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001.0000 Family Lycopodiaceae P.Beauv. ex Mirb.; Hist. Nat. Vég. [Lamarck & Mirbel] 4: 293 (1802)
17 genera, ca. 470 spp. Phylogeny: Chen et al. (2021).
Species listed in database: 477; hybrids: 20
= Huperziaceae Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 66: 236 (1962), nom. nov.
= Urostachyaceae Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 58 (1944) (nom. illeg.)
= Phylloglossaceae Kunze; Bot. Zeitung (Berlin) 1: 722 (1843)
= Lycopodiellaceae Val.N.Tikhom.; Novosti Sist. Vyssh. Rast. 49: 149 (2018)
001.3006 Genus Lycopodium L.; Sp. Pl. [Linnaeus] 2: 1100 (1753) s. str.
Distribution: Mainly temperate and subarctic regions
Species listed in database: 7
= Clopodium Raf.; Anal. Nat. 205 (1815) (nom. inval.)
= Oxynemum Raf.; Amer. Monthly Mag. & Crit. Rev. 2: 44 (1817)
species :
Lycopodium clavatum L.; Sp. Pl. [Linnaeus] 2: 1101 (1753)
subspecies :
subsp. clavatum
Name: Keulen-Bärlapp (DE), Common Clubmoss, Running Cedar (EN)
Distribution: Iceland; England (England, Wales, Scotland, Orkney Isl., Shetland Isl., Isle of Man); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Bulgaria; Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Ukraine; Moldova; Guinea; Sierra Leone; Ivory Coast; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Equatorial Guinea (Rio Muni); ?Gabon; Congo [Brazzaville]; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov., W-Cape Prov., E-Free State); Lesotho; Eswatini [Swaziland]; Madagascar; Comores (Grande Comore); Mauritius; La Réunion; Siberia (Altai, Buryatia, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Primorye, Sakhalin); Kazakhstan; Northern Caucasus; Georgia [Caucasus]; Turkey (Anatolia); China (Heilongjiang, Jilin, Liaoning, Nei Mongol); Mongolia; North Korea; South Korea (incl. Cheju Isl.); Japan; Alaska; Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon; USA (California, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Idaho, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Montana, North Carolina, New Hampshire, New Jersey, New Mexico, New York, Ohio, Oregon, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia); Ile Amsterdam; Ile St. Paul
10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO FRA POR SPA 13 BIH BUL CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR EST LAT LIT MLD RUS UKR 22 GUI IVO NGA SIE 23 BIO BUR CMN CON EQG RWA SAO ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 LES SAF SWZ 29 COM MAU MDG REU 30 SIB 31 RFE 32 KAZ 33 GEO NCS 34 TUR 36 CHI 37 MON 38 JAP KOR NKO 70 ASK 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA 91 ASP
Synonyms: hidden
= Lepidotis ciliata P.Beauv.; Prodr. Aethéogam. 108 (1805)
= Lepidotis clavata (L.) P.Beauv.; Prodr. Aethéogam. 108 (1805)
= Lycopodium aequale Charit.; Form. Rastitel´n. Pokrova Basseyna Irtysha 76, 87: f. 8.5 (2022) (tentative synonym)
= Lycopodium amplum Charit.; Form. Rastitel´n. Pokrova Basseyna Irtysha 77, 87: f. 8.7 (2022)
= Lycopodium aristatum var. robustius Grev. & Hook.; Bot. Misc. 2: 376 (1831)
= Lycopodium attenuatum Charit.; Form. Rastitel´n. Pokrova Basseyna Irtysha 77 (2022) (tentative synonym)
= Lycopodium ciliatum (P.Beauv.) Sw.; Syn. Fil. (Kiliae) [O.Swartz] 179 (1806), nom. illeg.
= Lycopodium clavatum f. brevipedunculatum Louis-Marie; Rev. Oka Agron. Inst. Agric. 14: 144, fig. 10, no. 2 (1940)
= Lycopodium clavatum f. decapitatum Louis-Marie; Rev. Oka Agron. Inst. Agric. 14: 144, fig. 10, no. 1 (1940)
= Lycopodium clavatum f. frondescens Vict.; Contr. Lab. Bot. Univ. Montréal 3: 25 (1925)
= Lycopodium clavatum f. frondosum Vict.; Contr. Lab. Bot. Univ. Montréal 3: 25 (1925)
= Lycopodium clavatum f. robustinus Nessel & Hoehne; Arch. Bot. São Paulo 1: 439 (1927)
= Lycopodium clavatum f. sterile House; Bull. New York State Mus. Nat. Hist. 243-244: 19 (1923)
= Lycopodium clavatum f. trilobum (Farw.) M.Broun; Index No. Amer. Ferns: 107 (1938)
= Lycopodium clavatum subvar. trilobum Farw.; Amer. Fern J. 27: 18 (1937)
= Lycopodium clavatum var. asiaticum Ching; Acta Bot. Yunnan. 4(3): 224 (1982)
= Lycopodium clavatum var. borbonicum Bory; Voy. Îles Afrique 2: 205 (1804)
= Lycopodium clavatum var. distachyum Spring; Flora 21(1): 171 (1838), nom. inval.
= Lycopodium clavatum var. kiboanum Nessel; Repert. Spec. Nov. Regni Veg. 36(12-15): 192, t. 177 (1934)
= Lycopodium clavatum var. laurentianum Vict.; Contr. Lab. Bot. Univ. Montreal 3: 23 (1925)
= Lycopodium clavatum var. macrostachyum Schur; Oesterr. Bot. Z. 8(2): 64 (1858)
= Lycopodium clavatum var. microstachyum Schur; Oesterr. Bot. Z. 8(2): 64 (1858)
= Lycopodium clavatum var. montanum Stehlé; Mém. Publiés Soc. Bot. France 1953-1954: 44 (1954)
= Lycopodium clavatum var. paucidivisum Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15 (Mém. Lycopod.): 91 (1842)
= Lycopodium clavatum var. preslianum Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15 (Mém. Lycopod.): 90 (1842)
= Lycopodium clavatum var. robustius (Grev. & Hook.) Nakai; Bot. Mag. (Tokyo) 39: 197 (1925)
= Lycopodium clavatum var. serpens (C.Presl) Spring; Flora 21(1): 172 (1838)
= Lycopodium clavatum var. sitchaense Spring; Flora 21(1): 173 (1838)
= Lycopodium clavatum var. subremotum Vict.; Contr. Lab. Bot. Univ. Montreal 3: 24 (1925)
= Lycopodium clavatum var. trichiatum (Bory) Nessel; Bärlappgewächse 301, t. 69, f. 47-48 (1939)
= Lycopodium compressum Charit.; Form. Rastitel´n. Pokrova Basseyna Irtysha 77, 87: f. 8.6 (2022) (tentative synonym)
= Lycopodium fasciculatum Charit.; Form. Rastitel´n. Pokrova Basseyna Irtysha 75, 87: f. 8.3 (2022) (tentative synonym)
= Lycopodium officinale Neck.; Meth. Musc. 150 (1771)
= Lycopodium preslii Grev. & Hook.; Bot. Misc. 2: 377 (1831)
= Lycopodium ramosum Charit.; Dopoln. Fl. i Rastitel´n. Yuga Zapadnoi Sibiri 94 (-95) (2020) (tentative synonym)
= Lycopodium raroramosum Charit.; Form. Rastitel´n. Pokrova Basseyna Irtysha 75 (-76, 87: f. 8.4) (2022) (tentative synonym)
= Lycopodium serpens C.Presl; Reliq. Haenk. 1(1): 81 (1825) (non Lycopodium serpens Desv. ex Poir. (1814))
= Lycopodium solidum Charit.; Form. Rastitel´n. Pokrova Basseyna Irtysha 74 (-75, 87: f. 8.1) (2022) (tentative synonym)
= Lycopodium trichiatum Bory; Voy. Îles Afrique 1: 350 (1804)
= Lycopodium trichiatum var. borbonicum (Bory) Spring; Mém. Acad. Roy. Sci. Belgique 15 (Monogr. Lycopod. 1): 92 (1842)
= Lycopodium trichophyes Spreng.; Syst. Veg., ed. 16 [Sprengel] 4(1): 13 (1827)
= Urostachys plutonis Herter; Index Lycopod. 75 (1949)
subspecies :
subsp. contiguum (Klotzsch) B.Øllg.; Fl. Ecuador 33: 126 (1988)
Distribution: Costa Rica; Panama; Colombia (Antioquia, Boyacá, Caldas, Casanare, Cauca, Cesar, Cundinamarca, Huila, La Guajira, Magdalena, Nariño, Norte de Santander, Risaralda, Santander, Tolima, Valle); Venezuela (Bolívar, Venezuela Distrito Federal, Merida, Tachira, Trujillo); Guyana; Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz)
80 COS PAN 82 CLM GUY VEN 83 BOL ECU PER
Synonyms: hidden
= Lycopodium clavatum var. pseudocontiguum Christ; Prim. Fl. Costaric. 3(1): 55 (1901)
= Lycopodium contiguum Klotzsch; Linnaea 18: 519 (1845)
= Lycopodium herbaceum (Spring) Hieron.; Bot. Jahrb. Syst. 34: 575 (1905)
= Lycopodium herbaceum var. majus Nessel; Repert. Spec. Nov. Regni Veg. 36: 192, pl. 177 (1934)
= Lycopodium mayoris Rosenst.; Mém. Soc. Sci. Nat. Neuchâtel 5: 56, t. 6p, f. 11 (1914)
= Lycopodium nidiforme Sessé & Moc.; Fl. Mexic., ed. 2: 239 (1894), nom. illeg.
= Lycopodium vestitum var. herbaceum Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 24: 45 (1849)
variant :
var. aristatum (Humb. & Bonpl. ex Willd.) Spring; Flora 21(1): 173 (1838)
Distribution: Mexico (Chiapas, Colima, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Oaxaca, Puebla, Queretaro, San Luis Potosi, Veracruz); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; St. Kitts; Guadeloupe; Martinique; Colombia; Venezuela (Amazonas, Anzoategui, Aragua, Barinas, Bolívar, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Monagas, Portuguesa, Sucre, Tachira, Trujillo); Ecuador; Galapagos; Peru; Brazil (Bahia, Espírito Santo, Goiás, Minas Gerais, Paraná, Rio de Janeiro, Roraima, Rio Grande do Sul, Santa Catarina, São Paulo); Paraguay (Caaguazu); Uruguay (Durazno, Maldonado); Argentina (Cordoba, Jujuy, Misiones, Salta)
79 MEX 80 COS ELS GUA HON NIC PAN 81 CUB GDL HIS JAM MAR PUE STK 82 CLM VEN 83 ECU GAL PER 84 BRA 85 ARG PAR URU
Synonyms: hidden
= Lycopodium aristatum Humb. & Bonpl. ex Willd.; Sp. Pl., ed. 4 [Willdenow] 5(1): 17 (1810)
= Lycopodium aristatum var. desvauxianum (Spring) Spring; Fl. Bras. [Martius] 1(2): 114 (1840)
= Lycopodium aristatum var. incurvum Grev. & Hook.; Bot. Misc. 2: 376 (1831)
= Lycopodium clavatum f. incurvum (Grev. & Hook.) Nessel; Bärlappgewächse 305 (1939)
= Lycopodium clavatum var. desvauxianum Spring; Flora 21(1): 173 (1838)
= Lycopodium clavatum var. equisetoides Schwacke; Arch. Bot. São Paulo 1: 438 (1927)
= Lycopodium clavatum var. eriostachys (Fée) Nessel & Hoehne; Arch. Bot. São Paulo 1: 435 (1927)
= Lycopodium clavatum var. jamaicense Spring; Flora 21(1): 174 (1838)
= Lycopodium clavatum var. minarum Christ; Pl. Nov. Mineir. 2: 41 (1900)
= Lycopodium clavatum var. piliferum (Raddi) Nessel & Hoehne; Arch. Bot. São Paulo 1: 435 (1927)
= Lycopodium clavatum var. raddianum Spring; Flora 21(1): 172 (1838)
= Lycopodium clavatum var. trichophyllum (Desv.) Nessel & Hoehne; Arch. Bot. São Paulo 1: 436 (1927)
= Lycopodium eriostachys Fée; Crypt. Vasc. Brésil 1: 224 (1869)
= Lycopodium piliferum Raddi; Pl. Bras. Nov. Gen. 1: 79 (1825)
= Lycopodium torridum Gaudich.; Voy. Uranie [Freycinet], Bot. 1: 283 (1828)
= Lycopodium trichiatum var. desvauxianum (Spring) Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15 (Mém. Lycopod.): 92 (1842)
= Lycopodium trichophyllum Desv.; Mém. Soc. Linn. Paris 6(2): 184 (1827)
species :
Lycopodium diaphanum (P.Beauv.) Sw.; Syn. Fil. (Kiliae) [O.Swartz] 179 (1806)
Distribution: Gough Isl.; Tristan da Cunha (incl. Inaccessible Isl.)
91 TDC
Synonyms: hidden
= Lepidotis diaphana P.Beauv.; Prodr. Aethéogam. 108 (1805)
species :
Lycopodium japonicum Thunb.; Syst. Veg., ed. 14 [J.A.Murray] 944 (1784)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Xinjiang, Yunnan, Zhejiang); Tibet; Japan; Taiwan; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, Tripura, Uttarakhand, West Bengal); Sri Lanka; Bangladesh; Myanmar; Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (Perak, Pahang); Borneo (Sabah, Mt. Kinabalu); Sulawesi; Philippines; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal); New Caledonia; Fiji (Viti Levu)
36 CHI TIB 38 JAP TAI 40 BAN BHU IND NEP SRL 41 CBD LAO MYA THA VIE 42 BOR MLY MOL NWG PHI SUL 43 SOL 60 FIJ NWC
Remarks: contains tropical Asian records for "L. clavatum"
Synonyms: hidden
= Lycopodium centrochinense Ching; Acta Bot. Yunnan. 4(3): 224-225 (1982) (fide Fraser-Jenkins 2008, 2017)
= Lycopodium clavatum var. nipponicum Nakai; Bot. Mag. (Tokyo) 39: 197 (1925)
= Lycopodium clavatum var. wallichianum Spring; Bull. Acad. Roy. Sci. Belgique 8: 90 (1841)
= Lycopodium divaricatum Wall. ex Hook. & Grev.; Bot. Misc. 2: 377 (1831)
= Lycopodium interjectum Ching & H.S.Kung; Acta Bot. Yunnan. 4(3): 221 (1982)
= Lycopodium japonicum f. yunnanense (Ching) W.M.Chu; Fl. Yunnan. 20: 25 (2006)
= Lycopodium japonicum var. wallichianum (Spring) Satou; Hikobia 12(3): 269 (1997)
= Lycopodium kinabaluense Ching; Acta Bot. Yunnan. 4(3): 223 (1982)
= Lycopodium pseudoclavatum Ching; Acta Bot. Yunnan. 4(3): 222-223 (1982) (fide Fraser-Jenkins 1997)
= Lycopodium pseudoclavatum var. yunnanense Ching; Acta Bot. Yunnan. 4(3): 223 (1982) (fide Fraser-Jenkins 1997)
= Lycopodium simulans Ching & H.S.Kung; Acta Bot. Yunnan. 4(3): 220-221 (1982)
= Lycopodium taliense Ching; Acta Bot. Yunnan. 4(3): 221 (1982)
= Stachygynandrum japonicum (Thunb.) P.Beauv.; Prodr. Aethéogam. 113 (1805)
species :
Lycopodium lagopus (Laest. ex C.Hartm.) Zinserl. ex Kuzen.; Fl. Murmansk. Obl. 1: 80 (1953)
Name: One-cone clubmoss
Distribution: England (Scotland); Norway; Sweden; Finland; Switzerland; Austria; N-Italy; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Kamchatka, Khabarovsk, Magadan, Primorye); Alaska; Greenland; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); St. Pierre et Miquelon; USA (Connecticut, Illinois, Massachusetts, Maine, Michigan, Minnesota, Montana, New Hampshire, New York, Ohio, Oregon, Pennsylvania, Vermont, Washington State, Wisconsin)
10 FIN GRB NOR SWE 11 AUT SWI 13 ITA 30 SIB 31 RFE 70 ASK CDN GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA
Synonyms: hidden
= Lycopodium clavatum f. furcatum (Luerss.) Vict.; Contr. Lab. Bot. Univ. Montreal 3: 25 (1925)
= Lycopodium clavatum f. monostachyum Hegi; Ill. Fl. Mitt.-Eur. [Hegi] 1: 67 (1906)
= Lycopodium clavatum subsp. lagopus (Laest.) Dostál
= Lycopodium clavatum subsp. megastachyon (Fernald & Bissell) Á.Löve & D.Löve; Nucleus (Calcutta) 1(1): 7 (1958)
= Lycopodium clavatum subsp. monostachyon (Hook. & Grev.) Selander; Acta Phytogeogr. Suec. 28: 22 (1950)
= Lycopodium clavatum var. brevispicatum Peck; Rep. (Annual) Regents Univ. State New York 54: 162 (1901)
= Lycopodium clavatum var. integerrimum Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles (Mém. Lycopod.) 15: 90 (1842)
= Lycopodium clavatum var. integrifolium (Hook. ex Goldie) Spring; Flora 21(1): 174 (1838)
= Lycopodium clavatum var. lagopus Laest. ex C.Hartm.; Handb. Skand. Fl. ed. 7: 313 (1858)
= Lycopodium clavatum var. megastachyon Fernald & Bissell; Rhodora 12(135): 53 (1910)
= Lycopodium clavatum var. monostachyon Hook. & Grev.; Bot. Misc. 2: 375 (1831)
= Lycopodium clavatum var. monostachyum Desv.; Mém. Soc. Linn. Paris 6(2): 184 (1827), nom. inval.
= Lycopodium clavatum var. monostachyum Schur; Oesterr. Bot. Z. 8(2): 64 (1858)
= Lycopodium integrifolium Hook. ex Goldie; Edinburgh New Philos. J. 6: 332 (1822)
= Lycopodium pseudoannotinum Schur; Enum. Pl. Transsilv. 826 (1866) (nom. inval.)
species :
Lycopodium papuanum Nessel; Repert. Spec. Nov. Regni Veg. 39(8): 64, t. 191a (1935)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Lycopodium venustulum Gaudich.; Voy. Uranie [Freycinet], Bot. 1: 283, t. 22 (1826)
Distribution: [Western] Samoa (Savai´i); Society Isl. (Tahiti); Austral Isl. (Rapa Iti); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
60 SAM 61 AUI SCI 63 HAW
Synonyms: hidden
= Lycopodium clavatum var. hochreutineri Nessel; Bärlappgewächse 300, t, f. 46 (1939) (nom. inval.)
= Lycopodium heterophyllum Hook. & Grev.; Icon. Filic. 1: t. 113 (1829 publ. 1831) (non Spreng. 1827)
= Lycopodium jovis Hedw.; Index Lycopod. 24 (1949)
= Lycopodium pinifolium var. hochreutineri Herter; Annuaire Conserv. Jard. Bot. Genève 15-16: 227 (1912)
= Lycopodium sirii Herter; Index Lycopod. 20: 41 (1949)
= Urostachys hochreutineri (Herter) Herter ex Nessel; Bärlappgewächse 234 (1939)
variant :
var. verticale W.H.Wagner; Contr. Univ. Michigan Herb. 22: 144 (1999)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
species :
Lycopodium vestitum Desv. ex Poir.; Encycl. [J.Lamarck et al.] Suppl. 3: 546 (1814)
Distribution: ?Venezuela; Ecuador; Peru
83 ECU PER
Synonyms: hidden
= Lycopodium albidum Baker; J. Bot. 25: 374 (1887)
= Lycopodium scariosum Hook.; Hooker´s Icon. Pl. 1: t. 89 (1836) (non G. Forst. 1786)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.04 (April 17th, 2025)

Covering 356,375 accepted species,
8,782 hybrids, 48,090 accepted infraspecific taxa and 1,002,897 synonyms,
in total 1,416,144* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

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With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

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