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045.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
3-34 genera (depending on genus concept), 1100-1200 spp. The genus concepts were wildly fluctuating and disputed, from the inclusion of everything into one monster genus Thelypteris to the split of over 30 genera. We use the new genus concept by Fawcett & Smith (2021), which is based on molecular results and adopts a compromise approachl
Species listed in database: 1181; hybrids: 39
045.2007 Genus Hoiokula S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 48 (2021)
Distribution: Hawaii
Species listed in database: 2
species :
Hoiokula pendens (D.D.Palmer) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 49 (2021)
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Cyclosorus pendens (D.D.Palmer) N.Snow; Bishop Mus. Occas. Pap. (2011)
= Pneumatopteris pendens D.D.Palmer; Amer. Fern J. 95(2): 81 (2005)
= Thelypteris pendens (D.D.Palmer) Christenh.; Global Fl. 4: 35 (2018)
species :
Hoiokula sandwicensis (Brack.) S.E.Fawc. & A.R.Sm.; Sida, Bot. Misc. 59: 49 (2021)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Cyclosorus sandwicensis (Brack.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 143 (1947)
= Dryopteris stegnogrammoides (Baker) C.Chr.; Index Filic. 294 (1905)
= Goniopteris sandwicensis (Brack.) J.Sm.; Hist. Fil. 192 (1875)
= Goniopteris stegnogrammoides (Baker) J.Sm.; Ferns Brit. Foreign, ed. 2: 303 (1877), nom. superfl.
= Nephrodium polycarpum Diels; Nat. Pflanzenfam. [Engler & Prantl] I, 4: 179 (1899)
= Phegopteris microdendron D.C.Eaton; Proc. Amer. Acad. Arts 7: 218 (1868)
= Phegopteris polycarpa Hillebr.; Fl. Hawaiian Isl. [Hillebrand] 560 (1888)
= Phegopteris stegnogrammoides (Baker) Luerss.; Flora 435 (1876)
= Pneumatopteris sandwicensis (Brack.) Holttum; Blumea 21: 303 (1973)
= Polypodium polycarpon Hook. & Arn.; Bot. Beechey Voy. 104 (1832) (non Sw. (1801))
= Polypodium sandwicense (Brack.) Hook.; Sp. Fil. [W.J.Hooker] 5: 5 (1863) (non 4: 267)
= Polypodium stegnogrammoides Baker; Syn. Fil. [W.J.Hooker & Baker] 317 (1867)
= Stegnogramma sandwicensis Brack.; U.S. Expl. Exped. 16 (Filic.): 26 (1854) (non Thelpyteris sandwicensis (Hook. & Arn.) Fosberg)
= Thelypteris hawaiiensis C.F.Reed; Phytologia 17: 282 (1968), nom. nov.
= Thelypteris stegnogrammoides (Baker) Fosberg; Occas. Pap. Bernice P. Bishop Mus. 23: 30 (1962)
= Thelypteris treebeardii Christenh.; Global Fl. 4: 39 (2018), nom. nov. for Stegnogramma sandwicensis
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.05 (May 20th, 2026)

Covering 359,646 accepted species,
8,890 hybrids, 47,679 accepted infraspecific taxa and 1,012,517 synonyms,
in total 1,428,732* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life