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492.0000 Family Rubiaceae Juss.; Gen. Pl. [Jussieu] 196 (1789), nom. cons.
589 genera, ca. 14.200 spp. mostly tropics, comparably few temperate. Phylogeny: Razafimandimbison & Rydin (2024)
Species listed in database: 14370; hybrids: 47
= Aparinaceae Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 2: 3, 38 (1813-1829)
= Asperulaceae Cham. ex Spenn.; Handb. Angew. Bot. 2: 493 (1835)
= Catesbaeaceae Martinov; Tekhno-Bot. Slovar 111 (1820)
= Cephalanthaceae Raf.; Ann. Gén. Sci. Phys. 6: 86 (1820)
= Cinchonaceae Batsch; Tab. Affin. Regni Veg. 234 (1802)
= Coffeaceae Batsch; Tab. Affin. Regni Veg. 233 (1802)
= Coutareaceae Martinov; Tekhno-Bot. Slovar 168 (1820)
= Dialypetalanthaceae Rizzini & Occhioni; Lilloa 17: 253 (1948), nom. cons.
= Galiaceae Lindl.; Intr. Nat. Syst. Bot., ed. 2: 249 (1836)
= Gardeniaceae Dumort.; Anal. Fam. Pl. 29, 32 (1829)
= Guettardaceae Batsch; Tab. Affin. Regni Veg. 235 (1802)
= Hameliaceae Mart.; Consp. Regn. Veg. [Martius] 31 (1835)
= Hedyotidaceae Dumort.; Commentat. Bot. [Dumortier] 57 (1822)
= Henriqueziaceae Bremek.; Acta Bot. Neerl. 6 (Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 141): 371 (1957)
= Houstoniaceae Raf.; Good Book 21 (1840)
= Hydrophylacaceae Martinov; Tekhno-Bot. Slovar 318 (1820)
= Lippayaceae Meisn.; Pl. Vasc. Gen. [Meisner], Tab. Diagn. 156, Comm. 112 (1838)
= Lygodisodeaceae Bartl.; Ord. Nat. Pl. 123, 207 (1830)
= Naucleaceae Wernham; New Phytol. 11: 225 (1912)
= Nonateliaceae Martinov; Tekhno-Bot. Slovar 420 (1820)
= Operculariaceae Juss. ex Perleb; Vers. Arzneikr. Pfl. 207 (1818)
= Pagamaeaceae Martinov; Tekhno-Bot. Slovar 447 (1820)
= Psychotriaceae F.Rudolphi; Syst. Orb. Veg. 49 (1830)
= Randiaceae Martinov; Tekhno-Bot. Slovar 534 (1820)
= Sabiceaceae Martinov; Tekhno-Bot. Slovar 555 (1820)
= Spermacoceaceae Bercht. & J.Presl; Přir. Rostlin 256 (1820)
= Theligonaceae Dumort.; Anal. Fam. Pl. 15, 17 (1829), nom. cons.
492.3605 Genus Leptodermis Wall.; Fl. Ind. [Carey & Wallich ed.] 2: 191 (1824)
Distribution: Himalayas (mostly China), N-India, Indochina
Species listed in database: 46

Search for: Leptodermis pilosa-var.-spicatiformis

Nothing found!
species :
Leptodermis amoena Spring.; Edinburgh J. Bot. 53(1): 99 (1996)
Distribution: Bhutan; NE-India (Arunachal Pradesh)
40 BHU IND
species :
Leptodermis beichuanensis H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 19 (1999)
Distribution: China (Sichuan)
36 CHI
species :
Leptodermis brevisepala H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 21 (1999)
Distribution: China (Sichuan)
36 CHI
species :
Leptodermis buxifolia H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 20 (1999)
Distribution: China (Sichuan, Gansu, Shaanxi)
36 CHI
Synonyms: hidden
= Leptodermis buxifolia f. strigosa H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 21 (1999)
species :
Leptodermis coriaceifolia Tao Chen; Fl. China 19: 201 (2011)
Distribution: China (Yunnan, Guangxi); Vietnam
36 CHI 41 VIE
species :
Leptodermis crassifolia Collett & Hemsl.; J. Linn. Soc., Bot. 28: 67 (1891)
Distribution: Myanmar (Shan); Thailand
41 MYA THA
species :
Leptodermis dielsiana H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 151 (1922)
Distribution: China (Yunnan)
36 CHI
species :
Leptodermis diffusa Batalin; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 373 (1894)
Distribution: China (SE-Gansu, S-Sichuan)
36 CHI
Synonyms: hidden
= Leptodermis diffusa var. umbellata (Batalin) Z.Y.Zhang; Fl. Tsinling. 1(5): 10 (1985)
= Leptodermis umbellata Batalin; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 374 (1894)
species :
Leptodermis forrestii Diels; Notes Roy. Bot. Gard. Edinburgh 5: 274 (1912)
Distribution: China (Sichuan, NW-Yunnan); SE-Tibet
36 CHI TIB
species :
Leptodermis glomerata Hutch.; Pl. Wilson. [Sargent] 3: 406 (1916)
Distribution: China (Yunnan)
36 CHI
species :
Leptodermis gracilis C.E.C.Fisch.; Bull. Misc. Inform. Kew 1940: 293 (1940 publ. 1941)
Distribution: SE-Tibet; Myanmar
36 TIB 41 MYA
variant :
var. longiflora H.S.Lo; Fl. Reipubl. Popularis Sin. 71(2): 338 (1999)
Distribution: China (Sichuan)
36 CHI
species :
Leptodermis griffithii Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 198 (1880)
Distribution: NE-India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim); Myanmar (Chin, Shan)
40 IND 41 MYA
species :
Leptodermis handeliana H.J.P.Winkl.; Symb. Sin. 7: 1025 (1936)
Distribution: China (Sichuan)
36 CHI
species :
Leptodermis hechiensis R.J.Wang; Phytotaxa 349(1): 68 (2018)
Distribution: China (Guangxi)
36 CHI
species :
Leptodermis hirsutiflora H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 19 (1999)
Distribution: Tibet
36 TIB
variant :
var. ciliata H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 20 (1999)
Distribution: SE-Tibet
36 TIB
species :
Leptodermis kumaonensis Parker; Indian Forester 48: 576 (1922)
Distribution: S-Tibet; Nepal; Bhutan; N-India (Uttarakhand, Kumaon, Garhwal)
36 TIB 40 BHU IND NEP
species :
Leptodermis lanata H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 19 (1999)
Distribution: China (NW-Yunnan)
36 CHI
species :
Leptodermis lanceolata Wall.; Fl. Ind. [Carey & Wallich ed.] 2: 191 (1824)
Distribution: Pakistan (Mirpur); Nepal; N-India (Himachal Pradesh, Kashmir, Meghalaya, Sikkim, Uttar Pradesh)
40 IND NEP PAK
Synonyms: hidden
= Hamiltonia fruticosa D.Don; Prodr. Fl. Nepal. 137 (1825)
= Hamiltonia lanceolata (Wall.) Baill.; Hist. Pl. [Baillon] 7: 273 (1880)
= Hamiltonia suaveolens D.Don; Prodr. Fl. Nepal. 137 (1825), nom. illeg.
species :
Leptodermis lecomtei Pit.; Fl. Indo-Chine [P.H.Lecomte et al.] 3: 409 (1924)
Distribution: Vietnam
41 VIE
species :
Leptodermis limprichtii H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg., Beih. 12: 490 (1922)
Distribution: China (C-Sichuan)
36 CHI
species :
Leptodermis ludlowii Spring.; Edinburgh J. Bot. 53(1): 102 (1996)
Distribution: Tibet; Bhutan; N-India (Sikkim, Chumbi)
36 TIB 40 BHU IND
Synonyms: hidden
= Leptodermis tubicalyx H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 17 (1999)
species :
Leptodermis oblonga Bunge; Mém. Acad. Imp. Sci. St.-Pétersbourg Divers Savans 2: 108 (1835)
Distribution: China (Gansu, Hebei, N-Henan, Ningxia, Shaanxi, Shanxi, Sichuan); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Hamiltonia oblonga (Bunge) Franch.; Nouv. Arch. Mus. Hist. Nat., Sér. 2, 6: 34 (1883)
= Leptodermis chanetii H.Lév.; Bull. Acad. Int. Géogr. Bot. 25: 47 (1915)
= Leptodermis huashanica H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 16 (1999)
= Leptodermis mongolica Buhse ex K.Schum.; Nat. Pflanzenfam. [Engler & Prantl] IV, 4: 126 (1891), nomen
= Leptodermis oblonga var. leptophylla H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 161 (1922)
species :
Leptodermis ordosica H.C.Fu & E.W.Ma; Fl. Intramongol. 5: 413 (1980 publ. 1981)
Distribution: China (Nei Mongol)
36 CHI
species :
Leptodermis parkeri Dunn; Bull. Misc. Inform. Kew 1920: 206 (1920)
Distribution: India (Himachal Pradesh, Punjab)
40 IND
species :
Leptodermis parvifolia Hutch.; Pl. Wilson. [Sargent] 3: 404 (1916)
Distribution: China (Sichuan)
36 CHI
species :
Leptodermis pilosa Diels; Notes Roy. Bot. Gard. Edinburgh 5: 276 (1912)
Distribution: China (Shaanxi, W-Hubei, Sichuan, Yunnan); SE-Tibet
36 CHI TIB
Synonyms: hidden
= Hamiltonia pilosa Franch. ex Diels; Notes Roy. Bot. Gard. Edinburgh 5: 276 (1912)
= Leptodermis fusca H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 158 (1922)
= Leptodermis microphylla H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 159 (1922)
= Leptodermis pilosa var. microphylla H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg., Beih. 12: 389 (1922)
variant :
var. acanthoclada H.S.Lo ex X.Y.Wen & Q.Lin; Acta Phytotax. Sin. 45(3): 410 (2007)
Distribution: China (W-Sichuan); Tibet
36 CHI TIB
Synonyms: hidden
= Leptodermis pilosa var. acanthoclada H.S.Lo; Fl. Reipubl. Popularis Sin. 71(2): 338 (1999)
variant :
var. glabrescens H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 160 (1922)
Distribution: China (?S-Sichuan, Yunnan)
36 CHI
Synonyms: hidden
= Leptodermis mairei H.Lév.; Repert. Spec. Nov. Regni Veg. 13: 179 (1914)
variant :
var. spicatiformis H.S.Lo ex X.Y.Wen & Q.Lin; Acta Phytotax. Sin. 45(3): 411 (2007)
Distribution: China (S-Gansu, SW-Shaanxi)
36 CHI
Synonyms: hidden
= Leptodermis pilosa var. spicatiformis H.S.Lo; Fl. Reipubl. Popularis Sin. 71(2): 338 (1999)
species :
Leptodermis potaninii Batalin; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 319 (1898)
Distribution: China (S-Shanxi, W-Hubei, Sichuan, W-Guizhou, Yunnan); Myanmar
36 CHI 41 MYA
Synonyms: hidden
= Leptodermis nigricans H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg., Beih. 12: 489 (1922)
variant :
var. angustifolia H.S.Lo; Fl. Reipubl. Popularis Sin. 71(2): 338 (1999)
Distribution: China (Yunnan)
36 CHI
variant :
var. glauca (Diels) H.J: P.Winkl.; Repert. Spec. Nov. Regni Veg., Beih. 12: 490 (1922)
Distribution: China (SW-Sichuan, SW-Guizhou, Yunnan)
36 CHI
Synonyms: hidden
= Hamiltonia glauca Franch. ex Diels; Notes Roy. Bot. Gard. Edinburgh 5: 275 (1912)
= Leptodermis esquirolii H.Lév.; Repert. Spec. Nov. Regni Veg. 9: 324 (1911)
= Leptodermis glauca Diels; Notes Roy. Bot. Gard. Edinburgh 5: 275 (1912)
= Leptodermis motsouensis H.Lév.; Bull. Acad. Int. Géogr. Bot. 25: 47 (1915)
variant :
var. tomentosa H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 153 (1922)
Distribution: China (SW-Sichuan, Yunnan)
36 CHI
Synonyms: hidden
= Leptodermis potanini var. rufa H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 154 (1922)
= Leptodermis tongchouanensis H.Lév.; Bull. Géogr. Bot. 25: 47 (1915)
species :
Leptodermis pulchella Yatabe; Bot. Mag. (Tokyo) 4: (2) t. 12 (1890)
Distribution: Japan (Honshu, Shikoku)
38 JAP
species :
Leptodermis pumila H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 18 (1999)
Distribution: China (Yunnan)
36 CHI
species :
Leptodermis purdomii Hutch.; Pl. Wilson. [Sargent] 3: 405 (1916)
Distribution: China (S-Gansu, Sichuan)
36 CHI
species :
Leptodermis rehderiana H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 157 (1922)
Distribution: China (Yunnan)
36 CHI
species :
Leptodermis riparia R.Parker; Indian Forester 50: 398 (1924)
Distribution: NW-India (Himachal Pradesh, Uttarakhand, ?Uttar Pradesh)
40 IND
species :
Leptodermis scabrida Hook.f.; Fl. Brit. India [J.D.Hooker] 3: 199 (1881)
Distribution: S-Tibet; NE-India (Arunachal Pradesh)
36 TIB 40 IND
species :
Leptodermis schneideri H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 156 (1922)
Distribution: China (?Sichuan, Yunnan); Tibet
36 CHI TIB
Synonyms: hidden
= Leptodermis schneideri var. hutchinsonii H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 157 (1922)
species :
Leptodermis scissa H.J.P.Winkl.; Symb. Sin. 7: 1024 (1936)
Distribution: China (Sichuan, Yunnan)
36 CHI
species :
Leptodermis stapfiana H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 164 (1922)
Distribution: W-Nepal; Bhutan; NE-India (Assam)
40 BHU IND NEP
species :
Leptodermis tomentella H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 159 (1922)
Distribution: China (Yunnan)
36 CHI
species :
Leptodermis trifida Craib; Bull. Misc. Inform. Kew 1914: 129 (1914)
Distribution: Thailand
41 THA
species :
Leptodermis velutiniflora H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 17 (1999)
Distribution: China (Sichuan)
36 CHI
variant :
var. tenera H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 17 (1999)
Distribution: China (Sichuan, Yunnan); Tibet
36 CHI TIB
species :
Leptodermis vestita Hemsl.; J. Linn. Soc., Bot. 23: 390 (1888)
Distribution: China (N-Guangdong)
36 CHI
Synonyms: hidden
= Leptodermis ovata H.J.P.Winkl.; Repert. Spec. Nov. Regni Veg. 18: 162 (1922)
species :
Leptodermis virgata Edgew. ex Aitch.; Cat. Punjab Pl. 70, et ex Fl. Brit. India [J.D.Hooker] 3: 198 (1881)
Distribution: Pakistan (Hazara, Salt Range, Murree, Mirpur); NW-India (Himachal Pradesh, Jammu & Kashmir, Punjab, Poonch)
40 IND PAK
species :
Leptodermis wardii C.E.C.Fisch. & Kaul; Bull. Misc. Inform. Kew 1940: 188 (1940)
Distribution: Myanmar (Kachin)
41 MYA
species :
Leptodermis wilsonii Diels; Notes Roy. Bot. Gard. Edinburgh 5: 275 (1912)
Distribution: China (Sichuan, Yunnan)
36 CHI
species :
Leptodermis xizangensis H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 16 (1999)
Distribution: Tibet
36 TIB
species :
Leptodermis yangshuoensis Tao Chen; Fl. China 19: 210 (2011)
Distribution: China (NE-Guangxi)
36 CHI
species :
Leptodermis yui H.S.Lo; J. Trop. Subtrop. Bot. 7(1): 15 (1999)
Distribution: China (SW-Sichuan)
36 CHI
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A complete, synonymic checklist of the Higher Plants of the World


Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life