134.0000
Family
Orchidaceae Juss.; Gen. Pl. [Jussieu] 64 (1789), nom. cons.
over 800 genera, more than 30.000 spp. worldwide, mostly tropical
Species listed in database: 30557; hybrids: 1460
= Apostasiaceae Lindl.; Nix. Pl. 22 (1833), nom. cons.
= Cypripediaceae Lindl.; Nix. Pl. 22 (1833)
= Limodoraceae Horan.; Char. Ess. Fam. 44 (1847)
= Liparidaceae Vines; Stud. Text-book Bot. 2: 567 (1895)
= Neottiaceae Horan.; Prim. Lin. Syst. Nat. 50 (1834)
= Neuwiediaceae R.Dahlgren ex Reveal & Hoogland; Bull. Mus. Natl. Hist. Nat., B, Adansonia, Sér. 4, 13: 91 (1991)
= Ophrydaceae Vines; Stud. Text-book Bot. 2: 566 (1895)
= Pycnanthaceae Ravenna; Onira 12(6): 29 (2011)
= Vanillaceae Lindl.; Key Bot. 73 (1835)
134.4330
Genus
Nigritella Rich.; Mém. Mus. Hist. Nat. 4: 48 (1818)
Distribution: Europe (mountains). Some authors include this genus in Gymnadenia based on molecular results, but this has been rejected (Hedren, Klein & Teppner 2002, Pillon et al. 2007). There are wildly fluctuating species concepts, either lumping all in two species (H. Baumann & R. Lorenz 2005) or in up to 40 apomictic microspecies. We use mostly the latest molecular analysis of Hedrén et al. (2017) and the rank applied there, except for the most recently described taxa for which there is no genetic analysis yet.
Species listed in database: 16; hybrids: 10
= Gymnadenia subgen.Nigritella (Rich.) O.Gerbaud & P.Quentin; Naturalistes Belges 80(3): 374 (1999)
species :
Nigritella bicolor
W.Foelsche; J. Eur. Orch. 42(1): 60 (2010)
Distribution: Switzerland; Austria; NW-Slovenia; N-Italy; Romania
11 AUT SWI 13 ITA ROM SLV
Synonyms: hidden
= Gymnadenia bicolor (W.Foelsche) W.Foelsche & O.Gerbaud; Bull. Soc. Franç. Orchidophilie Rhone-Alpes 24: 23 (2011)
= Nigritella bicolor var. rubida W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 32(1): 87 (2015)
species :
Nigritella carniolica
W.Foelsche, G.Foelsche, R.Wüest, E.Merz, C.Kreutz & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 37(2): 149 (2020)
Distribution: SE-Austria; Slovenia; Romania
11 AUT 13 ROM SLV
Remarks: provisionally accepted name, not yet confirmed by genetics
Synonyms: hidden
= Gymnadenia winkeliana Anghelescu, L.Balogh, Mih.Balogh & Kigyossy; Plants (Basel) 13(10)-1363: 38 (2024)
species :
Nigritella corneliana
Beauverd; Bull. Soc. Bot. Genève, Sér. 2, 17: 336 (1925 publ. 1926)
Distribution: SE-France; NW-Italy (W-Alps, Alpes Maritimes, SW-Alps)
12 FRA 13 ITA
Remarks: nigra agg.
Synonyms: hidden
= Gymnadenia corneliana (Beauverd) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Gymnadenia corneliana f. vesubiana (G.Keller) P.Delforge; Naturalistes Belges 87(Orchid. 19): 259 (2006)
= Gymnadenia corneliana var. vesubiana (G.Keller) G.Foelsche & W.Foelsche; J. Eur. Orch. 31(4): 803 (1999)
= Gymnadenia nigra f. vesubiana (G.Keller) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 276 (2011 publ. 2012)
= Gymnadenia nigra subsp. corneliana (Beauverd) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Nigritella angustifolia subsp. corneliana (Beauverd) Douin; Fl. Ill. France 11: 31 (1931)
= Nigritella corneliana (Beauverd) Gölz & H.R.Reinhard; Jahresber. Naturwiss. Vereins Wuppertal 39: 39 (1986)
= Nigritella corneliana var. vesubiana (G.Keller) W.Foelsche; Joannea Bot. 7: 48 (2009)
= Nigritella lithopolitanica subsp. corneliana (Beauverd) Teppner & E.Klein; Phyton (Horn) 25: 158 (1985)
= Nigritella nigra subsp. corneliana Beauverd; Bull. Soc. Bot. Genève, Sér. 2, 17: 337 (1925)
= Nigritella rubra subsp. corneliana (Beauverd) P.Fourn.; Quatre Fl. France 202 (1935)
= Nigritella rubra sublusus vesubiana G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A: 2-5: 268 (1935)
= Nigritella rubra var. corneliana (Beauverd) Soó; (1933)
variant :
var. bourneriasii
(E.Breiner & R.Breiner) E.Klein; Phyton (Horn) 36(1): 59 (1996)
Distribution: SE-France; NW-Italy
12 FRA 13 ITA
Synonyms: hidden
= Gymnadenia corneliana f. bourneriasii (E.Breiner & R.Breiner) P.Delforge; Naturalistes Belges 87(Orchids. 19): 259 (2006)
= Gymnadenia corneliana var. bourneriasii (E.Breiner & R.Breiner) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Gymnadenia corneliana var. bourneriasii (E.Breiner & R.Breiner) Pellic.; Bull. Mycol. Bot. Dauphiné-Savoie 176: 49 (2005), isonym
= Gymnadenia nigra f. bourneriasii (E.Breiner & R.Breiner) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 276 (2011 publ. 2012)
= Nigritella bourneriasii (E.Breiner & R.Breiner) Landolt; Fl. Indicativa 269 (2010)
= Nigritella corneliana subsp. bourneriasii E.Breiner & R.Breiner; Mitt. Arbeitskreises Heimische Orchideen 25(4): 480 (1993)
species :
Nigritella dulnigiana
W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 40(2): 118 (2024)
Distribution: Austria
11 AUT
Remarks: provisionally accepted name, not yet confirmed by genetics
Synonyms: hidden
= Gymnadenia dulnigiana (W.Foelsche) O.Gerbaud & W.Foelsche; Orchidophile (Paris) 55(4): 366 (2024)
species :
Nigritella graciliflora
W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 35(1): 166 (2018)
Distribution: SE-Austria (Steiermark)
11 AUT
Remarks: provisionally accepted name, not yet confirmed by genetics
species :
Nigritella karawankarum
W.Foelsche, R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 35(1): 91 (2018)
Distribution: SE-Austria (Karawanken, Koralpe)
11 AUT
Remarks: provisionally accepted name, not yet confirmed by genetics
species :
Nigritella kossutensis
W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic; J. Eur. Orch. 49(3-4): 505 (2017)
Distribution: NW-Slovenia (Karawanken)
13 SLV
Remarks: diploid taxon
Synonyms: hidden
= Gymnadenia kossutensis (W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic) J.M.H.Shaw; Orchid Rev., Suppl., 127 (1326): 45 (2019)
species :
Nigritella mauriennsis
W.Foelsche, G.Foelsche, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 39(1-2): 172 (2022)
Distribution: SE-France (Savoie)
12 FRA
Remarks: provisionally accepted name, not yet confirmed by genetics
species :
Nigritella nigra
(L.) Rchb.f.; Icon. Fl. Germ. Helv. [H.G.L.Reichenbach] 13-14: 102 (1851), s. str.
subspecies :
subsp. nigra
Name: Schwarzes Kohlröschen
Distribution: Norway; C-Sweden
10 NOR SWE
Remarks: triploid apomict (2n - 60)
Synonyms: hidden
= Gymnadenia nigra (L.) Rchb.f.; Bonplandia (Hannover) 4: 321 (1856)
= Gymnadenia nigra (L.) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998), isonym
= Gymnadenia nigra (L.) Wettst.; Verh. K.K. Zool.-Bot. Ges. Wien (1889) 83, et Ber. Deutsch. Bot. Ges. 7: 308 (1889), isonym
= Gymnadenia nigra var. longibracteata (Beck) Wettst.; Biblioth. Bot. 26: 92 (1892)
= Habenaria nigra (L.) R.Br.; Hort. Kew., ed. 2 [W.T.Aiton] 5: 192 (1813)
= Nigritella angustifolia f. longibracteata Beck; Ann. K.K. Naturhist. Hofmus. 5: 577 (1890)
= Nigritella angustifolia Rich.; De Orchid. Eur. 34 (1817)
= Nigritella brachystachya A.Kern.; Verh. K.K. Zool.-Bot. Ges. Wien 15: 224 (1865)
= Nigritella fragrans var. megastachya (A.Kern.) Rouy; Fl. France [Rouy & Foucaud] 13: 97 (1912)
= Nigritella hybrida Schur; Enum. Pl. Transsilv. 647 (1866)
= Nigritella megastachya A.Kern.; Verh. K.K. Zool.-Bot. Ges. Wien 15: 222 (1865)
= Nigritella nigra f. longibracteata (Beck) Soó; Arch. Bot. (Leipzig) 23: 93 (1928)
= Nigritella suaveolens W.D.J.Koch; Syn. Fl. Germ. Helv. 690 (1837)
= Orchis nigra (L.) Scop.; Fl. Carniol., ed. 2, 2: 200 (1771)
= Orchis variegata Schrank; Baier. Fl. 1: 241 (1789), nom. illeg.
= Satyrium nigrum L.; Sp. Pl. [Linnaeus] 2: 944 (1753)
= Sieberia nigra (L.) Spreng.; Anleit. Kenntn. Gew., ed. 2, 2: 282 (1817)
= xGymnigritella ×brachystachya (A.Kern.) E.G.Camus; Monogr. Orchid. 358 (1908)
= xGymnigritella ×megastachya (A.Kern.) E.G.Camus; Monogr. Orchid. 359 (1908)
subspecies :
subsp. austriaca
Teppner & E.Klein; Phyton (Horn) 31(1): 17 (1990)
Name: Österreichisches Kohlröschen (DE)
Distribution: Germany (Bayern); Austria; N-Italy; NW-Slovenia
11 AUT GER 13 ITA SLV
Remarks: tetraploid apomict (2n - 80)
Synonyms: hidden
= Gymnadenia austriaca (Teppner & E.Klein) P.Delforge; Naturalistes Belges 79: 254 (1998)
= Gymnadenia nigra subsp. austriaca (Teppner & E.Klein) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella austriaca (Teppner & E.Klein) P.Delforge; Naturalistes Belges 72(3): 100 (1991)
subspecies :
subsp. bucegiana
Hedrén, Anghelescu & R.Lorenz; J. Eur. Orch. 54(3-4): 293 (2022)
Distribution: Romania (Bucegi Mts.)
13 ROM
Remarks: tetraploid apomict (2n - 80)
subspecies :
subsp. carpatica
(Zapal.) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 717 (2005)
Distribution: Romania; Ukraine (E-Carpathians)
13 ROM 14 UKR
Synonyms: hidden
= Gymnadenia carpatica (Zapal.) Teppner & E.Klein; Phyton (Horn) 38(1): 221 (1998)
= Nigritella angustifolia var. carpatica Zapal.; Rozpr. Wydz. Mat.-Przyr. Akad. Umiejetn., Dzial B, Nauki Biol., Ser. 3, 6B(46B): 215 (1906)
= Nigritella carpatica (Zapal.) Teppner, E.Klein & Zagulskij; Phyton (Horn) 34(2): 171 (1994)
= Nigritella nigra var. carpatica (Zapal.) Pawl.; Bull. Int. Acad. Polon. Sci., Cl. Sci. Math., Sér. B 1, Bot. 1947: 85, 96 (1947)
= Nigritella rubra f. carpatica (Zapal.) Soó; Repert. Spec. Nov. Regni Veg. 24: 33 (1927)
subspecies :
subsp. gallica
E.Breiner & R.Breiner; Mitt. Arbeitskreises Heimische Orchideen 25(4): 471 (1993)
Distribution: Switzerland; France (Jura, Massif Central, Alps); NW-Italy
11 SWI 12 FRA 13 ITA
Remarks: tetraploid apomict (2n - 80)
Synonyms: hidden
= Gymnadenia austriaca var. gallica (E.Breiner & R.Breiner) P.Delforge; Naturalistes Belges 79(4): 255 (1998)
= Gymnadenia nigra subsp. gallica (E.Breiner & R.Breiner) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella gallica (E.Breiner & R.Breiner) Landolt; Fl. Indicativa 269 (2010)
= Nigritella nigra var. gallica (E.Breiner & R.Breiner) Wenker & S.Wenker; Jahresber. Naturwiss. Vereins Wuppertal 60: 200 (2007)
= Nigritella rhellicani subsp. gallica (E.Breiner & R.Breiner) Kerguélen; Bull. Assoc. Inform., Appl. Bot. (Marseille) 1: 183 (1994)
subspecies :
subsp. iberica
Teppner & E.Klein; Phyton (Horn) 33(2): 192 (1993)
Distribution: Spain; Andorra; France (Pyrénées)
12 ADO FRA SPA
Remarks: tetraploid apomict (2n - 80). Kreutz (2024) lists the species also from the W-Alps and C-France, which needs to be confirmed.
Synonyms: hidden
= Gymnadenia austriaca subsp. iberica (Teppner & E.Klein) L.Sáez & Aymerich; Orsis 29: 40 (2015)
= Gymnadenia austriaca var. iberica (Teppner & E.Klein) G.Foelsche & W.Foelsche; J. Eur. Orch. 31(4): 803 (1999)
= Gymnadenia nigra subsp. iberica (Teppner & E.Klein) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella austriaca subsp. iberica (Teppner & E.Klein) L.Sáez; Anales Jard. Bot. Madrid 61(1): 87 (2004)
= Nigritella iberica (Teppner & E.Klein) Kreutz; Guide Orchids Europe N. Afr. Middle E. 296 (2024)
= Nigritella nigra var. iberica (Teppner & E.Klein) Wenker & S.Wenker; Jahresber. Naturwiss. Vereins Wuppertal 60: 199 (2007)
= Nigritella rhellicani subsp. iberica (Teppner & E.Klein) Kerguélen; Bull. Assoc. Inform., Appl. Bot. (Marseille) 1: 183 (1994)
subspecies :
subsp. lithopolitanica
(Ravnik) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 717 (2005)
Name: Steineralpen-Kohlröschen
Distribution: Austria; NW-Slovenia (SE-Alps)
11 AUT 13 SLV
Conservation Status: IUCN R
Synonyms: hidden
= Gymnadenia lithopolitanica (Ravnik) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella lithopolitanica Ravnik; Acta Bot. Croat. 37: 226 (1978)
= Nigritella lithopolitanica var. luciae W.Foelsche, G.Foelsche, R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 158 (2020)
species :
Nigritella ravnikii
W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic; J. Eur. Orch. 49(3-4): 508 (2017)
Distribution: NW-Slovenia; NE-Italy
13 ITA SLV
Remarks: diploid taxon
Synonyms: hidden
= Gymnadenia ravnikii (W.Foelsche, R.Wüest, Dolinar, Daksk. & Pausic) J.M.H.Shaw; Orchid Rev., Suppl., 127 (1326): 45 (2019)
species :
Nigritella rhellicani
Teppner & E.Klein; Phyton (Horn) 31(1): 7 (1990)
subspecies :
subsp. rhellicani
Name: Rhellicanus-Kohlröschen, Alpen-Kohlröschen (DE)
Distribution: Germany (Bayern); Switzerland; Liechtenstein; Austria; Slovakia; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Albania; Romania; Bulgaria; Greece (very rare Rodopi, Mt. Falakron, Mt. Voras, +Mt. Karpenissio, Mt. Giona)
11 AUT GER LIE SLO SWI 12 FRA 13 ALB BIH BUL CRO GRC ITA ROM SLV
Synonyms: hidden
= Gymnadenia nigra subsp. rhellicani (Teppner & E.Klein) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Gymnadenia rhellicani (Teppner & E.Klein) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Gymnadenia rhellicani f. fulva (G.Keller) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella nigra lusus fulva G.Keller; Monogr. Iconogr. Orchid. Eur. 2: 267 (1933)
= Nigritella nigra subsp. rhellicani (Teppner & E.Klein) H.Baumann, Künkele & R.Lorenz; J. Eur. Orch. 36(3): 776 (2004)
= Nigritella nigra var. rosea Vis. & Sacc.; Atti Reale Ist. Veneto Sci. Lett. Arti 14: 321 (1868)
= Nigritella nigra var. rosea Vis. & Sacc. ex Goiran; Nuovo Giorn. Bot. Ital. 15: 38 (1883) [fide Baumann, Künkele & R. Lorenz 2002]
= Nigritella rhellicani f. fulva (G.Keller) Teppner & E.Klein; Phyton (Horn) 31(1): 10 (1990)
= Nigritella rhellicani var. rosea (Vis. & Sacc. ex Goiran) Teppner & E.Klein; Phyton (Horn) 31(1): 10 (1990)
subspecies :
subsp. cenisia
(G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) Kreutz; Eurorchis 17: 107 (2005)
Name: Mont-Cenis-Kohlröschen, Nigritelle du Mont Cenis
Distribution: SE-France (SW-Alps); NW-Italy
12 FRA 13 ITA
Synonyms: hidden
= Gymnadenia cenisia (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud; Naturalistes Belges 80(3): 374 (1999)
= Gymnadenia nigra subsp. cenisia (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Gymnadenia rhellicani var. robusta P.Delforge; Naturalistes Belges 84 (Orchid. 16): 127 (2003)
= Nigritella cenisia G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud; Orchidophile (Paris) 134: 248 (1998)
= Nigritella rhellicani var. cenisia (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) Wenker & S.Wenker; Jahresber. Naturwiss. Vereins Wuppertal 60: 196 (2007)
= Nigritella rhellicani var. robusta (P.Delforge) Kreutz; Kompend. Eur. Orchid. 78 (2004)
subspecies :
subsp. gabasiana
(Teppner & E.Klein) Kreutz; Eurorchis 17: 107 (2005)
Distribution: N-Spain (Pyrénées, Cordillera Cantabrica); Andorra; SW-France (Pyrénées)
12 ADO FRA SPA
Synonyms: hidden
= Gymnadenia gabasiana (Teppner & E.Klein) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Gymnadenia nigra subsp. gabasiana (Teppner & E.Klein) J.-M.Tison; Biocosme Mésogéen 27(4): 127 (2010)
= Nigritella gabasiana Teppner & E.Klein; Phyton (Horn) 33(2): 182 (1993)
= Nigritella nigra subsp. gabasiana (Teppner & E.Klein) H.Baumann & R.Lorenz; J. Eur. Orch. 37: 717 (2005)
= Nigritella nigra var. pyrenaica Schltr.; Repert. Spec. Nov. Regni Veg. 16: 271 (1919)
species :
Nigritella rubra
(Wettst.) K.Richt.; Pl. Eur. [K.Richter] 1: 278 (1890)
subspecies :
subsp. rubra
Name: Rotes Kohlröschen (DE)
Distribution: S-Germany (Bavarian Alps); Switzerland; Liechtenstein; Austria; Italy; NW-Slovenia
11 AUT GER LIE SWI 13 ITA SLV
Remarks: nigra agg.
Synonyms: hidden
= Gymnadenia miniata (Crantz) Hayek; Fl. Steiermark [Hayek] 2(2): 133 (1956)
= Gymnadenia miniata f. rosea (C.Boillat & V.Boillat) P.Delforge; Naturalistes Belges 96(Orchid. 28): 16 (2015)
= Gymnadenia nigra subsp. rubra (Wettst.) H.Sund.; Europ. Medit. Orchid., ed. 3: 41 (1980)
= Gymnadenia nigra var. rubra (Wettst.) Kraenzl.; Orchid. Gen. Sp. 1(2): 559 (1898)
= Gymnadenia rhellicani var. rosea (Vis. & Sacc. ex Goiran) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Gymnadenia rubra f. rosea (C.Boillat & V.Boillat) P.Delforge; Naturalistes Belges 92(Orchid. 24): 108 (2011)
= Gymnadenia rubra Wettst.; Verh. K.K. Zool.-Bot. Ges. Wien (1889) 83, et Ber. Deutsch. Bot. Ges. 7: 312 (1889)
= Gymnadenia rubra Wettst. ex Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella miniata (Crantz) Janch.; Phyton (Horn) 8: 232 (1959)
= Nigritella nigra subsp. rubra (Wettst.) Beauverd; Bull. Soc. Bot. Genève, Sér. 2, 17: 337 (1926)
= Nigritella nigra var. rubra (Wettst.) W.D.J.Koch; Syn. Deut. Schweiz. Fl., ed. 3 [Hallier & Brand]: 2434 (1905)
= Nigritella rubra f. carpatica Soó; Repert. Spec. Nov. Regni Veg. 24: 33 (1927)
= Nigritella rubra f. rosea C.Boillat & V.Boillat; J. Eur. Orch. 42(3-4): 491 (2010)
= Nigritella rubra f. wettsteinii Soó; Repert. Spec. Nov. Regni Veg. 24: 33 (1927)
= Orchis miniata Crantz; Stirp. Austr. Fasc., ed. 2, 2: 486 (1769), nom. rej. prop.
variant :
var. minor
(W.Foelsche & Zernig) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008)
Distribution: Austria (Oststeiermark, Mt. Trenchtling)
11 AUT
Synonyms: hidden
= Gymnadenia miniata f. minor (W.Foelsche & Zernig) P.Delforge; Naturalistes Belges 96(Orchid. 28): 16 (2015)
= Gymnadenia minor (W.Foelsche & Zernig) W.Foelsche, Zernig & O.Gerbaud; Bull. Soc. Franç. Orchidophilie Rhône-Alpes 24: 23 (2011)
= Gymnadenia minor var. obscurior (W.Foelsche & D.Foelsche) O.Gerbaud, W.Foelsche & D.Foelsche; Orchidophile (Paris) 55(4): 366 (2024)
= Gymnadenia rubra f. minor (W.Foelsche & Zernig) P.Delforge; Naturalistes Belges 92(Orchid. 24): 107 (2011)
= Nigritella minor var. obscurior W.Foelsche & D.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 40(2): 125 (2024)
= Nigritella minor W.Foelsche & Zernig; Joannea Bot. 6: 10 (2007)
subspecies :
subsp. buschmanniae
(Teppner & Ster) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 717 (2005)
Name: Buschmanns Kohlröschen
Distribution: N-Italy (Brenta)
13 ITA
Synonyms: hidden
= Gymnadenia buschmanniae (Teppner & Ster) Teppner & E.Klein; Phyton (Horn) 38(1): 222 (1998)
= Nigritella buschmanniae Teppner & Ster; Phyton (Horn) 36(2): 278 (1996)
subspecies :
subsp. dolomitensis
(Teppner & E.Klein) D.Wenker & S.Wenker; Jahresber. Naturwiss. Vereins Wuppertal 60: 177-250 (2007)
Name: Dolomiten-Kohlröschen (DE)
Distribution: N-Italy (Dolomiti)
13 ITA
Synonyms: hidden
= Gymnadenia dolomitensis Teppner & E.Klein; Phyton (Horn) 38: 223 (1998)
= Gymnadenia miniata var. dolomitensis (Teppner & E.Klein) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 16 (2015)
= Gymnadenia rubra var. dolomitensis (Teppner & E.Klein) P.Delforge; Naturalistes Belges 92(Orchid. 24): 108 (2011)
= Nigritella dolomitensis (Teppner & E.Klein) Hedrén, E.Klein & Teppner; Phyton (Horn) 40: 243 (2000)
= Nigritella miniata var. dolomitensis (Teppner & E.Klein) H.Baumann, R.Lorenz & Perazza; Gredleriana 12 (2012): 93 (2013)
= Nigritella rubra subsp. dolomitensis (Teppner & E.Klein) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 164 (2007), isonym
= Nigritella rubra var. dolomitensis (Teppner & E.Klein) R.Lorenz & Perazza; GIROS Notiz. 27: 8 (2004)
subspecies :
subsp. archiducis-johannis
(Teppner & E.Klein) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 717 (2005)
Name: Erzherzog-Johann-Kohlröschen
Distribution: Austria (Totes Gebirge); NW-Slovenia (Mt. Triglav)
11 AUT 13 SLV
Synonyms: hidden
= Gymnadenia archiducis-joannis (Teppner & E.Klein) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella archiducis-joannis Teppner & E.Klein; Phyton (Horn) 25(1): 168 (1985)
= Nigritella rubra subsp. archiducis-joannis (Teppner & E.Klein) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008)
subspecies :
subsp. hygrophila
(W.Foelsche & Heidtke) comb.ined.
Distribution: Austria (Steiermark); ?SE-France (?Mt. Cenis); Italy (Südtirol: Passo Pordoi); Slovenia (Triglav)
11 AUT 13 ITA SLV
Remarks: provisionally accepted name, status not yet confirmed by genetics
Synonyms: hidden
= Gymnadenia hygrophila (W.Foelsche & Heidtke) W.Foelsche, Heidtke & O.Gerbaud; Bull. Soc. Franç. Orchidophilie Rhone-Alpes 24: 23 (2011)
= Gymnadenia miniata f. hygrophila (W.Foelsche & Heidtke) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 16 (2015)
= Gymnadenia rubra f. hygrophila (W.Foelsche & Heidtke) P.Delforge; Naturalistes Belges 92(Orchid. 24): 107 (2011)
= Nigritella hygrophila var. pauciflora W.Foelsche, G.Foelsche, R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 164 (2020)
= Nigritella hygrophila W.Foelsche & Heidtke; J. Eur. Orch. 43(1): 143 (2011)
subspecies :
subsp. stiriaca
(Rech.) H.Baumann & R.Lorenz; J. Eur. Orch. 37(3): 718 (2005)
Name: Steirisches Kohlröschen
Distribution: Austria (Oberösterreich, Steiermark, Salzburg)
11 AUT
Synonyms: hidden
= Gymnadenia rubra var. stiriaca Rech.; Mitt. Naturwiss. Vereines Steiermark 42: 148 (1906)
= Gymnadenia stiriaca (Rech.) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella rubra subsp. stiriaca (Rech.) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008), isonym
= Nigritella stiriaca (Rech.) Teppner & E.Klein; Phyton (Horn) 25(1): 159 (1985)
species :
Nigritella savogiensis
W.Foelsche, G.Foelsche, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 39(1-2): 166 (2022)
Distribution: SE-France (Hte. Savoie)
12 FRA
Remarks: provisionally accepted name, not yet confirmed by genetics
species :
Nigritella sulzeri
W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 40(2): 111 (2024)
Distribution: Austria
11 AUT
Remarks: provisionally accepted name, not yet confirmed by genetics
Synonyms: hidden
= Gymnadenia sulzeri (W.Foelsche) O.Gerbaud & W.Foelsche; Orchidophile (Paris) 55(4): 366 (2024)
species :
Nigritella widderi
Teppner & E.Klein; Phyton (Horn) 25(2): 318 (1985)
Name: Widders Kohlröschen
Distribution: SE-Germany (Bavarian Alps); N-Austria (Steiermark); Italy (Abruzzes, Dolomiti)
11 AUT GER 13 ITA
Synonyms: hidden
= Gymnadenia widderi (Teppner & E.Klein) Teppner & E.Klein; Phyton (Horn) 38(1): 220-224 (1998)
= Nigritella rubra subsp. widderi (Teppner & E.Klein) H.Baumann & R.Lorenz; J. Eur. Orch. 37: 718 (2005)
= Nigritella rubra subsp. widderi (Teppner & E.Klein) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 20 (2008), isonym
species :
Nigritella suceveana
Kreutz, Bobocea & O.Gerbaud; Bull. S.F.O. Rhône-Alpes 44: 35 (2021)
Distribution: Romania; W-Ukraine (Carpathians)
13 ROM 14 UKR
Remarks: provisionally accepted name, not yet confirmed by genetics
species :
Nigritella ×delphineae
M.Gerbaud & O.Gerbaud; Orchidophile (Paris) 120: 33, 30 (1996)
Distribution: SE-France; N-Italy
12 FRA 13 ITA
Remarks: Nigritella corneliana x rhellicani
Synonyms: hidden
= Gymnadenia ×breinerorum O.Gerbaud & W.Foelsche; Naturalistes Belges 80(3): 381 (1999)
= Gymnadenia ×delphineae (M.Gerbaud & O.Gerbaud) M.Gerbaud & O.Gerbaud; Naturalistes Belges 80(3): 378 (1999)
= Gymnadenia ×nigra nothosubsp. delphineae (M.Gerbaud & O.Gerbaud) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 269 (2012)
= Nigritella ×breinerorum (O.Gerbaud & W.Foelsche) W.Foelsche; Joannea Bot. 7: 47 (2009)
species :
Nigritella ×eggeriana
(O.Gerbaud) W.Foelsche; Joannea Bot. 7: 47 (2009)
Distribution: France
12 FRA
Remarks: Nigritella austriaca (var. gallica) x rhellicani
Synonyms: hidden
= Gymnadenia nigra nothosubsp. eggeriana (O.Gerbaud) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 269 (2011 publ. 2012)
= Gymnadenia ×eggeriana O.Gerbaud; Naturalistes Belges 80(3): 378 (1999)
species :
Nigritella ×foelscheana
R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 135 (2020)
Distribution: Slovenia
13 SLV
Remarks: Nigritella kossutensis x N. ravnikii
species :
Nigritella ×fohringeri
Griebl; Ber. Arbeitskreis. Heimische Orchid. 28(2): 169 (166-170; fig.) (2011)
Distribution: Austria
11 AUT
Remarks: Nigritella lithopolitanica x miniata
Synonyms: hidden
= Gymnadenia ×fohringeri (Griebl) J.M.H.Shaw; Orchid Rev. 122 (1306): 39 (2014)
species :
Nigritella ×hennigsiana
R.Wüest, E.Merz, M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 130 (2020)
Distribution: Slovenia
13 SLV
Remarks: Nigritella lithopolitanica x karawankarum
species :
Nigritella ×petzenensis
F.Fohringer & Redl; Linzer Biol. Beitr. 36(1): 573 (2004)
Distribution: Austria; Slovenia
11 AUT 13 SLV
Remarks: Nigritella rhellicani x lithopolitanica
Synonyms: hidden
= Gymnadenia ×petzenensis (F.Fohringer & Redl) Oddone; GIROS Orch. Spont. Eur. 59(2): 357 (2016)
species :
Nigritella ×robatschiana
(O.Gerbaud & W.Foelsche) W.Foelsche; Joannea Bot. 7: 47 (2009)
Distribution: France; ?Italy
12 FRA
Remarks: Nigritella cenisia x rhellicani
Synonyms: hidden
= Gymnadenia ×robatschiana O.Gerbaud & W.Foelsche; Naturalistes Belges 80(3): 380 (1999)
species :
Nigritella ×teppneri
W.Foelsche; Ber. Arbeitskreis. Heimische Orchid. 32(1): 88 (2015)
Distribution: N-Italy
13 ITA
Remarks: Nigritella hygrophila x rhellicani
Synonyms: hidden
= Gymnadenia ×teppneri (W.Foelsche) Oddone; GIROS Orch. Spont. Eur. 59(2): 357 (2016)
species :
Nigritella ×wettsteiniana
(O.Abel) Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 3: 811 (1907)
Distribution: Switzerland; Austria; Italy; Romania
11 AUT SWI 13 ITA ROM
Remarks: Nigritella nigra s. l. x Nigritella rubra
Synonyms: hidden
= Gymnadenia ×bornmuelleri Dalla Torre & Sarnth.; Fl. Tirol [Dalla Torre & Sarnthein] 6(1): 531 (1906)
= Gymnadenia ×wettsteiniana O.Abel; Verh. K.K. Zool.-Bot. Ges. Wien 47: 611 (1897)
= Nigritella ×wettsteiniana (O.Abel) Schltr.; Repert. Spec. Nov. Regni Veg. 16: 271 (1919), isonym
species :
Nigritella ×wulfeniana
M.Gerbaud & O.Gerbaud; Ber. Arbeitskreis. Heimische Orchid. 37(2): 125 (2020)
Distribution: Slovenia
13 SLV
Remarks: Nigritella karawankarum x N. ravnikii
