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014.0000 Family Schizaeaceae Kaulf.; Wesen Farrenkr. 119 (1827)
3(-4) genera, 35 - 40 spp. Phylogeny: Ke et al. (2022)
Species listed in database: 39; hybrids: 1
= Schizaeeae Kaulf.; Hort. Reg. Monac. [Martius & Schrank] 4 (1829)
= Schizaeineae Bessey; Bot. High Schools: 377 (1880)
= Schizaeoideae Lindl.; Veg. Kingd. 80 (1846)
014.0002 Genus Actinostachys R.Br. ex Hook. & Baker; Gen. Fil. [Hooker] t. 111 (1842)
Distribution: Tropics, many endemics in New Caledonia
Species listed in database: 17
= Actinostachys R.Br. ex Wall.; Numer. List [Wallich] no. 1 (1829), nomen
= Schizaea subgen.Actinostachys (Wall.) J.W.Sturm; Fl. Bras. [Martius] 1(2): 179 (1859)
species :
Actinostachys balansae (E.Fourn.) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 131 (1947)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Schizaea balansae E.Fourn.; Ann. Sci. Nat., Bot., Sér. 5, 18: 353 (1873)
species :
Actinostachys confusa (Selling) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 173 (1947)
Distribution: Madagascar; Seychelles
29 MDG SEY
Synonyms: hidden
= Schizaea confusa Selling; Svensk Bot. Tidskr. 41: 432, f. 1-14 (1947)
species :
Actinostachys digitata (L.) Wall.; Numer. List [Wallich] no. 1 (1829)
Distribution: Seychelles; China (Hainan, Guangdong, Yunnan); Ryukyu Isl.; Ogasawara Isl. (Chichijima, Anijima, Hahajima); Taiwan; India (Assam State, Karnataka, Kerala, Manipur, Meghalaya, Mizoram, Tripura); Sri Lanka; Bangladesh; Andaman Isl.; Nicobar Isl.; Myanmar; Thailand (SE-Thailand, peninsular Thailand); Cambodia; Vietnam; peninsular Malaysia (widespread); Singapore; Sumatra; Borneo (Sabah, Sarawak); Brunei; Sulawesi; Palawan; Philippines; Moluccas (Seram, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.); Solomon Isl. (Guadalcanal, Kolombangara, New Georgia, Nggatokae, Ranongga, Santa Ysabel, Vangunu); Australia (Western Australia, Northern Territory, Queensland); Tonga (Vava´u); Palau (Babeldaob); Fed. States of Micronesia (Yap, Pohnpei)
29 SEY 36 CHI 38 NNS TAI 40 BAN IND SRL 41 AND CBD MYA NCB THA VIE 42 BOR BRN MLY MOL NWG PHI PLW SNG SUL SUM 43 BIS SOL 50 AUS 60 TON 62 MIC PLU
Conservation Status: endangered (Singapore)
Synonyms: hidden
= Acrostichum digitatum L.; Sp. Pl. [Linnaeus] 2: 1068 (1753)
= Actinostachys boninensis Nakai; J. Jap. Bot. 13: 140, f. 2 (1937)
= Actinostachys digitata (L.) Wall. ex C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 130 (1948)
= Actinostachys digitata var. boninensis (Nakai) Tagawa; J. Jap. Bot. 30: 280 (1955)
= Belvisia digitata (L.) Mirb.; Hist. Nat. Vég. [Lamarck & Mirbel] 3: 473 (1802)
= Schizaea boninensis (Nakai) H.Ohba; Sci. Rep. Tohoku Imp. Univ., Ser. 4, Biol. 36: 85 (1971)
= Schizaea digitata (L.) Sw.; Syn. Fil. (Kiliae) [O.Swartz] 150, 380 (1896)
species :
Actinostachys inopinata (Selling) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 130 (1947)
Distribution: peninsular Malaysia (Kelantan, Pahang, Selangor); Sumatra; Borneo (Sabah); Philippines (Bohol); New Guinea (Irian Jaya); Palau (Ulebsechel, Urukthapel)
42 BOR MLY NWG PHI SUM 62 PLU
Synonyms: hidden
= Schizaea inopinata Selling; Svensk Bot. Tidskr. 40: 274, f. 1-7 (1946)
species :
Actinostachys intermedia (Mett.) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 131 (1947)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Schizaea intermedia Mett.; Ann. Sci. Nat., Bot., Sér. 4, 15: 86 (1861)
species :
Actinostachys laevigata (Mett.) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 130 (1947)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Schizaea laevigata Mett.; Ann. Sci. Nat., Bot., Sér. 4, 15: 85 (1861)
species :
Actinostachys macrofunda Bierh.; Amer. J. Bot. 58: 638 (1971)
Distribution: New Caledonia
60 NWC
species :
Actinostachys melanesica (Selling) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 131 (1947)
Distribution: New Caledonia; Tonga (Late Isl.); Fiji (Viti Levu, Vanua Levu, Ovalau)
60 FIJ NWC TON
Synonyms: hidden
= Schizaea melanesica Selling; Svensk Bot. Tidskr. 38: 208, f. 1, t. 1(3-5), 2(10-13) 3(15-18, 22-24) (1944)
species :
Actinostachys minuta Amoroso & Coritico; PhytoKeys 151: 60, figs. 1-2 (2020)
Distribution: Philippines (Mindanao)
42 PHI
species :
Actinostachys oligostachys Bierh.; Amer. J. Bot. 55: 92, f.14-29, 32, 33, 48, 56, 69 (1968)
Distribution: New Caledonia
60 NWC
species :
Actinostachys pennula (Sw.) Hook.; Gen. Fil. [Hooker] plate 111a (1842)
Name: Ray Spiked Fern
Distribution: SE-USA (Florida: Pinella Co., Palm Beach Co.); Mexico (Chiapas, Quintana Roo); Guatemala; Belize; Nicaragua; Costa Rica; Cuba; Puerto Rico; Guadeloupe; Trinidad; Colombia (Amazonas, Atlantico, Caquetá, Casanare, Guainía, Guaviare, Magdalena, Meta, Santander, Valle, Vaupés, Vichada); Venezuela (Amazonas, Apure, Bolívar); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Beni, La Paz); Brazil (Acre, Alagoas, Amazonas, Amapá, Bahia, Ceará, Brasilia Distrito Federal, Espírito Santo, Goiás, Maranhão, Mato Grosso, Pará, Paraíba, Pernambuco, Paraná, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Rondônia, Roraima, Santa Catarina, Sergipe, São Paulo, Tocantins); Paraguay (Canindeyú); Argentina (Misiones)
78 USA 79 MEX 80 BLZ COS GUA NIC 81 CUB GDL PUE TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA 85 ARG PAR
Synonyms: hidden
= Acrostichum pennula (Sw.) Poir.; Encycl. [J.Lamarck et al.] Suppl. 1: 125 (1810)
= Actinostachys germanii Fée; Mém. Foug. 11 (Hist. Foug. Antil.): 123, t. 29 C 3 (1866)
= Actinostachys orbicularis (Baker) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 132 (1947)
= Actinostachys penicillata Maxon; Proc. Biol. Soc. Washington 46: 139 (1933), nom. superfl.
= Actinostachys trilateralis (Schkuhr) J.Sm.; London J. Bot. 1: 202 (1842)
= Schizaea digitata var. orbicularis Baker; J. Bot. 19: 208 (1881)
= Schizaea germanii (Fée) Prantl; Unters. Morph. Gefäßkrypt. 2: 132 (1881)
= Schizaea penicillata Humb. & Bonpl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 86 (1810), nom. superfl.
= Schizaea pennula Sw.; Syn. Fil. (Kiliae) [O.Swartz] 150, 379 (1806)
= Schizaea trilateralis Schkuhr; Deutschl. Krypt. Gew. 1: 137 (1809)
species :
Actinostachys plana (E.Fourn.) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 131 (1947)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Schizaea plana E.Fourn.; Ann. Sci. Nat., Bot., Sér. 5, 18: 353 (1873)
species :
Actinostachys simplex Amoroso & Coritico; Phytotaxa 508(1): 102 (2021)
Distribution: Philippines
42 PHI
species :
Actinostachys spirophylla (W.Troll) C.F.Reed; Bol. Soc. Brot., Ser. 2, 1: 131 (1948)
Distribution: Moluccas (Ambon); Palau (Babeldaob); Fed. States of Micronesia (Pohnpei)
42 MOL 62 MIC PLU
Synonyms: hidden
= Actinostachys ponapensis (Hosok.) H.Itô; Bot. Mag. (Tokyo) 67: 214 (1954)
= Schizaea ponapensis Hosok.; Trans. Nat. Hist. Soc. Formosa 31: 39 (1941)
= Schizaea spirophylla Troll; Flora 128: 34-3, fig. (1933)
species :
Actinostachys subtrijuga (Mart.) C.Presl; Suppl. Tent. Pterid. 73 (1845)
Distribution: Trinidad; Colombia (Amazonas, Guainía, Vaupés); Venezuela (Amazonas, Apure, Bolivar, Yaracuy); Guyana; Surinam; French Guiana; Brazil (Acre, Amazonas, Bahia, Mato Grosso, Pará, Paraíba, Pernambuco, Paraná, Rio Grande do Sul, Rondônia, Roraima, Santa Catarina, São Paulo)
81 TRT 82 CLM FRG GUY SUR VEN 84 BRA
Synonyms: hidden
= Schizaea penicillata Humb. & Bonpl. ex Bory; Voy. Monde, Crypt. 1: 250, t. 27 (1828)
= Schizaea pennula var. subtrijuga (Mart.) Baker; Syn. Fil. [W.J.Hooker & Baker] 430 (1868)
= Schizaea subtrijuga Mart.; Icon. Pl. Crypt. 117 (1834)
species :
Actinostachys tenuis (E.Fourn.) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 131 (1947)
Distribution: New Caledonia
60 NWC
Synonyms: hidden
= Schizaea tenuis E.Fourn.; Ann. Sci. Nat., Bot., Sér. 5, 18: 353 (1873)
species :
Actinostachys wagneri (Selling) C.F.Reed; Bol. Soc. Brot., Ser. 2, 21: 131 (1947)
Distribution: Thailand; Vietnam; peninsular Malaysia (Perak, Kedah); +Singapore; Borneo; Moluccas (Ambon, Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (Admiralty Isl.); Solomon Isl. (Kolombangara); Australia (NE-Queensland)
41 THA VIE 42 BOR MLY MOL NWG SNG 43 BIS SOL 50 AUS
Synonyms: hidden
= Schizaea paucijuga Holttum; Gard. Bull. Singapore 11: 267 (1947)
= Schizaea wagneri Selling; Svensk Bot. Tidskr. 40: 278, f. 8-11 (1946)
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.02 (Feb. 1st, 2026)

Covering 358,774 accepted species,
8,847 hybrids, 48,003 accepted infraspecific taxa and 1,006,442 synonyms,
in total 1,422,066* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

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Arranged in linear phylogenetic sequence reflecting true relationships

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