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Ver. 18.4
Feb. 11th, 2024

002.0000 Family Isoetaceae Rchb.; Bot. Damen: 309 (1828)
1 genus, ca. 190 spp.
Species listed in database: 204; hybrids: 27
= Bothriaceae Dulac; Fl. Dépt. Hautes-Pyrenees 38 (1867)
= Isoetineae Engl.; Syllabus, ed. 2: 65 (1898)
= Isoetoideae A.Gray; Bot. Text, ed. 5: 502 (1858)
002.0001 Genus Isoetes L.; Sp. Pl. 2: 1100 (1753)
Distribution: Worldwide. Many undescribed species remain (for China see Gu et al. 2023). Latest checklist: Troia et al. (2016). Taxonomy especially in North America and in the Mediterranean is quite disputed.
Species listed in database: 204; hybrids: 27
= Calamaria Boehm.; Def. Gen. Pl. ed. Boehmer 500 (1760)
= Cephaloceraton Gennari; Comment. Soc. Critt. Ital. 1: 111 (1862)
= Isoetella Gennari; Comment. Soc. Critt. Ital. 1: 114 (1862)
= Sivetes Raf.; Med. Repos. 11. 5: 358 (1808)
= Stylites Amstutz; Ann. Missouri Bot. Gard. 44: 121 (1957)
= Subularia Ray; Syn. Meth. Stirp. Brit., 283 (1696) (Pre-Linnaean)
species :
Isoetes abyssinica Chiov.; Atti Soc. Naturalisti Mat., Modena 64: 45-47 (1933)
Distribution: Ghana; Ethiopia; Eritrea; Kenya; Tanzania
22 GHA 24 ERI ETH 25 KEN TAN
species :
Isoetes aemulans J.P.Roux; Bothalia 43(2): 218 (2013)
Distribution: South Africa (Free State, Mpulamanga, KwaZulu-Natal)
27 SAF
species :
Isoetes aequinoctialis Welw. ex A.Braun; Sitz. Ges. Naturf. Freunde Berlin 7 (1867)
Distribution: Mali; Ghana; Niger; Central African Republic; Tanzania; Angola; Zambia; Zimbabwe; Namibia; South Africa (N-Cape Prov., Mpulamanga); Eswatini [Swaziland]
22 GHA MLI NGR 23 CAF 25 TAN 26 ANG ZAM ZIM 27 NAM SAF SWZ
Synonyms: hidden
= Calamaria aequinoctialis (Welw. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes erongensis Wannt.; Svensk Bot. Tidsskr. 64(2): 150-152 (1970)
species :
Isoetes afloramientorum D.F.Brunt. & J.B.S.Pereira; Phytotaxa 451(3): 219 (2020)
Distribution: Bolivia (Santa Cruz)
83 BOL
species :
Isoetes alcalophila Halloy; Lilloa 35(2): 68 (1980)
Distribution: Argentina (Catamarca, Tucuman)
85 ARG
species :
Isoetes alpina Kirk; Trans. & Proc. New Zealand Inst. 7: 377, t. 25 (1875)
Distribution: New Zealand (South Isl., North Isl.)
51 NZL
Synonyms: hidden
= Calamaria alpina (Kirk) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes kirkii var. alpina (Kirk) C.R.Marsden & Chinnock; comb. inval. (Thesis only)
= Isoetes multiangularis Colenso; Trans. & Proc. New Zealand Inst. 22: 449 (1890)
species :
Isoetes alstonii C.F.Reed & Verdc.; Kirkia 5: 19 (1965)
Distribution: Tanzania; S-Zambia; Zimbabwe; Mozambique; N-Namibia; Madagascar
25 TAN 26 MOZ ZAM ZIM 27 NAM 29 MDG
species :
Isoetes amazonica A.Braun; Baker, J. Bot. 18: 109 (1880)
Distribution: Brazil (Pará)
84 BRA
Synonyms: hidden
= Calamaria amazonica (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes anamariae J.B.C.Pereira; Amer. Fern J. 112(3): 194-199 (2022)
Distribution: Brazil (Bahia, Pernambuco)
84 BRA
species :
Isoetes anatolica Prada & Rolleri; Bot. J. Linn. Soc. 147(2): 215 (2005)
Distribution: Turkey (Anatolia)
34 TUR
species :
Isoetes andicola (Amstutz) L.D.Gómez; Brenesia 18: 4 (1980)
Distribution: Peru; Bolivia (La Paz)
83 BOL PER
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes andicola (Amstutz) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 177 (1981), isonym
= Isoetes andicola var. gemmifera (Rauh) L.D.Gómez; Brenesia 18: 4 (1980)
= Stylites andicola Amstutz; Ann. Missouri Bot. Gard. 44: 121 (1957)
= Stylites gemmifera Rauh; Sitzungsber. Heidelberg Akad. Wiss. Math. Naturwiss. Kl. 1959: 11 (1959)
species :
Isoetes andina Spruce ex Hook.; Brit. Ferns ad t. 55 (1861)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, Magdalena, Meta, Nariño, Putumayo, Risaralda); Venezuela (Merida, Tachira); Ecuador; Peru
82 CLM VEN 83 ECU PER
Synonyms: hidden
= Calamaria triquetra (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1893)
= Isoetes andina Spruce; Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 325 (1883), nomen
= Isoetes lechleri var. triquetra (A.Braun) L.D.Gómez; Brenesia 18: 5 (1980)
= Isoetes triquetra A.Braun; Verh. Bot. Ver. Brandenb. 4: 332 (1862)
species :
Isoetes appalachiana D.F.Brunt. & D.M.Britton; Rhodora 99: 118-133 (1997), nom. nov.
Name: Appalachian quillwort
Distribution: USA (Alabama, Delaware, Florida, Georgia, Kentucky, New Jersey, North Carolina, Pennsylvania, South Carolina, Tennessee, Virginia)
75 USA 78 USA
Synonyms: hidden
= Isoetes engelmannii f. fontana (A.A.Eaton) Clute; Fern-Allies 258 (1905)
= Isoetes engelmannii f. georgiana (Engelm.) Clute; Fern-Allies 258 (1905)
= Isoetes engelmannii var. fontana A.A.Eaton; Fern Bull. 13: 52 (1905)
= Isoetes engelmannii var. georgiana Engelm.; Trans. Acad. Sci. St. Louis 4: 384 (1882)
species :
Isoetes araucaniana Macluf & Hickey; Amer. Fern J. 97(4): 220 (2008)
Distribution: Chile (Biobío, Araucanía)
85 CLE
Synonyms: hidden
= Isoetes favulata Hickey; nom. nud.
species :
Isoetes asiatica (Makino) Makino; Bot. Mag. (Tokyo) 28: 184 (1914)
Distribution: Russian Far East (Kamchatka, Kuril Isl., Primorye, Sakhalin); ?China; Japan
31 RFE 38 JAP
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes echinospora subsp. asiatica (Makino) Á.Löve; Amer. Fern J. 52: 121 (1962)
= Isoetes echinospora var. asiatica Makino; Bot. Mag. (Tokyo) 18: 129 (1904) ex Iversen, Dansk Bot. Ark. 5(23): 2 (1928)
= Isoetes muricata subsp. asiatica (Makino) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982)
= Isoetes setacea subsp. asiatica (Makino) Holub; Acta Horti Bot. Prag. 1962: 55 (1962)
species :
Isoetes attenuata C.R.Marsden & Chinnock; Fl. Australia 48: 704 (1998)
Distribution: Australia (SE-South Australia)
50 AUS
species :
Isoetes australis S.Williams; Proc. Roy. Soc. Edinb. 62B(l). 1-8 (1943)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes azorica Durieu; Milde, Fil. Eur. 278 (1867)
Name: Azoren-Brachsenkraut
Distribution: Azores (Terceira, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.)
21 AZO
Conservation Status: IUCN E
Synonyms: hidden
= Calamaria azorica (Durieu) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes baculata Hickey & H.P.Fuchs; Syst. Bot. 11(2): 310 (1986)
Distribution: Colombia (Vaupés); Brazil (Amazonas)
82 CLM 84 BRA
species :
Isoetes baodongii Y.F.Gu, Y.H.Yan & Yi Jun Lu; Novon 29: 207 (2021)
Distribution: China (Zhejiang)
36 CHI
species :
Isoetes biafrana Alston; Bol. Soc. Broter. Ser. 2A, 30: 15 (1956)
Distribution: Benin; Cameroon; Bioko Isl. [Fernando Poo]
22 BEN 23 BIO CMN
species :
Isoetes bischlerae H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(1): 113 (1982)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Nariño)
82 CLM
Remarks: provisionally accepted name
species :
Isoetes bolanderi Engelm.; Amer. Naturalist 8: 214 (1874)
Name: Bolander´s Quillwort
Distribution: Canada (Alberta); USA (Arizona, California, Colorado, Idaho, Montana, New Mexico, Nevada, Oregon, Utah, Washington State, Wyoming)
71 CDN 73 USA 76 USA 77 USA
Remarks: not in Florida, obviously a BONAP error
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria bolanderi (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Calamaria pygmaea (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes bolanderi f. pygmaea (Engelm.) Clute; Fern Bull. 16: 54 (1908)
= Isoetes bolanderi var. parryi Engelm.; Amer. Naturalist 8: 214 (1874)
= Isoetes bolanderi var. pygmaea (Engelm.) Clute; Fern-Allies 228, 258 (1905)
= Isoetes bolanderi var. sonnei L.F.Hend.; Bull. Torrey Bot. Club 27(6): 358 (1900)
= Isoetes californica Engelm.; Gray´s Manual, ed. 5: 677 (1867)
= Isoetes parryi Engelm.; Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 338 (1883)
= Isoetes pygmaea Engelm.; Amer. Naturalist 8: 214 (1874)
species :
Isoetes boliviensis U.Weber; Hedwigia 63: 247, fig. 14-15 (1922)
Distribution: Peru; Bolivia (Cochabamba, La Paz)
83 BOL PER
species :
Isoetes boomii Luebke; Amer. Fern J. 82: 23 (1992)
Name: Boom´s quillwort
Distribution: USA (Florida, Alabama, C-Georgia: Laurens Co., Worth Co.)
75 USA 78 USA
Conservation Status: IUCN E
species :
Isoetes boryana Durieu; Bull. Soc. Bot. France 8: 164 (1861)
Distribution: +Spain; SW-France
12 FRA SPA
Conservation Status: IUCN V
Synonyms: hidden
= Calamaria boryana (Durieu) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes boryana subsp. brevis Gand.; Fl. Eur. 27: 195 (1891)
= Isoetes boryana subsp. condensata Gand.; Fl. Eur. 27: 195 (1891)
= Isoetes boryana subsp. frigida Gand.; Fl. Eur. 27: 195 (1891)
= Isoetes boryana subsp. gredensis Gand.; Fl. Eur. 27: 195 (1891)
= Isoetes boryana subsp. legitima Gand.; Fl. Eur. 27: 195 (1891)
= Isoetes boryana subsp. praetervisa Gand.; Fl. Eur. 27: 195 (1891)
= Isoetes variabilis proles boryana (Durieu) Rouy; Rouy & Foucaud, Fl. France 14: 477 (1913), nom. illeg.
= Isoetes variabilis subsp. boryana (Durieu) P.E.Fourn.; Quatre Fl. France: 5 (1934), nom. illeg.
= Isoetes velata var. boryana (Durieu) M.I.Romero & Real; Bot. J. Linn. Soc. 148(4): 464 (2005)
species :
Isoetes boyacensis H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(1): 107 (1982)
Distribution: Colombia (Boyacá, Cundinamarca, Meta, Nariño, Santander)
82 CLM
Remarks: provisionally accepted name
species :
Isoetes bradei Herter; Revista Sudamer. Bot. 8: 19 (1949)
Distribution: Brazil (São Paulo, ?Minas Gerais)
84 BRA
Conservation Status: considered extinct
species :
Isoetes brasiliensis H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 27 (1986)
Distribution: Brazil (Santa Catarina, Rio Grande do Sul)
84 BRA
Synonyms: hidden
= Isoetes reitzii H.P.Fuchs; (1979), nom. nud. pro syn.
species :
Isoetes brevicula E.R.L.Johnson; J. Roy. Soc. W. Australia 67(1): 34 (1984)]
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes butleri Engelm.; Bot. Gaz. 3: 1 (1878)
Name: Limestone quillwort
Distribution: USA (Alabama, Arkansas, Georgia, Illinois, Kansas, Kentucky, Missouri, Oklahoma, Tennessee, Texas)
74 USA 75 USA 77 USA 78 USA
Synonyms: hidden
= Calamaria butleri (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes butleri f. immaculata (Engelm.) Clute; Fern-Allies 258 (1905)
= Isoetes butleri var. immaculata Engelm.; Trans. Acad. Sci. St. Louis 4: 388 (1882)
species :
Isoetes cangae J.B.S.Pereira, Salino & Stützel; Phytotaxa 272(2): 145 (2016)
Distribution: Brazil (Pará)
84 BRA
species :
Isoetes caparaoensis J.B.S.Pereira; Bot. J. Linn. Soc. 189(1): 15 (2018)
Distribution: Brazil (Minas Gerais, Espírito Santo)
84 BRA
species :
Isoetes capensis A.V.Duthie; Trans. Roy. Soc. South Africa 17: 330 (1929)
Distribution: South Africa (W-Cape Prov.)
27 SAF
species :
Isoetes caroli E.R.L.Johnson; J. Roy. Soc. W. Australia 67(1): 34 (1984)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes caroliniana (A.A.Eaton) Luebke; Amer. Fern J. 82: 26 (1992)
Distribution: SE-USA
78 USA
Remarks: bona species fide Troia et al. 2016, not a synonym of I. valida
Synonyms: hidden
= Isoetes engelmannii var. caroliniana A.A.Eaton; Fern Bull. 8: 60 (1900)
species :
Isoetes chubutiana Hickey, Macluf & W.C.Taylor; Amer. Fern J. 93(3): 130 (2003)
Distribution: Argentina (Chubut, Neuquen, Rio Negro); Chile (Biobío, Los Lagos, Aisén, Magallanes)
85 ARG CLE
Synonyms: hidden
= Isoetes savatieri subsp. valdiviensis H.P.Fuchs; (1992), nom. nud.
= Isoetes valdiviensis H.P.Fuchs; (1960), nom. nud.
= Isoetes meyeri H.P.Fuchs; (1982), nom. nud.
species :
Isoetes cipoensis J.B.S.Pereira; Syst. Bot. 47(2): 301 (2022)
Distribution: Brazil (Minas Gerais)
84 BRA
species :
Isoetes coreana Y.H.Chung & H.K.Choi; Korean J. Pl. Taxon 16(1): 2 (1986)
Distribution: North Korea; South Korea; Japan
38 JAP KOR NKO
Synonyms: hidden
= Isoetes sinensis var. coreana (Y.H.Chung & H.K.Choi) M.Takamiya, M.Watanaba & K.Ono; Acta Phytotax. Geobot. 48(2) (1997 publ. 1998)
species :
Isoetes coromandelina L.f.; Suppl. Pl. Syst. Veg. ed. II: 447 (1781)
subspecies :
subsp. coromandelina
Distribution: Nepal; India (Andhra Pradesh, Bihar, Chhattisgarh, Goa, Gujarat, Haryana, Jharkand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Odisha, Rajasthan, Tamil Nadu, Uttar Pradesh, West Bengal); Sri Lanka; Bangladesh; Myanmar; Thailand; Vietnam
40 BAN IND NEP SRL 41 MYA THA VIE
Synonyms: hidden
= Calamaria coromandelina (L.f.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes brachyglossa A.Braun; Ver. Bot. Ver. Brandenb. 4: 328 (1862), nom. nov. for I. capsularis Roxb. ex Griff. (fide Fraser-Jenkins 1997, 2017)
= Isoetes capsularis Roxb. ex Griff.; Posth. Papers, Cryptog. Pl. 572-575, t. 116-118 (1849), non Roxb. 1844
= Isoetes coromandelina subsp. brachyglossa (A.Braun) Panigrahi; Biol. Mem. 6(2): 131 (1981) (fide Fraser-Jenkins 1997)
= Isoetes coromandelina subsp. thanensis S.K.Shukla, Sarv.K.Singh, P.K.Shukla, N.K.Dubey, H.Khanam & G.K.Srivast.; Taiwania 62(2): 122 (2017)
= Isoetes coromandelina var. raipurensis Unni; J. Bombay Nat. Hist. Soc. 64: 590-592 (1967) (fide Fraser-Jenkins 1997)
= Isoetes debii P.K.Shukla, G.K.Srivast. & S.K.Shukla; Frontiers Bot. 4-6: 69-72 (2002), nom. inval.
= Isoetes debii S.C.Sinha; Frontiers Bot. 4-6: 69 (1992)
= Isoetes fuchsii Bhu, H.K.Goswami, U.S.Sharma & A.K.Bajpai; J. Advancem. Pl. Sci. 2(1)-1: 4 (2019)
= Isoetes fuchsii H.K.Goswami & U.S.Sharma; Bionature (Bhopal) 21: 11-18 (2001), nom. inval.
= Isoetes indica D.D.Pant & G.K.Srivast.; Proc. Nat. Inst. Sci. India 28 B (3) 246, t. XIV9, f. 16-259, t. XV, f. 30-35, t. XIV, f. 30-40 text-fig. 4 A-D, 5 A-H, 7 A-M, 12 (1962) (fide Fraser-Jenkins 1997)
= Isoetes indica var. harotiensis Bhardwaja & Gena; Aspects Plant Sci. 14: 495-499 (1992), nom. nud.
= Isoetes mirzapurensis Panigrahi & R.D.Dixit; Proc. Indian Sci. Congr. pt. III, Abstract, Botany 243 (1965) (fide Fraser-Jenkins 1997)
= Isoetes narsinghgarhensis H.K.Goswami & Mazumdar; J. Advancem. Pl. Sci. 2(1)-1: 5 (2019), nom. nov. for I. pantii var. hybrida Goswami
= Isoetes pantii H.K.Goswami & B.S.Arya; J. Indian Bot. Soc. 49: 30 (1970) (fide Fraser-Jenkins 1997)
= Isoetes pantii var. hybrida H.K.Goswami; Indian Fern J. 21(1-2): 152(-153) (2004), nom. inval. [fide Fraser-Jenkins et al. 2020]
= Isoetes semilocularis Sm.; Rees, Cycl. 19: 150 (1819)
= Isoetes tuberculata Gena & Bhardwaja; J. Bombay Nat. Hist. Soc. 81(1): 167 (1984) (fide Fraser-Jenkins 1997)
= Isoetes unilocularis Roxb. ex Sm.; Rees, Cycl. 19: no. 3 (1811)
subspecies :
subsp. macrotuberculata C.R.Marsden; Contr. Herb. Austral. 24: 2 (1976)
Distribution: Australia (Western Australia, Northern Territory, Queensland)
50 AUS
species :
Isoetes creussensis Lazare & S.Riba; J. Bot. Soc. Bot. France 50: 21 (2010)
Distribution: Andorra; S-France
12 ADO FRA
Remarks: status somewhat uncertain, near I. lacustris but cytologically separate
species :
Isoetes cristata C.R.Marsden & Chinnock; Fl. Australia 48: 704 (1998)
Distribution: Australia (Northern Territory)
50 AUS
Conservation Status: Known from type locality only
species :
Isoetes cubana Engelm. ex Baker; J. Bot. 18: 110 (1880)
Distribution: ?Mexico (?Yucatan); Belize; Cuba
80 BLZ 81 CUB
Synonyms: hidden
= Calamaria cubana (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes delalandei J.Lloyd; Notes Fl. Ouest France: 25 (1851)
Distribution: NW-France
12 FRA
Synonyms: hidden
= Isoetes histrix proles delalandei (J.Lloyd) Rouy; Rouy & Foucaud, Fl. France 14: 481 (1913)
= Isoetes histrix subsp. delalandei (J.Lloyd) P.E.Fourn.; Quatre Fl. France 6 (1934)
= Isoetes histrix var. delalandei (J.Lloyd) Rouy; Rouy & Foucaud, Fl. France 14: 481 (1913)
species :
Isoetes delilei (Bory) Rothm.; Feddes Repert. 54: 72 (1944)
Distribution: Portugal; W-Spain; SE-France; ?Corsica; ?Sardinia; Morocco; ?Algeria
12 FRA POR SPA 20 MOR
Synonyms: hidden
= Isoetes setacea Bosc ex Delile; Mem. Mus. Paris 14(110): 117, t. 6-7 (1827) (non Lam.)
= Isoetes setacea var. delilei Bory; Compt. Rend. Acad. Paris 18: 1165 (1844)
= Isoetes palustris var. setacea F.Muell.; Milde, Fil. Eur. & Atl. 9: 280 (1867)
= Isoetes montana U.Weber; Hedwigia 63: 254 (1922)
species :
Isoetes dispora Hickey; Fieldiana, Bot. 34: 93 (1994)
Distribution: Peru
83 PER
Conservation Status: Known from type only
species :
Isoetes dixitii Shende; J. Univ. Bombay 14: 50 (1945)
Distribution: India (Chhattisgarh, Karnataka, Madhya Pradesh, Maharashtra)
40 IND
Remarks: bona sp. fide Fraser-Jenkins et al. 2017)
Synonyms: hidden
= Isoetes bilaspurensis Panigrahi; Biol. Mem. 6(2): 135, f. 1-10 (1981)
species :
Isoetes drummondii A.Braun; Monatsber. K. Akad. Berl. 593 (1863)
subspecies :
subsp. drummondii
Distribution: Australia (SW-Western Australia, SE-South Australia, SE-Queensland, Victoria, NE-Tasmania)
50 AUS
Synonyms: hidden
= Calamaria drummondii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
subspecies :
subsp. anomala C.R.Marsden & Chinnock; Muelleria 8(1): 58 (1993)
Distribution: Australia (SE-South Australia, SW-Western Australia, Victoria, S-New South Wales)
50 AUS
species :
Isoetes dubsii J.B.S.Pereira; PhytoKeys 131: 59 (2019)
Distribution: Brazil (Mato Grosso do Sul)
84 BRA
species :
Isoetes duriei Bory; Compt. Rend. Acad. Paris 18: 1166 (1844) (durieai auct., durieui auct.)
Name: Durieu-Brachsenkraut (DE)
Distribution: Portugal; Spain; Baleares; France; Corsica; Sardinia; Italy; Sicily; Pantelleria; Malta; Greece (Evvoia, Sporades, Kyklades); Crete (rare); ?Morocco; Algeria; Tunisia; East Aegaean Isl. (Lesvos, Rhodos, Karpathos, Samothraki); Turkey (Anatolia)
12 BAL COR FRA POR SAR SPA 13 GRC ITA KRI MLT PTL SIC 20 ALG TUN 34 EAI TUR
Synonyms: hidden
= Calamaria durieui (Bory) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetella duriei (Bory) Gennari; Comment. Soc. Critt. Ital. 1(3): 115 (1862)
= Isoetes duriaei subsp. balearica Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. felix Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. insularis Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. kabylica Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. leptophylla Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. ligustica De Not.; Kunze, Ind. Fil. Cult., 51 (1850)
= Isoetes duriaei subsp. litorea Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. lusitanica Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. maritima Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes duriaei subsp. sardoa Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes ligustica De Not. ex A.Braun; Monatsber. K. Akad. Berl., Sitzung 7 Dec (1863)
= Isoetes tridentata Durieu; Kuhn, Filic. Afr. 195 (1867)
species :
Isoetes echinospora Durieu; Bull. Soc. Bot. France 8: 164 (1861)
Name: Igelsporiges Brachsenkraut, Stachelsporiges Brachsenkraut (DE), Spiny-spore quillwort (EN)
Distribution: Iceland; England; Scotland; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Germany (Baden-Württemberg, Schleswig-Holstein+); Switzerland; Austria; Poland; Czech Republic; Slovakia; Spain; Andorra; S-France; Italy; ?Romania; Bulgaria; Estonia; Latvia; European Russia (N-European Russia, C-European Russia); Morocco; Siberia (Buryatia, Chita, Irkutsk, W-Siberia); Alaska; Greenland; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); St. Pierre et Miquelon; USA (California, Colorado, Connecticut, Delaware, Idaho, Massachusetts, Maine, Michigan, Minnesota, Montana, New Hampshire, New Jersey, New York, Ohio, Oregon, Pennsylvania, Rhode Island, Utah, Vermont, Washington State, Wisconsin, Wyoming)
10 DEN FIN GRB ICE IRE NOR SWE 11 AUT CZE GER NET POL SLO SWI 12 ADO FRA SPA 13 BUL ITA 14 EST LAT RUS 20 MOR 30 SIB 70 ASK CDN GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 76 USA 78 USA
Synonyms: hidden
= Calamaria echinospora (Durieu) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes ambigua A.Braun; Engelm., Trans. Acad. Sci. St. Louis 4: 380 (1882)
= Isoetes boottii A.Braun; Engelm., Gray´s Manual, ed. 5, 676 (1867)
= Isoetes braunii Durieu; Bull. Soc. Bot. France 11: 101 (1864)
= Isoetes braunii f. astoma (Iversen) M.Broun; Index N. Amer. Ferns 99 (1938)
= Isoetes braunii f. oligostoma (Iversen) M.Broun; Index N. Amer. Ferns 99 (1938)
= Isoetes braunii f. polystoma (Iversen) M.Broun; Index N. Amer. Ferns 99 (1938)
= Isoetes echinospora subf. angustivelata Iversen; Bot. Tidsskr. 40: 128, fig. a-c (1928)
= Isoetes echinospora subf. lativelata Iversen; Bot. Tidsskr. 40: 129 (1928)
= Isoetes echinospora f. astoma Iversen; Bot. Tidsskr. 40: 129 (1928)
= Isoetes echinospora f. boottii (Engelm.) Clute; Fern Allies 258 (1905)
= Isoetes echinospora f. oligostoma Iversen; Bot. Tidsskr. 40: 129 (1928)
= Isoetes echinospora f. polystoma Iversen; Bot. Tidsskr. 40: 128 (1928)
= Isoetes echinospora subsp. admixta Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. belgica Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. borealis Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. centralis Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. discedens Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. echinospora Durieu; Gand., Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. fennica Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. gothica Gand.; 27: 194 (1891)
= Isoetes echinospora subsp. montana Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. muricata (Durieu) Á.Löve & D.Löve; Bot. Notis. 114: 49 (1961)
= Isoetes echinospora subsp. oligophylla Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. petropolitana Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. praebens Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. pulchella Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. pygmaea Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. remissa Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora subsp. vogesiaca Gand.; Fl. Eur. 27: 194 (1891)
= Isoetes echinospora var. boothii Clute; Frye, Ferns Northwest 39 (pro syn.) (1934)
= Isoetes echinospora var. boottii Engelm.; Manual (Gray), ed. 5: 676 (1867)
= Isoetes echinospora var. braunii (Durieu) Á.Löve & D.Löve; 49 (1961), isonym
= Isoetes echinospora var. braunii (Durieu) Engelm.; Gray´s Manual, ed. 5: 676 (1867)
= Isoetes echinospora var. brittoni Cockerell; Muhlenbergia 3: 9 (1907)
= Isoetes echinospora var. elatior Fliche; Mem. Acad. Stanisl. IV, 11 (1878): 182 (1879)
= Isoetes echinospora var. hesperia (C.F.Reed) Á.Löve; Amer. Fern J. 52: 121 (1962)
= Isoetes echinospora var. muricata (Durieu) Engelm.; Gray´s Manual, ed. 5, 676 (1867)
= Isoetes lacustris subsp. curvifolia (Casp.) Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. echinospora (Durieu) Hook.f.; Student´s Fl. Brit. Isl., ed. 1, 471 (1870)
= Isoetes lacustris subsp. echinospora (Durieu) Syme & N.E.Br.; 7 (1886), isonym
= Isoetes lacustris subsp. echinospora (Durieu) T.Moore & More; Contr. Cybele Hiber.: 381 (1866)
= Isoetes lacustris var. crassa Léman; Dict. Sci. Nat. 24: 24 (1822)
= Isoetes lacustris var. elatior (Fliche) Mer; Bull. Soc. Bot. France 28: 73 (1881)
= Isoetes lacustris var. longifolia Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)
= Isoetes lacustris var. minor (A.Braun ex Milde) Motelay & Vendryès; Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. recurvata Motelay & Vendryès; Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes muricata Durieu; Bull. Soc. Bot. France 11: 100 (1864)
= Isoetes muricata var. braunii (Durieu) C.F.Reed; Amer. Fern J. 35: 83 (1945)
= Isoetes muricata var. hesperia C.F.Reed; Amer. Fern J. 35: 84 (1945)
= Isoetes setacea subsp. monspeliensis Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes setacea subsp. muricata (Durieu) Holub; Acta Horti Bot. Prag. (1964)
= Isoetes setacea subsp. stricta Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes setacea subsp. turgida Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes setacea subsp. valida Gand.; Fl. Eur. 27: 196 (1891)
= Isoetes setacea subsp. vicina Gand.; Fl. Eur. 27: 195 (1891)
= Isoetes tenella f. astoma (Iversen) Rothm.; Feddes Repert. Regni Veg. 54: 72 (1944)
= Isoetes tenella f. iversenii Rothm.; Feddes Repert. Regni Veg. 54: 73 (1944)
= Isoetes tenella f. lativelata (Iversen) Rothm.; Feddes Repert. Regni Veg. 54: 72 (1944)
= Isoetes tenella f. oligostoma (Iversen) Rothm.; Feddes Repert. Regni Veg. 54: 73 (1944)
= Isoetes tenella f. polystoma (Iversen) Rothm.; Feddes Repert. Regni Veg. 54: 73 (1944)
= Isoetes tenella var. braunii (Durieu) Rothm.; Feddes Repert. 54: 73 (1944)
= Isoetes tenella var. oligostoma (Iversen) Rothm.; Feddes Repert. 54: 72 (1944)
species :
Isoetes ecuadoriensis Aspl.; Bot. Notis. 1925: 357, f. (1925)
Distribution: Ecuador
83 ECU
Synonyms: hidden
= Isoetes keeleyorum H.P.Fuchs; (1988), nom. nud.
species :
Isoetes ekmanii U.Weber; Hedwigia 63: 251 (1922)
Distribution: Argentina (Buenos Aires, Corrientes, Misiones)
85 ARG
species :
Isoetes elatior F.Muell. & A.Braun; Linnaea 25: 722 (1852)
Distribution: Australia (Tasmania)
50 AUS
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria elatior (F.Muell.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes tasmanica F.Muell.; Durieu, Bull. Soc. Bot. France 11: 104 (1864)
species :
Isoetes eludens J.P.Roux, Hopper & Rhian J.Sm.; Kew Bull. 64(1): 124 (2009)
Distribution: South Africa (N-Cape: Namaqualand)
27 SAF
Conservation Status: extremely local in a single 2 meter diameter rock pool
species :
Isoetes engelmannii A.Braun; Flora 29: 178 (1846)
Name: Engelmann´s quillwort
Distribution: Canada (Ontario); USA (Alabama, Arkansas, Connecticut, Delaware, Georgia, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Missouri, Mississippi, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, West Virginia)
72 CDN 74 USA 75 USA 78 USA
Synonyms: hidden
= Calamaria engelmannii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes engelmannii var. gracilis Engelm.; A. Gray, Manual, ed. 5: 677 (1867)
= Isoetes lacustris var. microspora Engelm.; A. Braun, Flora 29: 179 (1846)
= Isoetes ×fairbrothersii J.D.Montgom. & W.C.Taylor; Amer. Fern J. 84(4): 120 (1994)
species :
Isoetes escondidensis Halloy; Lilloa 35(2): 67 (1980)
Distribution: Argentina (Tucuman)
85 ARG
Conservation Status: IUCN E
Synonyms: hidden
= Isoetes castillonii H.P.Fuchs; (1965), nom. nud.
= Isoetes halloyi H.P.Fuchs; (1981), nom. nud.
= Isoetes filipovichii H.P.Fuchs; (1965), nom. nud.
species :
Isoetes eshbaughii Hickey & H.P.Fuchs; Syst. Bot. 11: 310 (1986)
Distribution: Bolivia (Cochabamba)
83 BOL
species :
Isoetes fengii Y.F.Gu & Y.H.Yan; Pl. Sci. J. 41(2): 168 (2023)
Distribution: China (Yunnan)
36 CHI
species :
Isoetes flaccida Shuttlew. ex A.Braun; Flora 29: 178 (1846)
Name: Southern quillwort
Distribution: USA (Alabama, Florida, Georgia)
78 USA
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria flaccida (Shuttlew) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes alata (N.Pfeiff.) Small; Ferns Florida: 22 (1932)
= Isoetes flaccida f. rigida (Engelm.) Clute; Fern-Allies 258 (1905)
= Isoetes flaccida var. alata N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 137 (1922)
= Isoetes flaccida var. rigida Engelm.; Trans. Acad. Sci. St. Louis 4: 386 (1882)
variant :
var. chapmanii Engelm.; Trans. Acad. Sci. St. Louis 4: 386 (1882)
Name: Chapman´s quillwort
Distribution: USA (Florida, Georgia)
78 USA
Synonyms: hidden
= Isoetes chapmanii (Engelm.) Small; Ferns of Florida 222 (1931 publ. 1932)
= Isoetes flaccida f. chapmanii (Engelm.) Clute; Fern-Allies 258 (1905)
species :
Isoetes fluitans M.I.Romero; Bot. J. Linn. Soc. 146(2): 233 (2004)
Distribution: Spain
12 SPA
species :
Isoetes fuliginosa R.L.Small & Hickey; Amer. Fern J. 91(2): 55 (-57, figs. 5-7, 14-15) (2001)
Distribution: Colombia (C-Colombia)
82 CLM
species :
Isoetes fuscomarginata H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetac.): 12 (1986)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes gardneriana Kunze ex Mett.; Fil. Lechl. Fasc. 11: 36 (1859)
Distribution: Brazil (Goiás, Mato Grosso do Sul, Tocantins, Bahia, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Calamaria gardneriana (Kuntze ex Mett.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes georgiana Luebke; Amer. Fern J. 82: 24 (1992)
Name: Georgia quillwort
Distribution: USA (Georgia)
78 USA
species :
Isoetes giessii Launert; Mitt. Bot. Staatssamml. München 2: 309 (1957)
Distribution: Namibia
27 NAM
Conservation Status: known from two locations
species :
Isoetes gigantea U.Weber; Nova Hedwigia 63: 245 (1922)
Distribution: Brazil (Bahia, Minas Gerais, Rio de Janeiro, Mato Grosso)
84 BRA
species :
Isoetes graniticola D.F.Brunt.; Rhodora 118(975): 269 (2016)
Distribution: USA (Alabama)
78 USA
species :
Isoetes gunnii A.Braun; Monatsber. K. Akad. Berl. 535 (1868)
Distribution: Australia (Tasmania)
50 AUS
Synonyms: hidden
= Calamaria gunnii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes lacustris (L.) Rodway; Tasm. Fl. 279 (1903)
species :
Isoetes gymnocarpa (Gennari) A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 555 (1864)
Name: Nacktsamiges Brachsenkraut; Naked-fruited Quillwort
Distribution: S-Corsica; Italy; Sicily; Greece (incl. Kiklades); ?Algeria; East Aegaean Isl. (Karpathos, Lesvos, Rhodos)
12 COR 13 GRC ITA SIC 34 EAI
Synonyms: hidden
= Cephaloceraton gymnocarpum Gennari; Comment. Soc. Critt. Ital. 1(3): 113 (1862)
= Cephaloceraton histrix var. subinerme Gennari; Comment. Soc. Crittog. Ital. 1: 112 (1862)
= Cephaloceraton pseudohystrix Gennari; Ann. Agric. Siciliana 11: 175 (1865)
= Isoetes histrix proles sicula (Tod.) Rouy; Rouy & Foucaud, Fl. France 14: 482 (1913)
= Isoetes histrix f. subinermis (Gennari) Hayek; Repert. Spec. Nov. Regni Veg. Beih. 30(1): 12 (1924)
= Isoetes histrix subsp. sicula (Tod.) P.E.Fourn.; Quatre Fl. France: 6 (1934)
= Isoetes histrix var. subinermis (Gennari) Durieu; Ces., Comp. Fl. It. 25 (1867), et Motelay & Vendryès, Actes Soc. Linn. Bord. 36: 4009, t. 17), f. 1-5 (1883)
= Isoetes histrix var. subinermis (Gennari) Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 400 (1883)
= Isoetes hystrix var. subinermis Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes pseudohystrix (Gennari) Levier ex Barbey; Fl. Sard. Comp.: 195 (1885)
= Isoetes sicula Tod.; Syn. Pl. acot. vasc. Sic. 251 (1866)
= Isoetes subinermis (Gennari) Cesca & Peruzzi; Fl. Medit. 11: 308 (2001)
species :
Isoetes habbemensis Alston; J. Arnold Arbor. 26: 180 (1945)
Distribution: New Guinea (Irian Jaya: Mt. Wilhelmina-Lake Habbema)
42 NWG
Conservation Status: IUCN R (twice found)
species :
Isoetes hallasanensis H.K.Choi, Ch.Kim & J.Jung; J. Pl. Biol. 51(5): 357 (2008)
Distribution: South Korea (Cheju Isl.)
38 KOR
species :
Isoetes hallei Eb.Fisch. & Lobin; Willdenowia 52(3): 330 (2022)
Distribution: W-D.R.Congo [Zaire]
23 ZAI
Remarks: known only from type locality
species :
Isoetes harleyi J.B.C.Pereira; Amer. Fern J. 112(3): 199-200 (2022)
Distribution: Brazil (Bahia)
84 BRA
species :
Isoetes haussknechtii Troia & Greuter; Willdenowia 45(3): 395 (2015)
Name: Haussknechts Brachsenkraut
Distribution: Greece; East Aegaean Isl. (Lesvos)
13 GRC 34 EAI
Remarks: this is "I. echinospora", "I. setacea" or "I. delilei" sensu auct. mult. from Greece. Distribution not yet completely known, may be more widespread in the Mediterranean
species :
Isoetes hawaiiensis W.C.Taylor & W.H.Wagner; Amer. Fern J. 83(2): 67 (1993)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
species :
Isoetes heldreichii Wettst.; Verh. K. K. Zool.-Bot. Ges. Wien 36: 239, t. 8 (1886)
Name: Heldreichs Brachsenkraut
Distribution: +C-Greece (+Palaiokastro near Karditsa)
13 GRC
Remarks: extinct fide Troia & Greuter 2015
species :
Isoetes hemivelata R.L.Small & Hickey; Amer. Fern J. 91(2): 57 (60, figs. 9-11, 16-17) (2001)
Distribution: Colombia (C-Colombia)
82 CLM
species :
Isoetes herzogii U.Weber; Hedwigia 63: 250 (1922)
Distribution: Peru (Cuzco); Bolivia (Cochabamba, La Paz, Santa Cruz)
83 BOL PER
species :
Isoetes hewitsonii Hickey; Fieldiana, Bot. 34: 94 (1994)
Distribution: Peru (Cajamarca)
83 PER
species :
Isoetes hieronymi U.Weber; Hedwigia 63: 251, fig. 24-25 (1922)
Distribution: Argentina (Cordoba, Rio Negro); Chile (Coquimbo)
85 ARG CLE
Synonyms: hidden
= Isoetes argentina A.A.Eaton; Palmer, Amer. Fern J. 19: 18 (1929), nom. nud.
= Isoetes lechleri N.Pfeiff.; Ann. Missouri Bot. Gard. 9(2): 138 (1922), non Mett.
species :
Isoetes histrix Bory & Durieu; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1167 (1844)
Name: Stacheliges Brachsenkraut
Distribution: S-England; Portugal; Spain; France; Channel Isl.; Corsica; Sardinia; Italy; Sicily; Malta; Macedonia; Albania; Greece (scattered, incl. Kyklades); Crete (W-Crete); Morocco; Algeria; Tunisia; East Aegaean Isl. (Lesvos, Ikaria, Rhodos, Karpathos); Turkey (Anatolia); Lebanon
10 GRB 12 CIS COR FRA POR SAR SPA 13 ALB GRC ITA KRI MLT NMA SIC 20 ALG MOR TUN 34 EAI LEB TUR
Synonyms: hidden
= Calamaria histrix (Bory) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Cephaloceraton histrix (Bory) Gennari; Comment. Soc. Crittog. Ital. 1: 111 (1862)
= Isoetes chaeturetii Mendes; Agron. Lusit. 23: 7 (1961) (fide Musselman 2003)
= Isoetes histrix subvar. armata Batt. & Trab.; Fl. Alg. & Tunis. 407 (1902)
= Isoetes histrix subvar. inermis Batt. & Trab.; Fl. Alg. & Tunis. 407 (1902)
= Isoetes histrix unranked grandis A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)
= Isoetes histrix unranked longispina A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 618 (1864)
= Isoetes histrix unranked nana A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)
= Isoetes histrix unranked solitaria A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 618 (1864)
= Isoetes histrix f. desquamata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)
= Isoetes histrix f. longispina (A.Braun) Fiori; Fl. Ital. Crypt. 5: 425 (1943)
= Isoetes histrix f. loricata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)
= Isoetes histrix var. desquamata (A.Braun) Batt. & Trab.; Fl. Algérie Tunisie: 407 (1905)
= Isoetes histrix var. desquamata f.longispina (A.Braun) Fiori; Fl. Ital. Crypt., pars V: 425, f. 154 (1943)
= Isoetes histrix var. grandis (A.Braun) Batt. & Trab.; Fl. Algérie Tunisie: 407 (1905)
= Isoetes histrix var. lacunosa Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 251 (1866)
= Isoetes histrix var. loricata (A.Braun) Batt. & Trab.; Fl. Algérie Tunisie: 407 (1905)
= Isoetes histrix var. scutellata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)
= Isoetes histrix var. subinermis Durieu; Bull. Soc. Bot. France 8: 164 (1861), nom. nud.
= Isoetes histrix var. submersa Trab.; Batt. & Trab., Fl. Alg. & Tunis. 407 (1902)
= Isoetes hystrix subsp. algarbiensis Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. arcuata Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. cesatii Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. collina Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. corsica Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. hispanica Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. horridula Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. Iongifolia Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. longiseta Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. numidica Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. oceanica Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. oranensis Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. phrygia Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. todaroi Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes hystrix subsp. transiens Gand.; Fl. Eur. 27: 197 (1891)
= Isoetes setacea Moris & De Not.; Florula Capr. 161 (1839)
species :
Isoetes hopei J.R.Croft; Blumea 26: 177-190 (1980)
Distribution: New Guinea (Irian Jaya)
42 NWG
Conservation Status: EN in Papua New Guinea
species :
Isoetes howellii Engelm.; Trans. Acad. Sci. St. Louis 4: 385 (1882)
Name: Howell´s Quillwort
Distribution: Canada (British Columbia); USA (California, Idaho, Montana, Oregon, Utah, Washington State); Mexico (Baja California Norte, Baja California Sur)
71 CDN 73 USA 76 USA 79 MEX
Synonyms: hidden
= Isoetes melanopoda f. californica (A.A.Eaton) Clute; Fern-Allies 260 (1905)
= Isoetes melanopoda var. californica A.A.Eaton; Working List North American Pteridophytes 27 (1901)
= Isoetes nuda Engelm.; Trans. Acad. Sci. St. Louis 4: 385 (1882)
= Isoetes underwoodii L.F.Hend.; Bot. Gaz. 23: 124 (1897)
species :
Isoetes humilior F.Muell. ex A.Braun; Linnaea 25: 722 (1852)
Distribution: Australia (Tasmania)
50 AUS
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria humilior (F.Muell. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes hookeri A.Braun; Herb Melbourn (1866), Monatsber. K. Akad. Berl. 538 (1868)
= Isoetes tasmanica Durieu; Bull. Soc. Bot. France 11: 104 (1864), non F. Muell.
species :
Isoetes hyemalis D.F.Brunt.; Castanea 59(1): 13 (1994)
Name: Winter Quillwort
Distribution: E-USA (North Carolina, South Carolina, Virginia)
75 USA 78 USA
Remarks: sometimes included in I. riparia, but accepted by Troia et al. 2016
Synonyms: hidden
= Isoetes canadensis var. amesii (A.A.Eaton) Clute; Fern Bull. 16: 54 (1908)
= Isoetes riparia var. amesii (A.A.Eaton ex Gilbert) Proctor; Amer. Fern J. 39: 119 (1949)
= Isoetes saccharata var. amesii A.A.Eaton; Rhodora 5(59): 278-279 (1903)
species :
Isoetes hypsophila Hand.-Mazz.; Anzeiger Acad. Wiss. Wien. Nr. 13: 95 (1923)
Distribution: China (SW-Sichuan, NW-Yunnan)
36 CHI
species :
Isoetes inflata E.R.L.Johnson; J. Roy. Soc. W. Australia 67(1): 37 (1984)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes jaegeri Pitot; Bull. Inst. Franc. Afr. Noire, A, 30: 58, f. 1-19 (1968)
Distribution: Burkina Faso
22 BKN
species :
Isoetes jamaicensis Hickey; Amer. Fern J. 71(3): 69 (1981)
Distribution: Jamaica
81 JAM
species :
Isoetes japonica A.Braun; Monatsber. K. Akad. Berl. 1: 459 (1861)
Distribution: China; South Korea (Pyeongtaek area); Japan
36 CHI 38 JAP KOR
Synonyms: hidden
= Calamaria japonica (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes edulis Sieber ex Miq.; Prod. Fl. jap. 390 (1866)-67
species :
Isoetes jarmaniae D.F.Brunt., Kantvilas & M.Garrett; Phytotaxa 522: 29 (2021)
Distribution: Australia (Tasmania)
50 AUS
species :
Isoetes jejuensis H.K.Choi, Ch.Kim & J.Jung; J. Pl. Biol. 51(5): 356 (2008)
Distribution: South Korea (Cheju Isl.)
38 KOR
species :
Isoetes junciformis D.F.Brunt. & D.M.Britton; Amer. Fern J. 89(3): 187-197 (1999)
Name: Rush quillwort
Distribution: USA (S-Georgia)
78 USA
species :
Isoetes karstenii A.Braun; Verh. Bot. Ver. Brandenb. 4. 332 (1862)
Distribution: Colombia (Arauca, Boyacá, Cauca, Cundinamarca, Huila, Magdalena, Meta, Nariño, Risaralda, Santander, Tolima, Valle); Venezuela (Merida, Trujillo)
82 CLM VEN
Synonyms: hidden
= Isoetes colombiana (T.C.Palmer) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 176 (1981)
= Isoetes lechleri var. colombiana Palmer; Amer. Fern J. 19: 18 (1929)
= Isoetes steyermarkii H.P.Fuchs; (1960), nom. nud.
species :
Isoetes kashyapii M.Patel, M.N.Reddy & H.K.Goswami; Indian Fern J. 39(1): 231 (2022)
Distribution: India (Gujarat)
40 IND
species :
Isoetes killipii C.V.Morton; Amer. Fern J. 35: 49 (1945)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Meta, Nariño, Putumayo, Risaralda, Santander, Tolima, Valle); Venezuela (Amazonas, Apure, Bolívar, Tachira); Guyana (Mt. Roraima); Ecuador
82 CLM GUY VEN 83 ECU
Synonyms: hidden
= Isoetes roncadorensis H.P.Fuchs; (1981), nom. nud.
= Isoetes roraimensis H.P.Fuchs; (1981), nom. nud.
species :
Isoetes kirkii A.Braun; Monatsber. K. Akad. Berl. 649 (1869)
Distribution: New Zealand (North Isl., South Isl.)
51 NZL
Synonyms: hidden
= Calamaria kirkii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes labri-draconis N.R.Crouch; Ferns S. Afr. Compreh. Guide: 746 (2011)
Distribution: South Africa (KwaZulu-Natal: E-Drakensberg)
27 SAF
species :
Isoetes lacustris L.; Sp. Pl. 2: 1100 (1753)
Name: Gewöhnliches Brachsenkraut, See-Brachsenkraut (DE), Lake Quillwort (EN)
Distribution: England; Wales; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Germany (Baden-Württemberg, Bayern, Mecklenburg-Vorpommern+, Niedersachsen, Schleswig-Holstein); Switzerland; Austria; Poland; Czech Republic; Spain; Andorra; France; ?Romania; +Bulgaria; Estonia; Latvia; Lithuania; Belarus; European Russia (N-European Russia, C-European Russia); Siberia (Altai, Irkutsk, W-Siberia); ?Japan; Greenland; Canada (Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon; USA (Connecticut, Massachusetts, Maine, Michigan, Minnesota, New Hampshire, New Jersey, New York, Rhode Island, Tennessee, Virginia, Vermont, Wisconsin)
10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER NET POL SWI 12 ADO FRA SPA 13 BUL 14 BLR EST LAT LIT RUS 30 SIB 70 CDN GNL 71 CDN 72 CDN SPM 74 USA 75 USA 78 USA
Synonyms: hidden
= Calamaria lacustris (L.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Calamaria setacea (Lam.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes atrovirens Th.Fr.; Bot. Notis. (1863): 48 (1865)
= Isoetes hieroglyphica A.A.Eaton; Fernwort Papers 10 (1900)
= Isoetes lacustris subsp. abbreviata Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. arvernensis Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. cambrica Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. editorum Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. errabunda Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. hieroglyphica (A.A.Eaton) Taylor & Luebke
= Isoetes lacustris subsp. lacustris Hook.f.; Student´s Fl. Brit. Isl., ed. 1, 471 (1870)
= Isoetes lacustris subsp. lamottei Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. langei Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. latifolia Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. noltei Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. obscura Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. pervaga Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. rivalis Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. rossica Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. scanica Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. scotica Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. septentrionalis Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. spreta Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. succedanea Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris subsp. suecica Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris var. curvifolia Casp.; Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 10 (pro syn.) (1936)
= Isoetes lacustris var. elatior Fliche; Isoetes des Vosges: 7 (1879)
= Isoetes lacustris var. falcata Gay; Bull. Soc. Bot. France 10: 386 (1863)
= Isoetes lacustris var. falcatum Fliche; Isoetes des Vosges: 7 (1879)
= Isoetes lacustris var. fragilis Gray; Nat. Arr. Brit. Pl. 2: 24 (1821 publ. 1822)
= Isoetes lacustris var. gracilis Gray; Nat. Arr. Brit. Pl. 2: 24 (1821 publ. 1822)
= Isoetes lacustris var. humilior Motelay & Vendryès; Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. humilis Mer; Bull. Soc. Bot. France 28- 72 (1881)
= Isoetes lacustris var. intermedia Mer; Bull. Soc. Bot. France 28: 72 (1881)
= Isoetes lacustris var. leiospora Klinggr.; Schrift. Nat. Ges. Danzig 6, 1: 20 (1884)
= Isoetes lacustris var. minor Milde; Höh. Sporenpfl. Deutschl.: 141 (1865)
= Isoetes lacustris var. moorei (D.Moore) Hook.; J. D., Student´s Fl. Brit. Isl., ed. 3, 526 (1884)
= Isoetes lacustris var. moorei (D.Moore) Syme & N.E.Br.; 5 (1886)
= Isoetes lacustris var. rectifolia Casp.; Glück, Pascher, Die Süsswasser-Flora Mitteleuropas 15: 10 (1936)
= Isoetes lacustris var. recurva Motelay & Vendryès; Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. setacea Léman; Dict. Sci. Nat. 24: 24 (1822)
= Isoetes lacustris var. stricta (Gay) Mer; Bull. Soc. Bot. France 28: 72 (1881)
= Isoetes lacustris var. stricta Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)
= Isoetes lacustris var. strictum J.Gay; Bull. Soc. Bot. France 10: 392 (1863)
= Isoetes lacustris var. tenella Léman; Dict. Sci. Nat. 24: 24 (1822)
= Isoetes lacustris var. tenuifolia A.Braun ex Milde; Sporenpfl. 141 (1865)
= Isoetes lacustris var. terrestris Motelay & Vendryès; Actes Soc. Linn. Bord. 36: 327 (1883)
= Isoetes lacustris var. vulgaris Casp.; Luerss., Farnpfl. 855 (1889)
= Isoetes lacustris vergens Gand.; Fl. Eur. 27: 193 (1891)
= Isoetes lacustris × stricta Gay
= Isoetes leiospora Klinggr.; Schrift. Nat. Ges. Danzig 6(1): 20 (1884)
= Isoetes macrospora Durieu; Bull. Soc. Bot. France 11: 101 (1864)
= Isoetes macrospora f. hieroglyphica (A.A.Eaton) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 194 (1922)
= Isoetes moorei D.Moore; J. Bot. (London) 16 (n. s. 7): 353, t. 199 (1878)
= Isoetes setacea Lam.; Encycl. 3: 314 (1789)
= Isoetes tenella Lem. ex Desv.; Mém. Soc. Linn. Paris 6: 179 (1827)
= Isoetes tuckermani var. harveyi (A.A.Eaton) Clute; Fern-Allies 226, 227, 260 (1905)
species :
Isoetes laosiensis C.Kim & H.K.Choi; Amer. Fern J. 100(1): 48 (2010)
Distribution: Laos
41 LAO
species :
Isoetes laurentiana D.F.Brunt.; Botany (Ottawa) 97(11): 579 (2019)
Distribution: Canada (Québec)
72 CDN
species :
Isoetes lechleri Mett.; Fil. Lechl. 2: 36 (1859)
Distribution: Colombia (Arauca, Boyacá, Cauca, Cundinamarca, Huila, Meta, Nariño, Santander); Ecuador (Chimborazo, Pinchincha, Napo); Peru (Junín, Puno, Ayacucho, Ancash); Bolivia (La Paz)
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Calamaria lechleri (Mett.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes arumiana H.P.Fuchs; (1959), nom. nud.
= Isoetes glacialis Aspl.; Ark. Bot. 20A, no. 7: 35, fig. 7 (1926)
= Isoetes karstenii var. anomala (T.C.Palmer) R.L.Small & Hickey; Amer. Fern J. 22: 130 (1933)
= Isoetes laevis U.Weber; Hedwigia 63: 252, fig. 30-31 (1922)
= Isoetes lechleri var. anomala T.C.Palmer; Amer. Fern J. 22: 130 (1933)
= Isoetes peruviana U.Weber; Hedwigia 63: 246, fig. 12-13 (1922)
= Isoetes rimbachiana H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(2): 226 (1982), nom. nov. pro I. lechleri var. anomala
= Isoetes socia A.Braun; Verh. Bot. Ver. Brandenb. 4: 332 (1862)
species :
Isoetes libanotica Musselman, Bolin & R.D.Bray; Novon 21(3): 295 (2011)
Distribution: Lebanon
34 LEB
species :
Isoetes lithophila N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 135 (1922) (lithophylla)
Name: Rock Quillwort
Distribution: USA (SC-Texas: Burnett Co., Mason Co., Llano Co.)
73 USA 77 USA
Conservation Status: IUCN V
species :
Isoetes longifolia J.B.S.Pereira; Syst. Bot. 47(2): 304 (2022)
Distribution: Brazil (Bahia)
84 BRA
species :
Isoetes longissima Bory & Durieu (M.Laínz) L.Síez, Nava & Fern.Prieto; Naturalia Cantabricae 8(Especial): 64 (2020)
subspecies :
subsp. asturicensis
Distribution: Spain
12 SPA
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes asturicensis (M.Laínz) M.Laínz; Anuario Soc. Brot. 39: 118 (1973)
= Isoetes velata subsp. asturicensis (M.Laínz) Rivas Mart. & Prada; Lazaroa 2: 327 (1980)
= Isoetes boryana subsp. asturicensis M.Laínz; Bol. Inst. Estud. Astur., Suppl. Cienc. 15: 6 (1970)
= Isoetes velata var. asturicense (M.Laínz) E.Rico & X.Giráldez; Anal. Jard. Bot. Madrid 46(2): 585 (1989 publ. 1990)
subspecies :
subsp. intermedia (Trab.) Troia & Greuter; Willdenowia 44(2): 288 (2014)
Distribution: Morocco; Algeria
20 ALG MOR
Synonyms: hidden
= Isoetes velata subsp. intermedia (Trab.) Maire & Weiller; Maire, Fl. Afrique. N. 1: 89 (1952)
= Isoetes velata f. immaculata Maire; Encycl. Biol. 33: 90 (1952)
= Isoetes velata tax.infrasp.intermedia Trab.; Batt. & Trab., Fl. Alg. & Tunis. 407 (1905)
= Isoetes velata var. intermedia A.Braun; Bory & Durieu, Expl. Sci. Alg. (1848)
subspecies :
subsp. perralderiana (Milde) Troia & Greuter; Willdenowia 44(2): 288 (2014)
Distribution: Algeria
20 ALG
Synonyms: hidden
= Calamaria perralderiana (Durieu & Letourn. ex Milde) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes perralderiana Durieu & Letourn.; Kralik, Pl. Alg. Exsicc. 157 (1862)
= Isoetes perralderiana Durieu & Letourn. ex Milde; Fil. Eur.: 282 (1867)
= Isoetes velata subsp. perralderiana (Durieu & Letourn.) Trab.; Fl. Syn. 407 (1902), et Fl. Alg. & Tunis. 407 (1905)
= Isoetes velata var. perralderiana (Durieu & Letourn.) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 121 (1922)
species :
Isoetes longpingii Y.H.Yan, Y.F.Gu & J.P.Shu; Guihaia 42(10): 1625 (2022)
Distribution: China (Hunan)
36 CHI
species :
Isoetes louisianensis Thieret; Sida 5: 129 (1973)
Name: Louisiana Quillwort
Distribution: USA (S-Louisiana, Mississippi, Alabama)
78 USA
Remarks: may be a hybrid I. engelmannii x melanopoda
Conservation Status: IUCN E
species :
Isoetes luetzelburgii U.Weber ex Luetzelb.; Estud. Bot. Nordeste 3: 257 (1864)
Distribution: Brazil (Paraíba, Rio Grande do Norte)
84 BRA
species :
Isoetes malinverniana Ces. & De Not.; Index Sem. Hort. Genuensis 1858: 36 (1858)
Distribution: NW-Italy (Vercelli, Novara, Torino)
13 ITA
Conservation Status: IUCN E
Synonyms: hidden
= Calamaria malinverniana (Ces. & De Not.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes maritima Underw.; Bot. Gaz. 13: 94 (1888)
Name: Spiny-spored Quillwort
Distribution: Russian Far East (Commander Isl.); Alaska; Aleutes; C-Canada (Vancouver Isl., British Columbia); NW-USA (Washington)
31 RFE 70 ASK 71 CDN 73 USA
Synonyms: hidden
= Isoetes beringensis Kom.; Bull. Jard. Bot. Acad. Sci. USSR 30 (1-2): 196 (1932)
= Isoetes braunii var. maritima (Underw.) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 174 (1922)
= Isoetes echinospora f. maritima (Underw.) Clute; Fern. Bull. 16: 55 (1908)
= Isoetes echinospora subsp. maritima (Underw.) Á.Löve; Amer. Fern J. 52: 121 (1962)
= Isoetes echinospora var. maritima (Underw.) A.A.Eaton; Fern Bull. 13: 52 (1905)
= Isoetes echinospora var. savilei B.Boivin; Amer. Fern J. 51: 85 (1961)
= Isoetes macounii A.A.Eaton; Fern Bull. 8: 12 (1900)
= Isoetes muricata subsp. maritima (Underw.) Hultén; Ark. Bot. (n. s.) 7(1): 8 (1968)
= Isoetes setacea subsp. maritima (Underw.) Holub; Acta Horti bot. prag.: 55 (1964)
species :
Isoetes martii A.Braun ex Kuhn; Mart., Fl. Bras. 1(2): 646, t. 78 (1884)
Distribution: Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, São Paulo)
84 BRA
Synonyms: hidden
= Calamaria martii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes mattaponica Musselman & W.C.Taylor; Novon 11(2): 200 (-204, figs. 1-6) (2001)
Name: Mattaponi quillwort
Distribution: USA (Virginia: Mattaponica River)
78 USA
species :
Isoetes maxima Hickey, Macluf & Link-Pérez; Amer. Fern J. 99(3): 194 (2010)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes melanopoda Gay & Durieu; Bull. Soc. Bot. France 11: 102 (1864)
subspecies :
subsp. melanopoda
Name: Black-footed Quillwort
Distribution: USA (Alabama, Arkansas, Georgia, Iowa, Idaho, Illinois, Indiana, Kansas, Kentucky, Louisiana, Minnesota, Missouri, Mississippi, Montana, North Carolina, Nebraska, New Jersey, Oklahoma, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia)
73 USA 74 USA 75 USA 76 USA 77 USA 78 USA
Synonyms: hidden
= Calamaria melanopoda (J.Gay) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes melanopoda f. pallida (Engelm.) Fernald; Rhodora 51: 103 (1949)
= Isoetes melanopoda var. pallida Engelm.; Trans. Acad. Sci. St. Louis 4: 387 (1882)
subspecies :
subsp. silvatica D.F.Brunt. & D.M.Britton; Castanea 71(1): 26 (2006)
Distribution: E-USA (North Carolina)
75 USA 78 USA
species :
Isoetes melanospora Engelm.; Trans. Acad. Sci. St. Louis 3: 395 (1877)
Name: Blackspore Quillwort
Distribution: USA (C-Georgia, South Carolina: Lancaster Co.)
78 USA
Synonyms: hidden
= Calamaria melanospora (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes melanotheca Alston; Bol. Soc. Broter. Ser. 2A, 30: 15 (1956)
Distribution: Guinea-Bissau; Guinea; Togo; Benin
22 BEN GNB GUI TOG
Conservation Status: IUCN R
species :
Isoetes mexicana Underw.; Bot. Gaz. 13: 93 (1888)
Distribution: Mexico (Aguascalientes, Chihuahua, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Queretaro, Zacatecas); N-Guatemala
79 MEX 80 GUA
species :
Isoetes microvela D.F.Brunt.; Rhodora 100(903): 270-274, f. 1-3, 4e-h, 5 (1998)
Name: Thin-wall quillwort
Distribution: USA (North Carolina: coastal)
78 USA
species :
Isoetes minima A.A.Eaton; Fern Bull. 6: 30 (1898)
Name: Midget quillwort
Distribution: SW-Canada (British Columbia); USA (Washington State)
71 CDN 73 USA
Remarks: valid species fide Taylor et al. 2003
Synonyms: hidden
= Isoetes howellii var. minima (A.A.Eaton) N.Pfeiff.; Ann. Missouri Bot. Gard. 9(2): 142 (1922)
species :
Isoetes mississippiensis S.W.Leonard, W.C.Taylor, Musselman & R.D.Bray; PhytoKeys 2016(74): 97-106 (2016)
Distribution: USA (S-Mississippi)
78 USA
species :
Isoetes mongerensis E.R.L.Johnson; J. Roy. Soc. W. Australia 67(1): 39 (1984)
Distribution: Australia (SW-Western Australia)
50 AUS
species :
Isoetes montezumae A.A.Eaton; Fern Bull. 5: 25 (1897)
Distribution: Mexico
79 MEX
Remarks: recognized in Troia et al. 2016, included in I. mexicana by other authors
species :
Isoetes mourabaptistae J.B.S.Pereira, P.G.Windisch, Lorscheitt. & Labiak; Amer. Fern J. 102(2): 175 (2012)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes muelleri A.Braun; Monatsber. K. Akad. Berl. 541 (1868)
Distribution: Australia (Australian Capital Territory, Western Australia, Northern Territory, South Australia, Queensland, Victoria, Tasmania)
50 AUS
Synonyms: hidden
= Calamaria muelleri (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes stuartii A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1868: 539 (1868)
= Isoetes tenuissima F.Muell.; (1868) (non Boreau)
species :
Isoetes naipiana P.G.Windisch, Lorscheitt. & Nervo; Willdenowia 44: 394 (2014)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes nana J.B.S.Pereira; PhytoKeys 89: 93 (2017)
Distribution: Brazil (Rio de Janeiro)
84 BRA
species :
Isoetes neoguineensis Baker ex F.Muell.; Ann. Rep. Brit. N. Guinea 1897-8: 149, nomen (1898)
Distribution: New Guinea (Papua New Guinea: Owen Stanley Ranges)
42 NWG
Conservation Status: IUCN R (twice found)
Synonyms: hidden
= Isoetes neoguineensis Baker; Sadeb., Nat. Pflanzenfam. 1(4) 776 (1902)
= Isoetes neoguineensis var. rheophila J.R.Croft; Blumea 26: 177-190 (1980)
species :
Isoetes nigritiana A.Braun; Filic. Afr. 196 (1868)
Distribution: Mali; Burkina Faso; Ivory Coast; Nigeria; Central African Republic; Sudan
22 BKN IVO MLI NGA 23 CAF 24 SUD
Synonyms: hidden
= Isoetes garnieri Chev. & Monnier; Bull. Mus. Hist. Nat. Paris 18(1): 91-979 1 fig. (1946)
= Calamaria nigritiana (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes nigroreticulata Verdc.; Fl. Trop. E. Africa, Isoet. 2 (2005)
Distribution: Kenya
25 KEN
species :
Isoetes novogranadensis H.P.Fuchs; Caldasia 8: 314 (1969)
Distribution: Colombia (Bolívar, Boyacá, Cauca, Cundinamarca, Huila, Nariño, Putumayo, Valle); Ecuador; Peru
82 CLM 83 ECU PER
Synonyms: hidden
= Isoetes dichotoma L.E.Mora & W.Hagemann; Mutisia 43: 5 (1977)
= Isoetes hagemannii H.P.Fuchs; (1981), nom. nud.
= Isoetes leistii H.P.Fuchs; (1981), nom. nud.
= Isoetes narinensis H.P.Fuchs; (1981), nom. nud.
= Isoetes pichichensis H.P.Fuchs; (1981), nom. nud.
= Isoetes quilindanensis H.P.Fuchs; (1981), nom. nud.
species :
Isoetes nuttallii A.Braun ex Engelm.; Amer. Naturalist 8: 215 (1874)
Name: Nuttall´s Quillwort
Distribution: C-Canada (Vancouver Isl.); W-USA (California, Oregon, Washington)
71 CDN 73 USA 76 USA
Synonyms: hidden
= Calamaria nuttallii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Calamaria suksdorfii (Baker) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes opaca Nutt. ex Engelm.; Trans. Acad. Sci. St. Louis 4: 389 (1886), pro syn.
= Isoetes suksdorfii Baker; Handb. Fern-Allies 132 (1887)
species :
Isoetes occidentalis Hend.; Bull. Torrey Bot. Club 27: 358 (1900)
Name: Western Quillwort
Distribution: Alaska; Aleutes; C-Canada (British Columbia); NW-USA (Washington, Oregon, C-California, Idaho, Montana, Wyoming, Colorado)
70 ASK 71 CDN 73 USA 76 USA
Synonyms: hidden
= Isoetes echinospora f. flettii (A.A.Eaton) Clute; Fern Bull. 16: 55 (1908)
= Isoetes echinospora subsp. flettii (A.A.Eaton) Piper & Beattie; Fl. N. W. Coast [Piper & Beattie] 16 (1915)
= Isoetes echinospora var. flettii A.A.Eaton; Fern Bull. 13: 51 (1905)
= Isoetes flettii (A.A.Eaton) N.Pfeiff.; Ann. Missouri Bot. Gard. 9(2): 186 (1922)
= Isoetes howellii f. piperi (A.A.Eaton) Clute; Fern Bull. 16: 56 (1908)
= Isoetes howellii var. piperi (A.A.Eaton) Clute; Fern-Allies 246, 258 (1905)
= Isoetes lacustris subsp. paupercula (Engelm.) Feilberg; Nordic J. Bot. 4: 155 (1984)
= Isoetes lacustris var. paupercula Engelm.; Trans. Acad. Sci. St. Louis 4: 377 (1882)
= Isoetes macrospora f. paupercula (Engelm.) Clute; Fern Bull. 16: 56 (1908)
= Isoetes maritima var. flettii (A.A.Eaton) C.F.Reed; Amer. Fern J. 35: 85 (1945)
= Isoetes occidentalis subsp. paupercula (Engelm.) Feilberg; Nordic J. Bot. 4(2): 155 (1984)
= Isoetes occidentalis var. piperi (A.A.Eaton) A.Nelson & J.F.Macbr.; Bot. Gaz. 61(1): 30 (1916)
= Isoetes paupercula (Engelm.) A.A.Eaton; Proc. U.S. Nat. Herb. 23(1226): 649 (1901)
= Isoetes piperi A.A.Eaton; Fern Bull. 11: 85 (nom. nud.) (1903), 13: 51 (1905)
species :
Isoetes olympica A.Br.; Fil. Eur.: 285 (1867)
Distribution: Turkey (Anatolia); Lebanon; S-Syria
34 LEB SYR TUR
Remarks: histrix agg.
Synonyms: hidden
= Calamaria olympica (A.Br.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes orcuttii A.A.Eaton; Fern Bull. 6: 30 (1898) (nom. nud.), et 8: 13 (1900)
Name: Orcutt´s Quillwort
Distribution: USA (S-California); Mexico (Baja California Norte)
76 USA 79 MEX
Synonyms: hidden
= Isoetes nuttallii var. orcuttii (A.A.Eaton) Clute; Fern-Allies 253 (1905)
species :
Isoetes organensis U.Weber; Luetzelburg, Estud. Bot. Nordeste 3: 256 (no date)
Distribution: Brazil (Minas Gerais, Rio de Janeiro)
84 BRA
species :
Isoetes orientalis Hong Liu & Q.F.Wang; Novon 15(1): 164 (2005)
Distribution: China (Zhejiang)
36 CHI
Remarks: status uncertain, probably a hybrid Isoetes sinensis x yunguiensis
species :
Isoetes ovata N.Pfeiff.; Ann. Missouri Bot. Gard. 8(2): 109 (1922)
Distribution: Guyana; French Guiana
82 FRG GUY
Synonyms: hidden
= Isoetes clavata U.Weber; Ber. dt. Bot. Ges. 52: 121 (1934)
species :
Isoetes pallida Hickey; Amer. Fern J. 78(1): 35 (1988)
Distribution: Mexico (Campeche, Guerrero, Oaxaca); Belize
79 MEX 80 BLZ
Conservation Status: IUCN R
species :
Isoetes palmeri H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 168 (1981)
Distribution: Colombia (Arauca, Boyacá, Cauca, Cundinamarca); Venezuela (Merida); Ecuador (Chimborazo)
82 CLM VEN 83 ECU
Synonyms: hidden
= Isoetes cleeffii H.P.Fuchs; Proc. Kon. Akad. Wet. Amsterdam C 84(2): 177 (1981)
species :
Isoetes panamensis Maxon & C.V.Morton; Ann. Missouri Bot. Gard. 26(4): 272 (1939)
Distribution: Guatemala; Nicaragua; Costa Rica; Panama; Colombia; Ecuador; E-Peru; Bolivia (Cochabamba, La Paz); Brazil (Bahia, Maranhão, Mato Grosso do Sul, Mato Grosso, Paraná, Tocantins); Paraguay (Alto Paraná, Caaguazu)
80 COS GUA NIC PAN 82 CLM 83 BOL ECU PER 84 BRA 85 PAR
Synonyms: hidden
= Isoetes pacifica Svenson; Amer. Fern J. 34: 123 (1944)
= Isoetes savannarum L.D.Gómez; Phytologia 49(4): 339 (1981)
species :
Isoetes parvula Hickey; Fieldiana, Bot. 34: 92 (1994)
Distribution: Peru (Ayacucho)
83 PER
Conservation Status: Known from type only
species :
Isoetes pedersenii H.P.Fuchs ex E.I.Meza & Macluf; Anais Acad. Brasil. Ci. 82(2): 354 (2010)
Distribution: Brazil (Mato Grosso do Sul); Paraguay; Argentina (Corrientes)
84 BRA 85 ARG PAR
species :
Isoetes perrieriana Iversen; Dansk Bot. Ark. 7: 200-201 (1932)
Distribution: Madagascar
29 MDG
species :
Isoetes philippinensis Merr. & L.M.Perry; Amer. Fern J. 30: 19 (1940)
Distribution: Sulawesi; Philippines (Lanao del Norte)
42 PHI SUL
Conservation Status: CR in the Philippines, recently rediscovered (Amoroso et al. 2022)
species :
Isoetes phrygia Hausskn.; Mitt. Thüring. Bot. Vereins 13: 72 (1899)
Distribution: Greece (S-Peloponnisos, Kyklades); Crete (rare); Turkey (Anatolia)
13 GRC KRI 34 TUR
Synonyms: hidden
= Isoetes histrix var. phrygia Boiss.; Fl. Orient. 5: 748 (1884)
species :
Isoetes pichisermollii Eb.Fisch. & Lobin; Willdenowia 52(3): 330-331 (2022)
Distribution: Burundi
23 BUR
Remarks: cited by Pichi-Sermolli 1983/1985 as ´Isoetes abyssinica´
species :
Isoetes pirinica D.F.Brunt. & D.Ivanova; Fern Gaz. 21(6): 247 (2021)
Distribution: Bulgaria
13 BUL
species :
Isoetes pitotii Alston; Bol. Soc. Broter. Ser. 2, 30: 15 (1956)
Distribution: Guinea; Mali; Sudan
22 GUI MLI 24 SUD
species :
Isoetes precocia R.L.Small & Hickey; Amer. Fern J. 91(2): 61 (-62, figs. 12-13, 18-19) (2001)
Distribution: ?Colombia; Venezuela (Merida)
82 VEN
species :
Isoetes pringlei Underw.; Zoë 1: 98 (1890)
Distribution: Mexico (Jalisco, Mexico State, Chiapas)
79 MEX
Conservation Status: IUCN R
species :
Isoetes prototypus D.M.Britton & Goltz; Canad. J. Bot. 69: 278 (1991)
Name: Spike quillwort
Distribution: Canada (New Brunswick, Nova Scotia); USA (Maine, New Hampshire)
72 CDN 75 USA
Conservation Status: IUCN V
species :
Isoetes pseudojaponica M.Takamiya, Watan. & Ono; Acta Phytotax. Geobot. 48(2): 107 (1998)
Distribution: Japan
38 JAP
Conservation Status: VU in Japan
species :
Isoetes pusilla C.R.Marsden & Chinnock; Muelleria 8(1): 57 (1993)
Distribution: Australia (Victoria)
50 AUS
species :
Isoetes quiririensis J.B.S.Pereira & Labiak; Syst. Bot. 38(4): 869 (2014)
Distribution: Brazil (Paraná, Santa Catarina)
84 BRA
species :
Isoetes ramboi Herter; Revista Sudamer. Bot. 8 (1-2): 19-25 (1949)
Distribution: Brazil (Mato Grosso, Rio Grande do Sul, Santa Catarina)
84 BRA
species :
Isoetes riparia Engelm.; Flora (Regensburg) 29: 178 (1846)
Name: Shore Quillwort
Distribution: USA (Delaware, District of Columbia, Maryland, North Carolina, New Jersey, New York, Pennsylvania, Rhode Island, South Carolina, Vermont, Virginia, West Virginia)
75 USA 78 USA
Remarks: riparia agg.
Synonyms: hidden
= Calamaria riparia (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes amesii A.A.Eaton; Gilbert, Fern Bull. 11: 103 (1903)
= Isoetes foveolata var. plenospora A.A.Eaton; Rhodora 5: 280 (1903)
= Isoetes riparia f. saccharata (Engelm.) Clute; Fern Bull. 16: 57 (1908)
= Isoetes riparia f. saccharata (Engelm.) Proctor; Amer. Fern J. 39: 119 (1949)
= Isoetes riparia var. palmeri (A.A.Eaton) Proctor; Amer. Fern J. 39: 119 (1949)
= Isoetes riparia var. reticulata (A.A.Eaton) Proctor; Amer. Fern J. 39: 119 (1949)
= Isoetes riparia var. riparia Proctor; Amer. Fern J. 39: 119 (1949)
= Isoetes saccharata f. palmeri (A.A.Eaton) A.A.Eaton; Fern Bull. 11 : 25 (1903)
= Isoetes saccharata f. reticulata A.A.Eaton; Fern Bull. 11 : 25 (1903)
= Isoetes saccharata var. palmeri A.A.Eaton; Proc. Biol. Soc. Washington 14(10) 49 (1901)
= Isoetes saccharata var. reticulata A.A.Eaton; Proc. Biol. Soc. Washington 14(10): 49 (1901)
species :
Isoetes rwandensis Eb.Fisch. & Lobin; Willdenowia 52(3): 332 (2022)
Distribution: Rwanda
23 RWA
species :
Isoetes sabatina Troìa & Azzella; Pl. Biosystems 147(4): 1052 (2013)
Distribution: Italy
13 ITA
species :
Isoetes saccharata Engelm.; Gray´s Manual, ed. 5, 676 (1867)
Name: Sugar quillwort
Distribution: USA (District of Columbia, Delaware, Maryland, Virginia)
78 USA
Synonyms: hidden
= Calamaria saccharata (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
species :
Isoetes sahyadrii Mahab.; [Curr. Sci. 7(2): 62, f. 1 (1938), nom. inval.] ex L. N. Rao, Curr. Sci. 13(11): 286-287 (1944) (sahyadriensis auct.), nom. cons. prop.
Distribution: India (Andhra Pradesh, Karnataka, Maharashtra, Madhya Pradesh, Rajasthan)
40 IND
Synonyms: hidden
= Isoetes mahadevensis G.K.Srivast., D.D.Pant & P.K.Shukla; Amer. Fern J. 83(4): 106 (1993)
= Isoetes panchananii D.D.Pant & Srivast.; Proc. Nat. Inst. Sci. India 28: 243, t. 13, 14, f. 1, 2 (1962)
= Isoetes panchananii var. panchmarhiensis G.K.Srivast., Mritt.Srivast. & P.K.Shukla; Indian Fern J. 14(1-2): 44 (1998)
= Isoetes panchaganiensis G.K.Srivast., D.D.Pant & P.K.Shukla; Amer. Fern J. 83(4): 107 (1993) (fide Fraser-Jenkins 1997)
= Isoetes panchaganiensis var. kemangundiensis G.K.Srivast., D.D.Pant & P.K.Shukla; Amer. Fern J. 83(4): 114 (1993) (fide Fraser-Jenkins 1997)
= Isoetes rajasthanensis Gena & Bhardwaja; J. Bombay Nat. Hist. Soc. 81(1): 165 (1984)
= Isoetes reticulata Gena & Bhardwaja; J. Bombay Nat. Hist. Soc. 81(1): 166 (1984)
= Isoetes sampathkumaranii L.N.Rao; Curr. Sci. 13: 286 (1944)
species :
Isoetes santacruzensis J.B.S.Pereira; PhytoKeys 131: 61 (2019)
Distribution: Bolivia (Santa Cruz)
83 BOL
species :
Isoetes saracochensis H.P.Fuchs ex Hickey; Fieldiana, Bot. 34: 94 (1994)
Distribution: Peru
83 PER
Synonyms: hidden
= Isoetes saracochensis H.P.Fuchs; (1960), nom. nud.
species :
Isoetes savatieri Franch.; Bull. Soc. Bot. France 31: 395 (1884)
Distribution: Argentina (Chubut, Neuquen, Rio Negro, Tierra del Fuego); Chile (Valparaíso, Maule, Ñuble, Biobío, Araucanía, Los Lagos, Aisén, Magallanes)
85 ARG CLE
Conservation Status: IUCN R
Synonyms: hidden
= Calamaria savatieri (Franch.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes lechleri var. savatieri (Franch.) L.D.Gómez; Brenesia 18: 5 (1980)
= Isoetes chilensis H.P.Fuchs; (1981), nom. nud.
species :
Isoetes schweinfurthii A.Braun; J. Bot. 18: 108 (1880)]
Distribution: Senegal; Burkina Faso; Ivory Coast; Benin; Nigeria; Central African Republic; Chad; Sudan; Ethiopia; Tanzania; S-Angola; Zambia; Malawi; Zimbabwe; Mozambique; N-Namibia; S-Botswana; South Africa (Limpopo); Madagascar
22 BEN BKN IVO NGA SEN 23 CAF 24 CHA ETH SUD 25 TAN 26 ANG MLW MOZ ZAM ZIM 27 BOT NAM SAF 29 MDG
Synonyms: hidden
= Calamaria schweinfurthii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes rhodesiana Alston; Bol. Soc. Broter. Ser. 2, 30: 17 (1956)
= Isoetes kersii Wanntorp; Svensk Bot. Tidsskr. 64: 146, t. 1, 3 (1970)
species :
Isoetes sehnemii H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 38 (1986)
Distribution: Brazil (Rio Grande do Sul)
84 BRA
species :
Isoetes septentrionalis D.F.Brunt.; Canad. Field-Naturalist 129: 176 (2015)
Name: Northern Quillwort
Distribution: E-Canada (SE-Ontario, SW-Québec); USA (Connecticut, Massachusetts, New Hampshire, ?Maine, Pennsylvania, Vermont)
72 CDN 75 USA
Remarks: riparia agg.
species :
Isoetes serracarajensis J.B.S.Pereira, Salino & Stützel; Phytotaxa 272(2): 142 (2016)
Distribution: Brazil (Pará)
84 BRA
species :
Isoetes shangrilaensis Xiang Li, Yuqian Huang, X.Dai & Xing Liu; Phytotaxa 397(1): 71 (2019)
Distribution: China (Yunnan)
36 CHI
species :
Isoetes sinensis T.C.Palmer; Amer. Fern J. 17: 111 (1927)
Distribution: China (Anhui, Guangxi, Jiangsu, Zhejiang); S-Japan
36 CHI 38 JAP
Remarks: allotetraploid originated from I. yunguiensis x taiwanensis
species :
Isoetes smithii H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 17 (1986)
Distribution: Brazil (Rio Grande do Sul, Santa Catarina)
84 BRA
species :
Isoetes spannagelii H.P.Fuchs; Fl. Ilustr. Catar. 1 (Isoetaceae): 33 (1986)
Distribution: Brazil (Santa Catarina)
84 BRA
species :
Isoetes spinulospora C.Jermy & Schelpe; Contr. Bolus Herb. 10: 151 (1982)
Distribution: Bioko Isl. [Fernando Poo]
23 BIO
Conservation Status: IUCN R
species :
Isoetes stellenbossiensis A.V.Duthie; Trans. R. Soc. S. Afr. 17: 328 (1929)
Distribution: South Africa (W-Cape Prov., N-Cape Prov.)
27 SAF
species :
Isoetes stephanseniae A.V.Duthie; Trans. Roy. Soc. South Africa 17: 330 (1929)
Distribution: South Africa (W-Cape Prov.)
27 SAF
Synonyms: hidden
= Isoetes capensis var. stephanseniae (A.V.Duthie) Schelpe & N.C.Anthony; Bothalia 15(3-4): 555 (1985)
species :
Isoetes stevensii J.R.Croft; Blumea 26: 177-190 (1980)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: IUCN R
species :
Isoetes storkii T.C.Palmer; Amer. Fern J. 22(4): 136 (1932)
Distribution: Costa Rica; Panama
80 COS PAN
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes montana T.C.Palmer; Amer. Fern J. 21(4): 134 (1931) (nom. illeg., non Isoetes montana U. Weber (1922))
= Isoetes tryoniana Gómez Pompa; Revista Biol. Trop. 17: 108 (1970)
species :
Isoetes taiwanensis De Vol; Taiwania 17(1): 1-7, f. 1 (1972)
Distribution: Taiwan
38 TAI
Conservation Status: IUCN E (extremely rare, only in one single pond)
variant :
var. kinmenensis F.Y.Lu, H.H.Chen & Y.L.Hsueh; Flora of Kinmen 26 (2011)
Distribution: Taiwan
38 TAI
Conservation Status: IUCN E (extremely rare)
species :
Isoetes tamaulipana Mora-Olivo, A.Mend. & Mart.-Aval.; Phytotaxa 267(2): 115 (2016)
Distribution: Mexico (Tamaulipas)
79 MEX
species :
Isoetes tegetiformans Rury; Amer. Fern J. 68(4): 100 (1978)
Name: Mat-forming Merlin´s-grass
Distribution: USA (N-Georgia)
78 USA
Conservation Status: IUCN E. Extremely rare, known from only a few vernal pools on granite outcrops
species :
Isoetes tennesseensis Luebke & Budke; Amer. Fern J. 93(4): 184-190 (2003)
Name: Tennessee Quillwort
Distribution: USA (Tennessee)
78 USA
species :
Isoetes tenuifolia Jermy; J. Hall, J. Linn. Soc., Bot. 64: 118 (1971)
Distribution: Ghana; ?Kenya
22 GHA
Conservation Status: IUCN R (Ghana), E (Kenya)
species :
Isoetes texana Singhurst, Rushing & W.C.Holmes; Phytoneuron 22: 1 (2011)
Distribution: USA (Texas)
77 USA
species :
Isoetes tiguliana Gennari; Comment. Soc. Critt. Ital. 1(1): 42 (1861)
Distribution: Italy; Morocco; Algeria; Tunisia
13 ITA 20 ALG MOR TUN
Remarks: bona sp. fide Troia et al. 2016
Synonyms: hidden
= Calamaria dubia (Gennari) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Calamaria tegulensis (Genn.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes dubia Gennari; Comment. Soc. Critt. Ital. 1: 104 (1861)
= Isoetes tegulensis f. submersa Glück; Pascher, Die Süsswasser-Flora Mitteleuropas 15: 20 (1936)
= Isoetes tegulensis f. terrestris Glück; Pascher, Die Süsswasser-Flora Mitteleuropas 15: 20 (1936)
= Isoetes tegulensis Gennari; Comment. Soc. Critt. Ital. 1: 106 (1861) (sept.) (nom. illeg.)
= Isoetes velata subsp. dubia (Gennari) Batt. & Trab.; Fl. Syn. 407 (1902)
= Isoetes velata subsp. tegulensis (Gennari) Batt. & Trab.; Fl. Alg. & Tunis. 407 (1861), et Fl. Syn. 407 (1905)
= Isoetes velata var. tegulensis (Gennari) Fiori & Paol.; Fl. Anal. Ital. 1: 25 (1896)
= Isoetes velata var. tegulensis b.maculosa Gennari; Ann. Agric. Sci. 11: 175 (1865)
= Isoetes velata var. tegulensis c.emersa Gennari; Giorn. Bot. Ital. 2: 96 (1870)
= Isoetes velata var. tegulensis Fiori; Fiori & Paol., Fl. Anal. Ital. 1: 25 (1896)
species :
Isoetes todaroana Troìa & Raimondo; Amer. Fern J. 99(4): 238 (2009 publ. 2010)
Distribution: S-Italy; Sicily; Greece
13 GRC ITA SIC
Synonyms: hidden
= Isoetes iapygia Ernandes, Beccarisi & V.Zuccarello; Pl. Biosystems 144(4): 807 (2010) (fide Troia & Greuter 2015)
species :
Isoetes toximontana Musselman & J.P.Roux; Novon 12: 504-507 (2002)
Distribution: South Africa (N-Cape Prov., W-Cape: Gifberg to Kamiesberg)
27 SAF
species :
Isoetes transvaalensis C.Jermy & Schelpe; Contr. Bolus Herb. 10: 150 (1982)
Distribution: South Africa (Limpopo, Gauteng, Mpulamanga, E-Cape Prov., Free State, KwaZulu-Natal); Lesotho
27 LES SAF
species :
Isoetes triangula U.Weber; Hedwigia 63: 253, f. 32, 33 (1922)
Distribution: Mexico (Oaxaca); Venezuela (Amazonas, Bolívar, Cojedes, Guarico); Guyana; French Guiana; Brazil (Acre, Amazonas, Roraima)
79 MEX 82 FRG GUY VEN 84 BRA
Synonyms: hidden
= Isoetes schinzii H.P.Fuchs; (1960), nom. nud.
species :
Isoetes tripus A.Braun; Herb. Vindob. ex Hook. (1863)
Distribution: Australia (SW-Western Australia)
50 AUS
Synonyms: hidden
= Calamaria tripus (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes phaeospora Durieu; Bull. Soc. Bot. France 11: 103 (1864)
species :
Isoetes tuckermanii A.Braun ex Engelm.; A. Gray, Manual ed. 5: 676 (1867)
subspecies :
subsp. tuckermanii
Name: Tuckermans Quillwort
Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec); USA (Connecticut, Delaware, Massachusetts, Maryland, Maine, New Hampshire, New Jersey, New York, Rhode Island, Vermont)
72 CDN 75 USA 78 USA
Remarks: inclusion of I. acadiensis fide Troia et al. 2016
Synonyms: hidden
= Calamaria tuckermanii (A.Braun ex Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Isoetes tuckermanii var. borealis A.A.Eaton; Fern Bull. 6: 7 (1898)
subspecies :
subsp. acadiensis (Kott) D.F.Brunt.; Canad. Field-Naturalist 132(4): 366 (2018)
Distribution: E-Canada (Nova Scotia)
72 CDN
Synonyms: hidden
= Isoetes acadiensis Kott; Canad. J. Bot. 59: 2592 (1981)
species :
Isoetes tuerckheimii Brause; Urban, Symb. Ant. 7: 161 (1912)
Distribution: Hispaniola (Dominican Republic)
81 HIS
species :
Isoetes udupiensis P.K.Shukla, G.K.Srivast., S.K.Shukla & P.K.Rajagopal; Taxon 54(1): 112 (2005)
Distribution: India (Karnataka)
40 IND
Remarks: bona sp. fide Fraser-Jenkins et al. 2017
Synonyms: hidden
= Isoetes divyadarshanii P.K.Shukla, G.K.Srivast. & S.K.Shukla; Taxon 54(1): 113 (2005)
species :
Isoetes ulei U.Weber; Hedwigia 63: 255, fig. 40-41 (1922)
Distribution: Brazil (Rio de Janeiro)
84 BRA
Synonyms: hidden
= Isoetes goebelii U.Weber; Hedwigia 63: 247, fig. 16-17 (1922)
species :
Isoetes valida (Engelm.) Clute; Fern-Allies 236 (1905)
Name: Strong quillwort
Distribution: USA (Alabama, Delaware, Georgia, North Carolina, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)
75 USA 78 USA
Synonyms: hidden
= Isoetes engelmannii var. valida Engelm.; Manual (Gray), ed. 5. 677 (1867)
species :
Isoetes vanensis M.Keskin & G.Zare; Phytotaxa 269(4): 295 (2016)
Distribution: Turkey (E-Anatolia)
34 TUR
species :
Isoetes vermiculata Hickey; Ann. Missouri Bot. Gard. 76(4): 1160 (1989)
Distribution: Venezuela (Amazonas)
82 VEN
species :
Isoetes virginica N.Pfeiff.; Claytonia 3: 29 (1937) (fide Musselman 2003)
Name: Piedmont quillwort
Distribution: SE-USA (Alabama, Georgia, North Carolina, South Carolina, Virginia)
78 USA
Conservation Status: IUCN R
Synonyms: hidden
= Isoetes piedmontana (N.Pfeiff.) C.F.Reed; Phytologia 12: 392 (1965)
= Isoetes virginica var. piedmontana N.Pfeiff.; Bull. Torrey Bot. Club 66: 411 (1939)
species :
Isoetes viridimontana M.A.Rosenthal & W.C.Taylor; Amer. Fern J. 104(1): 8 (2014)
Distribution: USA (Vermont)
75 USA
species :
Isoetes weberi Herter ex U.Weber; Hedwigia 63: 255-256, fig. 44-45 (1922)
Distribution: Brazil (Rio Grande do Sul); Uruguay (Rocha)
84 BRA 85 URU
Synonyms: hidden
= Isoetes weberi Herter; Beih. Bot. Centralbl. 39(2): 254 (1922), nomen
species :
Isoetes welwitschii A.Braun ex Kuhn; Filic. Afr. 196 (1868)
Distribution: Ghana; Nigeria; Central African Republic; Sudan; Ethiopia; Eritrea; Kenya; Tanzania; Angola; Zambia; Botswana; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal); Madagascar
22 GHA NGA 23 CAF 24 ERI ETH SUD 25 KEN TAN 26 ANG ZAM 27 BOT SAF 29 MDG
Synonyms: hidden
= Isoetes natalensis Baker; Handb. Fern-Allies 132 (1887)
= Calamaria natalensis (Baker) Kuntze; Revis. Gen. Pl. 2: 828 (1891)
= Calamaria welwitschii (A.Braun ex Kuhn) Kunze; Revis. Gen. Pl. 2: 828 (1883)
species :
Isoetes wormaldii R.Sim; Trans. S. Afr. Phil. Soc. 16: 299, t. 5 (1906)
Distribution: South Africa (E-Cape: Grahamstown to East London)
27 SAF
Conservation Status: IUCN E
species :
Isoetes xiangfei Y.H.Yan, Y.F.Gu & J.P.Shu; Guihaia 42(10): 1626 (2022)
Distribution: China (Hunan)
36 CHI
species :
Isoetes yunguiensis C.F.Wang & W.C.Taylor; Novon 12(4): 587-591 (2002)
Distribution: China (Guizhou, Yunnan)
36 CHI
species :
Isoetes ×altonharvillii Musselman & R.D.Bray; Castanea 60(3): 246 (1995)
Distribution: USA (Delaware, Virginia)
78 USA
Remarks: Isoetes caroliniana x engelmannii
species :
Isoetes ×angeloi D.F.Brunton & K.L.McIntosh; Bot. Lett. 170(2): 281 (2022)
Distribution: Sicily
13 SIC
Remarks: Isoetes gymnocarpa x longissima
species :
Isoetes ×blondeaui D.F.Brunt. & P.C.Sokoloff; Botany (Ottawa) 98(10): 558 (2020)
Distribution: Canada (Quebec)
72 CDN
Remarks: Isoetes echinospora x laurentiana
species :
Isoetes ×brittonii D.F.Brunt. & W.C.Taylor; Amer. Fern J. 80: 82-89 (1990)
Distribution: USA (Connecticut, New Jersey, New York, Pennsylvania, South Carolina, Virginia)
75 USA 78 USA
Remarks: Isoetes engelmanii x riparia
species :
Isoetes ×brochonii Motelay; Actes Soc. Linn. Bord. 45: 45 (1892)
Distribution: Andorra; SW-France
12 ADO FRA
Remarks: Isoetes echinospora x creussensis
Synonyms: hidden
= Isoetes lacustris subsp. brochonii (Motelay) R.C.V.Douin; Bonnier, Fl. Ill. France 12: 117 (1934)
species :
Isoetes ×bruntonii Knepper & Musselman; Amer. Fern J. 86(1): 9 (1996)
Distribution: USA (Virginia)
78 USA
Remarks: Isoetes engelmannii x hyemalis
species :
Isoetes ×carltaylori Musselman., R.D.Bray & D.A.Knepper; Canad. J. Bot. 75(2): 301-309 (1997)
Distribution: USA
Remarks: Isoetes acadiensis x engelmannii
species :
Isoetes ×dodgei A.A.Eaton; Fern Bull. 6: 6 (1898)
Distribution: Canada (Ontario, Québec); USA (New Hampshire, New Jersey, New York, Pennsylvania, Vermont)
72 CDN 75 USA
Remarks: Isoetes echinospora x septentrionalis
Synonyms: hidden
= Isoetes canadensis A.A.Eaton; Proc. U.S. Nat. Herb. 23(1226): 650 (1901)
= Isoetes riparia f. canadensis (Engelm.) Proctor; Amer. Fern J. 39: 119 (1949)
= Isoetes riparia var. canadensis Engelm.; Trans. Acad. Sci. St. Louis 4: 383 (1882)
species :
Isoetes ×eatonii R.Dodge; Bot. Gaz. 23: 16, t. 4-5 (1897), pro sp.
Distribution: Canada (Ontario); USA (Connecticut, Massachusetts, New Hampshire, New Jersey, New York, Pennsylvania, Vermont)
72 CDN 75 USA
Remarks: Isoetes echinospora x engelmannii
Synonyms: hidden
= Isoetes eatoni R.Dodge; Ferns & Fern-Allies of New England 39 (1896)
= Isoetes ×gravesii A.A.Eaton; Fernwort Papers 2: 14 (1900)
= Isoetes eatoni f. gravesii (A.A.Eaton) C.F.Reed; Amer. Fern J. 35: 82 (1945)
= Isoetes eatoni var. gravesii (A.A.Eaton) Clute; Fern Bull. 16: 55 (1908)
= Isoetes valida var. gravesii (A.A.Eaton) Clute; Fern-Allies 243 (1905)
species :
Isoetes ×echtuckeri D.F.Brunt. & D.M.Britton; Canad. J. Bot. 77(11): 1662-1668 (1999)
Distribution: Canada (New Brunswick, Nova Scotia, Ontario, Québec); USA (Maine)
72 CDN 75 USA
Remarks: Isoetes tenella x tuckermanii
species :
Isoetes ×fernaldii D.F.Brunt. & P.C.Sokoloff; Webbia 75(1): 44 (2020)
Distribution: USA (Virginia)
78 USA
Remarks: Isoetes engelmannii x hyemalis
species :
Isoetes ×foveolata A.A.Eaton ex R.Dodge; Ferns & Fern-Allies of New England 38 (1896)
Distribution: Canada (Ontario); USA (Connecticut, Massachusetts, New Hampshire)
72 CDN 75 USA
Remarks: Isoetes engelmannii x tuckermanii
species :
Isoetes ×gopalkrishnae Sarv.K.Singh, P.K.Shukla & N.K.Dubey; Phytotaxa 362(1): 71 (2018)
Distribution: India (Uttar Pradesh)
40 IND
Remarks: Isoetes coromandelina x sampathkumarnii
species :
Isoetes ×harveyi A.A.Eaton; Fernwort Pap. 11 (1900)
Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario); USA (Maine)
72 CDN 75 USA
Remarks: Isoetes lacustris x tuckermanii
Synonyms: hidden
= Isoetes tuckermanii var. harveyi (A.A.Eaton) Clute; Fern Allies 260 (1905)
species :
Isoetes ×herb-wagneri W.C.Taylor; Amer. Fern J. 92(2): 162 (2002)
Distribution: USA (Montana)
73 USA
Remarks: Isoetes echinospora x bolanderi
species :
Isoetes ×heterospora A.A.Eaton; Fernwort Papers, 2: 8 (1900), pro sp.
Distribution: USA (New York)
75 USA
Remarks: Isoetes lacustris x tuckermannii
Synonyms: hidden
= Isoetes tuckermani var. heterospora (A.A.Eaton) Clute; Fern-Allies 226, 260 (1905)
= Isoetes macrospora var. heterospora A.A.Eaton; Rhodora 10: 42 (1908)
= Isoetes macrospora f. heterospora (A.A.Eaton) Clute; Fern Bull. 16: 56 (1908)
species :
Isoetes ×hickeyi W.C.Taylor & Luebke; Amer. Fern J. 78(1): 6 (1988)
Distribution: Scotland; Canada (Ontario, Saskatchewan); USA (Wisconsin)
10 GRB 71 CDN 72 CDN 74 USA
Remarks: Isoetes echinospora x lacustris
species :
Isoetes ×jeffreyi D.M.Britton & D.F.Brunt.; Canad. J. Bot. 70: 447-452 (1992)
Distribution: Canada (Ontario, Québec)
72 CDN
Remarks: Isoetes lacustris x riparia
species :
Isoetes ×jermyi D.F.Brunt. & Rumsey; Bot. Lett. 168(4): 507 (2021)
Distribution: Great Britain; France
10 GRB 12 FRA
Remarks: Isoetes echinospora x lacustris
species :
Isoetes ×kareniae D.F.Brunt. & P.C.Sokoloff; Webbia 75(1): 38 (2020)
Distribution: USA (Pennsylvania, Virginia)
75 USA 78 USA
Remarks: Isoetes appalachiana x engelmannii
species :
Isoetes ×marensis D.M.Britton & D.F.Brunt.; Canad. J. Bot. 73(9): 1352 (1995)
Distribution: Canada (British Columbia)
71 CDN
Remarks: Isoetes howellii x maritima
species :
Isoetes ×michinokuana M.Takamiya, M.Watanaba & K.Ono; Acta Phytotax. Geobot. 48(2): 109 (1997 publ. 1998)
Distribution: Japan
38 JAP
Remarks: Isoetes japonica x pseudojaponica
species :
Isoetes ×novae-angliae D.F.Brunt. & D.M.Britton; Rhodora 108(935): 238 (2006)
Distribution: USA (Massachusetts)
75 USA
Remarks: Isoetes septentrionalis x tuckermannii
Synonyms: hidden
= Isoetes canadensis var. robbinsii A.A.Eaton; Rhodora 5: 279 (1903)
= Isoetes chapmanii var. robbinsii (A.A.Eaton) M.Broun; Index N. Amer. Ferns 199 (1938)
= Isoetes dodgei var. robbinsii A.A.Eaton; Rhodora 10: 42 (1908)
= Isoetes riparia var. robbinsii (A.A.Eaton) Proctor; Amer. Fern J. 39: 119 (1949)
species :
Isoetes ×paratunica D.F.Brunt., Mochalova & A.A.Bobrov; Amer. Fern J. 105(2): 109, figs. 5-7 (2015)
Distribution: Russian Far East (Kamchatka)
31 RFE
Remarks: Isoetes asiatica x maritima
species :
Isoetes ×pseudotruncata D.M.Britton & D.F.Brunt.; Canad. J. Bot. 74(1): 58 (1996)
Distribution: Alaska; Canada (Alberta, British Columbia)
70 ASK 71 CDN
Remarks: Isoetes echinospora x maritima
species :
Isoetes ×robusta (Engelm.) D.F.Brunt.; Rhodora 120(984): 307 (2019)
Distribution: Canada; USA
Synonyms: hidden
= Isoetes braunii f. robusta (Engelm.) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 173 (1922)
= Isoetes echinospora f. robusta (Engelm.) Clute; Fern Allies 258 (1905)
= Isoetes echinospora var. robusta Engelm.; Trans. Acad. Sci. St. Louis 4: 380 (1882)
= Isoetes muricata f. robusta (Engelm.) C.F.Reed; Amer. Fern J. 35: 82 (1945)
= Isoetes tenella f. robusta (Engelm.) Rothm.; Feddes Repert. Regni Veg. 54: 73 (1944)
species :
Isoetes ×truncata (A.A.Eaton) Clute; Fern-Allies 260 (1905)
Name: Vancouver Quillwort (EN)
Distribution: SE-Alaska; Canada (Alberta)
70 ASK 71 CDN
Remarks: Isoetes maritima x occidentalis
Synonyms: hidden
= Isoetes echinospora var. truncata A.A.Eaton; Working List North American Pteridophytes 27 (1901)
= Isoetes muricata subsp. truncata (A.A.Eaton) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982)
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A complete, synonymic checklist of the Higher Plants of the World


Version 18.4 (Feb. 11th, 2024)

Covering 354,025 accepted species,
9,095 hybrids, 47,879 accepted infraspecific taxa and 995,101 synonyms,
in total 1,406,100* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life