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492.0000 Family Rubiaceae Juss.; Gen. Pl. [Jussieu] 196 (1789), nom. cons.
589 genera, ca. 14.200 spp. mostly tropics, comparably few temperate. Phylogeny: Razafimandimbison & Rydin (2024)
Species listed in database: 14370; hybrids: 47
= Aparinaceae Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 2: 3, 38 (1813-1829)
= Asperulaceae Cham. ex Spenn.; Handb. Angew. Bot. 2: 493 (1835)
= Catesbaeaceae Martinov; Tekhno-Bot. Slovar 111 (1820)
= Cephalanthaceae Raf.; Ann. Gén. Sci. Phys. 6: 86 (1820)
= Cinchonaceae Batsch; Tab. Affin. Regni Veg. 234 (1802)
= Coffeaceae Batsch; Tab. Affin. Regni Veg. 233 (1802)
= Coutareaceae Martinov; Tekhno-Bot. Slovar 168 (1820)
= Dialypetalanthaceae Rizzini & Occhioni; Lilloa 17: 253 (1948), nom. cons.
= Galiaceae Lindl.; Intr. Nat. Syst. Bot., ed. 2: 249 (1836)
= Gardeniaceae Dumort.; Anal. Fam. Pl. 29, 32 (1829)
= Guettardaceae Batsch; Tab. Affin. Regni Veg. 235 (1802)
= Hameliaceae Mart.; Consp. Regn. Veg. [Martius] 31 (1835)
= Hedyotidaceae Dumort.; Commentat. Bot. [Dumortier] 57 (1822)
= Henriqueziaceae Bremek.; Acta Bot. Neerl. 6 (Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 141): 371 (1957)
= Houstoniaceae Raf.; Good Book 21 (1840)
= Hydrophylacaceae Martinov; Tekhno-Bot. Slovar 318 (1820)
= Lippayaceae Meisn.; Pl. Vasc. Gen. [Meisner], Tab. Diagn. 156, Comm. 112 (1838)
= Lygodisodeaceae Bartl.; Ord. Nat. Pl. 123, 207 (1830)
= Naucleaceae Wernham; New Phytol. 11: 225 (1912)
= Nonateliaceae Martinov; Tekhno-Bot. Slovar 420 (1820)
= Operculariaceae Juss. ex Perleb; Vers. Arzneikr. Pfl. 207 (1818)
= Pagamaeaceae Martinov; Tekhno-Bot. Slovar 447 (1820)
= Psychotriaceae F.Rudolphi; Syst. Orb. Veg. 49 (1830)
= Randiaceae Martinov; Tekhno-Bot. Slovar 534 (1820)
= Sabiceaceae Martinov; Tekhno-Bot. Slovar 555 (1820)
= Spermacoceaceae Bercht. & J.Presl; Přir. Rostlin 256 (1820)
= Theligonaceae Dumort.; Anal. Fam. Pl. 15, 17 (1829), nom. cons.
492.1806 Genus Coccocypselum P.Browne; Civ. Nat. Hist. Jamaica 144 (1756), nom. cons.
Distribution: Tropical America, Caribbean
Species listed in database: 21
= Bellardia Schreb.; Gen. Pl., ed. 8[a], 2: 790 (1791), nom. illeg.
= Condalia Ruiz & Pav.; Fl. Peruv. Prodr. 11, t. 2 (1794)
= Lipostoma D.Don; Edinburgh New Philos. J. 11: 168 (1830)
= Sicelium P.Browne; Civ. Nat. Hist. Jamaica 144 (1756)
= Tontanea Aubl.; Hist. Pl. Guiane 1: 108 (1775)
species :
Coccocypselum anomalum K.Schum.; Fl. Bras. [Martius] 6(6): 311 (1889)
Distribution: NE-Brazil (Bahia); SE-Brazil (Espírito Santo, Rio de Janeiro)
84 BRA
species :
Coccocypselum aureum (Spreng.) Cham. & Schltdl.; Linnaea 4: 139 (1829)
Distribution: Costa Rica; Panama; Cuba; Colombia (Amazonas, Norte de Santander, Tolima, Valle); Venezuela (Anzoátegui, Bolívar); Guyana; French Guiana; Ecuador; Peru; Bolivia (Beni, La Paz, Santa Cruz); N-Brazil (Roraima, Tocantins); NE-Brazil (Pernambuco, Bahia); WC-Brazil (Mato Grosso, Goiás, Brasilia Distrito Federal); SE-Brazil (Minas Gerais, São Paulo, Rio de Janeiro)
80 COS PAN 81 CUB 82 CLM FRG GUY VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Coccocypselum aureum var. capitatum (Benth.) Steyerm.; Mem. New York Bot. Gard. 17: 307 (1967)
= Rondeletia capitata Benth.; J. Bot. [Hooker] 3: 217 (1841)
= Sabicea aurea (Spreng.) Steud.; Nomencl. Bot. [Steudel], ed. 2, 2: 712 (1840)
= Schwenkfelda aurea Spreng.; Neue Entdeck. Pflanzenk. [Sprengel] 1: 280 (1820)
species :
Coccocypselum bahiense C.B.Costa; Kew Bull. 62(4): 623 (2007)
Distribution: NE-Brazil (Bahia)
84 BRA
species :
Coccocypselum brevipetiolatum Steyerm.; Mem. New York Bot. Gard. 17: 305 (1967)
Distribution: Bolivia (La Paz)
83 BOL
species :
Coccocypselum capitatum (Graham) C.B.Costa & Mamede; Brittonia 58(2): 175 (2006)
Distribution: NE-Brazil (Bahia); SE-Brazil (Espírito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul)
84 BRA
Synonyms: hidden
= Acunaeanthus capitata Graham; Edinburgh New Philos. J. 1828: 389 (1828)
= Aeginetia capitata Graham; Edinburgh New Philos. J. 1828: 389 (1828)
= Coccocypselum campanuliflorum (Hook.) Cham. & Schltdl.; Linnaea 4: 140 (1829)
= Coccocypselum veronicoides C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 517 (1845)
= Coccocypselum violaceum Desf.; Tabl. École Bot., ed. 3 (Cat. Pl. Horti Paris.) 404 (1829)
= Hedyotis campanuliflora Hook.; Bot. Mag. 55: t. 2840 (1828)
= Lipostoma campanuliflorum (Hook.) D.Don ex Otto & A.Dietr.; Allg. Gartenzeitung [Otto & Dietrich] 1: 352 (1833)
= Lipostoma campanuliflorum (Hook.) Paxton; Pocket Bot. Dict. 187 (1840), isonym
= Lipostoma capitatum (Graham) D.Don; Edinburgh New Philos. J. 7: 168 (1830)
species :
Coccocypselum condalia Pers.; Syn. Pl. [Persoon] 1: 132 (1805)
Distribution: Belize; Costa Rica; Panama; Trinidad; Colombia (Caquetá); Venezuela (Apure, Bolívar); Guyana; Ecuador; Peru; Bolivia (Beni, Cochabamba, La Paz); N-Brazil; NE-Brazil (Bahia); WC-Brazil (Mato Grosso, Mato Grosso do Sul); SE-Brazil (Minas Gerais, Espírito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul); Paraguay (Alto Paraná)
80 BLZ COS PAN 81 TRT 82 CLM GUY VEN 83 BOL ECU PER 84 BRA 85 PAR
Synonyms: hidden
= Coccocypselum apurense Steyerm.; Ann. Missouri Bot. Gard. 74: 398 (1987)
= Coccocypselum brittonii Rusby; Bull. New York Bot. Gard. 4: 368 (1907)
= Coccocypselum condalia var. caaguazuense Hassl.; Repert. Spec. Nov. Regni Veg. 14: 169 (1915)
= Coccocypselum croatii Steyerm.; Ann. Missouri Bot. Gard. 74: 106 (1987)
= Coccocypselum decumbens K.Krause; Bot. Jahrb. Syst. 40: 323 (1908)
= Coccocypselum glabrum Britton; Bull. Torrey Bot. Club 18: 108 (1891), nom. illeg.
= Coccocypselum huberi Steyerm.; Ann. Missouri Bot. Gard. 74: 657 (1987)
= Coccocypselum trinitense Steyerm.; Mem. New York Bot. Gard. 17: 300 (1967)
= Coccocypselum umbellatum Poir.; Encycl. [J.Lamarck et al.], Suppl. 2: 307 (1811)
= Condalia repens Ruiz & Pav.; Fl. Peruv. [Ruiz & Pavon] 1: 54 (1798)
= Hedyotis cordata (Vell.) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 727 (1841)
= Manettia capitata Wernham; J. Bot. 57(Suppl.) 42 (1919)
= Manettia miersiana Wernham; J. Bot. 56(Suppl.): 14 (1918), nom. illeg.
= Oldenlandia cordata Vell.; Fl. Flumin. 53 (1825 publ. 1829)
species :
Coccocypselum cordifolium Nees & Mart.; Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12: 14 (1824)
Distribution: Mexico (Chiapas, Hidalgo, Oaxaca, Puebla, Tabasco, Veracruz); Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Guyana; French Guiana; N-Brazil (Pará); NE-Brazil (Maranhão, Pernambuco, Bahia, Alagoas); SE-Brazil (Minas Gerais, Espírito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul)
79 MEX 80 COS GUA HON NIC PAN 81 CUB 82 FRG GUY 84 BRA
Synonyms: hidden
= Coccocypselum nummulariifolium Cham. & Schltdl.; Linnaea 4: 145 (1829)
= Coccocypselum pleuropodum (Donn.Sm.) Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 4: 281 (1929)
= Coccocypselum rothschuhii Loes.; Bot. Jahrb. Syst. 60: 370 (1926)
= Geocardia pleuropoda (Donn.Sm.) Standl.; Contr. U.S. Natl. Herb. 17: 445 (1914)
= Geophila pleuropoda Donn.Sm.; Bot. Gaz. 52: 50 (1911)
= Tontanea pleuropoda (Donn.Sm.) Standl.; N. Amer. Fl. [Britton et al., eds.] 32: 148 (1921)
species :
Coccocypselum erythrocephalum Cham. & Schltdl.; Linnaea 4: 144 (1829)
Distribution: NE-Brazil (Bahia); WC-Brazil (Goiás); SE-Brazil (Minas Gerais, São Paulo, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Coccocypselum gardneri Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 11: 193 (1936)
= Coccocypselum montanum Mart.; Flora 24(2. Beibl.): 76 (1841)
species :
Coccocypselum geophiloides Wawra; Oesterr. Bot. Z. 31: 70 (1881)
Distribution: Peru; SE-Brazil (Minas Gerais, Espírito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul)
83 PER 84 BRA
Synonyms: hidden
= Coccocypselum krauseanum Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 8: 164 (1930)
= Coccocypselum sessiliflorum Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 11: 194 (1936)
species :
Coccocypselum glaberrimum Hadač; Folia Geobot. Phytotax. 5: 430 (1970)
Distribution: Cuba
81 CUB
species :
Coccocypselum glabrifolium Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 8: 165 (1930)
Distribution: N-Brazil (Pará); SE-Brazil (Minas Gerais, São Paulo); S-Brazil (Paraná)
84 BRA
Synonyms: hidden
= Coccocypselum hoehnei Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 176 (1940)
species :
Coccocypselum guianense (Aubl.) K.Schum.; Fl. Bras. [Martius] 6(6): 315 (1889)
Distribution: Mexico (Chiapas, Oaxaca, Puebla, Queretaro, San Luis Potosí, Tabasco, Tamaulipas, Veracruz); Guatemala; Belize; Honduras; Costa Rica; Cuba; Jamaica; Trinidad; Colombia (Antioquia, Chocó, Vichada); Venezuela (Amazonas, Anzoátegui, Bolívar, Delta Amacuro, Sucre); Guyana; Surinam; French Guiana; Ecuador; Peru; N-Brazil (Roraima, Amapá, Pará); NE-Brazil
79 MEX 80 BLZ COS GUA HON 81 CUB JAM TRT 82 CLM FRG GUY SUR VEN 83 ECU PER 84 BRA
Synonyms: hidden
= Bellardia guianensis (Aubl.) Forsyth f.; Bot. Nomencl. 106 (1794)
= Bellardia repens Willd.; Sp. Pl., ed. 4 [Willdenow] 1: 626 (1798)
= Bellardia tontanea Roem. & Schult.; Syst. Veg., ed. 15bis [Roemer & Schultes] 3: 205 (1818), nom. superfl.
= Coccocypselum glabrum Bartl. ex DC.; Prodr. [A.P. de Candolle] 4: 397 (1830)
= Coccocypselum guianense var. glabratum Proctor; J. Arnold Arbor. 63: 303 (1982)
= Coccocypselum guianense var. patens Steyerm.; Mem. New York Bot. Gard. 17: 303 (1967)
= Coccocypselum hirsutum var. glabrum (Bartl. ex DC.) L.O.Williams; Phytologia 25: 462 (1973)
= Coccocypselum tontanea Kunth; Nov. Gen. Sp. [H.B.K.] 3: 406 (1820)
= Tontanea glabra (Bartl. ex DC.) Standl.; J. Wash. Acad. Sci. 15: 104 (1925)
= Tontanea guianensis Aubl.; Hist. Pl. Guiane 108 (1775)
species :
Coccocypselum hasslerianum Chodat; Bull. Herb. Boissier, Sér. 2, 4: 169 (1904)
Distribution: Bolivia (La Paz); NE-Brazil (Bahia, Alagoas); WC-Brazil (Mato Grosso, Goiás, Brasilia Distrito Federal); SE-Brazil (Minas Gerais, Espírito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul); Paraguay (Alto Paraná, Cordillera); Argentina (Misiones)
83 BOL 84 BRA 85 ARG PAR
Synonyms: hidden
= Coccocypselum cordatum K.Krause; Anexos Mem. Inst. Butantan, Secç. Bot. 1(3): 13 (1922)
species :
Coccocypselum herbaceum P.Browne ex Aubl.; Hist. Pl. Guiane 1: 68 (1775)
Distribution: Mexico (Chiapas, Veracruz); Guatemala; Belize; El Salvador; ?Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola; Trinidad; Guyana
79 MEX 80 BLZ COS ELS GUA NIC PAN 81 CUB HIS JAM TRT 82 GUY
Synonyms: hidden
= Coccocypselum brownei Desv. ex Ham.; Prodr. Pl. Ind. Occid. [Hamilton] 17 (1825), nom. superfl.
= Coccocypselum glaberrimum (Borhidi & O.Muñiz) Borhidi; Acta Bot. Hung. 51: 275 (2009), nom. illeg.
= Coccocypselum pseudotontanea Griseb.; Fl. Brit. W. I. [Grisebach] 332 (1864)
= Coccocypselum pseudotontanea var. glaberrima Borhidi & O.Muñiz; Növenyrendsz. Novényföldr. Tansz., Eötvös Loránd Tudományegyet. Budapest 5: 40 (1977)
= Coccocypselum repens Sw.; Prodr. Veg. Ind. Occ. [O.P.Swartz] 31 (1788), nom. superfl.
= Coccocypselum tenue Urb.; Repert. Spec. Nov. Regni Veg. 13: 479 (1915)
= Tontanea herbacea Standl.; N. Amer. Fl. [Britton et al., eds.] 32(2): 147 (1921), nom. illeg.
= Tontanea repens (Sw.) Pers.; Syn. Pl. [Persoon] 1: 134 (1805)
= Tontanea tenuis (Urb.) Standl.; N. Amer. Fl. [Britton et al., eds.] 32(2): 146 (1921)
species :
Coccocypselum hirsutum Bartl. ex DC.; Prodr. [A.P. de Candolle] 4: 396 (1830)
Name: Yerba de guava
Distribution: USA [I] (Florida [I]); Mexico (Chiapas, Guerrero, Hidalgo, Jalisco, Nayarit, Oaxaca, Puebla, San Luis Potosí, Tabasco, Veracruz); Guatemala; Belize; Nicaragua; Costa Rica; Panama; Trinidad; Colombia (Amazonas, Antioquia, Boyacá, Caquetá, Chocó, Cundinamarca, Huila, Magdalena, Meta, Nariño, Risaralda, Santander, Tolima, Valle, Vichada); Venezuela (Bolívar); Guyana; French Guiana; Peru; Bolivia (La Paz, Santa Cruz); N-Brazil (Amazonas); NE-Brazil (Bahia); WC-Brazil (Mato Grosso, Goiás, Brasilia Distrito Federal); SE-Brazil (Minas Gerais)
78 usa 79 MEX 80 BLZ COS GUA NIC PAN 81 TRT 82 CLM FRG GUY VEN 83 BOL PER 84 BRA
Synonyms: hidden
= Coccocypselum ciliatum Cham. & Schltdl.; Linnaea 6: 414 (1831)
= Coccocypselum discolor Van Houtte ex Kunze; Linnaea 24: 192 (1851)
= Coccocypselum hirsutum var. uniflorum Hadač; Folia Geobot. Phytotax. 5: 431 (1970)
= Petesia repens Sessé & Moc.; Fl. Mexic. 21 (1893)
= Tontanea hirsuta (Bartl. ex DC.) Standl.; N. Amer. Fl. [Britton et al., eds.] 32: 147 (1921)
species :
Coccocypselum hispidulum (Standl.) Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 4: 281 (1929)
Distribution: Mexico (Chiapas); Nicaragua; Costa Rica; Panama; Cuba; Colombia (Antioquia, Chocó, Nariño, Valle)
79 MEX 80 COS NIC PAN 81 CUB 82 CLM
Synonyms: hidden
= Coccocypselum hispidum Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 7: 47 (1930)
= Tontanea hispidula Standl.; N. Amer. Fl. [Britton et al., eds.] 32(2): 147 (1921)
species :
Coccocypselum lanceolatum (Ruiz & Pav.) Pers.; Syn. Pl. [Persoon] 1: 132 (1805)
Distribution: Guatemala; Costa Rica; Panama; Cuba; Colombia (Antioquia, Boyacá, Caquetá, Cauca, Cesar, Chocó, Cundinamarca, Huila, Magdalena, Nariño, Norte de Santander, Quindío, Risaralda, Santander, Tolima, Valle); Venezuela (Amazonas, Anzoátegui, Aragua, Bolívar, Cojedes, Venezuela Distrito Federal, Falcón, Guárico, Lara, Mérida, Miranda, Monagas, Táchira, Trujillo, Yaracuy, Zulia); Guyana; Ecuador; Peru; Bolivia (Chuquisaca, La Paz); NE-Brazil (Ceará, Bahia); WC-Brazil (Mato Grosso, Goiás, Brasilia Distrito Federal); SE-Brazil (Minas Gerais, Espírito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Rio Grande do Sul, Santa Catarina); Paraguay (Alto Paraná, Amambay, Caazapa, Canindeyu, Cordillera, Guaira, Paraguari); Argentina (Misiones, Salta)
80 COS GUA PAN 81 CUB 82 CLM GUY VEN 83 BOL ECU PER 84 BRA 85 ARG PAR
Synonyms: hidden
= Bellardia mollis Willd. ex DC.; Prodr. [A.P. de Candolle] 4: 397 (1830)
= Coccocypselum canescens Willd. ex Schult. & Schult.; Mant. 3 [Schultes & Schultes f.] 130 (1827)
= Coccocypselum crassifolium Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 175 (1940)
= Coccocypselum dichroolasium Mart.; Flora 24(2. Beibl.): 76 (1841)
= Condalia lanceolata Ruiz & Pav.; Fl. Peruv. [Ruiz & Pavon] 1: 54 (1798)
= Schwenkfelda aspera Spreng.; Neue Entdeck. Pflanzenk. [Sprengel] 1: 281 (1820)
= Tontanea canescens (Willd. ex Schult. & Schult.f.) Standl.; N. Amer. Fl. [Britton et al., eds.] 32: 146 (1921)
species :
Coccocypselum lymansmithii Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 8: 165 (1930)
Distribution: WC-Brazil (Brasilia Distrito Federal); SE-Brazil (Minas Gerais, Espírito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Rio Grande do Sul, Santa Catarina); Paraguay (Amambay)
84 BRA 85 PAR
Synonyms: hidden
= Coccocypselum monanthos L.B.Sm. & Downs; Sellowia 7: 89 (1956)
= Coccocypselum pumilo Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 8: 346 (1931)
= Coccocypselum uniflorum Hassl.; Repert. Spec. Nov. Regni Veg. 14: 168 (1915), nom. superfl.
species :
Coccocypselum oblongatum Urb.; Symb. Antill. [Urban] 9: 523 (1928)
Distribution: Cuba
81 CUB
Remarks: provisionally accepted name
species :
Coccocypselum pedunculare Cham. & Schltdl.; Linnaea 4: 142 (1829)
Distribution: NE-Brazil; SE-Brazil (Minas Gerais)
84 BRA
species :
Coccocypselum pulchellum Cham.; Linnaea 9: 238 (1834)
Distribution: S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul)
84 BRA
Synonyms: hidden
= Coccocypselum reitzii L.B.Sm. & Downs; Sellowia 7: 89 (1956)
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life