132.0000
Family
Orchidaceae Juss.; Gen. Pl.: 64 (1789), nom. cons.
over 800 genera, more than 30.000 spp. worldwide, mostly tropical
Species listed in database: 30517; hybrids: 1413
= Apostasiaceae Lindl.; Nix. Pl.: 22 (1833), nom. cons.
= Cypripediaceae Lindl.; Nix. Pl.: 22 (1833)
= Limodoraceae Horan.; Char. Ess. Fam.: 44 (1847)
= Liparidaceae Vines; Stud. Text-book Bot. 2: 567 (1895)
= Neottiaceae Horan.; Prim. Lin. Syst. Nat.: 50 (1834)
= Neuwiediaceae R.Dahlgren ex Reveal & Hoogland; Bull. Mus. Natl. Hist. Nat., ser. 4, sect. B Adansonia 13: 91 (1991)
= Ophrydaceae Vines; Stud. Text-book Bot. 2: 566 (1895)
= Pycnanthaceae Ravenna; Onira 12(6): 29 (2011)
= Vanillaceae Lindl.; Key Bot.: 73 (1835)
132.4330
Genus
Nigritella Rich.; Orch. Eur. Annot. (Mem. Mus. Hist. Nat. Paris 4: 48: 1818) 26 (1817)
Distribution: Europe (mountains). Some authors include this genus in Gymnadenia based on molecular results, but this has been recently rejected (HEDREN, KLEIN & TEPPNER 2002, KREUTZ 2005, PILLON et al. 2007). There are wildly fluctuating species concepts, either lumping all in two species (H. Baumann & R. LORENZ 2005) or in up to 15 microspecies (TEPPNER & E. KLEIN).
Species listed in database: 14; hybrids: 10
= Gymnadenia subgen.Nigritella (Rich.) O.Gerbaud & P.Quentin; Naturalistes Belges 80(3): 374 (1999)
species :
Nigritella bicolor
W.Foelsche; J. Eur. Orch. 42(1): 60 (2010)
Distribution: Switzerland; Austria; NW-Slovenia; N-Italy; Romania
11 AUT SWI 13 ITA ROM SLV
Synonyms: hidden
= Gymnadenia bicolor (W.Foelsche) W.Foelsche & O.Gerbaud; Bull. Soc. Franc. Orchidophilie Rhone-Alpes 24: 23 (2011)
= Nigritella bicolor var. rubida W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 32(1): 87 (2015)
species :
Nigritella carniolica
W.Foelsche, G.Foelsche, R.Wüest, E.Merz, C.Kreutz & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 37(2): 149 (2020)
Distribution: Slovenia
13 SLV
Remarks: provisionally accepted name, not confirmed by genetics
species :
Nigritella corneliana
Beauverd; Bull. Soc. Bot. Geneve Ser. II 17: 336 (1925 publ. 1926)
Distribution: SE-France; NW-Italy (W-Alps, Alpes Maritimes, SW-Alps)
12 FRA 13 ITA
Remarks: nigra agg.
Synonyms: hidden
= Gymnadenia corneliana (Beauverd) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Gymnadenia corneliana f. vesubiana (G.Keller) P.Delforge; Naturalistes Belges 87(Orchid. 19): 259 (2006)
= Gymnadenia corneliana var. vesubiana (G.Keller) G.Foelsche & W.Foelsche; J. Eur. Orchid. 31(4): 803 (1999)
= Gymnadenia nigra f. vesubiana (G.Keller) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 276 (2011 publ. 2012)
= Gymnadenia nigra subsp. corneliana (Beauverd) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Nigritella angustifolia subsp. corneliana (Beauverd) Douin; Fl. Ill. France 11: 31 (1931)
= Nigritella corneliana (Beauverd) Gölz & H.R.Reinhard; Jahresber. Naturwiss. Ver. Wuppertal 39: 39 (1986)
= Nigritella corneliana var. vesubiana (G.Keller) W.Foelsche; Joannea Bot. 7: 48 (2009)
= Nigritella lithopolitanica subsp. corneliana (Beauverd) Teppner & E.Klein; Phyton (Horn) 25: 158 (1985)
= Nigritella nigra subsp. corneliana Beauverd; Bull. Soc. Bot. Genève, sér. 2, 17: 337 (1925)
= Nigritella rubra subsp. corneliana (Beauverd) P.Fourn.; Quatre Fl. France 202 (1935)
= Nigritella rubra sublusus vesubiana G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A 2-5: 268 (1935)
= Nigritella rubra var. corneliana (Beauverd) Soó; (1933)
variant :
var. bourneriasii
(E.Breiner & R.Breiner) E.Klein; Phyton (Horn) 36(1): 59 (1996)
Distribution: SE-France
12 FRA
Synonyms: hidden
= Gymnadenia corneliana f. bourneriasii (E.Breiner & R.Breiner) P.Delforge; Naturalistes Belges 87(Orchids. 19): 259 (2006)
= Gymnadenia corneliana var. bourneriasii (E.Breiner & R.Breiner) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Gymnadenia corneliana var. bourneriasii (E.Breiner & R.Breiner) Pellic.; Bull. Mycol. Bot. Dauphiné-Savoie 176: 49 (2005), isonym
= Gymnadenia nigra f. bourneriasii (E.Breiner & R.Breiner) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 276 (2011 publ. 2012)
= Nigritella bourneriasii (E.Breiner & R.Breiner) Landolt; Fl. Indicativa 269 (2010)
= Nigritella corneliana subsp. bourneriasii E.Breiner & R.Breiner; Mitt. Arbeitskr. Heim Orch. Baden-Württ. 25(4): 480 (1993)
species :
Nigritella graciliflora
W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 35(1): 166 (2018)
Distribution: SE-Austria
11 AUT
Remarks: provisionally accepted name, not confirmed by genetics
species :
Nigritella karawankarum
W.Foelsche, R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 35(1): 91 (2018)
Distribution: SE-Austria
11 AUT
Remarks: provisionally accepted name, not confirmed by genetics
species :
Nigritella kossutensis
W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic; J. Eur. Orch. 49(3-4): 505 (2017)
Distribution: NW-Slovenia
13 SLV
Remarks: diploid taxon
Synonyms: hidden
= Gymnadenia kossutensis (W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic) J.M.H.Shaw; Orchid Rev. Suppl., 127(1326): 45 (2019)
species :
Nigritella mauriennsis
W.Foelsche, G.Foelsche, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 39(1-2): 172 (2022)
Distribution: SE-France (Savoie)
12 FRA
Remarks: provisionally accepted name, not confirmed by genetics
species :
Nigritella nigra
(L.) Rchb.f.; Icon. Fl. Germ. 13-14: 102 (1851), s. str.
subspecies :
subsp. nigra
Name: Schwarzes Kohlröschen
Distribution: Norway; Sweden; ?Lithuania
10 NOR SWE
Remarks: triploid apomict (2n - 60)
Synonyms: hidden
= Gymnadenia nigra (L.) Rchb.f.; Bonplandia (Hannover) 4: 321 (1856)
= Gymnadenia nigra (L.) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998), isonym
= Gymnadenia nigra (L.) Wettst.; Verh. zool.-bot. Ges. Wien (1889) 83, et Ber. Deutsch. Bot. Ges. 7: 308 (1889), isonym
= Gymnadenia nigra var. longibracteata (Beck) Wettst.; Biblioth. Bot. 26: 92 (1892)
= Habenaria nigra (L.) R.Br.; Hort. Kew. ed. II 5: 192 (1813)
= Nigritella angustifolia f. longibracteata Beck; Ann. K. K. Naturhist. Hofmus. 5: 577 (1890)
= Nigritella angustifolia Rich.; De Orchid. Eur.: 34 (1817)
= Nigritella brachystachya A.Kern.; Verh. Zool.-Bot. Ges. Wien 15: 224 (1865)
= Nigritella fragrans var. megastachya (A.Kern.) Rouy; Fl. France 13: 97 (1912)
= Nigritella hybrida Schur; Enum. Pl. Transs. 647 (1866)
= Nigritella megastachya A.Kern.; Verh. Zool.-Bot. Ges. Wien 15: 222 (1865)
= Nigritella nigra f. longibracteata (Beck) Soó; Arch. Bot. (Leipzig) 23: 93 (1928)
= Nigritella suaveolens W.D.J.Koch; Syn. Fl. Germ. Helv.: 690 (1837)
= Orchis nigra (L.) Scop.; Fl. Carniol., ed. 2, 2: 200 (1771)
= Orchis variegata Schrank; Baier. Fl. 1: 241 (1789), nom. illeg.
= Satyrium nigrum L.; Sp. Pl. 944 (1753)
= Sieberia nigra (L.) Spreng.; Anleit. 2: 282 (1817)
= xGymnigritella ×brachystachya (A.Kern.) E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 358 (1908)
= xGymnigritella ×megastachya (A.Kern.) E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 359 (1908)
subspecies :
subsp. austriaca
Teppner & E.Klein; Phyton (Austria) 31(1): 17 (1990)
Name: Österreichisches Kohlröschen (DE)
Distribution: Germany (Bayern); Switzerland; Austria; SE-France; N-Italy; NW-Slovenia
11 AUT GER SWI 12 FRA 13 ITA SLV
Remarks: tetraploid apomict (2n - 80)
Synonyms: hidden
= Gymnadenia austriaca (Teppner & E.Klein) P.Delforge; Les Naturalistes Belges 79: 254 (1998)
= Gymnadenia nigra subsp. austriaca (Teppner & E.Klein) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella austriaca (Teppner & E.Klein) P.Delforge; Naturalistes Belges 72(3): 100 (1991)
subspecies :
subsp. bucegiana
Hedrén, Anghelescu & R.Lorenz; J. Eur. Orch. 54(3-4): 293 (2022)
Distribution: Romania (Bucegi Mts.)
13 ROM
Remarks: tetraploid apomict (2n - 80)
subspecies :
subsp. carpatica
(Zapal.) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 717 (2005)
Distribution: Romania; Ukraine (E-Carpathians)
13 ROM 14 UKR
Synonyms: hidden
= Gymnadenia carpatica (Zapal.) Teppner & E.Klein; Phyton (Austria) 38(1): 221 (1998)
= Nigritella angustifolia var. carpatica Zapal.; Consp. Fl. Galic. Crit. 1: 215 (1906)
= Nigritella carpatica (Zapal.) Teppner, E.Klein & Zagulskij; Phyton (Austria) 34(2): 171 (1994)
= Nigritella nigra var. carpatica (Zapal.) Pawl.; Bull. Int. Acad. Polon. Sci., Cl. Sci. Math., Sér. B 1, Bot. 1947: 85, 96 (1947)
= Nigritella rubra f. carpatica (Zapal.) Soó; Repert. Sp. Nov. Regni Veg. 24: 33 (1927)
subspecies :
subsp. gallica
E.Breiner & R.Breiner; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 25(4): 471 (1993)
Distribution: Switzerland; France (Jura, Massif Central, Alps); NW-Italy
11 SWI 12 FRA 13 ITA
Remarks: tetraploid apomict (2n - 80)
Synonyms: hidden
= Gymnadenia austriaca var. gallica (E.Breiner & R.Breiner) P.Delforge; Naturalistes Belges 79(4): 255 (1998)
= Gymnadenia nigra subsp. gallica (E.Breiner & R.Breiner) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella gallica (E.Breiner & R.Breiner) Landolt; Fl. Indicativa 269 (2010)
= Nigritella nigra var. gallica (E.Breiner & R.Breiner) Wenker & S.Wenker; Jahresber. Naturwiss. Vereins Wuppertal 60: 200 (2007)
= Nigritella rhellicani subsp. gallica (E.Breiner & R.Breiner) Kerguélen; Bull. Assoc. d´Informatique Appl. Bot. (Marseille) 1: 183 (1994)
= Nigritella rhellicani subsp. iberica (Teppner & E.Klein) Kerguélen; Bull. Assoc. d´Informatique Appl. Bot. (Marseille) 1: 183 (1994)
subspecies :
subsp. iberica
Teppner & E.Klein; Phyton (Austria) 33(2): 192 (1993)
Distribution: Spain; Andorra; France (Pyrenees)
12 ADO FRA SPA
Remarks: tetraploid apomict (2n - 80)
Synonyms: hidden
= Gymnadenia austriaca subsp. iberica (Teppner & E.Klein) L.Sáez & Aymerich; Orsis 29: 40 (2015)
= Gymnadenia austriaca var. iberica (Teppner & E.Klein) G.Foelsche & W.Foelsche; J. Eur. Orchid. 31(4): 803 (1999)
= Gymnadenia nigra subsp. iberica (Teppner & E.Klein) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella austriaca subsp. iberica (Teppner & E.Klein) L.Sáez; Anales Jard. Bot. Madrid 61(1): 87 (2004)
= Nigritella nigra var. iberica (Teppner & E.Klein) Wenker & S.Wenker; Jahresber. Naturwiss. Vereins Wuppertal 60: 199 (2007)
subspecies :
subsp. lithopolitanica
(Ravnik) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 717 (2005)
Name: Steineralpen-Kohlröschen
Distribution: Austria; NW-Slovenia (SE-Alps)
11 AUT 13 SLV
Conservation Status: IUCN R
Synonyms: hidden
= Gymnadenia lithopolitanica (Ravnik) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella lithopolitanica Ravnik; Acta Bot. Croat. 37: 226 (1978)
= Nigritella lithopolitanica var. luciae W.Foelsche, G.Foelsche, R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 158 (2020)
subspecies :
subsp. runei
(Teppner & E.Klein) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 717 (2005)
Distribution: N-Sweden (Lappland)
10 SWE
Conservation Status: IUCN R
Synonyms: hidden
= Gymnadenia runei (Teppner & E.Klein) Ericsson; Svensk Bot. Tidskr. 91: 142 (1997)
= Nigritella runei (Teppner & E.Klein) Kreutz; Kompend. Eur. Orchid. 79 (2004)
= xGymnigritella ×runei Teppner & E.Klein; Phyton (Austria) 29(2): 163 (1989)
species :
Nigritella ravnikii
W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic; J. Eur. Orch. 49(3-4): 508 (2017)
Distribution: NW-Slovenia
13 SLV
Remarks: diploid taxon
Synonyms: hidden
= Gymnadenia ravnikii (W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic) J.M.H.Shaw; Orchid Rev. Suppl., 127(1326): 45 (2019)
species :
Nigritella rhellicani
Teppner & E.Klein; Phyton (Austria) 31(1): 7 (1990)
subspecies :
subsp. rhellicani
Name: Rhellicanus-Kohlröschen, Alpen-Kohlröschen (DE)
Distribution: Germany (Bayern); Switzerland; Liechtenstein; Austria; Slovakia; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Albania; Romania; Bulgaria; Greece
11 AUT GER LIE SLO SWI 12 FRA 13 ALB BIH BUL CRO GRC ITA ROM SLV
Synonyms: hidden
= Gymnadenia nigra subsp. rhellicani (Teppner & E.Klein) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Gymnadenia rhellicani (Teppner & E.Klein) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Gymnadenia rhellicani f. fulva (G.Keller) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella nigra lusus fulva G.Keller; Keller, Monogr. Iconogr. Orchid. Eur. 2: 267 (1933)
= Nigritella nigra subsp. rhellicani (Teppner & E.Klein) H.Baumann, Künkele & R.Lorenz; J. Eur. Orch. 36(3): 776 (2004)
= Nigritella nigra var. rosea Vis. & Sacc.; Atti Reale Ist. Veneto Sci. Lett. Arti 14: 321 (1868)
= Nigritella nigra var. rosea Vis. & Sacc. ex Goiran; Nuov. Giorn. Bot. Ital. 15: 38 (1883) [fide Baumann, Künkele & R. Lorenz 2002]
= Nigritella rhellicani f. fulva (G.Keller) Teppner & E.Klein; Phyton (Austria) 31(1): 10 (1990)
= Nigritella rhellicani var. rosea (Vis. & Sacc. ex Goiran) Teppner & E.Klein; Phyton (Austria) 31(1): 10 (1990)
subspecies :
subsp. cenisia
(G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) Kreutz; Eurorchis 17: 107 (2005)
Name: Mont-Cenis-Kohlröschen, Nigritelle du Mont Cenis
Distribution: SE-France (SW-Alps); NW-Italy
12 FRA 13 ITA
Synonyms: hidden
= Gymnadenia cenisia (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud; Naturalistes Belges 80(3): 374 (1999)
= Gymnadenia nigra subsp. cenisia (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Gymnadenia rhellicani var. robusta P.Delforge; Naturalistes Belges 84 (Orchid. 16): 127 (2003)
= Nigritella cenisia G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud; L´Orchidophile 134: 248 (1998)
= Nigritella rhellicani var. cenisia (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) Wenker & S.Wenker; Jahresber. Naturwiss. Vereins Wuppertal 60: 196 (2007)
= Nigritella rhellicani var. robusta (P.Delforge) Kreutz; Kompend. Eur. Orchid. 78 (2004)
subspecies :
subsp. gabasiana
(Teppner & E.Klein) Kreutz; Eurorchis 17: 107 (2005)
Distribution: N-Spain (Pyrenees, Cordillera Cantabrica); Andorra; SW-France (Pyrenees)
12 ADO FRA SPA
Synonyms: hidden
= Gymnadenia gabasiana (Teppner & E.Klein) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Gymnadenia nigra subsp. gabasiana (Teppner & E.Klein) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Nigritella gabasiana Teppner & E.Klein; Phyton (Austria) 33(2): 182 (1993)
= Nigritella nigra subsp. gabasiana (Teppner & E.Klein) H.Baumann & R.Lorenz; J. Eur. Orch. 37: 717 (2005)
= Nigritella nigra var. pyrenaica Schltr.; Repert. Spec. Nov. Regni Veg. 16: 271 (1919)
species :
Nigritella rubra
(Wettst.) K.Richt.; Pl. Eur. 1: 278 (1890)
subspecies :
subsp. rubra
Name: Rotes Kohlröschen (DE)
Distribution: S-Germany (Bavarian Alps); Switzerland; Liechtenstein; Austria; Italy; NW-Slovenia
11 AUT GER LIE SWI 13 ITA SLV
Remarks: nigra agg.
Synonyms: hidden
= Gymnadenia miniata (Crantz) Hayek; Fl. Steiermark 2(2): 133 (1956)
= Gymnadenia miniata f. rosea (C.Boillat & V.Boillat) P.Delforge; Naturalistes Belges 96(Orchid. 28): 16 (2015)
= Gymnadenia nigra subsp. rubra (Wettst.) H.Sund.; Europ. Medit. Orchid., ed. 3: 41 (1980)
= Gymnadenia nigra var. rubra (Wettst.) Kraenzl.; Orchid. Gen. Sp. 1(2): 559 (1898)
= Gymnadenia rhellicani var. rosea (Vis. & Sacc. ex Goiran) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Gymnadenia rubra f. rosea (C.Boillat & V.Boillat) P.Delforge; Naturalistes Belges 92(Orchid. 24): 108 (2011)
= Gymnadenia rubra Wettst.; Verh. zool.-bot. Ges. Wien (1889) 83, et Ber. Deutsch. Bot. Ges. 7: 312 (1889)
= Gymnadenia rubra Wettst. ex Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella miniata (Crantz) Janch.; Phyton (Horn) 8: 232 (1959)
= Nigritella nigra subsp. rubra (Wettst.) Beauverd; Bull. Soc. Bot. Genéve ser. 2, 17: 337 (1926)
= Nigritella nigra var. rubra (Wettst.) W.D.J.Koch; Syn. Deut. Schweiz. Fl. ed. 3: 2434 (1905)
= Nigritella rubra f. carpatica Soó; Repert. Spec. Nov. Regni Veg. 24: 33 (1927)
= Nigritella rubra f. rosea C.Boillat & V.Boillat; J. Eur. Orch. 42(3-4): 491 (2010)
= Nigritella rubra f. wettsteinii Soó; Repert. Spec. Nov. Regni Veg. 24: 33 (1927)
= Orchis miniata Crantz; Stirp. Austr. Fasc., ed. 2, 2: 486 (1769), nom. rej. prop.
variant :
var. minor
(W.Foelsche & Zernig) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008)
Distribution: Austria (Oststeiermark, Mt. Trenchtling)
11 AUT
Synonyms: hidden
= Gymnadenia miniata f. minor (W.Foelsche & Zernig) P.Delforge; Naturalistes Belges 96(Orchid. 28): 16 (2015)
= Gymnadenia minor (W.Foelsche & Zernig) W.Foelsche, Zernig & O.Gerbaud; Bull. Soc. Franc. Orchidophilie Rhône-Alpes 24: 23 (2011)
= Gymnadenia rubra f. minor (W.Foelsche & Zernig) P.Delforge; Naturalistes Belges 92(Orchid. 24): 107 (2011)
= Nigritella minor W.Foelsche & Zernig; Joannea Bot. 6: 10 (2007)
subspecies :
subsp. buschmanniae
(Teppner & Ster) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 717 (2005)
Name: Buschmanns Kohlröschen
Distribution: N-Italy (Brenta)
13 ITA
Synonyms: hidden
= Gymnadenia buschmanniae (Teppner & Ster) Teppner & E.Klein; Phyton (Austria) 38(1): 222 (1998)
= Nigritella buschmanniae Teppner & Ster; Phyton (Austria) 36(2): 278 (1996)
subspecies :
subsp. dolomitensis
(Teppner & E.Klein) D.Wenker & S.Wenker; Jahresber. Naturwiss. Ver. Wuppertal 60: 177-250 (2007)
Name: Dolomiten-Kohlröschen (DE)
Distribution: N-Italy (Dolomiti)
13 ITA
Remarks: probably not in S-Germany
Synonyms: hidden
= Gymnadenia dolomitensis Teppner & E.Klein; Phyton (Horn) 38: 223 (1998)
= Gymnadenia miniata var. dolomitensis (Teppner & E.Klein) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 16 (2015)
= Gymnadenia rubra var. dolomitensis (Teppner & E.Klein) P.Delforge; Naturalistes Belges 92(Orchid. 24): 108 (2011)
= Nigritella dolomitensis (Teppner & E.Klein) Hedrén, E.Klein & Teppner; Phyton (Horn) 40: 243 (2000)
= Nigritella miniata var. dolomitensis (Teppner & E.Klein) H.Baumann, R.Lorenz & Perazza; Gredleriana 12(2012): 93 (2013)
= Nigritella rubra subsp. dolomitensis (Teppner & E.Klein) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 164 (2007), isonym
= Nigritella rubra var. dolomitensis (Teppner & E.Klein) R.Lorenz & Perazza; GIROS Notiz. 27: 8 (2004)
subspecies :
subsp. archiducis-johannis
(Teppner & E.Klein) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 717 (2005)
Name: Erzherzog-Johann-Kohlröschen
Distribution: Austria (Totes Gebirge); NW-Slovenia (Mt. Triglav)
11 AUT 13 SLV
Conservation Status: extremely rare, a few 100 specimens known
Synonyms: hidden
= Gymnadenia archiducis-joannis (Teppner & E.Klein) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella archiducis-joannis Teppner & E.Klein; Phyton (Austria) 25(1): 168 (1985)
= Nigritella rubra subsp. archiducis-joannis (Teppner & E.Klein) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008)
subspecies :
subsp. hygrophila
(W.Foelsche & Heidtke) comb.ined.
Distribution: Austria (Steiermark); ?SE-France (?Mt. Cenis); Italy (Südtirol: Passo Pordoi); Slovenia (Triglav)
11 AUT 13 ITA SLV
Remarks: provisionally accepted name, not confirmed by genetics
Synonyms: hidden
= Gymnadenia hygrophila (W.Foelsche & Heidtke) W.Foelsche, Heidtke & O.Gerbaud; Bull. Soc. Franc. Orchidophilie Rhone-Alpes 24: 23 (2011)
= Gymnadenia miniata f. hygrophila (W.Foelsche & Heidtke) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 16 (2015)
= Gymnadenia rubra f. hygrophila (W.Foelsche & Heidtke) P.Delforge; Naturalistes Belges 92(Orchid. 24): 107 (2011)
= Nigritella hygrophila var. pauciflora W.Foelsche, G.Foelsche, R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 164 (2020)
= Nigritella hygrophila W.Foelsche & Heidtke; J. Eur. Orch. 43(1): 143 (2011)
subspecies :
subsp. stiriaca
(Rech.) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 718 (2005)
Name: Steirisches Kohlröschen
Distribution: Austria (E-Alps)
11 AUT
Synonyms: hidden
= Gymnadenia rubra var. stiriaca Rech.; Mitt. Naturwiss. Vereins Steiermark 42: 148 (1906)
= Gymnadenia stiriaca (Rech.) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella rubra subsp. stiriaca (Rech.) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008), isonym
= Nigritella stiriaca (Rech.) Teppner & E.Klein; Phyton (Austria) 25(1): 159 (1985)
species :
Nigritella savogiensis
W.Foelsche, G.Foelsche, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 39(1-2): 166 (2022)
Distribution: SE-France (Savoie)
12 FRA
Remarks: provisionally accepted name, not confirmed by genetics
species :
Nigritella widderi
Teppner & E.Klein; Phyton (Austria) 25(2): 318 (1985)
Name: Widders Kohlröschen
Distribution: SE-Germany (Bavarian Alps); N-Austria (Steiermark); Italy (Abruzzes, Dolomiti); Bosnia & Hercegovina
11 AUT GER 13 BIH ITA
Synonyms: hidden
= Gymnadenia widderi (Teppner & E.Klein) Teppner & E.Klein; Phyton (Austria) 38(1): 220-224 (1998)
= Nigritella rubra subsp. widderi (Teppner & E.Klein) H.Baumann & R.Lorenz; J. Eur. Orch. 37: 718 (2005)
= Nigritella rubra subsp. widderi (Teppner & E.Klein) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008), isonym
species :
Nigritella suceveana
Kreutz, Bobocea & O.Gerbaud; Bull. S. F. O. Rhône-Alpes 44: 35 (2021)
Distribution: Romania
13 ROM
Remarks: provisionally accepted name, not confirmed by genetics
species :
Nigritella ×delphineae
M.Gerbaud & O.Gerbaud; L´Orchidophile 120: 33, 30 (1996)
Distribution: SE-France; N-Italy
12 FRA 13 ITA
Remarks: Nigritella corneliana x rhellicani
Synonyms: hidden
= Gymnadenia ×breinerorum O.Gerbaud & W.Foelsche; Naturalistes Belges 80(3): 381 (1999)
= Gymnadenia ×delphineae (M.Gerbaud & O.Gerbaud) M.Gerbaud & O.Gerbaud; Naturalistes Belges 80(3): 378 (1999)
= Gymnadenia ×nigra nothosubsp. delphineae (M.Gerbaud & O.Gerbaud) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 269 (2012)
= Nigritella ×breinerorum (O.Gerbaud & W.Foelsche) W.Foelsche; Joannea Bot. 7: 47 (2009)
species :
Nigritella ×eggeriana
(O.Gerbaud) W.Foelsche; Joannea Bot. 7: 47 (2009)
Distribution: France
12 FRA
Remarks: Nigritella austriaca (var. gallica) x rhellicani
Synonyms: hidden
= Gymnadenia nigra nothosubsp. eggeriana (O.Gerbaud) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 269 (2011 publ. 2012)
= Gymnadenia ×eggeriana O.Gerbaud; Naturalistes Belges 80(3): 378 (1999)
species :
Nigritella ×foelscheana
R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 135 (2020)
Distribution: Slovenia
13 SLV
Remarks: Nigritella kossutensis x N. ravnikii
species :
Nigritella ×fohringeri
Griebl; Ber. Arbeitskreis. Heimische Orchid. 28(2): 169 (166-170; fig.) (2011)
Distribution: Austria
11 AUT
Remarks: Nigritella lithopolitanica x miniata
Synonyms: hidden
= Gymnadenia ×fohringeri (Griebl) J.M.H.Shaw; Orchid Rev. 122(1306): 39 (2014)
species :
Nigritella ×hennigsiana
R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 130 (2020)
Distribution: Slovenia
13 SLV
Remarks: Nigritella lithopolitanica x karawankarum
species :
Nigritella ×petzenensis
F.Fohringer & Redl; Linzer Biol. Beitr. 36(1): 573 (2004)
Distribution: Austria; Slovenia
11 AUT 13 SLV
Remarks: Nigritella rhellicani x lithopolitanica
Synonyms: hidden
= Gymnadenia ×petzenensis (F.Fohringer & Redl) Oddone; GIROS Orch. Spont. Eur. 59(2): 357 (2016)
species :
Nigritella ×robatschiana
(O.Gerbaud & W.Foelsche) W.Foelsche; Joannea Bot. 7: 47 (2009)
Distribution: France; ?Italy
12 FRA
Remarks: Nigritella cenisia x rhellicani
Synonyms: hidden
= Gymnadenia ×robatschiana O.Gerbaud & W.Foelsche; Naturalistes Belges 80(3): 380 (1999)
species :
Nigritella ×teppneri
W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 32(1): 88 (2015)
Distribution: N-Italy
13 ITA
Remarks: Nigritella hygrophila x rhellicani
Synonyms: hidden
= Gymnadenia ×teppneri (W.Foelsche) Oddone; GIROS Orch. Spont. Eur. 59(2): 357 (2016)
species :
Nigritella ×wettsteiniana
(O.Abel) Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 811 (1907)
Distribution: Switzerland; Austria; Italy; Romania
11 AUT SWI 13 ITA ROM
Remarks: Nigritella nigra s. l. x Nigritella rubra
Synonyms: hidden
= Gymnadenia ×bornmuelleri Dalla Torre & Sarnth.; Fl. Tirol 6: I. 531 (1906)
= Gymnadenia ×wettsteiniana O.Abel; Verh. K. K. Zool.-Bot. Ges. Wien 47: 611 (1897)
= Nigritella ×wettsteiniana (O.Abel) Schltr.; Repert. Spec. Nov. Regni Veg. 16: 271 (1919), isonym
species :
Nigritella ×wulfeniana
M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 125 (2020)
Distribution: Slovenia
13 SLV
Remarks: Nigritella karawankarum x N. ravnikii
