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V. 12.5
14.10.2021

002.0000 Family Isoetaceae Rchb.; Bot. Damen: 309 (1828)
1 genus, ca. 190 spp.
Species listed in database: 191; hybrids: 24
= Bothriaceae Dulac, Fl. Dépt. Hautes-Pyrenees 38 (1867)
002.0001 Genus Isoetes L.; Sp. Pl. 2: 1100 (1753)
Distribution: Worldwide. Many undescribed species remain. Latest checklist: Troia et al. (2016)
Species listed in database: 191; hybrids: 24
= Calamaria Boehm., Def. Gen. Pl. ed. Boehmer 500 (1760)
= Cephaloceraton Gennari, Comment. Soc. Critt. Ital. 1: 111 (1862)
= Isoetella Gennari, Comment. Soc. Critt. Ital. 1: 114 (1862)
= Sivetes Raf., Med. Repos. 11. 5: 358 (1808)
= Stylites Amstutz, Ann. Missouri Bot. Gard. 44: 121 (1957)
= Subularia Ray, Syn. Meth. Stirp. Brit., 283 (1696) (Pre-Linnaean)
species :
Isoetes abyssinica Chiov.; Atti Soc. Nat. e Mat., Modena 64: 45-47, illus (1933)
Distribution: Ghana; Eritrea; Ethiopia; Burundi; Kenya; Tanzania
22 GHA 23 BUR 24 ERI ETH 25 KEN TAN
species :
Isoetes aemulans J.P.Roux; Bothalia 43(2): 218 (2013)
Distribution: South Africa (Free State, Mpulamanga, KwaZulu-Natal)
27 SAF
species :
Isoetes aequinoctialis Welw. ex A.Braun; Sitz. Ges. Naturf. Freunde Berl. 7 (1867), et Kuhn, Fil. Afr. 195 (1868)
Distribution: Mali; Ghana; Niger; Central African Republic; Tanzania; Angola; Zambia; Zimbabwe; Namibia; South Africa (N-Cape Prov., Mpulamanga); Eswatini [Swaziland]
22 GHA MLI NGR 23 CAF 25 TAN 26 ANG ZAM ZIM 27 NAM SAF SWZ
Synonyms: hidden
= Calamaria aequinoctialis (Welw. ex A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes erongoensis Wanntorp, Svensk Bot. Tidsskr. 64 (2): 150-152 (1970)
species :
Isoetes afloramientorum D.F.Brunt. & J.B.S.Pereira; Phytotaxa 451(3): 219 (2020)
Distribution: Bolivia (Santa Cruz)
83 BOL
species :
Isoetes alcalophila Halloy; Lilloa 35(2): 68 (1980)
Distribution: Argentina (Catamarca, Tucuman)
85 ARG
species :
Isoetes alpina Kirk; Tr. New Zeal. Inst. 7: 377, t. 25 (1875)
Distribution: New Zealand (South Isl., North Isl.)
51 NZL
Synonyms: hidden
= Calamaria alpina (Kirk) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes kirkii var. alpina (Kirk) C.R.Marsden & Chinnock
= Isoetes multiangularis Colenso, Tr. New Zeal. Inst. 22: 449 (1890)
species :
Isoetes alstonii C.F.Reed & Verdc.; Kirkia 5: 19 (1965)
Distribution: N-Namibia; S-Zambia; Tanzania; Mozambique; Zimbabwe; Madagascar
25 TAN 26 MOZ ZAM ZIM 27 NAM 29 MDG
species :
Isoetes amazonica A.Braun; Baker, J. Bot. 18: 109 (1880)
Distribution: N-Brazil (Pará)
84 BRA
Synonyms: hidden
= Calamaria amazonica (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes anatolica Prada & Rolleri; Bot. J. Linn. Soc. 147(2): 215 (2005)
Distribution: Turkey (Anatolia)
34 TUR
species :
Isoetes andicola (Amstutz) L.D.Gómez; Brenesia 18: 4 (1980)
Distribution: Peru; Bolivia (La Paz)
83 BOL PER
Conservation Status: IUCN R
Synonyms: hidden
= Stylites andicola Amstutz, Ann. Missouri Bot. Gard. 44: 121 (1957)
= Isoetes andicola (Amstutz) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 84(2): 177 (1981), comb. superfl.
= Isoetes andicola var. gemmifera (Rauh) L.D.Gómez, Brenesia 18: 4 (1980)
= Stylites gemmifera Rauh, Sitzungsber. Heidelberg Akad. Wiss. Math. Naturwiss. Kl. 1959: 11 (1959)
species :
Isoetes andina Spruce ex Hook.; Brit. Ferns ad t. 55 (1861)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, Magdalena, Meta, Nariño, Putumayo, Risaralda); Ecuador; Venezuela (Merida, Tachira); Peru
82 CLM VEN 83 ECU PER
Synonyms: hidden
= Calamaria triquetra (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1893)
= Isoetes andina Spruce, Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 325 (1883)
= Isoetes lechleri var. triquetra (A.Braun) L.D.Gómez, Brenesia 18: 5 (1980)
= Isoetes triquetra A.Braun, Verh. Bot. Ver. Brandenb. 4: 332 (1862)
species :
Isoetes appalachiana D.F.Brunt. & D.M.Britton; Rhodora 99: 118-133 (1997), nom. nov.
Name: Appalachian quillwort
Distribution: USA (Florida, Georgia, North Carolina, Pennsylvania, South Carolina, Virginia)
75 USA 78 USA
Synonyms: hidden
= Isoetes engelmannii var. georgiana Engelm., Trans. Acad. Sci. St. Louis 4: 384 (1882)
= Isoetes engelmannii f. georgiana (Engelm.) Clute, Fern-Allies 258 (1905)
= Isoetes engelmannii f. fontana (Eaton) Clute, Fern-Allies 258 (1905)
= Isoetes engelmannii var. fontana Eaton, Fern Bull. 13: 52 (1905)
species :
Isoetes araucaniana Macluf & Hickey; Amer. Fern J. 97(4): 220 (2008)
Distribution: Chile (Araucania)
85 CLE
Synonyms: hidden
= Isoetes favulata Hickey, ined.
species :
Isoetes asiatica (Makino) Makino; Bot. Mag. (Tokyo) 28: 184 (1914)
Distribution: Russian Far East (S-Kuril Isl., Sakhalin, Kamchatka); Japan; ?China
31 RFE 38 JAP
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes setacea ssp. asiatica (Makino) Holub, Acta Horti Bot. Prag (1964)
= Isoetes echinospora ssp. asiatica (Makino) Á.Löve, Amer. Fern J. 52: 121 (1962)
= Isoetes echinospora var. asiatica Makino, Bot. Mag. (Tokyo) 18: 129 (1904) ex Iversen, Dansk Bot. Ark. 5 (23): 2 (1928)
= Isoetes muricata ssp. asiatica (Makino) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982)
species :
Isoetes attenuata C.R.Marsden & Chinnock; Fl. Australia 48: 704 (1998)
Distribution: Australia (SE-South Australia)
50 AUS
species :
Isoetes australis S.Williams; Proc. Roy. Soc. Edinb. 62B(l). 1-8 (1943)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes azorica Durand; Milde, Fil. Eur. 278 (1867)
Name: Azoren-Brachsenkraut
Distribution: Azores (Terceira, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.)
21 AZO
Conservation Status: IUCN E
Synonyms: hidden
= Calamaria azorica (Durand) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes baculata Hickey & H.P.Fuchs; Syst. Bot. 11(2): 310 (1986)
Distribution: Colombia (Vaupés)
82 CLM
species :
Isoetes biafrana Alston; Bol. Soc. Broter. Ser. 2A, 30: 15 (1956)
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; Benin
22 BEN 23 BIO CMN
species :
Isoetes bischlerae H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(1): 113 (1982)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Nariño)
82 CLM
Remarks: provisionally accepted name
species :
Isoetes bolanderi Engelm.; Amer. Naturalist 8: 214 (1874)
Name: Bolander´s Quillwort
Distribution: USA (Arizona, California, Colorado, Florida, Idaho, Montana, New Mexico, Nevada, Oregon, Utah, Washington State, Wyoming); Canada (Alberta)
71 CDN 73 USA 76 USA 77 USA 78 USA
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria bolanderi (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Calamaria pygmaea (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes bolanderi f. pygmaea (Engelm.) Clute, Fern Bull. 16: 54 (1908)
= Isoetes bolanderi var. parryi Engelm., Amer. Naturalist 8: 214 (1874)
= Isoetes bolanderi var. pygmaea (Engelm.) Clute, Fern-Allies 228, 258 (1905)
= Isoetes bolanderi var. sonnei L.F.Hend., Bull. Torrey Bot. Cl. 27(6): 358 (1900)
= Isoetes californica Engelm., Gray´s Manual, ed. 5: 677 (1867)
= Isoetes parryi Engelm., Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 338 (1883)
= Isoetes pygmaea Engelm., Amer. Naturalist 8: 214 (1874)
species :
Isoetes boliviensis U.Weber; Hedwigia 63: 247, fig. 14-15 (1922)
Distribution: Bolivia (Cochabamba, La Paz); Peru
83 BOL PER
species :
Isoetes boomii Luebke; Amer. Fern J. 82: 23 (1992)
Name: Boom´s quillwort
Distribution: USA (C-Georgia: Laurens Co., Worth Co.)
75 USA 78 USA
Conservation Status: IUCN E
species :
Isoetes boryana Durand; Bull. Soc. Bot. France 8: 164 (1861)
Distribution: SW-France; +Spain
12 FRA SPA
Conservation Status: IUCN V
Synonyms: hidden
= Calamaria boryana (Durand) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes boryana ssp. brevis Gand., Fl. Eur. 27: 195 (1891)
= Isoetes boryana ssp. condensata Gand., Fl. Eur. 27: 195 (1891)
= Isoetes boryana ssp. frigida Gand., Fl. Eur. 27: 195 (1891)
= Isoetes boryana ssp. gredensis Gand., Fl. Eur. 27: 195 (1891)
= Isoetes boryana ssp. legitima Gand., Fl. Eur. 27: 195 (1891)
= Isoetes boryana ssp. praetervisa Gand., Fl. Eur. 27: 195 (1891)
= Isoetes variabilis ssp. boryana (Durieu) P.Fourn., Quatre Fl. France 5 (1934)
= Isoetes velata var. boryana (Durieu) M.I.Romero & Real, Bot. J. Linn. Soc. 148(4): 464 (2005)
species :
Isoetes boyacensis H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(1): 107 (1982)
Distribution: Colombia (Boyacá, Cundinamarca, Meta, Nariño, Santander)
82 CLM
Remarks: provisionally accepted name
species :
Isoetes bradei Herter; Rev. Südam. Bot. 8: 19 (1949)
Distribution: SE-Brazil (São Paulo)
84 BRA
Conservation Status: considered extinct
species :
Isoetes brasiliensis H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 27 (1986)
Distribution: Brazil (Santa Catarina, Rio Grande do Sul)
84 BRA
Synonyms: hidden
= Isoetes reitzii H.P.Fuchs, (1979), nom. nud. pro syn.
species :
Isoetes brevicula E.R.L.Johnson; J. Roy. Soc. W. Australia 67(1): 34 (1984)]
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes butleri Engelm.; Bot. Gaz. 3: 1 (1878)
Name: Limestone quillwort
Distribution: USA (Alabama, Arkansas, Georgia, Illinois, Kansas, Kentucky, Missouri, Oklahoma, Tennessee, Texas)
74 USA 75 USA 77 USA 78 USA
Synonyms: hidden
= Calamaria butleri (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes butleri f. immaculata (Engelm.) Clute, Fern-Allies 258 (1905)
= Isoetes butleri var. immaculata Engelm., Trans. Acad. Sci. St. Louis 4: 388 (1882)
species :
Isoetes cangae J.B.S.Pereira, Salino & Stützel; Phytotaxa 272(2): 145 (2016)
Distribution: Brazil (Pará)
84 BRA
species :
Isoetes caparaoensis J.B.S.Pereira; Bot. J. Linn. Soc. 189(1): 15 (2018)
Distribution: Brazil (Minas Gerais)
84 BRA
species :
Isoetes capensis Duthie; Trans. R. Soc. S. Afr. 17: 330 (1929)
Distribution: South Africa (W-Cape Prov.)
27 SAF
species :
Isoetes caroli E.R.L.Johnson; J. Roy. Soc. W. Austratia 67(1): 34 (1984)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes caroliniana (A.A.Eaton) Luebke; Amer. Fern J. 82: 26 (1992)
Distribution: SE-USA
78 USA
Remarks: bona species fide Troia et al. 2016, not a synonym of I. valida
Synonyms: hidden
= Isoetes engelmannii var. caroliniana A.A.Eaton, Fern Bull. 8: 60 (1900)
species :
Isoetes chubutiana Hickey, Macluf & W.C.Taylor; Amer. Fern J. 93(3): 130 (2003)
Distribution: Argentina (Chubut, Neuquen, Rio Negro); Chile (Los Lagos, Aisen)
85 ARG CLE
Synonyms: hidden
= Isoetes savatieri ssp. valdiviensis H.P.Fuchs, (1992), nom. nud.
= Isoetes valdiviensis H.P.Fuchs, (1960), nom. nud.
= Isoetes meyeri H.P.Fuchs, (1982), nom. nud.
species :
Isoetes coreana Y.H.Chung & H.K.Choi; Korean J. Pl. Taxon 16(1): 2 (1986)
Distribution: South Korea
38 KOR
Remarks: bona sp. fide Troia et al. 2016
Synonyms: hidden
= Isoetes sinensis var. coreana (Y.H.Chung & H.K.Choi) M.Takamiya, M.Watanaba & K.Ono, Acta Phytotax. Geobot. 48(2) (1997 publ. 1998)
species :
Isoetes coromandelina L.fil.; Suppl. Pl. Syst. Veg. ed. II: 447 (1781)
subspecies :
subsp. coromandelina
Distribution: India (Andhra Pradesh, Bihar, Chhattisgarh, Goa, Gujarat, Haryana, Jharkand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Odisha, Rajasthan, Tamil Nadu, Uttar Pradesh, West Bengal); Bangladesh; Thailand; Sri Lanka; Vietnam; Nepal; Myanmar
40 BAN IND NEP SRL 41 MYA THA VIE
Synonyms: hidden
= Calamaria coromandelina (L.fil.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes brachyglossa A.Braun, Ver. Bot. Ver. Brandenb. 4: 328 (1862), nom. nov. for I. capsularis Roxb. ex Griff. (fide Fraser-Jenkins 1997, 2017)
= Isoetes capsularis Roxb. ex Griff., Posth. Papers, Cryptog. Pl. 572-575, t. 116-118 (1849), non Roxb. 1844
= Isoetes coromandelina ssp. brachyglossa (A.Braun) Panigrahi, Biol. Mem. 6 (2): 131 (1981) (fide Fraser-Jenkins 1997)
= Isoetes coromandelina var. raipurensis Unni, J. Bombay Nat. Hist. Soc. 64: 590-592 (1967) (fide Fraser-Jenkins 1997)
= Isoetes debii P.K.Shukla, G.K.Srivast. & S.K.Shukla, Frontiers Bot. 4-6: 69-72 (2002)
= Isoetes fuchsii Goswami & U.S.Sharma, Bionature (Bhopal) 21: 11-18 (2001)
= Isoetes indica D.D.Pant & G.K.Srivast., Proc. Nat. Inst. Sci. India 28 B (3) 246, t. XIV9, f. 16-259, t. XV, f. 30-35, t. XIV, f. 30-40 text-fig. 4 A-D, 5 A-H, 7 A-M, 12 (1962) (fide Fraser-Jenkins 1997)
= Isoetes indica var. harotiensis Bhardwaja & Gena, Aspects Plant Sci. 14: 495-499 (1992), nom. nud.
= Isoetes mirzapurensis Panigrahi & R.D.Dixit, Proc. Indian Sci. Congr. pt. III, Abstract, Botany 243 (1965) (fide Fraser-Jenkins 1997)
= Isoetes narsinghgharensis Goswami & Mazumdar, nom. nov. for I. pantii var. hybrida Goswami
= Isoetes pantii Goswami & Arya, J. Indian Bot. Soc. 49: 30 (1970) (fide Fraser-Jenkins 1997)
= Isoetes pantii var. hybrida Goswami, [fide Fraser-Jenkins et al. 2020]
= Isoetes semilocularis J.E.Sm., Rees, Cycl. 19: 150 (1819)
= Isoetes tuberculata Gena & Bhardwaja, J. Bombay Nat. Hist. Soc. 81(1): 167 (1984) (fide Fraser-Jenkins 1997)
= Isoetes coromandeliana ssp. thanensis S.K.Shukla, Sarv.K.Singh, P.K.Shukla, N.K.Dubey, H.Khanam & G.K.Srivast., Taiwania 62(2): 122 (2017)
= Isoetes unilocularis J.E.Sm., Rees, Cycl. 19(1): 150 (1811)
subspecies :
subsp. macrotuberculata C.R.Marsden; Contr. Herb. Austral. 24: 2 (1976)
Distribution: Australia (Western Australia, Northern Territory, Queensland)
50 AUS
species :
Isoetes creussensis Lazare & S.Riba; J. Bot. Soc. Bot. France 50: 21 (2010)
Distribution: Andorra
12 ADO
species :
Isoetes cristata C.R.Marsden & Chinnock; Fl. Australia 48: 704 (1998)
Distribution: Australia (Northern Territory)
50 AUS
Conservation Status: Known from type locality only
species :
Isoetes cubana Engelm. ex Baker; J. Bot. 18: 110 (1880)
Distribution: Cuba; Belize; ?Mexico (?Yucatan)
80 BLZ 81 CUB
Synonyms: hidden
= Calamaria cubana (Engelm. ex Baker) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes delilei (Bory) Rothm.; Fedde, Repert. Spec. Nov. 54: 72 (1944)
Distribution: SE-France; Portugal; W-Spain; Morocco; ?Corsica; ?Sardinia
12 FRA POR SPA 20 MOR
Synonyms: hidden
= Isoetes setacea Bosc ex Delile, Mem. Mus. Paris 14: (110) 117, t. 6-7 (1827) (non Lam.)
= Isoetes setacea var. delilei Bory, Compt. Rend. Acad. Paris 18: 1165 (1844)
= Isoetes palustris var. setacea F.Muell., Milde, Fil. Eur. & Atl. 9: 280 (1867)
species :
Isoetes dispora Hickey; Fieldiana, Bot. 34: 93 (1994)
Distribution: Peru
83 PER
Conservation Status: Known from type only
species :
Isoetes dixitii Shende; Journ. Univ. Bombay 14: 50 (1945)
Distribution: India (Chhattisgarh, Karnataka, Madhya Pradesh, Maharashtra)
40 IND
Remarks: bona sp. fide Fraser-Jenkins et al. 2017)
Synonyms: hidden
= Isoetes bilaspurensis Panigrahi, Biol. Mem. 6 (2): 135, f. 1-10 (1981)
species :
Isoetes drummondii A.Braun; Monatsber. K. Akad. Berl. 593 (1863)
subspecies :
subsp. drummondii
Distribution: Australia (SW-Western Australia, SE-South Australia, SE-Queensland, Victoria, NE-Tasmania)
50 AUS
Synonyms: hidden
= Calamaria drummondii (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
subspecies :
subsp. anomala C.R.Marsden & Chinnock; Muelleria 8(1): 58 (1993)
Distribution: Australia (SE-South Australia, SW-Western Australia, Victoria, S-New South Wales)
50 AUS
species :
Isoetes dubsii J.B.S.Pereira; PhytoKeys 131: 59 (2019)
Distribution: Brazil (Mato Grosso do Sul)
84 BRA
species :
Isoetes duriei Bory; Compt. Rend. Acad. Paris 18: 1166 (1844) (durieai auct., durieui auct.)
Name: Durieus Brachsenkraut
Distribution: Portugal; Spain; Baleares; France; Corsica; Sardinia; Malta; Sicily; Italy; Greece (only Aegaean region and Kiklades); Crete; Tunisia; Algeria; ?Morocco; Turkey (Anatolia); East Aegaean Isl. (Lesvos, Rhodos, Karpathos)
12 BAL COR FRA POR SAR SPA 13 GRC ITA KRI MLT SIC 20 ALG TUN 34 EAI TUR
Synonyms: hidden
= Calamaria durieui (Bory) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetella duriei (Bory) Gennari, Comment. Soc. Critt. Ital. 1(3): 115 (1862)
= Isoetes duriaei ssp. balearica Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. felix Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. insularis Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. kabylica Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. leptophylla Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. ligustica De Not., Kunze, Ind. Fil. Cult., 51 (1850)
= Isoetes duriaei ssp. litorea Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. lusitanica Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. maritima Gand., Fl. Eur. 27: 196 (1891)
= Isoetes duriaei ssp. sardoa Gand., Fl. Eur. 27: 196 (1891)
= Isoetes ligustica De Not. ex A.Braun, Monatsber. K. Akad. Berl., Sitzung 7 Dec (1863)
= Isoetes tridentata Durand, ex Kuhn, Fil. Afr. 195 (1867)
species :
Isoetes echinospora Durand; Bull. Soc. Bot. France 8: 164 (1861)
Name: Stachelsporiges Brachsenkraut, Spiny-spore quillwort
Distribution: Austria; England; Scotland; Ireland; Czech Republic; Slovakia; Denmark; Finland; S-France; Andorra; Spain; Germany (Baden-Württemberg, Schleswig-Holstein+); Switzerland; Netherlands; Iceland; Italy; Norway; Poland; ?Romania; Sweden; Bulgaria; Morocco; European Russia (N-European Russia, C-European Russia); Estonia; Latvia; Siberia (W-Siberia, C-Siberia); Alaska; USA (California, Colorado, Connecticut, Delaware, Idaho, Massachusetts, Maine, Michigan, Minnesota, Montana, New Hampshire, New Jersey, New York, Ohio, Oregon, Pennsylvania, Rhode Island, Utah, Vermont, Washington State, Wisconsin, Wyoming); Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); Greenland; St. Pierre et Miquelon
10 DEN FIN GRB ICE IRE NOR SWE 11 AUT CZE GER NET POL SLO SWI 12 ADO FRA SPA 13 BUL ITA 14 EST LAT RUS 20 MOR 30 SIB 70 ASK CDN GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 76 USA 78 USA
Synonyms: hidden
= Calamaria echinospora (Durand) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes ambigua A.Braun, Engelm., Trans. Acad. Sci. St. Louis 4: 380 (1882)
= Isoetes boottii A.Braun, Engelm., Gray´s Manual, ed. 5, 676 (1867)
= Isoetes braunii Durand, Bull. Soc. Bot. France 11: 101 (1864)
= Isoetes braunii f. astoma (Iversen) M.Broun, Index N. Amer. Ferns 99 (1938)
= Isoetes braunii f. oligostoma (Iversen) M.Broun, Index N. Amer. Ferns 99 (1938)
= Isoetes braunii f. polystoma (Iversen) M.Broun, Index N. Amer. Ferns 99 (1938)
= Isoetes echinospora f. astoma Iversen, Bot. Tidsskr. 40: 129 (1928)
= Isoetes echinospora f. boottii (Engelm.) Clute, Fern Allies 258 (1905)
= Isoetes echinospora f. oligostoma Iversen, Bot. Tidsskr. 40: 129 (1928)
= Isoetes echinospora f. polystoma Iversen, Bot. Tidsskr. 40: 128 (1928)
= Isoetes echinospora ssp. admixta Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. belgica Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. borealis Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. centralis Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. discedens Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. echinospora Durand, Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. fennica Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. gothica Gand., 27: 194 (1891)
= Isoetes echinospora ssp. montana Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. muricata (Durieu) Á. & D.Löve, Bot. Notis. 114: 49 (1961)
= Isoetes echinospora ssp. oligophylla Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. petropolitana Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. praebens Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. pulchella Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. pygmaea Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. remissa Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora ssp. vogesiaca Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subf.angustivelata Iversen, Bot. Tidsskr. 40: 128, fig. a-c (1928)
= Isoetes echinospora subf.lativelata Iversen, Bot. Tidsskr. 40: 129 (1928)
= Isoetes echinospora var. boothii Clute, Frye, Ferns Northwest 39 (pro syn.) (1934)
= Isoetes echinospora var. boottii Engelm., Manual (Gray), ed. 5: 676 (1867)
= Isoetes echinospora var. braunii (Durieu) Engelm., Gray´s Manual, ed. 5: 676 (1867)
= Isoetes echinospora var. brittoni Cockerell, Muhlenbergia 3: 9 (1907)
= Isoetes echinospora var. elatior Fliche, Mem. Acad. Stanisl. IV, 11 (1878): 182 (1879)
= Isoetes echinospora var. hesperia (C.F.Reed) Á.Löve, Amer. Fern J. 52: 121 (1962)
= Isoetes echinospora var. muricata (Durand) Engelm., Gray´s Manual, ed. 5, 676 (1867)
= Isoetes lacustris ssp. curvifolia (Casp.) Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. echinospora Hook.fil., Student´s Fl. Brit. Isl., ed. 1, 471 (1870)
= Isoetes lacustris var. crassa Léman, Dict. Sci. Nat. 24: 24 (1822)
= Isoetes lacustris var. elatior (Fliche) Mer, Bull. Soc. Bot. France 28: 73 (1881)
= Isoetes lacustris var. longifolia Motelay & Vendryès, Actes Soc. Linn.
= Isoetes lacustris var. minor (A.Braun ex Milde) Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. recurvata Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes muricata Durand, Bull. Soc. Bot. France 11: 100 (1864)
= Isoetes muricata var. braunii (Durand) C.F.Reed, Amer. Fern J. 35: 83 (1945)
= Isoetes muricata var. hesperia C.F.Reed, Amer. Fern J. 35: 84 (1945)
= Isoetes setacea ssp. monspeliensis Gand., Fl. Eur. 27: 196 (1891)
= Isoetes setacea ssp. muricata (Durieu) Holub, Acta Horti Bot. Prag. (1964)
= Isoetes setacea ssp. normalis Gand., Fl. Eur. 27: 195 (1891)
= Isoetes setacea ssp. stricta Gand., Fl. Eur. 27: 196 (1891)
= Isoetes setacea ssp. turgida Gand., Fl. Eur. 27: 196 (1891)
= Isoetes setacea ssp. valida Gand., Fl. Eur. 27: 196 (1891)
= Isoetes setacea ssp. vicina Gand., Fl. Eur. 27: 195 (1891)
= Isoetes tenella f. astoma (Iversen) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 72 (1944)
= Isoetes tenella f. iversenii Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)
= Isoetes tenella f. lativelata (Iversen) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 72 (1944)
= Isoetes tenella f. oligostoma (Iversen) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)
= Isoetes tenella f. polystoma (Iversen) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)
= Isoetes tenella var. braunii (Durand) Rothm., Fedde, Repert. Spec. Nov. 54: 73 (1944)
= Isoetes tenella var. oligostoma (Iversen) Rothm., Fedde, Repert. Spec. Nov. 54: 72 (1944)
species :
Isoetes ecuadoriensis Aspl.; Bot. Notis. 1925: 357, f. (1925)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Isoetes keeleyorum H.P.Fuchs, (1988), nom. nud.
species :
Isoetes ekmanii Weber; Hedwigia 63: 251. fig. 26-27 (1922)
Distribution: Argentina (Buenos Aires, Corrientes, Misiones)
85 ARG
species :
Isoetes elatior F.Muell. & A.Braun; Linn. 25: 722 (1852)
Distribution: Australia (Tasmania)
50 AUS
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria elatior (F.Muell.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes tasmanica F.Muell., Durand, Bull. Soc. Bot. France 11: 104 (1864)
species :
Isoetes eludens J.P.Roux, Hopper & Rhian J.Sm.; Kew Bull. 64(1): 124 (2009)
Distribution: South Africa (N-Cape: Namaqualand)
27 SAF
Conservation Status: extremely local in a single 2 meter diameter rock pool
species :
Isoetes engelmannii A.Braun; Flora 29: 178 (1846)
Name: Engelmann´s quillwort
Distribution: USA (Alabama, Arkansas, Connecticut, Delaware, Georgia, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Missouri, Mississippi, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, West Virginia); Canada (Ontario)
72 CDN 74 USA 75 USA 78 USA
Synonyms: hidden
= Calamaria engelmannii (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes engelmannii var. gracilis Engelm., Gray´s Manual, ed. 5, 677 (1876)
= Isoetes lacustris var. microspora Engelm., A. Braun, Flora 29: 179 (1846)
= Isoetes ×fairbrothersii J.D.Montgom. & W.C.Taylor, Amer. Fern J. 84(4): 120 (1994)
species :
Isoetes escondidensis Halloy; Lilloa 35(2): 67 (1980)
Distribution: Argentina (Tucuman)
85 ARG
Conservation Status: IUCN E
Synonyms: hidden
= Isoetes castillonii H.P.Fuchs, (1965)
= Isoetes halloyi Fuchs, (1981), nom. nud.
= Isoetes filipovichii H.P.Fuchs, (1965), nom. nud.
species :
Isoetes eshbaughii Hickey; Syst. Bot. 11(2): 310 (1986)
Distribution: Bolivia (Cochabamba)
83 BOL
species :
Isoetes flaccida Shuttlew.; A. Braun, Flora 29: 178 (1846)
Name: Southern quillwort
Distribution: USA (Alabama, Florida, Georgia)
78 USA
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria flaccida (Shuttlew) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes flaccida f. rigida (Engelm.) Clute, Fern-Allies 258 (1905)
= Isoetes flaccida var. rigida Engelm., Trans. Acad. Sci. St. Louis 4: 386 (1882)
= Isoetes flaccida var. alata (Small) N.Pfeiff., Ann. Missouri Bot. Gard. 9(2): 137 (1922)
= Isoetes alata Small, Ferns of Florida 133 (1918) apud N. Pfeiff., Ann. Missouri Bot. Gard. 9: 137 (1922)
variant :
var. chapmanii Engelm.; Trans. Acad. Sci. St. Louis 4: 386 (1882)
Name: Chapman´s quillwort
Distribution: USA (Florida, Georgia)
78 USA
Synonyms: hidden
= Isoetes chapmanii (Engelm.) Small, Ferns of Florida 222 (1931 publ. 1932)
= Isoetes flaccida f. chapmanii (Engelm.) Clute, Fern-Allies 258 (1905)
species :
Isoetes fluitans M.I.Romero; Bot. J. Linn. Soc. 146(2): 233 (2004)
Distribution: Spain
12 SPA
species :
Isoetes fuchsii Bhu, H.K.Goswami, U.S.Sharma & A.K.Bajpai; J. Advancem. Pl. Sci. 2(1)-1: 4 (2019)
Distribution: India (Madhya Pradesh)
40 IND
Remarks: provisionally accepted name
species :
Isoetes fuliginosa R.L.Small & Hickey; Amer. Fern J. 91(2): 55 (-57, figs. 5-7, 14-15) (2001)
Distribution: C-Colombia
82 CLM
species :
Isoetes fuscomarginata H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetac.): 12 (1986)
Distribution: S-Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes gardneriana Kunze ex Mett.; Fil. Lechl. Fasc. 11: 36 (1859)
Distribution: WC-Brazil (Goiás)
84 BRA
Synonyms: hidden
= Calamaria gardneriana (Kuntze ex Mett.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes georgiana Luebke; Amer. Fern J. 82: 24 (1992)
Name: Georgia quillwort
Distribution: USA (Georgia)
78 USA
species :
Isoetes giessii Launert; Mitt. Bot. Staatssamml. München 2: 309 (1957)
Distribution: Namibia
27 NAM
Conservation Status: known from two locations
species :
Isoetes gigantea U.Weber; Nova Hedwigia 63: 245 (1922)
Distribution: SE-Brazil (Rio de Janeiro)
84 BRA
species :
Isoetes graniticola D.F.Brunt.; Rhodora 118(975): 269 (2016)
Distribution: USA (Alabama)
78 USA
species :
Isoetes gunnii A.Braun; Monatsber. K. Akad. Berl. 535 (1868)
Distribution: Australia (Tasmania)
50 AUS
Synonyms: hidden
= Calamaria gunnii (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes lacustris (L.) Rodway, Tasm. Fl. 279 (1903)
species :
Isoetes gymnocarpa (Gennari) A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 555 (1864)
Name: Nacktsamiges Brachsenkraut; Naked-fruited Quillwort
Distribution: Italy; Greece (incl. Kiklades); Sicily; East Aegaean Isl. (Karpathos, Lesvos, Rhodos)
13 GRC ITA SIC 34 EAI
Synonyms: hidden
= Cephaloceraton gymnocarpum Gennari, Comment. Soc. Critt. Ital. 1(3): 113 (1862)
= Cephaloceraton hystrix var. subinermis Gennari, Comment. Soc. Critt. Ital. 1 (3): 112 (1862)
= Isoetes histrix f. subinermis (Gennari) Hayek, Repert. Spec. Nov. Regni Veg. Beih. 30(1): 12 (1924)
= Isoetes histrix var. subinermis (Gennari) Durand, Ces., Comp. Fl. It. 25 (1867), et Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 4009, t. 17), f. 1-5 (1883)
= Isoetes hystrix var. subinermis Gand., Fl. Eur. 27: 197 (1891)
= Isoetes sicula Tod., Syn. Pl. acot. vasc. Sic. 251 (1866)
= Isoetes subinermis (Gennari) Cesca & Peruzzi, Fl. Medit. 11: 308 (2001)
species :
Isoetes habbemensis Alston; J. Arnold Arbor. 26: 180 (1945)
Distribution: New Guinea (Irian Jaya: Mt. Wilhelmina-Lake Habbema)
42 NWG
Conservation Status: IUCN R (twice found)
species :
Isoetes hallasanensis H.K.Choi, Ch.Kim & J.Jung; J. Pl. Biol. 51(5): 357 (2008)
Distribution: South Korea (Cheju Isl.)
38 KOR
species :
Isoetes haussknechtii Troia & Greuter; Willdenowia 45(3): 395 (2015)
Name: Haussknechts Brachsenkraut
Distribution: Greece; East Aegaean Isl. (Lesvos)
13 GRC 34 EAI
Remarks: this is I. echinospora, I. setacea or I. delilei sensu auct. mult. from Greece. Distribution not yet completely known, may be more widespread in the Mediterranean
species :
Isoetes hawaiiensis W.C.Taylor & W.H.Wagner; Amer. Fern J. 83(2): 67 (1993)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
species :
Isoetes heldreichii Wettst.; Verh. Zool.-Bot. Ges. Wien 36: 239, t. 8 (1886)
Name: Heldreichs Brachsenkraut
Distribution: +C-Greece (+Palaiokastro near Karditsa)
13 GRC
Remarks: extinct fide Troia & Greuter 2015
species :
Isoetes hemivelata R.L.Small & Hickey; Amer. Fern J. 91(2): 57 (60, figs. 9-11, 16-17) (2001)
Distribution: C-Colombia
82 CLM
species :
Isoetes herzogii Weber; Hedwigia 63: 250, fig. 22-23 (1922)
Distribution: Bolivia (Cochabamba, La Paz, Santa Cruz); Peru (Cuzco)
83 BOL PER
species :
Isoetes hewitsonii Hickey; Fieldiana, Bot. 34: 94 (1994)
Distribution: Peru (Cajamarca)
83 PER
species :
Isoetes hieronymi Weber; Hedwigia 63: 251, fig. 24-25 (1922)
Distribution: Argentina (Cordoba, Rio Negro); Chile (Coquimbo)
85 ARG CLE
Synonyms: hidden
= Isoetes argentina Eaton, Palmer, Amer. Fern J. 19: 18 (1929) (nom. nud.
= Isoetes lechleri N.Pfeiff., Ann. Missouri Bot. Gard. 9(2): 138 (1922) (non Mett.)
species :
Isoetes histrix Bory & Durand; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1167 (1844)]
Name: Stacheliges Brachsenkraut
Distribution: S-England; Channel Isl.; Portugal; Spain; France; Corsica; Sardinia; Malta; Sicily; Italy; Macedonia; Albania; Crete; Greece (Aegaean region); Tunisia; Algeria; Morocco; Turkey (Anatolia); Lebanon; East Aegaean Isl. (Lesvos)
10 GRB 12 CIS COR FRA POR SAR SPA 13 ALB GRC ITA KRI MLT NMA SIC 20 ALG MOR TUN 34 EAI LEB TUR
Synonyms: hidden
= Calamaria hystrix (Durand) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Cephaloceraton hystrix (Bory & Durand) Gennari, Comment. Soc. Critt. Ital. 1 (3): 111 (1862)
= Cephaloceraton pseudohystrix Gennari, Ann. Agric. Sci. 11: 175 (1865)
= Isoetes chaeturetii Mendes, Agron. Lusit. 23: 7 (1961) (fide Musselman 2003)
= Isoetes delalandei J.Lloyd, Notes Fl. Ouest France: 25 (1851)
= Isoetes histrix ssp. delalandei (J.Lloyd) P.Fourn., Quatre Fl. France 6 (1934)
= Isoetes histrix ssp. sicula (Tod.) P.Fourn., Quatre Fl. France 6 (1934)
= Isoetes histrix var. desquamata A.Braun, Monatsber. K. Akad. Berl., Sitzung 7 Dec (1863): 617 (1864)
= Isoetes histrix var. desquamata f.longispina (A.Braun) Fiori, Fl. Ital. Crypt., pars V: 425, f. 154 (1943)
= Isoetes histrix var. grandis Milde, Batt. & Trab., Fl. Alg. & Tunis. 407 (1902)
= Isoetes histrix var. lacunosa Tod., Syn. Pl. acot. vasc. Sic. 46 (1866)
= Isoetes histrix var. loricata A.Braun, Monatsber. K. Akad. Berl., Sitzung 7 Dec (1863): 617 (1863), et Batt. & Trab., Fl. Alg. & Tunis. 407 (1902)
= Isoetes histrix subvar. armata Batt. & Trab., Fl. Alg. & Tunis. 407 (1902)
= Isoetes histrix subvar. inermis Batt. & Trab., Fl. Alg. & Tunis. 407 (1902)
= Isoetes histrix var. scutellata A.Braun apud A.Braun, Monatsber. K. Akad. Berl., Sitzung 7 Dec (1863): 617 (1864), Arm. Sci. Nat. 2: 371 (1864)
= Isoetes histrix var. submersa Trab., Batt. & Trab., Fl. Alg. & Tunis. 407 (1902)
= Isoetes hystrix ssp. algarbiensis Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. arcuata Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. cesatii Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. collina Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. corsica Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. hispanica Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. horridula Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. Iongifolia Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. longiseta Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. numidica Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. oceanica Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. oranensis Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. phrygia Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. todaroi Gand., Fl. Eur. 27: 197 (1891)
= Isoetes hystrix ssp. transiens Gand., Fl. Eur. 27: 197 (1891)
= Isoetes pseudohystrix Gennari, Levier, Barb., Fl. Sardoae Comp., 74, 195 (1885)
= Isoetes setacea Moris & De Not., Florula Capr. 161 (1839)
species :
Isoetes hopei J.R.Croft; Blumea 26: 177-190 (1980)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Conservation Status: EN in Papua New Guinea
species :
Isoetes howellii Engelm.; Trans. Acad. Sci. St. Louis 4: 385 (1882)
Name: Howell´s Quillwort
Distribution: USA (California, Idaho, Montana, Oregon, Utah, Washington State); Canada (British Columbia); Mexico (Baja California Norte, Baja California Sur)
71 CDN 73 USA 76 USA 79 MEX
Synonyms: hidden
= Isoetes melanopoda f. californica (Eaton) Clute, Fern-Allies 260 (1905)
= Isoetes melanopoda var. californica A.A.Eaton, Working List North American Pteridophytes 27 (1901)
= Isoetes nuda Engelm., Trans. Acad. Sci. St. Louis 4: 385 (1882)
= Isoetes underwoodii L.F.Henderson, Bot. Gaz. 23: 124 (1897)
species :
Isoetes humilior F.Muell. ex A.Braun; Linn. 25: 722 (1852)
Distribution: Australia (Tasmania)
50 AUS
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria humilior (F.Muell. ex A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes hookeri A.Braun, Herb Melbourn (1866), Monatsber. K. Akad. Berl. 538 (1868)
species :
Isoetes hyemalis D.F.Brunt.; Castanea 59(1): 13 (1994)
Name: Winter Quillwort
Distribution: E-USA (North Carolina, South Carolina, Virginia)
75 USA 78 USA
Remarks: sometimes included in I. riparia, but accepted by Troia et al. 2016
species :
Isoetes hypsophila Hand.-Mazz.; Anzeiger Acad. Wiss. Wien. Nr. 13: 95 (1923)
Distribution: China (SW-Sichuan, NW-Yunnan)
36 CHI
species :
Isoetes inflata E.R.L.Johnson; J. Roy. Soc. W. Australia 67(1): 37 (1984)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes jaegeri Pitot; Bull. Inst. Franc. Afr. Noire, A, 30: 58, f. 1-19 (1968)
Distribution: Burkina Faso
22 BKN
species :
Isoetes jamaicensis Hickey; Amer. Fern J. 71(3): 69 (1981)
Distribution: Jamaica
81 JAM
species :
Isoetes japonica A.Braun; Monatsber. K. Akad. Berl. 1: 459 (1861)
Distribution: Japan
38 JAP
Synonyms: hidden
= Calamaria japonica (A.Braun ) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes edulis Sieber ex Miq., Prod. Fl. jap. 390 (1866)-67
species :
Isoetes jejuensis H.K.Choi, Ch.Kim & J.Jung; J. Pl. Biol. 51(5): 356 (2008)
Distribution: South Korea (Cheju Isl.)
38 KOR
species :
Isoetes junciformis D.F.Brunt. & D.M.Britton; Amer. Fern J. 89 (3): 187-197 (1999)
Name: Rush quillwort
Distribution: USA (S-Georgia)
78 USA
species :
Isoetes karstenii A.Braun; Verh. Bot. Ver. Brandenb. 4. 332 (1862)
Distribution: Venezuela (Merida, Trujillo); Colombia (Arauca, Boyacá, Cauca, Cundinamarca, Huila, Magdalena, Meta, Nariño, Risaralda, Santander, Tolima, Valle)
82 CLM VEN
Synonyms: hidden
= Isoetes colombiana (T.C.Palmer) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 84(2): 176 (1981)
= Isoetes lechleri var. colombiana Palmer, Amer. Fern J. 19: 18 (1929)
= Isoetes steyermarkii Fuchs, (1960), nom. nud.
variant :
var. anomala (T.C.Palmer) R.L.Small & Hickey; Amer. Fern J. 22: 130 (1933)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Isoetes rimbachiana H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 85(2): 226 (1982), nom. nov. pro I. lechleri var. anomala
= Isoetes lechleri var. anomala T.C.Palmer, Amer. Fern J. 22: 130 (1933)
species :
Isoetes killipii C.V.Morton; Amer. Fern J. 35: 49 (1945)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Meta, Nariño, Putumayo, Risaralda, Santander, Tolima, Valle); Venezuela (Amazonas, Apure, Bolívar, Tachira); Ecuador; Guyana (Mt. Roraima)
82 CLM GUY VEN 83 ECU
Synonyms: hidden
= Isoetes roncadorensis H.P.Fuchs, (1981), nom. nud.
= Isoetes roraimensis H.P.Fuchs, (1981), nom. nud.
species :
Isoetes kirkii A.Braun; Monatsber. K. Akad. Berl. 649 (1869)
Distribution: New Zealand (North Isl., South Isl.)
51 NZL
Synonyms: hidden
= Calamaria kirkii (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes labri-draconis N.R.Crouch; Ferns Southern Africa 746 (2012)
Distribution: South Africa (KwaZulu-Natal: E-Drakensberg)
27 SAF
species :
Isoetes lacustris L.; Sp. Pl. 2: 1100 (1753)
Name: See-Brachsenkraut, Lake Quillwort
Distribution: Austria; Belgium; England; Wales; +Bulgaria; Czech Republic; Denmark; Finland; France; Germany (Baden-Württemberg, Bayern, Mecklenburg-Vorpommern+, Niedersachsen, Schleswig-Holstein); Ireland; Switzerland; Netherlands; Spain; Norway; Poland; ?Romania; Sweden; European Russia (N-European Russia, C-European Russia); Estonia; Latvia; Lithuania; Belarus; Siberia (W-Siberia, C-Siberia); ?Japan; USA (Connecticut, Massachusetts, Maine, Michigan, Minnesota, New Hampshire, New Jersey, New York, Rhode Island, Tennessee, Virginia, Vermont, Wisconsin); Canada (Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Ontario, Prince Edward Isl., Québec, Saskatchewan); Greenland; St. Pierre et Miquelon
10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER NET POL SWI 12 FRA SPA 13 BUL 14 BLR EST LAT LIT RUS 30 SIB 70 CDN GNL 71 CDN 72 CDN SPM 74 USA 75 USA 78 USA
Synonyms: hidden
= Calamaria lacustris (L.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes atrovirens Th.Fr., Bot. Notis. (1863): 48 (1865)
= Isoetes brochonii Motelay, Actes Soc. Linn. Bord. 45: 45 (1892) (fide Prelli & Bock 1989)
= Isoetes harveyi Eaton, Fernwort Papers 2: 11 (1900)
= Isoetes hieroglyphica A.A.Eaton, Fernwort Papers 10 (1900)
= Isoetes lacustris ssp. abbreviata Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. arvernensis Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. cambrica Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. editorum Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. errabunda Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. hieroglyphica (A.A.Eaton) Taylor & Luebke
= Isoetes lacustris ssp. lacustris Hook.fil., Student´s Fl. Brit. Isl., ed. 1, 471 (1870)
= Isoetes lacustris ssp. lamottei Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. langei Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. latifolia Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. noltei Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. obscura Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. pervaga Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. rivalis Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. rossica Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. scanica Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. scotica Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. septentrionalis Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. spreta Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. succedanea Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ssp. suecica Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris var. curvifolia Casp., Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 10 (pro syn.) (1936)
= Isoetes lacustris var. falcata Gay, Bull. Soc. Bot. France 10: 386 (1863)
= Isoetes lacustris var. humilior Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. humilis Mer, Bull. Soc. Bot. France 28- 72 (1881)
= Isoetes lacustris var. intermedia Mer, Bull. Soc. Bot. France 28: 72 (1881)
= Isoetes lacustris var. leiospora Klinggr., Schrift. Nat. Ges. Danzig 6, 1: 20 (1884)
= Isoetes lacustris var. morei (Moore) Hook., J. D., Student´s Fl. Brit. Isl., ed. 3, 526 (1884)
= Isoetes lacustris var. rectifolia Casp., Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 10 (1936)
= Isoetes lacustris var. recurva Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. setacea Léman, Dict. Sci. Nat. 24: 24 (1822)
= Isoetes lacustris var. stricta (Gay) Mer, Bull. Soc. Bot. France 28: 72 (1881)
= Isoetes lacustris var. tenuifolia A.Braun ex Milde, Sporenpfl. 141 (1865)
= Isoetes lacustris var. terrestris Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. vulgaris Casp., Luerss., Farnpfl. 855 (1889)
= Isoetes lacustris vergens Gand., Fl. Eur. 27: 193 (1891)
= Isoetes lacustris ×, stricta Gay
= Isoetes leiospora Klinggr., Schrift. Nat. Ges. Danzig 6 (1): 20 (1884)
= Isoetes macrospora Durand, Bull. Soc. Bot. France 11: 101 (1864)
= Isoetes macrospora f. hieroglyphica (A.A.Eaton) N.Pfeiff., Ann. Missouri Bot. Gard. 9: 194 (1922)
= Isoetes moorei Moore, J. Bot. (London) 16 (n. s. 7): 353, t. 199 (1878)
= Calamaria setacea (Lam.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes setacea Lam., Encycl. 3: 314 (1789)
= Isoetes tenella Lem. ex Desv., Mém. Soc. Linn. Paris 6: 179 (1827)
= Isoetes tuckermani var. harveyi (Eaton) Clute, Fern-Allies 226, 227, 260 (1905)
= Isoetes lacustris var. tenella Léman, Dict. Sci. Nat. 24: 24 (1822)
species :
Isoetes laosiensis C.Kim & H.K.Choi; Amer. Fern J. 100(1): 48 (2010)
Distribution: Laos
41 LAO
species :
Isoetes laurentiana D.F.Brunt.; Botany (Ottawa) 97(11): 579 (2019)
Distribution: Canada (Québec)
72 CDN
species :
Isoetes lechleri Mett.; Fil. Lechl. 2: 36 (1859)
Distribution: Peru (Junín, Puno, Ayacucho, Ancash); Colombia (Arauca, Boyacá, Cauca, Cundinamarca, Huila, Meta, Nariño, Santander); Ecuador (Chimborazo, Pinchincha, Napo); Bolivia (La Paz)
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Calamaria lechleri (Mett.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes arumiana Fuchs, (1959), nom. nud.
= Isoetes glacialis Aspl., Ark. Bot. 20A, no. 7: 35, fig. 7 (1926)
= Isoetes laevis Weber, Hedwigia 63: 252, fig. 30-31 (1922)
= Isoetes peruviana Weber, Hedwigia 63: 246, fig. 12-13 (1922)
= Isoetes socia A.Braun, Verh. Bot. Ver. Brandenb. 4: 332 (1862)
species :
Isoetes libanotica Musselman, Bolin & R.D.Bray; Novon 21(3): 295 (2011)
Distribution: Lebanon
34 LEB
species :
Isoetes lithophila N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 135 (1922) (lithophylla)
Name: Rock Quillwort
Distribution: USA (SC-Texas: Burnett Co., Mason Co., Llano Co.)
73 USA 77 USA
Conservation Status: IUCN V
species :
Isoetes longissima Bory; Compt. Rend. Acad. Sci. 18: 1165 (1844)
subspecies :
subsp. longissima
Distribution: Spain; Portugal; Baleares; France; Corsica; Sardinia; Sicily; Italy; Libya; Tunisia; Algeria; Morocco
12 BAL COR FRA POR SAR SPA 13 ITA SIC 20 ALG LBY MOR TUN
Synonyms: hidden
= Calamaria longissima (Bory & Durand) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes baetica Willk., Willk. & Lange, Prodr. Fl. Hispan. 1: 14 (1861)
= Isoetes boryana var. lereschii Rchb.fil.apud Ler. & Lév., Deux Exc. Bot. Esp. 118 (1880)
= Isoetes chaboissaei Nym., Consp. Fl. Eur. Suppl. 1: 871 (1883)-84
= Isoetes decipiens Bory, Flora 29: 719 (1846) (nom. prov)
= Isoetes dubia var. emersa Gennari, Nuov. Giorn. Bot. Ital. 2: 96 (1870)
= Isoetes dubia var. maculosa Gennari, Comment. Soc. Critt. Ital. 1(2): 105 (1861)
= Isoetes grammontii Bory, Milde, Fil. Eur. & Atl. 280 (1867)
= Isoetes iluliana Gennari, Comment. Soc. Critt. Ital. 1: 4 (1861) (febr.) (syn. subst.)
= Isoetes lacustris Moris, Stirp. Sard. EI. 3: 12 (1829)
= Isoetes setacea ssp. longissima (Bory) Greuter & Burdet, Willdenowia 14(1): 37 (1984)
= Isoetes sicula Tod. ex Ces. & De Not.apud Gennari, Comm. Soc. Critt. It. 1 (2): 103 (1861)
= Isoetes tenuissima Mart., Cat. 341
= Isoetes tenuissima var. chaboissaei Le Grand, Bull. Soc. Bot. France 42: 50 (1895)
= Isoetes variabilis Le Grand, Mém. Soc. Hist. Cher ser. 4, 1896: 11 (1896) (nom. illeg.)
= Isoetes velata A.Braun, Bory & Durand, Expl. Sci. Alg., Atlas: tab. fig. 1 (1849)
= Isoetes velata f. baetica (Willk.) Prada, Acta Bot. Malac. 8: 88 (1983)
= Isoetes velata f. lereschii (Rchb.fil. ex Leresche & Levier) Prada, Lazaroa 3: 33 (1982)
= Isoetes velata ssp. algeriensis Gand., Fl. Eur. 27: 195 (1891)
= Isoetes velata ssp. australis Gand., Fl. Eur. 27: 195 (1891)
= Isoetes velata ssp. barbara Gand., Fl. Eur. 27: 194 (1891)
= Isoetes velata ssp. calaostiensis Gennari, Spec. e Var. Fl. Sarda 31 (1867)
= Isoetes velata ssp. divergens Gand., Fl. Eur. 27: 195 (1891)
= Isoetes velata ssp. gallica Gand., Fl. Eur. 27: 194 (1891)
= Isoetes velata ssp. longissima (Bory) Greuter & Burdet, Willdenowia 14(1): 37 (1984)
= Isoetes velata ssp. redux Gand., Fl. Eur. 27: 195 (1891)
= Isoetes velata ssp. robusta Gand., Fl. Eur. 27: 194 (1891)
= Isoetes velata ssp. tenuicula Gand., Fl. Eur. 27: 195 (1891)
= Isoetes velata ssp. trinacria Gand., Fl. Eur. 27: 195 (1891)
= Isoetes velata var. brevifolia (A.Braun) Franch., Bull. Soc. Bot. France 31: 349 (1884)
= Isoetes velata var. brevifolia A.Braun, Sitzungsber. Akad. Wiss. Wien 605 (1863), et Batt. & Trab., Fl. Alg. & Tunis. 407 (1902)
= Isoetes velata var. humilior A.Braun ex Mares & Vigin., Pl. Vasc. des Iles Baleares, 322 (1890)
= Isoetes velata var. lereschii (Rchb.fil. ex Leresche & Levier) Molina Abril & Sard.Rosc., Guineana 10: 346 (2004)
= Isoetes velata var. longissima A.Braun, Bory & Durand, Expl. Sci. Alg., t. 37, fig. 2 (1848)
= Isoetes velata var. sicula Gennari, Milde, Fil. Eur. & Atl. 282 (1867)
= Isoetes velata var. sicula Tod., Syn. Pl. acot. vasc. Sic. 250 (1866)
= Isoetes vellata var. boryana (Durieu) M.I.Romero & Real, Bot. J. Linn. Soc. 148(4): 464 (2005)
subspecies :
subsp. adspersa (A.Braun) Troia & Greuter; Willdenowia 44(2): 288 (2014)
Distribution: France; Tunisia; Algeria; Morocco
12 FRA 20 ALG MOR TUN
Synonyms: hidden
= Isoetes velata ssp. adspersa (A.Braun) Batt. & Trab., Fl. Alg. & Tunis. 407 (1905)
= Calamaria adspersa (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes adspersa A.Braun, Bory & Durand, Expl. Sci. Alg., Atlas: tab. 37, rig. 3 (1849)
= Isoetes adspersa var. perreymondii A.Braun, Schoenefeld, Bull. Soc. Bot. France 12: 261 (1865)
= Isoetes aspersum Asch. & Graebn., Syn. Mitteleur. Fl. 1: 169 Hauptreg. 23 (1897)
= Isoetes capillacea Bory, Compt. Rend. Acad. Sci. 23: 620 (1846)
= Isoetes lineolata Durand ex Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 381 (1883)
= Isoetes perreymondii (Bory) Duval-Jouve, Bull. Soc. Bot. France 16: 213 (1869)
= Isoetes setacea Bory, Compt. Rend. Acad. Sci. 18: 1165 (1844)
= Isoetes setacea var. perreymondii Bory, Compt. Rend. Acad. Paris 18: 1165 (1844)
= Isoetes velata perreymondii (A.Braun) Franch., Bull. Soc. Bot. France 31: 349 (1884)
= Isoetes velata var. perrymondii (Bory) Franch., Die Süsswasser-Flora Mitteleuropas 15: 17 (1936)
subspecies :
subsp. asturicensis (M.Laínz) comb.ined.
Distribution: Spain
12 SPA
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes asturicensis (M.Laínz) M.Laínz, Anuario Soc. Brot. 39: 118 (1973)
= Isoetes velata ssp. asturicensis (M.Laínz) Rivas Mart. & Prada, Lazaroa 2: 327 (1980)
= Isoetes boryana ssp. asturicensis M.Laínz, Bol. Inst. Estud. Astur., Suppl. Cienc. 15: 6 (1970)
= Isoetes velata var. asturicense (M.Laínz) E.Rico & X.Giráldez, Anal. Jard. Bot. Madrid 46(2): 585 (1989 publ. 1990)
subspecies :
subsp. intermedia (Trab.) Troia & Greuter; Willdenowia 44(2): 288 (2014)
Distribution: Morocco; Algeria
20 ALG MOR
Synonyms: hidden
= Isoetes velata ssp. intermedia (Trab.) Maire & Weiller, Maire, Fl. Afrique. N. 1: 89 (1952)
= Isoetes velata f. immaculata Maire, Encycl. Biol. 33: 90 (1952)
= Isoetes velata tax.infrasp.intermedia Trab., Batt. & Trab., Fl. Alg. & Tunis. 407 (1905)
= Isoetes velata var. intermedia A.Braun, Bory & Durand, Expl. Sci. Alg. (1848)
subspecies :
subsp. perralderiana (Milde) Troia & Greuter; Willdenowia 44(2): 288 (2014)
Distribution: Algeria
20 ALG
Synonyms: hidden
= Calamaria perralderiana (Durand & Letourn.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes perralderiana Durand & Letourn., Kralik, Pl. Alg. Exsicc. 157 (1862)
= Isoetes perralderiana Milde, Fil. Eur. 282 (1867) (basion.)
= Isoetes velata ssp. perralderiana (Durand & Letourn.) Trab., Fl. Syn. 407 (1902), et Fl. Alg. & Tunis. 407 (1905)
= Isoetes velata var. perralderiana (Durand & Letourn.) N.Pfeiff., Ann. Missouri Bot. Gard. 9: 121 (1922)
subspecies :
subsp. tenuissima (Boreau) Troia & Greuter; Willdenowia 44(2): 288 (2014)
Distribution: Baleares; France
12 BAL FRA
Conservation Status: IUCN V
Synonyms: hidden
= Isoetes velata ssp. tenuissima (Boreau) O.Bolòs & Vigo, Butll. Inst. Catalana Hist. Nat. 38: 64 (1974)
= Calamaria tenuissima (Boreau) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes chaboissaei Hy, Morot, Journ. de Bot. 8: 97 (1894)
= Isoetes tenuissima Boreau, Bull. Soc. Industr. Angers 21: 269 (1850) (basion.)
= Isoetes tenuissima var. capillifolia Le Grand, Bull. Soc. Bot. France 42: 50 (1895)
= Isoetes tenuissima var. latifolia Le Grand, Bull. Soc. Bot. France 42: 50 (1895)
= Isoetes tenuissima var. longifolia Le Grand, Bull. Soc. Bot. France 42: 50 (1895)
= Isoetes tenuissima var. tenuissima Le Grand, Bull. Soc. Bot. France 42: 50 (1895)
= Isoetes viollaei F.Hy, Bull. Herb. Boiss. ser. 3, App. 1: 23 (1894)
species :
Isoetes louisianensis Thieret; Sida 5: 129 (1973)
Name: Louisiana Quillwort
Distribution: USA (S-Louisiana, Mississippi, Alabama)
78 USA
Remarks: may be a hybrid I. engelmannii x melanopoda
Conservation Status: IUCN E
species :
Isoetes luetzelburgii Weber ex Luetzelb.; Estud. Bot. Nordeste 3: 257 (1864)
Distribution: Brazil (Rio Grande do Norte)
84 BRA
species :
Isoetes malinverniana Ces. & De Not.; Ind. Sem. Hort. Genuensis 1858: 36 (1858)
Distribution: NW-Italy (Vercelli, Novara, Torino)
13 ITA
Conservation Status: IUCN E
Synonyms: hidden
= Calamaria malinverniana (Ces. & De Not.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes maritima Underw.; Bot. Gaz. 13: 94 (1888)
Name: Spiny-spored Quillwort
Distribution: Russian Far East (Commander Isl.); Aleutes; Alaska; W-Canada (Vancouver Isl., British Columbia); NW-USA (Washington)
31 RFE 70 ASK 71 CDN 73 USA
Synonyms: hidden
= Isoetes beringensis Kom., Bull. Jard. Bot. Acad. Sci. USSR 30 (1-2): 196 (1932)
= Isoetes braunii var. maritima (Underw.) N.Pfeiff., Ann. Missouri Bot. Gard. 9: 174 (1922)
= Isoetes echinospora f. maritima (Underw.) Clute
= Isoetes echinospora ssp. maritima (Underw.) Á.Löve, Amer. Fern J. 52: 121 (1962)
= Isoetes echinospora var. maritima (Underw.) Eaton, Fern Bull. 13: 52 (1905)
= Isoetes echinospora var. savilei B.Boivin, Amer. Fern J. 51: 85 (1961)
= Isoetes macounii Eaton, Fern Bull. 8: 12 (1900)
= Isoetes muricata ssp. maritima (Underw.) Hultén, Ark. Bot. (n. s.) 7(1): 8 (1968)
= Isoetes setacea ssp. maritima (Underw.) Holub, Acta Horti bot. prag.: 55 (1964)
species :
Isoetes martii A.Braun ex Kuhn; Mart., Fl. Bras. 1 (2): 646, t. 78 (1884)
Distribution: SE-Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Calamaria martii (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes mattaponica Musselman & W.C.Taylor; Novon 11(2): 200 (-204, figs. 1-6) (2001)
Name: Mattaponi quillwort
Distribution: USA (Virginia: Mattaponica River)
78 USA
species :
Isoetes maxima Hickey, Macluf & Link-Pérez; Amer. Fern J. 99(3): 194 (2010)
Distribution: S-Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes melanopoda Gay & Durand; Bull. Soc. Bot. France 11: 102 (1864)
subspecies :
subsp. melanopoda
Name: Black-footed Quillwort
Distribution: USA (Alabama, Arkansas, Georgia, Iowa, Idaho, Illinois, Indiana, Kansas, Kentucky, Louisiana, Minnesota, Missouri, Mississippi, Montana, North Carolina, Nebraska, New Jersey, Oklahoma, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia)
73 USA 74 USA 75 USA 76 USA 77 USA 78 USA
Synonyms: hidden
= Calamaria melanopoda (Gay & Durand) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes melanopoda f. pallida (Engelm.) Fernald, Rhodora 51: 103 (1949)
= Isoetes melanopoda var. pallida Engelm., Trans. Acad. Sci. St. Louis 4: 387 (1882)
subspecies :
subsp. silvatica D.F.Brunt. & D.M.Britton; Castanea 71(1): 26 (2006)
Distribution: E-USA (North Carolina)
75 USA 78 USA
species :
Isoetes melanospora Engelm.; Trans. Acad. Sci. St. Louis 3: 395 (1877)
Name: Blackspore Quillwort
Distribution: USA (C-Georgia, South Carolina: Lancaster Co.)
78 USA
Synonyms: hidden
= Calamaria melanospora (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes melanotheca Alston; Bol. Soc. Broter. Ser. 2A, 30: 15 (1956)
Distribution: Guinea; Guinea-Bissau; Benin
22 BEN GNB GUI
Conservation Status: IUCN R
species :
Isoetes mexicana Underw.; Bot. Gaz. 13: 93 (1888)
Distribution: N-Guatemala; Mexico (Aguascalientes, Chihuahua, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Queretaro, Zacatecas)
79 MEX 80 GUA
species :
Isoetes microvela D.F.Brunt & D.M.Britton; Rhodora 100(903): 270-274, f. 1-3, 4e-h, 5 (1998)
Name: Thin-wall quillwort
Distribution: USA (North Carolina: coastal)
78 USA
species :
Isoetes minima A.A.Eaton; Fern Bull. 6: 30 (1898)
Name: Midget quillwort
Distribution: USA (Washington State); SW-Canada (British Columbia)
71 CDN 73 USA
Remarks: valid species fide Taylor & al. 2003
Synonyms: hidden
= Isoetes howellii var. minima (A.A.Eaton) N.Pfeiff., Ann. Missouri Bot. Gard. 9(2): 142 (1922)
species :
Isoetes mississippiensis S.W.Leonard, W.C.Taylor, Musselman & R.D.Bray; PhytoKeys 2016(74): 97-106 (2016)
Distribution: USA (S-Mississippi)
78 USA
species :
Isoetes mongerensis E.R.L.Johnson; J. Roy. Soc. W. Australia 67(1): 39 (1984)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes montezumae A.A.Eaton; Fern Bull. 5: 25 (1897)
Distribution: Mexico
79 MEX
Remarks: recognized in Troia et al. 2016, included in I. mexicana by other authors
species :
Isoetes mourabaptistae J.B.S.Pereira, P.G.Windisch, Lorscheitt. & Labiak; Amer. Fern J. 102 (2): 175 (2012)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes muelleri A.Braun; Monatsber. K. Akad. Berl. 541 (1868)
Distribution: Australia (Western Australia, Northern Territory, South Australia, Queensland, Victoria, Tasmania)
50 AUS
Synonyms: hidden
= Calamaria muelleri (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes stuartii A.Braun, Monatsber. K. Akad. Berl. 539 (1868)
= Isoetes tenuissima F.Muell., (1868) (non Boreau)
species :
Isoetes naipiana P.G.Windisch, Lorscheitt. & Nervo; Willdenowia 44: 394 (2014)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes nana J.B.S.Pereira; PhytoKeys 89: 93 (2017)
Distribution: Brazil (Rio de Janeiro)
84 BRA
species :
Isoetes narsinghgarhensis H.K.Goswami & Mazumdar; J. Advancem. Pl. Sci. 2(1)-1: 5 (2019)
Distribution: India (Madhya Pradesh)
40 IND
Remarks: provisionally accepted name
species :
Isoetes neoguineensis Baker ex F.Muell.; Ann. Rep. Brit. N. Guinea 1897-8: 149, nomen (1898)
Distribution: New Guinea (Papua New Guinea: Owen Stanley Ranges)
42 NWG
Conservation Status: IUCN R (twice found)
Synonyms: hidden
= Isoetes neoguineensis Baker, Sadeb., Nat. Pfl. 1(4) 776 (1902)
= Isoetes neoguineensis var. rheophila J.R.Croft, Blumea 26: 177-190 (1980)
species :
Isoetes nigritiana A.Braun; Fil. Afr. 196 (1868)
Distribution: Sudan; Burkina Faso; Mali; Nigeria; Central African Republic; Ivory Coast
22 BKN IVO MLI NGA 23 CAF 24 SUD
Synonyms: hidden
= Isoetes garnieri Chev. & Monnier, Bull. Mus. Hist. Nat. Paris 18(1): 91-979 1 fig. (1946)
= Calamaria nigritiana (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes nigroreticulata Verdc.; Fl. Trop. E. Africa, Isoet. 2 (2005)
Distribution: Kenya
25 KEN
species :
Isoetes novogranadensis H.P.Fuchs; Caldasia 8: 314 (1969)
Distribution: Colombia (Bolívar, Boyacá, Cauca, Cundinamarca, Huila, Nariño, Putumayo, Valle); Ecuador; Peru
82 CLM 83 ECU PER
Synonyms: hidden
= Isoetes dichotoma L.E.Mora & W.Hagemann, Mutisia 43: 5 (1977)
= Isoetes hagemannii H.P.Fuchs, (1981), nom. nud.
= Isoetes narinensis H.P.Fuchs, (1981), nom. nud.
= Isoetes leistii H.P.Fuchs, (1981), nom. nud.
= Isoetes pichichensis H.P.Fuchs, (1981), nom. nud.
= Isoetes quilindanensis H.P.Fuchs, (1981), nom. nud.
species :
Isoetes nuttallii A.Braun ex Engelm.; Amer. Naturalist 8: 215 (1874)
Name: Nuttall´s Quillwort
Distribution: W-USA (California, Oregon, Washington); W-Canada (Vancouver Isl.)
71 CDN 73 USA 76 USA
Synonyms: hidden
= Calamaria nuttallii (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Calamaria suksdorfii (Baker) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes opaca Nutt., ex Engelm., Trans. St. Louis Acad. Sci. 4: 388 (1882)
= Isoetes suksdorfii Baker, Handb. Fern-Allies 132 (1887)
species :
Isoetes occidentalis Hend.; Bull. Torrey Bot. Cl. 27: 358 (1900)
Name: Western Quillwort
Distribution: Aleutes; Alaska; W-Canada (British Columbia); NW-USA (Washington, Oregon, C-California, Idaho, Montana, Wyoming, Colorado)
70 ASK 71 CDN 73 USA 76 USA
Synonyms: hidden
= Isoetes echinospora f. flettii (A.A.Eaton) Clute, Fern Bull. 16: 55 (1908)
= Isoetes echinospora ssp. flettii (A.A.Eaton) Piper & Beattie, Fl. N. W. Coast [Piper & Beattie] 16 (1915)
= Isoetes echinospora var. flettii A.A.Eaton, Fern Bull. 13: 51 (1905)
= Isoetes flettii (A.A.Eaton) N.Pfeiff., Ann. Missouri Bot. Gard. 9(2): 186 (1922)
= Isoetes howellii f. piperi (Eaton) Clute, Fern Bull. 16: 56 (1908)
= Isoetes howellii var. piperi (Eaton) Clute, Fern-Allies 246, 258 (1905)
= Isoetes lacustris ssp. paupercula (Engelm.) Feilberg, Nordic J. Bot. 4: 155 (1984)
= Isoetes lacustris var. paupercula Engelm., Trans. Acad. Sci. St. Louis 4: 377 (1882)
= Isoetes macrospora f. paupercula (Engelm.) Clute, Fern Bull. 16: 56 (1908)
= Isoetes maritima var. flettii (Eaton) C.F.Reed, Amer. Fern J. 35: 85 (1945)
= Isoetes occidentalis ssp. paupercula (Engelm.) Feilberg, Nordic J. Bot. 4(2): 155 (1984)
= Isoetes occidentalis var. piperi (A.A.Eaton) A.Nelson & J.F.Macbr., Bot. Gaz. 61(1): 30 (1916)
= Isoetes paupercula (Engelm.) A.A.Eaton, Proc. U. S. Nat. Herb. 23(1226): 649 (1901)
= Isoetes piperi A.A.Eaton, Fern Bull. 11: 85 (nom. nud.) (1903), 13: 51 (1905)
species :
Isoetes olympica A.Braun; Milde, Fil. Eur. 285 (1867)
Distribution: Turkey (Anatolia); S-Syria; Lebanon
34 LEB SYR TUR
Remarks: histrix agg.
Synonyms: hidden
= Calamaria olympica (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes orcuttii Eaton; Fern Bull. 6: 30 (1898) (nom. nud.), et 8: 13 (1900)
Name: Orcutt´s Quillwort
Distribution: USA (S-California); Mexico (Baja California Norte)
76 USA 79 MEX
Synonyms: hidden
= Isoetes nuttallii var. orcuttii (A.A.Eaton) Clute, Fern-Allies 253 (1905)
species :
Isoetes organensis Weber; Luetzelburg, Estud. Bot. Nordeste 3: 256 (no date)
Distribution: SE-Brazil (Rio de Janeiro)
84 BRA
species :
Isoetes orientalis Hong Liu & Q.F.Wang; Novon 15(1): 164 (2005)
Distribution: China (Zhejiang)
36 CHI
Remarks: probably a hybrid Isoetes sinensis x yunguiensis
species :
Isoetes ovata N.Pfeiff.; Ann. Missouri Bot. Gard. 8(2): 109 (1922)
Distribution: Guyana; French Guiana
82 FRG GUY
Synonyms: hidden
= Isoetes clavata Weber, Ber. dt. Bot. Ges. 52: 121 (1934)
species :
Isoetes pallida Hickey; Amer. Fern J. 78(1): 35 (1988)
Distribution: Mexico (Campeche, Guerrero, Oaxaca); Belize
79 MEX 80 BLZ
Conservation Status: IUCN R
species :
Isoetes palmeri H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 168 (1981)
Distribution: Venezuela (Merida); Colombia (Arauca, Boyacá, Cauca, Cundinamarca); Ecuador (Chimborazo)
82 CLM VEN 83 ECU
Synonyms: hidden
= Isoetes cleeffii H.P.Fuchs, Proc. Kon. Akad. Wet. Amsterdam C 84(2): 177 (1981)
species :
Isoetes panamensis Maxon & C.V.Morton; Ann. Missouri Bot. Gard. 26(4): 272 (1939)
Distribution: Nicaragua; Costa Rica; Panama; Guatemala; Colombia; Ecuador; E-Peru; NE-Brazil (Maranhao, Bahia); WC-Brazil (Mato Grosso); Paraguay (Alto Paraná, Caaguazu); Bolivia (Cochabamba, La Paz)
80 COS GUA NIC PAN 82 CLM 83 BOL ECU PER 84 BRA 85 PAR
Synonyms: hidden
= Isoetes pacifica Svenson, Amer. Fern J. 34: 123 (1944)
= Isoetes savannarum L.D.Gómez, Phytologia 49(4): 339 (1981)
species :
Isoetes parvula Hickey; Fieldiana, Bot. 34: 92 (1994)
Distribution: Peru (Ayacucho)
83 PER
Conservation Status: Known from type only
species :
Isoetes pedersenii H.P.Fuchs ex E.I.Meza & Macluf; Anais Acad. Brasil. Ci. 82(2): 354 (2010)
Distribution: Argentina (Corrientes); S-Brazil; Paraguay
84 BRA 85 ARG PAR
species :
Isoetes perrieriana Iversen; Dansk Bot. Ark. 7: 200-201 (1932)
Distribution: Madagascar
29 MDG
species :
Isoetes philippinensis Merr. & Perry; Amer. Fern J. 30: 19, fig. (1940)
Distribution: Philippines (Lanao del Norte); Sulawesi
42 PHI SUL
Conservation Status: CR in the Philippines, possibly extinct in known locality
species :
Isoetes phrygia Hausskn.; Mitt. Thüring. Bot. Vereins 13: 72 (1899)
Distribution: Crete; Turkey (Anatolia)
13 KRI 34 TUR
Synonyms: hidden
= Isoetes histrix var. phrygia Boiss., Fl. Orient. 5: 748 (1884)
species :
Isoetes pitotii Alston; Bol. Soc. Broter. Ser. 2, 30: 15 (1956)
Distribution: Mali; Guinea; Sudan
22 GUI MLI 24 SUD
species :
Isoetes precocia R.L.Small & Hickey; Amer. Fern J. 91(2): 61 (-62, figs. 12-13, 18-19) (2001)
Distribution: Venezuela (Merida); ?Colombia
82 VEN
species :
Isoetes pringlei Underw.; Zoë 1: 98 (1890)
Distribution: Mexico (Jalisco, Mexico State, Chiapas)
79 MEX
Conservation Status: IUCN R
species :
Isoetes prototypus D.M.Britton; Canad. J. Bot. 69: 278 (1991)
Name: Spike quillwort
Distribution: USA (Maine, New Hampshire); Canada (New Brunswick, Nova Scotia)
72 CDN 75 USA
Conservation Status: IUCN V
species :
Isoetes pseudojaponica M.Takamiya, Watan. & Ono; Acta Phytotax. Geobot. 48(2): 107 (1998)
Distribution: Japan
38 JAP
Conservation Status: VU in Japan
species :
Isoetes pusilla C.R.Marsden & Chinnock; Muelleria 8(1): 57 (1993)
Distribution: Australia (Victoria)
50 AUS
species :
Isoetes quiririensis J.B.S.Pereira & Labiak; Syst. Bot. 38(4): 869 (2014)
Distribution: S-Brazil (Santa Catarina)
84 BRA
species :
Isoetes ramboi Herter; Rev. Südam. Bot. 8 (1-2): 19-25 (1949)
Distribution: S-Brazil (Rio Grande do Sul, Santa Catarina)
84 BRA
species :
Isoetes riparia Engelm. ex A.Braun; Flora (Regensburg) 29: 178 (1846)
Name: Shore Quillwort
Distribution: USA (Delaware, District of Columbia, Maryland, North Carolina, New Jersey, New York, Pennsylvania, Rhode Island, South Carolina, Vermont, Virginia, West Virginia)
75 USA 78 USA
Remarks: riparia agg.
Synonyms: hidden
= Calamaria riparia (Engelm. ex A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes foveolata var. plenospora Eaton, Rhodora 5: 280 (1903)
= Isoetes riparia var. riparia Proctor, Amer. Fern J. 39: 119 (1949)
= Isoetes riparia var. amesii (Eaton) Proctor, Amer. Fern J. 39: 119 (1949)
= Isoetes amesii Eaton, Gilbert, Fern Bull. 11: 103 (1903)
= Isoetes canadensis var. amesii (Eaton) Clute, Fern Bull. 16: 54 (1908)
= Isoetes riparia f. saccharata (Engelm.) Proctor, Amer. Fern J. 39: 119 (1949)
= Isoetes saccharata var. amesii A.A.Eaton, Rhodora 5(59): 278-279 (1903)
= Isoetes dodgei Eaton, Fern Bull. 6: 6 (1898)
= Isoetes riparia f. saccharata (Engelm.) Clute, Fern Bull. 16: 57 (1908)
= Isoetes riparia var. palmeri (Eaton ) Proctor, Amer. Fern J. 39: 119 (1949)
= Isoetes saccharata f. palmeri Eaton, Fern Bull. 11 : 25 (1903)
= Isoetes saccharata var. palmeri A.A.Eaton, Proc. Biol. Soc. Wash. 14(10) 49 (1901)
= Isoetes riparia var. reticulata (Eaton) Proctor, Amer. Fern J. 39: 119 (1949)
= Isoetes saccharata f. reticulata Eaton, Fern Bull. 11 : 25 (1903)
= Isoetes saccharata var. reticulata A.A.Eaton, Proc. Biol. Soc. Wash. 14(10): 49 (1901)
= Isoetes saccharata var. reticulata Eaton, Steele, Proc. Biol. Soc. Wash. 14: 49 (1901)
species :
Isoetes sabatina Troìa & Azzella; Pl. Biosystems 147(4): 1052 (2013)
Distribution: Italy
13 ITA
species :
Isoetes saccharata Engelm.; Gray´s Manual, ed. 5, 676 (1867)
Name: Sugar quillwort
Distribution: USA (District of Columbia, Delaware, Maryland, Virginia)
78 USA
Synonyms: hidden
= Calamaria saccharata (Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
species :
Isoetes sahyadrii Mahab.; [Curr. Sci. 7 (2): 62, f. 1 (1938), nom. inval.] ex L. N. Rao, Curr. Sci. 13(11): 286-287 (1944) (sahyadriensis auct.)
Distribution: India (Andhra Pradesh, Karnataka, Maharashtra, Madhya Pradesh, Rajasthan)
40 IND
Synonyms: hidden
= Isoetes mahadevensis G.K.Srivast., D.D.Pant & P.K.Shukla, Amer. Fern J. 83(4): 106 (1993)
= Isoetes panchananii D.D.Pant & Srivast., Proc. Nat. Inst. Sci. India 28: 243, t. 13, 14, f. 1, 2 (1962)
= Isoetes panchananii var. panchmarhiensis G.K.Srivast., Mritt.Srivast. & P.K.Shukla, Indian Fern J. 14(1-2): 44 (1998)
= Isoetes panchganiensis G.K.Srivast., D.D.Pant & P.K.Shukla, Amer. Fern J. 83(4): 107 (1993) (fide Fraser-Jenkins 1997)
= Isoetes panchganiensis var. kemangundiensis G.K.Srivast., D.D.Pant & P.K.Shukla, Amer. Fern J. 83(4): 114 (1993) (fide Fraser-Jenkins 1997)
= Isoetes rajasthanensis Gena & Bhardwaja, J. Bombay Nat. Hist. Soc. 81(1): 165 (1984)
= Isoetes reticulata Gena & Bhardwaja, J. Bombay Nat. Hist. Soc. 81(1): 166 (1984)
= Isoetes sampathkumarnii L.N.Rao, Curr. Sci. 13: 286, f. 1-3 (1944)
species :
Isoetes santacruzensis J.B.S.Pereira; PhytoKeys 131: 61 (2019)
Distribution: Bolivia
83 BOL
species :
Isoetes saracochensis Fuchs ex Hickey; Fieldiana, Bot. 34: 94 (1994)
Distribution: Peru
83 PER
Synonyms: hidden
= Isoetes saracochensis Fuchs, (1960), nom. nud.
species :
Isoetes savatieri Franch.; Bull. Soc. Bot. France 31: 395 (1884)
Distribution: Argentina (Chubut, Neuquen, Rio Negro, Tierra del Fuego); Chile (Bio Bio, Araucania, Los Lagos, Aisen, Magallanes)
85 ARG CLE
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria savatieri (Franch.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes lechleri var. savatieri (Franch.) L.D.Gómez, Brenesia 18: 5 (1980)
= Isoetes chilensis H.P.H.P.Fuchs, (1981), nom. nud.
species :
Isoetes schweinfurthii A.Braun; J. Bot. 18: 108 (1880)]
Distribution: Senegal; Burkina Faso; Chad; Ivory Coast; Benin; Central African Republic; Sudan; Ethiopia; Tanzania; S-Angola; Zambia; Malawi; Mozambique; Zimbabwe; S-Botswana; N-Namibia; South Africa (Limpopo); Madagascar
22 BEN BKN IVO SEN 23 CAF 24 CHA ETH SUD 25 TAN 26 ANG MLW MOZ ZAM ZIM 27 BOT NAM SAF 29 MDG
Synonyms: hidden
= Calamaria schweinfurthii (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes rhodesiana Alston, Bol. Soc. Broter. Ser. 2, 30: 17 (1956)
= Isoetes kersii Wanntorp, Svensk Bot. Tidsskr. 64: 146, t. 1, 3 (1970)
species :
Isoetes sehnemii H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 38 (1986)
Distribution: S-Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes septentrionalis D.F.Brunt.; Canad. Field-Naturalist 129: 176 (2015)
Name: Northern Quillwort
Distribution: USA (Connecticut, Massachusetts, New Hampshire, ?Maine, Pennsylvania, Vermont); E-Canada (SE-Ontario, SW-Québec)
72 CDN 75 USA
Remarks: riparia agg.
species :
Isoetes serracarajensis J.B.S.Pereira, Salino & Stützel; Phytotaxa 272(2): 142 (2016)
Distribution: Brazil (Pará)
84 BRA
species :
Isoetes shangrilaensis Xiang Li, Yuqian Huang, X.Dai & Xing Liu; Phytotaxa 397(1): 71 (2019)
Distribution: China (Yunnan)
36 CHI
species :
Isoetes sinensis Palmer; Amer. Fern J. 17: 112 (1927)
Distribution: China (Anhui, Guangxi, Jiangsu, Zhejiang); S-Japan; South Korea
36 CHI 38 JAP KOR
Remarks: allotetraploid originated from I. yunguiensis x taiwanensis
species :
Isoetes smithii H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 17 (1986)
Distribution: S-Brazil (Rio Grande do Sul, Santa Catarina)
84 BRA
species :
Isoetes spannagelii H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 33 (1986)
Distribution: S-Brazil (Rio Grande do Sul, Santa Catarina)
84 BRA
species :
Isoetes spinulospora Jermy & Schelpe; Contr. Bolus Herb. 10: 151 (1982)
Distribution: Bioko Isl. [Fernando Poo]
23 BIO
Conservation Status: IUCN R
species :
Isoetes stellenbossiensis A.V.Duthie; Trans. R. Soc. S. Afr. 17: 328 (1929)
Distribution: South Africa (W-Cape Prov., N-Cape Prov.)
27 SAF
species :
Isoetes stephanseniae Duthie; Trans. R. Soc. S. Afr. 17: 330 (1929)
Distribution: South Africa (W-Cape Prov.)
27 SAF
Synonyms: hidden
= Isoetes capensis var. stephansenii (A.V.Duthie) Schelpe & N.C.Anthony, Bothalia 15(3-4): 555 (1985)
species :
Isoetes stevensii J.R.Croft; Blumea 26: 177-190 (1980)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: IUCN R
species :
Isoetes storkii T.C.Palmer; Amer. Fern J. 22(4): 136 (1932)
Distribution: Costa Rica; Panama
80 COS PAN
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes montana T.C.Palmer, Amer. Fern J. 21(4): 134 (1931) (nom. illeg., non Isoetes montana U. Weber (1922))
= Isoetes tryoniana L.D.Gómez, Rev. Biol. Trop. 17(1): 108, f. 8 (1972)
species :
Isoetes taiwanensis De Vol; Taiwania 17(1): 1-7, f. 1 (1972)
Distribution: Taiwan
38 TAI
Conservation Status: IUCN E (extremely rare, only in one single pond)
variant :
var. kinmenensis F.Y.Lu, H.H.Chen & Y.L.Hsueh; Flora of Kinmen 26 (2011)
Distribution: Taiwan
38 TAI
Conservation Status: IUCN E (extremely rare)
species :
Isoetes tamaulipana Mora-Olivo, A.Mend. & Mart.-Aval.; Phytotaxa 267(2): 115 (2016)
Distribution: Mexico (Tamaulipas)
79 MEX
species :
Isoetes tegetiformans Rury; Amer. Fern J. 68(4): 100 (1978)
Name: Mat-forming Merlin´s-grass
Distribution: USA (N-Georgia)
78 USA
Conservation Status: IUCN E. Extremely rare, known from only a few vernal pools on granite outcrops
species :
Isoetes tennesseensis Luebke & Budke; Amer. Fern J. 93(4): 184-190 (2003)
Name: Tennessee Quillwort
Distribution: USA (Tennessee)
78 USA
species :
Isoetes tenuifolia Jermy; J. Hall, J. Linn. Soc. Bot. 64: 118 (1971)
Distribution: Ghana; ?Kenya
22 GHA
Conservation Status: IUCN R (Ghana), E (Kenya)
species :
Isoetes texana Singhurst, Rushing & W.C.Holmes; Phytoneuron 22: 1 (2011)
Distribution: USA (Texas)
77 USA
species :
Isoetes tiguliana Gennari; Comment. Soc. Critt. Ital. 1 (1): 42 (1861)
Distribution: Italy; Morocco; Algeria; Tunisia
13 ITA 20 ALG MOR TUN
Remarks: bona sp. fide Troia et al. 2016
Synonyms: hidden
= Calamaria dubia (Gennari) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Calamaria tegulensis (Genn.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes dubia Gennari, Comment. Soc. Critt. Ital. 1: 104 (1861) (basion.)
= Isoetes tegulensis f. submersa Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 20 (1936)
= Isoetes tegulensis f. terrestris Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 20 (1936)
= Isoetes tegulensis Gennari, Comment. Soc. Critt. Ital. 1: 106 (1861) (sept.) (nom. illeg.)
= Isoetes velata ssp. dubia (Gennari) Batt. & Trab., Fl. Syn. 407 (1902)
= Isoetes velata ssp. tegulensis (Gennari) Batt. & Trab., Fl. Alg. & Tunis. 407 (1861), et Fl. Syn. 407 (1905)
= Isoetes velata var. tegulensis (Gennari) Fiori & Paol., Fl. Anal. Ital. 1: 25 (1896)
= Isoetes velata var. tegulensis b.maculosa Gennari, Ann. Agric. Sci. 11: 175 (1865)
= Isoetes velata var. tegulensis c.emersa Gennari, Giorn. Bot. Ital. 2: 96 (1870)
= Isoetes velata var. tegulensis Fiori, Fiori & Paol., Fl. Anal. Ital. 1: 25 (1896)
species :
Isoetes todaroana Troìa & Raimondo; Amer. Fern J. 99(4): 238 (2009 publ. 2010)
Distribution: Sicily; Greece; S-Italy
13 GRC ITA SIC
Synonyms: hidden
= Isoetes iapygia Ernandes, Beccarisi & V.Zuccarello, Pl. Biosystems 144(4): 807 (2010) (fide Troia & Greuter 2015)
species :
Isoetes toximontana Musselman & J.P.Roux; Novon 12: 504-507 (2002)
Distribution: South Africa (N-Cape Prov., W-Cape: Gifberg to Kamiesberg)
27 SAF
species :
Isoetes transvaalensis Jermy & Schelpe; Contr. Bolus Herb. 10: 150 (1982)
Distribution: South Africa (Limpopo, Gauteng, Mpulamanga, E-Cape Prov., Free State, KwaZulu-Natal); Lesotho
27 LES SAF
species :
Isoetes triangula U.Weber; Hedwigia 63: 253, f. 32, 33 (1922)
Distribution: N-Brazil (Amazonas); Venezuela (Amazonas, Bolívar, Cojedes, Guarico); Guyana; French Guiana; Mexico (Oaxaca)
79 MEX 82 FRG GUY VEN 84 BRA
Synonyms: hidden
= Isoetes schinzii H.P.Fuchs, (1960), nom. nud.
species :
Isoetes tripus A.Braun; Herb. Vindob. ex Hook. (1863)
Distribution: Australia (SW-Western Australia)
50 AUS
Synonyms: hidden
= Calamaria tripus (A.Braun) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes phaeospora Durand, Bull. Soc. Bot. France 11: 103 (1864)
species :
Isoetes tuckermanii A.Braun ex Engelm.; A. Gray, Manual ed. 5: 676 (1867)
subspecies :
subsp. tuckermanii
Name: Tuckermans Quillwort
Distribution: USA (Connecticut, Delaware, Massachusetts, Maryland, Maine, New Hampshire, New Jersey, New York, Rhode Island, Vermont); Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec)
72 CDN 75 USA 78 USA
Remarks: inclusion of I. acadiensis fide Troia et al. 2016
Synonyms: hidden
= Calamaria tuckermanii (A.Braun ex Engelm.) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Isoetes tuckermanii var. borealis Eaton, Fern Bull. 6: 7 (1898)
subspecies :
subsp. acadiensis (Kott) D.F.Brunt.; Canad. Field-Naturalist 132(4): 366 (2018)
Distribution: E-Canada (Nova Scotia)
72 CDN
Synonyms: hidden
= Isoetes acadiensis Kott, Canad. J. Bot. 59: 2592 (1981)
species :
Isoetes tuerckheimii Brause; Urban, Symb. Ant. 7: 161 (1912)
Distribution: Hispaniola (Haiti, Santo Domingo)
81 HIS
species :
Isoetes udupiensis P.K.Shukla, G.K.Srivast., S.K.Shukla & P.K.Rajagopal; Taxon 54(1): 112 (2005)
Distribution: India (Karnataka)
40 IND
Remarks: bona sp. fide Fraser-Jenkins et al. 2017
Synonyms: hidden
= Isoetes divyadarshanii P.K.Shukla, G.K.Srivast. & S.K.Shukla, Taxon 54(1): 113 (2005)
species :
Isoetes ulei Weber; Hedwigia 63: 255, fig. 40-41 (1922)
Distribution: Brazil (Rio de Janeiro)
84 BRA
Synonyms: hidden
= Isoetes goebelii Weber, Hedwigia 63: 247, fig. 16-17 (1922)
species :
Isoetes valida (Engelm.) Clute; Fern-Allies 236 (1905)
Name: Strong quillwort
Distribution: USA (Alabama, Delaware, Georgia, North Carolina, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)
75 USA 78 USA
Synonyms: hidden
= Isoetes engelmannii var. valida Engelm., Manual (Gray), ed. 5. 677 (1867)
species :
Isoetes vanensis M.Keskin & G.Zare; Phytotaxa 269(4): 295 (2016)
Distribution: Turkey (E-Anatolia)
34 TUR
species :
Isoetes vermiculata Hickey; Ann. Missouri Bot. Gard. 76(4): 1160 (1989)
Distribution: Venezuela (Amazonas)
82 VEN
species :
Isoetes virginica N.Pfeiff.; Claytonia 3: 29 (1937) (fide Musselman 2003)
Name: Piedmont quillwort
Distribution: SE-USA (Alabama, Georgia, North Carolina, South Carolina, Virginia)
78 USA
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes piedmontana (N.Pfeiff.) C.F.Reed, Phytologia 12: 392 (1965)
= Isoetes virginica var. piedmontana N.Pfeiff., Bull. Torrey Bot. Cl. 66: 411 (1939)
species :
Isoetes viridimontana M.A.Rosenthal & W.C.Taylor; Amer. Fern J. 104(1): 8 (2014)
Distribution: USA (Vermont)
75 USA
species :
Isoetes weberi Herter ex Weber; Hedwigia 63: 255-256, fig. 44-45 (1922)
Distribution: S-Brazil (Rio Grande do Sul); Uruguay (Rocha)
84 BRA 85 URU
species :
Isoetes welwitschii A.Braun ex Kuhn; Fil. Afr. 196 (1868)
Distribution: Ghana; Nigeria; Central African Republic; Sudan; Eritrea; Ethiopia; Kenya; Tanzania; Angola; Zambia; Botswana; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal); Madagascar
22 GHA NGA 23 CAF 24 ERI ETH SUD 25 KEN TAN 26 ANG ZAM 27 BOT SAF 29 MDG
Synonyms: hidden
= Isoetes natalensis Baker, Handb. Fern-Allies 132 (1887)
= Calamaria natalensis (Baker) Kuntze, Rev. Gen. Pl. 2: 828 (1891)
= Calamaria welwitschii (A.Braun ex Kuhn) Kunze, Rev. Gen. Pl. 2: 828 (1883)
species :
Isoetes wormaldii R.Sim; Trans. S. Afr. Phil. Soc. 16: 299, t. 5 (1906)
Distribution: South Africa (E-Cape: Grahamstown to East London)
27 SAF
Conservation Status: IUCN E
species :
Isoetes yunguiensis C.F.Wang & W.C.Taylor; Novon 12 (4): 587-591 (2002)
Distribution: China (Guizhou, Yunnan)
36 CHI
species :
Isoetes ×altonharvillii Musselman & R.D.Bray; Castanea 60(3): 246 (1995)
Distribution: USA (Delaware, Virginia)
78 USA
Remarks: Isoetes caroliniana x engelmannii
species :
Isoetes ×blondeaui D.F.Brunt. & P.C.Sokoloff; Botany (Ottawa) 98(10): 558 (2020)
Distribution: Canada (Quebec)
72 CDN
species :
Isoetes ×brittonii D.F.Brunt. & W.C.Taylor; Amer. Fern J. 80: 82-89 (1990)
Distribution: USA (Connecticut, New Jersey, New York, Pennsylvania, South Carolina, Virginia)
75 USA 78 USA
Remarks: Isoetes engelmanii x Isoetes riparia
species :
Isoetes ×bruntonii Knepper & Musselman; Amer. Fern J. 86(1): 9 (1996)
Distribution: USA (Virginia)
78 USA
Remarks: Isoetes engelmannii x hyemalis
species :
Isoetes ×carltaylori Musselman., R.D.Bray & D.A.Knepper; Canad. J. Bot. 75 (2): 301-309 (1997)
Distribution: USA
Remarks: Isoetes acadiensis x Isoetes engelmannii
species :
Isoetes ×dodgei A.A.Eaton; Fern Bull. 6: 6 (1898)
Distribution: USA (New Hampshire, New Jersey, New York, Pennsylvania, Vermont); Canada (Ontario, Québec)
72 CDN 75 USA
Remarks: Isoetes echinospora x Isoetes septentrionalis
Synonyms: hidden
= Isoetes canadensis A.A.Eaton, Proc. U. S. Nat. Herb. 23(1226): 650 (1901)
= Isoetes riparia f. canadensis (Engelm.) Proctor, Amer. Fern J. 39: 119 (1949)
= Isoetes riparia var. canadensis Engelm., Trans. Acad. Sci. St. Louis 4: 383 (1882)
species :
Isoetes ×eatonii R.Dodge; Bot. Gaz. 23: 16, t. 4-5 (1897), pro sp.
Distribution: USA (Connecticut, Massachusetts, New Hampshire, New Jersey, New York, Pennsylvania, Vermont); Canada (Ontario)
72 CDN 75 USA
Remarks: Isoetes echinospora x engelmannii
Synonyms: hidden
= Isoetes eatoni R.Dodge, Ferns & Fern-Allies of New England 39 (1896)
= Isoetes ×gravesii Eaton, Fernwort Papers 2: 14 (1900)
= Isoetes eatoni f. gravesii (Eaton) C.F.Reed, Amer. Fern J. 35: 82 (1945)
= Isoetes eatoni var. gravesii (Eaton) Clute, Fern Bull. 16: 55 (1908)
= Isoetes valida var. gravesii (Eaton) Clute, Fern-Allies 243 (1905)
species :
Isoetes ×echtuckeri D.F.Brunt. & D.M.Britton; Canad. J. Bot. 77 (11): 1662-1668 (1999)
Distribution: USA (Maine); Canada (New Brunswick, Nova Scotia, Ontario, Québec)
72 CDN 75 USA
species :
Isoetes ×fernaldii D.F.Brunt. & P.C.Sokoloff; Webbia 75(1): 44 (2020)
Distribution: USA (Virginia)
78 USA
species :
Isoetes ×foveolata A.A.Eaton ex R.Dodge; Ferns & Fern-Allies of New England 38 (1896)
Distribution: USA (Connecticut, Massachusetts, New Hampshire); Canada (Ontario)
72 CDN 75 USA
Remarks: Isoetes engelmannii x Isoetes tuckermanii
species :
Isoetes ×gopalkrishnae Sarv.K.Singh, P.K.Shukla & N.K.Dubey; Phytotaxa 362(1): 71 (2018)
Distribution: India (Uttar Pradesh)
40 IND
species :
Isoetes ×harveyi A.A.Eaton; Fernwort Pap. 11 (1900)
Distribution: USA (Maine); Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario)
72 CDN 75 USA
Remarks: Isoetes lacustris x Isoetes tuckermanii
Synonyms: hidden
= Isoetes tuckermanii var. harveyi (A.A.Eaton) Clute, Fern Allies 260 (1905)
species :
Isoetes ×herb-wagneri W.C.Taylor; Amer. Fern J. 92(2): 162 (2002)
Distribution: USA (Montana)
73 USA
Remarks: Isoetes echinospora x bolanderi
species :
Isoetes ×heterospora Eaton; Fernwort Papers, 2: 8 (1900), pro sp.
Distribution: USA (New York)
75 USA
Remarks: Isoetes lacustris x tuckermannii
Synonyms: hidden
= Isoetes tuckermani var. heterospora (Eaton) Clute, Fern-Allies 226, 260 (1905)
= Isoetes macrospora var. heterospora Eaton, Rhodora 10: 42 (1908)
= Isoetes macrospora f. heterospora (Eaton) Clute, Fern Bull. 16: 56 (1908)
species :
Isoetes ×hickeyi W.C.Taylor & Luebke; Amer. Fern J. 78(1): 6 (1988)
Distribution: USA (Wisconsin); Canada (Ontario, Saskatchewan); Scotland
10 GRB 71 CDN 72 CDN 74 USA
Remarks: Isoetes echinospora x Isoetes lacustris
species :
Isoetes ×jeffreyi D.M.Britton & D.F.Brunt.; Canad. J. Bot. 70: 447-452 (1992)
Distribution: Canada (Ontario, Québec)
72 CDN
Remarks: Isoetes lacustris x Isoetes riparia
species :
Isoetes ×kareniae D.F.Brunt. & P.C.Sokoloff; Webbia 75(1): 38 (2020)
Distribution: USA (Pennsylvania, Virginia)
75 USA 78 USA
species :
Isoetes ×marensis D.M.Britton & D.F.Brunt.; Canad. J. Bot. 73(9): 1352 (1995)
Distribution: Canada (British Columbia)
71 CDN
Remarks: Isoetes howellii x maritima
species :
Isoetes ×michinokuana M.Takamiya, M.Watanaba & K.Ono; Acta Phytotax. Geobot. 48(2): 109 (1997 publ. 1998)
Distribution: Japan
38 JAP
Remarks: Isoetes japonica x pseudojaponica
species :
Isoetes ×novae-angliae D.F.Brunt. & D.M.Britton; Rhodora 108(935): 238 (2006)
Distribution: USA (Massachusetts)
75 USA
Remarks: Isoetes septentrionalis x tuckermannii
Synonyms: hidden
= Isoetes riparia var. robbinsii (Eaton) Proctor, Amer. Fern J. 39: 119 (1949)
= Isoetes canadensis var. robbinsii A.A.Eaton, Rhodora 5: 279 (1903)
= Isoetes dodgei var. robbinsii Eaton, Rhodora 10: 42 (1908)
= Isoetes chapmanii var. robbinsii (Eaton) M.Broun, Ind. N. Amer. Ferns 199 (1938)
species :
Isoetes ×paratunica D.F.Brunt., Mochalova & A.A.Bobrov; Amer. Fern J. 105(2): 109, figs. 5-7 (2015)
Distribution: European Russia
14 RUS
species :
Isoetes ×pseudotruncata D.M.Britton & D.F.Brunt.; Canad. J. Bot. 74(1): 58 (1996)
Distribution: Alaska; Canada (Alberta, British Columbia)
70 ASK 71 CDN
Remarks: Isoetes echinospora x maritima
species :
Isoetes ×robusta (Engelm.) D.F.Brunt.; Rhodora 120(984): 307 (2019)
Distribution: USA; Canada
Synonyms: hidden
= Isoetes braunii f. robusta (Engelm.) N.Pfeiff., Ann. Missouri Bot. Gard. 9: 173 (1922)
= Isoetes echinospora f. robusta (Engelm.) Clute, Fern Allies 258 (1905)
= Isoetes echinospora var. robusta Engelm., Trans. Acad. Sci. St. Louis 4: 380 (1882)
= Isoetes muricata f. robusta (Engelm.) C.F.Reed, Amer. Fern J. 35: 82 (1945)
= Isoetes tenella f. robusta (Engelm.) Rothm., Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)
species :
Isoetes ×truncata (A.A.Eaton) Clute; Fern-Allies 260 (1905)
Name: Vancouver Quillwort
Distribution: SE-Alaska; Canada (Alberta)
70 ASK 71 CDN
Remarks: Isoetes maritima x Isoetes occidentalis
Synonyms: hidden
= Isoetes echinospora var. truncata A.A.Eaton, Working List North American Pteridophytes 27 (1901)
= Isoetes muricata ssp. truncata (A.A.Eaton) H.P.Fuchs, Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982)
...

Lilium philadelphicum (Liliaceae) (Photo: M. Hassler)