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V. 12.1
14.3.2021

001.0000 Family Lycopodiaceae P.Beauv. ex Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 4: 293 (1802)
(3-)16 genera, ca. 490 spp. According to Øllgaard (2012) and molecular results, all genera should be accepted.
Species listed in database: 489; hybrids: 18
= Huperziaceae Rothm.; Fedde, Repert. 66: 236 (1962), nom. nov.
= Urostachyaceae Rothm.; Fedde, Repert. Spec. Nov. 54: 58 (1944) (nom. illeg.)
= Phylloglossaceae Kunze; Bot. Zeit. (Berlin) 1: 722 (1843)
= Lycopodiellaceae Val.N.Tikhom.; Novosti Sist. Vyssh. Rast. 49: 149 (2018)
001.1001 Genus Lycopodium L.; Sp. Pl. 2: 1100 (1753) s. str.
Distribution: Mainly temperate and subarctic regions
Species listed in database: 9
= Clopodium Raf.; Anal. nat. 205 (1815) (nom. inval.)
= Oxynemum Raf.; Amer. Monthly Mag. & Crit. Rev. 2: 44 (1817)
species :
Lycopodium clavatum L.; Sp. Pl. 2: 1101 (1753)
subspecies :
subsp. clavatum
Name: Keulenbärlapp, Common Clubmoss, Running Cedar
Distribution: Austria; Belgium; Luxembourg; England; Wales; Scotland; Ireland; Bulgaria; Czech Republic; Slovakia; Denmark; Finland; France; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Netherlands; Spain; Andorra; Hungary; Iceland; Italy; Bosnia & Hercegovina; Montenegro; Croatia; Macedonia; Slovenia; Serbia; Kosovo; Portugal; Norway; Poland; Romania; Sweden; N-European Russia; C-European Russia; W-European Russia; E-European Russia; Estonia; Latvia; Lithuania; Belarus; Ukraine; Kazakhstan; Northern Caucasus; Georgia [Caucasus]; Siberia (W-Siberia, C-Siberia); Russian Far East; Mongolia; China (Heilongjiang, Jilin, Liaoning, Nei Mongol); Japan; North Korea; South Korea; Turkey (Anatolia); Guinea; Sierra Leone; Ivory Coast; Nigeria; Cameroon; ?Gabon; Bioko Isl. [Fernando Poo]; São Tomé; Equatorial Guinea (Rio Muni); D.R.Congo [Zaire]; Congo [Brazzaville]; Sudan; Ethiopia; Uganda; Rwanda; Burundi; Kenya; Tanzania; Angola; Zambia; Malawi; Mozambique; E-Zimbabwe; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov., W-Cape Prov., E-Free State); Lesotho; Eswatini [Swaziland]; Comores (Grande Comore); Madagascar; Mauritius; La Réunion; Alaska; USA (California, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Idaho, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Montana, North Carolina, New Hampshire, New Jersey, New Mexico, New York, Ohio, Oregon, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia); Canada (British Columbia, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon
10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO FRA POR SPA 13 BIH BUL CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR EST LAT LIT RUS UKR 22 GUI IVO NGA SIE 23 BIO BUR CMN CON EQG RWA SAO ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 LES SAF SWZ 29 COM MAU MDG REU 30 SIB 31 RFE 32 KAZ 33 GEO NCS 34 TUR 36 CHI 37 MON 38 JAP KOR NKO 70 ASK 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA
Synonyms: hidden
= Lepidotis ciliata P.Beauv.; Prodr. aethéogam. 108 (1805)
= Lepidotis clavata (L.) P.Beauv.; Prodr. aethéogam. 108 (1805)
= Lycopodium aristatum var. robustius Hook. & Grev.; Bot. Misc. 2: 376 (1831)
= Lycopodium ciliatum (P.Beauv.) Sw.; Syn. Fil. 179 (1805)
= Lycopodium clavatum f. brevipedunculatum Louis-Marie; Rev. Oka Agron. Inst. Agric. 14: 144, fig. 10, no. 2 (1940)
= Lycopodium clavatum f. decapitatum Louis-Marie; Rev. Oka Agron. Inst. Agric. 14: 144, fig. 10, no. 1 (1940)
= Lycopodium clavatum f. frondescens Vict.; Contr. Lab. Bot. Univ. Montréal 3: 25 (1925)
= Lycopodium clavatum f. frondosum Vict.; Contr. Lab. Bot. Univ. Montréal 3: 25 (1925)
= Lycopodium clavatum f. robustinus Nessel & Hoehne; Arch. Bot. Sao Paulo 1: 439 (1927)
= Lycopodium clavatum f. sterilis House; Bull. New York State Mus. Nat. Hist. 243-244: 19 (1923)
= Lycopodium clavatum f. trilobum (Farw.) M.Broun; Index N. Amer. Ferns 107 (1938)
= Lycopodium clavatum subvar. trilobum Farw.; Amer. Fern J. 27: 18 (1937)
= Lycopodium clavatum var. asiaticum Ching; Acta Bot. Yunnan. 4(3): 224 (1982)
= Lycopodium clavatum var. borbonicum Bory; Voy. Iles Afrique 2: 205 (1804)
= Lycopodium clavatum var. distachyum Spring; Flora 21(1): 171 (1838), nom. inval.
= Lycopodium clavatum var. kiboanum Nessel; Fedde, Repert. 36(12-15): 192, t. 177 (1934)
= Lycopodium clavatum var. laurentianum Vict.; Contr. Lab. Bot. Univ. Montreal 3: 23 (1925)
= Lycopodium clavatum var. macrostachyum Schur; (1858)
= Lycopodium clavatum var. microstachyum Schur; (1858)
= Lycopodium clavatum var. montanum Stehlé; Bull. Soc. Bot. France Mém. 1953-1954: 44 (1954)
= Lycopodium clavatum var. nipponicum Nakai; Bot. Mag. (Tokyo) 39: 197 (1925)
= Lycopodium clavatum var. paucidivisum Spring; Nouv. Mém. Acad. Roy. Sci. Bruxelles 15: 91 (1842)
= Lycopodium clavatum var. preslianum Spring; Mém. Acad. Roy. Soc. Bruxelles 15: 90 (1842)
= Lycopodium clavatum var. robustius (Hook. & Grev.) Nakai; Bot. Mag. 39(463): 197 (1925)
= Lycopodium clavatum var. serpens (C.Presl) Spring; Flora 21(1): 172 (1838)
= Lycopodium clavatum var. sitchaense Spring; Flora 21(1): 173 (1838)
= Lycopodium clavatum var. subremotum Vict.; Contr. Lab. Bot. Univ. Montreal 3: 24 (1925)
= Lycopodium clavatum var. trichiatum (Bory) Nessel; Bärlappgew. 301, t. 69, f. 47-48 (1939)
= Lycopodium clavatum var. tristachyum (Pursh) Hook.; Fl. Bot.-Amer. (Hooker) 2: 267 (1840)
= Lycopodium officinale Neck.; Meth. musc. 150 (1771)
= Lycopodium preslii Grev. & Hook.; Bot. Misc. 2: 377 (1831)
= Lycopodium serpens C.Presl; Rel. Haenk. 1(1): 81 (1825) (non Lycopodium serpens Desv. ex Poir. (1814))
= Lycopodium tamariscispica Ces.; Rendic. Accad. Reale Sci. Fis. 16: 28, 30 (1877)
= Lycopodium torridum Gaudich.; Voy. Uranie [Freycinet] 1: 283 (1828)
= Lycopodium trichiatum Bory; Voy. Iles Afrique 1: 350 (1804)
= Lycopodium trichiatum var. borbonicum (Bory) Spring; Mémoires de l’Académie Royale des Sciences, Lettres et Beaux Arts de Belgique 15: 92 (1842)
= Lycopodium trichophyes Spreng.; Syst. Veg. 4(1): 13 (1827)
= Urostachys plutonis Herter; Index Lyc. 75 (1949)
subspecies :
subsp. contiguum (Klotzsch) B.Øllg.; Fl. Ecuador 33: 126 (1988)
Distribution: Costa Rica; Panama; Colombia (Antioquia, Boyacá, Caldas, Casanare, Cauca, Cesar, Cundinamarca, Huila, La Guajira, Magdalena, Nariño, Norte de Santander, Risaralda, Santander, Tolima, Valle); Venezuela (Bolívar, Venezuela Distrito Federal, Merida, Tachira, Trujillo); Ecuador; Peru; Guyana; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz)
80 COS PAN 82 CLM GUY VEN 83 BOL ECU PER
Synonyms: hidden
= Lycopodium clavatum var. pseudocontiguum Christ; Primit. Fl. Costar. 3(1): 55 (1901)
= Lycopodium contiguum Klotzsch; Linn. 18: 519 (1844)
= Lycopodium herbaceum (Spring) Hieron.; Bot. Jahrb. Syst. 34: 575 (1905)
= Lycopodium mayoris Rosenst.; Mém. Soc. neuchateloise 5: 56, t. 6p, f. 11 (1914)
= Lycopodium vestitum var. herbaceum Spring; Mém. Acad. Roy. Soc. Bruxelles 24: 45 (1849)
variant :
var. aristatum (Humb. & Bonpl. ex Willd.) Spring; Flora 21(1): 173 (1838)
Distribution: Costa Rica; Puerto Rico; Cuba; Hispaniola; Jamaica; Galapagos; Nicaragua; Mexico (Chiapas, Colima, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Oaxaca, Puebla, Queretaro, San Luis Potosi, Veracruz); Guatemala; Honduras; El Salvador; Panama; Colombia; Venezuela (Amazonas, Anzoategui, Aragua, Barinas, Bolívar, Venezuela Distrito Federal, Falcon, Lara, Merida, Miranda, Monagas, Portuguesa, Sucre, Tachira, Trujillo); Ecuador; Peru; SE-Brazil (Minas Gerais, Espirito Santo, São Paulo, Rio de Janeiro); St. Kitts; Guadeloupe; Martinique; Argentina (Cordoba, Jujuy, Misiones, Salta); S-Brazil (Paraná, Rio Grande do Sul, Santa Catarina); Paraguay (Caaguazu)
79 MEX 80 COS ELS GUA HON NIC PAN 81 CUB GDL HIS JAM MAR PUE STK 82 CLM VEN 83 ECU GAL PER 84 BRA 85 ARG PAR
Synonyms: hidden
= Lycopodium aristatum Humb. & Bonpl. ex Willd.; Sp. Pl. ed. IV, 5: 17 (1810)
= Lycopodium aristatum var. desvauxianum (Spring) Spring; Mart., Fl. Bras. 1(2): 114 (1840)
= Lycopodium aristatum var. incurvum Hook. & Grev.; Bot. Misc. 2: 376 (1831)
= Lycopodium clavatum var. equisetoides Schwacke; Arch. Bot. Sao Paulo 1: 438 (1927)
= Lycopodium clavatum var. eriostachys (Fée) Nessel & Hoehne; Arch. Bot. Sao Paulo 1: 435 (1927)
= Lycopodium clavatum var. jamaicense Spring; Flora 21(1): 174 (1838)
= Lycopodium clavatum f. incurvum (Grev. & Hook.) Nessel; Bärlappgewächse 305 (1939)
= Lycopodium clavatum var. desvauxianum Spring; Flora 21(1): 173 (1838)
= Lycopodium clavatum var. minarum Christ; Pl. Nov. Mineiras 2: 41 (1900)
= Lycopodium clavatum var. piliferum (Raddi) Nessel & Hoehne; Arch. Bot. Sao Paulo 1: 435 (1927)
= Lycopodium clavatum var. trichophyllum (Desv.) Nessel & Hoehne; Arch. Bot. Sao Paulo 1: 436 (1927)
= Lycopodium eriostachys Fée; Cr. vasc. Br. 1: 224 (1869)
= Lycopodium piliferum Raddi; Pl. Bras. 79 (1825)
= Lycopodium trichiatum var. desvauxianum (Spring) Spring; Mém. Acad. Roy. Sci. Bruxelles 15: 92 (1842)
= Lycopodium trichophyllum Desv.; Mém. Soc. Linn. Paris 6: 184 (1827)
species :
Lycopodium diaphanum (P.Beauv.) Sw.; Syn. Fil. 179 (1806)
Distribution: Tristan da Cunha (incl. Inaccessible Isl.); Gough Isl.
91 TDC
Synonyms: hidden
= Lepidotis diaphana P.Beauv.; Prodr. aethéogam. 108 (1805)
species :
Lycopodium hygrophilum Alderw.; Bull. Jard. Bot. Buitenz. II, 24: 5 (1917)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
species :
Lycopodium japonicum Thunb.; Syst. Veg. ed. 14. 944: (1784)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Xinjiang, Yunnan, Zhejiang); Tibet; Taiwan; Japan; India (Arunachal Pradesh, Assam State, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, Tripura, Uttarakhand, West Bengal); Nepal; Bhutan; Myanmar; Sri Lanka; Bangladesh; Cambodia; Laos; Vietnam; Sulawesi; Fiji (Viti Levu); New Caledonia; peninsular Malaysia (Perak, Pahang); Borneo (Sabah, Mt. Kinabalu); Moluccas (Seram); Philippines; Thailand; Vietnam; Solomon Isl. (Guadalcanal); New Guinea (Irian Jaya)
36 CHI TIB 38 JAP TAI 40 BAN BHU IND NEP SRL 41 CBD LAO MYA THA VIE 42 BOR MLY MOL NWG PHI SUL 43 SOL 60 FIJ NWC
Remarks: contains tropical Asian records for "L. clavatum"
Synonyms: hidden
= Lycopodium centrochinense Ching; Acta Bot. Yunnan. 4(3): 224-225 (1982) (fide Fraser-Jenkins 2008, 2017)
= Lycopodium clavatum var. nipponicum Nakai; Bot. Mag. 39(463): 197 (1925)
= Lycopodium divaricatum Wall. ex Hook. & Grev.; Bot. Misc. 2: 377 (1831)
= Lycopodium clavatum var. wallichianum Spring; Monogr. Lyc. 1: 90 (1842)
= Lycopodium interjectum Ching & H.S.Kung; Acta Bot. Yunnan. 4(3): 221 (1982)
= Lycopodium japonicum f. yunnanense (Ching) W.M.Chu; Fl. Yunnan. 20: 25 (2006)
= Lycopodium kinabaluense Ching; Acta Bot. Yunnan. 4(3): 223 (1982)
= Lycopodium pseudoclavatum Ching; Acta Bot. Yunnan. 4(3): 222-223 (1982) (fide Fraser-Jenkins 1997)
= Lycopodium pseudoclavatum var. yunnanense Ching; Acta Bot. Yunnan. 4(3): 223 (1982) (fide Fraser-Jenkins 1997)
= Lycopodium simulans Ching & H.S.Kung; Acta Bot. Yunnan. 4(3): 220-221 (1982)
= Lycopodium taliense Ching; Acta Bot. Yunnan. 4: 221 (1982)
= Stachygynandrum japonicum (Thunb.) P.Beauv.; Prodr. aethéogam. 113 (1805)
species :
Lycopodium lagopus (Laest. ex C.Hartm.) Zinserl. ex Kuzen.; Fl. Murmansk Obl. 1: 80 (1953)
Name: One-cone clubmoss
Distribution: Scotland; Norway; Sweden; Finland; Austria; Switzerland; N-Italy; N-European Russia; Siberia (W-Siberia, C-Siberia); Russian Far East; Alaska; USA (Connecticut, Illinois, Massachusetts, Maine, Michigan, Minnesota, Montana, New Hampshire, New York, Ohio, Oregon, Pennsylvania, Vermont, Washington State, Wisconsin); Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); Greenland; St. Pierre et Miquelon
10 FIN GRB NOR SWE 11 AUT SWI 13 ITA 14 RUS 30 SIB 31 RFE 70 ASK CDN GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA
Synonyms: hidden
= Lycopodium clavatum f. furcatum (Luerss.) Vict.; Contr. Lab. Bot. Univ. Montreal 3: 25 (1925)
= Lycopodium clavatum ssp. lagopus (Laest.) Dostál
= Lycopodium clavatum var. monostachyum Schur; (1858)
= Lycopodium clavatum ssp. monostachyon (Hook. & Grev.) Selander; Acta Phytogeogr. Suec. 28: 22 (1950)
= Lycopodium clavatum var. brevispicatum Peck; Rep. (Ann.) Regents Univ. State N. Y. 54: 162 (1901)
= Lycopodium clavatum var. integerrimum Spring; Nouv. Mem. Acad. Roy. Sci. Bruxelles (Mon. Fam. Lycopodiaceae) 15: 90 (1842)
= Lycopodium clavatum var. lagopus Laest. ex C.Hartm.; Handb. Skand. Fl. ed. 7: 313 (1858)
= Lycopodium clavatum var. megastachyon Fernald & Bissell; Rhodora 12(135): 53 (1910)
= Lycopodium clavatum ssp. megastachyon (Fernald & Bissell) Á.Löve & D.Löve; Nucleus (Calcutta) 1(1): 7 (1958)
= Lycopodium clavatum var. monostachyon Hook. & Grev.; Bot. Misc. 2: 375 (1831)
= Lycopodium integrifolium Hook. ex Goldie; Edinburgh New Philos. J. 6: 332 (1822)
= Lycopodium clavatum var. integrifolium (Hook. ex Goldie) Spring; Flora 21(1): 174 (1838)
= Lycopodium pseudoannotinum Schur; Enum. pl. Transsilv. 826 (1866) (nom. inval.)
species :
Lycopodium papuanum Nessel; Repert. Spec. Nov. Regni Veg. 39(8): 64, t. 191a (1935)
Distribution: New Guinea
42 NWG
species :
Lycopodium pullei Alderw.; Bull. Jard. Bot. Buitenz. II, 24: 6 (1917)
Distribution: New Guinea
42 NWG
species :
Lycopodium venustulum Gaudich.; Voy. Uranie [Freycinet] 1: 283, t. 22 (1826)
Distribution: [Western] Samoa (Savaii); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.); Society Isl. (Tahiti); Austral Isl. (Rapa Iti)
60 SAM 61 AUI SCI 63 HAW
Synonyms: hidden
= Lycopodium sirii Herter; Est. Bot. Reg. Uruguaya (Index Lycop.) 20: 41 (Ind. Lycopod.) (1949)
= Lycopodium jovis Hedw.; Index Lyc. 24 (1949)
= Lycopodium heterophyllum Hook. & Grev.; Icon. Fil. 1: t. 113 (1829 publ. 1831) (non Spreng. 1827)
variant :
var. verticale W.H.Wagner; Contr. Univ. Michigan Herb. 22: 144 (1999)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
species :
Lycopodium vestitum Desv. ex Poir.; Encycl. suppl. 3: 546 (1814)
Distribution: Ecuador; Peru; ?Venezuela
83 ECU PER
Synonyms: hidden
= Lycopodium albidum Baker; J. Bot. 25: 374 (1887)
= Lycopodium scariosum Hook.; (1836) (non G. Forst.)
...

Lilium philadelphicum (Liliaceae) (Photo: M. Hassler)