Cheilanthes canescens (Kunze) Mett.; Abh. Senckenberg. Naturf. Ges. 3(1) [Aspl.]: 25, no. 18 (1859)
= Cheilanthes canescens Kunze; Linnaea 18: 143 (1839)
= Physematium canescens (Kunze) Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
species :
Woodsia cinnamomea
Christ; Bull. Acad. Int. Géogr. Bot., Sér. 3, 16: 122 (1906)
Distribution: China (SW-Sichuan)
36 CHI
Synonyms: hidden
= Eriosoriopsis cinnamomea (Christ) Shmakov; Turczaninowia 18(2): 15 (2015)
species :
Woodsia cochisensis
Windham; Contr. Univ. Michigan Herb. 19: 54 (1993)
Name: Cochise cliff fern (EN)
Distribution: USA (SE-Arizona, SW-New Mexico); Mexico (Chihuahua, Sonora, ?Baja California, Sur, Durango, Sinaloa)
76 USA 77 USA 79 MEX
Remarks: subgen. Physematium
Conservation Status: IUCN R
Synonyms: hidden
= Physematium cochisense (Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
= Woodsiopsis cochisensis (Windham) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Woodsia cycloloba
Hand.-Mazz.; Symb. Sin. 6: 19, t. 1, f. 5 (1929)
Distribution: China (Yunnan, Sichuan, Shaanxi); Tibet; ?Pakistan (?Swat); Nepal; Bhutan; India (Uttarakhand, Sikkim)
36 CHI TIB 40 BHU IND NEP
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Eriosoriopsis cycloloba (Hand.-Mazz.) Shmakov; Turczaninowia 18(2): 15 (2015)
species :
Woodsia cystopteroides
Windham & Mickel; Mem. New York Bot. Gard. 88 [Pterid. Mexico]: 688 (2004)
Distribution: Mexico (Chihuahua, Durango, Sinaloa, Sonora)
79 MEX
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium cystopteroides (Windham & Mickel) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
= Woodsiopsis cystopteroides (Windham & Mickel) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Woodsia elongata
Hook.; Sp. Fil. [W.J.Hooker] 1: 62, t. 21 c (1844)
Distribution: China (NW-Yunnan); Tibet; Nepal; Bhutan; India (Himachal Pradesh, Uttarakhand, Sikkim, West Bengal, Darjeeling, Arunachal Pradesh); Myanmar
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: subgen. Cheilanthopsis
Synonyms: hidden
= Cheilanthopsis elongata (Hook.) Copel.; Univ. Calif. Publ. Bot. 12: 395 (1931)
= Physematium elongatum (Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
species :
Woodsia fragilis
(Trevir.) T.Moore; Index Fil. [T. Moore] 283 (1861), nom. illeg.
Distribution: Northern Caucasus; Georgia [Caucasus] (Abkhazia, Samegrelo, Adzharia, Imereti, Shiga Kartli, Mtiuleti, Tush-Pshav-Khevsureti); Azerbaijan
33 AZE GEO NCS
Remarks: subgen. Physematium
Synonyms: hidden
= Cystopteris caucasica (C.A.Mey.) Fée; Mém. Foug. 5 (Gen. Filic.) 299 (1852)
= Diacalpe fragilis (Trevir.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1875)
= Dicksonia fragilis Trevir.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 7: 155, t. 3, f. 18-19 (1816)
= Hymenocystis caucasica C.A.Mey.; Verz. Pfl. Casp. Meer [C.A. von Meyer] 229 (1831)
= Hymenocystis fragilis (Trevir.) Askerov; Izv. Akad. Nauk Azerbajdzansk. S.S.R., Ser. Biol. Med. Nauk 1986(3): 52 (1986)
= Physematium fragile (Trevir.) Kunze; Analecta Pteridogr. 42 (1837)
= Woodsia caucasica (C.A.Mey.) J.Sm.; J. Bot. [Hooker] 4: 191 (1841)
species :
Woodsia glabella
R.Br.; Narr. Journey Polar Sea [Franklin] 754 (1823)
Name: Nordischer Wimperfarn (DE), Smooth Cliff Fern (EN)
Distribution: Svalbard; Norway; Sweden; Finland; Romania; C-European Russia; N-European Russia (incl. N-Urals); Siberia (W-Siberia/Siberian Urals, NW-Siberia, SW-Siberia, Altay-Yenisei region, Tunguska-Lena region, Baikal region, N-Yakutia); Russian Far East (Upper Amur, Russian Manchuria/Primorye, Sakhalin, Kamchatka, Okhotsk / Magadan, Chukotka); Northern Caucasus; Azerbaijan; China (Gansu, Hebei, Jilin, Qinghai, Xinjiang, Yunnan); Mongolia; North Korea; Japan; Nepal; NW-India (Uttarakhand, Himachal Pradesh, Jammu & Kashmir); Alaska; Greenland; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); USA (Massachusetts, Maine, Minnesota, New Hampshire, New York, Vermont)
10 FIN NOR SVA SWE 13 ROM 14 RUS 30 SIB 31 RFE 33 AZE NCS 36 CHI 37 MON 38 JAP NKO 40 IND NEP 70 ASK CDN GNL 71 CDN 72 CDN 75 USA
Remarks: subgen. Woodsia s. str., glabella agg.
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Woodsia alpina var. glabella (R.Br. ex Richardson) D.C.Eaton; Canad. Naturalist & Quart. J. Sci., Ser. 2: 90 (1865)
= Woodsia glabella f. rufescens Abrom.; Biblioth. Bot. 8: 73 (1897)
= Woodsia hyperborea var. glabella (R.Br. ex Richardson) Watt; Canad. Naturalist & Quart. J. Sci., Ser. 3: 160 (1866)
= Woodsia ilvensis var. glabella (R.Br.) Gelert; Fl. Arct. [Ostenfeld] 7 (1902)
= Woodsia lapponica Ångstr.; Handb. Skand. Fl., ed. 9: 267 (1864)
= Woodsia yazawae Makino; Bot. Mag. (Tokyo) 18: 132 (1904)
species :
Woodsia gorovoii
Krestsch. & Shmakov; Turczaninowia 10(1): 9 (2007)
Distribution: Russian Far East (Lower Amur)
31 RFE
Remarks: ilvensis agg.
species :
Woodsia guizhouensis
P.S.Wang, Q.Luo & Li Bing Zhang; Novon 22(2): 191 (2012)
Distribution: China (W-Guizhou)
36 CHI
Synonyms: hidden
= Eriosoriopsis guizhouensis (P.S.Wang, Q.Luo & Li Bing Zhang) Shmakov; Turczaninowia 18(2): 14 (2015)
species :
Woodsia hancockii
Baker; Ann. Bot. (Oxford) 5: 196 (1891)
Distribution: China (Hebei, Heilongjiang, Jilin, Nei Mongol, Shaanxi, Shanxi); North Korea; Japan; Nepal; India (Himachal Pradesh, Uttarakhand, Garhwal)
36 CHI 38 JAP NKO 40 IND NEP
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Woodsia gracillima C.Chr.; Acta Horti Gothob. 1: 42, t. 16 (1924)
= Woodsia tsurugisanensis Makino; Bot. Mag. (Tokyo) 28: 177 (1914)
species :
Woodsia hieronymi
(Ponce & Arana) comb.ined.
Distribution: Argentina (Catamarca, Jujuy, Salta, Tucumán)
85 ARG
Remarks: subgen. Physematium. To be expected in Bolivia
Synonyms: hidden
= Physematium hieronymi Ponce & Arana; Willdenowia 55(1): 34 (2025)
species :
Woodsia ilvensis
(L.) R.Br.; Trans. Linn. Soc. London 11: 173 (1815)
Name: Rostroter Wimperfarn, Südlicher Wimperfarn (DE), Rusty Cliff Fern (EN)
Distribution: Iceland; England (C-Scotland, NW-Wales, NW-England); Norway; Sweden; Finland; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, +Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Austria; Poland; Czech Republic; Slovakia; Hungary; France; Italy; Slovenia; Croatia; Romania; Estonia; C-European Russia; N-European Russia; W-European Russia; Ukraine; Crimea; Siberia (W-Siberia/Siberian Urals, NW-Siberia, SW-Siberia, Altay-Yenisei region, Tunguska-Lena region, Baikal region, N-Yakutia); Russian Far East (Upper Amur, Russian Manchuria/Primorye, Sakhalin, Kamchatka, Okhotsk / Magadan, Chukotka); Kazakhstan; Northern Caucasus; ?Georgia [Caucasus]; China (Hebei, Heilongjiang, Jilin, Nei Mongol, Xinjiang); Mongolia; Mongolia; North Korea; Japan; Alaska; Greenland; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); USA (Connecticut, Iowa, Illinois, Massachusetts, Maryland, Maine, Michigan, Minnesota, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Virginia, Vermont, Wisconsin, West Virginia)
10 FIN GRB ICE NOR SWE 11 AUT CZE GER HUN POL SLO SWI 12 FRA 13 CRO ITA ROM SLV 14 EST KRY RUS UKR 30 SIB 31 RFE 32 KAZ 33 NCS 36 CHI 37 MON 38 JAP NKO 70 ASK CDN GNL 71 CDN 72 CDN 74 USA 75 USA 78 USA
Remarks: subgen. Woodsia s. str., ilvensis agg.
Synonyms: hidden
= Acrostichum ilvense L.; Sp. Pl. [Linnaeus] 2: 1071 (1753)
= Acrostichum marantae Pall.; Reise Russ. Reich. 3(1): 293 (1776)
= Aspidium rufidulum (Michx.) Sw.; Syn. Fil. (Kiliae) [O.Swartz] 58 (1806)
= Athyrium rufidulum (Michx.) Eaton; Man. Bot. [A.Eaton] 122 (1817)
= Cincinalis setigera Desv.; Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 311 (1811)
= Lastrea rufidula C.Presl; Tent. Pterid. 76 (1836)
= Nephrodium rufidulum Michx.; Fl. Bor.-Amer. [Michaux] 2: 269 (1803)
= Notholaena rufidula (Michx.) Desv.; Mém. Soc. Linn. Paris 6(2): 221 (1827)
= Notholaena setigera Desv.; J. Bot. [Morot] 1: 93 (1813)
= Polypodium ilvense (L.) Sw.; J. Bot. [Schrader] 1800(2): 27 (1801)
= Polypodium ilvense (L.) Vill.; Hist. Pl. Dauphiné [Villars] 3: 848 (1789)
= Polypodium marantae (Pall.) Hoffm.; Deutschl. Fl. [Röhling] 2: 5 (1759)
= Preslia paleacea Opiz; Kratos 2(1): 5 (1820), pro syn.
= Woodsia frigida Gand.; Oesterr. Bot. Z. 31: 18 (1881)
= Woodsia hyperborea proles rufidula (Michx.) Rouy; Fl. France [Rouy & Foucaud] 14: 387 (1913)
= Woodsia hyperborea subsp. ilvensis (L.) Bonnier & Layens; 378 (1894), nom. illeg.
= Woodsia hyperborea subsp. ilvensis (L.) Hook.f.; Student´s Fl. Brit. Isl., ed. 1: 515 (1870), nom. illeg.
= Woodsia hyperborea var. ilvensis (L.) Jess.; Deut. Excurs.-Fl. 620 (1879), nom. illeg.
= Woodsia hyperborea var. rufidula (Michx.) W.D.J.Koch; Syn. Fl. Germ. Helv., ed. 2, 3: 975 (1845)
= Woodsia ilvensis subsp. rufidula (Michx.) Asch.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 1: 45 (1896)
= Woodsia ilvensis var. mangutica Galanin; Fl. Daurii 1: 43 (2008), contrary to Art. 36.1b
= Woodsia ilvensis var. rufidula (Michx.) Stefánsson; 4 (1901)
= Woodsia ilvensis var. rufidula (Michx.) Gelert; Fl. Arct. [Ostenfeld] 7 (1902)
= Woodsia paleacea Opiz; Kratos 2(1): 6 (1820)
= Woodsia raiana Newman; Hist. Brit. Ferns, ed. 2: 140 (1844)
= Woodsia rufidula (Michx.) L.C.Beck; Bot. North. Middle States [Beck] 458 (1833)
= Woodsia rupicola Hoffmanns.; Preiss-Verzeichn. Pflanzen 7: 60 (1832), nom. illeg.
= Woodsia uralensis Gand.; Oesterr. Bot. Z. 31: 18 (1881)
= Woodsia vestita Spreng.; Novi Provent. 44 (1819)
species :
Woodsia indusiosa
Christ; Notul. Syst. (Paris) 1: 44 (1909)
Distribution: China (NW-Yunnan, SW-Sichuan); ?Bhutan
36 CHI
Remarks: subgen. Cheilanthopsis. Not in India
Synonyms: hidden
= Cheilanthes straminea Brause; Hedwigia 54: 205, t. 4 H (1914)
= Cheilanthopsis indusiosa (Christ) Ching; Sinensia 8: 154 (1932) (indusiora)
= Cheilanthopsis straminea (Brause) Hieron. ex Copel.; Univ. Calif. Publ. Bot. 16: 95 (1929)
= Physematium indusiosum (Christ) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
species :
Woodsia intermedia
Rupr.; Beitr. Pflanzenk. Russ. Reiches 3: 54 (1845)
Distribution: Siberia (Siberian Urals)
30 SIB
Remarks: alpina agg.; often included in W. alpina but kept separate by Russian authors
species :
Woodsia jujuiensis
Copel.; Univ. Calif. Publ. Bot. 19: 297, t. 50 (1941)
Distribution: Peru (Cuzco); Bolivia (Cochabamba); Argentina (Catamarca, Jujuy, La Rioja, San Juan, Salta, Tucumán)
83 BOL PER 85 ARG
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium jujuiense (Copel.) Ponce & Arana; Willdenowia 55(1): 38 (2025)
= Woodsia montevidensis var. fuscipes Hieron.; Hedwigia 46: 322 (1907)
species :
Woodsia kangdingensis
H.S.Kung, Li Bing Zhang & X.S.Guo; Acta Bot. Yunnan. 17(4): 421 (1995)
Distribution: China (Sichuan)
36 CHI
Remarks: subgen. Cheilanthopsis
Synonyms: hidden
= Cheilanthopsis kangdingensis (H.S.Kung, Li Bing Zhang & X.S.Guo) Shmakov; Pteridol. New Millennium 64 (2003)
= Physematium kangdingense (H.S.Kung, Li Bing Zhang & X.S.Guo) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
species :
Woodsia kungiana
Li Bing Zhang, N.T.Lu & X.F.Gao; Phytotaxa 397(3): 253 (2019)
Distribution: China (Sichuan)
36 CHI
species :
Woodsia lanosa
Hook.; Syn. Fil. [W.J.Hooker & Baker] 47 (1866)
Distribution: China (Yunnan, Sichuan); Tibet; Nepal; Bhutan; India (Uttarakhand, Sikkim)
36 CHI TIB 40 BHU IND NEP
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Eriosoriopsis lanosa (Hook.) Shmakov; Turczaninowia 18(2): 15 (2015)
= Woodsia pellaeopsis Hand.-Mazz.; Symb. Sin. 6: 18, t. 1, f. 6 (1929)
species :
Woodsia macrochlaena
Mett. ex Kuhn; J. Bot. 6: 270 (1868)
Distribution: Russian Far East (Primorye); China (Hebei, Liaoning, Shandong); South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI 38 JAP KOR
Remarks: subgen. Woodsia s. str.
Synonyms: hidden
= Woodsia frondosa Christ; Repert. Spec. Nov. Regni Veg. 6: 12 (1908) (bona sp. fide Fl. Reipubl. Popularis Sin. 4(2))
= Woodsia japonica Makino; Bot. Mag. (Tokyo) 18: 134 (1904)
= Woodsia sinuata Makino; Bot. Mag. (Tokyo) 11: 64 (1897) (non Christ (1902))
species :
Woodsia macrospora
C.Chr. & Maxon; J. Wash. Acad. Sci. 17: 499 (1927)
Distribution: China (S-Gansu)
36 CHI
Remarks: sometimes included in W. cinnamomea, but kept separate by Flora of China
species :
Woodsia manchuriensis
Hook.; Sec. Cent. Ferns, t. 98 (1861)
Distribution: Russian Far East (Khabarovsk, Primorye); China (Anhui, Guizhou, Hebei, Heilongjiang, Henan, Jiangxi, Jilin, Liaoning, Shandong, Sichuan, Zhejiang); North Korea; South Korea (incl. Cheju Isl.); Japan
31 RFE 36 CHI 38 JAP KOR NKO
Remarks: subgen. Cheilanthopsis
Synonyms: hidden
= Diacalpe manchuriensis (Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1876)
= Physematium manchuriense (Hook.) Nakai; Bot. Mag. (Tokyo) 39: 176 (1925)
= Protowoodsia manchuriensis (Hook.) Ching; Lingnan Sci. J. 21: 37 (1945)
= Woodsia insularis Hance; Ann. Sci. Nat., Bot., Sér. 4, 15: 228 (1861)
species :
Woodsia mexicana
Fée; Mém. Foug. 7 (Icon. Esp. Nouv.): 66, t. 26, f. 3 (1857)
Distribution: Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chihuahua, Coahuila, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Jalisco, México State, Nuevo Leon, Queretaro, San Luis Potosi, Sinaloa, Sonora, Veracruz, Zacatecas)
79 MEX
Remarks: subgen. Physematium
Synonyms: hidden
= Woodsia oregana f. pusilla (E.Fourn.) B.Boivin; Bull. Soc. Bot. France 113: 408 (1967)
= Woodsia oregana var. mexicana (Fée) B.Boivin; Bull. Soc. Bot. France 113: 408 (1967)
= Woodsia pusilla E.Fourn.; Bull. Soc. Bot. France 27: 329 (1880)
= Woodsia pusilla var. mexicana (Fée) T.M.C.Taylor; Amer. Fern J. 37: 86 (1947)
= Woodsiopsis mexicana (Fée) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Woodsia mollis
(Kaulf.) J.Sm.; J. Bot. [Hooker] 4: 191 (1841)
Name: Mexican Cliff Fern
Distribution: Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México State, Michoacán, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala; El Salvador; Honduras; Nicaragua
79 MEX 80 ELS GUA HON NIC
Remarks: subgen. Physematium
Synonyms: hidden
= Diacalpe guatemalensis (Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1875)
= Lastrea bulbosa C.Presl; Tent. Pterid. 75 (1836)
= Physematium molle Kaulf.; Flora 1829(1): 341 (1829)
= Woodsia fragilis Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 5, 1: 280 (1849) (non T. Moore (1857))
= Woodsia guatemalensis Hook.; Sp. Fil. [W.J.Hooker] 1: 60, t. 21 (1844)
species :
Woodsia montevidensis
(Spreng.) Hieron.; Bot. Jahrb. Syst. 22: 363 (1896)
Distribution: Brazil (Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); Uruguay (Lavalleja, Maldonado, Minas, Rocha, Treinta y Tres); Argentina (Buenos Aires, Cordoba, La Rioja, Mendoza, Rio Negro, San Juan, San Luis, Tucuman)
84 BRA 85 ARG URU
Remarks: subgen. Physematium. Confirmed by Ponce et al. 2025 only for S-Brazil, Uruguay and Argentina. The formerly recorded distribution in the northern part of South America as well as the synonymy need further checking
Synonyms: hidden
= Cheilanthes andina Griseb.; Abh. Königl. Ges. Wiss. Göttingen 24: 342 (1879) (unconfirmed synonym)
= Cheilanthes crenata Kunze; Linnaea 9: 84 (1834) (unconfirmed synonym)
= Cystopteris vestita C.Presl; Tent. Pterid. 93 (1836), nom. nud. (unconfirmed synonym)
= Dicksonia montevidensis Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 122 (1827)
= Physematium euporolepis Trevis.; Nuovo Giorn. Bot. Ital. 7: 155 (1875) (unconfirmed synonym)
= Physematium incisum (Gillies ex Hook. & Grev.) C.Presl; Tent. Pterid. 66 (1836)
= Physematium montevidense (Spreng.) Shmakov; Turczaninowia 18(2): 12 (2015)
= Polypodium flabellatum Klotzsch; Linnaea 20: 382 (1847) (unconfirmed synonym)
= Woodsia crenata (Kunze) Hieron.; Bot. Jahrb. Syst. 34: 440 (1904) (unconfirmed synonym)
= Woodsia crenata var. pallidipes Hieron.; Bot. Jahrb. Syst. 34: 440 (1904) (unconfirmed synonym)
= Woodsia euporolepis (Trevis.) Trevis.; Rendiconti Reale Ist. Lombardo Sci. 2(9): 812 (1876) (unconfirmed synonym)
= Woodsia incisa Gillies ex Hook. & Grev.; Icon. Filic. 2: t. 191 (1830)
= Woodsia obtusa var. andina Hook.; Syn. Fil. [W.J.Hooker & Baker] 48 (1866) (unconfirmed synonym)
= Woodsia obtusa var. montevidensis (Spreng.) Hicken
species :
Woodsia neomexicana
Windham; Contr. Univ. Michigan Herb. 19: 52 (1993)
Name: New Mexican Cliff Fern (EN)
Distribution: USA (Arizona, New Mexico, S-Colorado, W-Texas, C-Oklahoma, S-Dakota); Mexico (Coahuila, Nuevo Leon, Puebla, Tamaulipas, Zacatecas)
73 USA 74 USA 76 USA 77 USA 79 MEX
Synonyms: hidden
= Physematium neomexicanum (Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
= Woodsiopsis neomexicana (Windham) Shmakov; Turczaninowia 18(2): 12 (2015)
species :
Woodsia nikkoensis
H.Ogura & Nakaike; J. Jap. Bot. 77(5): 296 (2002)
Distribution: Japan (Honshu)
38 JAP
species :
Woodsia oblonga
Ching & S.H.Wu; Fl. Tsinling. 2: 221 (1974)
Distribution: China (Hebei, Henan, Shandong)
36 CHI
species :
Woodsia obtusa
(Spreng.) Torr.; Rep. Bot. Dep. Surv. New York, no. 50, Assembly: 195 (1840)
subspecies :
subsp. obtusa
Name: Blunt-lobed Cliff Fern
Distribution: Canada (Ontario, Québec); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, Nebraska, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee, Texas, Virginia, Vermont, Wisconsin, West Virginia)
72 CDN 74 USA 75 USA 77 USA 78 USA
Remarks: subgen. Physematium
Synonyms: hidden
= Alsophila perriniana Spreng.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 10: 232 (1821)
= Aspidium obtusum (Spreng.) Sw.; Syn. Fil. (Kiliae) [O.Swartz] 420 (1806)
= Aspidium pennsylvanicum F.Dietr.; Nachtr. Vollst. Lex. Gärtn. 9: 374 (1838), nom. superfl.
= Cystopteris albescens Link; Fil. Spec. 47 (1841)
= Cystopteris obtusa (Spreng.) C.Presl; Tent. Pterid. 93 (1836)
= Cystopteris perriniana (Spreng.) Link; Hort. Berol. [Link] 2: 131 (1833)
= Physematium obtusum (Spreng.) Hook.; Fl. Bor.-Amer. [Hooker] 2: 259 (1840)
= Physematium perrinianum (Spreng.) C.Presl; Tent. Pterid. 66 (1836)
= Physematium vestitum (C.Presl) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 6 [Epim. Bot.]: 66 (1851)
= Polypodium obtusum Spreng.; Anleit. Kenntn. Gew. 3: 93 (1804)
= Woodsia obtusa f. darlingtonii (Stowell) M.Broun; Index No. Amer. Ferns: 166 (1938)
= Woodsia obtusa var. angusta Peck; Working List N. Amer. Pterid. 39 (1901)
= Woodsia obtusa var. darlingtonii Stowell; J. Trenton Nat. Hist. Soc. 1: 25 (1887)
= Woodsia perriniana (Spreng.) Hook. & Grev.; Icon. Filic. t. 68 (1828)
= Woodsiopsis obtusa (Spreng.) Shmakov; Turczaninowia 18(2): 13 (2015)
subspecies :
subsp. occidentalis
Windham; Contr. Univ. Michigan Herb. 19: 56 (1993)
Name: Western cliff fern (EN)
Distribution: USA (Arkansas, Kansas, Oklahoma, Texas)
74 USA 77 USA 78 USA
Synonyms: hidden
= Physematium obtusum subsp. occidentale (Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
= Woodsiopsis obtusa subsp. occidentalis (Windham) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Woodsia okamotoi
Tagawa; Acta Phytotax. Geobot. 7: 185 (1938)
Distribution: Taiwan
38 TAI
Conservation Status: CR in Taiwan
Synonyms: hidden
= Eriosoriopsis okamotoi (Tagawa) Shmakov; Turczaninowia 18(2): 15 (2015)
species :
Woodsia oregana
D.C.Eaton; Canad. Naturalist & Quart. J. Sci., Ser. 2: 90 (1865)
subspecies :
subsp. oregana
Name: Oregon cliff fern (EN)
Distribution: Canada (Alberta, British Columbia, Saskatchewan); USA (California, Idaho, Montana, Nevada, Oregon, Utah, Washington State, Wyoming)
71 CDN 73 USA 76 USA
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium oreganum (D.C.Eaton) Trevis.; Nuovo Giorn. Bot. Ital. 7: 162 (1875)
= Woodsiopsis oregana (D.C.Eaton) Shmakov; Turczaninowia 18(2): 13 (2015)
subspecies :
subsp. cathcartiana
(B.L.Rob.) Windham; Contr. Univ. Michigan Herb. 19: 58 (1993)
Name: Oregon cliff fern (EN)
Distribution: Canada (Manitoba, Ontario, Québec, Saskatchewan); USA (Arizona, California, Colorado, Iowa, Idaho, Kansas, Michigan, Minnesota, Montana, North Dakota, Nebraska, New Mexico, Nevada, New York, Oklahoma, South Dakota, Utah, Wisconsin, Wyoming)
71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA
Synonyms: hidden
= Physematium oreganum subsp. cathcartianum (B.L.Rob.) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
= Woodsia cathcartiana B.L.Rob.; Rhodora 10: 30 (1908)
= Woodsia oregana f. cathcartiana (B.L.Rob.) B.Boivin; Bull. Soc. Bot. France 113: 408 (1966)
= Woodsia oregana f. glandulosa T.M.C.Taylor; Amer. Fern J. 37: 85 (1947)
= Woodsia oregana var. cathcartiana (B.L.Rob.) C.V.Morton; Amer. Fern J. 40: 222 (1950)
= Woodsia pusilla var. cathcartiana (B.L.Rob.) T.M.C.Taylor; Amer. Fern J. 37: 86 (1947)
= Woodsia scopulina f. cathcartiana (B.L.Rob.) Clute; Our Ferns 133 (1901)
= Woodsia scopulina var. cathcartiana (B.L.Rob.) Clute; Fern Bull. 15: 127 (1908)
= Woodsiopsis oregana subsp. cathcartiana (B.L.Rob.) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Woodsia pallida
Copel.; Univ. Calif. Publ. Bot. 19: 2917, t. 51 (1941)
Distribution: Argentina (Catamarca, Jujuy, La Rioja, San Juan, Salta, Tucumán); Bolivia (Chuquisaca, Potosí); Chile (Antofagasta, Arica y Parinacota, Tarapacá)
83 BOL 85 ARG CLE
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium pallidum (Copel.) Ponce & Arana; Willdenowia 55(1): 45 (2025)
species :
Woodsia peruviana
Hook.; Sp. Fil. [W.J.Hooker] 1: 61, t. 21 (1844)
Distribution: Ecuador; Peru; Argentina (Catamarca, Jujuy, Salta, Tucumán)
83 ECU PER 85 ARG
Remarks: subgen. Physematium
Synonyms: hidden
= Diacalpe peruviana (Hook.) Trevis.; Nuovo Giorn. Bot. Ital. 7: 160 (1875)
= Physematium peruvianum (Hook.) Ponce & Arana; Willdenowia 55(1): 36 (2025)
species :
Woodsia phillipsii
Windham; Contr. Univ. Michigan Herb. 19: 52 (1993)
Name: Philipps´s Cliff Fern (EN)
Distribution: USA (S-Arizona, S-New Mexico, W-Texas); Mexico (Chihuahua, Coahuila, Durango, Jalisco, México State, Sonora)
76 USA 77 USA 79 MEX
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium phillipsii (Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
= Woodsiopsis phillipsii (Windham) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Woodsia pilosa
Ching; Fl. Tsinling. 2: 222 (1974)
Distribution: China (Henan)
36 CHI
species :
Woodsia pinnatifida
(Fomin) Shmakov; Obzor Vidov. Sem. Woodsiaceae Evraz. 55 (1995)
Distribution: Siberia (W-Siberia/Siberian Urals, SW-Siberia, Altay-Yenisei region, Tunguska-Lena region, Baikal region); Russian Far East (Upper Amur, Russian Manchuria/Primorye, Sakhalin, Okhotsk / Magadan, Chukotka); Mongolia
30 SIB 31 RFE 37 MON
Remarks: glabella agg.
Synonyms: hidden
= Woodsia glabella subsp. pinnatifida (Fomin) Stepanov; Sibirsk. Biol. Zhurn. 1993(1): 47 (1993)
= Woodsia glabella var. heterophylla Turcz. ex Fomin; Izv. Kievsk. Bot. Sada 3: 5 (1925)
= Woodsia glabella var. pinnatifida Fomin; Izv. Kievsk. Bot. Sada 3: 6 (1925)
= Woodsia heterophylla (Turcz. ex Fomin) Shmakov; Obzor Vidov. Sem. Woodsiaceae Evraz. 54 (1995) (fide Fraser-Jenkins et al. 2018)
species :
Woodsia plummerae
Lemmon; Bot. Gaz. 7: 6 (1882)
Name: Plummer´s Cliff Fern (EN)
Distribution: USA (Arizona, California, Colorado, New Mexico, Oklahoma, Texas); Mexico (Baja California Norte, Baja California Sur, Chihuahua, Jalisco, Sonora)
73 USA 74 USA 76 USA 77 USA 79 MEX
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium plummerae (Lemmon) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
= Woodsia obtusa var. glandulosa D.C.Eaton & Faxon; Bull. Torrey Bot. Club 9: 50 (1882)
= Woodsia obtusa var. plummerae (Lemmon) Maxon; Proc. U.S. Natl. Mus. 23: 644 (1901)
= Woodsia pusilla var. glandulosa (D.C.Eaton & Faxon) T.M.C.Taylor; Amer. Fern J. 37: 86 (1947)
= Woodsiopsis plummerae (Lemmon) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Woodsia polystichoides
D.C.Eaton; Proc. Amer. Acad. Arts 4: 110 (1858)
Distribution: Russian Far East (Lower Amur, Khabarovsk, Kuril Isl., Primorye, Sakhalin); China (Jilin, Liaoning, Henan, Beijing, Shandong, Shaanxi, Zhejiang, Anhui, Jiangxi, Hubei, Sichuan, Yunnan, etc.); North Korea; South Korea (incl. Cheju Isl.); Japan; Taiwan
31 RFE 36 CHI 38 JAP KOR NKO TAI
Remarks: subgen. Woodsia s. str.
Synonyms: hidden
= Physematium polystichoides (D.C.Eaton) Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
= Woodsia brandtii Franch. & Sav.; Enum. Pl. Jap. 2: 205, 616 (1877)
= Woodsia polystichoides f. veitchii (Hance) Kitag.; Neolin. Fl. Manshur. 41 (1979)
= Woodsia polystichoides var. incisa Ching & Y.T.Hsieh; Fl. Tsinling. 2: 222 (1974)
= Woodsia polystichoides var. nudiuscula Hook.; Gard. Ferns no. 8: sub t. 32 (1861)
= Woodsia polystichoides var. veitchii Hance; Ann. Sci. Nat., Bot., Sér. 4, 4: 229 (1861)
= Woodsia polystichoides var. veitchii Hook.; Gard. Ferns no. 8: t. 32, figs. 1, 2, 4--7 (1861)
species :
Woodsia pseudopolystichoides
(Fomin) Kiselev & Shmakov; Obzor Vidov. Sem. Woodsiaceae Evraz. 29 (1995)
Distribution: Siberia (Primorye)
30 SIB 31 RFE
Synonyms: hidden
= Woodsia ilvensis var. pseudopolystichoides Fomin; Fl. Sibir. Orient. Extremi 5: 21 (1930)
species :
Woodsia pubescens
Spreng.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 10: 233, t. 16, f. 5-7 (1821)
Distribution: Jamaica
81 JAM
Remarks: subgen. Physematium
Synonyms: hidden
= Physematium pubescens (Spreng.) Bevis; Hort. Brit. [Loudon], ed. 3: 656 (1839)
species :
Woodsia pulchella
Bertol.; Fl. Ital. Crypt. 1: 111 (1858)
Name: Zierlicher Wimperfarn (DE)
Distribution: Germany (Bayern); S-Switzerland; Austria; Spain; France; Italy; Slovenia; Romania
11 AUT GER SWI 12 FRA SPA 13 ITA ROM SLV
Synonyms: hidden
= Woodsia glabella subsp. pulchella (Bertol.) Á.Löve & D.Löve; Bot. Not. 114: 49 (1961)
species :
Woodsia rosthorniana
Diels; Bot. Jahrb. Syst. 29: 187 (1900)
Distribution: China (Hebei, Liaoning, Shaanxi, Sichuan, Yunnan); Tibet; Bhutan; NE-India (Arunachal Pradesh)
36 CHI TIB 40 BHU IND
Synonyms: hidden
= Eriosoriopsis rosthorniana (Diels) Ching & S.H.Wu; Fern Fam. Gen. China 402 (1991) (without bas. ref.)
= Woodsia delavayi Christ; Mém. Soc. Bot. France 1 [Hist. Acrostich.]: 45 (1905)
= Woodsia jeholensis Nakai & Kitag.; Rep. Exped. Manchoukuo, Sect. IV, 1 (Pl. Nov. Jehol.): 17, f. 1 (1934)
= Woodsia lanosa var. attenuata Christ; Bot. Gaz. 56: 338 (1913)
species :
Woodsia scopulina
D.C.Eaton; Canad. Naturalist & Quart. J. Sci., Ser. 2: 91 (1865)
subspecies :
subsp. scopulina
Name: Rocky Mountain woodsia (EN)
Distribution: Alaska; Canada (Alberta, British Columbia, Manitoba, Saskatchewan, Yukon); USA (California, Colorado, Idaho, Montana, Nevada, Oregon, Utah, Washington State, Wyoming)
70 ASK CDN 71 CDN 73 USA 76 USA
Synonyms: hidden
= Woodsia obtusa f. nana (Lemmon ex Gilbert) Clute; Fern Bull. 15: 126 (1908)
= Woodsia obtusa var. lyallii Hook.; Syn. Fil. [W.J.Hooker & Baker], ed. 1: 48 (1866)
= Woodsia obtusa var. nana Lemmon ex Gilbert; Working List N. Amer. Pterid. 21, 40 (1901)
= Woodsia oregana var. lyallii (Hook.) B.Boivin; Bull. Soc. Bot. France 113: 409 (1966)
= Physematium scopulinum Trevis.; Nuovo Giorn. Bot. Ital. 7: 161 (1875)
= Woodsia scopulina f. nana (Lemmon) M.Broun; Index No. Amer. Ferns: 167 (1938)
= Woodsiopsis scopulina (D.C.Eaton) Shmakov; Turczaninowia 18(2): 13 (2015)
subspecies :
subsp. appalachiana
(T.M.C.Taylor) Windham; Contr. Univ. Michigan Herb. 19: 58 (1993)
Name: Appalachian cliff fern (EN)
Distribution: USA (Arkansas, Georgia, Kentucky, North Carolina, Oklahoma, Tennessee, Virginia, West Virginia)
74 USA 75 USA 78 USA
Synonyms: hidden
= Physematium scopulinum subsp. appalachianum (T.M.C.Taylor) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
= Woodsia appalachiana T.M.C.Taylor; Amer. Fern J. 37: 88 (1947)
= Woodsia oregana var. appalachiana (T.M.C.Taylor) B.Boivin; Bull. Soc. Bot. France 113: 409 (1967)
= Woodsia scopulina var. appalachiana (T.M.C.Taylor) C.V.Morton; Amer. Fern J. 40: 224 (1950)
= Woodsiopsis appalachiana (T.M.C.Taylor) Shmakov; Turczaninowia 18(2): 12 (2015)
subspecies :
subsp. laurentiana
Windham; Contr. Univ. Michigan Herb. 19: 59 (1993)
Name: Laurentian cliff fern (EN)
Distribution: Canada (Alberta, British Columbia, Ontario, Québec); USA (Arizona, California, Colorado, Idaho, Minnesota, Montana, Nevada, Oregon, South Dakota, Utah, Washington State, Wyoming)
71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA
Synonyms: hidden
= Physematium scopulinum subsp. laurentianum (Windham) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
= Woodsiopsis scopulina subsp. laurentiana (Windham) Shmakov; Turczaninowia 18(2): 13 (2015)
species :
Woodsia shennongensis
D.S.Jiang & D.M.Chen; J. Hunan Agric. Univ. 26(2): 88 (2000)
Distribution: China (Hubei)
36 CHI
Remarks: provisionally accepted name
species :
Woodsia shensiensis
Ching; Sinensia 8: 141 (1932)
Distribution: China (Nei Mongol, Shaanxi, E-Sichuan)
36 CHI
Remarks: probably a synonym of W. hancockii fide Fraser-Jenkins et al. 2018
species :
Woodsia sinica
Ching; Sinensia 8: 145 (1932)
Distribution: China (Shanxi)
36 CHI
species :
Woodsia subcordata
Turcz.; Bull. Soc. Imp. Naturalistes Moscou 5: 206 (1832)
Distribution: Russian Far East (Upper Amur, Russian Manchuria/Primorye, Sakhalin); China (Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Shanxi); Mongolia; North Korea; South Korea; Japan
31 RFE 36 CHI 37 MON 38 JAP KOR NKO
Remarks: subgen. Woodsia s. str., subcordata agg.; W. kitadakensis and W. longifolia are often kept separate.
Synonyms: hidden
= Woodsia commixta Ching; Bull. Fan Mem. Inst. Biol., Bot., Ser. 2, 7: 314 (1949), nom. nov. for W. sinuata (Hook.) Christ
= Woodsia eriosora Christ; Repert. Spec. Nov. Regni Veg. 6: 12 (1908)
= Woodsia kitadakensis Ohwi; Bot. Mag. (Tokyo) 44: 572 (1930)
= Woodsia longifolia Tagawa; Acta Phytotax. Geobot. 5: 252 (1936) (fide Flora of Korea)
= Woodsia microsora Kodama; Icon. Pl. Koisikav. 8: 101, t. 196 (1927) (fide Flora of Korea)
= Woodsia pilosella Rupr.; Dist. Crypt. Vasc. Ross. 54 (1845)
= Woodsia polystichoides f. sinuata (Hook.) Kitag.; Neolin. Fl. Manshur. 41 (1979)
= Woodsia polystichoides var. sinuata Hook.; Gard. Ferns, t. 32, f. 3 (1862)
= Woodsia pseudoilvensis Tagawa; Acta Phytotax. Geobot. 5: 251 (1936)
= Woodsia saitoana Tagawa; Acta Phytotax. Geobot. 5: 250 (1936) (fide Flora of Korea)
= Woodsia sinuata (Hook.) Christ; Bull. Herb. Boissier, Sér. 2, 2: 830 (1902) (non Makino)
= Woodsia subcordata var. kitadakensis (Ohwi) Mitsuta; Acta Phytotax. Geobot. 30(4-6): 100 (1979)
= Woodsia subcordata var. longifolia (Tagawa) Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 85 (1991)
= Woodsia viridis Ching; Sinensia 3: 146 (1932)
species :
Woodsia taigischensis
(Stepanov) Kuznetsov; Sist. Zametki Mater. Gerb. Krylova Tomsk. Gosud. Univ. Kuybysheva 102: 8 (2010)
Distribution: Siberia (W-Sayan)
30 SIB
Remarks: ilvensis agg.
Synonyms: hidden
= Woodsia ilvensis var. taigischensis Stepanov; Sibirsk. Biol. Zhurn. 1993(1): 48 (1993)
species :
Woodsia taishanensis
F.Z.Li & C.K.Ni; Acta Phytotax. Sin. 20(3): 343 (1982)
Distribution: Russian Far East (Kuril Isl., Primorye, Sakhalin); China (Hebei, Heilongjiang, Henan, Jilin, Liaoning, Shandong, Shanxi); South Korea; Japan
31 RFE 36 CHI 38 JAP KOR
Synonyms: hidden
= Woodsia conmixta Ching; Bull. Fan Mem. Inst. Biol., Bot., Ser. 2, 1: 314 (1949)
= Woodsia intermedia Tagawa; Acta Phytotax. Geobot. 5: 250 (1936) (non Rupr. (1845))
= Woodsia subintermedia Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 83 (1991), nom. nov. superfl.
species :
Woodsia ×abbeae
Butters; Amer. Fern J. 31: 18 (1941)
Distribution: Canada (Ontario, Saskatchewan); USA (Michigan, Minnesota, Wisconsin)
71 CDN 72 CDN 74 USA 75 USA
Remarks: Woodsia ilvensis x Woodsia scopulina
Synonyms: hidden
= Woodsia confusa T.M.C.Taylor; Amer. Fern J. 37: 87 (1947)
= Woodsia oregana var. squammosa B.Boivin; Naturaliste Canad. 87: 49 (1960)
= xWoodsimatium ×abbeae (Butters) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
species :
Woodsia ×gracilis
(G.Lawson) Butters; Amer. Fern J. 31: 15 (1941)
Distribution: Canada (New Brunswick, Ontario, Québec); USA (Minnesota, Vermont)
72 CDN 75 USA
Remarks: Woodsia alpina x ilvensis
Synonyms: hidden
= Woodsia ilvensis var. gracilis G.Lawson; Edinburgh New Philos. J., Ser. 2, 19: 281 (1864)
species :
Woodsia ×kansana
R.E.Brooks; Amer. Fern J. 72(3): 81 (1982)
Distribution: USA (Kansas)
74 USA
Remarks: Woodsia obtusa x oregana subsp. oregana
Synonyms: hidden
= Physematium ×kansanum (R.E.Brooks) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1166 (2020)
species :
Woodsia ×maxonii
R.M.Tryon; Amer. Fern J. 38: 169 (1949)
Distribution: Canada (Ontario); USA (Minnesota)
72 CDN 75 USA
Remarks: Physematium oreganum subsp. cathcartianum x P. scopulinum subsp. laurentianum
Synonyms: hidden
= Physematium ×maxonii (R.M.Tryon) Li Bing Zhang, N.T.Lu & X.F.Gao; Taxon 68(6): 1167 (2020)
species :
Woodsia ×tryonis
B.Boivin; Naturaliste Canad. 93: 273 (1966)
Name: Tryon´s woodsia
Distribution: Canada (Ontario)
72 CDN
Remarks: Woodsia ilvensis x Woodsia glabella
