338.0000 Family Phyllanthaceae Martinov; Tekhno-Bot. Slovar 369 (1820), nom. cons.

58 genera, ca. 2200 spp.

Species listed in database: 2113; hybrids: 5

338.2416 Genus Glochidion J.R.Forst. & G.Forst.; Char. Gen. Pl., ed. 1: 57 (1775), nom. cons.

Distribution: Tropical Asia, Malesia, tropical Australia, Pacific

Species listed in database: 332

species :

Glochidion acuminatissimum Airy Shaw; Kew Bull. 27(1): 55 (1972)

Distribution: New Guinea (Irian Jaya)

42 NWG

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus lalongatanus Govaerts; Skvortsovia 4(3): 95 (2018), nom. nov.

species :

Glochidion acustylum Airy Shaw; Kew Bull. 34(3): 591 (1980)

Distribution: Sulawesi; New Guinea (Papua New Guinea)

42 NWG SUL

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus acustylus (Airy Shaw) Govaerts; Skvortsovia 4(3): 94 (2018)

species :

Glochidion alstonii Airy Shaw; Kew Bull. 36(3): 600 (1981)

Distribution: Sulawesi

42 SUL

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus alstonii (Airy Shaw) Govaerts; Skvortsovia 4(3): 94 (2018)

species :

Glochidion alticola Airy Shaw; Kew Bull. 27(1): 8 (1972)

Distribution: Sumatra

Error

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus alticola (Airy Shaw) Govaerts; Skvortsovia 4(3): 94 (2018)

species :

Glochidion aluminescens Airy Shaw; Kew Bull. 27(1): 16 (1972)

Distribution: Borneo (Sarawak)

42 BOR

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus aluminescens (Airy Shaw) Govaerts; Skvortsovia 4(3): 94 (2018)

species :

Glochidion ambiguum Airy Shaw; Kew Bull. 27(1): 62 (1972)

Distribution: New Guinea (Papua New Guinea); Bismarck Arch.; Solomon Isl.

42 NWG 43 BIS SOL

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus ambiguus (Airy Shaw) Govaerts; Skvortsovia 4(3): 94 (2018)

species :

Glochidion amentuligerum (Müll.Arg.) Croizat; Sargentia 1: 46 (1942)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus amentuliger (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Phyllanthus amentuliger Müll.Arg.; Flora 48: 390 (1865)

species :

Glochidion anamiticum (Kuntze) Merr.; Brittonia 2: 195 (1936)

Distribution: Vietnam; etc.

41 VIE

Synonyms: hidden

= Diasperus anamiticus Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Phyllanthus anamiticus (Kuntze) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

= Glochidion annamense Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 627 (1927)

species :

Glochidion andamanicum Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 42(2): 238 (1873)

Distribution: Andaman Isl. (South Andaman Isl.)

41 AND

Synonyms: hidden

= Phyllanthus andamanicobaricus Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

variant :

var. desmogyne (Hook.f.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 186 (1995)

Distribution: Andaman Isl. (North Andaman Isl.); Myanmar (Taninthayi)

41 AND MYA

Synonyms: hidden

= Diasperus desmogyne (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion airyshawii N.P.Balakr. & Chakr.; Bull. Bot. Surv. India 25: 220 (1983 publ. 1985)

= Glochidion desmogyne Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 310 (1887)

= Phyllanthus andamanicobaricus var. desmogyne (Hook.f.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion andersonii Airy Shaw; Kew Bull. 29(2): 287 (1974)

Distribution: Borneo (Sarawak)

42 BOR

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus sarawakensis Govaerts; Skvortsovia 4(3): 95 (2018), nom. nov.

species :

Glochidion anfractuosum Gibbs; J. Linn. Soc., Bot. 39: 168 (1909)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus anfractuosus (Gibbs) W.L.Wagner & Lorence; PhytoKeys 4: 69 (2011)

species :

Glochidion angulatum C.B.Rob.; Philipp. J. Sci., C 4: 91 (1909)

Distribution: Sumatra; Borneo; Brunei; Sulawesi; Philippines (Panay, Basilan, Leyte, Mindanao); Moluccas; New Guinea (Papua New Guinea); Solomon Isl.

42 BOR BRN MOL NWG PHI SUL 43 SOL

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus malesianus Govaerts; Skvortsovia 4(3): 95 (2018), nom. nov.

species :

Glochidion anthopotamicus (Hand.-Mazz.) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: China (Guangdong, Guangxi, Guizhou, Yunnan); Vietnam

36 CHI 41 VIE

Remarks: subgen. Phyllanthodendron

Synonyms: hidden

= Phyllanthus anthopotamicus Hand.-Mazz.; Symb. Sin. 7(2): 223-224 (1931)

= Phyllanthodendron anthopotamicum (Hand.-Mazz.) Croizat; J. Arnold Arbor. 23(1): 37 (1942)

species :

Glochidion apodogynum Airy Shaw; Kew Bull. 27(1): 44 (1972)

Distribution: New Guinea (Papua New Guinea); Australia (Northern Territory, Queensland)

42 NWG 50 AUS

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus apodogynus (Airy Shaw) Govaerts; Skvortsovia 4(3): 94 (2018)

species :

Glochidion arachnodes (Govaerts & Radcl.-Sm.) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: Cambodia

41 CBD

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Arachnodes chevalieri Gagnep.; Notul. Syst. (Paris) 14: 32 (1950)

= Phyllanthodendron chevalieri (Gagnep.) Airy Shaw; Kew Bull. 14: 470 (1960)

= Phyllanthus arachnodes Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 175 (1996), nom. nov.

= Phyllanthus chevalieri (Gagnep.) G.L.Webster; Ann. Missouri Bot. Gard. 81: 45 (1994), nom. illeg.

species :

Glochidion arborescens Blume; Bijdr. Fl. Ned. Ind. 584 (1825)

Distribution: China (Yunnan); India (Arunachal Pradesh, Assam, Mizoram Odisha, West Bengal); Bangladesh; Andaman Isl. (South Andaman Isl., Middle Andaman Isl.); Nicobar Isl. (Central Nicobar Isl.); Myanmar; Thailand; peninsular Malaysia (Langkawi Isl., Pahang, Selangor, Melaka); Singapore; Sumatra; Borneo; Java; Lesser Sunda Isl.

36 CHI 40 BAN IND 41 AND MYA NCB THA 42 BOR LSI MLY SNG

Synonyms: hidden

= Bradleia arborescens (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Diasperus arborescens (Blume) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus sclerophyllus (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus silheticus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus teysmannii (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion bancanum Miq.; Fl. Ned. Ind., Eerste Bijv. 450 (1861)

= Glochidion sclerophyllum Hook.f.; Fl. Brit. India [J.D.Hooker] 5(14): 313-314 (1887)

= Glochidion silheticum (Müll.Arg.) Croizat; J. Arnold Arbor. 21(4): 492 (1940)

= Glochidion zeylanicum var. arborescens (Blume) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 228 (1995)

= Phyllanthus arborescens (Blume) Müll.Arg.; Flora 48: 370 (1865)

= Phyllanthus obliquus var. mollis Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 284 (1866)

= Phyllanthus silheticus Müll.Arg.; Flora 48: 378 (1865)

= Phyllanthus teysmannii Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1270 (1866)

= Phyllanthus zeylanicus var. arborescens (Blume) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

species :

Glochidion atalotrichum A.C.Sm.; Contr. U.S. Natl. Herb. 37: 74 (1967)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus atalotrichus (A.C.Sm.) W.L.Wagner & Lorence; PhytoKeys 4: 70 (2011)

species :

Glochidion atrovirens A.C.Sm.; Fl. Vit. Nova 2: 491, 481 (1981)

Distribution: Fiji

60 FIJ

Synonyms: hidden

= Phyllanthus atrovirens (A.C.Sm.) W.L.Wagner & Lorence; PhytoKeys 4: 70 (2011)

species :

Glochidion auii Airy Shaw; Kew Bull. 27(1): 19 (1972)

Distribution: Borneo (Sarawak)

42 BOR

Synonyms: hidden

= Phyllanthus auii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

species :

Glochidion azaleon Airy Shaw; Kew Bull. 27(1): 15 (1972)

Distribution: Borneo (Kalimantan, Sabah)

42 BOR

Synonyms: hidden

= Phyllanthus azaleon (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

species :

Glochidion bachmaense Thin; Euphorb. Vietnam 49 (1995 publ. 1996)

Distribution: Vietnam

41 VIE

Synonyms: hidden

= Phyllanthus bachmaensis (Thin) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

species :

Glochidion balansae Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 628 (1927)

Distribution: Laos; N-Vietnam

41 LAO VIE

Synonyms: hidden

= Phyllanthus tricalysioides Govaerts & Christenh.; Nelumbo 68(1): 147 (2026), nom. nov.

species :

Glochidion barronense Airy Shaw; Kew Bull. 31(2): 343 (1976)

Distribution: Australia (Queensland)

50 AUS

Synonyms: hidden

= Phyllanthus barronensis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

species :

Glochidion beccarii Airy Shaw; Kew Bull. 23: 10 (1969)

Distribution: Sumatra

Error

Synonyms: hidden

= Phyllanthus beccarii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

species :

Glochidion beehleri W.N.Takeuchi; Sida 18(4): 945 (1999)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus beehleri (W.N.Takeuchi) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

species :

Glochidion beguinii Airy Shaw; Kew Bull. 27(1): 27 (1972)

Distribution: Moluccas (Halmaheira)

42 MOL

Synonyms: hidden

= Phyllanthus beguinii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 136 (2026)

species :

Glochidion benguetense Elmer; Leafl. Philipp. Bot. 1: 304 (1908)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Glochidion sablanense Elmer; Leafl. Philipp. Bot. 1: 306 (1908)

= Phyllanthus sablanense (Elmer) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion benthamianum Domin; Biblioth. Bot. 22(89): 318 (1927)

Distribution: New Guinea (Irian Jaya, Papua New Guinea); Australia (Queensland)

42 NWG 50 AUS

Synonyms: hidden

= Glochidion obscurum var. papuanum J.J.Sm.; (1910)

= Glochidion capitis-york Airy Shaw; Kew Bull. 23: 25 (1969)

= Phyllanthus capitis-york (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion billardierei Baill.; Adansonia 2: 241 (1861 publ. 1862)

Distribution: New Caledonia (incl. Loyalty Isl.)

60 NWC

Synonyms: hidden

= Bradleia glauca Labill.; Sert. Austro-Caledon. 76, t. 77 (1825)

= Diasperus billardierei (Baill.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus kanalophilus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus wagapensis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion glaucum (Labill.) Müll.Arg.; Linnaea 32: 62 (1863), nom. illeg.

= Glochidion heterolobum Müll.Arg.; Linnaea 32: 63 (1863)

= Glochidion kanalense Baill.; Adansonia 2: 241 (1861 publ. 1862)

= Glochidion wagapense (Müll.Arg.) Briq.; Annuaire Conserv. Jard. Bot. Genève 4: 226 (1900)

= Phyllanthus billardierei (Baill.) Müll.Arg.; Flora 48: 375 (1865)

= Phyllanthus glaucus (Labill.) Müll.Arg.; Flora 48: 386 (1865), nom. illeg.

= Phyllanthus kanalophilus Müll.Arg.; Flora 48: 375 (1865)

= Phyllanthus wagapensis Müll.Arg.; Flora 48: 372 (1865)

species :

Glochidion borgmannii Airy Shaw; Kew Bull. 27(1): 49 (1972)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus borgmannii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion borneense (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

Distribution: peninsular Malaysia (Penang, Perak, Johor); Singapore; Sumatra (Bangka Isl.); Borneo; Brunei; Java

42 BOR BRN MLY SNG

Synonyms: hidden

= Diasperus borneensis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus microbotrys (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus polycarpus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion microbotrys Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 319 (1887)

= Glochidion trilobum Ridl.; Bull. Misc. Inform. Kew 1923: 364 (1923)

= Phyllanthus borneensis Müll.Arg.; Flora 48: 377 (1865)

= Phyllanthus polycarpus Müll.Arg.; Flora 48: 387 (1865)

species :

Glochidion bourdillonii Gamble; Bull. Misc. Inform. Kew 1925: 330 (1925)

Distribution: S-India (Karnataka, Kerala, Tamil Nadu)

40 IND

Synonyms: hidden

= Phyllanthus bourdillonii (Gamble) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

variant :

var. bhutanicum (D.G.Long) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 190 (1995)

Distribution: Bhutan

40 BHU

Synonyms: hidden

= Glochidion bhutanicum D.G.Long; Notes Roy. Bot. Gard. Edinburgh 44(1): 169 (1986)

= Glochidion bourdillonii subsp. bhutanicum (D.G.Long) Chakr. & N.P.Balakr.; Indo-Burmese Phyllanthaceae 204 (2018)

= Phyllanthus bourdillonii var. bhutanicus (D.G.Long) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion bracteatum Gillespie; Bull. Bernice P. Bishop Mus. 91: 15 (1932)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus bracteatus (Gillespie) W.L.Wagner & Lorence; PhytoKeys 4: 70 (2011)

species :

Glochidion breyniopsis Esser & R.W.Bouman; Phytotaxa 540(1): 64 (2022) nom. nov.

Distribution: China (Guangxi)

36 CHI

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthodendron breynioides P.T.Li; Bull. Bot. Lab. N. E. Forest. Inst., Harbin 7(3): 6 (1987)

= Phyllanthus breynioides (P.T.Li) Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 176 (1996)

species :

Glochidion brideliifolium Airy Shaw; Kew Bull. 23: 15 (1969)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus brideliifolius (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion brooksii Ridl.; Bull. Misc. Inform. Kew 1925: 88 (1925)

Distribution: peninsular Malaysia; Sumatra; Borneo

42 BOR MLY

Synonyms: hidden

= Glochidion laevigatum var. cuspidatum Ridl.; Fl. Malay Penins. 3: 215 (1924)

species :

Glochidion brothersonii J.Florence; Fl. Polynésie Franç. [Florence] 1: 68 (1997)

Distribution: Society Isl. (Raiatea)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus brothersonii (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 70 (2011)

species :

Glochidion brunnescens A.C.Sm.; Fl. Vit. Nova 2: 491, 482 (1981)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus brunnescens (A.C.Sm.) W.L.Wagner & Lorence; PhytoKeys 4: 71 (2011)

species :

Glochidion bullatissimum Airy Shaw; Kew Bull. 32(2): 376 (1978)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus bullatissimus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion butonicum Airy Shaw; Kew Bull. 29(2): 291 (1974)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus butonicus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion cacuminum Müll.Arg.; Linnaea 32: 60 (1863)

Distribution: Java; Lesser Sunda Isl. (Sumbawa)

42 LSI

Synonyms: hidden

= Phyllanthus cacuminum (Müll.Arg.) Müll.Arg.; Flora 48: 371 (1865)

= Diasperus cacuminum (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

species :

Glochidion cagayanense C.B.Rob.; Philipp. J. Sci., C 6: 328 (1911)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus cagayanensis (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion calciphilum Croizat; Sargentia 1: 46 (1942)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus calciphilus (Croizat) W.L.Wagner & Lorence; PhytoKeys 4: 71 (2011)

species :

Glochidion caledonicum Müll.Arg.; Linnaea 32: 62 (1863)

Distribution: New Caledonia (incl. Loyalty Isl.)

60 NWC

Synonyms: hidden

= Bradleia zeylanica Labill.; Sert. Austro-Caledon. 76, t. 76 (1825), nom. illeg.

= Diasperus caledonicus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Glochidion diospyroides Schltr.; Bot. Jahrb. Syst. 39: 147 (1906)

= Phyllanthus caledonicus (Müll.Arg.) Müll.Arg.; Flora 48: 376 (1865)

species :

Glochidion calocarpum Kurz; J. Bot. 13: 330 (1875)

Distribution: Andaman Isl. (scattered, Barren Isl.); Nicobar Isl. (Car Nicobar Isl., North Nicobar Isl., Central Nicobar Isl., Great Nicobar Isl., Little Nicobar Isl.)

41 AND NCB

Synonyms: hidden

= Diasperus calocarpus (Kurz) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Phyllanthus calocarpus (Kurz) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion caloneurum Airy Shaw; Kew Bull. 23: 16 (1969)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus caloneurus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion calospermum Airy Shaw; Kew Bull. 27(1): 13 (1972)

Distribution: Borneo (Sarawak, Sabah, Kalimantan)

42 BOR

Synonyms: hidden

= Phyllanthus calospermus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion camiguinense Merr.; Philipp. J. Sci., C 3: 414 (1909)

Distribution: Philippines (Camiguin de Babuyanes)

42 PHI

Synonyms: hidden

= Phyllanthus camiguinensis (Merr.) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion candolleanum (Wight & Arn.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 191 (1995)

Distribution: S-India (Andhra Pradesh, Karnataka, Maharashtra, Kerala, Tamil Nadu); Sri Lanka

40 IND SRL

Synonyms: hidden

= Diasperus arboreus (Wight) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus neilgerrhensis (Wight) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus perrottetianus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus pycnocarpus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion arboreum var. pauciflorum Hook.f. ex Gamble; Fl. Madras 914 (1922), pro syn.

= Glochidion arboreum Wight; Icon. Pl. Ind. Orient. [Wight] 5: t. 1907 (1852)

= Glochidion neilgherrense Wight; Icon. Pl. Ind. Orient. [Wight] 5: t. 1907 (1852)

= Glochidion pachycarpum Alston; Ann. Roy. Bot. Gard. (Peradeniya) 11: 5 (1928)

= Glochidion pauciflorum (Hook.f.) Gamble; Fl. Madras 1307 (1925)

= Glochidion perrottetianum Bedd.; Forester´s Man. Bot. 194 (1872)

= Glochidion pycnocarpum (Müll.Arg.) Bedd.; Forester´s Man. Bot. 194 (1872)

= Glochidion pycnocarpum var. ellipticum Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 316 (1887)

= Glochidion sisparense Gamble; Fl. Madras 1307 (1925)

= Lobocarpus candolleanus Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 7 (1834)

= Phyllanthus arboreus (Wight) Müll.Arg.; Flora 48: 385 (1865)

= Phyllanthus candolleanus (Wight & Arn.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

= Phyllanthus neilgherrensis (Wight) Müll.Arg.; Flora 48: 385 (1865)

= Phyllanthus pachycarpus (Alston) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

= Phyllanthus perrottetianus Müll.Arg.; Flora 48: 387 (1865)

= Phyllanthus pycnocarpus Müll.Arg.; Flora 48: 386 (1865)

species :

Glochidion canescens Elmer; Leafl. Philipp. Bot. 8: 3082 (1919)

Distribution: Philippines (Luzon); Lesser Sunda Isl. (Timor); Moluccas (Seram)

42 LSI MOL PHI

Synonyms: hidden

= Phyllanthus canescens (Elmer) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion carinatum (Beille) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: Vietnam

41 VIE

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus carinatus Beille; Bull. Soc. Bot. France 72: 160 (1925), et Fl. Indo-Chine [P.H.Lecomte et al.] 5: 591 (1927)

= Phyllanthodendron carinatum (Beille) Croizat; J. Arnold Arbor. 23: 36 (1942)

species :

Glochidion carrickii Airy Shaw; Kew Bull. 25(3): 482 (1971)

Distribution: peninsular Malaysia (twice from Ulu Gombak F. R., Selangor)

42 MLY

Synonyms: hidden

= Phyllanthus carrickii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion carrii Airy Shaw; Kew Bull. 23: 20 (1969)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus carrii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 137 (2026)

species :

Glochidion castaneum Airy Shaw; Kew Bull. 27(1): 36 (1972)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus castaneus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion caudatifolium (P.T.Li) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: China (Guangxi)

36 CHI

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus lii Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 177 (1996), nom. nov.

= Phyllanthodendron caudatifolium P.T.Li; Bull. Bot. Lab. N. E. Forest. Inst., Harbin 7(3): 7 (1987)

species :

Glochidion cauliflorum Merr.; Philipp. J. Sci. 16: 545 (1920)

Distribution: Philippines (Mindanao, Leyte)

42 PHI

Synonyms: hidden

= Phyllanthus caulanthus Govaerts & Christenh.; Nelumbo 68(1): 138 (2026), nom. nov.

species :

Glochidion celastroides (Müll.Arg.) Pax; Nat. Pflanzenfam. [Engler & Prantl] III, 5: 24 (1890)

Distribution: Sumatra (Bangka Isl., Belitung); Borneo; Brunei

42 BOR BRN

Synonyms: hidden

= Diasperus celastroides (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891), as ´celastrodes´

= Phyllanthus celastroides Müll.Arg.; Flora 48: 390 (1865)

species :

Glochidion cenabrei Merr.; Philipp. J. Sci. 26: 463 (1925)

Distribution: Palawan

42 PLW

Synonyms: hidden

= Phyllanthus cenabrei (Merr.) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion chevalieri Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 615 (1927)

Distribution: Laos; Cambodia; Vietnam

41 CBD LAO VIE

Synonyms: hidden

= Phyllanthus laoticus Govaerts & Christenh.; Nelumbo 68(1): 141 (2026), nom. nov.

species :

Glochidion chlamydogyne Airy Shaw; Kew Bull. 23: 14 (1969)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus chlamydogyne (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion chodoense C.S.Lee & H.T.Im; Korean J. Pl. Taxon. 24: 13 (1994)

Distribution: South Korea

38 KOR

Synonyms: hidden

= Phyllanthus chodoensis (C.S.Lee & H.T.Im) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion chondrocarpum Airy Shaw; Kew Bull. 32(2): 372 (1978)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus chondrocarpus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion christophersenii Croizat; Occas. Pap. Bernice P. Bishop Mus. 17: No. 16, 213 (1943)

Distribution: [Western] Samoa (Savai´i)

60 SAM

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus christophersenii (Croizat) W.L.Wagner & Lorence; PhytoKeys 4: 71 (2011)

species :

Glochidion cleistanthoides Fosberg; Willdenowia 20(1-2): 263 (1991)

Distribution: Fed. States of Micronesia

62 MIC

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus cleistanthoides (Fosberg) W.L.Wagner & Lorence; PhytoKeys 4: 71 (2011)

species :

Glochidion coccineum (Banks) Müll.Arg.; Linnaea 32: 60 (1863)

Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Yunnan); India (Assam, ?West Bengal); Myanmar; Thailand; Laos; Cambodia; Vietnam

36 CHI 40 IND 41 CBD LAO MYA THA VIE

Synonyms: hidden

= Agyneia coccinea Banks; Account Embassy Kingd. Ava [Symes] 479 (1800)

= Bradleia coccinea Benth.; Numer. List [Wallich] no. 7868 (1847), nomen

= Diasperus coccineus (Banks) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Episteira coccinea (Banks) Raf.; Sylva Tellur. 20 (1838)

= Phyllanthus coccineus (Banks) Müll.Arg.; Flora 48: 370 (1865)

species :

Glochidion collectorum Airy Shaw; Kew Bull. 33(1): 33 (1978)

Distribution: Solomon Isl.

43 SOL

Synonyms: hidden

= Phyllanthus collectorum (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion collinum A.C.Sm.; Fl. Vit. Nova 2: 494, 486 (1981)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus vitilevuensis W.L.Wagner & Lorence; PhytoKeys 4: 90 (2011), nom. nov.

species :

Glochidion comitum Florence; Novon 7: 29 (1997)

Distribution: Pitcairn Isl.

61 PIT

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus comitus (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 72 (2011)

species :

Glochidion concolor Müll.Arg.; Linnaea 32: 63 (1863)

Distribution: Tonga (´Eua, Lifuka, Vava´u); Tonga (Late Isl., Tongatapu, ´Eua, Nomuka, Lifuka, Vava´u, Niuafo´ou); Niue; Fiji; Wallis & Futuna; Cook Isl. (Rarotonga, Mangaia); Society Isl. (Tahiti); Tuamotu Arch. (Makatea Isl.); Austral Isl. (Rapa Iti)

60 FIJ NUE TON WAL 61 AUI COO SCI TUA

Remarks: see also G. ramiflorum which occurs only west of Vanuatu fide Smith 1981, and G. tahitense

Synonyms: hidden

= Diasperus concolor (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion ramiflorum var. lanceolatum Müll.Arg.; Linnaea 32: 63 (1863)

= Phyllanthus concolor (Müll.Arg.) Müll.Arg.; Flora 48: 374 (1865)

= Phyllanthus concolor var. ellipticus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 290 (1866)

= Phyllanthus ramiflorus var. lanceolatus Müll.Arg.; Flora 48: 374 (1865)

species :

Glochidion conostylum Airy Shaw; Kew Bull. 23: 23 (1969)

Distribution: Bismarck Arch.

43 BIS

Synonyms: hidden

= Phyllanthus conostylus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion cordatum Seem. ex Müll.Arg.; Linnaea 32: 64 (1863)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus cordatus (Seem. ex Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion cordatum Seem.; Bonplandia (Hannover) 9: 259 (1861), nomen

= Phyllanthus cordatus (Seem. ex Müll.Arg.) Müll.Arg.; Flora 48: 376 (1865)

species :

Glochidion coriaceum Thwaites; Enum. Pl. Zeyl. [Thwaites] 285 (1859)

Distribution: Sri Lanka

40 SRL

Synonyms: hidden

= Bradleia coriacea Benth.; Numer. List [Wallich] no. 7872 (1847), nomen

= Diasperus brachylobus (Thwaites) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus coriaceus (Thwaites) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus permixtus Kuntze; Revis. Gen. Pl. 2: 597 (1891)

= Glochidion brachylobum (Thwaites) Müll.Arg.; Linnaea 32: 62 (1863)

= Phyllanthus brachylobus (Thwaites) Müll.Arg.; Flora 48: 373 (1865)

= Phyllanthus coriaceus (Thwaites) Müll.Arg.; Flora 48: 386 (1865)

species :

Glochidion coronulatum C.B.Rob.; Philipp. J. Sci., C 4: 94 (1909)

Distribution: Palawan; Philippines (Luzon, Leyte)

42 PHI PLW

Synonyms: hidden

= Glochidion balsahanense Elmer; Leafl. Philipp. Bot. 4: 1289 (1911)

= Phyllanthus coronulatus (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 138 (2026)

species :

Glochidion cupreum Airy Shaw; Kew Bull. 27(1): 15 (1972)

Distribution: Borneo (Kalimantan)

42 BOR

Synonyms: hidden

= Phyllanthus cupreus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion curranii C.B.Rob.; Philipp. J. Sci., C 4: 102 (1909)

Distribution: Palawan; Philippines (Culion)

42 PHI PLW

Synonyms: hidden

= Phyllanthus culionanthus Govaerts & Christenh.; Nelumbo 68(1): 138 (2026), nom. nov.

species :

Glochidion cuspidatum (Müll.Arg.) Pax; Bot. Jahrb. Syst. 25: 645 (1898)

Distribution: [Western] Samoa (Savai´i, ´Upolu); American Samoa (Tutuila)

60 ASA SAM

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus cuspidatus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus cuspidatus Müll.Arg.; Flora 48: 377 (1865)

species :

Glochidion cyrtophyllum Miq.; Fl. Ned. Ind. 1(2): 378 (1859)

Distribution: Sumatra

Error

Synonyms: hidden

= Diasperus cyrtophyllus (Miq.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus cyrtophyllus (Miq.) Müll.Arg.; Flora 48: 390 (1865)

species :

Glochidion cyrtostylum Miq.; Fl. Ned. Ind. 1(2): 378 (1859)

Distribution: Borneo; Java; Lesser Sunda Isl. (Bali)

42 BOR LSI

Synonyms: hidden

= Diasperus cyrtostylus (Miq.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus cyrtostylus (Miq.) Müll.Arg.; Flora 48: 387 (1865)

species :

Glochidion daltonii (Müll.Arg.) Kurz; Prelim. Rep. Forest Pegu, App. A: cv. (1875), App. B. 77, in clavi, et Forest Fl. Burma 2: 344 (1877)

Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Shandong, Sichuan, Yunnan, Zhejiang); Nepal; India (Assam, Meghalaya, Sikkim, West Bengal); Bangladesh; Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia

36 CHI 40 BAN IND NEP 41 CBD LAO MYA THA VIE 42 MLY

Synonyms: hidden

= Diasperus daltonii (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus gamblei (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion gamblei Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 310 (1887)

= Phyllanthus daltonii Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 310 (1866)

species :

Glochidion dasystylum Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 42(2): 237 (1873)

Distribution: Myanmar

41 MYA

Synonyms: hidden

= Diasperus dasystylus (Kurz) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Phyllanthus dasystylus (Kurz) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion daviesii W.N.Takeuchi; Harvard Pap. Bot. 8(1): 131 (2003)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus daviesii (W.N.Takeuchi) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion decorum J.J.Sm.; Nova Guinea 8: 222 (1910)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus decorus (J.J.Sm.) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion delticola Airy Shaw; Kew Bull. 29(2): 292 (1974)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus delticola (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion dichromum Airy Shaw; Kew Bull. 27(1): 23 (1972)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus dichromus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion discogyne Airy Shaw; Kew Bull. 32(2): 374 (1978)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus discogyne (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion disparilaterum S.Moore; J. Bot. 63(Suppl.): 95 (1925)

Distribution: Sumatra

Error

Synonyms: hidden

= Phyllanthus disparilaterus (S.Moore) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion disparipes Airy Shaw; Kew Bull. 27(1): 43 (1972)

Distribution: New Guinea (Papua New Guinea); Australia (Western Australia, Northern Territory, Queensland)

42 NWG 50 AUS

Synonyms: hidden

= Phyllanthus disparipes (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion dodecapterum Airy Shaw; Kew Bull. 36(3): 602 (1981)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus dodecapterus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion dolichostylum Merr.; Philipp. J. Sci., C 9: 483 (1914 publ. 1915)

Distribution: Palawan

42 PLW

Synonyms: hidden

= Phyllanthus dolichostylus (Merr.) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion dongmoensis (Thin) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: Vietnam

41 VIE

Remarks: subgen. Phyllanthodendron

Synonyms: hidden

= Phyllanthus dongmoensis Thin; J. Biol. (Vietnam) 14(2): 16 (1992)

species :

Glochidion drypetifolium Airy Shaw; Kew Bull. 27(1): 34 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus drypetifolius (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion dumicola Airy Shaw; Kew Bull. 27(1): 47 (1972)

Distribution: Borneo (Kalimantan); New Guinea (Irian Jaya)

42 BOR NWG

Synonyms: hidden

= Phyllanthus dumicola (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion dunnianum (H.Lév.) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: China (Guizhou, Yunnan, Guangxi); Vietnam

36 CHI 41 VIE

Remarks: subgen. Phyllanthodendron

Synonyms: hidden

= Phyllanthodendron cavaleriei H.Lév.; Repert. Spec. Nov. Regni Veg. 9(222-226): 454 (1911)

= Phyllanthodendron dunnianum H.Lév.; Repert. Spec. Nov. Regni Veg. 9(214-216): 324 (1911)

= Phyllanthodendron dunnianum var. hypoglaucum H.Lév.; Fl. Kouy-Tcheou 166 (1914)

= Phyllanthus dunnianus (H.Lév.) Hand.-Mazz. ex Rehder; J. Arnold Arbor. 14(3): 230-231 (1933)

species :

Glochidion elaphrocarpum Airy Shaw; Kew Bull. 27(1): 35 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus elaphrocarpus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion ellipticum Wight; Icon. Pl. Ind. Orient. [Wight] 5: t. 1906 (1852)

Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Taiwan; Nepal; India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Karnataka, Kerala, Maharashtra, Meghalaya, Nagaland Odisha, Sikkim, Tamil Nadu, Tripura, Uttar Pradesh, West Bengal); Bangladesh; Myanmar; Thailand; Laos; Vietnam

36 CHI 38 TAI 40 BAN IND NEP 41 LAO MYA THA VIE

Synonyms: hidden

= Bradleia wightiana Benth.; Numer. List [Wallich] no. 7862 (1847), nomen

= Diasperus assamicus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus malabaricus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion assamicum (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5(14): 319-320 (1887)

= Glochidion assamicum var. brevipedicellatum Hurus. & Yu.Tanaka; Fl. E. Himalaya 177 (1966)

= Glochidion assamicum var. magnicapsulum Croizat & H.Hara; J. Jap. Bot. 16: 319 (1940)

= Glochidion balakrishnanii Jothi, Manickam, Sundaresan & Josephine; J. Econ. Taxon. Bot. 26(1): 114 (2002)

= Glochidion diversifolium Bedd.; Forester´s Man. Bot. 193 (1872)

= Glochidion diversifolium var. wightianum (Müll.Arg.) Bedd.; Fl. Sylv. S. India: cxciii (1873)

= Glochidion ellipticum var. wightiana (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 321 (1887)

= Glochidion malabaricum (Müll.Arg.) Bedd.; Forester´s Man. Bot. 194 (1872)

= Glochidion mandakamdevi Borthakur & Kalita; J. Econ. Taxon. Bot. 30: 490 (2006)

= Glochidion ovalifolium F.Y.Lu & Y.S.Hsu; Quart. J. Forest Res. 25(4): 87 (2003)

= Phyllanthus assamicus Müll.Arg.; Flora 48: 378. 1865

= Phyllanthus diversifolius var. longifolius Müll.Arg.; Flora 48: 378 (1865)

= Phyllanthus diversifolius var. wightianus Müll.Arg.; Flora 48: 378 (1865)

= Phyllanthus malabaricus Müll.Arg.; Linnaea 34: 69 (1865 publ. 1866)

species :

Glochidion elmeri Merr.; Univ. Calif. Publ. Bot. 15: 139 (1929)

Distribution: Sumatra; Borneo

42 BOR

Synonyms: hidden

= Phyllanthus elmeri (Merr.) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion emarginatum J.W.Moore; Bull. Bernice P. Bishop Mus. 102: 30 (1933)

Distribution: Society Isl. (Raiatea)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Glochidion raiateense J.W.Moore; Bull. Bernice P. Bishop Mus. 102: 30 (1933)

= Phyllanthus emarginatus (J.W.Moore) W.L.Wagner & Lorence; PhytoKeys 4: 73 (2011)

species :

Glochidion eriocarpum Champ. ex Benth.; Hooker´s J. Bot. Kew Gard. Misc. 6: 6 (1854)

Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hongkong, Hunan, Yunnan); Taiwan; Myanmar; Thailand; Laos; Vietnam; peninsular Malaysia; Sulawesi

36 CHI 38 TAI 41 LAO MYA THA VIE 42 MLY SUL

Synonyms: hidden

= Diasperus eriocarpus (Champ. ex Benth.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus villicaulis (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion esquirolii H.Lév.; Repert. Spec. Nov. Regni Veg. 12: 186 (1913)

= Glochidion villicaule Hook.f.; Fl. Brit. India [J.D.Hooker] 5(14): 326 (1887)

= Phyllanthus eriocarpus (Champ. ex Benth.) Müll.Arg.; Linnaea 32: 67 (1863)

species :

Glochidion eucleoides S.Moore; J. Bot. 61: Suppl. 44 (1923)

Distribution: New Guinea (Irian Jaya, Papua New Guinea, d´Entrecasteaux Arch., Louisiade Arch.)

42 NWG

Synonyms: hidden

= Phyllanthus eucleoides (S.Moore) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion euryoides A.C.Sm.; J. Arnold Arbor. 33: 373 (1952)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus euryoides (A.C.Sm.) W.L.Wagner & Lorence; PhytoKeys 4: 74 (2011)

species :

Glochidion falcatilimbum Merr.; Philipp. J. Sci. 16: 542 (1920)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus falcatilimbus (Merr.) Govaerts & Christenh.; Nelumbo 68(1): 139 (2026)

species :

Glochidion ferdinandi (Müll.Arg.) F.M.Bailey; Queensl. Fl. 5: 1423 (1902)

Distribution: Australia (Queensland, New South Wales)

50 AUS

Synonyms: hidden

= Diasperus ferdinandi (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion ferdinandi (Müll.Arg.) Pax & K.Hoffm.; Nat. Pflanzenfam. [Engler & Prantl], 2. Aufl., 19c: 57 (1931)

= Glochidion ferdinandi var. pubens Maiden ex Airy Shaw; Kew Bull. 31: 346 (1976)

= Phyllanthus ferdinandi Müll.Arg.; Flora 48: 379 (1865)

= Phyllanthus ferdinandi var. minor Benth.; Fl. Austral. [Bentham] 6: 96 (1873)

species :

Glochidion flavidum Kurz ex Teijsm. & Binn.; Tijdschr. Ned.-Indië 27: 46 (1864)

Distribution: Java

Error

Synonyms: hidden

= Diasperus flavidus (Kurz ex Teijsm. & Binn.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus flavidus (Kurz ex Teijsm. & Binn.) Müll.Arg.; Flora 48: 390 (1865)

species :

Glochidion formanii Airy Shaw; Kew Bull. 27(1): 25 (1972)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus formanii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion frodinii Airy Shaw; Kew Bull. 27(1): 51 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus pyriformis Govaerts & Christenh.; Nelumbo 68(1): 145 (2026), nom. nov.

species :

Glochidion fulvirameum Miq.; Fl. Ned. Ind. 1(2): 376 (1859)

Distribution: Java; Moluccas (Aru Isl.); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl.

42 MOL NWG 43 SOL

Synonyms: hidden

= Diasperus fulvirameus (Miq.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion salomonis Briq.; Annuaire Conserv. Jard. Bot. Genève 4: 226 (1900)

= Phyllanthus fulvirameus (Miq.) Müll.Arg.; Flora 48: 385 (1865)

species :

Glochidion galorii Airy Shaw; Kew Bull. 25(3): 484 (1971)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus galorii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion gaudichaudii (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

Distribution: Moluccas (Aru Isl., Gorong Isl.)

42 MOL

Synonyms: hidden

= Diasperus gaudichaudii (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus gaudichaudii Müll.Arg.; Flora 48: 379 (1865)

species :

Glochidion geoffrayi Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 619 (1927)

Distribution: Cambodia; Vietnam

41 CBD VIE

Synonyms: hidden

= Phyllanthus minusculus Govaerts & Christenh.; Nelumbo 68(1): 143 (2026), nom. nov.

species :

Glochidion gigantifolium (Vidal) J.J.Sm.; Nova Guinea 8(1): 223 (1910)

Distribution: Philippines (Luzon, Negros, Mindanao)

42 PHI

Synonyms: hidden

= Phyllanthus gigantifolius Vidal; Revis. Pl. Vasc. Filip. 236 (1886)

= Glochidion gigantifolium (Vidal) Merr.; Enum. Philipp. Fl. Pl. 2: 399 (1923), isonym

species :

Glochidion gillespiei Croizat; Sargentia 1: 46 (1942)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus gillespiei (Croizat) W.L.Wagner & Lorence; PhytoKeys 5: 74 (2011)

species :

Glochidion gimi (K.Schum.) Pax & K.Hoffm.; Nat. Pflanzenfam. [Engler & Prantl], 2. Aufl., 19c: 58 (1931)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus gimi (K.Schum.) Govaerts; Skvortsovia 4(3): 94 (2018)

= Tetraglochidion gimi K.Schum.; Nachtr. Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 291 (1905)

species :

Glochidion glabrum J.J.Sm.; Nova Guinea 8: 224 (1910)

Distribution: Sulawesi; Lesser Sunda Isl.; New Guinea (Irian Jaya)

42 LSI NWG SUL

Synonyms: hidden

= Glochidion mindorense subsp. glabrum (J.J.Sm.) Airy Shaw; Kew Bull. 27(1): 22 (1972)

= Phyllanthus glaber (J.J.Sm.) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion glaucescens Merr.; Philipp. J. Sci., C 8: 381 (1913)

Distribution: Philippines (Leyte)

42 PHI

Synonyms: hidden

= Phyllanthus metallicus Govaerts & Christenh.; Nelumbo 68(1): 142 (2026), nom. nov.

species :

Glochidion glaucops Airy Shaw; Kew Bull. 27(1): 10 (1972)

Distribution: Sumatra

Error

Synonyms: hidden

= Phyllanthus glaucops (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion glomerulatum (Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

Distribution: Thailand; Cambodia; Vietnam; peninsular Malaysia; +Singapore; Sumatra (incl. Bangka Isl.); Borneo; Brunei; Java

41 CBD THA VIE 42 BOR BRN MLY SNG

Synonyms: hidden

= Agyneia glomerulata Miq.; Fl. Ned. Ind., Eerste Bijv. 447 (1861)

= Bridelia heterantha Benth.; Numer. List [Wallich] no. 7878 (1847) (nom. inval.)

= Diasperus glomerulatus (Miq.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus heteranthus (Wall.) Kuntze; Revis. Gen. Pl. 2: 597 (1891)

= Diasperus nanogynus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion nanogynum (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 318 (1887)

= Glochidion palustre Koord.; Bull. Jard. Bot. Buitenzorg, Sér. 3, 1: 145 (1918)

= Phyllanthus glomerulatus (Miq.) Müll.Arg.; Flora 48: 375 (1865)

= Phyllanthus nanogynus Müll.Arg.; Flora 48: 376 (1865)

species :

Glochidion goniocarpum Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 309 (1887)

Distribution: peninsular Malaysia; +Singapore

42 MLY SNG

Synonyms: hidden

= Diasperus goniocarpus (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Phyllanthus goniocarpus (Hook.f.) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion goniocladum Airy Shaw; Kew Bull. 34(3): 592 (1980)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus giluwensis Govaerts & Christenh.; Nelumbo 68(1): 140 (2026), nom. nov.

species :

Glochidion gracile Airy Shaw; Kew Bull. 36(3): 601 (1981)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus crocotillus Govaerts & Christenh.; Nelumbo 68(1): 138 (2026), nom. nov.

species :

Glochidion gracilentum (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

Distribution: Java

Error

Synonyms: hidden

= Diasperus gracilentus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus gracilentus Müll.Arg.; Flora 48: 380 (1865)

species :

Glochidion grantii J.Florence; Bull. Mus. Natl. Hist. Nat., B, Adansonia 18: 250, fig. 3 (1996)

Distribution: Society Isl. (Raiatea, Tahaa)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus grantii (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 74 (2011)

species :

Glochidion granulare Airy Shaw; Kew Bull. 27(1): 42 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Glochidion granulare var. heterogyne Airy Shaw; Kew Bull. 36(3): 602 (1981)

= Phyllanthus granularis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion grayanum (Müll.Arg.) J.Florence; Bull. Mus. Natl. Hist. Nat., B, Adansonia 18: 253 (1996)

Distribution: Society Isl. (Tahiti)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus grayanus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus grayanus Müll.Arg.; Flora 48: 380 (1865)

species :

Glochidion grossum Airy Shaw; Kew Bull. 25(3): 483 (1971)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus grossus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion harveyanum Domin; Biblioth. Bot. 22(89): 319 (1927)

Distribution: Australia (Queensland)

50 AUS

Synonyms: hidden

= Glochidion mindorense subsp. harveyanum (Domin) Airy Shaw; Kew Bull. 27(1): 23 (1972)

= Glochidion harveyanum var. pubescens Airy Shaw; Kew Bull. 31(2): 347 (1976)

= Phyllanthus harveyanus (Domin) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion helferi (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 311 (1887)

Distribution: Andaman Isl.; Myanmar (Taninthayi)

41 AND MYA

Synonyms: hidden

= Diasperus andamanicus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus helferi (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion helferi (Müll.Arg.) Pax; Nat. Pflanzenfam. [Engler & Prantl] III, 5: 24 (1896)

= Glochidion subscandens Kurz; Forest Fl. Burma 2: 344 (1877), nom. illeg.

= Phyllanthus andamanicus Kurz; Rep. Veg. Andaman Isl., ed. 2: 47 (1870) (pro syn.)

= Phyllanthus helferi Müll.Arg.; Flora 48: 372 (1865)

species :

Glochidion heterocalyx Airy Shaw; Kew Bull. 27(1): 57 (1972)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus heterocalyx (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion heyneanum (Wight & Arn.) Wight; Icon. Pl. Ind. Orient. [Wight] t. 1908 (1852) ex Müll. Arg., Linnaea 32(Heft 1): 67 (1863)

Distribution: China (Yunnan); Pakistan (Hazara, Murree); Nepal; India (throughout, incl. Jammu & Kashmir); Bangladesh; Myanmar; Laos; Cambodia; Vietnam

36 CHI 40 BAN IND NEP PAK 41 CBD LAO MYA VIE

Synonyms: hidden

= Bradleia ovata Wall.; Numer. List [Wallich] no. 7052 (1832), nomen

= Diasperus asperus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus heyneanus (Wight & Arn.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus nepalensis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus velutinus (Wight) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Eriococcus glaucescens Zoll.; Tijdschr. Ned.-Indië 14: 173 (1857)

= Glochidion asperum (Müll.Arg.) Bedd.; Fl. Sylv. S. India: cxciii (1873)

= Glochidion nepalense (Müll.Arg.) Kurz; Forest Fl. Burma 2: 344 (1877)

= Glochidion shanense Gang Yao & S.X.Luo; PhytoKeys 96: 59 (2018)

= Glochidion velutinum Wight; Icon. Pl. Ind. Orient. [Wight] 5(2): t. 1907, f. 2 (1852)

= Gynoon heyneanum Wight & Arn.; Edinburgh New Philos. J. 14: 300 (1833)

= Phyllanthus asperus Müll.Arg.; Flora 48: 377 (1865)

= Phyllanthus heyneanus (Wight & Arn.) Müll.Arg.; Flora 48: 389 (1865), nom. illeg.

= Phyllanthus nepalensis Müll.Arg.; Flora 48: 375 (1865)

= Phyllanthus velutinus (Wight) Müll.Arg.; Flora 48: 380 (1865)

variant :

var. arunachalense (Chakr. & M.Gangop.) Chakr. & N.P.Balakr.; Indo-Burmese Phyllanthaceae 219 (2018)

Distribution: India (Assam, Arunachal Pradesh)

40 IND

Synonyms: hidden

= Glochidion mishmiense var. arunachalense (Chakr. & M.Gangop.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 39: 556 (2015)

= Glochidion zeylanicum var. arunachalense Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 229 (1995)

= Phyllanthus velutinus var. arunachalensis (Chakrab. & M.Gangop.) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

= Phyllanthus zeylanicus var. arunachalensis (Chakr. & M.Gangop.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

variant :

var. mishmiense (Hook.f.) Chakr. & N.P.Balakr.; Indo-Burmese Phyllanthaceae 221 (2018)

Distribution: India (Arunachal Pradesh, Tamil Nadu)

40 IND

Synonyms: hidden

= Diasperus mishmiensis (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion mishmiense Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 327 (1887)

= Phyllanthus velutinus var. mishmiensis (Hook.f.) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

variant :

var. tirupathiense (Rasingam, Chorghe, Prasanna & Sankara Rao) Chakr. & N.P.Balakr.; Indo-Burmese Phyllanthaceae 221 (2018)

Distribution: India (Andhra Pradesh)

40 IND

Synonyms: hidden

= Glochidion tirupathiense Rasingam, Chorghe, Prasanna & Sankara Rao; Taiwania 59(1): 9 (2014)

= Phyllanthus velutinus var. tirupathiensis (Rasingam, Chorghe, Prasanna & Sankara Rao) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion hivaoaense J.Florence; Fl. Polynésie Franç. [Florence] 1: 74 (1997)

Distribution: Marquesas Isl. (Hiva Oa, Tahuata)

61 MRQ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus hivaoaensis (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 75 (2011)

species :

Glochidion hohenackeri (Müll.Arg.) Bedd.; Forester´s Man. Bot. 193 (1872)

Distribution: S-India (Goa, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu)

40 IND

Synonyms: hidden

= Bradleia lanceolaria Wall.; Numer. List [Wallich] no. 7855 (1832)

= Bridelia sinica J.Graham; Cat. Pl. Bombay [Graham] 179 (1839)

= Diasperus fagifolius (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus hohenackeri (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus ralphii (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion fagifolium (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5(14): 312-313 (1887)

= Glochidion hohenackeri var. kothayarense Jothi & Manickam; Indian J. Forest. 27(1): 41 (2004)

= Glochidion ralphii Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 314 (1887)

= Phyllanthus fagifolius Müll.Arg.; Flora 48: 373 (1865)

= Phyllanthus fagifolius var. caesius Müll.Arg.; Flora 48: 373 (1865)

= Phyllanthus hohenackeri Müll.Arg.; Flora 48: 373 (1865)

variant :

var. johnstonei (Hook.f.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 210 (1995)

Distribution: S-India (Karnataka, Kerala)

40 IND

Synonyms: hidden

= Diasperus johnstonei (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion johnstonei Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 314 (1887)

= Phyllanthus hohenackeri var. johnstonei (Hook.f.) N.P.Balakr. & Chakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion hosokawae Fosberg; Willdenowia 20(1-2): 261 (1991)

Distribution: Fed. States of Micronesia

62 MIC

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus hosokawae (Fosberg) W.L.Wagner & Lorence; PhytoKeys 4: 75 (2011)

species :

Glochidion huahineense J.Florence; Fl. Polynésie Franç. [Florence] 1: 74 (1997)

Distribution: Society Isl. (Huahine)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus huahineensis (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 77 (2011)

species :

Glochidion humile Merr.; Philipp. J. Sci. 16: 544 (1920)

Distribution: Philippines (Mindanao)

42 PHI

Synonyms: hidden

= Phyllanthus mindanaoanthus Govaerts & Christenh.; Nelumbo 68(1): 142 (2026), nom. nov.

species :

Glochidion huntii Airy Shaw; Kew Bull. 27(1): 61 (1971)

Distribution: Solomon Isl.

43 SOL

Synonyms: hidden

= Phyllanthus solomonanthus Govaerts & Christenh.; Nelumbo 68(1): 146 (2026), nom. nov.

species :

Glochidion hylandii Airy Shaw; Kew Bull. 31(2): 347 (1976)

Distribution: Australia (Queensland)

50 AUS

Synonyms: hidden

= Phyllanthus hylandii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 140 (2026)

species :

Glochidion impuber (Roxb.) Govaerts; World Checkl. & Bibliogr. Euphorbiaceae 951 (2000)

Distribution: Moluccas

42 MOL

Synonyms: hidden

= Agyneia impuber (Roxb.) Miq.; Fl. Ned. Ind. 1(2): 367 (1859)

= Bradleia impuber Roxb.; Hort. Bengal. 104 (1814), nomen, Fl. Ind. [Roxburgh] 3: 698 (1832)

= Diasperus lambertianus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion lambertianum Müll.Arg.; Linnaea 32: 60 (1863)

= Phyllanthus impuber (Roxb.) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

= Phyllanthus lambertianus (Müll.Arg.) Müll.Arg.; Flora 48: 372 (1865)

species :

Glochidion insectum Airy Shaw; Kew Bull. 27(1): 54 (1972)

Distribution: Sulawesi; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.

42 NWG SUL 43 BIS

Synonyms: hidden

= Phyllanthus insectus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion insigne (Müll.Arg.) J.J.Sm. ex Koord. & Valeton; Meded. Dept. Landb. Ned.-Indië 10: 160 (1910)

Distribution: Sumatra; Sulawesi; Java

42 SUL

Synonyms: hidden

= Phyllanthus insignis Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1271 (1866)

= Diasperus insignis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

species :

Glochidion insulanum Müll.Arg.; Linnaea 32: 67 (1863)

Distribution: Moluccas

42 MOL

Synonyms: hidden

= Diasperus insulanus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus insulanus (Müll.Arg.) Müll.Arg.; Flora 48: 387 (1865)

species :

Glochidion intercastellanum Airy Shaw; Kew Bull. 23: 24 (1969)

Distribution: New Guinea (d´Entrecasteaux Arch.)

42 NWG

Synonyms: hidden

= Phyllanthus intercastellanus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion inusitatum A.C.Sm.; Fl. Vit. Nova 2: 493, 486 (1981)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus inusitatus (A.C.Sm.) W.L.Wagner & Lorence; PhytoKeys 4: 77 (2011)

species :

Glochidion jarawae Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 26(3): 527 (2002)

Distribution: Andaman Isl. (South Andaman Isl.)

41 AND

Synonyms: hidden

= Phyllanthus jarawae (Chakr. & N.P.Balakr.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion kanehirae Hosok.; Trans. Nat. Hist. Soc. Taiwan 25: 22 (1935)

Distribution: Palau (Urukthapel); Fed. States of Micronesia (Yap, Truk, Moen, Tol, Udot, Uman)

62 MIC PLU

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus kanehirae (Hosok.) W.L.Wagner & Lorence; PhytoKeys 4: 77 (2011)

species :

Glochidion karnaticum Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 211 (1995)

Distribution: India (Karnataka)

40 IND

Synonyms: hidden

= Phyllanthus karnaticus (Chakr. & M.Gangop.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion katikii Airy Shaw; Kew Bull. 32(2): 373 (1978)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus katikii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion kaweesakii (Pornp., Chantar. & J.Parn.) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: NE-Thailand

41 THA

Remarks: subgen. Phyllanthodendron

Synonyms: hidden

= Phyllanthus kaweesakii Pornp., Chantar. & J.Parn.; Botany (Ottawa) 95(6): 572 (2017)

species :

Glochidion kerangae Airy Shaw; Kew Bull. 27(1): 18 (1972)

Distribution: Borneo (Sarawak, Sabah); Brunei

42 BOR BRN

Synonyms: hidden

= Phyllanthus kerangae (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion kerrii Craib; Bull. Misc. Inform. Kew 1911: 458 (1911)

Distribution: Thailand; Cambodia

41 CBD THA

Synonyms: hidden

= Glochidion dasystylum var. kerrii (Craib) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 13(3): 710 (1989)

= Phyllanthus craibii Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009), nom. nov.

species :

Glochidion khasicum (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5(14): 324 (1887)

Distribution: China (Guangxi, Yunnan); Bhutan; India (Arunachal Pradesh, Assam, Meghalaya, Sikkim, West Bengal); Bangladesh

36 CHI 40 BAN BHU IND

Synonyms: hidden

= Phyllanthus khasicus Müll.Arg.; Flora 48: 389 (1865)

= Diasperus khasicus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

variant :

var. bilobulatum (Vasudeva Rao & Chakr.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 213 (1995)

Distribution: Andaman Isl. (North Andaman Isl.)

41 AND

Synonyms: hidden

= Glochidion bilobulatum Vasudeva Rao & Chakr.; J. Econ. Taxon. Bot. 5(4): 936 (1984)

= Glochidion kingii M.V.Ramana, Sanjappa, Venu & Chorghe; Nordic J. Bot. 33(1): 38 (2014)

= Phyllanthus khasicus var. bilobulatus (Vasudeva Rao & Chakr.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

species :

Glochidion kopiaginis Airy Shaw; Kew Bull. 33(1): 30 (1978)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus kopiaginis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion korthalsii (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

Distribution: Borneo

42 BOR

Synonyms: hidden

= Diasperus korthalsii (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus korthalsii Müll.Arg.; Flora 48: 377 (1865)

species :

Glochidion kostermansii Airy Shaw; Kew Bull. 27(1): 29 (1972)

Distribution: Lesser Sunda Isl. (Sumbawa, Flores)

42 LSI

Synonyms: hidden

= Phyllanthus tomentellus Govaerts & Christenh.; Nelumbo 68(1): 147 (2026), nom. nov.

species :

Glochidion kunstlerianum Gage; Rec. Bot. Surv. India 9: 220 (1922)

Distribution: peninsular Malaysia (Perak, Selangor, Johor); +Singapore; Sumatra; Borneo

42 BOR MLY SNG

Synonyms: hidden

= Phyllanthus kunstlerianus (Gage) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion lalae Airy Shaw; Kew Bull. 23: 21 (1969)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus lalae (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion lambiricum Airy Shaw; Kew Bull. 36(3): 599 (1981)

Distribution: Borneo (Sarawak)

42 BOR

Synonyms: hidden

= Phyllanthus lambiricus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion lanceilimbum Merr.; Philipp. J. Sci. 26: 462 (1925)

Distribution: Borneo; Sulawesi; Philippines (Mindanao); Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Solomon Isl.; Australia (Queensland)

42 BOR MOL NWG PHI SUL 43 BIS SOL 50 AUS

Synonyms: hidden

= Phyllanthus lanceilimbus (Merr.) Merr.; Philipp. J. Sci. 30: 402 (1926)

species :

Glochidion lanceolarium (Roxb.) Voigt; Hort. Suburb. Calcutt. 153 (1845)

Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Yunnan); Ryukyu Isl.; Nepal; India (Assam, Chhattisgarh, Jharkhand, Karnataka, Meghalaya, Odisha, Sikkim,Uttarakhand, West Bengal); Bangladesh; Myanmar; Thailand; Laos; Cambodia; Vietnam

36 CHI 38 NNS 40 BAN IND NEP 41 CBD LAO MYA THA VIE

Synonyms: hidden

= Bradleja lanceolaria Roxb.; Fl. Ind. [Roxburgh] 3: 697-698 (1832)

= Diasperus benthamianus Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus lanceolarius (Roxb.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion cantoniense Hance; Ann. Sci. Nat., Bot., Sér. 5, 5: 241 (1866)

= Glochidion macrophyllum Benth.; London J. Bot. 1: 491 (1842)

= Glochidion subsessile var. birmanicum Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 13: 716 (1989)

= Glochisandra acuminata Wight; Icon. Pl. Ind. Orient. [Wight] 5(2): 28, t. 1905 (1852)

= Phyllanthus benthamianus Müll.Arg.; Flora 48: 371 (1865), nom. illeg.

= Phyllanthus fraxinifolius G.Lodd.; Bot. Cab. 9: t. 839 (1824), nom. subnud.

= Phyllanthus lanceolarius (Roxb.) Müll.Arg.; Flora 48: 371 (1865)

species :

Glochidion lanceolatum Hayata; J. Coll. Sci. Imp. Univ. Tokyo 20: 16, t. 20 (1904)

Distribution: Taiwan (incl. Lan Yu Isl.)

38 TAI

Synonyms: hidden

= Glochidion kotoense Hayata; Icon. Pl. Formosan. 9: 96 (1920)

= Glochidion kusukusense Hayata; Icon. Pl. Formosan. 9: 96 (1920)

= Glochidion sphaerostigmum Hayata; Icon. Pl. Formosan. 9: 96-97 (1920)

= Glochidion zeylanicum var. lanceolatum (Hayata) M.J.Deng & J.C.Wang; Fl. Taiwan, ed. 2, 3: 480 (1993)

= Phyllanthus keelungensis T.Kuros.; J. Jap. Bot. 91(2): 119 (2016), nom. nov.

= Phyllanthus kotoensis (Hayata) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion lancisepalum Merr.; J. Straits Branch Roy. Asiat. Soc. 86: 319 (1922)

Distribution: Borneo

42 BOR

Synonyms: hidden

= Phyllanthus lancisepalus (Merr.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

species :

Glochidion lanyuense Gang Yao & S.X.Luo; PhytoKeys 159: 147, figs. 4, 5B, D, F, H, J, K, M (2020)

Distribution: Taiwan (Lan Yu Isl.)

38 TAI

Synonyms: hidden

= Phyllanthus lanyuensis (Gang Yao & S.X.Luo) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion latistylum C.B.Rob.; Philipp. J. Sci., C 4: 93 (1909)

Distribution: Philippines (Mindanao)

42 PHI

Synonyms: hidden

= Phyllanthus latistylus (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion lativenium (Croizat) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: China (Guizhou)

36 CHI

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus lativenius (Croizat) Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 177 (1996)

= Phyllanthodendron lativenium Croizat; J. Arnold Arbor. 23: 36 (1942)

species :

Glochidion leptostylum Airy Shaw; Kew Bull. 27(1): 37 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus leptostylus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion leucocarpum Airy Shaw; Kew Bull. 36(2): 303 (1981)

Distribution: Sumatra

Error

Synonyms: hidden

= Phyllanthus leucocarpus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion leucogynum Miq.; Fl. Ned. Ind. 1(2): 376 (1859)

Distribution: Java

Error

Synonyms: hidden

= Diasperus leucogynus (Miq.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus leucogynus (Miq.) Müll.Arg.; Flora 48: 75 (1865)

species :

Glochidion liangkwangense Gang Yao & Z.Q.Song; Phytotaxa 677(1): 96 (2024)

Distribution: China (Guangdong, Guangxi)

36 CHI

Synonyms: hidden

= Phyllanthus liangkwangense (Gang Yao & Z.Q.Song) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion ligulatum C.B.Rob.; Philipp. J. Sci., C 6: 329 (1911)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus ligulatus (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion lingulatum (Beille) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: N-Thailand; Laos; Cambodia; N-Vietnam; S-Vietnam

41 CBD LAO THA VIE

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus lingulatus Beille; Bull. Soc. Bot. France 72: 161 (1925), et Fl. Indo-Chine [P.H.Lecomte et al.] 5: 592 (1927)

= Phyllanthodendron lingulatum (Beille) Croizat; J. Arnold Arbor. 23: 34 (1942)

species :

Glochidion littorale Blume; Bijdr. Fl. Ned. Ind. 585 (1826)

Distribution: S-India (Kerala, Tamil Nadu); Maldives; Sri Lanka; Thailand; Cambodia; Vietnam; peninsular Malaysia (Perak, Pahang, Johor); Singapore; Sumatra; Borneo; Brunei; Sulawesi; Palawan; Philippines (Luzon, Polillo, Mindoro, Panay, Leyte, Jolo, Mindanao); Java; Palau (Koror)

40 IND MDV SRL 41 CBD THA VIE 42 BOR BRN MLY PHI PLW SNG SUL 62 PLU

Synonyms: hidden

= Agyneia impuber Benth.; Numer. List [Wallich] no. 7869 (1847), pro syn.

= Bradleia littorea (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Bradleia obtusa Benth.; Numer. List [Wallich] no. 7869 (1847)

= Diasperus litoralis (Blume) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion littorale var. caudatum Airy Shaw; Kew Bull. 29(2): 290 (1974)

= Glochidion littorale var. culminicola Airy Shaw; Kew Bull. 29(2): 290 (1974)

= Phyllanthus littoralis (Blume) Müll.Arg.; Flora 48: 370 (1865)

species :

Glochidion lobocarpum (Benth.) F.M.Bailey; Queensl. Fl. 5: 1424 (1902)

Distribution: New Guinea (Papua New Guinea); Australia (Queensland)

42 NWG 50 AUS

Synonyms: hidden

= Phyllanthus lobocarpus Benth.; Fl. Austral. [Bentham] 6: 97 (1873)

species :

Glochidion loerzingii Airy Shaw; Kew Bull. 23: 9 (1969)

Distribution: Sumatra

Error

Synonyms: hidden

= Phyllanthus loerzingii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion longfieldiae (Riley) F.Br.; Bull. Bernice P. Bishop Mus. 130: 141 (1935)

Distribution: Austral Isl. (Rapa Iti)

61 AUI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus longfieldiae Riley; Bull. Misc. Inform. Kew 1926: 55 (1926)

species :

Glochidion longistylum C.B.Rob.; Philipp. J. Sci., C 4: 96 (1909)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus zambales Govaerts & Christenh.; Nelumbo 68(1): 148 (2026), nom. nov.

species :

Glochidion lucidum Blume; Bijdr. Fl. Ned. Ind. 584 (1826)

Distribution: peninsular Malaysia (Terengganu, Pahang); Sumatra; Borneo; Sulawesi; Philippines; Java; Moluccas; New Guinea (Irian Jaya)

42 BOR MLY MOL NWG PHI SUL

Synonyms: hidden

= Bradleia lucida (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Diasperus lucidus (Blume) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus lucidus (Blume) Müll.Arg.; Flora 48: 370 (1865)

species :

Glochidion lutescens Blume; Bijdr. Fl. Ned. Ind. 585 (1825)

Distribution: China (Guangdong, Guizhou, Sichuan, Yunnan); Myanmar; Thailand; Laos; Vietnam; peninsular Malaysia; Singapore; Sumatra; Borneo; Brunei; Sulawesi; Palawan; Philippines (Luzon, Mindoro, Negros, Leyte, Dinagat); Java; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.

36 CHI 41 LAO MYA THA VIE 42 BOR BRN MLY MOL NWG PHI PLW SNG SUL 43 BIS

Synonyms: hidden

= Anisonema hypoleucum Miq.; Fl. Ned. Ind., Eerste Bijv. 449 (1861)

= Bradleia laevigata Benth.; Numer. List [Wallich] no. 7853 (1847), nomen

= Bradleia lutescens (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Diasperus glaucifolius (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus kollmannianus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus laevigatus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus lutescens (Blume) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Glochidion breynioides C.B.Rob.; Philipp. J. Sci., C 4(1): 95-96 (1909)

= Glochidion glaucifolium Müll.Arg.; Linnaea 32: 65 (1863), et Linnaea 34: 65 (1865 publ. 1866)

= Glochidion hollandianum J.J.Sm.; Nova Guinea 12: 544, t. 228A (1917)

= Glochidion hypoleucum (Miq.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 275 (1900)

= Glochidion kollmannianum (Müll.Arg.) J.J.Sm.; Bijdr. Boomsoort. Java 12: 166 (1910)

= Glochidion laevigatum (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 319 (1887)

= Glochidion pusilliflorum S.Moore; J. Bot. 63(Suppl.): 93 (1925)

= Phyllanthus glaucifolius Wall.; Numer. List [Wallich] no. 7923 (1832), nomen

= Phyllanthus hypoleucus (Miq.) Müll.Arg.; Flora 48: 374 (1865), nom. illeg.

= Phyllanthus kollmannianus Müll.Arg.; Flora 48: 378 (1865)

= Phyllanthus laevigatus Müll.Arg.; Flora 48: 374 (1865)

= Phyllanthus lutescens (Blume) Müll.Arg.; Flora 48: 77 (1865)

species :

Glochidion luzonense Elmer; Leafl. Philipp. Bot. 1: 301 (1908)

Distribution: Sulawesi; Philippines (Luzon)

42 PHI SUL

Synonyms: hidden

= Phyllanthus aeta Govaerts & Christenh.; Nelumbo 68(1): 136 (2026), nom. nov.

species :

Glochidion macphersonii Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 175 (1996), nom. nov.

Distribution: New Caledonia

60 NWC

Synonyms: hidden

= Bradleia macrophylla Labill.; Sert. Austro-Caledon. 77, t. 78 (1825)

= Diasperus macrophyllus (Labill.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion macrophyllum (Labill.) Müll.Arg.; Linnaea 32: (1863) 61 (non Benth.)

= Phyllanthus macrophyllus (Labill.) Müll.Arg.; Flora 48: 370 (1865)

species :

Glochidion macrocarpum Blume; Bijdr. Fl. Ned. Ind. 584 (1826)

subspecies :

subsp. macrocarpum

Distribution: Sumatra; Java

Error

Synonyms: hidden

= Bradleia macrocarpa (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Diasperus macrocarpus (Blume) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Phyllanthus macrocarpus (Blume) Müll.Arg.; Flora 48: 370 (1865)

= Glochidion macrocarpum f. depressoglobosum J.J.Sm.; Bijdr. Boomsoort. Java 12: 156 (1910)

variant :

var. sumbawanum Airy Shaw; Kew Bull. 33(1): 28 (1978)

Distribution: Lesser Sunda Isl. (Sumbawa)

42 LSI

Synonyms: hidden

= Phyllanthus macrocarpus subsp. sumbawanus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

subspecies :

subsp. orientale Airy Shaw; Kew Bull. 23: 12 (1969)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus macrocarpus subsp. orientalis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 141 (2026)

species :

Glochidion macrosepalum Hosok.; Trans. Nat. Hist. Soc. Taiwan 25: 21 (1935)

Distribution: Palau (Babeldaob)

62 PLU

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus macrosepalus (Hosok.) W.L.Wagner & Lorence; PhytoKeys 4: 78 (2011)

species :

Glochidion macrostigma Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 313 (1887)

Distribution: peninsular Malaysia (Kedah, Penang, Perak); Sumatra; Borneo; Java

42 BOR MLY

Synonyms: hidden

= Diasperus macrostigma (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion capitatum J.J.Sm. ex Koord. & Valeton; Meded. Dept. Landb. Ned.-Indië 10: 133 (1910)

= Phyllanthus macrostigma (Hook.f.) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion maingayi Gage; Rec. Bot. Surv. India 9: 221 (1922)

Distribution: peninsular Malaysia

42 MLY

Synonyms: hidden

= Phyllanthus maingayi (Gage) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion malindangense C.B.Rob.; Philipp. J. Sci., C 4: 101 (1909)

Distribution: Philippines (Mindanao)

42 PHI

Synonyms: hidden

= Phyllanthus malindangensis (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion malingheense C.Y.Deng & Gang Yao; Phytotaxa 561(1): 105 (2022)

Distribution: China (Guizhou)

36 CHI

Synonyms: hidden

= Phyllanthus malingheensis (C.Y.Deng & Gang Yao) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion manono Baill. ex Müll.Arg.; Linnaea 32: 65 (1863)

Distribution: Fiji; Society Isl. (Moorea, Tahiti, Bora Bora)

60 FIJ 61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus manono (Baill.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus manono (Baill.) Müll.Arg.; Flora 48: 377 (1865)

species :

Glochidion marchionicum F.Br.; Bull. Bernice P. Bishop Mus. 130: 142 (1935)

Distribution: Marquesas Isl. (Nuku Hiva, Ua Huka, Ua Pou, Hiva Oa, Tahuata, Fatu Hiva)

61 MRQ

Remarks: subgen. Glochidion

Synonyms: hidden

= Glochidion ×marquesanum (F.Br.) Croizat; Sargentia 1: 48 (1942)

= Glochidion ramiflorum var. marquesanum F.Br.; Bull. Bernice P. Bishop Mus. 130: 144 (1935)

= Glochidion tooviianum J.Florence; Bull. Mus. Natl. Hist. Nat., B, Adansonia 18: 260 (1996)

= Phyllanthus marchionicus (F.Br.) W.L.Wagner & Lorence; PhytoKeys 4: 79 (2011)

species :

Glochidion marianum Müll.Arg.; Linnaea 32: 65 (1863)

Distribution: Guam; Fed. States of Micronesia (Pohnpei)

62 GUM MIC

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus mariannensis W.L.Wagner & Lorence; PhytoKeys 4: 79 (2011), nom. nov.

species :

Glochidion medogense T.L.Chin; Acta Phytotax. Sin. 18(2): 251, f. 1, 1-3 (1980)

Distribution: SE-Tibet

36 TIB

Synonyms: hidden

= Phyllanthus medogensis (T.L.Chin) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion mehipitense Pax & K.Hoffm.; Mitt. Inst. Allg. Bot. Hamburg 7: 225 (1931)

Distribution: Borneo

42 BOR

Synonyms: hidden

= Phyllanthus mehipitensis (Pax & K.Hoffm.) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion meijeri Airy Shaw; Kew Bull. 27(1): 11 (1972)

Distribution: Sumatra

Error

Synonyms: hidden

= Glochidion physocarpum Airy Shaw; Kew Bull. 32(2): 370 (1978)

= Phyllanthus meijeri (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion melvilleorum Airy Shaw; Kew Bull. 25(3): 487 (1971), as ´melvilliorum´

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus melvilleorum (Airy Shaw) W.L.Wagner & Lorence; PhytoKeys 4: 81 (2011), as ´melvilliorum´

species :

Glochidion merrillii C.B.Rob.; Philipp. J. Sci., C 4: 100 (1909)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus merrillii (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 142 (2026)

species :

Glochidion mindorense C.B.Rob.; Philipp. J. Sci., C 4: 98 (1909)

Distribution: Borneo; Sulawesi; Palawan; Philippines (Luzon, Polillo, Mindoro, Leyte, Samar); Java; Lesser Sunda Isl.; Moluccas

42 BOR LSI MOL PHI PLW SUL

Synonyms: hidden

= Phyllanthus mangyananthus Govaerts & Christenh.; Nelumbo 68(1): 142 (2026), nom. nov.

species :

Glochidion minutiflorum (Ridl.) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: SW- & peninsular Thailand; peninsular Malaysia (Kelantan, Perak)

41 THA 42 MLY

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Cleistanthus minutiflorus Ridl.; J. Straits Branch Roy. Asiat. Soc. 59: 169 (1911)

= Phyllanthodendron coriaceum Gage; Rec. Bot. Surv. India 9: 219 (1922)

= Phyllanthodendron minutiflorum (Ridl.) Airy Shaw; Kew Bull. 14: 471 (1960)

= Phyllanthus ridleyanus Airy Shaw; Kew Bull. 26(2): 323 (1972), nom. nov.

species :

Glochidion mirabilis (Müll.Arg.) R.W.Bouman; Phytotaxa 540(1): 64 (2022)

Distribution: Thailand; Laos

41 LAO THA

Remarks: subgen. Phyllanthodendron

Synonyms: hidden

= Phyllanthus mirabilis Müll.Arg.; Flora 47: 513 (1864), et Prodr. [A.P. de Candolle] 15(2): 355 (1866)

= Diasperus mirabilis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthodendron mirabile (Müll.Arg.) Hemsl.; Hooker´s Icon. Pl. 26: t. 2563-2564 (1898)

species :

Glochidion mitrastylum Airy Shaw; Kew Bull. 33(1): 31 (1978)

Distribution: New Guinea (Papua New Guinea); Bismarck Arch. (New Britain)

42 NWG 43 BIS

Synonyms: hidden

= Phyllanthus mitrastylus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion moi (P.T.Li) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: China (Guangxi)

36 CHI

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus moi P.T.Li; Guihaia 3(3): 167-168, f. 1 (1983)

= Phyllanthodendron moi (P.T.Li) P.T.Li; Bull. Bot. Res., Harbin 7(3): 9 (1987)

species :

Glochidion molle Blume; Bijdr. Fl. Ned. Ind. 586 (1826)

Distribution: Vietnam; Borneo; Sulawesi; Palawan; Philippines (Jolo, Basilan, Mindanao); Java; Lesser Sunda Isl. (Bali); Moluccas

41 VIE 42 BOR LSI MOL PHI PLW SUL

Synonyms: hidden

= Bradleia blumei Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841), nom. superfl.

= Bradleia mollis (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Diasperus mollis (Blume) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion villosum Miq.; Fl. Ned. Ind. 1(2): 376 (1859)

= Phyllanthus mollis (Blume) Müll.Arg.; Flora 48: 387 (1865)

= Phyllanthus tomentosus Noronha; Verh. Batav. Genootsch. Kunst. 5(4): 22 (1790), nomen

species :

Glochidion moluccanum Blume; Bijdr. Fl. Ned. Ind. 587 (1825)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Bradleia moluccana (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Diasperus moluccanus (Blume) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus moluccanus (Blume) Müll.Arg.; Flora 48: 380 (1865)

variant :

var. glabrescens Airy Shaw; Kew Bull. 27(1): 25 (1972)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus moluccanus var. glabrescens (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion monostylum Airy Shaw; Kew Bull. 29(2): 288 (1974)

Distribution: Borneo (Sabah)

42 BOR

Synonyms: hidden

= Phyllanthus monostylus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion moonii Thwaites; Enum. Pl. Zeyl. [Thwaites] 286 (1859)

Distribution: Sri Lanka

40 SRL

Synonyms: hidden

= Diasperus glaucogynus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus moonii (Thwaites) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus symplocoides (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion glaucogynum (Müll.Arg.) Bedd.; Forester´s Man. Bot. 195 (1872)

= Glochidion montanum Thwaites; Enum. Pl. Zeyl. [Thwaites] 286 (1859)

= Glochidion moonii f. glaucogynum (Müll.Arg.) Alston; Ann. Roy. Bot. Gard. (Peradeniya) 11: 6 (1928)

= Glochidion moonii var. glaucogynum (Müll.Arg.) Trimen; Handb. Fl. Ceylon 4: 52 (1898)

= Glochidion nemorale var. moonii (Thwaites) Chakr. & N.P.Balakr.; Indo-Burmese Phyllanthaceae 236 (2018)

= Glochidion symplocoides (Müll.Arg.) Bedd.; Forester´s Man. Bot. 195 (1872)

= Gynoon hirsutum Wight; Icon. Pl. Ind. Orient. [Wight] t. 1909 (1852)

= Phyllanthus glaucogynus Müll.Arg.; Flora 48: 389 (1865)

= Phyllanthus moonii (Thwaites) Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 312 (1866)

= Phyllanthus pubescens Moon; Cat. Pl. Ceylon. 65 (1824)

= Phyllanthus srilankanus Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009), nom. nov.

= Phyllanthus symplocoides Müll.Arg.; Flora 48: 389 (1865)

species :

Glochidion moorei P.T.Li; Acta Phytotax. Sin. 20(1): 117 (1982), nom. nov.

Distribution: Society Isl. (Raiatea)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Glochidion salicifolium J.W.Moore; Bull. Bernice P. Bishop Mus. 226: 13 (1963), nom. illeg.

= Phyllanthus raiateaensis W.L.Wagner & Lorence; PhytoKeys 4: 84 (2011), nom. nov.

species :

Glochidion mop Airy Shaw; Kew Bull. 27(1): 50 (1972)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus mop (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion muelleri Briq.; Annuaire Conserv. Jard. Bot. Genève 4: 225 (1900)

Distribution: Java

Error

Synonyms: hidden

= Phyllanthus muelleri (Briq.) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion multilobum A.C.Sm.; Fl. Vit. Nova 2: 493, 484 (1981)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus multilobus (A.C.Sm.) W.L.Wagner & Lorence; PhytoKeys 4: 81 (2011)

species :

Glochidion multiloculare (Roxb. ex Willd.) Voigt; Hort. Suburb. Calcutt. 152 (1845)

Distribution: Nepal; India (Arunachal Pradesh, Assam, Bihar, Chattisgarh, Jharkhand, Manipur, Meghalaya, Nagaland, Tripura, Sikkim, Uttar Pradesh, West Bengal); Bangladesh; N-Myanmar

40 BAN IND NEP 41 MYA

Synonyms: hidden

= Agyneia multilocularis Rottler ex Willd.; Neue Schriften Ges. Naturf. Freunde Berlin 4: 206 (1803)

= Agyneia pubera Benth.; Numer. List [Wallich] no. 7870 (1847)

= Bradleia multilocularis (Rottler ex Willd.) Roxb.; Hort. Bengal. 69 (1814)

= Bradleia multilocularis (Rottler ex Willd.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 3: 19 (1826), isonym

= Diasperus multilocularis (Rottler ex Willd.) Kuntze; Revis. Gen. Pl. 600 et 602 (1891)

= Phyllanthus multilocularis (Rottler ex Willd.) Müll.Arg.; Flora 48: 370 (1865)

variant :

var. pubescens Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 14(3): 720 (1990)

Distribution: N-India (Sikkim, Assam)

40 IND

Synonyms: hidden

= Phyllanthus multilocularis var. pubescens (Chakr. & M.Gangop.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

species :

Glochidion muscisilvae Airy Shaw; Kew Bull. 33(1): 29 (1978)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus muscisilvae (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion myrtifolium J.W.Moore; Bull. Bernice P. Bishop Mus. 226: 10 (1963)

Distribution: Society Isl. (Raiatea, Tahaa)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Glochidion longipedicellatum J.W.Moore; Bull. Bernice P. Bishop Mus. 226: 9 (1963), nom. illeg.

= Glochidion longipes P.T.Li; Acta Phytotax. Sin. 20(1): 117 (1982), nom. nov.

= Phyllanthus st-johnii W.L.Wagner & Lorence; PhytoKeys 4: 86 (2011), nom. nov.

species :

Glochidion nadeaudii J.Florence; Bull. Mus. Natl. Hist. Nat., B, Adansonia 18: 253, fig. 4 (1996)

Distribution: Society Isl. (Moorea)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus nadeaudii (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 81 (2011)

species :

Glochidion namilo Guillaumin; Bull. Mus. Natl. Hist. Nat., Sér. 2, 9: 300 (1937)

Distribution: Vanuatu

60 VAN

Synonyms: hidden

= Phyllanthus namilo (Guillaumin) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion nemorale Thwaites; Enum. Pl. Zeyl. [Thwaites] 286 (1859)

Distribution: Sri Lanka

40 SRL

Synonyms: hidden

= Diasperus gardneri (Thwaites) Kuntze; Revis. Gen. Pl. 2: 597 (1891)

= Diasperus nemoralis (Thwaites) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion acutifolium Alston; Ann. Roy. Bot. Gard. (Peradeniya) 11: 7 (1928)

= Glochidion gardneri Thwaites; Enum. Pl. Zeyl. [Thwaites] 286 (1859)

= Glochidion gardneri var. acuminatum Trimen; Handb. Fl. Ceylon 4: 31 (1898)

= Glochidion leptogynum (Müll.Arg.) Bedd.; Forester´s Man. Bot. 195 (1872)

= Glochidion montanum f. subglabrum (Trimen) Alston; Ann. Roy. Bot. Gard. (Peradeniya) 11: 6 (1928)

= Glochidion montanum var. glaberrimum Alston; Ann. Roy. Bot. Gard. (Peradeniya) 11: 6 (1928)

= Glochidion moonii var. subglabrum Trimen; Handb. Fl. Ceylon 4: 31 (1898)

= Phyllanthus leptogynus Müll.Arg.; Flora 48: 389 (1865)

= Phyllanthus moonii var. subglaber (Trimen) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

= Phyllanthus nemoralis (Thwaites) Müll.Arg.; Flora 48: 389 (1865)

species :

Glochidion nervosum Alston; Handb. Fl. Ceylon 6: Suppl., 260 (1931), in clavi.

Distribution: Sri Lanka

40 SRL

Synonyms: hidden

= Phyllanthus sinhalanthus Govaerts & Christenh.; Nelumbo 68(1): 146 (2026), nom. nov.

species :

Glochidion nesophilum Airy Shaw; Kew Bull. 27(1): 45 (1972)

Distribution: New Guinea (d´Entrecasteaux Isl., Louisiade Arch.); Bismarck Arch.

42 NWG 43 BIS

Synonyms: hidden

= Phyllanthus nesophilus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion nobile Airy Shaw; Kew Bull. 32(2): 371 (1978)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus boumanii Govaerts & Christenh.; Nelumbo 68(1): 137 (2026), nom. nov.

species :

Glochidion nothofageticum Airy Shaw; Kew Bull. 34(3): 593 (1980)

Distribution: Bismarck Arch. (New Britain)

43 BIS

Synonyms: hidden

= Phyllanthus nothofageticus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion novae-georgiae Airy Shaw; Kew Bull. 23: 23 (1969)

Distribution: Bismarck Arch. (New Georgia); Solomon Isl.

43 BIS SOL

Synonyms: hidden

= Phyllanthus novae-georgiae (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion novoguineense K.Schum.; Nachtr. Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 287 (1905)

Distribution: Sulawesi; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.

42 MOL NWG SUL 43 BIS

Synonyms: hidden

= Glochidion globosum J.J.Sm.; Nova Guinea 8: 783 (1912)

= Glochidion platycladum Airy Shaw; Kew Bull. 23: 18 (1969)

= Glochidion vinkiorum Airy Shaw; Kew Bull. 23: 19 (1969)

= Phyllanthus novoguineensis (K.Schum.) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

= Phyllanthus submollis var. glabra Lauterb. & K.Schum.; Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 391 (1900)

species :

Glochidion nubigenum Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 315 (1887)

Distribution: SE-Tibet; Nepal; India (Arunachal Pradesh, Assam, Sikkim, West Bengal); Myanmar

36 TIB 40 IND NEP 41 MYA

Synonyms: hidden

= Diasperus nubigenus (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion metanubigenum Hurus. & Yu.Tanaka; Fl. E. Himalaya 177 (1966)

= Glochidion velutinum var. nubigenum (Hook.f.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 14(3): 720 (1990)

= Phyllanthus nubigenus (Hook.f.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

species :

Glochidion oblatum Hook.f.; Fl. Brit. India [J.D.Hooker] 5(14): 312 (1887)

Distribution: China (Yunnan); India (Assam, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Bangladesh; Myanmar; Thailand

36 CHI 40 BAN IND 41 MYA THA

Synonyms: hidden

= Diasperus oblatus Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Phyllanthus oblatus (Hook.f.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

species :

Glochidion oblongifolium Airy Shaw; Kew Bull. 33(1): 28 (1978)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus oblongifolius (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion obovatum Siebold & Zucc.; Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(2): 143 (1843)

Distribution: China (Fujian, Zhejiang); Japan (Honshu, Shikoku, Kyushu); Ryukyu Isl.; Taiwan

36 CHI 38 JAP NNS TAI

Synonyms: hidden

= Bradleia sinensis Siebold ex Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 128 (1867)

= Diasperus obovatus (Siebold & Zucc.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus obovatus (Siebold & Zucc.) Müll.Arg.; Flora 48: 387 (1865)

= Phyllanthus sieboldianus T.Kuros.; J. Jap. Bot. 91(2): 119 (2016), nom. nov.

species :

Glochidion obscurum (Roxb. ex Willd.) Blume; Bijdr. Fl. Ned. Ind. 585 (1826)

Distribution: Myanmar; S- & peninsular Thailand; Cambodia; Vietnam; peninsular Malaysia (common throughout); Sumatra; Borneo; Brunei; Sulawesi; Java; Lesser Sunda Isl. (Bali); Moluccas

41 CBD MYA THA VIE 42 BOR BRN LSI MLY MOL SUL

Synonyms: hidden

= Bradleia glaucophylla Hassk.; Cat. Hort. Bot. Bogor. [Hasskarl] 242 (1844)

= Bradleia kipareh Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Bradleia pinnata Roxb.; Hort. Bengal. 69 (1814), nom. superfl., et, Fl. Ind. [Roxburgh] 3: 700 (1832)

= Diasperus dasyanthus (Kurz ex Teijsm. & Binn.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus kipareh (Steud.) Kuntze; Revis. Gen. Pl. 599 et 602 (1891)

= Diasperus obscurus (Roxb. ex Willd.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus rhabdocarpus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus subobscurus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion blumeanum Müll.Arg.; Linnaea 32: 65 (1863)

= Glochidion dasyanthum Kurz ex Teijsm. & Binn.; Natuurk. Tijdschr. Ned.-Indië 27: 47 (1864)

= Glochidion glaucum Blume; Bijdr. Fl. Ned. Ind. 587 (1826)

= Glochidion obscurum var. macrocalyx J.J.Sm.; Bijdr. Boomsoort. Java 12: 130 (1910)

= Glochidion pinnatum (Roxb.) Voigt; Hort. Suburb. Calcutt. 153 (1845)

= Glochidion roxburghianum Müll.Arg.; Linnaea 32: 61 (1863)

= Phyllanthus dasyanthus (Kurz ex Teijsm. & Binn.) Müll.Arg.; Flora 48: 390 (1865)

= Phyllanthus kipareh (Steud.) Müll.Arg.; Flora 48: 373 (1865)

= Phyllanthus obscurus Roxb. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 581 (1805)

= Phyllanthus rhabdocarpus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1271 (1866)

= Phyllanthus subobscurus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1271 (1866)

species :

Glochidion oligotrichum (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 1(1): 3: 276 (1900)

Distribution: Java

Error

Synonyms: hidden

= Diasperus oligotrichus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus oligotrichus Müll.Arg.; Flora 48: 379 (1865)

species :

Glochidion oogynum Airy Shaw; Kew Bull. 27(1): 46 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus oogynus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion orbicularifolium (P.T.Li) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: China (Guangxi)

36 CHI

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus orbicularifolius (P.T.Li) Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 177 (1996)

= Phyllanthodendron orbicularifolium P.T.Li; Bull. Bot. Lab. N. E. Forest. Inst., Harbin 7(3): 5 (1987)

species :

Glochidion ornatum Kurz ex Teijsm. & Binn.; Tijdschr. Ned.-Indië 27: 46 (1864)

Distribution: Sumatra

Error

Synonyms: hidden

= Diasperus ornatus (Kurz ex Teijsm. & Binn.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus ornatus (Kurz ex Teijsm. & Binn.) Müll.Arg.; Flora 48: 390 (1865)

species :

Glochidion orohenense J.W.Moore; Occas. Pap. Bernice P. Bishop Mus. 16: 6 (1940)

Distribution: Society Isl. (Tahiti)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus orohenensis (J.W.Moore) W.L.Wagner & Lorence; PhytoKeys 4: 81 (2011)

species :

Glochidion oxycarpum (Müll.Arg.) R.W.Bouman; Blumea 63(2): 170 (2018)

Distribution: Sumatra

Error

Synonyms: hidden

= Diasperus oxycarpus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus oxycarpus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1270 (1866)

species :

Glochidion oxygonum Airy Shaw; Kew Bull. 23: 12 (1969)

Distribution: Moluccas (Halmaheira)

42 MOL

Synonyms: hidden

= Phyllanthus oxygonus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion pachyconum Airy Shaw; Kew Bull. 27(1): 52 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus pachyconus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion palauense Hosok.; Trans. Nat. Hist. Soc. Taiwan 25: 22 (1935)

Distribution: Palau (Ulebsechel)

62 PLU

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus otobedii W.L.Wagner & Lorence; PhytoKeys 4: 82 (2011), nom. nov.

species :

Glochidion palawanense Elmer; Leafl. Philipp. Bot. 4: 1290 (1911)

Distribution: Palawan

42 PLW

Synonyms: hidden

= Phyllanthus palawanensis (Elmer) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion paludicola (Airy Shaw) Airy Shaw; Kew Bull., Addit. Ser. 8: 111 (1980)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Glochidion mindorense subsp. paludicola Airy Shaw; Kew Bull. 27(1): 22 (1972)

= Phyllanthus paludicola (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion papenooense J.Florence; Bull. Mus. Natl. Hist. Nat., B, Adansonia 18: 253 (1996)

Distribution: Society Isl. (Tahiti)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus papenooensis (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 83 (2011)

species :

Glochidion peltiferum S.Moore; J. Bot. 63(Suppl.): 94 (1925)

Distribution: Sumatra

Error

Synonyms: hidden

= Phyllanthus peltiferum (S.Moore) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion petraeum (Li) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: China (SW-Guangxi)

36 CHI

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthodendron petraeum P.T.Li; Bull. Bot. Lab. N. E. Forest. Inst., Harbin 7(3): 4 (1987)

= Phyllanthus guanxiensis Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 176 (1996), nom. nov.

= Phyllanthus petraeus A.Chev. ex Beille; Compt. Rend. Hebd. Séances Acad. Sci. 145: 1294 (1907)

species :

Glochidion phellocarpum Airy Shaw; Kew Bull. 35(2): 385 (1980)

Distribution: peninsular Malaysia (Johor); Sumatra; ?Java

42 MLY

Synonyms: hidden

= Phyllanthus phellocarpus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion philippicum (Cav.) C.B.Rob.; Philipp. J. Sci., C 4(1): 103 (1909)

Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Sichuan, Yunnan); Taiwan; Sumatra (Enggano Isl.); Borneo; Sulawesi; Palawan; Philippines (Luzon, Mindanao); Java; Lesser Sunda Isl. (Bali, Timor, etc.); East Timor; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Solomon Isl.; Australia (Queensland); Vanuatu

36 CHI 38 TAI 42 BOR LSI MOL NWG PHI PLW SUL 43 BIS SOL 50 AUS 60 VAN

Synonyms: hidden

= Bradleia philippensis Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 592 (1805)

= Bradleia philippica Cav.; Icon. [Cavanilles] 4: 48 (1798)

= Coccoglochidion erythrococcus K.Schum.; Nachtr. Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 293 (1905)

= Diasperus compressicaulis (Kurz ex Teijsm. & Binn.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus kurzianus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus philippinensis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus quercinus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion compressicaule Kurz ex Teijsm. & Binn.; Natuurk. Tijdschr. Ned.-Indie 27: 45 (1864)

= Glochidion ferdinandi var. mollis (Benth.) F.M.Bailey; Queensl. Fl. 5: 1424 (1905)

= Glochidion formosanum Hayata; J. Coll. Sci. Imp. Univ. Tokyo 20(3): 20, t. 2G (1904)

= Glochidion philippinense Benth.; Fl. Hongk. 314 (1861), nom. illeg.

= Glochidion quercinum (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 1(2): 276 (1900)

= Glochidion sieboldianum (Miq.) Koidz.; Fl. Symb. Orient.-Asiat. 25 (1930)

= Phyllanthus compressicaulis (Kurz ex Teijsm. & Binn.) Müll.Arg.; Flora 48: 376 (1865)

= Phyllanthus ferdinandi var. mollis Benth.; Fl. Austral. [Bentham] 6: 97 (1873)

= Phyllanthus kurzianus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1272 (1866)

= Phyllanthus philippica (Cav.) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

= Phyllanthus philippinensis (Willd.) Müll.Arg.; Flora 48: 376 (1865)

= Phyllanthus quercinus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 306 (1866)

species :

Glochidion phyllanthoides Merr.; Philipp. J. Sci. 16: 541 (1920)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus phyllanthoides (Merr.) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion phyllochlamys Airy Shaw; Kew Bull. 33(1): 30 (1978)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus phyllochlamys (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion pilosum (Lour.) Merr.; Trans. Amer. Philos. Soc., n. s. 24(2): 10, 232 (1935)

Distribution: S-Vietnam

41 VIE

Synonyms: hidden

= Diasperus pilosus (Lour.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Emblica pilosa (Lour.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 3: 20 (1826)

= Nymphanthus pilosus Lour.; Fl. Cochinch. 544 (1790)

= Phyllanthus pilosus (Lour.) Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 432 (1866)

species :

Glochidion pitcairnense (F.Br.) H.St.John; Trans. Roy. Soc. New Zealand, Bot. 1: 187 (1962)

Distribution: Gambier Isl.; Pitcairn Isl.; Henderson Isl.

61 GBI HEN PIT

Remarks: subgen. Glochidion

Synonyms: hidden

= Glochidion taitense var. pitcairnense F.Br.; Bull. Bernice P. Bishop Mus. 130: 142 (1935)

= Phyllanthus pitcairnensis (F.Br.) W.L.Wagner & Lorence; PhytoKeys 4: 83 (2011), as ´(H. St. John) W. L. Wagner & Lorence´

species :

Glochidion plagiophyllum Airy Shaw; Kew Bull. 37(1): 23 (1982)

Distribution: Lesser Sunda Isl. (Sumba)

42 LSI

Synonyms: hidden

= Phyllanthus plagiophyllus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion pleiosepalum Airy Shaw; Kew Bull. 27(1): 56 (1972)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus pleiosepalus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion podocarpum (Müll.Arg.) C.B.Rob.; Philipp. J. Sci., C 6: 330 (1911)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus podocarpus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus podocarpus Müll.Arg.; Flora 48: 388 (1865)

species :

Glochidion poilanei (Beille) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: S-Vietnam

41 VIE

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthodendron poilanei (Beille) Croizat; J. Arnold Arbor. 23: 34 (1942)

= Phyllanthus lignulatus var. tonkinensis Baill.; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 593 (1927)

= Phyllanthus poilanei Beille; Bull. Soc. Bot. France 72: 162 (1925), et Fl. Indo-Chine [P.H.Lecomte et al.] 5: 593 (1927)

species :

Glochidion pomiferum Airy Shaw; Kew Bull. 23: 13 (1969)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus maliformis Govaerts & Christenh.; Nelumbo 68(1): 142 (2026), nom. nov.

species :

Glochidion ponapense Hosok.; Trans. Nat. Hist. Soc. Taiwan 25: 24 (1935)

Distribution: Fed. States of Micronesia (Pohnpei)

62 MIC

Remarks: subgen. Glochidion

Synonyms: hidden

= Glochidion excorticans var. calvum Fosberg; Willdenowia 20: 261 (1991)

= Phyllanthus ponapensis (Hosok.) W.L.Wagner & Lorence; PhytoKeys 4: 84 (2011)

species :

Glochidion praeclarum Airy Shaw; Kew Bull. 33(1): 27 (1978)

Distribution: Borneo (Sarawak)

42 BOR

Synonyms: hidden

= Phyllanthus praeclarus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion prinoides S.Moore; J. Bot. 63(Suppl.): 95 (1925)

Distribution: Java

Error

Synonyms: hidden

= Phyllanthus prinoides (S.Moore) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion pruinosum Airy Shaw; Muelleria 4(3): 211 (1980)

Distribution: Australia (Queensland)

50 AUS

Synonyms: hidden

= Phyllanthus wundu Govaerts & Christenh.; Nelumbo 68(1): 148 (2026), nom. nov.

species :

Glochidion psidioides C.B.Rob.; Philipp. J. Sci., C 4: 92 (1909)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus psidioides (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion puberulum Hosok.; Trans. Nat. Hist. Soc. Taiwan 25: 23 (1935)

Distribution: Fed. States of Micronesia (Truk, Moen, Dublon, Udot, Uman, Pohnpei)

62 MIC

Synonyms: hidden

= Glochidion excorticans Fosberg; Willdenowia 20: 260 (1991)

= Glochidion senyavinianum Glassman; Bull. Bernice P. Bishop Mus. 209: 71 (1952)

= Phyllanthus senyavinianus (Glassman) W.L.Wagner & Lorence; PhytoKeys 4: 87 (2011)

species :

Glochidion puberum (L.) Hutch.; Pl. Wilson. [Sargent] 2(3): 518 (1916)

Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hongkong, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang); Tibet; Japan (Kyushu); Taiwan; USA [I] (Alabama [I])

36 CHI TIB 38 JAP TAI 78 usa

Synonyms: hidden

= Agyneia impubes L.; Mant. Pl. Altera [Linnaeus] 296 (1771)

= Agyneia pinnata Miq.; Fl. Ned. Ind. 1(2): 368 (1859)

= Agyneia pubera L.; Mant. Pl. Altera [Linnaeus] 161, 296, 576 (1771)

= Agyneia sinica Miq.; Fl. Ned. Ind. 1(2): 368 (1859)

= Bradleia pubera (L.) Roxb.; Hort. Bengal. 104 (1814), nomen, Fl. Ind. [Roxburgh] 3: 698 (1832)

= Bradleja sinica Gaertn.; Fruct. Sem. Pl. 2: 127, t. 109, f. 1 (1791)

= Diasperus impubes (L.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus villosus (Poir.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion distichum Hance; Ann. Sci. Nat., Bot., Sér. 4, 18: 228 (1862)

= Glochidion hayatae var. tsushimense Hurus.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 6: 332 (1954)

= Glochidion pseudoobscurum Pamp.; Nuovo Giorn. Bot. Ital., n. s. 17(3): 411-413 (1910)

= Glochidion sinicum (Gaertn.) Hook. & Arn.; Bot. Beechey Voy. 210 (1837)

= Glochidion sinicum (Gaertn.) S.M.Almeida; Fl. Savantwadi 1: 387 (1990), isonym

= Nymphanthus chinensis Lour.; Fl. Cochinch. 2: 544 (1790)

= Phyllanthus puber (L.) Müll.Arg.; Flora 48: 387 (1865)

= Phyllanthus puberus var. sinicus (Hook. & Arn.) Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 307 (1866)

= Phyllanthus villosus Poir.; Encycl. [J.Lamarck et al.] 5: 297 (1804), nom. nov.

species :

Glochidion pubicapsa Airy Shaw; Kew Bull., Addit. Ser. 4: 130 (1975)

Distribution: Sumatra; Borneo; Sulawesi; Palawan; Java

42 BOR PLW SUL

Synonyms: hidden

= Glochidion pubicarpum Elmer; Leafl. Philipp. Bot. 10: 3733 (1939), anglice.

= Phyllanthus pubicapsa (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

variant :

var. brunneiforme Airy Shaw; Kew Bull., Addit. Ser. 4: 131 (1975)

Distribution: Borneo (Sarawak)

42 BOR

Synonyms: hidden

= Phyllanthus pubicapsa var. brunneiformis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion pulgarense Elmer; Leafl. Philipp. Bot. 4: 1291 (1911)

Distribution: Palawan

42 PLW

Synonyms: hidden

= Phyllanthus pulgarensis (Elmer) Govaerts & Christenh.; Nelumbo 68(1): 144 (2026)

species :

Glochidion punctatum Pax & K.Hoffm.; Mitt. Inst. Allg. Bot. Hamburg 7: 226 (1931)

Distribution: Borneo

42 BOR

Synonyms: hidden

= Phyllanthus punctatus (Pax & K.Hoffm.) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion pungens Airy Shaw; Kew Bull. 31(2): 349 (1976)

Distribution: Australia (Queensland)

50 AUS

Synonyms: hidden

= Phyllanthus pungens (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion pyriforme Airy Shaw; Kew Bull. 27(1): 33 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus pyriformis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion raivavense F.Br.; Bull. Bernice P. Bishop Mus. 130: 142 (1935)

Distribution: Austral Isl. (Raivavae, Rurutu, Tubuai)

61 AUI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus raivavensis (F.Br.) W.L.Wagner & Lorence; PhytoKeys 4: 85 (2011)

species :

Glochidion ramiflorum J.R.Forst. & G.Forst.; Char. Gen. Pl., ed. 1: 57 (1775)

Distribution: China [I] (W-Guangdong [I]); Vanuatu; Rotuma Isl.; Fiji; Wallis & Futuna (Alofi Isl., Futuna Isl., ´Uvea Isl. and islets [Wallis]); Palau (Babeldaob); Fed. States of Micronesia (Yap, Truk, Pohnpei, ?Namonuito, ?Nomwin)

36 chi 60 FIJ ROT VAN WAL 62 MIC PLU

Remarks: Plants east of Vanuatu should not be called G. ramiflorum fide Smith 1981 but probably belong to G. concolor.

Synonyms: hidden

= Bradleia glochidion Gaertn.; Fruct. Sem. Pl. 2: 128 (1791), nom. superfl.

= Diasperus ramiflorus (J.R.Forst. & G.Forst.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Glochidion tannaense Guillaumin; J. Arnold Arbor. 13: 90 (1932)

= Phyllanthus ramiflorus (J.R.Forst. & G.Forst.) Müll.Arg.; Flora 48: 374 (1865), nom. illeg.

= Phyllanthus tannaensis (Guillaumin) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion rapaense J.Florence; Bull. Mus. Natl. Hist. Nat., B, Adansonia 18: 258, fig. 6 (1996)

Distribution: Austral Isl. (Rapa Iti)

61 AUI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus rapaensis (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 85 (2011)

species :

Glochidion reinwardtii (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

Distribution: Java

Error

Synonyms: hidden

= Diasperus reinwardtii (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus reinwardtii Müll.Arg.; Flora 48: 379 (1865)

species :

Glochidion reticulatum Elmer; Leafl. Philipp. Bot. 1: 302 (1908)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus cancellatus Govaerts & Christenh.; Nelumbo 68(1): 137 (2026), nom. nov.

species :

Glochidion retinerve Airy Shaw; Kew Bull. 27(1): 39 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus reticulatovenosus Govaerts & Christenh.; Nelumbo 68(1): 145 (2026), nom. nov.

species :

Glochidion robinsonii Elmer; Leafl. Philipp. Bot. 7: 2643 (1915)

Distribution: Philippines (Luzon, Panay, Leyte, Mindanao)

42 PHI

Synonyms: hidden

= Glochidion sorsogonense Elmer ex Merr.; Enum. Philipp. Fl. Pl. 2: 402 (1923), in obs., pro syn., Elmer, Leafl. Philipp. Bot. 10: 3734 (1939), descr. angl.

= Phyllanthus sorsogonensis (Elmer ex Merr.) Govaerts & Christenh.; Nelumbo 68(1): 146 (2026)

species :

Glochidion roseum (Craib & Hutch.) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: China (Yunnan); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (Langkawi Isl., Kedah, Penang, Terengganu, Perak)

36 CHI 41 CBD LAO THA VIE 42 MLY

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Cleistanthus dubius Ridl.; J. Straits Branch Roy. Asiat. Soc. 59: 168 (1911)

= Glochidion flavum Ridl.; J. Straits Branch Roy. Asiat. Soc. 59: 173 (1911)

= Phyllanthodendron album Craib & Hutch.; Bull. Misc. Inform. Kew 1910(8): 279-280 (1910)

= Phyllanthodendron dubium (Ridl.) Gage; Rec. Bot. Surv. India 9: 219 (1922)

= Phyllanthodendron roseum Craib & Hutch.; Bull. Misc. Inform. Kew 1910(1): 23 (1910)

= Phyllanthodendron roseum var. glabrum Craib ex Hosseus; Beih. Bot. Centralbl. 28(2): 406-407 (1911)

= Phyllanthodendron roseum var. siamense (Pax & K.Hoffm.) Craib; Bull. Misc. Inform. Kew 1911: 460 (1911)

= Phyllanthodendron siamense (Pax & K.Hoffm.) Hosseus; Repert. Spec. Nov. Regni Veg. 10: 116 (1911)

= Phyllanthus albus (Craib ex Hosseus) Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 590 (1927), nom. illeg.

= Phyllanthus roseus (Craib & Hutch.) Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 590 (1927)

= Phyllanthus roseus var. glaber (Craib ex Hosseus) Craib ex Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 590 (1927)

= Uranthera siamensis Pax & K.Hoffm.; Pflanzenr. [Engler] IV.147.III, Heft 47 (Euph.-Cluytieae): 95 (1911)

species :

Glochidion rubicundum (Beille) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: Vietnam

41 VIE

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus rubicundus Beille; Bull. Soc. Bot. France 72: 162 (1925), et Fl. Indo-Chine [P.H.Lecomte et al.] 5: 592 (1927)

species :

Glochidion rubrum Blume; Bijdr. Fl. Ned. Ind. 586 (1825)

Distribution: China (Fujian); Ryukyu Isl.; Taiwan; Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Singapore; Sumatra; Borneo; Brunei; Sulawesi; Palawan; Philippines (Luzon, Mindanao); Java; Lesser Sunda Isl. (Bali, etc.); Moluccas

36 CHI 38 NNS TAI 41 CBD LAO MYA THA VIE 42 BOR BRN LSI MLY MOL PHI PLW SNG SUL

Synonyms: hidden

= Bradleia coronata Benth.; Numer. List [Wallich] no. 7857 (1847), nomen

= Bradleia rubra (Blume) Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Bridelia glauca Benth.; Numer. List [Wallich] no. 7875 (1847), nomen

= Diasperus coronatus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 597 (1891)

= Diasperus leiostylus (Kurz) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus subscandens (Zoll. & Moritzi) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus wightianus Kuntze; Revis. Gen. Pl. 2: 603 (1891), nom. illeg.

= Glochidion chademenosocarpum Hayata; Icon. Pl. Formosan. 9: 94-95 (1920)

= Glochidion coronatum Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 326 (1887)

= Glochidion diversifolium (Miq.) Merr.; Bull. Bur. Forest. Philipp. Islands 1: 29 (1903)

= Glochidion foliosum S.Moore; J. Bot. 63(Suppl.): 96 (1925)

= Glochidion fortunei Hance; Ann. Sci. Nat., Bot., Sér. 4, 18: 228 (1862)

= Glochidion fortunei var. longistylum H.Keng; J. Wash. Acad. Sci. 41(6): 200 (1951)

= Glochidion fortunei var. megacarpum H.Keng; J. Wash. Acad. Sci. 41(6): 200 (1951)

= Glochidion grave S.Moore; J. Bot. 63(Suppl.): 96 (1925)

= Glochidion insulare Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 310 (1887)

= Glochidion insulare var. brevistylum Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 13(3): 715 (1989)

= Glochidion leiostylum Kurz; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 42(2): 237 (1873)

= Glochidion penangense (Müll.Arg.) Airy Shaw; Kew Bull. 23: 6 (1969)

= Glochidion rubrum f. longistylis J.J.Sm.; Bijdr. Boomsoort. Java 12: 152 (1910)

= Glochidion rubrum var. brevistylum (Chakr. & M.Gangop.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 223 (1995)

= Glochidion subscandens Zoll. & Moritzi; Natuur- Geneesk. Arch. Ned.-Indië 2: 584 (1845)

= Glochidion suishaense Hayata; Icon. Pl. Formosan. 9: 97 (1920)

= Glochidion thorelii Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 622 (1927)

= Glochidion versicolor S.Moore; J. Bot. 63(Suppl.): 94 (1925)

= Phyllanthus diversifolius Miq.; Fl. Ned. Ind., Eerste Bijv. 448 (1861)

= Phyllanthus diversifolius var. brevistylus (Chakr. & M.Gangop.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 713 (2009)

= Phyllanthus nitidulus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1272 (1866)

= Phyllanthus penangensis Müll.Arg.; Flora 48: 388 (1865)

= Phyllanthus puberus var. fortunei (Hance) Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 307 (1866)

= Phyllanthus ruber (Blume) T.Kuros.; J. Jap. Bot. 91: 119 (2016), nom. illeg.

= Phyllanthus subscandens (Zoll. & Moritzi) Müll.Arg.; Flora 48: (1865) 372. 48: 372 (1865)

= Phyllanthus subscandens var. brevistylus (Chakrab. & M.Gangop.) Govaerts & Christenh.; Nelumbo 68(1): 146 (2026)

species :

Glochidion rufoglaucum (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 275 (1900)

Distribution: Java

Error

Synonyms: hidden

= Diasperus rufoglaucus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus rufoglaucus Müll.Arg.; Flora 48: 372 (1865)

species :

Glochidion rugulosum Airy Shaw; Kew Bull. 23: 17 (1969)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus rugulosum (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion runikerae Airy Shaw; Kew Bull. 33(1): 33 (1978)

Distribution: Solomon Isl.

43 SOL

Synonyms: hidden

= Phyllanthus runikerae (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion saccocarpum Airy Shaw; Kew Bull. 25(3): 487 (1971)

Distribution: Solomon Isl.

43 SOL

Synonyms: hidden

= Phyllanthus saccocarpus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion samoanus (Müll.Arg.) R.W.Bouman; Phytotaxa 540(1): 63 (2022)

Distribution: American Samoa (Olosega, Tutuila, Ta´u); [Western] Samoa (´Upolu, Savai´i)

60 ASA SAM

Remarks: subgen. Glochidion

Synonyms: hidden

= Glochidion cuspidatum var. samoanum (Müll.Arg.) Pax; Bot. Jahrb. Syst. 25: 645 (1898)

= Glochidion ramiflorum var. samoanum (Müll.Arg.) Pax; Bot. Jahrb. Syst. 25: 645 (1898)

= Phyllanthus gaudichaudii var. samoanus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 300 (1866)

= Phyllanthus ramiflorus var. samoanus Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 289 (1866)

= Phyllanthus samoanus (Müll.Arg.) W.L.Wagner & Lorence; PhytoKeys 4: 86 (2011)

species :

Glochidion santisukii Airy Shaw; Kew Bull. 35(2): 385 (1980), nom. nov.

Distribution: peninsular Thailand

41 THA

Synonyms: hidden

= Glochidion cataractarum Airy Shaw; Kew Bull. 32: 76 (1977), nom. illeg.

= Phyllanthus santisukii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion seemannii Müll.Arg.; Linnaea 32: 63 (1863)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus seemannii (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus venulosus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion venulosum (Müll.Arg.) P.T.Li; Guihaia 14(2): 131 (1994)

= Phyllanthus seemannianus (Müll.Arg.) Müll.Arg.; Flora 48: 374 (1865)

= Phyllanthus seemannii (Müll.Arg.) Müll.Arg.; Flora 48: 374 (1865)

= Phyllanthus venulosus Müll.Arg.; Flora 48: 374 (1865)

species :

Glochidion semicordatum (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

Distribution: Java

Error

Synonyms: hidden

= Diasperus semicordatus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Phyllanthus semicordatus Müll.Arg.; Flora 48: 376 (1865)

species :

Glochidion sericeum (Blume) Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 326 (1887)

Distribution: Cambodia; Vietnam; peninsular Thailand; peninsular Malaysia (Terengganu and Perak southward); Singapore; Sumatra; Borneo; Brunei; Java

41 CBD THA VIE 42 BOR BRN MLY SNG

Synonyms: hidden

= Diasperus sericeus (Blume) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion sericeum (Blume) Pax; Nat. Pflanzenfam. [Engler & Prantl] III, 5: 24 (1890)

= Glochidionopsis sericea Blume; Bijdr. Fl. Ned. Ind. 588 (1825)

= Phyllanthus sericeus (Blume) Müll.Arg.; Flora 48: 390 (1865)

species :

Glochidion sessiliflorum Airy Shaw; Kew Bull. 31(2): 350 (1976)

Distribution: Australia (Queensland)

50 AUS

Synonyms: hidden

= Glochidion sessiliflorum var. pedicellatum Airy Shaw; Kew Bull. 35: 640 (1980)

= Glochidion sessiliflorum var. stylosum Airy Shaw; Kew Bull. 35: 641 (1980)

= Phyllanthus sessiliflorus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 145 (2026)

species :

Glochidion sessilis (Warb.) R.W.Bouman; Phytotaxa 540(1): 63 (2022)

Distribution: New Guinea (Irian Jaya)

42 NWG

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus sessilis Warb.; Bot. Jahrb. Syst. 13: 357 (1891)

species :

Glochidion singaporense Gage; Rec. Bot. Surv. India 9: 221 (1922)

Distribution: peninsular Malaysia (Selangor, Negeri Sembilan, Melaka, Johor); Singapore; Sumatra; Borneo

42 BOR MLY SNG

Synonyms: hidden

= Phyllanthus singaporensis (Gage) Govaerts & Christenh.; Nelumbo 68(1): 146 (2026)

species :

Glochidion societatis J.Florence; Fl. Polynésie Franç. [Florence] 1: 68 (1997)

Distribution: Society Isl. (Huahine, Maupiti, Raiatea, Tahaa); Austral Isl. (Rimatara)

61 AUI SCI

Synonyms: hidden

= Phyllanthus florencei W.L.Wagner & Lorence; PhytoKeys 4: 74 (2011), nom. nov.

species :

Glochidion sphaerogynum (Müll.Arg.) Kurz; Forest Fl. Burma 2: 346 (1877)

Distribution: China (Guangdong, Guangxi, Hainan, Yunnan); India (Arunachal Pradesh, Assam, Meghalaya, Mizoram, Sikkim, West Bengal); Bangladesh; Myanmar; Thailand; Laos; Vietnam

36 CHI 40 BAN IND 41 LAO MYA THA VIE

Synonyms: hidden

= Diasperus sphaerogynus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion fagifolium Miq. ex Bedd.; Forester´s Man. Bot. 193 (1872), nom. illeg.

= Glochidion mandakatense Kalita & Borthakur; J. Econ. Taxon. Bot. 30(3): 488 (2006)

= Phyllanthus sphaerogynus Müll.Arg.; Flora 48: 375 (1865)

species :

Glochidion stellatum (Retz.) Bedd.; Forester´s Man. Bot. 194 (1872)

Distribution: Sri Lanka; Vietnam

40 SRL 41 VIE

Synonyms: hidden

= Diasperus jussieuianus (Wight) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus stellatus (Retz.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion jussieuianum (Wight) Thwaites; Enum. Pl. Zeyl. [Thwaites] 285 (1861)

= Glochidion rigidum (A.Juss.) Müll.Arg.; Linnaea 32: 67 (1863)

= Glochidion thwaitesii Müll.Arg.; Linnaea 32: 66 (1863)

= Gynoon jussieuianum Wight; Icon. Pl. Ind. Orient. [Wight] 5(2): 29 (1852)

= Gynoon rigidum A.Juss.; Euphorb. Gen. 107, t. 3 (1824)

= Gynoon tetrandrum D.Dietr.; Syn. Pl. [Dietrich] 5: 388 (1852)

= Gynoon triandrum Wight & Arn.; Edinburgh New Philos. J. 14: 300 (1833)

= Phyllanthus jussieuianus (Wight) Müll.Arg.; Flora 48: 386 (1865)

= Phyllanthus stellatus Retz.; Observ. Bot. [Retzius] 5: 29 (1788)

species :

Glochidion stenophyllum Airy Shaw; Kew Bull. 27(1): 31 (1972)

Distribution: Sulawesi; Moluccas; New Guinea (Irian Jaya)

42 MOL NWG SUL

Synonyms: hidden

= Phyllanthus bambusifolius Govaerts & Christenh.; Nelumbo 68(1): 136 (2026), nom. nov.

species :

Glochidion stilpnophyllum Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 175 (1996), nom. nov.

Distribution: Borneo; Philippines (Luzon, Polillo, Samar)

42 BOR PHI

Synonyms: hidden

= Glochidion nitidum Merr.; Philipp. J. Sci., C 9: 483 (1914 publ. 1915)

= Glochidion subfalcatum var. nitidum (Merr.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 232 (1995)

= Phyllanthus stilpnophyllus (Govaerts & Radcl.-Sm.) Govaerts & Christenh.; Nelumbo 68(1): 146 (2026)

species :

Glochidion stipulare Airy Shaw; Kew Bull. 32(2): 377 (1978)

Distribution: Vanuatu

60 VAN

Synonyms: hidden

= Phyllanthus vanuatuanthus Govaerts & Christenh.; Nelumbo 68(1): 148 (2026), nom. nov.

species :

Glochidion striatum J.J.Sm.; Nova Guinea 8: 784 (1912)

Distribution: New Guinea (Irian Jaya)

42 NWG

Synonyms: hidden

= Phyllanthus striatus (J.J.Sm.) Govaerts & Christenh.; Nelumbo 68(1): 146 (2026)

species :

Glochidion styliferum J.J.Sm.; Bull. Jard. Bot. Buitenzorg, Sér. 3, 1: 391 (1920)

Distribution: Borneo

42 BOR

Synonyms: hidden

= Phyllanthus styliferus (J.J.Sm.) Govaerts & Christenh.; Nelumbo 68(1): 146 (2026)

species :

Glochidion stylosum Ridl.; Bull. Misc. Inform. Kew 1923: 364 (1923)

Distribution: peninsular Malaysia (Semangkok Pass, Selangor); Borneo

42 BOR MLY

Synonyms: hidden

= Phyllanthus malayanthus Govaerts & Christenh.; Nelumbo 68(1): 142 (2026), nom. nov.

species :

Glochidion subfalcatum Elmer; Leafl. Philipp. Bot. 1: 305 (1908)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus subfalcatus (Elmer) Govaerts & Christenh.; Nelumbo 68(1): 146 (2026)

species :

Glochidion submolle (Lauterb. & Schum.) Airy Shaw; Kew Bull. 32(2): 377 (1978)

Distribution: New Guinea (Irian Jaya, Papua New Guinea)

42 NWG

Synonyms: hidden

= Glochidion magnificum K.Schum.; Nachtr. Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 288 (1905)

= Phyllanthus submollis K.Schum. & Lauterb.; Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 390 (1900)

species :

Glochidion subsessile N.P.Balakr. & Chakr.; Proc. Indian Acad. Sci., Pl. Sci. 92(4): 359 (1983)

subspecies :

subsp. subsessile

Distribution: Andaman Isl. (South Andaman Isl.)

41 AND

Synonyms: hidden

= Phyllanthus subsessilis (N.P.Balakr. & Chakr.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

subspecies :

subsp. birmanicum Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 13(3): 716 (1989)

Distribution: Myanmar

41 MYA

species :

Glochidion superbum Baill. ex Müll.Arg.; Linnaea 32: 64 (1863)

Distribution: Myanmar; S-Thailand; peninsular Malaysia (common throughout); Singapore; Sumatra (incl. Bangka Isl.); Borneo; Java

41 MYA THA 42 BOR MLY SNG

Synonyms: hidden

= Bradleia finlaysoniana Benth.; Numer. List [Wallich] no. 7860 (1847), nomen

= Diasperus finlaysonianus (Wall.) Kuntze; Revis. Gen. Pl. 2: 602 (1891)

= Glochidion dasyphyllum Miq.; Fl. Ned. Ind., Eerste Bijv. 451 (1861), nom. illeg.

= Phyllanthus finlaysonianus Benth.; Numer. List [Wallich] no. 7907 (1847), nomen

= Phyllanthus superbus (Baill. ex Müll.Arg.) Müll.Arg.; Flora 48: 375 (1865)

species :

Glochidion symingtonii Airy Shaw; Kew Bull. 27(1): 6 (1972)

Distribution: peninsular Malaysia (Keledang Saiong and Gunung Peninjau, Perak)

42 MLY

Synonyms: hidden

= Phyllanthus symingtonii (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion taitense Baill. ex Müll.Arg.; Linnaea 32: 66 (1863) (tahitense auct.)

Distribution: Cook Isl. (Mangaia, Ngaputoru Group); Society Isl. (Moorea, Tahiti)

61 COO SCI

Remarks: see also G. concolor and G. ramiflorum

Synonyms: hidden

= Diasperus taitensis (Baill. ex Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion majus Baill.; Étude Euphorb. 637 (1858), nomen

= Glochidion ramiflorum var. macrophyllum Müll.Arg.; Linnaea 32: 63 (1863)

= Glochidion taitense Baill.; Étude Euphorb. 637 (1858), nomen

= Phyllanthus ramiflorus var. macrophyllus (Müll.Arg.) Müll.Arg.; Flora 48: 374 (1865)

= Phyllanthus taitensis (Baill. ex Müll.Arg.) Müll.Arg.; Flora 48: 380 (1865)

= Phyllanthus taitensis var. glabrescens Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 301 (1866)

species :

Glochidion talakonense Sankara Rao, J.Swamy, Nagaraju, S.B.Padal, M.Tarak.Naidu, K.C.Mohan & Thulasiah; Bangladesh J. Pl. Taxon. 23(1): 59 (2016)

Distribution: India (Andhra Pradesh)

40 IND

Synonyms: hidden

= Phyllanthus talakonensis (Sankara Rao, J.Swamy, Nagaraju, S.B.Padal, M.Tarak.Naidu, K.C.Mohan & Thulasiah) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion talmyanum Beille; Fl. Indo-Chine [P.H.Lecomte et al.] 5: 614 (1927)

Distribution: S-Vietnam

41 VIE

Synonyms: hidden

= Phyllanthus talmyanus (Beille) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion temehaniense J.W.Moore; Bull. Bernice P. Bishop Mus. 226: 15 (1963)

Distribution: Society Isl. (Raiatea, Huahine, Tahaa)

61 SCI

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus temehaniensis (J.W.Moore) W.L.Wagner & Lorence; PhytoKeys 4: 88 (2011)

species :

Glochidion tenuistylum Stapf; Trans. Linn. Soc. London, Bot., Ser. 2, 4: 223 (1894)

Distribution: Borneo

42 BOR

Synonyms: hidden

= Glochidion penangense var. tenuistylum (Stapf) Airy Shaw; Kew Bull. 23: 7 (1969)

= Phyllanthus tenuistylus (Stapf) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion ternateum Airy Shaw; Kew Bull. 27(1): 28 (1972)

Distribution: Moluccas (Ternate Isl.)

42 MOL

Synonyms: hidden

= Phyllanthus ternateus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion tetrapteron Gage; Rec. Bot. Surv. India 9: 223 (1922)

Distribution: peninsular Malaysia (Pahang); Sumatra

42 MLY

Synonyms: hidden

= Phyllanthus tetrapteros (Gage) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion thesioides H.Eichl.; Fl. S. Austral. [J.M.Black], ed. 2, Suppl.: 210 (1965)

Distribution: Australia (Northern Territory, New South Wales)

50 AUS

Synonyms: hidden

= Phyllanthus thesioides Benth.; Fl. Austral. [Bentham] 6: 98 (1873), non Müll. Arg. 1863

= Sauropus thesioides (H.Eichl.) Airy Shaw; Kew Bull. 35(3): 684 (1980)

species :

Glochidion thomsonii (Müll.Arg.) Hook.f.; Fl. Brit. India [J.D.Hooker] 5(14): 318 (1887)

Distribution: SE-Tibet; India (Assam, Meghalaya); Bangladesh

36 TIB 40 BAN IND

Synonyms: hidden

= Diasperus thomsonii (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion rubidulum T.L.Chin; Acta Phytotax. Sin. 18(2): 251-252, f. 1, 4-6 (1980)

= Phyllanthus thomsonii Müll.Arg.; Flora 48: 375 (1865)

species :

Glochidion timorense Airy Shaw; Kew Bull. 27(1): 31 (1972)

Distribution: Lesser Sunda Isl. (Timor)

42 LSI

Synonyms: hidden

= Phyllanthus timorensis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion tomentosum Dalzell; Hooker´s J. Bot. Kew Gard. Misc. 2: 38 (1851)

Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Yunnan); Tibet; Ryukyu Isl.; Taiwan; India (throughout); Sri Lanka; Bangladesh; Andaman Isl. (common); Nicobar Isl. (common); Thailand

36 CHI TIB 38 NNS TAI 40 BAN IND SRL 41 AND NCB THA

Synonyms: hidden

= Agyneia hirsuta Miq.; Fl. Ned. Ind. 1(2): 368 (1859)

= Bradleia hirsuta Roxb.; Fl. Ind. [Roxburgh] 3: 699 (1832)

= Bradleia mollis Benth.; Numer. List [Wallich] no. 7858 (1847), nomen

= Bradleia mollis Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Diasperus arnottianus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 598 (1891)

= Diasperus hirsutus (Roxb.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus tomentosus (Dalzell) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion arnottianum Müll.Arg.; Linnaea 32: 60 (1863)

= Glochidion dasyphyllum K.Koch; Hort. Dendrol. 85 (1853)

= Glochidion dasyphyllum var. iriomatense Hurus.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 6: 334 (1954)

= Glochidion hirsutum (Roxb.) Voigt; Hort. Suburb. Calcutt. 153 (1845)

= Glochidion hongkongense var. puberulum Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 13(3): 712 (1989)

= Glochidion molle Hook. & Arn.; Bot. Beechey Voy. 210 (1837), nom. illeg.

= Glochidion tomentosum var. talbotii Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 371 (1887)

= Glochidion zeylanicum var. talbotii (Hook.f.) Haines; Bot. Bihar Orissa 2: 132 (1921)

= Glochidion zeylanicum var. tomentosum (Dalzell) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 229 (1995)

= Glochidion zeylanicum var. tomentosum (Dalzell) Trimen; Cat. Pl. Ceylon. 79 (1824)

= Phyllanthus arnottianus (Müll.Arg.) Müll.Arg.; Flora 48: 370 (1865)

= Phyllanthus hirsutus (Roxb.) Müll.Arg.; Flora 48: 371 (1865)

= Phyllanthus obliquus var. tomentosus (Dalzell) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

= Phyllanthus tomentosus (Dalzell) Müll.Arg.; Flora 48: 371 (1865)

= Phyllanthus zeylanicus var. tomentosus (Dalzell) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

species :

Glochidion triandrum (Blanco) C.B.Rob.; Philipp. J. Sci., C 4(1): 92 (1909)

Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hunan, Sichuan, Yunnan); Ryukyu Isl.; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam, Meghalaya, Mizoram. Nagaland, Sikkim, West Bengal); Bangladesh; Cambodia; Philippines (Luzon, Polillo, Mindoro, Sibuyan, Negros, Guimaras, Biliran)

36 CHI 38 NNS TAI 40 BAN BHU IND NEP 41 CBD 42 PHI

Synonyms: hidden

= Bradleia acuminata Benth.; Numer. List [Wallich] no. 7855 (1847), nomen

= Bridelia acuminata Benth.; Numer. List [Wallich] no. 7885 (1847)

= Diasperus triandrus (Blanco) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion acuminatum Müll.Arg.; Linnaea 32: 68 (1863)

= Glochidion bicolor (Müll.Arg.) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 20: 18, t. 2E (1904)

= Glochidion eleutherostylum Müll.Arg.; Linnaea 32: 69 (1863)

= Glochidion hayatae Croizat & H.Hara; J. Jap. Bot. 16: 316 (1940)

= Glochidion hypoleucum Hayata; Icon. Pl. Formosan. 9: 95-96 (1920)

= Glochidion quinquestylum Elmer; Leafl. Philipp. Bot. 1: 303 (1908)

= Kirganelia triandra Blanco; Fl. Filip. [Blanco] 711 (1837)

= Phyllanthus bicolor Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 311 (1866)

= Phyllanthus meghalayensis Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009), nom. nov.

= Phyllanthus triandrus (Blanco) Müll.Arg.; Flora 48: 379 (1865)

variant :

var. siamense (Airy Shaw) P.T.Li; Acta Phytotax. Sin. 26(1): 62 (1988)

Distribution: China (Yunnan); W-Thailand; Laos

36 CHI 41 LAO THA

Synonyms: hidden

= Glochidion acuminatum var. siamense Airy Shaw; Kew Bull. 26(2): 273 (1972)

= Phyllanthus meghalayensis var. siamensis (Airy Shaw) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33: 714 (2009)

species :

Glochidion trichogynum Müll.Arg.; Linnaea 32: 66 (1863)

Distribution: Palawan; Philippines (Luzon, Mindoro, Leyte, Mindanao)

42 PHI PLW

Synonyms: hidden

= Diasperus trichogynus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Phyllanthus trichogynus (Müll.Arg.) Müll.Arg.; Flora 48: 380 (1865)

species :

Glochidion trichophorum Merr.; Philipp. J. Sci., C 9: 484 (1914 publ. 1915)

Distribution: Philippines (Luzon)

42 PHI

Synonyms: hidden

= Phyllanthus trichophorus (Merr.) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion trusanicum Airy Shaw; Kew Bull. 27(1): 20 (1972)

Distribution: Borneo (Sarawak)

42 BOR

Synonyms: hidden

= Phyllanthus trusanicus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 147 (2026)

species :

Glochidion tuamotuense Florence; Fl. Polynésie Franç. [Florence] 1: 98 (1997)

Distribution: Gambier Isl. (Taravai); Tuamotu Arch. (Niau)

61 GBI TUA

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus tuamotuensis (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 89 (2011)

species :

Glochidion ultrabasicola Airy Shaw; Kew Bull. 36(3): 601 (1981)

Distribution: Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus ultrabasicola (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion urceolare Airy Shaw; Kew Bull. 27(1): 40 (1972)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus urceolaris (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion urophylloides Elmer; Leafl. Philipp. Bot. 1: 300 (1908)

Distribution: Cambodia; Vietnam; Borneo; Philippines (Batan, Luzon, Polillo, Mindoro, Panay, Leyte, Mindanao, Basilan)

41 CBD VIE 42 BOR PHI

Synonyms: hidden

= Glochidion fenicis Merr.; Philipp. J. Sci., C 3: 414 (1909)

= Glochidion lanceifolium C.B.Rob.; Philipp. J. Sci., C 4: 90 (1909)

= Glochidion subangulatum Elmer; Leafl. Philipp. Bot. 7: 2644 (1915)

= Phyllanthus urophylloides (Elmer) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion varians Miq.; Fl. Ned. Ind., Eerste Bijv. 450 (1861)

Distribution: peninsular Malaysia (Perak, Johor); +Singapore; Sumatra

42 MLY SNG

Synonyms: hidden

= Diasperus fuscus (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus varians (Miq.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion fuscum (Müll.Arg.) Boerl.; Handl. Fl. Ned. Ind. [Boerlage] 3: 276 (1900)

= Phyllanthus fuscus Müll.Arg.; Flora 48: 380 (1865)

= Phyllanthus varians (Miq.) Müll.Arg.; Flora 48: 380 (1865)

species :

Glochidion vitiense (Müll.Arg.) Gillespie; Bull. Bernice P. Bishop Mus. 91: 17 (1932)

Distribution: Fiji

60 FIJ

Remarks: subgen. Glochidion

Synonyms: hidden

= Diasperus vitiensis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion concolor var. obovatum Müll.Arg.; Linnaea 32: 62 (1863)

= Phyllanthus concolor var. obovatus (Müll.Arg.) Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 290 (1866)

= Phyllanthus vitiensis Müll.Arg.; Flora 48: 374 (1865)

species :

Glochidion wallichianum Müll.Arg.; Linnaea 32: 67 (1863)

Distribution: Myanmar; Thailand; peninsular Malaysia; Sumatra; Borneo

41 MYA THA 42 BOR MLY

Synonyms: hidden

= Diasperus curtisii (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus desmocarpus (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Glochidion curtisii Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 327 (1887)

= Glochidion desmocarpum Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 318 (1887)

= Glochidion glomerulatum var. wallichianum (Müll.Arg.) Chakr. & M.Gangop.; J. Econ. Taxon. Bot. 19(1): 204 (1995)

= Phyllanthus wallichianus (Müll.Arg.) Müll.Arg.; Flora 48: 387 (1865)

species :

Glochidion weberi C.B.Rob.; Philipp. J. Sci., C 6: 330 (1911)

Distribution: Philippines (Mindanao)

42 PHI

Synonyms: hidden

= Phyllanthus weberi (C.B.Rob.) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion websteri Fosberg; Willdenowia 20(1-2): 262 (1991)

Distribution: Fed. States of Micronesia

62 MIC

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus websteri (Fosberg) W.L.Wagner & Lorence; PhytoKeys 4: 90 (2011)

species :

Glochidion welzenii W.N.Takeuchi; Blumea 53(2): 399 (2008)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Synonyms: hidden

= Phyllanthus welzenii (W.N.Takeuchi) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion wilderi Florence; Fl. Polynésie Franç. [Florence] 1: 99 (1997)

Distribution: Gambier Isl. (Mangareva); Tuamotu Arch. (Makatea)

61 GBI TUA

Remarks: subgen. Glochidion

Synonyms: hidden

= Phyllanthus wilderi (J.Florence) W.L.Wagner & Lorence; PhytoKeys 4: 90 (2011)

species :

Glochidion williamsii C.B.Rob.; Philipp. J. Sci., C 3: 199 (1908)

Distribution: Borneo; Brunei; Sulawesi; Philippines (Luzon)

42 BOR BRN PHI SUL

Synonyms: hidden

= Phyllanthus coccinicoccus Govaerts & Christenh.; Nelumbo 68(1): 138 (2026), nom. nov.

species :

Glochidion wilsonii Hutch.; Pl. Wilson. [Sargent] 2(3): 518-519 (1916)

Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hubei, Jiangxi, Sichuan, Zhejiang)

36 CHI

Synonyms: hidden

= Glochidion bodinieri H.Lév.; Repert. Spec. Nov. Regni Veg. 12(317-321): 183 (1913)

= Glochidion pseudoobscurum var. glabrum Pamp.; Nuovo Giorn. Bot. Ital., n. s. 17(3): 412 (1910)

= Glochidion pseudoobscurum var. lanceolatum Pamp.; Nuovo Giorn. Bot. Ital., n. s. 17(3): 412-413 (1910)

= Phyllanthus wilsonii (Hutch.) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion wisselense Airy Shaw; Kew Bull. 27(1): 38 (1972)

Distribution: Philippines; New Guinea (Irian Jaya)

42 NWG PHI

Synonyms: hidden

= Glochidion soegengii Airy Shaw; Kew Bull. 29(2): 293 (1974)

= Phyllanthus wisselensis (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion wonenggau Airy Shaw; Kew Bull. 27(1): 26 (1972)

Distribution: NE-Sulawesi

42 SUL

Synonyms: hidden

= Phyllanthus wonenggau (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion woodii Merr.; Philipp. J. Sci. 26: 464 (1925)

Distribution: Philippines (Mindanao)

42 PHI

Synonyms: hidden

= Phyllanthus mindanawensis Govaerts & Christenh.; Nelumbo 68(1): 142 (2026), nom. nov.

species :

Glochidion wrightii Benth.; Fl. Hongk. 313-314 (1861)

Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Yunnan)

36 CHI

Synonyms: hidden

= Diasperus wrightii (Benth.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Phyllanthus wrightii (Benth.) Müll.Arg.; Flora 48: 378 (1865)

species :

Glochidion xerocarpum (O.Schwarz) Airy Shaw; Kew Bull., Addit. Ser. 8: 115 (1980), et in Muelleria 4(3): 212 (1980)

Distribution: Borneo; Sulawesi; Philippines; Java; Lesser Sunda Isl. (Timor, etc.); East Timor; Moluccas; New Guinea (Papua New Guinea); Australia (Northern Territory)

42 BOR LSI MOL NWG PHI SUL 50 AUS

Synonyms: hidden

= Phyllanthus xerocarpus Schwarz; Repert. Spec. Nov. Regni Veg. 24: 87 (1927)

species :

Glochidion xestophyllum Airy Shaw; Kew Bull. 23: 11 (1969)

Distribution: peninsular Malaysia (once from Banang F. R., Johor); Borneo

42 BOR MLY

Synonyms: hidden

= Phyllanthus xestophyllus (Airy Shaw) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion yangchunense Z.Q.Song & Gang Yao; PhytoKeys 239: 230 (2024)

Distribution: China (Guangdong)

36 CHI

Synonyms: hidden

= Phyllanthus yangchunensis (Z.Q.Song & Gang Yao) Govaerts & Christenh.; Nelumbo 68(1): 148 (2026)

species :

Glochidion yunnanense (Croizat) R.W.Bouman; Phytotaxa 540(1): 65 (2022)

Distribution: China (Guizhou, Yunnan)

36 CHI

Remarks: subgen. Pseudoactephila

Synonyms: hidden

= Phyllanthus yunnanensis (Croizat) Govaerts & Radcl.-Sm.; Kew Bull. 51(1): 178 (1996)

= Phyllanthodendron yunnanense Croizat; J. Arnold Arbor. 23: 36 (1942)

species :

Glochidion zeylanicum (Gaertn.) A.Juss.; Euphorb. Gen. 107 (1824)

Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hongkong, Yunnan); Japan (Yakushima Isl., Tanegashima Isl.); Ryukyu Isl.; Taiwan; India (Andhra Pradesh,Assam, Chhattisgarh, Goa, Karnataka, Kerala, Maharashtra, Odisha, Tamil Nadu, Tripura); Sri Lanka; Bangladesh; Andaman Isl. (South Andaman Isl.); Nicobar Isl. (North Nicobar Isl., Great Nicobar Isl., Little Nicobar Isl.); Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (Perak, Pahang, Melaka, Johor); Singapore; Sumatra; Borneo; Brunei; Sulawesi; Java; Lesser Sunda Isl. (Bali, etc.); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Solomon Isl.; Australia (Northern Territory, Queensland, New South Wales); Vanuatu

36 CHI 38 JAP NNS TAI 40 BAN IND SRL 41 AND CBD LAO MYA NCB THA VIE 42 BOR BRN LSI MLY NWG SNG SUL 43 BIS SOL 50 AUS 60 VAN

Conservation Status: vulnerable (Singapore)

Synonyms: hidden

= Agyneia flexuosa B.Heyne ex Benth.; Numer. List [Wallich] no. 7863 (1847), nomen

= Agyneia obliqua Willd.; Sp. Pl., ed. 4 [Willdenow] 4: 568 (1805)

= Bradleia nitida Roxb.; Hort. Bengal. 69 (1814), nomen, Fl. Ind. [Roxburgh] 3: 699 (1832)

= Bradleia obliqua (Willd.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 3: 19 (1826)

= Bradleia timoriensis Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 222 (1841)

= Bradleja zeylanica Gaertn.; Fruct. Sem. Pl. 2: 128 (1791)

= Diasperus brunneus (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus cyrtophylloides (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus hongkongensis (Müll.Arg.) Kuntze; Revis. Gen. Pl. 2: 599 (1891)

= Diasperus nitidus (Roxb.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus obliquus (Willd.) Kuntze; Revis. Gen. Pl. 2: 600 (1891)

= Diasperus perakensis (Hook.f.) Kuntze; Revis. Gen. Pl. 2: 601 (1891)

= Diasperus zeylanicus (Gaertn.) Kuntze; Revis. Gen. Pl. 601 et 603 (1891)

= Glochidion brunneum Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 312 (1887)

= Glochidion brunneum subsp. andamanicum N.P.Balakr. & Chakr.; Proc. Indian Acad. Sci., Pl. Sci. 92(4): 357 (1983)

= Glochidion canaranum (Müll.Arg.) Bedd.; Fl. Sylv. S. India cxcii (1873)

= Glochidion ferdinandi var. supra-axillare (Benth.) F.M.Bailey; Queensl. Fl. 5: 1424 (1902)

= Glochidion glaberrimum Ridl.; Bull. Misc. Inform. Kew 1923: 363 (1923)

= Glochidion hongkongense Müll.Arg.; Linnaea 32: 60 (1863)

= Glochidion lanceolatum var. liukiuense (Hayata) Hurus.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 6: 332 (1954)

= Glochidion littorale Benth.; Fl. Hongk. 314 (1861), nom. illeg.

= Glochidion liukiuense Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: Art. I, 265 (1911)

= Glochidion nitidum (Roxb.) Voigt; Hort. Suburb. Calcutt. 153 (1845)

= Glochidion obliquum (Willd.) Decne.; Nouv. Ann. Mus. Hist. Nat. 3: 481 (1834)

= Glochidion pedunculatum Merr.; Philipp. J. Sci., C 11: 67 (1916)

= Glochidion perakense Hook.f.; Fl. Brit. India [J.D.Hooker] 5: 317 (1887)

= Glochidion perakense var. supraaxillare (Benth.) Airy Shaw; Kew Bull. 27(1): 72 (1972)

= Glochidion sumatranum Miq.; Fl. Ned. Ind., Eerste Bijv. 450 (1861)

= Glochidion supraaxillare (Benth.) Domin; Biblioth. Bot. 22(89): 318 (1927)

= Glochidion zeylanicum var. malayanum J.J.Sm.; Bijdr. Boomsoort. Java 12: 118 (1910)

= Glochidion zeylanicum var. nitidum (Roxb.) Haines; Bot. Bihar Orissa 2: 132 (1921)

= Phyllanthus canaranus Müll.Arg.; Flora 48: 871 (1865)

= Phyllanthus cyrtophylloides Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 1270 (1866)

= Phyllanthus ferdinandi var. supraaxillaris Benth.; Fl. Austral. [Bentham] 6: 96 (1873)

= Phyllanthus gaertneri T.Kuros.; J. Jap. Bot. 91: 119 (2016)

= Phyllanthus hongkongensis (Müll.Arg.) Müll.Arg.; Flora 48: 371 (1865)

= Phyllanthus nitidus (Roxb.) Müll.Arg.; Flora 48: 371 (1865)

= Phyllanthus obliquus (Willd.) Müll.Arg.; Flora 48: 371 (1865)

= Phyllanthus zeylanicus (Gaertn.) Müll.Arg.; Prodr. [A.P. de Candolle] 15(2): 281 (1866), nom. illeg.

variant :

var. paucicarpum Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 28(1): 123 (2004)

Distribution: Andaman Isl. (Middle Andaman Isl.)

41 AND

Synonyms: hidden

= Phyllanthus zeylanicus var. paucicarpus (Chakr. & N.P.Balakr.) Chakr. & N.P.Balakr.; J. Econ. Taxon. Bot. 33(3): 714 (2009)

= Phyllanthus obliquus var. paucicarpus (Chakrab. & N.P.Balakr.) Govaerts & Christenh.; Nelumbo 68(1): 143 (2026)

species :

Glochidion zollingeri Miq.; Fl. Ned. Ind. 1: t. 686 (1856)

Distribution: Java; Lesser Sunda Isl. (Bali); East Timor

42 LSI

Synonyms: hidden

= Diasperus zollingeri (Miq.) Kuntze; Revis. Gen. Pl. 601 et 603 (1891)

= Phyllanthus zollingeri (Miq.) Müll.Arg.; Flora 48: 371 (1865), nom. illeg.

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A complete, synonymic checklist of the Higher Plants of the World

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A complete, synonymic checklist of the Higher Plants of the World

Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
**in total 1,428,487* names**

[about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

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Search

+ Family

Filter

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Family (36)

Genus (67)

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A complete, synonymic checklist of the Higher Plants of the World

Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
**in total 1,428,487* names**

[about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]