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001.0000 Family Lycopodiaceae P.Beauv. ex Mirb.; Hist. Nat. Vég. [Lamarck & Mirbel] 4: 293 (1802)
17 genera, ca. 470 spp. Phylogeny: Chen et al. (2021).
Species listed in database: 477; hybrids: 21
= Huperziaceae Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 66: 236 (1962), nom. nov.
= Urostachyaceae Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 58 (1944) (nom. illeg.)
= Phylloglossaceae Kunze; Bot. Zeitung (Berlin) 1: 722 (1843)
= Lycopodiellaceae Val.N.Tikhom.; Novosti Sist. Vyssh. Rast. 49: 149 (2018)
001.3002 Genus Dendrolycopodium A.Haines; Fam. Huperziac. Lycopodiac. New England 84 (2003)
Distribution: North America, E-Asia. Frequently re-included in Lycopodium s. lat. but accepted by PPG-I
Species listed in database: 5
species :
Dendrolycopodium dendroideum (Michx.) A.Haines; Fam. Huperziac. Lycopodiac. New England 84 (2003)
Name: Round-branch Ground-Pine, Tree groundpine
Distribution: Alaska; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); St. Pierre et Miquelon; USA (Alabama, Connecticut, Iowa, Idaho, Illinois, Indiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Montana, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Dakota, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia, Wyoming)
70 ASK CDN 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 78 USA
Synonyms: hidden
= Lepidotis dendroidea (Michx.) P.Beauv.; Prodr. Aethéogam. 108 (1805)
= Lycopodium dendroideum Michx.; Fl. Bor.-Amer. [Michaux] 2: 282 (1803)
= Lycopodium obscurum f. dendroideum (Michx.) H.L.Blomq. & Correll; J. Elisha Mitchell Sci. Soc. 56: 101 (1940)
= Lycopodium obscurum var. dendroideum (Michx.) D.C.Eaton; Manual [A.Gray], ed. 6: 696 (1890)
= Lycopodium obscurum var. hybridum Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 91 (1916 publ. 1917)
species :
Dendrolycopodium hickeyi (W.H.Wagner, Beitel & R.C.Moran) A.Haines; Fam. Huperziac. Lycopodiac. New England 84 (2003)
Name: Pennsylvania clubmoss
Distribution: Canada (New Brunswick, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); USA (Connecticut, Delaware, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia)
71 CDN 72 CDN 73 USA 74 USA 75 USA 78 USA
Synonyms: hidden
= Lycopodium hickeyi W.H.Wagner, Beitel & R.C.Moran; Amer. Fern J. 79: 121 (1989), nom. nov.
= Lycopodium obscurum var. isophyllum Hickey; Amer. Fern J. 67: 47 (1977)
species :
Dendrolycopodium juniperoideum (Sw.) A.Haines; Fam. Huperziac. Lycopodiac. New England 84 (2003)
Distribution: Siberia (Tunguska-Lena region, Baikal region); Russian Far East (Upper Amur, Russian Manchuria/Primorye, Sakhalin, Kamchatka, Okhotsk / Magadan); Japan; Taiwan
30 SIB 31 RFE 38 JAP TAI
Remarks: Not mentioned in Flora of China but recorded for Taiwan by Knapp (Ferns of Taiwan, 2011 ff.)
Synonyms: hidden
= Lycopodium juniperinum Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 15 (1827)
= Lycopodium juniperoideum Sw.; Syn. Fil. (Kiliae) [O.Swartz] 401 (1806)
= Lycopodium obscurum f. juniperoideum (Sw.) Takeda; Bot. Mag. (Tokyo) 23(274): 213 (1909)
species :
Dendrolycopodium obscurum (L.) A.Haines; Fam. Huperziac. Lycopodiac. New England 84 (2003)
Name: Ground Pine, Eastern Tree Clubmoss, Princess Pine
Distribution: Russian Far East (Upper Amur, Russian Manchuria/Primorye, Sakhalin); NE-China; North Korea; South Korea; Japan; Alaska; Canada (New Brunswick, Nova Scotia, Ontario, Québec); USA (Alabama, Connecticut, District of Columbia, Delaware, Georgia, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia)
31 RFE 36 CHI 38 JAP KOR NKO 70 ASK 72 CDN 74 USA 75 USA 78 USA
Synonyms: hidden
= Lycopodium dendroideum var. obscurum (L.) Torr. ex Beck; Bot. North. Middle States [Beck] 460 (1833)
= Lycopodium obscurum subvar. breve Farw.; Amer. Fern J. 27: 19 (1937)
= Lycopodium obscurum subvar. brevispicatum Farw.; Amer. Fern J. 27: 19 (1937)
= Lycopodium obscurum subvar. foliaceum Farw.; Amer. Fern J. 27: 19 (1937)
= Lycopodium obscurum subvar. parvispicatum Farw.; Amer. Fern J. 27: 19 (1937)
= Lycopodium obscurum subvar. proliferum (Vict.) Farw.; Amer. Fern J. 27: 19 (1937)
= Lycopodium obscurum f. breve (Farw.) M.Broun; Index No. Amer. Ferns: 110 (1938)
= Lycopodium obscurum f. brevispicatum (Farw.) M.Broun; Index No. Amer. Ferns: 111 (1938)
= Lycopodium obscurum f. exsertum Vict.; Contr. Lab. Bot. Univ. Montréal 3: 32 (1925)
= Lycopodium obscurum f. exsertum-furcatum Vict.; Contr. Lab. Bot. Univ. Montréal 3: 33 (1925)
= Lycopodium obscurum f. foliaceum (Farw.) M.Broun; Index No. Amer. Ferns: 110 (1938)
= Lycopodium obscurum f. monostachyon Vict.; Contr. Lab. Bot. Univ. Montréal 3: 33 (1925)
= Lycopodium obscurum f. parvispicatum (Farw.) M.Broun; Index No. Amer. Ferns: 110 (1938)
= Lycopodium obscurum f. proliferum Vict.; Contr. Lab. Bot. Univ. Montréal 3: 32 (1925)
= Lycopodium obscurum L.; Sp. Pl. [Linnaeus] 2: 1102 (1753)
species :
Dendrolycopodium verticale (Li Bing Zhang) Li Bing Zhang & X.M.Zhou; Phytotaxa 295(2): 199 (2017)
Distribution: China (Anhui, Chongqing, Guizhou, Hubei, Hunan, Jiangxi, Shaanxi, Shanxi, Sichuan, NE-Yunnan, Zhejiang); E-Tibet; ?Taiwan; Bhutan; NE-India (Arunachal Pradesh)
36 CHI TIB 40 BHU IND
Synonyms: hidden
= Lycopodium dendroideum f. strictum Milde; Fil. Eur. 254 (1867)
= Lycopodium obscurum f. strictum (Milde) Nakai ex H.Hara; Bot. Mag. (Tokyo) 48(574): 706 (1934)
= Lycopodium verticale Li Bing Zhang; Fl. China 2-3: 29 (2013)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.12 (Dec. 21st, 2025)

Covering 358,226 accepted species,
8,847 hybrids, 48,003 accepted infraspecific taxa and 1,006,142 synonyms,
in total 1,421,218* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life