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235.0000 Family Fabaceae Lindl.; Edwards´s Bot. Reg. 22: ad t. 1845 (1836), nom. cons., nom. alt.
728 genera, ca. 22.300 spp. Phylogeny and new subfamily system: Azani et al. (LPWG) (2017)
Species listed in database: 22404; hybrids: 149
= Acaciaceae E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 1: 164 (1836)
= Aspalathaceae Martinov; Tekhno-Bot. Slovar 51 (1820)
= Astragalaceae Bercht. & J.Presl; Prír. Rostlin 229 (1820)
= Bauhiniaceae Martinov; Tekhno-Bot. Slovar 67 (1820)
= Caesalpiniaceae R.Br.; Voy. Terra Austral. 2: 551 (1814), nom. cons.
= Cassiaceae Vest; Anleit. Stud. Bot. 270, 291 (1818)
= Ciceraceae Steele; Handb. Field Bot. [Steele] xx, 63 (1847)
= Coronillaceae Martinov; Tekhno-Bot. Slovar 162 (1820)
= Cytisaceae Bercht. & J.Presl; Prír. Rostlin 229 (1820)
= Dalbergiaceae Mart.; Consp. Regn. Veg. [Martius] 34 (1835)
= Daleaceae Bercht. & J.Presl; Prír. Rostlin 230 (1820)
= Detariaceae (DC.) J.Hess; Uebers. Phan. Nat. Pfl.-Fam. 46 (1832)
= Galedupaceae Martinov; Tekhno-Bot. Slovar 277 (1820)
= Geoffroeaceae Mart.; Consp. Regn. Veg. [Martius] 34 (1835)
= Hedysaraceae Bercht. & J.Presl; Prír. Rostlin 230 (1820)
= Inocarpaceae Zoll.; Syst. Verz. Ind. Archip. [Zollinger] 2: 117 (1854-1855)
= Lathyraceae Burnett; Outlines Bot. [Burnett] 660, 1092, 1138 (1835)
= Leguminosae Juss.; Gen. Pl. [Jussieu] 345 (1789), nom. cons.
= Lotaceae Oken; Lehrb. Naturgesch. 2(2.2): xv, 727 (1826)
= Mimosaceae R.Br.; Voy. Terra Austral. 2: 551 (1814), nom. cons.
= Papilionaceae Giseke; Prael. Ord. Nat. Pl. 415 (1792), nom. cons., nom. alt.
= Phaseolaceae Mart.; Consp. Regn. Veg. [Martius] 34 (1835)
= Robiniaceae Vest; Anleit. Stud. Bot. 270, 291 (1818)
= Sophoraceae Bercht. & J.Presl; Prír. Rostlin 229 (1820)
= Swartziaceae Bartl.; Ord. Nat. Pl. 231, 413 (1830)
= Tamarindaceae Martinov; Tekhno-Bot. Slovar 622 (1820)
= Trifoliaceae Bercht. & J.Presl; Prír. Rostlin 230 (1820)
= Viciaceae Oken; Lehrb. Naturgesch. 2(2.2): xv (1826)
235.8337 Genus Grona Lour.; Fl. Cochinch. 459 (1790)
Distribution: tropics and subtropics
Species listed in database: 49
species :
Grona adscendens (Sw.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 108 (2018)
Name: Budbud, Caddy, Kuzen, Pistache Marron, Sweet Heart, Wild Ground Nut (Trinidad & Tobago); Amor seco (Dominican Republic), Zarzabacoa galana (Puerto Rico)
Distribution: Senegal; Guinea-Bissau; Guinea; Sierra Leone; Liberia; Burkina Faso; Ivory Coast; Ghana; Togo; Benin; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; Annobon Isl.; Equatorial Guinea; Gabon; Central African Republic; D.R.Congo [Zaire]; Rwanda; Burundi; South Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Limpopo, KwaZulu-Natal); Madagascar; Mauritius; La Réunion; Seychelles; S-India; Sri Lanka; Myanmar; Thailand; Laos; Vietnam; peninsular Malaysia (Perak, Pahan); Sulawesi [I]; Philippines (Panay, Mindanao); Java; New Caledonia; Vanuatu [I]; Rotuma Isl.; Niue; Fiji; Cook Isl. [I]; Mexico (Campeche, Chiapas, Guerrero, Hidalgo, Oaxaca, Queretaro, Quintana Roo, San Luis Potosi, Tabasco, Tamaulipas, Veracruz); Guatemala; Belize; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola; Puerto Rico; St. Kitts; Antigua; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Tobago; Trinidad; Colombia (Amazonas, Antioquia, Caldas, Caquetá, Casanare, Cauca, Chocó, Cundinamarca, Guainía, Huila, La Guajira, Magdalena, Meta, Nariño, Putumayo, Santander, Valle, Vaupés, Vichada); Venezuela (Amazonas, Anzoategui, Apure, Aragua, Barinas, Bolívar, Delta Amacuro, Venezuela Distrito Federal, Lara, Merida, Miranda, Monagas, Sucre, Tachira, Trujillo, Yaracuy, Zulia); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Beni, Chuquisaca, Cochabamba, La Paz, Santa Cruz); N-Brazil (Roraima, Amapa, Pará, Amazonas, Acre, Rondônia); NE-Brazil (Maranhao, Ceara, Paraíba, Alagoas, Sergipe); WC-Brazil (Mato Grosso, Mato Grosso do Sul); SE-Brazil (Minas Gerais, Espirito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Rio Grande do Sul, Santa Catarina); Argentina (Misiones)
22 BEN BKN GHA GNB GUI IVO LBR NGA SEN SIE TOG 23 ANN BIO BUR CAF CMN EQG GAB PRI RWA SAO ZAI 24 ETH SSU 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 SAF 29 MAU MDG REU SEY 40 IND SRL 41 LAO MYA THA VIE 42 JAW MLY PHI sul 60 FIJ NUE NWC ROT van 61 coo 79 MEX 80 BLZ COS GUA HON NIC PAN 81 ABA CUB DOM GDL GRE HIS JAM MAR MTS PUE STK STL STV TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA 85 ARG
Synonyms: hidden
= Desmodium adscendens (Sw.) DC.; Prodr. [A.P. de Candolle] 2: 332 (1825)
= Desmodium adscendens f. glabrescens Schindl.; Repert. Spec. Nov. Regni Veg. 21: 8 (1925)
= Desmodium adscendens var. robustum B.G.Schub.; Bull. Jard. Bot. État Bruxelles 22: 290 (1954)
= Desmodium adscendens var. trifoliastrum (Miq.) Schindl.; Candollea 6: 484 (1936)
= Desmodium arinense Hoehne; Relat. Commiss. Linhas Telegr. Estratég. Matto Grosso Amazonas 8: 74 (1919)
= Desmodium caespitosum (Poir.) DC.; Prodr. [A.P. de Candolle] 2: 333 (1825)
= Desmodium coeruleum (Lindl.) G.Don; Gen. Hist. 2: 293 (1832)
= Desmodium ellipticum Macfad.; Fl. Jamaica [Macfadyen] 1: 268 (1837)
= Desmodium glaucescens Miq.; Linnaea 18: 549 (1844)
= Desmodium obovatum Vogel; Linnaea 12: 106 (1838)
= Desmodium ovalifolium Guill. & Perr.; Fl. Seneg. Tent. 208 (1832)
= Desmodium oxalidifolium Blume ex Miq.; Fl. Ned. Ind. 1(1): 249 (1855), nomen
= Desmodium oxalidifolium G.Don; Gen. Hist. 2: 294 (1832)
= Desmodium racemiferum DC.; Prodr. [A.P. de Candolle] 2: 331 (1825)
= Desmodium simplex G.Don; Gen. Hist. 2: 294 (1832)
= Desmodium thwaitesii Baker; Fl. Brit. India [J.D.Hooker] 2: 169 (1876)
= Desmodium trifoliastrum Miq.; Fl. Ned. Ind. 1(1): 248 (1858)
= Desmodium vogelii Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 496 (1841), nom. superfl.
= Grona adscendens var. robusta (B.G.Schub.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 108 (2018)
= Hedysarum adscendens Sw.; Prodr. [O.P.Swartz] 106 (1788)
= Hedysarum ascendens Lindl.; Bot. Reg. 10: t. 815 (1824), orth. var.
= Hedysarum caespitosum Poir.; Encycl. [J.Lamarck et al.] 6: 421 (1804)
= Hedysarum lindleyi Mart.; Ausw. Merkw. Pfl. 2: 25 (1830)
= Meibomia adscendens (Sw.) Hoehne; Album Secc. Bot. Mus. Paulista, São Paulo, 130 (1925), et Araucarilandia 100 (1930), isonym
= Meibomia adscendens (Sw.) Kuntze; Revis. Gen. Pl. 2: 195 (1891)
= Meibomia adscendens var. obovata (Vogel) Kuntze; Revis. Gen. Pl. 1: 195 (1891)
= Meibomia racemifera (DC.) Kuntze; Revis. Gen. Pl. 1: 198 (1891)
= Meibomia thwaitesii (Baker) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
= Meibomia trifoliastra (Miq.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
species :
Grona appressipila (B.G.Schub.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 108 (2018)
Distribution: Tanzania; Zambia
25 TAN 26 ZAM
Synonyms: hidden
= Desmodium appressipilum B.G.Schub.; Kew Bull. 25(1): 61 (1971)
species :
Grona auricoma (Graham ex Benth.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 109 (2018)
Distribution: Andaman Isl. [I]; Nicobar Isl. [I]; Myanmar (Mon, Taninthayi); Thailand; Laos; Cambodia; Vietnam; Java
41 and CBD LAO MYA ncb THA VIE 42 JAW
Synonyms: hidden
= Desmodium auricomum Graham; Numer. List [Wallich] no. 5704 (1831), nomen
= Desmodium auricomum Graham ex Benth.; Pl. Jungh. [Miquel] 2: 223 (1852)
= Meibomia auricoma (Graham ex Benth.) Kuntze; Revis. Gen. Pl. 1: 195 (1891)
variant :
var. pseudoauricoma (H.Ohashi) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 109 (2018)
Distribution: Cambodia; Vietnam
41 CBD VIE
Synonyms: hidden
= Desmodium auricomum var. pseudoauricomum H.Ohashi; J. Jap. Bot. 69(1): 26 (1994)
species :
Grona barbata (L.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 109 (2018)
Name: Zarzabacoa peluda
Distribution: Mexico (Chiapas, Colima, Guerrero, Jalisco, Mexico State, Michoacán, Nayarit, Oaxaca, Sinaloa, Tabasco, Veracruz); Guatemala; Belize; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola; Puerto Rico; Virgin Isl. (Tortola); Antigua; St. Lucia; Barbados; Trinidad; Colombia (Amazonas, Antioquia, Arauca, Caquetá, Casanare, Cauca, Córdoba, Cundinamarca, Guainía, Guaviare, Huila, Magdalena, Meta, Nariño, Santander, Tolima, Valle, Vichada); Venezuela (Amazonas, Anzoategui, Apure, Aragua, Barinas, Bolívar, Carabobo, Cojedes, Delta Amacuro, Venezuela Distrito Federal, Falcon, Guarico, Lara, Merida, Miranda, Monagas, Portuguesa, Sucre, Tachira, Trujillo, Zulia); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Beni, La Paz, Santa Cruz); N-Brazil (Pará, Acre, Rondônia); NE-Brazil (Maranhao, Ceara, Bahia, Sergipe); WC-Brazil (Mato Grosso, Goiás, Brasilia Distrito Federal, Mato Grosso do Sul); SE-Brazil (Minas Gerais, Espirito Santo, São Paulo, Rio de Janeiro); S-Brazil (Paraná, Rio Grande do Sul, Santa Catarina); Paraguay (Caazapa, Canindeyu, Central, Guaira, Itapua); Argentina (Chaco, Corrientes, Misiones)
79 MEX 80 BLZ COS GUA HON NIC PAN 81 ABA BBD CUB HIS JAM PUE STL TRT VIR 82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA 85 ARG PAR
Synonyms: hidden
= Desmodium barbatum (L.) Benth. & Oerst.; Vidensk. Meddel. Naturhist. Foren. Kjøbenhavn, Ser. 1, 3: 18 (1853)
= Desmodium barbatum (L.) Benth.; Pl. Jungh. [Miquel] 1: 224 (1852)
= Desmodium barbatum var. venustulum (Kunth) Griseb.; Fl. Brit. W. I. [Grisebach] 186 (1859)
= Desmodium coeruleoviolaceum (G.Mey.) DC.; Prodr. [A.P. de Candolle] 2: 331 (1825)
= Desmodium venustulum (Kunth) Hemsl.; Biol. Centr.-Amer., Bot. 1: 290 (1880)
= Desmodium villosum (Cham. & Schltdl.) Hemsl.; Biol. Centr.-Amer., Bot. 1: 291 (1880), nom. illeg.
= Desmodium villosum (Mill.) DC.; Prodr. [A.P. de Candolle] 2: 338 (1825)
= Hedysarum barbatum L.; Syst. Nat., ed. 10, 2: 1170 (1759)
= Hedysarum coeruleoviolaceum G.Mey.; Prim. Fl. Esseq. 246 (1818)
= Hedysarum ildefonsianum Mure; Doctrine Ecole Rio de Janeiro Pathog. Brésil. 352 (1849)
= Hedysarum lagocephalum Link; Enum. Hort. Berol. Alt. 2: 248 (1822)
= Hedysarum procumbens Vell.; Fl. Flumin. 319 (1829), nom. illeg.
= Hedysarum venustulum Kunth; Nov. Gen. Sp. [H.B.K.] 6: 519 (1824)
= Hedysarum villosum Mill.; Gard. Dict., ed. 8: no. 9 (1768)
= Meibomia barbata (L.) Hoehne; Araucarilandia 100 (1930), isonym
= Meibomia barbata (L.) Kuntze; Revis. Gen. Pl. 2: 195 (1891)
= Meibomia cayennensis (DC.) Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia neglecta Kuntze; Revis. Gen. Pl. 2: 198 (1891)
= Meibomia villosa (Mill.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
= Nicolsonia barbata (L.) DC.; Prodr. [A.P. de Candolle] 2: 325
= Nicolsonia cayennensis DC.; Mém. Légum. 7: 314, t. 51 (1825)
= Nicolsonia cayennensis var. laxiuscula DC.; Prodr. [A.P. de Candolle] 2: 325 (1825)
= Nicolsonia cayennensis var. obovata DC.; Mém. Légum. 7: 314 (1825)
= Nicolsonia major Steud.; Flora 26: 757 (1843)
= Nicolsonia radicans Steud.; Flora 26: 757 (1843)
= Nicolsonia venustula (Kunth) DC.; Prodr. [A.P. de Candolle] 2: 325 (1825)
= Nicolsonia villosa (Mill.) Cham. & Schltdl.; Linnaea 5: 584 (1830)
= Perrottetia barbata (L.) DC.; Ann. Sci. Nat. (Paris) 4: 96 (1825)
= Perrottetia cayennensis DC.; Ann. Sci. Nat. (Paris) 4: 96 (1825)
= Perrottetia venustula (Kunth) DC.; Ann. Sci. Nat. (Paris) 4: 96 (1825)
= Urania barbata (L.) Desv.; Mém. Soc. Linn. Paris 4: 309 (1826)
= Uraria lagocephala (Link) DC.; Prodr. [A.P. de Candolle] 2: 324 (1825)
variant :
var. argyrea (Baker) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 109 (2018)
Distribution: Angola; Namibia
26 ANG 27 NAM
Synonyms: hidden
= Desmodium barbatum var. argyreum (Welw. ex Baker) B.G.Schub.; Bull. Jard. Bot. État Bruxelles 22: 298 (1952)
= Desmodium dimorphum var. argyreum Welw. ex Baker; Fl. Trop. Afr. [Oliver et al.] 2: 161 (1871)
= Nicolsonia barbata var. argyrae (Welw. ex Baker) Schindl.
variant :
var. dimorpha (Welw. ex Baker) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 109 (2018)
Distribution: Guinea; Burkina Faso; Benin; Nigeria; Cameroon; Gabon; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; South Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; Botswana; South Africa (Limpopo, Mpulamanga); Madagascar
22 BEN BKN GUI NGA 23 BUR CMN GAB RWA ZAI 24 ETH SSU SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 BOT SAF 29 MDG
Synonyms: hidden
= Desmodium barbatum subsp. dimorphum (Welw. ex Baker) J.R.Laundon
= Desmodium barbatum var. dimorphum (Welw. ex Baker) B.G.Schub.; Bull. Jard. Bot. État Bruxelles 22: 298 (1952)
= Desmodium barbatum var. emirneense Baker; (1882)
= Desmodium dimorphum Welw. ex Baker; Fl. Trop. Afr. [Oliver et al.] 2: 161 (1871)
= Meibomia dimorpha (Welw. ex Baker) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
= Nicolsonia barbata var. dimorpha (Welw. ex Baker) Schindl.
variant :
var. procumbens (B.G.Schub.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 109 (2018)
Distribution: trop. Africa; Madagascar
29 MDG
Synonyms: hidden
= Desmodium campenonii Drake; Hist. Phys. Madagascar [Grandidier] 1: 172 (1903)
= Desmodium barbatum var. nummularifolium Baker; (1882)
= Desmodium barbatum var. procumbens B.G.Schub.; Bull. Jard. Bot. État Bruxelles 22: 297 (1952)
species :
Grona brachystachya (Graham ex Benth.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 110 (2018)
Distribution: India (Andhra Pradesh, Bihar, Chhattisgarh, Jharkhand, Maharashtra, Tamil Nadu, Uttarakhand, Uttar Pradesh, West Bengal)
40 IND
Synonyms: hidden
= Desmodium benthamii N.P.Balakr.; J. Bombay Nat. Hist. Soc. 63: 328 (1967), nom. superfl.
= Desmodium brachystachyum Graham ex Benth.; Pl. Jungh. [Miquel] 223 (1852)
= Meibomia brachystachya (Graham ex Benth.) Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Nicolsonia brachystachya (Graham ex Benth.) Schindl.; Repert. Spec. Nov. Regni Veg. 23: 359 (1927)
species :
Grona brevipedicellata (W.Saisorn, Chantar. & Balslev) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 110 (2018)
Distribution: Thailand
41 THA
Synonyms: hidden
= Desmodium brevipedicellatum W.Saisorn, Chantar. & Balslev; Phytotaxa 263(2): 154 (2016)
species :
Grona brownii (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 110 (2018)
Distribution: Australia (Western Australia, Northern Territory)
50 AUS
Synonyms: hidden
= Desmodium neurocarpum var. gracile Benth.; Fl. Austral. [Bentham] 2: 234 (1864)
= Desmodium brownii Schindl.; Notes Roy. Bot. Gard. Edinburgh 15: 131 (1926)
species :
Grona afra (E.Mey.) H.Ohashi & K.Ohashi; J. Jap. Bot. 94(3): 143 (2019)
Distribution: Burundi; Uganda; Kenya; Tanzania; Angola; Zambia; Mozambique; South Africa (Mpulamanga, KwaZulu-Natal, E-Cape Prov.)
23 BUR 25 KEN TAN UGA 26 ANG MOZ ZAM 27 SAF
Synonyms: hidden
= Desmodium afrum (E.Mey.) Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 619 (1916 publ. 1917), nom. illeg.
= Desmodium afrum (E.Mey.) Schindl.; Repert. Spec. Nov. Regni Veg. 23: 360 (1927)
= Desmodium dregeanum Benth.; Pl. Jungh. [Miquel] 222 (1852)
= Meibomia afra (E.Mey.) Kuntze; Revis. Gen. Pl. 2: 197 (1891)
= Nicolsonia afra E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 123 (1836)
species :
Grona cordifolia (Harms) H.Ohashi & K.Ohashi; J. Jap. Bot. 94(3): 144 (2019)
Distribution: Central African Republic; D.R.Congo [Zaire]; Tanzania; Angola; Zambia; Malawi
23 CAF ZAI 25 TAN 26 ANG MLW ZAM
Synonyms: hidden
= Desmodium cordifolium (Harms) Schindl.; Repert. Spec. Nov. Regni Veg. 22: 257 (1926)
= Desmodium homblei De Wild.; Repert. Spec. Nov. Regni Veg. 13: 114 (1914)
= Glycine cordifolia Harms; Bot. Jahrb. Syst. 49: 441 (1913)
species :
Grona ferruginea (Wall. ex Thwaites) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 110 (2018)
Distribution: S-India (Andhra Pradesh, Karnataka, Kerala, Madhya Pradesh, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Desmodium ferrugineum Wall.; Numer. List [Wallich] no. 5732 (1831), nomen
= Desmodium ferrugineum Wall. ex Thwaites; Enum. Pl. Zeyl. [Thwaites] 87 (1859)
species :
Grona filiformis (Zoll. & Moritzi) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 111 (2018)
Distribution: Java; Lesser Sunda Isl.; New Guinea (Irian Jaya, Papua New Guinea); Australia (Western Australia, Northern Territory, Queensland)
42 JAW LSI NWG 50 AUS
Synonyms: hidden
= Desmodium filiforme Zoll. & Moritzi; Natuur- Geneesk. Arch. Ned.-Indië 3: 77 (1846)
= Desmodium neurocarpum Benth.; Fl. Austral. [Bentham] 2: 234 (1864)
= Desmodium neurocarpum var. queenslandicum Domin; Biblioth. Bot. 22(89): 214 (1926)
= Meibomia filiformis (Zoll. & Moritzi) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
= Meibomia neurocarpa (Benth.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
species :
Grona flagellaris (Benth.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 111 (2018)
Distribution: Australia (Western Australia, Northern Territory, Queensland)
50 AUS
Synonyms: hidden
= Desmodium flagellare Benth.; Fl. Austral. [Bentham] 2: 233 (1864)
= Meibomia flagellaris (Benth.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
species :
Grona fulvescens (B.G.Schub.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 111 (2018)
Distribution: D.R.Congo [Zaire]; Zambia
23 ZAI 26 ZAM
Synonyms: hidden
= Desmodium fulvescens B.G.Schub.; Bull. Jard. Bot. État Bruxelles 22: 296 (1952)
species :
Grona glareosa (Pedley) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 111 (2018)
Distribution: Australia (Northern Territory, Western Australia)
50 AUS
Synonyms: hidden
= Desmodium glareosum Pedley; Austrobaileya 5(2): 242 (1999)
species :
Grona griffithiana (Benth.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 111 (2018)
Distribution: China (Guizhou, Sichuan, Yunnan); India (Assam, Manipur, Meghalaya, Nagaland, Tripura); Myanmar; Thailand; Vietnam
36 CHI 40 IND 41 MYA THA VIE
Synonyms: hidden
= Desmodium griffithianum Benth.; Pl. Jungh. [Miquel] 222 (1852)
= Desmodium griffithianum var. leiocarpum X.F.Gao & C.Chen; Acta Bot. Yunnan. 20: 270 (1998)
= Desmodium oxalidifolium H.Lév.; Fl. Kouy-Tcheou 233. 1914-15
= Meibomia griffithiana (Benth.) Kuntze; Revis. Gen. Pl. 1: 198 (1891)
species :
Grona hannii (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 111 (2018)
Distribution: peninsular Malaysia
42 MLY
Synonyms: hidden
= Desmodium hannii Schindl.; Repert. Spec. Nov. Regni Veg. (1925)
species :
Grona harmsii (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 112 (2018)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Desmodium harmsii Schindl.; Bot. Jahrb. Syst. 54: 55 (1916)
species :
Grona helenae (Buscal. & Muschl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 112 (2018)
Distribution: D.R.Congo [Zaire]; Angola; Zambia
23 ZAI 26 ANG ZAM
Synonyms: hidden
= Desmodium helenae Buscal. & Muschl.; Bot. Jahrb. Syst. 49: 472 (1913)
species :
Grona heterocarpa (L.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 112 (2018)
Distribution: Ghana [I]; Tanzania [I]; Seychelles; China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); South Korea; Japan (Honshu, Shikoku, Kyushu); Ryukyu Isl.; Volcano Isl. [I] (Iwojima [I]); Taiwan; Nepal; Bhutan; India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Dadra-Nagar-Haveli, Delhi, Goa, Gujarat, Haryana, Himachal Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Pondicherry, Punjab, Rajasthan, Tamil Nadu, Tripura, Uttar Pradesh, West Bengal, Yanan, Sikkim, Jammu & Kashmir); Sri Lanka; Bangladesh; Andaman Isl. (North Andamans, Middle Andamans, South Andamans, Little Andaman Isl., Barren Isl.); Nicobar Isl. (Car Nicobar Isl., North Nicobars, Central Nicobars, Great Nicobar Isl., Little Nicobar Isl.); Myanmar (Magway, Mandalay, Sagaing); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Singapore [I]; Sumatra; Borneo; Sulawesi; Philippines (Luzon, Mindanao); Java; Lesser Sunda Isl.; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Solomon Isl.; Australia (Queensland); New Caledonia; Vanuatu; Tonga (´Eua, Kao, Vava´u, Niuafo´ou); Society Isl. (Tahiti, Mehetia, Huahine, Bora Bora, Raiatea); Gambier Isl. (Mangareva); Tuamotu Arch. (Makatea Isl.); Austral Isl. (Raivavae, Rurutu, Tubuai); Palau (Babeldaob, Koror, Ngarakabesang); Nicaragua [I]; Costa Rica [I]; St. Vincent [I]; Trinidad & Tobago [I]; Ecuador [I]; Peru [I]; Bolivia [I]
22 gha 25 tan 29 SEY 36 CHI 38 JAP KOR kzn NNS TAI 40 BAN BHU IND NEP SRL 41 AND CBD LAO MYA NCB THA VIE 42 BOR JAW LSI MLY NWG PHI sng SUL SUM 43 BIS SOL 50 AUS 60 NWC TON VAN 61 AUI GBI SCI TUA 62 PLU 80 cos nic 81 stv trt 83 bol ecu per
Synonyms: hidden
= Desmodium buergeri Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 45 (1867)
= Desmodium heterocarpon (L.) DC.; Prodr. [A.P. de Candolle] 2: 334 (1825)
= Desmodium heterocarpon f. albiflorum (Ridl.) H.Ohashi; J. Jap. Bot. 66: 20 (1991)
= Desmodium heterocarpon var. buergeri (Miq.) Hosok.; J. Soc. Trop. Agric. 4: 201 (1932)
= Desmodium ovalifolium Wall.; Numer. List [Wallich] no. 5730 (1831) ex Ridl., Fl. Malay. Peninsula 1: 610 (1922)
= Desmodium polycarpon f. hirsutum Pamp.; Nuovo Giorn. Bot. Ital., n. s. 17: 15 (1910)
= Desmodium polycarpon var. albiflorum Ridl.; Fl. Malay Penins. 1: 609 (1922)
= Desmodium polycarpon var. trichocaulum (DC.) Benth.; Fl. Brit. India [J.D.Hooker] 2: 172 (1876)
= Desmodium spectabile Sweet; Hort. Brit. [Sweet] 479 (1826), nomen
= Desmodium toppinii Schindl.; Repert. Spec. Nov. Regni Veg. 21: 5 (1925)
= Desmodium trichocaulon DC.; Ann. Sci. Nat. (Paris) 4: 101 (1825)
= Desmodium trinerve Graham; Numer. List [Wallich] no. 5720 (1831)
= Desmodium tuberculatum G.Don; Gen. Hist. 2: 298 (1832)
= Grona repens Lour.; Fl. Cochinch. 459 (1790)
= Hedysarum heterocarpon L.; Sp. Pl. [Linnaeus] 2: 747 (1753)
= Hedysarum patens Roxb.; Fl. Ind. [Roxburgh] 2: 362 (1832)
= Hedysarum purpureum Roxb.; Fl. Ind. [Roxburgh] 2: 358 (1832), nom. illeg.
= Hedysarum spectabile Bojer ex Sweet; Hort. Brit. [Sweet] 479 (1826), pro syn.
= Hedysarum tuberculosum Labill.; Sert. Austro-Caledon. 71 (1825)
= Hippocrepis barbata Lour.; Fl. Cochinch. 453 (1790)
= Meibomia buergeri (Miq.) Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia heterocarpon (L.) Kuntze; Revis. Gen. Pl. 1: 196 (1891)
= Pleurolobus heterocarpus (L.) J.St.-Hil.; J. Bot. [Desvaux] 1: 61 (1813)
variant :
var. gymnocarpa (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 112 (2018)
Distribution: S-India (Karnataka, Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Desmodium heterocarpum var. gymnocarpum Schindl.; Repert. Spec. Nov. Regni Veg. 21: 5 (1925)
variant :
var. strigosa (Meeuwen) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 112 (2018)
Distribution: China (Guangdong, Guangxi, Hainan, S-Yunnan); Taiwan; Nepal; Bhutan; India (widespread); Sri Lanka; Myanmar (widespread); Thailand; Laos; Vietnam; peninsular Malaysia; Philippines; Australia (Northern Territory, Queensland); ?New Caledonia; Niue [I]; Fiji; Tonga; American Samoa (Ta´u, Ofu, Tutuila); [Western] Samoa (Savai´i, ´Upolu); Cook Isl. (Rarotonga, Mangaia, Ngaputoru); Marquesas Isl. [I] (Nuku Hiva [I], Ua Huka [I], Hiva Oa [I]); Fed. States of Micronesia (Yap, Truk, Aloen, Pohnpei); Hawaii [I] (Oahu [I])
36 CHI 38 TAI 40 BHU IND NEP SRL 41 LAO MYA THA VIE 42 MLY PHI 50 AUS 60 ASA FIJ nue SAM TON 61 COO mrq 62 MIC 63 haw
Remarks: native status in Pacific disputed
Synonyms: hidden
= Desmodium distans Royle ex Baker; Fl. Brit. India [J.D.Hooker] 2: 172 (1876)
= Desmodium gyroides Hassk.; Pl. Jav. Rar. [Hasskarl] 362 (1848), nom. illeg.
= Desmodium heterocarpon f. strigosum (Meeuwen) Fosberg; Micronesica 2: 145 (1966 publ. 1967)
= Desmodium heterocarpon var. strigosum Meeuwen; Reinwardtia 6: 95 (1961)
= Desmodium hippocrepis DC.; Prodr. [A.P. de Candolle] 2: 338 (1825)
= Desmodium nervosum Vogel; Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19, Suppl. 1: 28 (1843)
= Desmodium patens G.Don ex Wight; Icon. Pl. Ind. Orient. [Wight] 2: t. 407 (1840)
= Desmodium polycarpum (Poir.) DC.; Prodr. [A.P. de Candolle] 2: 334 (1825)
= Desmodium purpureum (Roxb.) Hook. & Arn.; Bot. Beechey Voy. 62 (1833)
= Desmodium serpens Wall.; Numer. List [Wallich] no. 5733 (1831), nomen
= Desmodium siliquosum (Burm.f.) DC.; Prodr. [A.P. de Candolle] 2: 336 (1825)
= Hedysarum polycarpon Poir.; Encycl. [J.Lamarck et al.] 6: 413 (1805)
= Hedysarum siliquosum Burm.f.; Fl. Ind. [N.L.Burman] 169, t. 55, f. 2 (1768)
species :
Grona heterophylla (Willd.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 114 (2018)
Distribution: Madagascar [I]; China (Anhui, Fujian, Guangdong, Guangxi, Hainan, Jiangxi, Yunnan); Ryukyu Isl.; Bonin Isl. (Chichijima); Taiwan; Nepal; India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Goa, Gujarat, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Pondicherry, Sikkim, Tamil Nadu, Tripura, West Bengal); Sri Lanka; Bangladesh; Nicobar Isl. (North Nicobars, Central Nicobars); Myanmar (Yangon); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Singapore [I]; Sumatra; South China Sea Isl.; Borneo; Brunei; Philippines (throughout); New Guinea (Irian Jaya, Papua New Guinea); Australia (Queensland); New Caledonia [I]; Fiji [I]; Cook Isl. [I] (Rarotonga [I], Atiu [I], ?Aitutaki [I]); [Western] Samoa [I] (´Upolu [I], Savai´i [I]); Northern Marianas [I] (Pagan [I], Alamagan [I], Saipan [I], Tinian [I], Rota [I]); Guam [I]; Fed. States of Micronesia [I] (Yap [I], Truk [I], Pohnpei [I]); Hawaii [I] (Molokai [I], Maui [I], Hawaii Isl. [I])
29 mdg 36 CHI 38 NNS OGA TAI 40 BAN IND NEP SRL 41 CBD LAO MYA NCB THA VIE 42 BOR BRN MLY NWG PHI SCS sng SUM 50 AUS 60 fij nwc sam 61 coo 62 gum mic mrn 63 haw
Synonyms: hidden
= Desmodium heterophyllum (Willd.) DC.; Prodr. [A.P. de Candolle] 2: 334 (1825)
= Desmodium triflorum var. majus Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 229 (1834)
= Desmodium triflorum Wall. ex Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 229 (1834), nom. illeg.
= Hedysarum heterophyllum Willd.; Sp. Pl., ed. 4 [Willdenow] 3(2): 1202-1203 (1802)
= Hedysarum reptans Roxb.; (Hort. Bengal. 57 (1814), nomen), Fl. Ind. [Roxburgh] 3: 354 (1832)
= Meibomia heterophylla (Willd.) Kuntze; Revis. Gen. Pl. 1: 196 (1891)
species :
Grona hirta (Guill. & Perr.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 114 (2018)
Distribution: Cape Verde Isl. (Ilha de Maio, Fogo Isl.); Senegal; Gambia; Guinea-Bissau; Guinea; Sierra Leone; Mali; Burkina Faso; Ghana; Togo; Benin; Niger; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Equatorial Guinea; Gabon; Central African Republic; D.R.Congo [Zaire]; Burundi; Chad; South Sudan; Ethiopia; Tanzania; Angola; Zambia; Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Eswatini [Swaziland]; Madagascar
21 CVI 22 BEN BKN GAM GHA GNB GUI MLI NGA NGR SEN SIE TOG 23 BIO BUR CAF CMN EQG GAB ZAI 24 CHA ETH SSU 25 TAN 26 ANG MOZ ZAM ZIM 27 SAF SWZ 29 MDG
Synonyms: hidden
= Desmodium boivinianum Baill.; Bull. Mens. Soc. Linn. Paris 1: 431 (1884)
= Desmodium hirtum Guill. & Perr.; Fl. Seneg. Tent. 209 (1832)
= Desmodium humblotianum Baill.; Bull. Mens. Soc. Linn. Paris 1: 431 (1884)
= Desmodium monospermum Boivin ex Baill.; Bull. Mens. Soc. Linn. Paris 1: 431 (1884)
= Meibomia hirta (Guill. & Perr.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
variant :
var. delicatula (A.Rich.) H.Ohashi & K.Ohashi; J. Jap. Bot. 94(3): 144 (2019)
Distribution: D.R.Congo [Zaire]; Ethiopia; Zambia; Mozambique; Madagascar
23 ZAI 24 ETH 26 MOZ ZAM 29 MDG
Synonyms: hidden
= Desmodium delicatulum A.Rich.; Tent. Fl. Abyss. 1: 205 (1847)
= Desmodium hirtum f. compressum B.G.Schub.; Bull. Jard. Bot. État Bruxelles 22: 288 (1952)
= Desmodium hirtum var. delicatulum (A.Rich.) Harms ex Baker f.; Legum. Trop. Africa [E.G.Baker] 329 (1929)
= Meibomia delicatula (A.Rich.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
species :
Grona jucunda (Thwaites) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 114 (2018)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Desmodium jucundum Thwaites; Enum. Pl. Zeyl. [Thwaites] 411 (1859)
= Meibomia jucunda (Thwaites) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
species :
Grona juruenensis (Hoehne) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
Distribution: N-Brazil (Rondônia); WC-Brazil (Mato Grosso)
84 BRA
Synonyms: hidden
= Desmodium juruenense Hoehne; Comiss. Linhas Telegr. Estratég. Mato Grosso Amazonas, Annexo 5, Bot. 8: 73, t. 148, f. 1 (1919)
= Nicolsonia juruenensis (Hoehne) Schindl.; Repert. Spec. Nov. Regni Veg. 23: 359 (1927)
= Meibomia juruenensis (Hoehne) Hoehne; Anexos Mem. Inst. Butantan, Secc. Bot. 1(1): 33 (1921)
species :
Grona laosensis H.Ohashi, K.Ohashi & Tagane; J. Jap. Bot. 96(5): 270 (2021)
Distribution: Laos
41 LAO
species :
Grona muelleri (Benth.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
Distribution: Australia (Western Australia, Northern Territory, Queensland)
50 AUS
Synonyms: hidden
= Meibomia muelleri (Benth.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
= Desmodium muelleri Benth.; Fl. Austral. [Bentham] 2: 235 (1864)
species :
Grona nemorosa (F.Muell. ex Benth.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
subspecies :
subsp. nemorosa
Distribution: New Guinea (Papua New Guinea); Australia (Queensland, New South Wales)
42 NWG 50 AUS
Synonyms: hidden
= Desmodium archboldianum Baker f. [Brittonia 2: 318 (1937)]
= Desmodium nemorosum F.Muell. ex Benth.; Fl. Austral. [Bentham] 2: 234 (1864)
= Desmodium nemorosum subvar. eboracense Schindl.; Repert. Spec. Nov. Regni Veg. 21: 10 (1925)
= Desmodium nemorosum var. simplex Schindl.; Repert. Spec. Nov. Regni Veg. 21: 10 (1925)
= Meibomia nemorosa (F.Muell. ex Benth.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
subspecies :
subsp. whitfordii (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
Distribution: Philippines (Luzon); Australia (Queensland)
42 PHI 50 AUS
Synonyms: hidden
= Desmodium nemorosum subsp. whitfordii (Schindl.) H.Ohashi; J. Jap. Bot. 79(2): 128 (2004)
= Desmodium nemorosum subvar. whitfordii (Schindl.) H.Ohashi; Ginkgoana 1: 220 (1973)
= Desmodium nemorosum var. whitfordii (Schindl.) H.Ohashi; Ginkgoana 1: 220 (1973)
= Desmodium whitfordii (Schindl.) Pedley; Austrobaileya 5(2): 247 (1999)
species :
Grona orinocensis (DC.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
Distribution: Colombia (Amazonas, Caquetá, Guainía, Putumayo, Vichada); Venezuela (Amazonas)
82 CLM VEN
Synonyms: hidden
= Desmodium orinocense (DC.) Cuello; Novon 4(2): 98 (1994)
= Desmodium adscendens var. orinocense DC.; Prodr. [A.P. de Candolle] 2: 332 (1825)
= Nicolsonia orinocensis (DC.) Schindl.; Repert. Spec. Nov. Regni Veg. 23: 358 (1927)
species :
Grona ovalifolia (Prain) H.Ohashi, K.Ohashi & Tagane; J. Jap. Bot. 96(5): 270 (2021)
Distribution: Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia; Sulawesi; Philippines (Luzon)
41 CBD LAO THA VIE 42 MLY PHI SUL
Synonyms: hidden
= Desmodium heterocarpon subsp. ovalifolium (Prain) H.Ohashi; J. Jap. Bot. 66: 21 (1991)
= Desmodium heterocarpon var. ovalifolium (Prain) Rugayah; Berita Biol. 3: 205 (1986)
= Desmodium ovalifolium (Prain) Wall. ex Ridl.; Fl. Malay Penins. 1: 610 (1922)
= Desmodium polycarpum var. ovalifolium Prain; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 66: 141 (1897)
= Grona heterocarpos subsp. ovalifolia (Prain) H.Ohashi & K.Ohashi; J. Jap. Bot. 93: 114 (2018)
species :
Grona pullenii (Pedley) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
Distribution: Lesser Sunda Isl.; Australia (Northern Territory, Western Australia, Queensland)
42 LSI 50 AUS
Synonyms: hidden
= Desmodium pullenii Pedley; Austrobaileya 5(2): 243 (1999)
species :
Grona pycnotricha (Pedley) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
Distribution: Australia (Northern Territory, Western Australia)
50 AUS
Synonyms: hidden
= Desmodium pycnotrichum Pedley; Austrobaileya 5(2): 236 (1999)
species :
Grona ramosissima (G.Don) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 115 (2018)
Distribution: Guinea-Bissau; Guinea; Sierra Leone; Liberia; Mali; Burkina Faso; Ivory Coast; Ghana; Togo; Benin; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; Annobon Isl.; Equatorial Guinea; Gabon; Central African Republic; D.R.Congo [Zaire]; Rwanda; Burundi; Chad; South Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Mozambique; Madagascar; Comores (Mayotte, etc.); Mauritius; La Réunion; Rodrigues; Seychelles [I]
22 BEN BKN GHA GNB GUI IVO LBR MLI NGA SIE TOG 23 ANN BIO BUR CAF CMN EQG GAB PRI RWA SAO ZAI 24 CHA ETH SSU 25 KEN TAN UGA 26 ANG MOZ ZAM 29 COM MAU MDG REU ROD sey
Synonyms: hidden
= Desmodium fruticulosum Desv.; Mém. Soc. Linn. Paris 4: (1826) 319 (non Walp. 1842)
= Desmodium ramosissimum G.Don; Gen. Hist. 2: 294 (1832)
= Hedysarum fruticulosum Desv.; Mém. Soc. Linn. Paris 4: 319 (1826), nom. illeg.
= Hedysarum fruticulosum Schumach. & Thonn.; Beskr. Guin. Pl. [Schumacher] 363 (1827)
species :
Grona reticulata (Champ. ex Benth.) H.Ohashi & K.H.Ohashi; J. Jap. Bot. 96(1): 13 (2021)
Distribution: China (Guangdong, Guangxi, Hainan, S-Yunnan); Myanmar (Bago); Thailand; Cambodia
36 CHI 41 CBD MYA THA
Synonyms: hidden
= Desmodium heterocarpon f. pilosum (Craib) H.Ohashi; J. Jap. Bot. 66(1): 20 (1991)
= Desmodium heterocarpon subsp. angustifolium (Benth. ex Craib) H.Ohashi; Ginkgoana 1: 212 (1973) (nom. nov., as ´comb. nov.´)
= Desmodium polycarpon var. angustifolium Craib; Bull. Misc. Inform. Kew 1911: 38 (1911), nomen
= Desmodium polycarpon var. rigidum Ridl.; J. Straits Branch Roy. Asiat. Soc. 59: 97 (1911)
= Desmodium reticulatum Champ. ex Benth.; Hooker´s J. Bot. Kew Gard. Misc. 4: 46 (1852)
= Desmodium reticulatum var. pilosum Craib; Fl. Siam 1: 416 (1928)
= Grona heterocarpa subsp. angustifolia (H.Ohashi) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 114 (2018)
= Meibomia reticulata (Champ. ex Benth.) Kuntze; Revis. Gen. Pl. 1: 198 (1891)
species :
Grona retroflexa (L.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(6): 385 (2018)
Distribution: India (Assam, Himachal Pradesh, Karnataka, Kerala, Manipur, Meghalaya, Sikkim, Tamil Nadu, West Bengal); Sri Lanka; Thailand
40 IND SRL 41 THA
Remarks: sometimes included in G. styracifolia
Synonyms: hidden
= Codariocalyx capitatus (Burm.f.) Hassk.; Flora 25(2. Beibl.): 49 (1842)
= Desmodium capitatum (Burm.f.) DC.; Prodr. [A.P. de Candolle] 2: 336 (1825)
= Desmodium retroflexum (L.) DC.; Prodr. [A.P. de Candolle] 2: 336 (1825)
= Grona capitata (Burm.f.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 110 (2018)
= Hedysarum capitatum Burm.f.; Fl. Ind. [N.L.Burman] 167 (1768)
= Hedysarum retroflexum L.; Mant. Pl. 1: 103 (1767)
= Meibomia capitata (Burm.f.) Kuntze; Revis. Gen. Pl. 1: 195 (1891)
= Meibomia retroflexa (L.) Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Pseudarthria capitata (Burm.f.) Hassk.; Cat. Hort. Bot. Bogor. [Hasskarl] 281 (1844)
= Uraria retroflexa (L.) Drake; J. Bot. [Morot] 5: 192 (1891)
species :
Grona rubra (Lour.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: China (Guangdong, Guangxi, Hainan); Laos; Cambodia; Vietnam; peninsular Malaysia
36 CHI 41 CBD LAO VIE 42 MLY
Synonyms: hidden
= Desmodium carlesii Schindl.; Bot. Jahrb. Syst. 54(1): 56 (1917)
= Hedysarum rubrum (Lour.) Spreng.; Syst. Veg., ed. 16 [Sprengel] 3: 317 (1826)
= Meibomia rubra (Lour.) Kuntze; Revis. Gen. Pl. 1: 198 (1891)
= Ornithopus ruber Lour.; Fl. Cochinch. 2: 452-453 (1790)
= Desmodium rubrum (Lour.) DC.; Prodr. [A.P. de Candolle] 2: 327 (1825)
variant :
var. macrocarpa (H.Ohashi) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Desmodium rubrum var. macrocarpum H.Ohashi; Ginkgoana 1: 222 (1973)
variant :
var. uncinata (H.Ohashi) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Desmodium rubrum var. uncinatum H.Ohashi; Fl. Cambodge Laos Vietnam 27: 128 (1994)
species :
Grona saulierei (Schindl.) Kottaim.; Int. J. Curr. Res. Biosci. Pl. Biol. 6(10): 38 (2019)
Distribution: S-India (Tamil Nadu)
40 IND
Synonyms: hidden
= Desmodium barbatum subsp. saulierei (Schindl.) H.Ohashi; Ginkgoana 1: 203 (1973)
= Grona barbata subsp. saulierei (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 109 (2018)
= Nicolsonia barbata var. saulierei Schindl.; Repert. Spec. Nov. Regni Veg. 23: 359 (1927)
species :
Grona schubertiae (H.Ohashi) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: Cambodia; Vietnam
41 CBD VIE
Synonyms: hidden
= Desmodium schubertiae H.Ohashi; J. Arnold Arbor. 71(3): 381 (1990)
species :
Grona schweinfurthii (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: Sierra Leone; Nigeria; Central African Republic; D.R.Congo [Zaire]; Chad; South Sudan; Ethiopia
22 NGA SIE 23 CAF ZAI 24 CHA ETH SSU
Synonyms: hidden
= Desmodium schweinfurthii Schindl.; Bot. Jahrb. Syst. 54: 57 (1916)
species :
Grona setigera (E.Mey.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: Senegal; Gambia; Equatorial Guinea; S-Chad; South Sudan; Ethiopia; Zimbabwe; Mozambique; Madagascar
22 GAM SEN 23 EQG 24 CHA ETH SSU 26 MOZ ZIM 29 MDG
Synonyms: hidden
= Desmodium setigerum (E.Mey.) Benth. ex Harv.; Fl. Cap. [Harvey & auct. succ.] 2: 229 (1862)
= Nicolsonia setigera E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 124 (1836)
species :
Grona stolzii (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: D.R.Congo [Zaire]; Burundi; Tanzania; Zambia; Malawi; Zimbabwe
23 BUR ZAI 25 TAN 26 MLW ZAM ZIM
Synonyms: hidden
= Desmodium stolzii Schindl.; Repert. Spec. Nov. Regni Veg. 21: 9 (1925)
species :
Grona strigillosa (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(5): 304 (2018)
Distribution: China (S-Guangxi); Myanmar; Laos; Cambodia; Vietnam; Australia [I]
36 CHI 41 CBD LAO MYA VIE 50 aus
Synonyms: hidden
= Desmodium pendenticarpum C.Z.Gao & Q.R.Lai; Bull. Bot. Res., Harbin 9(4): 41 (1989) (with two types cited) ex H. Ohashi, J. Jap. Bot. 70(2): 115 (1995)
= Desmodium strigillosum Schindl.; Bot. Jahrb. Syst. 54: 57 (1916)
= Desmodium strigillosum var. pendenticarpum (C.Z.Gao & Q.R.Lai) P.H.Huang; Bull. Bot. Res., Harbin 18(1): 6 (1998)
species :
Grona styracifolia (Osbeck) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 116 (2018)
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Hubei, S-Yunnan); India; Sri Lanka; Bangladesh; Myanmar (Taninthayi); Laos; Cambodia; Vietnam; peninsular Malaysia; Sumatra; Sulawesi; Java; Lesser Sunda Isl.; Moluccas; Palau [I]; Fed. States of Micronesia [I] (Yap [I])
36 CHI 40 BAN IND SRL 41 CBD LAO MYA VIE 42 JAW LSI MLY MOL SUL SUM 62 mic plu
Synonyms: hidden
= Desmodium celebicum Schindl.; Repert. Spec. Nov. Regni Veg. 21: 6 (1925)
= Desmodium rotundifolium Wall.; Numer. List [Wallich] no. 5696 (1831)
= Desmodium strigillosum subsp. celebicum (Schindl.) H.Ohashi; J. Jap. Bot. 79(2): 130 (2004)
= Desmodium styracifolium (Osbeck) Merr.; Amer. J. Bot. 3: 580 (1916)
= Desmodium styracifolium (Osbeck) Schindl.; Repert. Spec. Nov. Regni Veg. 21: 5 (1925)
= Hedysarum conicum Poir.; Encycl. [J.Lamarck et al.] 6: 419 (1805)
= Hedysarum decumbens Dennst.; Schlüssel Hortus Malab. 217 (1818)
= Hedysarum pilosum Roxb. ex Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 223 (1834)
= Hedysarum spinosissimum DC.; Fl. Franç. [de Candolle & Lamarck], éd. 3, 6: 583 (1815), nom. illeg.
= Hedysarum styracifolium Osbeck; Dagb. Ostind. Resa 247 (1757)
= Nicolsonia styracifolia (Osbeck) Desv.; Ann. Sci. Nat. (Paris) 9: 418 (1826)
= Uraria styracifolia (Osbeck) Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 222 (1834)
species :
Grona tiwiensis (Pedley) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 117 (2018)
Distribution: Australia (Northern Territory)
50 AUS
Synonyms: hidden
= Desmodium tiwiense Pedley; Austrobaileya 5(2): 248 (1999)
species :
Grona trichostachya (Benth.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 117 (2018)
Distribution: New Guinea (Irian Jaya, Papua New Guinea); Australia (Western Australia, Northern Territory, Queensland)
42 NWG 50 AUS
Synonyms: hidden
= Meibomia trichostachya (Benth.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
= Desmodium trichostachyum Benth.; Fl. Austral. [Bentham] 2: 234 (1864)
species :
Grona triflora (L.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 117 (2018)
Name: Threeflower ticktrefoil, Zarzabacoa de tres flores
Distribution: Gambia; Guinea-Bissau; Guinea; Sierra Leone; Liberia; Ivory Coast; Ghana; Togo; Benin; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; Principe Isl.; São Tomé; Equatorial Guinea; Central African Republic; D.R.Congo [Zaire]; Burundi; S-Chad; Sudan; South Sudan; Soqotra; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe [I]; Madagascar; Mauritius; La Réunion; Rodrigues; Seychelles; China (Fujian, Guangdong, Guangxi, Hainan, Jiangxi, Yunnan, Zhejiang); Ryukyu Isl.; Volcano Isl. (Iwojima); Taiwan; Pakistan (Rawalpindi, Hazara); Nepal; Bhutan; India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Dadra-Nagar-Haveli, Daman, Delhi, Diu, Goa, Gujarat, Haryana, Himachal Pradesh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Nagaland, Odisha, Pondicherry, Punjab, Rajasthan, Sikkim, Tamil Nadu, Tripura, Uttar Pradesh, West Bengal, Yanan, Sikkim, Kashmir); Lakshadweep Isl. [Laccadives]; Maldives; Sri Lanka; Bangladesh; Andaman Isl. (North Andamans, Middle Andamans, South Andamans, Little Andaman Isl.); Nicobar Isl. (Car Nicobar Isl., North Nicobars, Central Nicobars, Great Nicobar Isl., Little Nicobar Isl.); Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (throughout); Singapore; Sumatra; South China Sea Isl.; Borneo; Brunei; Sulawesi; Philippines; Java; Lesser Sunda Isl. (Bali, etc.); Moluccas; Cocos [Keeling] Isl. [I]; Christmas Isl. [Austr.] [I]; New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch.; Australia (Northern Territory, Queensland); Vanuatu [I]; Rotuma Isl. [I]; Tonga [I] (´Eua [I], Niuafo´ou [I]); Niue [I]; Fiji [I]; [Western] Samoa [I] (´Upolu [I], Savai´i [I], Aleipata Isl. [I]); American Samoa [I] (Tutuila [I], Ta´u [I]); N-Line Isl. [I] (Tabueran [I]); Cook Isl. [I] (Rarotonga [I], Aitutaki [I], Mangaia [I]); Marquesas Isl. [I] (Nuku Hiva [I], Ua Huka [I], Hiva Oa [I], Tahuata [I], Fatu Hiva [I]); Society Isl. [I] (Mopelia Atoll [I], Tahiti [I], Raiatea [I], Mopiha´a Atoll [I]); Tuamotu Arch. [I] (Rangiroa Atoll [I]); Palau [I] (Koror [I], Ngarakabesang [I]); Northern Marianas [I] (Asuncion [I], Agrihan [I], Pagan [I], Alamagan [I], Guguan [I], Saipan [I], Tinian [I], Rota [I]); Guam [I]; Fed. States of Micronesia [I] (Malakal [I], Yap [I], Fais [I], Moen [I], Pohnpei [I]); Hawaii [I] (Kauai [I], Oahu [I], Molokai [I], Lanai [I], Maui [I], Kahoolawe Isl. [I], Hawaii Isl. [I]); USA [I] (Florida [I], Louisiana [I]); Mexico [I] (Chiapas [I], Colima [I], Guerrero [I], Jalisco [I], Michoacán [I], Nayarit [I], Oaxaca [I], Puebla [I], San Luis Potosi [I], Sinaloa [I], Tabasco [I], Tamaulipas [I], Veracruz [I], Zacatecas [I]); Guatemala [I]; Belize [I]; Honduras [I]; Nicaragua [I]; Costa Rica [I]; Panama [I]; Cuba [I]; Cayman Isl. [I]; Jamaica [I]; Hispaniola [I]; Puerto Rico [I]; Virgin Isl. [I] (Guana [I], St. Croix [I], St. John [I], St. Thomas [I], Tortola [I], Virgin Gorda [I]); St. Martin/St. Maarten [I]; St. Barthélémy [I]; Saba [I]; St. Eustatius [I]; St. Kitts [I]; Antigua [I]; Guadeloupe [I]; Dominica [I]; Martinique [I]; St. Lucia [I]; Grenadines [I]; St. Vincent [I]; Grenada [I]; Barbados [I]; Tobago [I]; Trinidad [I]; Curacao [I]; Colombia [I] (Antioquia [I], Cauca [I], Chocó [I], Meta [I], Nariño [I], Tolima [I], Valle [I]); Venezuela [I] (Amazonas [I], Anzoategui [I], Apure [I], Barinas [I], Bolívar [I], Carabobo [I], Cojedes [I], Miranda [I], Monagas [I], Portuguesa [I], Sucre [I]); Guyana [I]; Surinam [I]; French Guiana [I]; Bolivia [I] (Beni [I], Cochabamba [I], La Paz [I], Santa Cruz [I]); N-Brazil [I] (Roraima [I], Amapa [I], Amazonas [I]); NE-Brazil [I] (Maranhao [I], Rio Grande do Norte [I], Paraíba [I], Pernambuco [I], Bahia [I]); WC-Brazil [I] (Mato Grosso [I], Mato Grosso do Sul [I]); SE-Brazil [I] (Minas Gerais [I], Espirito Santo [I], São Paulo [I], Rio de Janeiro [I])
22 BEN GAM GHA GNB GUI IVO LBR NGA SIE TOG 23 BIO BUR CAF CMN EQG PRI SAO ZAI 24 CHA SOC SSU SUD 25 KEN TAN UGA 26 ANG MLW ZAM zim 29 MAU MDG REU ROD SEY 36 CHI 38 KZN NNS TAI 40 BAN BHU IND LDV MDV NEP PAK SRL 41 AND CBD LAO MYA NCB THA VIE 42 BOR BRN cki JAW LSI MLY MOL NWG PHI SCS SNG SUL SUM xms 43 BIS 50 AUS 60 asa fij nue rot sam ton van 61 coo lin mrq sci tua 62 gum mic mrn plu 63 haw 78 usa 79 mex 80 blz cos gua hon nic pan 81 aba bbd cay cub cur dom gdl gre his jam mar pue sab sba ste stk stl stm stv trt vir 82 clm frg guy sur ven 83 bol 84 bra
Synonyms: hidden
= Aeschynomene triflora (L.) Poir.; Encycl. [J.Lamarck et al.] 4(2): 451 (1798)
= Desmodium albiflorum Cordem.; Fl. Réunion [Cordemoy] 403 (1895)
= Desmodium bullamense G.Don; Gen. Hist. 2: 294 (1832)
= Desmodium granulatum (Schumach. & Thonn.) Walp.; Repert. Bot. Syst. [Walpers] 1: 737 (1842)
= Desmodium stipulaceum (Burm.f.) Hassk.; Cat. Hort. Bot. Bogor. [Hasskarl] 274 (1844), nom. illeg.
= Desmodium triflorum (L.) DC.; Prodr. [A.P. de Candolle] 2: 334 (1825)
= Desmodium triflorum var. adpressum Ohwi; J. Jap. Bot. 26: 234 (1951)
= Desmodium triflorum var. minimus Stehlé; Bull. Mus. Natl. Hist. Nat., Sér. 2, 18: 104 (1946)
= Desmodium triflorum var. minus Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 229 (1834)
= Desmodium triflorum var. pygmaeum Hoehne; Relat. Commiss. Linhas Telegr. Estratég. Matto Grosso Amazonas 5, Bot. pt. 8: 73 (1919)
= Desmodium triflorum var. villosum Wight & Arn.; Prodr. Fl. Ind. Orient. [Wight & Arnott] 1: 229 (1834)
= Hedysarum biflorum Willemet; Ann. Bot. [Usteri] 18: 49 (1796)
= Hedysarum granulatum Schumach. & Thonn.; Beskr. Guin. Pl. [Schumacher] 362 (1827)
= Hedysarum stipulaceum Burm.f.; Fl. Ind. [N.L.Burman] 168 (1768)
= Hedysarum triflorum L.; Sp. Pl. [Linnaeus] 2: 749-750 (1753)
= Hippocrepis humilis Blanco; Fl. Filip. [Blanco] 585 (1837)
= Meibomia triflora (L.) Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia triflora f. coerulescens Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia triflora f. flavescens Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia triflora f. purpurea Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia triflora f. violacea Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia triflora f. virescens Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia triflora var. glabrescens Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Meibomia triflora var. pilosa Kuntze; Revis. Gen. Pl. 1: 197 (1891)
= Nicolsonia reptans Meisn.; Linnaea 21: 260 (1848)
= Nicolsonia triflora (L.) Griseb.; Abh. Königl. Ges. Wiss. Göttingen 7: 202 (1857)
= Onobrychis heterophylla Schrank ex Steud.; Nomencl. Bot. [Steudel], ed. 2, 2: 213 (1841), nomen
= Pleurolobus triflorus (L.) J.St.-Hil.; Nouv. Bull. Sci. Soc. Philom. Paris 3: 192 (1812)
= Sagotia triflora (L.) Duchass. & Walp.; Linnaea 23: 738 (1850)
species :
Grona varians (Labill.) H.Ohashi & K.H.Ohashi; J. Jap. Bot. 96(1): 11 (2021)
Distribution: Australia (Queensland, New South Wales, Victoria, Tasmania); New Caledonia
50 AUS 60 NWC
Synonyms: hidden
= Desmodium dietrichiae Domin; Biblioth. Bot. 22(89): 213 (1926)
= Desmodium spartioides DC.; Prodr. [A.P. de Candolle] 2: 337 (1825)
= Desmodium varians (Labill.) G.Don; Gen. Hist. 2: 298 (1832)
= Desmodium varians var. angustifolium Benth.; Fl. Austral. [Bentham] 2: 233 (1864)
= Hedysarum varians Labill.; Sert. Austro-Caledon. 71, t. 71 (1825)
= Meibomia varians (Labill.) Kuntze; Revis. Gen. Pl. 2: 198 (1891)
species :
Grona vidalii (H.Ohashi) H.Ohashi & K.Ohashi; J. Jap. Bot. 95(1): 3 (2020)
Distribution: Laos; Vietnam
41 LAO VIE
Synonyms: hidden
= Desmodium vidalii H.Ohashi; J. Jap. Bot. 67(6): 320 (1992)
species :
Grona wittei (B.G.Schub.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 117 (2018)
Distribution: D.R.Congo [Zaire]
23 ZAI
Synonyms: hidden
= Desmodium wittei B.G.Schub.; Bull. Jard. Bot. État Bruxelles 22: 292 (1952)
species :
Grona wynaadensis (Bedd. ex Gamble) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 117 (2018)
Distribution: S-India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Desmodium ferrugineum subsp. wynaadense (Bedd. ex Gamble) H.Ohashi; Ginkgoana 1: 206 (1973)
= Desmodium wynaadense Bedd. ex Gamble; Fl. Madras 343, 346 (1918)
species :
Grona zenkeri (Schindl.) H.Ohashi & K.Ohashi; J. Jap. Bot. 93(2): 119 (2018)
Distribution: Cameroon
23 CMN
Synonyms: hidden
= Desmodium zenkeri Schindl.; Repert. Spec. Nov. Regni Veg. 21: 8 (1925)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.04 (April 17th, 2025)

Covering 356,375 accepted species,
8,782 hybrids, 48,090 accepted infraspecific taxa and 1,002,897 synonyms,
in total 1,416,144* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

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With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

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