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V. 12.5
14.10.2021

125.0000 Family Orchidaceae Juss.; Gen. Pl.: 64 (1789), nom. cons.
over 800 genera, more than 30.000 spp. worldwide, mostly tropical
Species listed in database: 30547; hybrids: 1294
= Apostasiaceae Lindl., Nix. Pl.: 22 (1833), nom. cons.
= Cypripediaceae Lindl., Nix. Pl.: 22 (1833)
= Limodoraceae Horan., Char. Ess. Fam.: 44 (1847)
= Liparidaceae Vines, Stud. Text-book Bot. 2: 567 (1895)
= Neottiaceae Horan., Prim. Lin. Syst. Nat.: 50 (1834)
= Neuwiediaceae R.Dahlgren ex Reveal & Hoogland, Bull. Mus. Natl. Hist. Nat., ser. 4, sect. B Adansonia 13: 91 (1991)
= Ophrydaceae Vines, Stud. Text-book Bot. 2: 566 (1895)
= Pycnanthaceae Ravenna, Onira 12(6): 29 (2011)
= Vanillaceae Lindl., Key Bot.: 73 (1835)
125.4356 Genus Ophrys L.; Sp. Pl. 2: 945, 948 (1753)
Distribution: mostly Mediterranean and Anatolia, few to W-Asia and central Europe. The taxonomy of the genus is currently very chaotic due to many recently described microspecies and various conflicting species concepts. Some authors recognize almost 400 microtaxa. Molecular results indicate that possibly only 10-50 very variable species are involved.
Species listed in database: 110; hybrids: 483
= Arachnites F.W.Schmidt, Fl. Boem. 1: 74 (1793)
= Myodium Salisb., Trans. Hort. Soc. 1: 289 (1812)
species :
Ophrys aegilica Engelen; Fl. Medit. 30: 281, figs. 2-3, 6-7 (2020)
Distribution: Greece
13 GRC
Remarks: provisionally accepted name
species :
Ophrys aesculapii Renz; Fedde, Repert. 25: 249 (1928)
Name: Äskulap-Ragwurz
Distribution: C-Greece; S-Greece
13 GRC
Remarks: mammosa agg.
Conservation Status: IUCN R
Synonyms: hidden
= Ophrys aesculapii ssp. pseudoaranifera Renz, Repert. Spec. Nov. Regni Veg. 25: 250 (1928)
= Ophrys aranifera ssp. aesculapii (Renz) Soó, Acta Bot. Acad. Sci. Hung. 16: 383 (1970)
= Ophrys aranifera ssp. renzii Soó, Repert. Spec. Nov. Regni Veg. 26: 278 (1929)
= Ophrys renzii (Soó) Soó, Fedde, Repert. 26: 278 (1929)
= Ophrys sphegodes ssp. aesculapii (Renz) Soó ex J.J.Wood, Orchidee 31 (6): 232 (1980)
species :
Ophrys alasiatica Kreutz, Segers & Walraven; Jour. Eur. Orch. 34 (3): 480 (2002)
Distribution: Cyprus
34 CYP
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys cypricola Paulus, [ms.]
= Ophrys sphegodes ssp. alasiatica (Kreutz, Segers & Walraven) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 728 (2005)
species :
Ophrys algeriensis Kreutz & L.Lewis; Ber. Arbeitskreis. Heimische Orchid. 36(1): 250 (2019)
Distribution: Algeria
20 ALG
Remarks: provisionally accepted name
species :
Ophrys alpujata Riech. & H.Kohlmüller; Ber. Arbeitskreis. Heimische Orchid. 36(1): 268 (2019)
Distribution: Spain
12 SPA
Remarks: provisionally accepted name
species :
Ophrys amanensis (Nelson ex Renz & Taubenheim) P.Delforge; Nat. Belg. 72 (3): 101 (1991)
subspecies :
subsp. amanensis
Distribution: S-Turkey
34 TUR
Remarks: mammosa agg.
Conservation Status: IUCN R
Synonyms: hidden
= Ophrys transhyrcana ssp. amanensis Nelson ex Renz & Taubenheim, Notes Roy. Bot. Gard. Edinb. 41 (2): 271 (1983)
= Ophrys sphegodes ssp. amanensis (E.Nelson ex Renz & Taubenheim) H.A.Pedersen, P.J.Cribb & Rolf Kühn, Field Guide Orchids Europe Medit. 242 (2019)
subspecies :
subsp. antalyensis (Kreutz & Seckel) Kreutz; Kompend. Eur. Orchid. 80 (2004)
Distribution: S-Turkey (Antalya)
34 TUR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys antalyensis Kreutz & Seckel, Kreutz, Orchideen Türkei 752 (1998)
subspecies :
subsp. iceliensis (Kreutz) Kreutz; Kompend. Eur. Orchid. 80 (2004)
Distribution: C-Turkey
34 TUR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys iceliensis Kreutz, J. Eur. Orchid. 32 (3/4): 527-537 (2000)
species :
Ophrys apifera Huds.; Fl. Angl. ed. I, 340 (1762)
Name: Bienen-Ragwurz (DE); Bee Orchid (EN)
Distribution: Albania; Austria; Belgium; Luxembourg; Baleares; England; Channel Isl.; Corsica; Crete; Czech Republic; Slovakia; France; Germany; Greece (incl. Kyklades); Ireland; Luxembourg; Switzerland; Liechtenstein; Malta; Netherlands; Spain; Gibraltar; Hungary; Italy; Serbia & Kosovo; Montenegro; Macedonia; Bosnia & Hercegovina; Slovenia; Croatia; Portugal; Romania; Sardinia; Sicily; European Turkey; Crimea; W-Northern Caucasus; Georgia [Caucasus] (Abkhazia); East Aegaean Isl. (including Rhodos, Karpathos, Nisyros, Tilos); Cyprus; Israel; Syria; Morocco; Algeria; Tunisia; Libya
10 GRB IRE 11 AUT BGM CZE GER HUN LIE LUX NET SLO SWI 12 BAL CIS COR FRA GIB POR SAR SPA 13 ALB BIH CRO GRC ITA KRI MLT MNT NMA ROM SIC SLV SRB TUE 14 KRY 20 ALG LBY MOR TUN 33 GEO NCS 34 CYP EAI ISR SYR
Remarks: apifera agg.
Synonyms: hidden
= Arachnites apifera (Huds.) Hoffm., Deutsch. Fl. ed. 2, 2: 180 (1895)
= Ophrys albiflora Spruner ex Boiss., Fl. Orient. 5: 79 (1884)
= Ophrys apifera ssp. austriaca (Wiesb. ex Dichtl) K.Richt., Pl. Eur. 1: 264 (1890)
= Ophrys apifera var. austriaca (Wiesb. ex Dichtl) Beck, Fl. Nieder-Österr. 198 (1890)
= Ophrys apifera ssp. muteliae (Mutel) K.Richt., (1905)
= Ophrys apifera var. muteliae Mutel, Ann. Sci. Nat., Bot. II, 3: 243 (1835)
= Ophrys apifera f. flaveola Klaver & Kreutz, GIROS Orch. Spont. Eur. 59(1): 48 (2016)
= Ophrys aquisgranensis Kaltenb., Verh. Rheinl. 7: 519, t. 7 (1850)
= Ophrys arachnites Mill., Gard. Dict. ed. 8: 7 (1768)
= Ophrys austriaca Wiesb. ex Dichtl, Deutsch. Bot. Monatsschr. 1: 148 (1883)
= Ophrys broteroi Welw. ex Nyman, Consp. 698 (1882)
= Ophrys chlorantha Hegetschw., Fl. Schweiz 876 (1839)
= Ophrys pseudoapifera Caldesi, (1880)
= Ophrys ripaensis Porta, Atti Acc. Rovereto, Ser. 3: 11: fasc. 2, 7 (1905), Fedde, Repert. Nov. Sp. 8: 485
= Ophrys rostrata Ten., Ind. Sem. Hort. Neap. (1830) 15
= Ophrys sphegodes Mill., Gard. Dict. ed. 8, n. 8 (1768)
= Ophrys trollii Hegetschw., Fl. Schweiz 874, t. 8 (1839)
= Orchis apifera Salisb., Prod. 7 (1796)
variant :
var. almaracensis Pérez-Chisc., Durán Oliva & Gil Llano; Stud. Bot. (Salamanca), 9: 114, 113 (1990 publ. 1991)
Distribution: Spain
12 SPA
Synonyms: hidden
= Ophrys apifera f. almaracensis (Pérez-Chisc., Durán Oliva & Gil Llano) P.Delforge, Naturalistes Belges 88(Orchid. 20): 246 (2007)
variant :
var. atrofusca Kreutz; J. Eur. Orch. 48(2-4): 329 (2016)
Distribution: Belgium
11 BGM
variant :
var. aurita (Moggr.) Gremli; Neue Beitr. Fl. Schweiz 4: 31 (1887)
Distribution: Switzerland; France; Germany; etc.
11 GER SWI 12 FRA
Synonyms: hidden
= Ophrys insectifera subvar. aurita Moggr., Nova Acta Acad. Leop. Carol. 35: 13, tab. 4 (1869)
variant :
var. badensis Lewis & Kreutz; J. Eur. Orch. 44(2): 403 - 412 (2012)
Distribution: rare; but throughout the range of the species
Remarks: this form was treated usually as "var. friburgensis", but the name friburgensis applies to var. botteronii
Synonyms: hidden
= Ophrys apifera f. badensis (L.Lewis & Kreutz) P.Delforge, Naturalistes Belges 93(Orchid. 25): 242 (2012)
variant :
var. basiliensis S.Schwegler & Matthies; Orchid Rev. 112(1258): 214 (2004)
Distribution: Switzerland
11 SWI
Synonyms: hidden
= Ophrys apifera f. basiliensis (S.Schwegler & Matthies) P.Delforge, Naturalistes Belges 87(Orchid. 19): 259 (2006)
variant :
var. belgarum Turner Ettl.; Watsonia 22 (1): 107 (1998)
Distribution: England; Belgium; etc.
10 GRB 11 BGM
Synonyms: hidden
= Ophrys apifera f. belgarum (Turner Ettl.) P.Delforge, Naturalistes Belges 88(Orchid. 20): 246 (2007)
variant :
var. botteronii (Chodat) Brand; Koch, Syn. Deutsch. Schweiz. Fl. ed 3, 3: 2438 (1905)
Distribution: rare; but throughout the range of the species
Synonyms: hidden
= Ophrys apifera f. friburgensis (Freyhold) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
= Ophrys apifera ssp. friburgensis (Freyhold) P.Fourn., Fl. Compl. plaine franc. 516 (1928)
= Ophrys friburgensis (Freyhold) Nägeli, Ber. Schweiz. Bot. Ges. 21: 178 (1912)
= Ophrys ×botteronii Chodat, [Notice sur Polyg. d´Eur. & d´Or. (1887)], Bull. Soc. Bot. Geneve v. (1889) 187
= Ophrys apifera f. botteronii (Chodat) P.D.Sell, Fl. Gr. Brit. Ireland 5: 365 (1996)
= Ophrys apifera ssp. botteronii (Chodat) Hegi, Ill. Fl. Mitteleur. ed. 1, 2: 334 (1909)
= Ophrys ×epeirophora Peter, Flora 66: 10 (1883)
variant :
var. brevilabellata Kreutz, R.Selig & Zelesny; Ber. Arbeitskreis. Heimische Orchid. 24(1): 93 (2007)
Distribution: France (Cevennes)
12 FRA
Synonyms: hidden
= Ophrys apifera f. brevilabellata (Kreutz, R.Selig & Zelesny) P.Delforge, Naturalistes Belges 89(Orchid. 21): 251 (2008)
variant :
var. cambrensis M.J.Clark; J. Eur. Orch. 46(3/4): 726 (2014)
Distribution: Wales
10 GRB
variant :
var. chlorantha (Hegetschw.) K.Richt.; Pl. Eur. 1: 264 (1890)
Distribution: rare throughout the range of the species
Synonyms: hidden
= Ophrys apifera ssp. chlorantha (Hegetschw.) Arcang., Comp. fl. ital. ed. 1: 661 (1882)
variant :
var. curviflora A.Soulié; Orchidophile (Asnieres) 163: 296 (2004)
Distribution: France (Aveyron); W-Germany
11 GER 12 FRA
Synonyms: hidden
= Ophrys apifera f. curviflora (A.Soulié) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
variant :
var. fulvofusca M.P.Grasso & Scrugli; Orchidee 38 (1): 47 (1987)
Distribution: Sardinia
12 SAR
Synonyms: hidden
= Ophrys apifera f. fulvofusca (M.P.Grasso & Scrugli) P.Delforge, Naturalistes Belges 88(Orchid. 20): 246 (2007)
variant :
var. immaculata Bréb.; Fl. Normandie ed. 2: 261 (1849)
Distribution: very rare through the range of the species
Synonyms: hidden
= Ophrys apifera f. bicolor (Nägeli) P.D.Sell, Fl. Gr. Brit. Ireland 5: 365 (1997)
= Ophrys apifera var. bicolor (Nägeli) E.Nelson, Monogr. Ikon. Ophrys 178 (1962) [nom. illeg.]
= Ophrys bicolor Nägeli, Ber. Schweiz. Bot. Ges. 23: 64 (1914)
= Ophrys apifera f. bicolor (Nägeli) P.D.Sell, Fl. Great Britain & Ireland 5: 365 (1996)
= Ophrys immaculata (Bréb.) Nägeli, Jahresh. vaterl. Ver. Naturk. Württ. 72: 217 (1916)
= Ophrys mangini Tallon, Poucel, Découv. Orch. France 131 (1942)
= Ophrys ×luizetii E.G.Camus, Bull. Soc. Bot. Fr. 38: 39, 43 (1891)
variant :
var. jenensis Voelckel & E.Theel; Orchidee (Hamburg) 3(14)E: 95 (2017)
Distribution: Germany
11 GER
variant :
var. jurana (Ruppert) J.Schuster; Repert. Spec. Nov. Regni Veg., Beih. 60: 435 (1935)
Distribution: rare through the range of the species
Synonyms: hidden
= Ophrys ×jurana W.Zimm. ex A.Fuchs, Repert. Spec. Nov. Regni Veg., Beih. 51: 136 (1928)
= Ophrys jurana (Ruppert) Neuberger, Schulfl. Baden ed. 7-8: 57 (1925)
= Ophrys apifera ssp. jurana Ruppert, Allg. Bot. Zeitschr. 17: 2 (1911)
variant :
var. laetitiae Klaver; GIROS Notiz. 45: 16 (2010)
Distribution: Italy
13 ITA
variant :
var. patinata U.Grabner & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 27(2): 62 (2010 publ. 2011)
Distribution: Croatia
13 CRO
Synonyms: hidden
= Ophrys apifera f. patinata (U.Grabner & Kreutz) P.Delforge, Naturalistes Belges 93(Orchid. 25): 242 (2012)
variant :
var. renatae K.Barbieri, L.Barbieri & Contorni; GIROS Orch. Spont. Eur. 62(2): 448 (2019)
Distribution: Italy
13 ITA
variant :
var. saraepontana (Ruppert) H.Baumgartner bis & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 27(2): 215 (2010 publ. 2011)
Distribution: Germany
11 GER
Synonyms: hidden
= Ophrys ×saraepontana Ruppert, Verh. Nat. Ver. Preuss. Rheinl., 81: 181 (1925), et Fedde, Repert. 31: 325 (1924 publ. 1926)
variant :
var. tilaventina Nonis & Liverani; Caesiana 9: 57 (1997)
Distribution: Italy (Pordenone)
13 ITA
Synonyms: hidden
= Cephalophrys integra (Sacc.) Cox, (1912) [pro hybr.]
= Ophrys apifera f. tilaventina (Nonis & Liverani) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
= Ophrys apifera ssp. purpurea (Tausch) K.Richt., Pl. Eur. 1: 264 (1890)
= Ophrys integra Sacc., Nuov. Giorn. Bot. Ital. 3: 165 (1871)
= Ophrys purpurea Tausch, Flora 14: 222 (1831)
variant :
var. trollii (Hegetschw.) Rchb.fil.; Icon. fl. germ. helv. 13/14: 97 (1851)
Distribution: very rare through the range of the species
Synonyms: hidden
= Ophrys apifera ssp. trollii (Hegetschw.) K.Richt., Pl. Eur. 1: 264 (1890)
= Ophrys apifera ssp. trollii (Hegetschw.) O.Bolòs, Bull. Inst. Catalana Hist. Nat., 59 Bot., 8: 145 (1991)
= Ophrys apifera f. trollii (Hegetschw.) P.D.Sell, Fl. Great Britain & Ireland 5: 365 (1996)
= Ophrys penedensis Diettr.-Kalkh., Allg. Bot. Zeitschr. 20: 81 (1914)
= Ophrys trollii Hegetschw., Fl. Schweiz 874 (1839)
variant :
var. urbinensis Klaver; GIROS Orch. Spont. Eur. 59(2): 288 (2016)
Distribution: Italy
13 ITA
species :
Ophrys arachnitiformis Gren. & Philippe; Mem. Soc. Emul. Doubs, ser. 3, 4: 391 (1860)
subspecies :
subsp. arachnitiformis
Distribution: SE-France; NW-Italy
12 FRA 13 ITA
Remarks: sphegodes agg., probably a synonym of O. exaltata
Synonyms: hidden
= Ophrys arachnites ssp. arachnitiformis (Gren. & Philippe) K.Richt., Pl. Eur. 1: 262 (1890)
= Ophrys aranifera var. nicaeensis Barla, Fl. Illus. Nice, Orchid. 66 (1868)
= Ophrys arachnitiformis f. nicaeensis (Barla) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
= Ophrys aranifera var. specularia Rchb.fil., Icon. Fl. Germ. Helv. 13/14: 90, 91 (1851)
= Ophrys arachnitiformis f. specularia (Rchb.fil.) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
= Ophrys exaltata ssp. arachnitiformis (Gren. & Philippe) Del Prete, Webbia 37 (2): 251 (1984)
= Ophrys nicaeensis (Barla) Ruppert, Verh. Naturhist. Ver. Preuß. Rheinl. Westf. 83: 302 (1926)
= Ophrys specularia (Rchb.fil.) Lojac., Fl. Sic. 3: 35 (1909)
= Ophrys sphegodes var. arachnitiformis Gren. & Philippe, Mém. Soc. Émul. Doubs, ser. 3, 4: 399 (1859)
subspecies :
subsp. archipelagi (Gölz & H.R.Reinhard) Kreutz; Kompend. Eur. Orchid. 83 (2004)
Distribution: EC-Italy (Adriatic coast); Croatia (Korcula); S-Italy; Albania; NW-Greece (incl. Ionian Isl.); Aegaean Isl. (Kefallenia)
13 ALB CRO GRC ITA 34 EAI
Remarks: sphegodes agg. Pollinator Colletes cunicularius (Paulus 2014)
Synonyms: hidden
= Ophrys archipelagi Gölz & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (4): 731 (1986)
= Ophrys exaltata ssp. archipelagi (Gölz & Reinhard) Del Prete, Del Prete & G. Tosi, Orchidee Spontanee d´Italia: 37 (1988)
= Ophrys arachnitiformis ssp. mateolana (Medagli, D´Emerico, Bianco & Ruggiero) Kreutz, Eurorchis 17: 108 (2005)
= Ophrys exaltata ssp. mateolana (Medagli, D´Emerico, Bianco & Ruggiero) Paulus & Gack, J. Eur. Orchid. 31 (2), 373 (1999)
= Ophrys mateolana Medagli, D´Emerico, Bianco & Ruggiero, L´Orchidophile 97: 109 (1991)
= Ophrys arachnitiformis ssp. cephalonica (B. & H.Baumann) Kreutz, Eurorchis 17: 107 (2005)
= Ophrys archipelagi ssp. cephalonica (B. & H.Baumann) Paulus & M.Hirth, J. Eur. Orch. 46(2): 242 (2014)
= Ophrys cephalonica (B. & H.Baumann) Devillers-Tersch. & Devillers, Nat. Belg. 69 (2): 110 (1988)
= Ophrys exaltata ssp. cephalonica (B. & H.Baumann) Soca, Monde Pl. 97(475): 27 (2002)
= Ophrys sphegodes ssp. cephalonica B. & H.Baumann, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 16 (1): 124 (1984)
= Ophrys ×gilsoniorum P.Delforge, Nat. Belg. 73 (3): 99 (1992)
= Ophrys ×joukoffiorum P.Delforge, Nat. Belg. 73 (3): 101 (1992)
= Ophrys ×poelmansiana P.Delforge, Nat. Belg. 73 (3): 101 (1992)
subspecies :
subsp. tyrrhena (Gölz & H.R.Reinhard) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 718 (2005)
Distribution: W-Italy; WC-Italy
13 ITA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys ×montis-leonis O. & E.Danesch, Orchid. Eur. Ophrys-Hybr. 225. 196 (1972) (pro hybr.)
= Ophrys exaltata ssp. montis-leonis (O.Danesch & E.Danesch) Soca, Monde Pl. 97(475): 27 (2002)
= Ophrys exaltata ssp. tyrrhena (Gölz & H.R.Reinhard) Del Prete, Del Prete & G. Tosi, Orchidee Spontanee d´Italia: 36 (1988)
= Ophrys tyrrhena Gölz & H.R.Reinhard, Pl. Syst. Evol. 136 (1-2): 32 (1980)
subspecies :
subsp. maritima (Romolini & Soca) comb.ined.
Distribution: C-Italy (Liguria, Toscana)
13 ITA
Remarks: sphegodes agg., related to ssp. arachnitiformis and ssp. montis-leonis
Synonyms: hidden
= Ophrys maritima Pacifico & Soca, J. Eur. Orch. 43(4): 770 (-772
= Ophrys sphegodes ssp. maritima (Pacifico & Soca) Kreutz, GIROS Orch. Spont. Eur. 58: 7 (2015)
subspecies :
subsp. cilentana (Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 83 (2004)
Distribution: Italy (Campania); Croatia
13 CRO ITA
Remarks: sphegodes agg., pollinator Andrena florentina
Synonyms: hidden
= Ophrys cilentana Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 351 (2000)
= Serapias cilentana (Presser) P.Delforge, Naturalistes Belges 96(Orchid. 28): 20 (2015), comb. superfl.
= Serapias cilentana (Presser) Presser, J. Eur. Orch. 44(3): 620 (2012)
subspecies :
subsp. gallica (van Looken) Kreutz; Eurorchis 17: 108 (2005)
Distribution: France
12 FRA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys arachnitiformis E.G.Camus, Jour. Bot. Paris 24: 7, 116 (1893) [non Gren. & Philippe]
= Ophrys arachnitiformis ssp. occidentalis Scappat., Orchidophile 152: 132 (2002)
= Ophrys arachnitiformis var. occidentalis (Scappat.) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
= Ophrys aranifera ssp. arachnitiformis E.G.Camus & A.Camus, Iconogr. Orchid. Europe, Text (1): 337 (1928) [non Gren. & Philippe]
= Ophrys aranifera ssp. arachnitiformis E.G.Camus, Bergon & A.Camus, Monogr. Orchid. 287 (1908) [non Gren. & Philippe]
= Ophrys aranifera var. exaltata (Tod.) E.G.Camus, Jour. Bot. Paris 24: 7, 113 (1893)
= Ophrys exaltata Gren., Mem. Soc. Emul. Dept. Doubs ser. 3, 4: 379 (1859) [non Ten.]
= Ophrys gallica Looken, Liparis 6.2 (extra number): 66, 65 (2000)
= Ophrys insectifera ssp. integra Moggr. & Rchb.fil., Nova Acta Acad. Leop.-Carol. Germ. Nat. Cur. 35: 11 (1869)
= Ophrys integra (Moggr. & Rchb.fil.) Paulus & Gack, Israel J. Bot. 39(1-2): 79 (1990) [non Sacc. 1871]
= Ophrys occidentalis (Scappat.) Scappat. & M.Demange, L´Orchidophile (Asnieres) 36(165): 105 (2005)
= Ophrys sphecodes ssp. arachnitiformis Malag., Acta Phytotax. Barcin., 18: 9 (1976 publ. 1977) [non Gren. & Philippe]
= Ophrys sphegodes ssp. arachnitiformis H.Sund., Europ. Medit. Orchid., ed. 3: 39 (1980) [non Gren. & Philippe]
= Ophrys sphegodes ssp. integra (Moggr. & Rchb.fil.) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 380 (1986)
subspecies :
subsp. marzuola (Geniez, Melki & Soca) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 165 (2007)
Distribution: France
12 FRA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys exaltata ssp. marzuola Geniez, Melki & Soca, Monde Pl. 97(475): 27 (2002)
subspecies :
subsp. splendida (Gölz & H.R.Reinhard) H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 718 (2005)
Name: Prächtige Ragwurz
Distribution: SE-France (Arles, to, Cannes)
12 FRA
Remarks: sphegodes agg., pollinator Andrena squalida
Conservation Status: IUCN V
Synonyms: hidden
= Ophrys splendida Gölz & H.R.Reinhard, Pl. Syst. Evol. 136 (1-2): 30 (1980)
= Ophrys exaltata ssp. splendida (Gölz & H.R.Reinhard) Soca, Monde Pl. 97(475): 27 (2002)
species :
Ophrys araneola Rchb.; Pl. Crit. 9: 22 (1831), Fl. Germ. Excurs. 140 (19) (1830)
subspecies :
subsp. araneola
Name: Kleine Spinnen-Ragwurz
Distribution: Croatia; Italy; Spain; France; Switzerland; Germany
11 GER SWI 12 FRA SPA 13 CRO ITA
Remarks: sphegodes agg., pollinator Andrena combinata, A. florivaga, ?A. lathyri
Synonyms: hidden
= Ophrys aranifera ssp. araneola (Rchb.) K.Richt., Pl. Eur. 1: 263 (1890)
= Ophrys aranifera ssp. litigiosa (E.G.Camus) E.G.Camus, Bergon & A.Camus, Monogr. Orchid. 285 (1908)
= Ophrys aranifera var. araneola (Rchb.) Rchb.fil., Icon. Fl. Germ. Helv. 13/14: 89 (1851)
= Ophrys aranifera var. pseudospeculum Rchb.fil., Icon. Fl. Germ. Helv. 13/14: 89 (1851)
= Ophrys litigiosa Camus, Mem. Herb. Boiss. no. 20: 49 (1899)
= Ophrys pseudospeculum var. araneola (Rchb.) Mutel, Fl. France 3: 255 (1836)
= Ophrys sphegodes ssp. araneola (Rchb.) M.Laínz, An. Jard. Bot. Madrid, 40 (1): 279 (1983)
= Ophrys sphegodes ssp. litigiosa (E.G.Camus) Bech., Beitr. Pflanzengeogr. Nordschweiz 46 (1925)
= Ophrys sphegodes ssp. pseudospeculum (Schinz & Thell.) Kelh., Fl. Kant. Schaffhausen: 74 (1920)
= Ophrys sphegodes var. pseudospeculum Schinz & Thell., Fl. Schweiz ed. 3, 2: 72 (1914)
= Ophrys tommasinii ssp. araneola (Rchb.) Soó, Acta Bot. Acad. Sci. Hung. 25 (3-4): 362 (1979 publ. 1980)
= Ophrys tommasinii ssp. litigiosa (Camus) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 383 (1973) [without basionym page]
subspecies :
subsp. argentaria (Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 84 (2004)
Distribution: WC-Italy (Tuscany); ?Croatia
13 ITA
Remarks: sphegodes agg., pollinator Andrena fulvata
Synonyms: hidden
= Ophrys argentaria Devillers-Tersch. & Devillers, Nat. Belg. 72 (3): 99 (1991)
= Ophrys sphegodes var. argentaria (Devillers-Tersch. & Devillers) Faurh., Orchidee (Hamburg) 53(3): 345 (2002)
= Ophrys sphegodes ssp. argentaria (Devillers-Tersch. & Devillers) Biagioli & Grünanger, GIROS Orch. Spont. Eur. 58: 6 (2015)
subspecies :
subsp. ausonia (Devillers, Devillers-Tersch. & P.Delforge) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 167 (2007)
Distribution: Italy
13 ITA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys ausonia Devillers, Devillers-Tersch. & P.Delforge, Naturalistes Belges 85 (Orchid. 17): 148 (2004)
= Ophrys sphegodes ssp. ausonia (Devillers, Devillers-Tersch. & P.Delforge) Biagioli & Grünanger, GIROS Orch. Spont. Eur. 58: 6 (2015)
subspecies :
subsp. minipassionis (Romolini & Soca) comb.ined.
Distribution: C-Italy (Emilia-Romagna, Lazio, Puglia, Toscana, Umbria)
13 ITA
Remarks: sphegodes agg., related to ssp. passionis
Synonyms: hidden
= Ophrys minipassionis Romolini & Soca, J. Eur. Orch. 43(4): 772 (2011)
= Ophrys sphegodes ssp. minipassionis (Romolini & Soca) Biagioli & Grünanger, GIROS Orch. Spont. Eur. 58: 7 (2015)
subspecies :
subsp. gravinensis (D´Alonzo) comb.ined.
Distribution: Italy (Puglia)
13 ITA
Synonyms: hidden
= Ophrys gravinensis D´Alonzo, GIROS Orch. Spont. Eur. 58: 100 (2015)
subspecies :
subsp. cretensis (H.Baumann & Künkele) Kreutz; Kompend. Eur. Orchid. 84 (2004)
Distribution: Crete; Greece (Kyklades)
13 GRC KRI
Remarks: sphegodes agg. Pollinator Andrena vachali
Synonyms: hidden
= Ophrys sphegodes ssp. cretensis H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 375 (1986)
= Ophrys cretensis (Baumann & Künkele) Paulus, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (4): 855 (1988)
subspecies :
subsp. illyrica (S.Hertel & K.Hertel) Kreutz; Kompend. Eur. Orchid. 84 (2004)
Distribution: Croatia (Istria); ?Italy
13 CRO
Remarks: sphegodes agg. Pollinator Andrena pandellei (Paulus 2014)
Synonyms: hidden
= Ophrys illyrica S. & K.Hertel, Jour. Eur. Orch. 34 (3): 512 (2002)
subspecies :
subsp. incantata (Devillers & Devillers-Tersch.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 167 (2007)
Distribution: Croatia (Dalmatia, ?Istria)
13 CRO
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys incantata Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid. 17): 147 (2004)
subspecies :
subsp. quadriloba (Rchb.fil.) Kreutz; Kompend. Eur. Orchid. 84 (2004)
Distribution: N-Spain (La Rioja, Navarra, Ebro basin); ?+France
12 SPA
Remarks: sphegodes agg., pollinator Andrena hypopolia
Synonyms: hidden
= Ophrys aranifera var. quadriloba Rchb.fil., Icon. Fl. Germ. Helv. 13/14: 89 (1851)
= Ophrys quadriloba (Rchb.fil.) E.G.Camus, Monogr. Orch. 295 (1908), pro hybr.
= Ophrys riojana C.E.Hermos., J. Eur. Orchid. 31 (4): 881 (1999)
= Ophrys sphegodes ssp. riojana (C.E.Hermos.) Biagioli & Grünanger, GIROS Orch. Spont. Eur. 58: 7 (2015)
subspecies :
subsp. tommasinii (Vis.) Kreutz; Kompend. Eur. Orchid. 84 (2004)
Distribution: Italy (Abruzzi, Majella, ?S-Italy); Croatia (Istria, Cres, Krk)
13 CRO ITA
Remarks: sphegodes agg. Pollinator Andrena vulpecula (Paulus 2014)
Synonyms: hidden
= Ophrys tommasinii Vis., Fl. Dalm. 3: 354 (1852)
= Ophrys sphegodes ssp. tommasinii (Vis.) Soó, Acta Bot. Acad. Sci. Hung. 16 (3-4): 382 (1970 publ. 1971)
= Ophrys aranifera ssp. tommasinii (Vis.) E.G.Camus, Bergon & A.Camus, Monogr. Orch. 289 (1908) [nom. inval.]
= Ophrys aranifera var. tommasinii Rchb.fil., Icon. Fl. Germ. Helv. 13/14: 178 (1851)
species :
Ophrys argolica H.Fleischm. ex Vierh.; Verh. zool.-bot. Ges. Wien 69: 295 (1919)
subspecies :
subsp. argolica
Distribution: S-Greece (Attiki, Peloponnisos)
13 GRC
Remarks: mammosa agg., argolica subagg., pollinator Anthophora plagiata
Synonyms: hidden
= Ophrys ferrum-equinum ssp. argolica (H.Fleischm.) Soó, Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 907 (1926)
subspecies :
subsp. icariensis (M.Hirth & H.Spaeth) Kreutz; Kompend. Eur. Orchid. 85 (-86) (2004)
Distribution: East Aegaean Isl. (Ikaria, Pserimos); Greece (Kyklades: Naxos, Herakleia)
13 GRC 34 EAI
Remarks: mammosa agg., argolica subagg.
Synonyms: hidden
= Ophrys bornmuelleri ssp. icariensis (M.Hirth & H.Spaeth) Biel & H.Kretzschmar
= Ophrys icariensis M.Hirth & H.Spaeth, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 22 (4): 699 (1990)
subspecies :
subsp. olympiotissa (Paulus) comb.ined.
Distribution: Greece (Tsaritsani in S-Olympos Mts.)
13 GRC
Remarks: mammosa agg., argolica subagg. Pollinator Anthophora plagiata.
Synonyms: hidden
= Ophrys olympiotissa Paulus, J. Eur. Orch. 43(3): 503 (2011)
subspecies :
subsp. atargatis Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 97 (2007)
Distribution: Syria
34 SYR
Remarks: mammosa agg., argolica subagg.
Synonyms: hidden
= Ophrys lesbis f. atargatis (Kreutz) P.Delforge, Naturalistes Belges 89(Orchid. 21): 251 (2008)
= Ophrys atargatis (Kreutz) P.Delforge, Naturalistes Belges 96(Orchid. 28): 16 (2015)
subspecies :
subsp. climacis (Heimeier & Perschke) H.A.Pedersen & P.J.Cribb; Field Guide Orchids Europe Medit. 270 (2019)
Distribution: S-Turkey (Antalya)
34 TUR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys ferrum-equinum ssp. climacis (Heimeier & Perschke) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 721 (2005)
= Ophrys climacis Heimeier & Perschke, J. Eur. Orchid. 30 (1): 205 (1998)
subspecies :
subsp. elegans (Renz) E.Nelson; Monogr. Ophrys 153 (1962)
Distribution: Cyprus
34 CYP
Remarks: mammosa agg., argolica subagg., pollinator Anthophora erschowi
Conservation Status: IUCN V
Synonyms: hidden
= Ophrys elegans (Renz) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13 (3): 350 (1981)
= Ophrys gottfriediana ssp. elegans Renz, Repert. Spec. Nov. Regni Veg. 27: 206 (1929)
subspecies :
subsp. lucis (Kalteisen & H.R.Reinhard) H.A.Pedersen & Faurh.; Orchidee (Hamburg) 53(3): 345 (2002)
Name: Rhodische Hufeisen-Ragwurz (DE); Rhodian Horseshoe Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, Symi, Chalki, Tilos); S-Turkey; NW-Syria
34 EAI SYR TUR
Remarks: mammosa agg., argolica subagg., pollinator Anthophora ?mucida
Synonyms: hidden
= Ophrys ferrum-equinum ssp. lucis (Kalteisen & H.R.Reinhard) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 721 (2005)
= Ophrys lucis (Kalteisen & H.R.Reinhard) Paulus & Gack, Israel J. Bot. 39(1-2): 78 (1990)
= Ophrys aegaea ssp. lucis Kalteisen & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 19 (4): 922 (1987)
subspecies :
subsp. mandalyana (B. & H.Baumann) Kreutz; Orchidee (Hamburg) 57(1): 100 (2006)
Distribution: SW-Turkey
34 TUR
Remarks: mammosa agg., argolica subagg.
Synonyms: hidden
= Ophrys ferrum-equinum ssp. mandalyana B. & H.Baumann, Jour. Eur. Orch. 37(4): 949 (2005)
= Ophrys lesbis f. mandalyana (B. & H.Baumann) P.Delforge, Naturalistes Belges 89(Orchid. 21): 251 (2008)
= Ophrys ferrum-equinum ssp. mandalyana B. & H.Baumann, J. Eur. Orch. 37(4): 949 (2005)
= Ophrys lesbis var. mandalyana (B. & H.Baumann) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
subspecies :
subsp. crabronifera (Sebast. & Mauri) Faurh.; Orchidee (Hamburg) 53(3): 345 (2002)
Distribution: C-Italy (Tuscany, Latium)
13 ITA
Remarks: mammosa agg., argolica subagg.
Synonyms: hidden
= Ophrys crabronifera Sebast. & Mauri, Rom. Pl. Cent. 13, t. 2 (1813)
subspecies :
subsp. morisii (Martelli) Kreutz; Kompend. Eur. Orchid. 86 (2004)
Name: Moris´ Ragwurz
Distribution: Sardinia; Corsica
12 COR SAR
Remarks: mammosa agg., argolica subagg., pollinator Anthophora sicheli
Synonyms: hidden
= Ophrys morisii (Martelli) G.Keller & Soó, Monogr. Iconogr. Orchid. Eur. 2(2-3): 42 (1931)
= Ophrys aranifera var. morisii Martelli, Fl. Italia 1: 232 (1896)
= Ophrys exaltata ssp. morisii (Martelli) Del Prete, Webbia 37: 251 (1984)
= Ophrys morisii (Martelli) Cif. & Giacom., Nomencl. Fl. Ital. 1: 157 (1950)
= Ophrys sphegodes ssp. morisii (Martelli) Del Prete, Webbia 37 (2): 251 (1984)
= Ophrys crabronifera ssp. morisii (Martelli) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 719 (2005)
subspecies :
subsp. biscutella (O.Danesch & E.Danesch) Kreutz; Kompend. Eur. Orchid. 85 (2004)
Distribution: SE-Italy (Apulia, Calabria); Croatia (Dalmatia)
13 CRO ITA
Remarks: mammosa agg., argolica subagg.
Synonyms: hidden
= Ophrys biscutella O. & E.Danesch, Die Orchidee 21: 358 (1970)
= Ophrys crabronifera ssp. biscutella (O.Danesch & E.Danesch) Klaver & Kreutz, GIROS Notiz. 53: 4 (2013)
subspecies :
subsp. pollinensis (O.Danesch & E.Danesch) Kreutz; Kompend. Eur. Orchid. 86 (2004)
Name: Brillen-Ragwurz
Distribution: S-Italy (Campania, Calabria)
13 ITA
Remarks: mammosa agg., argolica subagg., pollinator Anthophora retusa
Synonyms: hidden
= Ophrys crabronifera ssp. pollinensis (Nelson) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 719 (2005)
= Ophrys crabronifera ssp. sundermannii (Soó) Del Prete, Comp. Rend. Coll. S. F. O. 5: 62 (1982)
= Ophrys exaltata ssp. sundermannii Soó, Acta Bot. Acad. Sci. Hung. 16: 392 (1970)
= Ophrys fuciflora ssp. pollinensis Nelson, Monogr. Ophrys 167, 168 (1962)
= Ophrys holosericea ssp. pollinensis (Nelson) Landwehr, Orch. Europa 2: 454 (1977)
= Ophrys holosericea ssp. pollinensis (Nelson) O. & E.Danesch, Pl. Syst. Evol. 124 (2): 130 (1975)
= Ophrys holosericea var. sundermannii (Soó) Landwehr, Orchid. Europa 2: 454 (1977)
= Ophrys pollinensis E.Nelson ex Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 352 (2000)
subspecies :
subsp. virescens (Sommier) comb.ined.
Distribution: S-France; Spain; ?Italy
12 FRA SPA
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys virescens Phil. ex Gren., Mem. Soc. Emul. Doubs ser. 3, 4: 401 (1859)
= Ophrys crabronifera ssp. virescens (Sommier) Klaver & Kreutz, GIROS Notiz. 53: 4 (2013)
= Ophrys araneola ssp. virescens (Philippe ex Gren.) Kreutz, Kompend. Eur. Orchid. 85 (2004)
= Ophrys araneoloides Paulus
= Ophrys aranifera [beta] virescens Gren., Mem. Soc. Emul. Doubs Ser. 3, 4: 396 (1860)
= Ophrys aranifera var. virescens E.G.Camus, Mém. Soc. d´Emul. Dept. Doubs ser. 3, 4: 398 (1859)
species :
Ophrys atlantica Munby; Bull. Soc. Bot. Fr. 3: 108 (1856)
Name: Atlas-Ragwurz
Distribution: S-Spain (Málaga); Tunisia; Algeria; Morocco
12 SPA 20 ALG MOR TUN
Remarks: fusca agg., atlantica subagg., pollinator Chalicodoma parietina
Synonyms: hidden
= Ophrys atlantica ssp. durieui (Rchb.fil.) Maire & Weiller, Fl. Afr. Nord 6: 246 (1959)
= Ophrys fusca ssp. atlantica (Munby) E.G.Camus, Bergon & A.Camus, Monogr. Orchid. 250 (1928)
= Ophrys fusca ssp. durieui (Rchb.fil.) Soó, Fedde, Repert. 24: 26 (1927)
= Ophrys fusca var. durieui Rchb., Icon. Fl. Germ. Helv. 13-14: 74 (1851)
species :
Ophrys aveyronensis (J.J.Wood) P.Delforge; Guide Orchid. Eur. 189 (1984)
subspecies :
subsp. aveyronensis
Distribution: SC-France (Aveyron)
12 FRA
Remarks: sphegodes agg., pollinator Andrena hattorfiana
Conservation Status: IUCN V
Synonyms: hidden
= Ophrys aveyronensis (J.J.Wood) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 329 (1986) [isonym]
= Ophrys aveyronensis (J.J.Wood) P.Delforge & D.Tyteca, Eur. Wildorch. 2:110 (1981) [nom. inval.]
= Ophrys aveyronensis (J.J.Wood) P.Delforge, L´Orchidophile (Asnieres) 61: 582 (1984) [nom. inval.]
= Ophrys sphegodes ssp. aveyronensis J.J.Wood, Orchidee 34 (3): 106 (1983)
subspecies :
subsp. vitorica (Kreutz) Paulus; J. Eur. Orch. 49(3-4): 440 (2017)
Distribution: N-Spain (Navarra, Burgos, La Rioja)
12 SPA
Synonyms: hidden
= Ophrys vitorica Kreutz, Ber. Arbeitskreis. Heimische Orchid. 24(1): 123 (2007)
species :
Ophrys aymoninii (Breistr.) Buttler; Willdenowia 16 (1): 115 (1986)
Name: Aymonins Ragwurz
Distribution: SC-France (Aveyron)
12 FRA
Remarks: insectifera agg., pollinator Andrena combinata. Valid species according to molecular studies
Synonyms: hidden
= Ophrys insectifera ssp. aymoninii Breistr., Bull. Soc. Bot. Fr., Lett. Bot. 128 (1­2): 71 (1981)
species :
Ophrys beirana D.Tyteca, J.Pessoa & Luísa Borges; J. Eur. Orch. 52(2-4): 339 (2020)
Distribution: Portugal
12 POR
Remarks: provisionally accepted name
species :
Ophrys bertolonii Moretti; Dec. 6: 9, in Brugn. Giorn. Fis. 2 (1823)
subspecies :
subsp. bertolonii
Name: Bertolonis Ragwurz
Distribution: Albania; Bosnia & Hercegovina; NW-Greece (Korfu); Malta; Sicily; C-Italy; S-Italy; ?Corsica
13 ALB BIH GRC ITA MLT SIC
Remarks: bertolonii agg., pollinator Chalicodoma parietina, C. pyrenaica, Dasyscolia ciliata
Synonyms: hidden
= Arachnites bertolonii (Moretti) Tod., Orch. Sicul. 79 (1842)
= Ophrys insectifera ssp. bertolonii (Moretti) Moggr. & Rchb.fil., Nova Acta Leop.-Carol. Akad. 35: 11 (1869)
= Ophrys inzengae Nyman, Syll. Suppl. 61 (1865)
= Ophrys scolopax Ten., Fl. Nap. 1: Prod. 53 (1811)
= Ophrys speculum Bertol., Amoen. Ital. 201 (1804) [nec Link (1800)]
= Ophrys bertolonii var. ferrequinoides Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 281 (1986)
subspecies :
subsp. aurelia (P.Delforge, Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 87 (2004)
Distribution: NW-Italy; SE-France; NE-Spain
12 FRA SPA 13 ITA
Remarks: bertolonii agg., pollinator Chalicodoma parietina
Synonyms: hidden
= Ophrys aurelia P.Delforge, Devillers-Tersch. & Devillers, Nat. Belg. 70 (3): 77 (1989)
= Ophrys bertolonii ssp. magniflora (Geniez & Melki) Soca, Monde Pl. 96 (472): 13 (2001)
= Ophrys catalaunica f. magniflora (Geniez & Melki) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
= Ophrys magniflora Geniez & Melki, Actes XIIeme Colloque SFO (Clermont-Ferrand) 59 (1991)
= Ophrys ×pseudobertolonii var. aurelia (P. Delforge, Devillers-Tersch. & Devillers) Hennecke & S.Munzinger, Ber. Arbeitskreis. Heimische Orchid. 31(2): 227 (2015)
subspecies :
subsp. balearica (P.Delforge) L.Sáez & Rosselló; Flora Montiberica 7: 89 (1997)
Distribution: Baleares (incl. Mallorca, Ibiza)
12 BAL
Remarks: bertolonii agg., pollinator Chalicodoma sicula
Synonyms: hidden
= Ophrys balearica P.Delforge, Mém. Soc. Roy. Bot. Belg. 11: 15 (1989 publ. 1990)
= Ophrys bertolonii ssp. balearica (P.Delforge) Soca, Monde Pl. 96 (472): 13 (2001) [isonym]
= Ophrys bertolonii f. triloba Renz, Repert. Spec. Nov. Regni Veg. 28: 260 (1930) [nom. nud.]
= Ophrys bertolonii f. triloba (Renz) Soca, Monde Pl. 96 (472): 13 (2001)
subspecies :
subsp. benacensis (Reisigl) P.Delforge; Bull. Soc. Roy. Bot. Belg. 115 (2): 276 (1982)
Distribution: Italy (Veneto)
13 ITA
Remarks: bertolonii agg., pollinator Chalicodoma parietina
Synonyms: hidden
= Ophrys benacensis (Reisigl) O. & E.Danesch, Pl. Syst. Evol. 124 (2): 108 (1975)
= Ophrys bertoloniiformis ssp. benacensis Reisigl, Orchidee 23 (4): 163 (1972)
= Ophrys saratoi ssp. benacensis (Reisigl) Del Prete, Webbia 37 (2): 252 (1984)
= Ophrys ×pseudobertolonii var. benacensis (Reisigl) Hennecke & S.Munzinger, Ber. Arbeitskreis. Heimische Orchid. 31(2): 227 (2015)
subspecies :
subsp. bertoloniiformis (O. & E.Danesch) H.Sund.; Europ. Medit. Orchid., ed. 3: 39 (1980)
Distribution: S-Italy (Mte. Gargano)
13 ITA
Remarks: bertolonii agg., pollinator Chalicodoma benoisti. Hybridogen, origin Ophrys bertolonii x sphegodes s. lat.
Synonyms: hidden
= Ophrys arachnites var. speculum Durazzo, (1804)
= Ophrys bertoloniiformis O. & E.Danesch, Orchidee 22: 117 (1971)
= Ophrys pseudobertolonii ssp. bertoloniiformis (O. & E.Danesch) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 367 (1986)
subspecies :
subsp. catalaunica (O. & E.Danesch) Soca; Monde Pl. 96 (472): 13 (2001)
Distribution: NE-Spain (Catalunya); SW-France (Aude, Corbières)
12 FRA SPA
Remarks: bertolonii agg.
Synonyms: hidden
= Ophrys bertolonii var. catalaunica (O. & E.Danesch) D.Tyteca & B.D.Tyteca, Bull. Soc. Roy. Bot. Belg. 117 (1): 61 (1984)
= Ophrys pseudobertolonii ssp. catalaunica (O. & E.Danesch) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 368 (1986)
= Ophrys catalaunica O. & E.Danesch, Orchid. Eur. Ophrys-Hybr. 230, 196 (1972)
= Ophrys bertolonii ssp. catalaunica (O.Danesch & E.Danesch) O.Bolòs & Vigo, Fl. Països Catalans 4: 666 (2001) [isonym]
subspecies :
subsp. drumana (P.Delforge) Kreutz; Kompend. Eur. Orchid. 88 (2004)
Distribution: France (Drome)
12 FRA
Remarks: bertolonii agg.
Synonyms: hidden
= Ophrys drumana P.Delforge, Nat. Belg. 69 (5­6): 188 (1988)
subspecies :
subsp. explanata (Lojac.) Soca; Monde Pl. 96 (472): 12 (2001)
Distribution: Sicily
13 SIC
Remarks: bertolonii agg., pollinator Chalicodoma sicula
Synonyms: hidden
= Ophrys explanata (Lojac.) P.Delforge, Mém. Soc. Roy. Bot. Belg. 11: 18 (1989 publ. 1990)
= Ophrys bertolonii var. explanata Lojac., Fl. Sicul. 3: 36 (1908)
= Ophrys bertoloniiformis ssp. explanata (Lojac.) Hennecke & S.Munzinger, Ber. Arbeitskreis. Heimische Orchid. 31(2): 227 (2015)
subspecies :
subsp. flavicans (Vis.) K.Richt.; Pl. Eur. 1: 263 (1890)
Distribution: Croatia (Dalmatia)
13 CRO
Remarks: bertolonii agg., hybridogenous origin Ophrys bertolonii x tommasinii. Pollinator Chalicodoma manicata.
Synonyms: hidden
= Ophrys bertolonii var. dalmatica Murr, Dt. Bot. Monatsschr. 19: 72 (1901)
= Ophrys flavicans Vis., Fl. Dalm. 1: 178 (1842)
= Ophrys dalmatica (Murr) Soó, Notizbl. Bot. Gart. Berlin 9: 902 (1926) [pro hybr.]
subspecies :
subsp. romolinii (Soca) Véla & R.Martin; Étude Cartogr. Prélim. Orchid. Sicile 91 (2015)
Distribution: Sicily
13 SIC
Synonyms: hidden
= Ophrys romolinii Soca, Le monde des Plantes 96 (472): 11 (2001) [nom. nov. superfl.]
subspecies :
subsp. saratoi (E.G.Camus) Soca; Monde Pl. 96 (472): 12 (2001)
Distribution: SE-France (Var, Alpes-Maritimes)
12 FRA
Remarks: bertolonii agg., pollinator Chalicodoma albonotata
Synonyms: hidden
= Ophrys bertolonii var. bilineata Barla, (1868)
= Ophrys gervannensis Dekker, (1988) [nom. nud.]
= Ophrys grassensis Jauvy ex Steud., Nomencl. Bot. ed. 2: 219 (1841) [nom. nud.]
= Ophrys saratoi E.G.Camus, J. Bot. (Morot) 7: 159 (1893) [pro hybr.]
= Ophrys ×araniferiformis Dalla Torre & Sarnth., Fl. Tirol 6: t. 522 (1906)
= Ophrys ×barlae E.G.Camus, J. Bot. (Morot) 7: 159 (1893)
= Ophrys ×bilineata (Barla) Fiori, Fl. Italia 1(1): 235 (1896)
= Ophrys ×disjecta Murr, Deutsch. Bot. Monatsschr. 19: 14, Asch. & Graebn. Syn. Mitteleur. Fl. 3: 646 (1901)
= Ophrys ×sordida Ruppert, (1926)
species :
Ophrys bingoelensis Yild. & Kiliç; Ot Sist. Bot. Dergisi 26(2): 40 (2019)
Distribution: Turkey
34 TUR
Remarks: provisionally accepted name
species :
Ophrys bombyliflora Link; Schrader, Journ. für die Bot. 1799 (2): 325 (1800)
Name: Drohnen-Ragwurz (DE); Bumblebee Orchid (EN); Orquidea abejonaflor (ES)
Distribution: Crete; Greece (except North, incl. Kyklades); Albania; Montenegro; Serbia & Kosovo; Croatia; Malta (incl. Gozo, Comino Isl.); Sicily; Italy; Sardinia; Corsica; Baleares; Spain; Gibraltar; Portugal; France; Tunisia; Algeria; Morocco; Libya (Cyrenaica); East Aegaean Isl. (including Rhodos, Karpathos, Tilos); W-Turkey; Canary Isl. (Tenerife, Gran Canaria)
12 BAL COR FRA GIB POR SAR SPA 13 ALB CRO GRC ITA KRI MLT MNT SIC SRB 20 ALG LBY MOR TUN 21 CNY 34 EAI TUR
Remarks: tenthredinifera agg. Pollinator Eucera nigrescens
Synonyms: hidden
= Arachnites bombyliflora (Link) Tod., Orch. Sicul. 91 (1842)
= Ophrys bombylifera Willd., Sp. Pl. 4: 68 (1805)
= Ophrys bombyliflora f. buccheriana De Langhe & R.D´Hose, Bull. Soc. Roy. Bot. Belg. 118 (1): 70 (1985)
= Ophrys bombyliflora f. parviflora Mifsud, Jour. Eur. Orch. 41(3/4), 616 (2009)
= Ophrys bombyliflora ssp. canariensis G.Kunkel, Kanarischen Ins. Pflanzenwelt: 158 (1980), nom. inval.
= Ophrys bombyliflora var. albarranii P.P.Ferrer, E.Laguna & Gómez-Montbl., Flora Montiber. 61: 4 (2015)
= Ophrys bombyliflora var. parviflora (Mifsud) Mifsud, J. Eur. Orch. 46(3/4): 684 (2014)
= Ophrys canaliculata Viv., App. Fl. Cors. Prod. 7 (1825)
= Ophrys disthoma Biv., Sic. pl. Cent. 1: 59 (1806)
= Ophrys hiulca Sebast. & Mauri, Cent. Rom. Pl. 13: 43 (1820)
= Ophrys labrofossa Brot., Phyt. Lusit. 2: 29, t. 88 (1827)
= Ophrys picta Salzm. ex Ball, Journ. Linn. Soc. 16: 673 (1878)
= Ophrys picta ssp. nafarroana Robles & M.Becerra, Acta Bot. Malac. 46e10589: [4] (2019)
= Ophrys pulla Ten., Fl. Nap. Prod. 1: p. 53 (1811)
= Ophrys tabanifera Willd., Sp. Pl. 4: 68 (1805)
species :
Ophrys bornmuelleri M.Schultze; Mitt. Thüring. Bot. Ver. n. s. 13-14: 127 (1899)
subspecies :
subsp. bornmuelleri
Distribution: Israel; Syria; Lebanon; SE-Turkey; N-Iraq
34 IRQ ISR LEB SYR TUR
Remarks: holosericea agg., bornmuelleri-group, pollinator Eucera penicillata (syn. paulusi)
Synonyms: hidden
= Ophrys fuciflora ssp. bornmuelleri (M.Schultze) B. & E.Willing, Orchidee 26 (2): 78 (1975)
= Ophrys holosericea ssp. bornmuelleri (M.Schultze) H.Sund., Taxon 24 (5-6): 625 (1975)
subspecies :
subsp. aphrodite (Devillers & Devillers-Tersch.) comb.ined.
Distribution: Cyprus
34 CYP
Synonyms: hidden
= Ophrys aphrodite Devillers & Devillers-Tersch., Naturalistes Belges 93(Orchid. 25): 160 (2012)
subspecies :
subsp. carduchorum Renz & Taubenheim; Orchidee 31 (6): 242 (1980)
Distribution: SE-Turkey
34 TUR
Remarks: holosericea agg., bornmuelleri-group
Conservation Status: IUCN E
Synonyms: hidden
= Ophrys carduchorum (Renz & Taubenheim) P.Delforge, Nat. Belg. 71 (3): 127 (1990)
subspecies :
subsp. ziyaretiana (Kreutz & Ruedi Peter) Kreutz; Kompend. Eur. Orchid. 89 (2004)
Distribution: S-Turkey (Hatay); NW-Syria; Lebanon; N-Israel
34 ISR LEB SYR TUR
Remarks: holosericea agg., bornmuelleri-group
Synonyms: hidden
= Ophrys bornmuelleri var. ziyaretiana (Kreutz & Ruedi Peter) P.Delforge, Naturalistes Belges 85 (Orchid. 17): 251 (2004)
= Ophrys ziyaretiana Kreutz & Ruedi Peter, J. Eur. Orchid. 30 (1): 104 (1997)
= Ophrys pseudolevantina M. & H.Schönfelder, Ber. Arbeitskr. Heim. Orch. 18(2): 71 (2001) [nom. prov.]
= Ophrys fuciflora var. ziyaretiana (Kreutz & Ruedi Peter) Faurh. & H.A.Pedersen, Journ. Eur. Orch. 40(4): 695 (2008)
species :
Ophrys candica (E.Nelson ex Soó) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13 (3): 349 (1981)
subspecies :
subsp. candica
Name: Weißglanz-Ragwurz (DE); Enamel Bee Orchid (EN)
Distribution: Crete; Greece (very rare: S-Peloponnisos, Kythera Isl.); East Aegaean Isl. (Rhodos)
13 GRC KRI 34 EAI
Remarks: holosericea agg., bornmuelleri-group, pollinator ?Eucera ephippia
Synonyms: hidden
= Ophrys candica (E.Nelson ex Soó) Greuter, Matthäs & Risse, Willdenowia 15 (1): 53 (1985)
= Ophrys candica f. minoa (C. & A.Alibertis) P.Delforge, Naturalistes Belges 87(Orchid. 19): 261 (2006)
= Ophrys candica ssp. cytherea B. & H.Baumann, Jour. Eur. Orch. 37(3): 718 (2005)
= Ophrys candica ssp. minoa C. & A.Alibertis, L´Orchidophile 87: 111 (1989) [fide Kretzschmar & Eccarius 2001]
= Ophrys candica var. minoa (C. & A.Alibertis) Paulus, Gügel, D. & U.Rückbr., Ber. Arbeitskr. Heim. Orch. 18 (1): 21 (2001) [fide Kretzschmar & Eccarius 2001]
= Ophrys fuciflora ssp. candica E.Nelson, Monogr. Ophrys 165 (1962) [nom. inval.]
= Ophrys fuciflora ssp. candica E.Nelson ex Soó, Bot. Journ. Linn. Soc. 76 (4): 368 (1978)
= Ophrys fuciflora var. candica (E.Nelson ex Soó) H.Sund., Europ. Medit. Orchideen 45 (1975)
= Ophrys graeca (B. & H.Baumann) S.Hertel & H.Weyland, Ber. Arbeitskreis. Heimische Orchid. 34(1): 182 (2017)
= Ophrys holosericea ssp. candica (E.Nelson ex Soó) Renz & Taubenheim, Notes Roy. Bot. Gard. Edinb. 41 (2): 276 (1983)
= Ophrys holosericea ssp. candica (Greuter, Matthäs & Risse) Pedersen & Faurh., Jour. Eur. Orch. 37(2): 288 (2005)
= Ophrys holosericea ssp. cerigona B. & H.Baumann, J. Eur. Orch. 39(1): 115 (2007)
= Ophrys holosericea ssp. graeca B. & H.Baumann, Jour. Eur. Orch. 37(3): 723 (2005)
= Ophrys holosericea var. candica (Nelson) H.Sund., Taxon 24 (5-6): 625 (1975)
= Ophrys lacaena f. cytherea (B. & H.Baumann) P.Delforge, Naturalistes Belges 87(Orchid. 19): 261 (2006)
= Ophrys lacaena f. graeca (B. & H.Baumann) P.Delforge, Naturalistes Belges 87(Orchid. 19): 261 (2006)
= Ophrys minoa (C. & A.Alibertis) P.Delforge, Nat. Belg. 72 (3): 101 (1991) [fide Kretzschmar & Eccarius 2001]
subspecies :
subsp. calliantha (Bartolo & Pulv.) Kreutz; Kompend. Eur. Orchid. 89 (2004)
Distribution: Sicily
13 SIC
Remarks: holosericea agg., bornmuelleri-group, hybridogenous origin Ophrys candica x oxyrrhynchos
Synonyms: hidden
= Ophrys oxyrrhynchos ssp. calliantha (Bartolo & Pulv.) Galesi, Cristaudo & Maugeri, J. Eur. Orchideen 36(2): 506 (2004)
= Ophrys calliantha Bartolo & Pulv., Caesiana 9: 41 (1997)
subspecies :
subsp. lacaena (P.Delforge) Kreutz; Kompend. Eur. Orchid. 89 (2004)
Distribution: S-Greece (S-Peloponnisos)
13 GRC
Remarks: holosericea agg., "bornmuelleri-like"-group
Synonyms: hidden
= Ophrys holosericea ssp. lacaena (P.Delforge) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 723 (2005)
= Ophrys lacaena P.Delforge, Naturalistes Belges 85 (Orchid. 17): 239 (2004)
species :
Ophrys ciliata Biv.; SIc. pl. Cent. 1: 60 (1806)
subspecies :
subsp. ciliata
Name: Spiegel-Ragwurz (DE); Mirror Bee Orchid (EN)
Distribution: Greece; European Turkey; Turkey; East Aegaean Isl. (Rhodos, Tilos); Albania; Malta (incl. Gozo, Comino Isl.); Sicily; Italy; Sardinia; Corsica; Baleares; Spain; Gibraltar; Portugal; Croatia; France; Tunisia; Algeria; Morocco; Libya (Cyrenaica)
12 BAL COR FRA GIB POR SAR SPA 13 ALB CRO GRC ITA MLT SIC TUE 20 ALG LBY MOR TUN 34 EAI TUR
Remarks: ciliata agg., pollinator Dasiscolia ciliata ssp. ciliata
Synonyms: hidden
= Arachnites speculum (Link) Tod., Orch. Sicul. 93 (1842)
= Ophrys scolopax Willd., Sp. Pl. 4: 69 (1805)
= Ophrys speculum Link, Schrader, Journ. für die Bot. 1799 (2): 324 (1800)
= Ophrys vernixia ssp. ciliata (Biv.) Kerguélen, Lejeunia 120: 135 (1987), isonym
= Ophrys vernixia ssp. ciliata (Biv.) Del Prete, Webbia 37 (2): 252 (1984)
subspecies :
subsp. orientalis (Paulus) Kreutz; Orchidee (Hamburg) 57(1): 101 (2006)
Name: Östliche Spiegel-Ragwurz (DE); Eastern Mirror Bee Orchid (EN)
Distribution: Greece; Crete [I]; Turkey
13 GRC kri 34 TUR
Remarks: ciliata agg., pollinator Dasiscolia ciliata ssp. araratensis
Synonyms: hidden
= Ophrys ciliata var. orientalis (Paulus) Kreutz, Kompend. Eur. Orchid. 91 (2004)
= Ophrys vernixia ssp. orientalis Paulus, Ber. AHO 18 (1): 43 (2001)
= Ophrys speculum ssp. orientalis (Paulus) Paulus & Salk., Ber. Arbeitskreis. Heimische Orchid. 24(2): 7 (2007)
= Ophrys speculum var. orientalis (Paulus) Kreutz, Ber. Arbeitskreis. Heimische Orchid. 24(1): 178 (2007)
= Ophrys eos Devillers & Devillers-Tersch., Naturalistes Belges 90(Orchid.22): 289 (2009), nom. nov. for O. ciliata ssp. orientalis
subspecies :
subsp. lusitanica (O.Danesch & E.Danesch) H.Baumann, Künkele & R.Lorenz; J. Eur. Orchideen 36(2): 571 (2004)
Name: Iberische Spiegel-Ragwurz (DE); Iberian Mirror Bee Orchid (EN)
Distribution: S-Spain; Portugal
12 POR SPA
Remarks: ciliata agg.
Synonyms: hidden
= Arachnites speculum Tod., (1842)
= Ophrys speculum ssp. lusitanica O. & E.Danesch, Orchidee 20: 21 (1969)
= Ophrys vernixia ssp. lusitanica (O. & E.Danesch) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 391 (1986)
= Ophrys lusitanica (O.Danesch & E.Danesch) Paulus & Gack, Israel J. Bot. 39(1-2): 77 (1990)
= Ophrys vernixia Brot., Fl. Lusit. 1: 24 (1800 publ. 1804) [nom. illeg.]
= Ophrys vernixia ssp. occidentalis Paulus, Ber. AHO 18 (1) (2001)
species :
Ophrys cilicica Schltr.; Fedde, Repert. 19: 45 (1923)
Name: Cilicische Ragwurz
Distribution: S-Turkey; NW-Syria; Iran; N-Iraq
34 IRN IRQ SYR TUR
Remarks: reinholdii agg.
Synonyms: hidden
= Ophrys aranifera ssp. boissieri (Soó) Soó, Repert. Spec. Nov. Regni Veg. 26: 279 (1929)
= Ophrys aranifera ssp. mammosa var.boissieri Soó, Repert. Spec. Nov. Regni Veg. 24: 35 (1927)
= Ophrys kurdica D. & U.Rückbr., Orchidee 26 (4): 164 (1975)
= Ophrys kurdica ssp. kurdistanica (Renz) Soó, Acta Bot. Acad. Sci. Hung. 25 (3-4): 362 (1979 publ. 1980)
= Ophrys kurdistanica Renz, Fl. Iranica 126: 83 (1978)
= Ophrys mammosa ssp. boissieri (Soó) Soó, Acta Bot. Acad. Sci. Hung. 18: 383 (1973)
= Ophrys sphegodes ssp. boissieri (Soó) Soó, Acta Bot. Acad. Sci. Hung. 16(3/4): 390 (1970 publ. 1971)
species :
Ophrys cretica (Vierh.) E.Nelson; Gestalt. & Arth. Orch. Eur. & Mittelmeerl. 146 (1962)
subspecies :
subsp. cretica
Name: Kretische Ragwurz
Distribution: Crete; Greece (S-Peloponnisos, Kyklades)
13 GRC KRI
Remarks: reinholdii agg., pollinator Melecta tuberculata
Synonyms: hidden
= Ophrys cretica f. albovirescens E.Klein, L´Orchidophile 34: 1136 (1978) [without latin descr. or type]
= Ophrys cretica var. albovirescens E.Klein, Orchidee 29 (1): 29 (1978) [without type]
= Ophrys kotschyi ssp. cretica (Vierh.) H.Sund., Europ. Medit. Orchideen 45 (1975)
= Ophrys spruneri ssp. cretica (Vierh.) Renz, Repert. Spec. Nov. Regni Veg. 30: 109 (1932)
= Ophrys spruneri var. cretica (Vierh.) Vierh. ex Soó, Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 907 (1926)
= Ophrys spruneri f. cretica Vierh., Österr. Bot. Zeitschr. 66: 164 (1916)
subspecies :
subsp. ariadnae (Paulus) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 130 (2002)
Distribution: Crete; S-Greece (S-Peloponnisos, Attiki, Kythira, Kyklades); East Aegaean Isl. (Karpathos, ?Kasos, Paros, Naxos)
13 GRC KRI 34 EAI
Remarks: reinholdii agg., pollinator Melecta albifrons
Synonyms: hidden
= Ophrys kotschyi ssp. ariadnae (Paulus) Faurh., Orchidee (Hamburg) 53(3): 345 (2002)
= Ophrys ariadnae Paulus, J. Eur. Orchid. 26 (3­4): 635 (1994)
= Ophrys cretica ssp. naxia Nelson, Monogr. Ophrys 146 (1962) [nom. inval.]
= Ophrys karpathensis (Nelson) Paulus & Gack, [nom. inval.]
= Ophrys cretica ssp. karpathensis Nelson, Monogr. Ophrys 146 (1962) [nom. inval.]
subspecies :
subsp. beloniae G. & H.Kretzschmar; Ber. Arbeitskr. Heim. Orch. 13 (1): 10 (1996)
Name: Naxos-Ragwurz (DE); Naxos Bee Orchid (EN)
Distribution: East Aegaean Isl. (Naxos, S-Rhodos); ?Greece (Kyklades: Paros)
34 EAI
Remarks: reinholdii agg.
Synonyms: hidden
= Ophrys beloniae (G. & H.Kretzschmar) Devillers & Devillers-Tersch., Naturalistes Belges 90(Orchid.22): 290 (2009)
= Ophrys cretica f. beloniae (G. & H.Kretzschmar) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
subspecies :
subsp. bicornuta G.Kretzschmar & H.Jahn; Ber. Arbeitskr. Heim. Orch. 18 (2) (2001)
Distribution: E-Crete
13 KRI
Remarks: reinholdii agg.
Synonyms: hidden
= Ophrys cretica f. bicornuta (H.Kretzschmar & R.L.Jahn) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
species :
Ophrys delphinensis O. & E.Danesch; Orchid. Eur. Ophrys-Hybr. 222, 54 (1972), pro hybr.
Name: Delphi-Ragwurz
Distribution: C-Greece (Delphi, Patras etc.); S-Greece (Peloponnisos)
13 GRC
Remarks: mammosa agg., argolica subagg., pollinator Anthophora plagiata. Hybridogenous origin O. argolica x cornuta
species :
Ophrys dimidiata Rakosy, Paulus & M.Hirth; J. Eur. Orch. 52(1): 212 (2020)
Distribution: Greece
13 GRC
Remarks: provisionally accepted name
species :
Ophrys eretriae M.Hirth & Paulus; J. Eur. Orch. 52(1): 133 (2020)
Distribution: Greece
13 GRC
Remarks: provisionally accepted name
species :
Ophrys exaltata Ten.; Cat. Pl. Hort. Neap. (1819) App. 2: 83
subspecies :
subsp. exaltata
Name: Hornissen-Ragwurz, Hohe Ragwurz
Distribution: S-Italy; Sicily
13 ITA SIC
Remarks: sphegodes agg., pollinator Colletes cunicularius. O. exaltata and O. arachnitiformis might be two extreme forms of a widespread and variable species, which would then be called O. exaltata.
Synonyms: hidden
= Ophrys fuciflora ssp. exaltata (Ten.) E.Nelson, Gestaltwandel 169 (1962)
= Ophrys aranifera ssp. exaltata (Ten.) E.G.Camus, Bergon & A.Camus, Monogr. Orchid. 288 (1908) nom. illeg.
= Ophrys exaltata ssp. sicula (E.Nelson ex Soó) Del Prete, Webbia 37 (2): 251 (1984)
= Ophrys sphecodes ssp. sicula E.Nelson, Monogr. Ophrys 193 (1962) [nom. inval.]
= Ophrys sphecodes ssp. sicula E.Nelson ex Soó, Bot. Journ. Linn. Soc. 76 (4): 367 (1978)
= Ophrys trinacrica Del Prete, Comp. Rend. Coll. S. F. O. 5: 61 (1982)
= Ophrys sphegodes var. sicula (E.Nelson ex Soó) Galesi, Cristaudo & Maugeri, J. Eur. Orchideen 36(2): 505 (2004)
= Ophrys holosericea ssp. exaltata (Ten.) H.Sund., Taxon 24 (5-6): 625 (1975)
= Ophrys sipontensis R.Lorenz & Gembardt, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 19 (3): 472 (1987), nom. illeg., non Gumpr.
subspecies :
subsp. castellana (Devillers-Tersch. & Devillers) Soca; Monde Pl. 97(475): 27 (2002)
Distribution: C-Spain (Aragon); N-Spain (Navarra, etc.)
12 SPA
Remarks: sphegodes agg., pollinator probably Andrena (Opandrena) schencki
Synonyms: hidden
= Ophrys castellana Devillers-Tersch. & Devillers, Nat. Belg. 69 (2): 108 (1988)
species :
Ophrys ferrum-equinum Desf.; Ann. Mus. Hist. Nat. (Paris) 10: 226 (1807)
subspecies :
subsp. ferrum-equinum
Name: Hufeisen-Ragwurz (DE); Horseshoe Orchid (EN)
Distribution: Albania; Greece (except Northeast, incl. Kyklades); East Aegaean Isl. (Samos, Naxos, Rhodos, Karpathos, Chalki, Kastellorizo, Pserimos, Tilos, etc.); S-Turkey
13 ALB GRC 34 EAI TUR
Remarks: mammosa agg., pollinator Chalicodoma parietina
Synonyms: hidden
= Ophrys andrachnites Bory & Chaub., Nouv. Fl. Pelop. 62, t. 34 (1838)
= Ophrys ferrum-equinum f. anafiensis (Biel) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
= Ophrys ferrum-equinum f. convexa (B. & H.Baumann) P.Delforge, Naturalistes Belges 87(Orchid. 19): 261 (2006)
= Ophrys ferrum-equinum f. labiosa (Kreutz) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
= Ophrys ferrum-equinum f. minor (Biel) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
= Ophrys ferrum-equinum ssp. convexa B. & H.Baumann, Jour. Eur. Orch. 37(4): 948 (2005)
= Ophrys ferrum-equinum ssp. labiosa (Kreutz) Kreutz, Kompend. Eur. Orchid. 92 (2004)
= Ophrys ferrum-equinum var. anafiensis Biel, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 98 (2001)
= Ophrys ferrum-equinum var. labiosa (Kreutz) Hennecke, Ber. Arbeitskreis. Heimische Orchid. 31(2): 241 (2015)
= Ophrys ferrum-equinum var. minor Biel, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 98 (2001)
= Ophrys ferrum-equinum var. pseudogottfriediana Paulus, J. Eur. Orch. 46(2): 246 (2014)
= Ophrys labiosa Kreutz, J. Eur. Orchid. 29 (4): 683 (1997)
subspecies :
subsp. aegaea (Kalteisen & H.R.Reinhard) H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 720 (2005)
Distribution: East Aegaean Isl. (Karpathos); Greece (Kyklades: Kasos, Amorgos, Iraklia Isl.)
13 GRC 34 EAI
Remarks: mammosa agg., pollinator Anthophora orientalis
Synonyms: hidden
= Ophrys argolica ssp. aegaea (Kalteisen & H.R.Reinhard) H.A.Pedersen & Faurh., Orchidee (Hamburg) 53(3): 345 (2002)
= Ophrys aegaea Kalteisen & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 19 (4): 918 (1987)
subspecies :
subsp. lycia (Renz & Taubenheim) comb.ined.
Distribution: SW-Turkey (Antalya)
34 TUR
Remarks: mammosa agg., argolica subagg.
Conservation Status: IUCN E
Synonyms: hidden
= Ophrys lycia Renz & Taubenheim, Orchidee 31 (6): 237 (1980)
= Ophrys sphegodes ssp. lycia (Renz & Taubenheim) H.A.Pedersen, P.J.Cribb & Rolf Kühn, Field Guide Orchids Europe Medit. 255 (2019)
subspecies :
subsp. gottfriediana (Renz) E.Nelson; Gestaltwandel 141 (1962)
Distribution: W-Greece (Kefallonia, Ithaca, Zante Isl.)
13 GRC
Remarks: mammosa agg., pollinator Chalicodoma parietina
Conservation Status: IUCN R
Synonyms: hidden
= Ophrys gottfriediana Renz, Fedde, Repert. 25: 255 (1928)
= Ophrys ferrum-equinum var. gottfriediana (Renz) Biel, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 87-127 (2001)
= Ophrys spruneri ssp. gottfriediana (Renz) Soó, Repert. Spec. Nov. Regni Veg. 26: 279 (1929)
= Ophrys ferrum-equinum f. parnassica (Vierh.) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
= Ophrys ferrum-equinum f. subtriloba Hayek
subspecies :
subsp. lesbis (Gölz & H.R.Reinhard) H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 721 (2005)
Distribution: East Aegaean Isl. (Lesvos, Samos); SW-Turkey
34 EAI TUR
Remarks: mammosa agg., pollinator Andrena curiosa
Synonyms: hidden
= Ophrys argolica ssp. lesbis (Gölz & H.R.Reinhard) H.A.Pedersen & Faurh., Orchidee (Hamburg) 53(3): 345 (2002)
= Ophrys lesbis Gölz & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 21 (1): 19 (1989)
= Ophrys aegaea ssp. lesbis (Gölz & H.R.Reinhard) Biel & H.Kretzschmar
species :
Ophrys fusca Link; Schrader, Journ. für die Bot. 1799 (2): 324 (1800) s. str.
subspecies :
subsp. fusca
Name: Braune Ragwurz
Distribution: Spain; Gibraltar; Baleares; France; Corsica; ?Italy; ?Sardinia; Tunisia; Algeria; ?Morocco
12 BAL COR FRA GIB SPA 20 ALG TUN
Remarks: fusca agg., fusca s. str., pollinator Andrena (Melandrena) nigroaenea
Synonyms: hidden
= Arachnites fusca (Link) Tod., W. Schmidt, Fl. Boem. 1: 98 (1793)
= Ophrys fusca var. maculata Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 281 (1986)
= Ophrys fusca var. rubescens Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 281 (1986)
= Ophrys laetea Willk. & Lange, Prod. Fl. Hisp. 1: 174 (1861)
= Ophrys lutea Biv., Cent. 2: 41, t. 5 (1807) [nom. illeg.]
= Ophrys myodes Lapeyr., Hist. Abr. Pl. Pyren. 2: 551 (1813)
= Ophrys tricolor Desf. ex Nyman, Consp. 699 (1882)
= Ophrys fusca var. distincta F.M.Vázquez, Folia Bot. Extremadur. 3: 253 (2009)
= Ophrys fusca var. lutea F.M.Vázquez, Folia Bot. Extremadur. 3: 257 (2009)
= Ophrys fusca var. lutescens F.M.Vázquez, Folia Bot. Extremadur. 3: 249 (2009)
subspecies :
subsp. clara (F.M.Vázquez & S.Ramos) F.M.Vázquez; J. Eur. Orch. 40(4): 710 (2008)
Distribution: S-Spain
12 SPA
Remarks: fusca agg., probably fusca s. str.
Synonyms: hidden
= Ophrys clara Vázquez & Ramos, Jour. Eur. Orch. 37(4): 818 (2005)
subspecies :
subsp. limensis F.M.Vázquez; Folia Bot. Extremadur. 3: 269 (2009)
Distribution: Spain
12 SPA
Remarks: fusca agg., probably fusca s. str.
subspecies :
subsp. delforgei (Devillers-Tersch. & Devillers) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 169 (2007)
Distribution: S-France (Provence, Bouches-du-Rhone)
12 FRA
Remarks: fusca agg., fusca s. str.
Synonyms: hidden
= Ophrys delforgei Devillers-Tersch. & Devillers, Naturalistes Belges 87(Orchid. 19): 62 (2006)
= Ophrys forestieri auct., non (Rchb. fil.) Lojac.
subspecies :
subsp. maghrebiaca Kreutz, Rebbas, Babali, Miara & Ait-Hamm.; Ber. Arbeitskreis. Heimische Orchid. 30(2): 195 (2013)
Distribution: Algeria
20 ALG
Synonyms: hidden
= Ophrys maghrebiaca (Kreutz, Rebbas, Babali, Miara & Ait-Hamm.) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
subspecies :
subsp. lucifera (Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 95 (2004)
Distribution: Italy; Sicily; Malta
13 ITA MLT SIC
Remarks: fusca agg., fusca s. str.
Synonyms: hidden
= Ophrys lucifera Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 322 (2000)
subspecies :
subsp. creticola (Paulus) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 182 (2002)
Distribution: Crete
13 KRI
Remarks: fusca agg., fusca s. str., early flowering taxon
Synonyms: hidden
= Ophrys creticola Paulus, J. Eur. Orchid. 30 (1): 177 (1998)
subspecies :
subsp. cressa (Paulus) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 178 (2002)
Distribution: C-Crete; E-Crete
13 KRI
Remarks: fusca agg., fusca s. str., pollinator Andrena merula. Status and separation of ssp. cressa and ssp. creberrima highly disputed, as well as further splitting by Paulus (2007)
Synonyms: hidden
= Ophrys cressa Paulus, J. Eur. Orchid. 30 (1): 176 (1998)
= Ophrys phaidra Paulus, Jahresber. Naturw. Ver. Wuppertal 60: 101-151 (2007)
= Ophrys kedra Paulus, Jahresber. Naturw. Ver. Wuppertal 60: 101-151 (2007)
= Ophrys pallidula Paulus, Jahresber. Naturwiss. Vereins Wuppertal 60: 101-151 (2007)
subspecies :
subsp. peraiolae (G.Foelsche, W.Foelsche, M.Gerbaud & O.Gerbaud) Kreutz; Kompend. Eur. Orchid. 95 (2004)
Distribution: NE-Corsica
12 COR
Remarks: fusca agg., fusca s. str.
Synonyms: hidden
= Ophrys peraiolae G. & W.Foelsche, M. & O.Gerbaud, J. Eur. Orchid. 32 (3/4): 403-455 (2000)
= Ophrys peraiolae var. rubra G. & W.Foelsche, M. & O.Gerbaud, J. Eur. Orchid. 32 (3/4): 403-455 (2000)
subspecies :
subsp. forestieri (Rchb.fil.) Kreutz; Kompend. Eur. Orchid. 94 (2004)
Distribution: SW-France (Gers, Hautes-Pyrenees); N-Spain; E-Spain; Italy; Baleares; Sicily; NW-Greece (Korfu); S-Greece; ?Albania; ?Montenegro; Corsica; Portugal; Croatia; etc.
12 BAL COR FRA POR SPA 13 CRO GRC ITA SIC
Remarks: fusca agg., fusca s. str., pollinator Andrena nigroaenea. Taxonomy see Paulus 2014
Synonyms: hidden
= Ophrys fusca ssp. lupercalis (Devillers-Tersch. & Devillers) Kreutz, Orchidee (Hamburg) 57(1): 101 (2006)
= Ophrys lupercalis Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 373 (1994)
= Ophrys forestieri (Rchb.fil.) Lojac., Fl. Sic. 3: 44 (1909)
= Ophrys fusca var. forestieri Rchb.fil., Icon. Fl. Germ. Helv. 13-14: t. 464, f. 12 (1851)
= Ophrys intermedia Tineo ex Lojac., Fl. Sic. 3: 45 (1909)
= Ophrys gackiae P.Delforge, Naturalistes Belges 85 (Orchid. 17): 118 (2004) (fide Paulus 2014)
= Ophrys fusca ssp. gackiae (P.Delforge) Kreutz, GIROS Orch. Spont. Eur. 58: 6 (2015)
= Ophrys decembris S.Moingeon & J.-M.Moingeon, J. Eur. Orch. 43(1): 75 (2011) (fide Paulus 2014)
= Ophrys fusca ssp. laureotica Kalog., Delipetrou & A.Alibertis, J. Eur. Orch. 43(2): 369 (2011) (fide Paulus 2014)
= Ophrys forestieri ssp. laureotica (Kalog., Delipetrou & A.Alibertis) Paulus, J. Eur. Orch. 52(2-4): 368 (2020)
subspecies :
subsp. malacitana (M.R.Lowe, I.Phillips & Paulus) comb.ined.
Distribution: S-Spain (Malaga, Granada, W-Almeria)
12 SPA
Remarks: fusca agg., fusca s. str., pollinator Colletes cunicularius
Synonyms: hidden
= Ophrys malacitana M.R.Lowe, I.Phillips & Paulus, J. Eur. Orch. 42(3-4): 547 (2010)
subspecies :
subsp. fabrella (Paulus & Ayasse ex P.Delforge) Kreutz; Kompend. Eur. Orchid. 94 (2004)
Distribution: Baleares (incl. Mallorca, Ibiza)
12 BAL
Remarks: fusca agg., fusca s. str., pollinator Andrena fabrella
Synonyms: hidden
= Ophrys fabrella Paulus & Ayasse ex P.Delforge, Naturalistes Belges 85 (Orchid. 17): 119 (-120) (2004)
subspecies :
subsp. vasconica (O.Danesch & E.Danesch) Kreutz; Kompend. Eur. Orchid. 111 (2004)
Distribution: S-France; Spain; ?Portugal
12 FRA SPA
Remarks: fusca agg., fusca s. str., pollinators Andrena nigroaenea, Andrena flavipes
Synonyms: hidden
= Ophrys vasconica (O. & E.Danesch) P.Delforge, Nat. Belg. 72 (3): 101 (1991)
= Ophrys fusca ssp. vasconica O. & E.Danesch, Die Orchidee 20(6): 258 (1969)
= Ophrys ×brigittae H.Baumann, Beih. Veröff. Naturschutz Landschaftspflege Baden-Württemberg 19: 142 (1981)
subspecies :
subsp. minima Balayer; Bull. Soc. Bot. France, Lett. Bot. 133: 281 (1986)
Distribution: N-Italy; N-Spain; S-France; Croatia
12 FRA SPA 13 CRO ITA
Remarks: fusca agg., fusca s. str., pollinator Andrena wilkella, A. flavipes, A. ornata
Synonyms: hidden
= Ophrys sulcata Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 375 (1994)
subspecies :
subsp. lucana (P.Delforge, Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 94 (2004)
Distribution: S-Italy
13 ITA
Remarks: fusca agg., fusca s. str., pollinator Andrena labialis
Synonyms: hidden
= Ophrys lucana P.Delforge, Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 264 (2000)
subspecies :
subsp. marmorata (G.Foelsche & W.Foelsche) Kreutz; Kompend. Eur. Orchid. 95 (2004)
Distribution: S-Corsica; ?Croatia; Algeria
12 COR 20 ALG
Remarks: fusca agg., fusca s. str., pollinator Andrena wilkella
Synonyms: hidden
= Ophrys marmorata G. & W.Foelsche, L´Orchidophile (133) 177-178 (1998) [fide Paulus 2001]
subspecies :
subsp. obaesa (Lojac.) E.G.Camus & A.Camus; Iconogr. orchid. Europe, Text(1): 293 (1928)
Distribution: Sicily
13 SIC
Remarks: fusca agg., bilunulata-group, pollinator Andrena (Zonandrena) flavipes
Synonyms: hidden
= Ophrys obaesa Lojac., Fl. Sic. 3: 46 (1909)
= Ophrys ×ficuzzana H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 468 (1986)
= Ophrys caerulea Geniez & Melki, Actes Coll. S. F. O. 12: 51 (1991) [nom. nud.]
subspecies :
subsp. bilunulata (Risso) Kreutz; Kompend. Eur. Orchid. 93 (2004)
Distribution: S-France; N-Italy; C-Italy; N-Spain
12 FRA SPA 13 ITA
Remarks: fusca agg., bilunulata-group, pollinators Andrena (Euandrena) vulpecula, Andrena (Taeniandrena) wilkella, Andrena flavipes (Spain)
Synonyms: hidden
= Ophrys fusca Link ssp. bilunulata (Risso) Aldasoro & L.Sáez, Fl. Iber. 21: 177 (2005), isonym
= Ophrys bilunulata Risso, Fl. Nice 463 (1844) [fide Paulus (2001)]
= Ophrys dianica M.R.Lowe, Piera, M.B.Crespo & J.E.Arnold, J. Eur. Orchid. 33 (2): 552 (2001)
= Ophrys fusca ssp. bilunulata (Risso) Aldasoro & L.Sáez, Fl. Iber. 21: 177 (2005) [isonym]
subspecies :
subsp. kalirachiensis (Paulus, M.Hirth & Dimadis) comb.ined.
Distribution: N-Greece (Grevena)
13 GRC
Synonyms: hidden
= Ophrys bilunulata ssp. kalirachiensis Paulus, M.Hirth & Dimadis, J. Eur. Orch. 48(1): 66 (2016)
subspecies :
subsp. zonata (Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 96 (2004)
Distribution: Sardinia; Corsica
12 COR SAR
Remarks: fusca agg., bilunulata-group, pollinator Andrena (Zonandrena) flavipes
Synonyms: hidden
= Ophrys zonata Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 374 (1994)
subspecies :
subsp. caesiella (P.Delforge) comb.ined.
Distribution: SW-France; E-Spain; C-Spain; S-Portugal; Baleares; Malta (incl. Comino Isl.); Sicily; Algeria; Tunisia; Morocco
12 BAL FRA POR SPA 13 MLT SIC 20 ALG MOR TUN
Remarks: fusca agg., bilunulata-group, pollinator Andrena flavipes (Paulus 2014). Paulus (2014) transfers the taxon from subfusca to fusca (bilunulata-group) because of the same pollinator
Synonyms: hidden
= Ophrys fusca ssp. caesiella (P.Delforge) Kreutz, Kompend. Eur. Orchid. 93 (2004)
= Ophrys marmorata ssp. caesiella (P.Delforge) Véla & R.Martin, Cat. Syn. Fl. Tunisie 367 (2011)
= Ophrys bilunulata ssp. caesiella (P.Delforge) Paulus, J. Eur. Orch. 46(2): 272 (2014)
= Ophrys caesiella P.Delforge, Nat. Belg. 81 (3): 233 (2000)
= Ophrys fusca ssp. gazella (Devillers-Tersch. & Devillers) Kreutz, Kompend. Eur. Orchid. 94 (2004)
= Ophrys gazella Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 322 (2000)
= Ophrys africana G. & W.Foelsche, J. Eur. Orchid. 33 (2): 656 (2001)
subspecies :
subsp. punctulata (Renz) comb.ined.
Distribution: W-Greece (Korfu, Ithaka, Kefallonia, Attika, Peloponnisos); C-Aegaean Isl.; ?Montenegro; ?Albania
13 GRC 34 EAI
Remarks: fusca agg., bilunulata-group, pollinator Andrena flavipes (Paulus 2014). Was formerly treated as leucadica, name change see Paulus 2014. Ophrys "flavipes-fusca" sensu Paulus.
Synonyms: hidden
= Ophrys bilunulata ssp. punctulata (Renz) Paulus, J. Eur. Orch. 46(2): 272 (2014)
= Ophrys ×punctulata Renz, Repert. Spec. Nov. Regni Veg. 25: 265 (1928)
subspecies :
subsp. sancti-isidorii A.Saliaris, Saliaris & A.Alibertis; J. Eur. Orch. 42(2): 329 (2010)
Name: Sankt Isidors Ragwurz (DE); Saint Isidors Bee Orchid (EN)
Distribution: East Aegaean Isl. (Chios, Lesvos, Samos, Kos, Rhodos, Karpathos); SW-Turkey
34 EAI TUR
Remarks: fusca agg., bilunulata-group, pollinator Andrena flavipes (Paulus 2014)
Synonyms: hidden
= Ophrys bilunulata ssp. sancti-isidorii (A.Saliaris, Saliaris & A.Alibertis) Paulus, J. Eur. Orch. 46(2): 272 (2014)
= Ophrys sancti-isidorii (A.Saliaris, Saliaris & A.Alibertis) P.Delforge, Naturalistes Belges 96(Orchid. 28): 19 (2015)
subspecies :
subsp. leucadica (Renz) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 184 (2002)
Distribution: NW-Greece (Ionian Isl.: Kefallonia, Ithaki); ?W-Greece; ?C-Greece; Croatia
13 CRO GRC
Remarks: fusca agg., bilunulata-group, pollinator Andrena fimbriata. Restricted distribution fide Paulus (2014)
Synonyms: hidden
= Ophrys ×leucadica Renz, Fedde, Repert. Spec. Nov. Regni Veg. 26: 265 (1928)
subspecies :
subsp. israelitica (H.Baumann & Künkele) comb.ined.
Distribution: Greece (Kyklades: Tinos, Andros, Paros, Naxos); Cyprus; Israel; Lebanon; Syria; S-Turkey
13 GRC 34 CYP ISR LEB SYR TUR
Remarks: fusca agg., bilunulata-group, pollinator Andrena (Zonandrena) flavipes. Paulus (2014) transfers the taxon to fusca (bilunulata-group) because of the same pollinator
Synonyms: hidden
= Ophrys israelitica H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (3): 612 (1988)
= Ophrys omegaifera ssp. israelitica (H.Baumann & Künkele) G. & K.Morschek, Orch. Cyprus: 126 (1996)
subspecies :
subsp. funerea (Viv.) Arcang.; Comp. Fl. Ital. ed. 1: 662 (1882)
Distribution: Sardinia; Corsica; Algeria
12 COR SAR 20 ALG
Remarks: fusca agg., funerea-group, pollinator Andrena (Zonandrena) flavipes
Synonyms: hidden
= Ophrys fusca var. funerea (Viv.) Barla, Fl. Illus. Nice, Orchid. 75 (1872)
= Ophrys fusca f. funerea (Viv.) Briq., Prodr. Fl. Corse 1: 348 (1910)
= Ophrys fusca ssp. funerea (Viv.) Camus, Icon. Orch. Eur. 293 (1929) [comb. illeg.]
= Ophrys funerea Viv., Fl. Cors. Sp. Nov. 15 (1824)
= Ophrys ortubae
= Ophrys lutea ssp. funerea Batt. & Trab., Fl. Alger. 23 (1895)
subspecies :
subsp. hespera (Devillers-Tersch. & Devillers) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 169 (2007)
Distribution: C-Italy (Mte. Argentario); ?Croatia
13 ITA
Remarks: fusca agg., funerea-group
Synonyms: hidden
= Ophrys hespera Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 322 (2000)
subspecies :
subsp. arnoldii (P.Delforge) Kreutz; Orchidee (Hamburg) 57(1): 101 (2006)
Distribution: NE-Spain (Catalunya); S-France; Baleares (Ibiza)
12 BAL FRA SPA
Remarks: fusca agg., funerea-group. Pollinator Andrena nigroaenea
Synonyms: hidden
= Ophrys arnoldii P.Delforge, Nat. Belg. 80 (3): 247 (1999)
subspecies :
subsp. calocaerina (Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 93 (2004)
Distribution: S-Greece; C-Greece; ?Sicily; EC-Italy
13 GRC ITA
Remarks: fusca agg., funerea-group, pollinator ?Andrena labialis. Late-flowering taxon
Synonyms: hidden
= Ophrys calocaerina Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7,suppl.): 374 (1994) [fide Paulus 2001]
subspecies :
subsp. perpusilla (Devillers-Tersch. & Devillers) Kreutz; Kompend. Eur. Orchid. 95 (-96) (2004)
Distribution: W-Greece (Lakonia, Kyklades, W-Peloponnisos)
13 GRC
Remarks: fusca agg., bilunulata-group
Synonyms: hidden
= Ophrys perpusilla Devillers-Tersch. & Devillers, Naturalistes Belges 85 (Orchid. 17): 249 (2004)
subspecies :
subsp. creberrima (Paulus) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 174 (2002)
Distribution: Crete; Greece (Kyklades)
13 GRC KRI
Remarks: fusca agg., funerea-group. Status highly disputed
Synonyms: hidden
= Ophrys creberrima Paulus, J. Eur. Orchid. 30 (1): 168 (1998)
subspecies :
subsp. ortuabis (M.P.Grasso & Manca) Kreutz; Kompend. Eur. Orchid. 95 (2004)
Distribution: Sardinia
12 SAR
Remarks: fusca agg., funerea-group. Pollinator Andrena hypopolia
Synonyms: hidden
= Ophrys ortuabis Manca & M.P.Grasso, L´Orchidophile 151(33): 81-86 (2002)
subspecies :
subsp. parvula (Paulus) Kreutz; Kompend. Eur. Orchid. 95 (2004)
Name: Kleine Rhodische Ragwurz (DE); Small Rhodian Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, Kos)
34 EAI
Remarks: fusca agg., funerea-group. Pollinator Andrena tomora.
Synonyms: hidden
= Ophrys parvula Paulus, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 38-86 (2001)
subspecies :
subsp. thriptiensis (Paulus) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 188 (2002)
Distribution: E-Crete
13 KRI
Remarks: fusca agg., attaviria-group. Pollinator Andrena (Euandrena) bicolor
Synonyms: hidden
= Ophrys thriptiensis Paulus, J. Eur. Orchid. 30 (1): 173 (1998)
subspecies :
subsp. attaviria (D.Rückbr., U.Rückbr., Wenker & S.Wenker) Kreutz; Kompend. Eur. Orchid. 93 (2004)
Name: Attaviros-Ragwurz (DE); Attaviros Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, Lesvos, Leipsoi, ?Samos, ?Chios, etc.); SW-Turkey
34 EAI TUR
Remarks: fusca agg., attaviria-group
Synonyms: hidden
= Ophrys attaviria D. & U.Rückbr. & D. & S.Wenker, Ber. Arbeitskr. Heim. Orch. 7 (2): 10 (1990)
subspecies :
subsp. eptapigiensis (Paulus) Kreutz; Kompend. Eur. Orchid. 93 (2004)
Name: Sieben-Quellen-Ragwurz (DE); Seven Springs Bee Orchid (EN)
Distribution: East Aegaean Isl. (Lesvos, Rhodos, ?Karpathos); SW-Turkey
34 EAI TUR
Remarks: fusca agg., attaviria-group, status uncertain, probably a variant or synonym of ssp. attaviria
Synonyms: hidden
= Ophrys attaviria f. eptapigiensis (Paulus) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
= Ophrys attaviria var. eptapigiensis (Paulus) P.Delforge, (2005)
= Ophrys eptapigiensis Paulus, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 38-86 (2001)
subspecies :
subsp. lindia (Paulus) Kreutz; Kompend. Eur. Orchid. 94 (2004)
Name: Lindos-Ragwurz (DE); Lindos Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos)
34 EAI
Remarks: fusca agg., attaviria-group, status uncertain, probably a synonym of ssp. attaviria or ssp. leucadica
Synonyms: hidden
= Ophrys lindia Paulus, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 38-86 (2001)
subspecies :
subsp. phaseliana (D. & U.Rückbr.) Kreutz; Kompend. Eur. Orchid. 96 (2004)
Distribution: SW-Turkey; East Aegaean Isl. (Paros, Kos, Samos, Kythira, Naxos, Amorgos, Astipaleia, Pserimos, Tilos); Malta; Greece (Kyklades: Kea)
13 GRC MLT 34 EAI TUR
Remarks: fusca agg., attaviria-group. Pollinator Andrena similis
Synonyms: hidden
= Ophrys phaseliana D. & U.Rückbr., J. Eur. Orchid. 28 (2): 395 (1996)
= Ophrys fusca ssp. parosica (P.Delforge) Kreutz, Kompend. Eur. Orchid. 95 (2004)
= Ophrys parosica P.Delforge, Nat. Belg. 76 (3): 160 (1995)
= Ophrys parosica var. phaseliana (D. & U.Rückbr.) P.Delforge, Naturalistes Belges 88(Orchid. 20): 247 (2007)
subspecies :
subsp. sabulosa (Paulus & Gack ex P.Delforge) Kreutz; Kompend. Eur. Orchid. 96 (2004)
Distribution: Sicily
13 SIC
Remarks: fusca agg., attaviria-group
Synonyms: hidden
= Ophrys sabulosa Paulus & Gack ex P.Delforge, Naturalistes Belges 85 (Orchid. 17): 114 (2004)
subspecies :
subsp. cesmeensis (Kreutz) Kreutz; Kompend. Eur. Orchid. 93 (2004)
Distribution: SW-Turkey (Cesme peninsula)
34 TUR
Remarks: fusca agg., attaviria-group
Synonyms: hidden
= Ophrys attaviria ssp. cesmeensis Kreutz, Feldführer türk. Orchid. 202 (2003)
= Ophrys cesmeensis (Kreutz) P.Delforge, Naturalistes Belges 85 (Orchid. 17): 252 (2004)
= Ophrys attaviria var. cesmeensis (Kreutz) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
subspecies :
subsp. pelinaea (P.Delforge) comb.ined.
Distribution: East Aegaean Isl. (Lesvos, Chios, Kos, Samos, Leipsoi, Ikaria)
34 EAI
Remarks: fusca agg., attaviria-group
Synonyms: hidden
= Ophrys pelinaea P.Delforge, Naturalistes Belges 88(Orchid. 20): 177 (2007)
= Ophrys meropes P.Delforge, Naturalistes Belges 90(Orchid.22): 184 (2009)
= Ophrys theophrasti Devillers & Devillers-Tersch., Naturalistes Belges 90(Orchid.22): 289 (2009)
subspecies :
subsp. thracica Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 102 (2007)
Distribution: Turkey
34 TUR
Remarks: fusca agg., attaviria-group
Synonyms: hidden
= Ophrys thracica (Kreutz) Devillers & Devillers-Tersch., Naturalistes Belges 90(Orchid.22): 289 (2009)
= Ophrys attaviria f. thracica (Kreutz) P.Delforge, Naturalistes Belges 89(Orchid. 21): 251 (2008)
subspecies :
subsp. migoutiana (H.Gay) Kreutz; Kompend. Eur. Orchid. 95 (2004)
Distribution: Algeria; Tunisia
20 ALG TUN
Remarks: fusca agg., migoutiana-group
Synonyms: hidden
= Ophrys ×migoutiana H.Gay, Mem. Assoc Franc. avanc. sci. Paris (1888) 502
subspecies :
subsp. cinereophila (Paulus & Gack) Faurh.; Orchidee (Hamburg) 53(3): 345 (2002)
Name: Schattenliebende Ragwurz (DE); Shadow Bee Orchids (EN)
Distribution: Greece (Evvoia, Peloponnisos, Ionian Isl.: Kefallenia, Zakynthos, Ithaki); Crete (incl. Gavdos Isl.); East Aegaean Isl. (Amorgos, Naxos, Rhodos, Lesvos, Karpathos, Kasos, Chalki, Kastellorizo, Pserimos, Tilos, etc.); SW-Turkey; Cyprus
13 GRC KRI 34 CYP EAI TUR
Remarks: fusca agg., probably to O. subfusca. Pollinator Andrena (Chlorandrena) cinereophila
Synonyms: hidden
= Ophrys cinereophila Paulus & Gack, J. Eur. Orchid. 30 (1): 170 (1998)
= Ophrys subfusca ssp. cinereophila (Paulus & Gack) Kreutz, Orchidee (Hamburg) 57(1): 102 (2006)
subspecies :
subsp. akhdarensis B. & H.Baumann; Journ. Eur. Orchid. 33 (2): 691-725 (2001)
Distribution: Libya (Cyrenaica)
20 LBY
Remarks: fusca agg., group uncertain
Synonyms: hidden
= Ophrys akhdarensis (B. & H.Baumann) P.Delforge, Naturalistes Belges 85 (Orchid. 17): 251 (2004)
subspecies :
subsp. pseudomigoutiana (R.Martin, Véla & Ouni) comb.ined.
Distribution: Tunisia
20 TUN
Synonyms: hidden
= Ophrys pseudomigoutiana R.Martin, Véla & Ouni, Acta Bot. Gallica Bot. Lett. 162(4): 259 (2015)
species :
Ophrys garganica E.Nelson; Monogr. Ophrys 195 (1962)
subspecies :
subsp. garganica
Name: Gargano-Ragwurz
Distribution: C-Italy; S-Italy; Sicily; Sardinia; Croatia (Korcula Isl.)
12 SAR 13 CRO ITA SIC
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys sphegodes ssp. garganica E.Nelson, Monogr. Ophrys 195 (1962) [nom. nud.]
= Ophrys passionis ssp. garganica (E.Nelson) Kreutz, Eurorchis 17: 109 (2005)
= Ophrys garganica (E.Nelson) O. & E.Danesch, Pl. Syst. Evol. 124: 94 (1975)
= Ophrys incubacea ssp. garganica (E.Nelson ex O.Danesch & E.Danesch) Galesi, Cristaudo & Maugeri, J. Eur. Orchideen 36(2): 505 (2004)
= Ophrys passionis var. garganica (E.Nelson ex O.Danesch & E.Danesch) P.Delforge, Naturalistes Belges 85 (Orchid. 17): 252 (2004)
= Ophrys passionis ssp. garganica E.Nelson ex H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 727 (2005)
subspecies :
subsp. caloptera (Devillers-Tersch. & Devillers) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 170 (2007)
Distribution: NE-Spain; S-France; SE-France; NW-Italy
12 FRA SPA 13 ITA
Remarks: sphegodes agg., pollinator Andrena pilipes (carbonaria)
Synonyms: hidden
= Ophrys arachnitiformis var. passionis (Sennen) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
= Ophrys sphegodes var. garganicoides Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 282 (1986)
= Ophrys garganica ssp. passionis (Sennen ex Devillers & Devillers-Tersch.) Paulus & Gack, Journ. Eur. Orchid. 31 (2), 375 (1999)
= Ophrys caloptera Devillers-Tersch. & Devillers, Natural. belges 87 (Orchid. 19): 228-251 (2006), nom. nov. (superfl.?)
= Ophrys passionis Sennen, Pl. Espagne no. 5881 (1926) et Treb. Mus. Cienc. Nat. Barcelona, 15: Ser. Bot. No. 1, 35 (1931) ex Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 379 (1994)
subspecies :
subsp. majellensis (Helga Daiss & Herm.Daiss) Kreutz; Kompend. Eur. Orchid. 96 (2004)
Distribution: C-Italy (Majella)
13 ITA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys passionis ssp. majellensis (Helga Daiss & Herm.Daiss) Kreutz, Eurorchis 17: 109 (2005)
= Ophrys majellensis (Herm.Daiss & Helga Daiss) P.Delforge, Nat. Belg. 79 (3): 104 (1998)
= Ophrys passionis ssp. majellensis (Herm.Daiss & Helga Daiss) Romolini & Soca, Caesiana 15: 23 (2000)
= Ophrys sphegodes ssp. majellensis Herm.Daiss & Helga Daiss, J. Eur. Orch. 28: 614 (1996 publ. 1997)
subspecies :
subsp. pseudoatrata (S.Hertel & Presser) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 172 (2007)
Distribution: Italy (Basilicata: Potenza)
13 ITA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys pseudoatrata S.Hertel & Presser, J. Eur. Orch. 38(3): 516 (2006)
= Ophrys passionis ssp. pseudoatrata (S.Hertel & Presser) Biagioli & Grünanger, GIROS Orch. Spont. Eur. 58: 6 (2015)
subspecies :
subsp. sipontensis (R.Lorenz & Gembardt) Kreutz; Kompend. Eur. Orchid. 97 (2004)
Distribution: EC-Italy (Mte. Gargano); SE-Italy (Altamura)
13 ITA
Remarks: sphegodes agg., pollinator Xylocopa iris
Synonyms: hidden
= Ophrys sphegodes ssp. sipontensis (Lorenz & Gembardt) Pedersen & Faurh., Jour. Eur. Orch. 37(2): 289 (2005)
= Ophrys garganica ssp. sipontensis (Gumpr.) Del Prete, Webbia 37 (2): 251 (1984)
= Ophrys sphegodes ssp. sipontensis Gumpr., Die Orchidee 18(2): 62 (1967) [nom. inval.]
= Ophrys sipontensis Gumpr. ex O. & E.Danesch, Ophrys Hybr. 43 (1972) [nom. inval.]
= Ophrys sphegodes ssp. garganica var.sipontensis Gumpr. ex Fen., Webbia 29: 281 (1974) [comb. inval.]
= Ophrys sipontensis R.Lorenz & Gembardt, Mitt. Arbeitskr. Heim. Orch. Baden-Württ. 19(3): 474 (1987)
= Ophrys murgiana Cillo, Medagli & Margh., GIROS Notiz. 41: 23 (2009) [southern population of sipontensis]
species :
Ophrys hebes (Kalopissis) B. & E.Willing; Orchidee 31 (4): 159 (1980)
Name: Hebes Ragwurz
Distribution: Greece (widespread except Northeast and Cyclades); Albania; Macedonia
13 ALB GRC NMA
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys sphegodes ssp. hebes Kalopissis, Sci. Ann. Forest Univ. 133 (1975)
variant :
var. negadensis (G.Thiele & W.Thiele) Kreutz; Kompend. Eur. Orchid. 97 (2004)
Distribution: N-Greece
13 GRC
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys negadensis G. & W.Thiele, Journ. Eur. Orchid. 33 (3): 897 (2001)
species :
Ophrys heldreichii Schltr.; Fedde, Repert. 19: 46 (1923)
subspecies :
subsp. heldreichii
Distribution: Crete; East Aegaean Isl. (Karpathos)
13 KRI 34 EAI
Remarks: holosericea agg., oestrifera-group, pollinator Tetralonia berlandi
Synonyms: hidden
= Ophrys cornuta ssp. heldreichii (Schltr.) Renz, Repert. Spec. Nov. Regni Veg. 28: 248 (1930)
= Ophrys oestrifera ssp. heldreichii (Schltr.) Hayek
= Ophrys orientalis ssp. heldreichii (Schltr.) Soó, Monogr. Iconogr. Orchid. Eur. 2(2/3): 65 (1931), et 2: 65, 448 (1931)
= Ophrys scolopax ssp. heldreichii (Schltr.) E.Nelson, Gestaltwandel 160 (1962)
= Ophrys scolopax ssp. heldreichii (Schltr.) E.G.Camus, Icon. Orchid. Europe, Text(1): 315 (1928)
= Ophrys heldreichii var. schlechterana (Soó) Soó
= Ophrys fuciflora f. pseudooestrifera Renz
= Ophrys heldreichii ssp. pusilla B. & H.Baumann, Jour. Eur. Orch. 37(4): 951 (2005)
subspecies :
subsp. calypsus (M.Hirth & H.Spaeth) Kreutz; Kompend. Eur. Orchid. 98 (2004)
Distribution: East Aegaean Isl. (Leipsoi, Pserimos, etc., Karpathos, Kasos); SW-Turkey; Greece (Kyklades, Kythira)
13 GRC 34 EAI TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys calypsus M.Hirth & H.Spaeth, J. Eur. Orchid. 26 (3­4): 433, 430 (1994)
= Ophrys heldreichii var. calypsus (M.Hirth & H.Spaeth) P.Delforge, Nat. Belg. 76 (3): 176 (1995)
variant :
var. pseudoapulica P.Delforge; Nat. Belg. 76 (3): 176 (1995)
Name: Calypso-Ragwurz (DE); Calypso Bee Orchid (EN)
Distribution: Greece; East Aegaean Isl. (Rhodos); W-Turkey
13 GRC 34 EAI TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys calypsus var. pseudoapulica (P.Delforge) P.Delforge, Naturalistes Belges 85 (Orchid. 17): 251 (2004)
= Ophrys heldreichii ssp. pseudoapulica (P.Delforge) Kreutz, Kompend. Eur. Orchid. 98 (2004)
variant :
var. scolopaxoides P.Delforge; Nat. Belg. 76 (3): 176 (1995)
Name: Calypso-Ragwurz (DE); Calypso Bee Orchid (EN)
Distribution: East Aegaean Isl. (Leipsoi, Rhodos)
34 EAI
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys calypsus var. scolopaxoides (P.Delforge) P.Delforge, Naturalistes Belges 85 (Orchid. 17): 252 (2004)
subspecies :
subsp. chiosica (P.Delforge, Onckelinx & Saliaris) comb.ined.
Distribution: East Aegaean Isl. (Chios)
34 EAI
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys chiosica P.Delforge, Onckelinx & Saliaris, Naturalistes Belges 88(Orchid. 20): 178 (2007)
subspecies :
subsp. homeri (M.Hirth & H.Spaeth) comb.ined.
Distribution: East Aegaean Isl. (Chios, Lesvos, Samos, Leipsoi, Kos); SW-Turkey (Bodrum peninsula)
34 EAI TUR
Remarks: holosericea agg., oestrifera-group. Pollinator Cubitalia parvicornis.
Synonyms: hidden
= Ophrys homeri M.Hirth & H.Spaeth, J. Eur. Orchid. 30 (1): 9, 7 (prepr. 30 Sept. 1997)
= Ophrys holosericea ssp. homeri (M.Hirth & H.Spaeth) Kreutz, Kompend. Eur. Orchid. 101 (2004)
subspecies :
subsp. pharia (Devillers & Devillers-Tersch.) Kreutz; Kompend. Eur. Orchid. 98 (2004)
Distribution: Croatia (Hvar Isl.)
13 CRO
Remarks: holosericea agg., oestrifera-group. Taxonomic status disputed
Synonyms: hidden
= Ophrys pharia Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid. 17): 233 (2004)
subspecies :
subsp. ptolemaea (P.Delforge) comb.ined.
Distribution: East Aegaean Isl. (Kos)
34 EAI
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys ptolemaea P.Delforge, Naturalistes Belges 90(Orchid.22): 185 (2009)
subspecies :
subsp. polyxo (J.Mast, M.-A.Garnier, Devillers-Tersch. & Devillers) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 173 (2007)
Name: Rhodische Gehörnte Ragwurz (DE); Rhodian Horned Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos)
34 EAI
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys polyxo J.Mast, Garnier, Devillers-Tersch. & Devillers, Les Naturalistes Belges 86: 147-156 (2006)
subspecies :
subsp. schlechteriana Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 909 (1926)
Distribution: C-Greece (Evvoia, Attiki); S-Greece (Peloponnisos)
13 GRC
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys oestrifera ssp. schlechteriana (Soó) Kreutz, Kompend. Eur. Orchid. 109 (2004)
= Ophrys heldreichii var. schlechteriana (Soó) Soó, Monogr. Iconogr. Orchid. Eur. 2: 64: 447 (1938)
= Ophrys schlechteriana (Soó) Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 378 (1994)
= Ophrys schlechteriana Soó, Notizbl. Bot. Gart. Berlin 9: 909 (1926)
= Ophrys ×skopelii Renz, Fedde, Repert. 25: 264 (1928)
subspecies :
subsp. samiotissa (Hirth & Paulus) comb.ined.
Distribution: East Aegaean Isl. (Samos)
34 EAI
Remarks: holosericea agg., oestrifera-group. Pollinator Eucera punctulata.
Synonyms: hidden
= Ophrys samiotissa M.Hirth & Paulus, J. Eur. Orch. 43(4): 863-873 (2011)
species :
Ophrys helenae Renz; Fedde, Repert. 25: 251 (1928)
Name: Helenes Ragwurz
Distribution: Greece (NW-Greece, NW-Peloponnisos); Albania
13 ALB GRC
Remarks: mammosa agg.
Conservation Status: IUCN R
Synonyms: hidden
= Ophrys aranifera ssp. helenae (Renz) Soó, Repert. Spec. Nov. Regni Veg. 26: 278 (1929)
= Ophrys mammosa ssp. helenae (Renz) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 380, 383 (1973)
= Ophrys mammosa ssp. helenae (Renz) Soó, Fedde, Repert. 8517-8): 446 (1974), with basionym ref.
= Ophrys sphecodes ssp. helenae (Renz) Soó & D.M.Moore, Bot. Journ. Linn. Soc. 76 (4): 367 (1978)
= Ophrys sphegodes var. helenae (Renz) H.Sund., Europ. Medit. Orchideen 45 (1975)
species :
Ophrys holosericea (Burm.fil.) Greuter; Boissiera 13: 185 (1967)
subspecies :
subsp. holosericea
Name: Hummel-Ragwurz
Distribution: Austria; Belgium; Luxembourg; France; Germany; Luxembourg; +Netherlands; Switzerland; Liechtenstein; Hungary
11 AUT BGM GER HUN LIE LUX NET SWI 12 FRA
Remarks: holosericea agg., holosericea s. str., pollinator Eucera longicornis, E. nigrescens
Synonyms: hidden
= Arachnites fuciflora F.W.Schmidt, Fl. Boehm. 1: 76 (1793)
= Arachnites oxyrhynchus Tod., Orch. Sicul. 81 (1842)
= Epipactis arachnites (Scop.) F.W.Schmidt, Mayer, Phys. Aufs. 1: 249 (1791)
= Ophrys apiculata Rich., Mem. Mus. Par. 4: 55 (1818)
= Ophrys apifera ssp. arachnites (Scop.) Watson, (1867)
= Ophrys arachnites (Scop.) Reichard, Fl. Moeno-Franc. 2: 89 (1778) [nom. illeg.]
= Ophrys arachnoides Andr., Bot. Repos. 7: tab. 470 (1807)
= Ophrys arachnoides Peterm., Flora 27: 544 (1844)
= Ophrys bombyliflora Spreng., Syst. 3: 702 (1826)
= Ophrys flavescens Sassenf., Trierisch. Fl. ed. 3, 79 (1903)
= Ophrys fuciflora (F.W.Schmidt) Moench, Meth. Suppl. 311 (1802) [non Crantz 1769]
= Ophrys fuciflora f. tallosii Soó, Acta Bot. Acad. Sci. Hung. 18 (1-2): 173 (1973)
= Ophrys fuciflora f. triloboviridis Soó, Acta Bot. Acad. Sci. Hung. 18 (1-2): 173 (1973)
= Ophrys fuciflora var. apiformoides Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 282 (1986)
= Ophrys fuciflora var. carmeloides Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 282 (1986)
= Ophrys fucus J.P.Bergeret, Phytonom. Univ. 2: 65 (1784-1786)
= Ophrys holoserica f. chlorantha Lumare, Medagli, Dura & Anelli, GIROS Orch. Spont. Eur. 62(1): 91 (2019)
= Ophrys holoserica var. aliceae Argiolas, GIROS Orch. Spont. Eur. 61(2): 350 (2018)
= Ophrys holosericea var. bicolor E.Breiner & R.Breiner, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13 (2): 142 (1981)
= Ophrys holosericea var. flavescens (Rosbach) H.Sund., Taxon 24 (5-6): 625 1975)
= Ophrys holosericea var. scolopaxioides P.Haffner, Faunist.-florist. Not. Saarland 28: 517-527 (1996) [not validly published]
= Ophrys insectifera ssp. arachnites (Scop.) Moggr. & Rchb.fil., Nova Acta Acad. Leop.-Carol. German. Nat. Cur. 35: 12 (1869)
= Ophrys oestrifera Rchb., Fl. Germ. Excurs. 128 (1830)
= Ophrys ×pulchra E.G.Camus, Bull. Soc. Bot. Fr. 38: 43 (1891)
= Orchis arachnites Scop., Fl. Carn. ed. II, 2: 194 (1772)
= Orchis fuciflora Crantz, Stirp. Austr. ed. II, fasc. 6: 483 (1769)
= Orchis fuciflora Haller ex Schrank, Baier. Fl. 242 (1789) [nom. illeg.]
= Orchis holosericea Burm.fil., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur., 4 (Appendix): 237 (1770)
subspecies :
subsp. parvimaculata (O. & E.Danesch) O. & E.Danesch; Pl. Syst. Evol. 124 (2): 129 (1975)
Distribution: C-Italy; SE-Italy (Mte. Gargano to Lecce); Croatia (Istria: Bale, Rovinj)
13 CRO ITA
Remarks: holosericea agg., "bornmuelleri-like"-group. Pollinator Eucera nigrescens. Populations in Istria are probably an undescribed taxon (Paulus 2014)
Synonyms: hidden
= Ophrys parvimaculata (O.Danesch & E.Danesch) Paulus & Gack, Israel J. Bot. 39(1-2): 78 (1990)
= Ophrys fuciflora ssp. parvimaculata O. & E.Danesch, Die Orchidee 21(1): 21 (1970)
= Ophrys peucetiae Lozito, D´Emerico, Medagli & Turco, GIROS Notiz. 2: 6 (2012)
subspecies :
subsp. montis-aviarii (O.Hirschy & L.Bennery) comb.ined.
Distribution: S-France (Provence)
12 FRA
Synonyms: hidden
= Ophrys montis-aviarii O.Hirschy & L.Bennery, L´Orchidophile 203: 352 (2014)
subspecies :
subsp. aegirtica (P.Delforge) comb.ined.
Distribution: SW-France; C-France
12 FRA
Remarks: holosericea agg., tetraloniae-group. Pollinator Eucera ephippia
Synonyms: hidden
= Ophrys aegirtica P.Delforge, Nat. Belg. 77 (4): 205 (1996)
= Ophrys truncata Dulac, Fl. Hautes-Pyr. 128 (1867) [nom. illeg.]
subspecies :
subsp. aestuariensis (J.-P.Ring, J.-C.Querré, Biron & Fouquet) comb.ined.
Distribution: France
12 FRA
Synonyms: hidden
= Ophrys fuciflora ssp. aestuariensis J.-P.Ring, J.-C.Querré, Biron & Fouquet, Orchidophile (Argenteuil) 51(225): 166 (2020)
subspecies :
subsp. holubyana (András.) J.Dostál; Folia Mus. Rer. Nat. Bohem. Occid., Bot., 21: 17 (1984)
Distribution: Czech Republic; Slovakia; Hungary
11 CZE HUN SLO
Remarks: holosericea agg., group uncertain. Hybridogenous origin Ophrys oestrifera ssp. bicornis x holosericea. Pollinator Eucera longicornis
Conservation Status: IUCN I
Synonyms: hidden
= Ophrys holosericea ssp. holubyana (András.) Kreutz, Kompend. Eur. Orchid. 101 (2004) [isonym]
= Ophrys arachnites ssp. holubyana (András.) Holub, Folia Geobot. Phytotax. 18 (2): 205 (1983)
= Ophrys holubyana András., Magyar Bot. Lapok 16: 11 (1917) [pro syn.]
subspecies :
subsp. brachyotus (Rchb.) comb.ined.
Distribution: N-Italy (Lake Garda)
13 ITA
Remarks: holosericea agg., tetraloniae-group.
Synonyms: hidden
= Ophrys brachyotus Rchb., Fl. Germ. Excurs. 128 (1830)
subspecies :
subsp. serotina (Rolli ex Paulus) Kreutz; Kompend. Eur. Orchid. 102 (2004)
Distribution: NE-Italy; S-France; Switzerland; Croatia
11 SWI 12 FRA 13 CRO ITA
Remarks: holosericea agg., tetraloniae-group. Pollinator Eucera clypeata.
Synonyms: hidden
= Ophrys fuciflora ssp. lorenae E.De Martino & Centur., Jour. Eur. Orch. 34(3): 589 (2002)
= Ophrys lorenae (E.De Martino & Centur.) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
= Ophrys fuciflora var. untchjii M.Schultze, Asch. & Graebn., Syn. 3: 631 (1907)
= Ophrys holosericea ssp. lorenae (E.De Martino & Centur.) Kreutz, Kompend. Eur. Orchid. 101 (2004)
= Ophrys holosericea ssp. medea (Devillers & Devillers-Tersch.) Kreutz, Eurorchis 17: 109 (2005)
= Ophrys holosericea ssp. untchjii (M.Schultze) Kreutz, Kompend. Eur. Orchid. 102 (2004)
= Ophrys medea Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid. 17): 48 (2004)
= Ophrys serotina Rolli ex Paulus, Ber. Arbeitskr. Heim. Orch. 17 (2): 23, 24 (2000) [fide Martino & Centur. 2002]
= Ophrys serotina Rolli ex Cortesi, Ann. Bot., Roma, 8: 213 (1910) [fide Martino & Centur. 2002]
= Ophrys untchjii (M.Schultze) P.Delforge, Nat. Belg. 81 (4): 398 (2000)
= Ophrys ×gerstlaueri A.Fuchs, Mitt. Bayer. Bot. Ges. 3: 388 (1917)
= Ophrys istriensis S.Hertel, Paulus & Weyland, Ber. Arbeitskreis. Heimische Orchid. 33(1): 79 (2016) [tentative synonym]
subspecies :
subsp. pseudoscolopax (Moggr.) comb.ined.
Distribution: S-France; SE-France; ?NW-Italy; ?Croatia
12 FRA
Remarks: holosericea agg., tetraloniae-group. Pollinator ?Tetralonia crassa
Synonyms: hidden
= Ophrys fuciflora ssp. demangei Scappat., Bull. S. F. O. Rhône-Alpes 15: 88 (2015)
= Ophrys demangei (Scappat.) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
= Ophrys arachnites var. filiforme Risso
= Ophrys fuciflora var. linearis (Moggr.) E.G.Camus, Bergon & A.Camus, (1908)
= Ophrys fuciflora var. linearis Moggr., Nov. Actorum Acad. Caes. Leop.-Carol. German. Nat. Cur. 35: 12 (1869)
= Ophrys holosericea ssp. linearis (Moggr.) Kreutz, Kompend. Eur. Orchid. 101 (2004)
= Ophrys holosericea var. linearis (Moggr.) Landwehr, Orchid. Sauvages France & Europe 2: 472 (1983)
= Ophrys insectifera ssp. arachnites var.pseudoscolopax Moggr., Verh. Kaiserl. Leopoldino-Carolinischen deutsch. Acad. Naturforscher (Novo Acta Physico Medica) 35: 12, Abb.: Farbtafel III, fig. 24-26 (1869), nom. dub.
= Ophrys insectifera subvar. linearis Moggr., Nova Acta Acad. Leop.-Carol. 35: 12 (1869)
= Ophrys linearis (Moggr.) P.Delforge, Devillers & Devillers-Tersch., Nat. Belg. 81 (3): 153 (2000)
= Ophrys pseudoscolopax (Moggr.) Paulus & Gack, J. Eur. Orchid. 31 (2), 394 (1999)
subspecies :
subsp. querciphila (Nicole, Hervy & Soca) comb.ined.
Distribution: France (Languedoc)
12 FRA
Synonyms: hidden
= Ophrys querciphila Nicole, Hervy & Soca, Orchidophile (Asnières) 212: 94 (2017)
subspecies :
subsp. gracilis (Büel, O. & E.Danesch) O. & E.Danesch; Pl. Syst. Evol, 124 (2): 129 (1975)
Distribution: Italy (Tuscany to Basilicata, ?Apulia); ?Croatia
13 ITA
Remarks: holosericea agg., holosericea s. str., pollinator ?Eucera clypeata. Possibly identical with ssp. tetraloniae
Synonyms: hidden
= Ophrys fuciflora ssp. gracilis Büel, O. & E.Danesch, Orchidee 23 (4): 160 (1972), without type, et Orchidee 24 (1): 36 (1973) [with type]
= Ophrys fuciflora var. gracilis (Büel, O.Danesch & E.Danesch) Faurh., J. Eur. Orch. 40(1): 57 (2008)
= Ophrys gracilis (Büel, O.Danesch & E.Danesch) Paulus, Ber. Arbeitskr. Heim. Orch. 13(2): 9 (1996) [comb. superfl.]
= Ophrys gracilis (Büel, O. & E.Danesch) Englmaier, Abh. zool.-bot. Ges. Österr. 22: 140 (1984)
subspecies :
subsp. elatior (Gumpr.) H.Baumann & Künkele; Farn- u. Blütenpfl. Baden-Württemberg 8: 416 (1998)
Distribution: E-France (Alsace, Jura); SW-Germany; SW-Switzerland
11 GER SWI 12 FRA
Remarks: holosericea agg., tetraloniae-group. Pollinator Tetraloniella salicariae
Synonyms: hidden
= Ophrys holosericea ssp. elatior (Gumpr.) Gumpr., Orchidee 34 (5): 179 (1983) [nom. inval.]
= Ophrys fuciflora ssp. elatior Gumpr. ex R.Engel & P.Quentin, L´Orchidophile 123: 168 (1996)
= Ophrys fuciflora ssp. elatior Gumpr., Orchidee 31 (2): 60 (1980)
= Ophrys elatior Gumpr. ex Paulus, Ber. Arbeitskrs. Heim. Orchid. 14 (1996)
subspecies :
subsp. tetraloniae (W.P.Teschner) Kreutz; Kompend. Eur. Orchid. 102 (2004)
Distribution: Croatia (Istria, Dalmatia); Slovenia; ?C-Italy; Bosnia & Hercegovina
13 BIH CRO SLV
Remarks: holosericea agg., tetraloniae-group. Pollinator Tetraloniella fulvescens (dufouri), Tetraloniella inulae
Synonyms: hidden
= Ophrys tetraloniae W.P.Teschner, Orchidee 38 (5): 223 (1987)
= Ophrys fuciflora ssp. tetraloniae (W.P.Teschner) Faurh., J. Eur. Orch. 40(1): 54 (2008)
= Ophrys holosericea ssp. laxiflora Zelesny & Kreutz, Jour. Eur. Orch. 39(3/4): 449 (2007)
subspecies :
subsp. annae (Devillers-Tersch. & Devillers) H.Baumann, Giotta, Künkele, R.Lorenz & Piccitto; J. Eur. Orchid. 27 (2): 217 (1995)
Distribution: Sardinia; S-Corsica
12 COR SAR
Remarks: holosericea agg., tetraloniae-group. Pollinator unknown
Synonyms: hidden
= Ophrys fuciflora ssp. annae (Devillers-Tersch. & Devillers) Engel & Quentin, Orchidophile 124: 205 (1996)
= Ophrys annae Devillers-Tersch. & Devillers, Nat. Belg. 73 (3): 110 (1992)
subspecies :
subsp. robertiarum D´Alonzo & Perilli; GIROS Notiz. 60(1): 19 (2017) [nothossp.]
Distribution: Italy
13 ITA
Remarks: Ophrys apulica x cinnabarina
subspecies :
subsp. pinguis (Romolini & Soca) Kreutz; GIROS Orch. Spont. Eur. 58: 6 (2015)
Distribution: C-Italy (Abruzzo, Lazio, Toscana)
13 ITA
Remarks: holosericea agg., ?tetraloniae-group, related to ssp. serotina and ssp. linearis
Synonyms: hidden
= Ophrys pinguis Romolini & Soca, J. Eur. Orch. 43(4): 766 (2011)
subspecies :
subsp. cinnabarina (Romolini & Soca) Kreutz; GIROS Orch. Spont. Eur. 58: 6 (2015)
Distribution: C-Italy (Abruzzo, Lazio, Toscana, Latium)
13 ITA
Remarks: holosericea agg., ?tetraloniae-group, related to ssp. serotina and ssp. linearis
Synonyms: hidden
= Ophrys cinnabarina Romolini & Soca, J. Eur. Orch. 43(4): 767-768 (2011)
subspecies :
subsp. appennina (Romolini & Soca) Kreutz; GIROS Orch. Spont. Eur. 58: 6 (2015)
Distribution: C-Italy (Abruzzo, Basilicata, Calabria, Campania, Lazio, Liguria, Molise, Toscana, Umbria)
13 ITA
Remarks: holosericea agg., ?tetraloniae-group, related to ssp. serotina and ssp. linearis
Synonyms: hidden
= Ophrys appennina Romolini & Soca, J. Eur. Orch. 43(4): 764 (2011)
subspecies :
subsp. posidonia (P.Delforge) Kreutz; Kompend. Eur. Orchid. 102 (2004)
Distribution: SC-Italy; S-Italy
13 ITA
Remarks: holosericea agg., tetraloniae-group
Synonyms: hidden
= Ophrys posidonia P.Delforge, Nat. Belg. 81 (3): 173 (2000)
= Ophrys fuciflora var. posidonia (P.Delforge) Faurh., J. Eur. Orch. 40(1): 56 (2008)
subspecies :
subsp. chestermanii J.J.Wood; Orchidee 33 (2): 67 (1982)
Distribution: C-Sardinia
12 SAR
Remarks: holosericea agg., "bornmuelleri-like"-group. Pollinator Psithyrus vestalis
Synonyms: hidden
= Ophrys chestermanii (J.J.Wood) Gölz & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (1): 115 (1988)
= Ophrys fuciflora ssp. chestermanii (J.J.Wood) H.Blatt & W.Wirth, Ber. Arbeitskr. Heim. Orch. 7 (1): 7 (1990)
subspecies :
subsp. aeoli (P.Delforge) Kreutz; Kompend. Eur. Orchid. 99 (2004)
Distribution: Greece (Kyklades: Amorgos, Astypaleia)
13 GRC
Remarks: holosericea agg., "bornmuelleri-like"-group
Synonyms: hidden
= Ophrys aeoli P.Delforge, Natural. Belges 78 (3) (Orchid. 10): 153-176 (1997)
subspecies :
subsp. halkionis (G. & H.Kretzschmar) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 175 (2007)
Distribution: Greece (Kyklades: Naxos, Tinos, Kimolos, ?Kea, ?Syros, ?Serifos)
13 GRC
Remarks: holosericea agg., "bornmuelleri-like"-group
Synonyms: hidden
= Ophrys andria ssp. halkionis (G. & H.Kretzschmar) Kreutz, Eurorchis 17: 107 (2005)
= Ophrys andria var. halkionis G. & H.Kretzschmar, Ber. Arbeitskr. Heim. Orch. 13 (1): 9 (1996)
= Ophrys thesei P.Delforge, Natural. Belges 78 (3) (Orchid. 10): 153-176 (1997)
subspecies :
subsp. andria (P.Delforge) Faurh.; Orchidee (Hamburg) 53(3): 345 (2002)
Distribution: Greece (Andros)
13 GRC
Remarks: holosericea agg., "bornmuelleri-like"-group
Synonyms: hidden
= Ophrys andria P.Delforge, Nat. Belg. 75 (4): 136 (1994)
= Ophrys fuciflora ssp. andria (P.Delforge) Faurh., Ber. Arbeitskreis. Heimische Orchid. 22(2): 7 (2005 publ. 2006)
subspecies :
subsp. episcopalis (Poir.) Kreutz; Kompend. Eur. Orchid. 100 (2004)
Distribution: Crete
13 KRI
Remarks: holosericea agg., "bornmuelleri-like"-group. Pollinator Eucera cressa. In strict sense restricted to Crete
Synonyms: hidden
= Ophrys episcopalis Poir., Encyc. Suppl. 4: 170 (1810)
= Ophrys fuciflora ssp. maxima (H.Fleischm.) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 382, 384 (1973)
= Ophrys fuciflora var. maxima H.Fleischm., Österr. Bot. Zeitschr. 74: 188 (1925)
= Ophrys holosericea ssp. maxima (H.Fleischm.) Greuter, Boissiera 13: 185 (1967)
= Ophrys maxima (H.Fleischm.) Paulus & Gack, Jahresber. Naturwiss. Ver. Wuppertal 39: 79 (1986)
subspecies :
subsp. halia (Paulus) Kreutz; Kompend. Eur. Orchid. 100 (2004)
Name: Halia-Ragwurz (DE); Halia Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos)
34 EAI
Remarks: holosericea agg., "bornmuelleri-like"-group. Status uncertain, very similar to ssp. episcopalis and possibly a synonym
Synonyms: hidden
= Ophrys halia Paulus, Ber. Arbeitskr. Heim. Orch. 18 (2) (2001)
subspecies :
subsp. saliarisii (Paulus & Hirth) comb.ined.
Name: Saliarisis Ragwurz (DE); Saliarisi´s Bee Orchid (EN)
Distribution: East Aegaean Isl. (Chios, Rhodos)
34 EAI
Remarks: holosericea agg., "bornmuelleri-like"-group. Status uncertain, probably only a color variant of ssp. halia. Pollinator Eucera plumigera
Synonyms: hidden
= Ophrys saliarisii Paulus & M.Hirth, Journ. Eur. Orch. 41(3/4): 671 (2009)
subspecies :
subsp. aramaeorum (P.Delforge) Kreutz; Kompend. Eur. Orchid. 99 (2004)
Distribution: SC-Turkey (Anatolia: Mersin, Adana, Hatay, Diyarbakir, Siirt)
34 TUR
Remarks: holosericea agg., "bornmuelleri-like"-group.
Synonyms: hidden
= Ophrys aramaeorum P.Delforge, Nat. Belg. 81 (3): 228 (2000)
subspecies :
subsp. libanotica H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 724 (2005)
Distribution: Lebanon; W-Syria; N-Israel; W-Jordania
34 ISR JOR LEB SYR
Remarks: holosericea agg., "bornmuelleri-like"-group.
Synonyms: hidden
= Ophrys libanotica (B. & H.Baumann) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
= Ophrys episcopalis ssp. libanotica (B. & H.Baumann) Véla & Viglione, Acta Bot. Gallica Bot. Lett. 162(4): 278 (2015)
subspecies :
subsp. mesopotamica Kreutz & H.Baumgartner bis; J. Eur. Orch. 46(2): 449 (2014)
Distribution: Turkey (SE-Anatolia: Siirt, Sirnak); ?Syria
34 TUR
Remarks: holosericea agg., "bornmuelleri-like"-group.
Synonyms: hidden
= Ophrys mesopotamica (Kreutz & H.Baumgartner bis) P.Delforge, Naturalistes Belges 96(Orchid. 28): 18 (2015)
subspecies :
subsp. oblita (Kreutz, Gügel & W.Hahn) comb.ined.
Distribution: W-Turkey; ?Aegaean Isl. (Samos)
34 TUR
Remarks: holosericea agg., "bornmuelleri-like"-group
Synonyms: hidden
= Ophrys oblita Kreutz, Gügel & W.Hahn, Ber. Arbeitskreis. Heimische Orchid. 26(2): 55 (2009)
= Ophrys fuciflora ssp. oblita (Kreutz, Gügel & W.Hahn) Faurh., H.A.Pedersen & S.G.Christ., Ber. Arbeitskreis. Heimische Orchid. 28(2): 208 (2011)
= Ophrys episcopalis var. samia P.Delforge, Naturalistes Belges 89(Orchid. 21): 205 (2008)
= Ophrys samia (P.Delforge) P.Delforge, Naturalistes Belges 92(Orchid. 24): 203 (2011)
subspecies :
subsp. helios (Kreutz) Kreutz; Kompend. Eur. Orchid. 100 (2004)
Distribution: East Aegaean Isl. (Karpathos); E-Crete
13 KRI 34 EAI
Remarks: holosericea agg., "bornmuelleri-like"-group. Status uncertain, maybe a synonym of ssp. episcopalis
Synonyms: hidden
= Ophrys helios Kreutz, Journ. Eur. Orchid. 33 (3): 872 (2001) [fide Kretzschmar 2002]
= Ophrys holoserica ssp. taloniensis Kreutz, Ber. Arbeitskreis. Heimische Orchid. 24(1): 111 (2007)
= Ophrys helios f. taloniensis (Kreutz) P.Delforge, Naturalistes Belges 89(Orchid. 21): 251 (2008)
subspecies :
subsp. halicarnassia H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 478 (1986) [nothossp.]
Distribution: Turkey
34 TUR
Remarks: holosericea agg., "bornmuelleri-like"-group. Origin Ophrys holosericea ssp. heterochila x holosericea ssp. holosericea
subspecies :
subsp. heterochila Renz & Taubenheim; Orchidee 31 (6): 237 (1980)
Name: Dodekanes-Ragwurz (DE); Dodekanes Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, Nisyros, Samos, Naxos, Kalymnos, Leipsoi); SW-Turkey
34 EAI TUR
Remarks: holosericea agg., "bornmuelleri-like"-group. Pollinator Eucera cypria
Conservation Status: IUCN E (Anatolia)
Synonyms: hidden
= Ophrys dodekanensis H.Kretzschmar & Kreutz, Ber. Arbeitskr. Heim. Orchid. 18 (1) (2001)
= Ophrys fuciflora ssp. heterochila (Renz & Taubenheim) H.Blatt & W.Wirth, Ber. Arbeitskr. Heim. Orch. 7 (1): 7 (1990)
= Ophrys heterochila (Renz & Taubenheim) Paulus & Gack, Jahresber. Naturwiss. Ver. Wuppertal 43: 94 (1990)
= Ophrys heterochila (Renz & Taubenheim) P.Delforge, Nat. Belg. 72 (3): 101 (1991) [isonym]
= Ophrys heterochila ssp. dodekanensis (H.Kretzschmar & Kreutz) Kreutz, Kompend. Eur. Orchid. 109 (2004)
subspecies :
subsp. oreas (Devillers & Devillers-Tersch.) comb.ined.
Name: Profitis-Ilias-Hummelragwurz (DE); Profitis Ilias Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos)
34 EAI
Remarks: holosericea agg., "bornmuelleri-like"-group
Synonyms: hidden
= Ophrys oreas Devillers & Devillers-Tersch., Naturalistes Belges 90(Orchid.22): 290 (2009)
subspecies :
subsp. lyciensis (Paulus, Gügel, D. & U.Rückbr.) H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 724 (2005)
Distribution: SW-Turkey; East Aegaean Isl. (Chios, Kastellorizo)
34 EAI TUR
Remarks: holosericea agg., "bornmuelleri-like"-group. Pollinator Eucera graeca
Synonyms: hidden
= Ophrys candica ssp. lyciensis (Paulus, Gügel, D. & U.Rückbr.) Kreutz, Kompend. Eur. Orchid. 90 (2004)
= Ophrys lyciensis Paulus, Gügel & Rückbrodt, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 19-33 (2001)
subspecies :
subsp. colossaea (Delforge) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 173 (2007)
Name: Große rhodische Hummelragwurz (DE); Colossal Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, Kos); SW-Turkey
34 EAI TUR
Remarks: holosericea agg., group uncertain. Pollinator Eucera dalmatica. A fairly distinctive, late flowering taxon
Synonyms: hidden
= Ophrys colossaea P.Delforge, Naturalistes Belges 87(Orchid. 19): 214 (2006)
subspecies :
subsp. ethemeae Kreutz; Ber. Arbeitskreis. Heimische Orchid. 28(2): 291 (289-293; figs.) (2011)
Distribution: East Aegaean Isl.
34 EAI
Remarks: holosericea agg., group uncertain
Synonyms: hidden
= Ophrys ethemeae (Kreutz) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
subspecies :
subsp. toroslaria Kreutz, Gügel & Zaiss; Ber. Arbeitskreis. Heimische Orchid. 28(2): 286 (285-289; figs.) (2011)
Distribution: Turkey
34 TUR
Remarks: holosericea agg., group uncertain
subspecies :
subsp. gresivaudanica (O.Gerbaud) Kreutz; Kompend. Eur. Orchid. 100 (2004)
Distribution: France
12 FRA
Remarks: holosericea agg., tetraloniae-group
Synonyms: hidden
= Ophrys gresivaudanica O.Gerbaud, Jour. Eur. Orch. 34 (1): 63-86 (2002)
subspecies :
subsp. dinarica (Kranjcev & P.Delforge) Kreutz; Kompend. Eur. Orchid. 99 (2004)
Distribution: Croatia (Dalmatinska Zagora); Italy
13 CRO ITA
Remarks: holosericea agg., tetraloniae-group
Synonyms: hidden
= Ophrys holosericea f. dinarica (Kranjcev & P.Delforge) Faurh., J. Eur. Orch. 41(3/4): 637 (2009)
= Ophrys dinarica Kranjcev & P.Delforge, Naturalistes Belges 85 (Orchid. 17): 32 (2004)
= Ophrys fuciflora var. dinarica (Kranjcev & P.Delforge) Faurh., J. Eur. Orch. 41(3-4): 637 (2009)
subspecies :
subsp. personata (P.Delforge) comb.ined.
Distribution: Italy (Umbria to Abruzzi)
13 ITA
Synonyms: hidden
= Ophrys personata P.Delforge, Naturalistes Belges 96(Orchid. 28): 18 (2015)
subspecies :
subsp. druentica (P.Delforge & Viglione) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 174 (2007)
Distribution: SE-France (Provence)
12 FRA
Remarks: holosericea agg., tetraloniae-group
Synonyms: hidden
= Ophrys druentica P.Delforge & Viglione, Natural. belges 87 (Orchid. 19): 121-140 (2006)
subspecies :
subsp. delfuocoi D´Alonzo & Perilli; GIROS Orch. Spont. Eur. 61(1): 65 (2018) [nothossp.]
Distribution: Italy
13 ITA
subspecies :
subsp. apulica (O. & E.Danesch) Buttler; Willdenowia 16(1): 115 (1986)
Name: Apulische Ragwurz
Distribution: S-Italy
13 ITA
Remarks: holosericea agg., holosericea s. str., pollinator Tetralonia berlandi
Synonyms: hidden
= Ophrys fuciflora ssp. apulica O. & E.Danesch, Orchidee 21: 20 (1970)
= Ophrys apulica (O. & E.Danesch) O. & E.Danesch ex Gölz & H.R.Reinhard, Orchidee 29 (3) (1978)
= Ophrys holosericea ssp. apulica (O. & E.Danesch) Buttler, Willdenowia 16 (1): 115 (1986)
= Ophrys apulica (O. & E.Danesch) O. & E.Danesch, Pl. Syst. Evol. 124 (2): 82 (1975)
= Ophrys holosericea ssp. apulica (O. & E.Danesch) H.Sund., Taxon 24 (5-6): 625 (1975)
subspecies :
subsp. paolina Liverani & Romolini; J. Eur. Orch. 42(3-4): 469 (2010)
Distribution: Italy (N-Mte. Gargano)
13 ITA
Remarks: holosericea agg., holosericea s. str.
Conservation Status: Very rare, known from only a few populations and threatened by grazing
subspecies :
subsp. cyrenaica Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 106 (2007)
Distribution: Libya
20 LBY
Remarks: holosericea agg., group uncertain
Synonyms: hidden
= Ophrys cyrenaica (Kreutz) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
subspecies :
subsp. montiliensis (Aubenas & Scappat.) comb.ined.
Distribution: Italy
13 ITA
Remarks: holosericea agg., group uncertain
Synonyms: hidden
= Ophrys fuciflora ssp. montiliensis Aubenas & Scappat., Bull. Mens. Soc. Linn. Lyon 81(9-10): 297 (2012)
= Ophrys montiliensis (Aubenas & Scappat.) P.Delforge, Naturalistes Belges 93(Orchid. 25): 243 (2012)
subspecies :
subsp. malvasiana (S.Hertel & Weyland) comb.ined.
Distribution: S-Greece (Monemvasia)
13 GRC
Remarks: holosericea agg., group uncertain, possibly also a ssp. of heldreichii
Conservation Status: Extremely rare, only about 80 specimens known
Synonyms: hidden
= Ophrys malvasiana S.Hertel & Weyland, J. Eur. Orch. 41(2-3): 687 (2009)
subspecies :
subsp. vanbruggeniana J.Essink, L.Essink & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 27(2): 220 (2010 publ. 2011)
Distribution: Turkey
34 TUR
Remarks: holosericea agg., group uncertain
Synonyms: hidden
= Ophrys vanbruggeniana (J.Essink, L.Essink & Kreutz) P.Delforge, Naturalistes Belges 96(Orchid. 28): 19 (2015)
subspecies :
subsp. pallidiconi (Faurh.) comb.ined.
Distribution: Turkey (Antalya)
34 TUR
Remarks: holosericea agg., group uncertain
Synonyms: hidden
= Ophrys fuciflora ssp. pallidiconi Faurh., J. Eur. Orch. 43(3): 563 (2011)
species :
Ophrys incubacea Bianca ex Tod.; Orch. Sicul. 75 (1842)
subspecies :
subsp. incubacea
Name: Schwarze Ragwurz
Distribution: Greece; Albania; Slovenia; Croatia; Bosnia & Hercegovina; Sicily; Italy; Sardinia; Corsica; Baleares; Spain; Portugal; France
12 BAL COR FRA POR SAR SPA 13 ALB BIH CRO GRC ITA SIC SLV
Remarks: sphegodes agg., pollinator Andrena morio
Synonyms: hidden
= Arachnites atrata (Rchb.fil.) Tod., Hort. Panor. 2: t. 28 (1875)
= Ophrys incubacea f. albina De Bellard, Calzado, Hervás, J.C.Huertas, Reyes Carr. & Ruíz Cano, Micobot. Jaén 14(1): [27] (2019)
= Ophrys aranifera var. atrata Rchb.fil., Icon. Fl. Germ. Helv. 13-14: 91 (1851)
= Ophrys aranifera ssp. atrata (Rchb.fil.) Arcang., Comp. Fl. Ital. ed. 1: 660 (1882)
= Ophrys aranifera ssp. atrata (Rchb.fil.) E.G.Camus, Monogr. Orchid. 286 (1908) [isonym]
= Ophrys aranifera ssp. incubacea (Bianca) Soó, Monogr. Iconogr. Orchid. Eur. 2: 388 (1938)
= Ophrys aranifera ssp. incubacea (Bianca) E.G.Camus & A.Camus, Iconogr. Orchid. Europe, Text(2): 337 (1928)
= Ophrys atrata Lindl., Bot. Reg. t. (1807) [nom. illeg.]
= Ophrys biceratia Delile ex Nyman, Consp. 697 (1882)
= Ophrys cyanogramma Welw. ex Nyman, Consp. 697 (1882)
= Ophrys sphegodes ssp. atrata (Lindl.) Malag., Acta Phytotax. Barcin., 18: 9 (1976 publ. 1977)
= Ophrys sphegodes ssp. atrata (Arcang.) E.Mayer, Sezn. Prapr. Cvet. Slov. 387 (1952)
variant :
var. dianensis Perazza & Doro; J. Eur. Orchideen 35(4): 726 (2003)
Distribution: Italy; Spain
12 SPA 13 ITA
Remarks: sphegodes agg.
variant :
var. fucifera (Rchb.fil.) Hennecke; Ber. Arbeitskreis. Heimische Orchid. 33(1): 108 (2016)
Distribution:
Synonyms: hidden
= Ophrys aranifera var. fucifera Rchb.fil., Icon. Fl. Germ. Helv. (H. G. L. Reichenbach) 13-14: 89 (1851)
variant :
var. rubriflora (Rivas Goday) F.M.Vázquez; Folia Bot. Extremadur. 3: 297 (2009)
Distribution: Spain
12 SPA
variant :
var. septentrionalis Perazza & R.Lorenz; Orchid. Italia 318 (2013)
Distribution: Italy
13 ITA
subspecies :
subsp. brutia (P.Delforge) Kreutz; Kompend. Eur. Orchid. 102 (2004)
Distribution: S-Italy (Calabria); ?Croatia
13 ITA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys brutia P.Delforge, Naturalistes Belges 84 (Orchid. 16): 74 (-75) (2003)
subspecies :
subsp. molisana (P.Delforge) comb.ined.
Distribution: C-Italy
13 ITA
Remarks: provisionally accepted name, status unverified
Synonyms: hidden
= Ophrys molisana P.Delforge, Naturalistes Belges 96(Orchid. 28): 18 (2015)
subspecies :
subsp. castri-caesaris Van Looken; Liparis 11: 58 (2005)
Distribution: SE-France (Alpes-Maritimes); NW-Italy (Liguria)
12 FRA 13 ITA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys garganica ssp. castri-caesaris (Looken) Kreutz, Ber. Arbeitskreis. Heimische Orchid. 24(1): 171 (2007)
= Ophrys passionis ssp. castri-caesaris (Van Looken) Kreutz, Eurorchis 17: 109 (2005)
= Ophrys ligustica Romolini & Soca, J. Eur. Orch. 43(4): 773-774 (2011), nom. nov. superfl.
subspecies :
subsp. ionica Gennaio, Gargiulo, Medagli & Chetta; GIROS Orch. Spont. Eur. 59(2): 391 (2016) [nothossp.]
Distribution: Italy
13 ITA
Remarks: Ophrys brutia x incubacea
subspecies :
subsp. pacensis F.M.Vázquez; Folia Bot. Extremadur. 3: 302 (2009)
Distribution: Spain
12 SPA
Remarks: sphegodes agg.
species :
Ophrys insectifera L.; Sp. Pl. 948 (1753)
subspecies :
subsp. insectifera
Name: Fliegen-Ragwurz
Distribution: Austria; Belgium; Luxembourg; England; Czech Republic; Slovakia; Denmark; Finland; France; Germany; Ireland; Luxembourg; Switzerland; Liechtenstein; Netherlands; Spain; Hungary; Italy; Serbia & Kosovo; Montenegro; Macedonia; Bosnia & Hercegovina; Slovenia; Croatia; NW-Greece; Norway; Poland; Romania; European Russia (N-European Russia, C-European Russia); Estonia; Lithuania; Latvia; Belarus; Sweden
10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 FRA SPA 13 BIH CRO GRC ITA MNT NMA ROM SLV SRB 14 BLR EST LAT LIT RUS
Remarks: insectifera agg., pollinator Argogorytes mystaceus, A. fargei and A. combinata
Synonyms: hidden
= Arachnites musciflora F.W.Schmidt, Fl. Boem. 1: 75 (1793)
= Epipactis myodes F.W.Schmidt, Mayer, Phys. Aufs. 1: 248 (1791)
= Malaxis myodes Bernh., Syst. Verz. Erf. 315 (1800)
= Ophrys ambusta Picard, Zeitschr. Naturw. 77: 363 (1905)
= Ophrys insectifera ssp. myodes Ehrh., Hann. Mag. 1780 (15): 232 (1780) [nom. illeg.]
= Ophrys insectifera var. myodes L., Sp. Pl. 2: 948 (1753) [pro syn.]
= Ophrys muscaria Lam., Fl. Fr. 3: 515 (1779)
= Ophrys muscifera Huds., Fl. Angl. ed. I, 340 (1762)
= Ophrys musciflora Schrank, Baier. Fl. 242 (1789) [nom. illeg.]
= Ophrys musciflora Moritzi, Fl. Schweiz 503 (1844) [nom. illeg.]
= Ophrys myodes Jacq., Misc. 2: 273 (1781) [nom. inval.]
= Orchis insectifera (L.) Crantz, Stirp. Austr. ed. II, fasc. 6: 481 (1769)
= Orchis muscaria Scop., Fl. Carn. ed. II, 2: 193 (1772)
= Orchis muscifera (Huds.) Salisb., Prod. Stirp. Chap. Allerton 7 (1796)
= Orchis musciflora Schrank, Baier. Fl. 1: 242 (1789)
= Orchis myodes Bernh., Syst. Verz. Erf. 315 (1800) [in syn.]
= Ophrys insectifera var. luteomarginata (L.Lewis) Kreutz, Ber. Arbeitskreis. Heimische Orchid. 24(1): 175 (2007)
= Ophrys insectifera f. luteomarginata L.Lewis, J. Eur. Orch. 38(1): 190 (2006)
subspecies :
subsp. subinsectifera (C.E.Hermos. & Sabando) O.Bolòs & Vigo; Fl. Països Catalans 4: 661 (2001)
Distribution: N-Spain
12 SPA
Remarks: insectifera agg., pollinator Sterictiphora gastrica (Argidae)
Synonyms: hidden
= Ophrys subinsectifera Amardeilh, L´Orchidophile 27 (123): 149-154 (1996) [nom. illeg.]
= Ophrys subinsectifera C.E.Hermos. & Sabando, Estud. Mus. Cienc. Nat. Alava 10-11: 133, 135 (1995-96 publ. 1996)
species :
Ophrys iricolor Desf.; Ann. Mus. Par. 10: 224, t. 13 (1807)
subspecies :
subsp. iricolor
Name: Regenbogen-Ragwurz (DE); Rainbow Bee Orchid (EN)
Distribution: Albania; Croatia; Crete; Greece (incl. Kyklades); European Turkey; Turkey; Lebanon; Syria; East Aegaean Isl. (including Rhodos, Karpathos, Pserimos); Cyprus; Israel
13 ALB CRO GRC KRI TUE 34 CYP EAI ISR LEB SYR TUR
Remarks: fusca agg., iricolor subagg., pollinator Andrena morio
Synonyms: hidden
= Arachnites iricolor (Desf.) Bubani, Fl. Pyrenees IV: 49 (1902)
= Ophrys fusca ssp. iricolor (Desf.) K.Richt., Pl. Eur. 1: 261 (1890)
= Ophrys fusca ssp. iricolor (Desf.) O.Schwarz, Fedde, Repert. 36: 77 (1934)
= Ophrys fusca var. iricolor (Desf.) Rchb., Deutschl. Fl. 13: 95 (1851)
= Ophrys achillis P.Delforge, Naturalistes Belges 94(Orchid. 26): 238 (2013)
subspecies :
subsp. astypalaeica (P.Delforge) Kreutz; Kompend. Eur. Orchid. 103 (2004)
Distribution: Greece (Kyklades: Astypaleia Isl.)
13 GRC
Remarks: fusca agg., iricolor subagg.
Synonyms: hidden
= Ophrys astypalaeica P.Delforge, Natural. Belges 78 (3) (Orchid. 10): 153-176 (1997)
subspecies :
subsp. mesaritica (Paulus, C. & A.Alibertis) Kreutz; Kompend. Eur. Orchid. 104 (2004)
Distribution: S-Crete; NW-Greece (Kefallonia Isl., Lefkada Isl., Korfu); S-Greece (Kythera Isl.)
13 GRC KRI
Remarks: fusca agg., iricolor subagg., pollinator Andrena nigroaenea (Paulus 2014)
Synonyms: hidden
= Ophrys mesaritica Paulus, C. & A.Alibertis, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 22 (4): 774 (1990)
= Ophrys iricolor ssp. mesaritica C. & A.Alibertis, Orch. Kreta 79 (1989) [nom. prov.]
subspecies :
subsp. hospitalis (P.Delforge) Mifsud & L.Lewis; J. Eur. Orch. 45(1): 117 (2013)
Distribution: Malta (incl. Comino Isl.)
13 MLT
Synonyms: hidden
= Ophrys hospitalis P.Delforge, Naturalistes Belges 93(Orchid. 25): 69 (2012)
subspecies :
subsp. vallesiana (Devillers & Devillers-Tersch.) Paulus & Gack; Jahresber. Naturwiss. Ver. Wuppertal 48: 199 (1995)
Distribution: Tunisia; Algeria
20 ALG TUN
Remarks: fusca agg., iricolor subagg.
Synonyms: hidden
= Ophrys vallesiana Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 373 (1994)
subspecies :
subsp. maxima (A.Terracc.) Paulus & Gack; Jahresber. Naturwiss. Ver. Wuppertal 48: 199 (1995), et Journ. Eur. Orchid. 31 (2), 361 (1999)
Distribution: Corsica; Sardinia; Tunisia; S-Italy (Mte. Gargano, to, Brindisi)
12 COR SAR 13 ITA 20 TUN
Remarks: fusca agg., iricolor subagg.
Synonyms: hidden
= Ophrys fusca f. maxima A.Terracc., Bol. Soc. Brot. Ital. 22 (1910)
= Ophrys fusca var. maxima (A.Terracc.) E.G. & A.Camus, (1908)
= Ophrys iricolor ssp. eleonorae (Devillers-Tersch. & Devillers) Paulus & Gack, Jahresber. Naturwiss. Ver. Wuppertal 48: 199 (1995) [comb. illeg., non Ophrys x eleonorae Devillers & Devillers-Tersch. (1994)]
= Ophrys ×eleonorae Devillers-Tersch. & Devillers, Nat. Belg. 72 (3): 100 (1994)
= Ophrys iricolor ssp. eleonorae (Devillers-Tersch. & Devillers) Kreutz, Kompend. Eur. Orchid. 103 (2004)
subspecies :
subsp. lojaconoi (P.Delforge) Kreutz; Kompend. Eur. Orchid. 103 (2004)
Distribution: Italy (Mte. Gargano)
13 ITA
Remarks: fusca agg., iricolor subagg., pollinator Andrena nigroaenea. Probably a synonym of ssp. maxima (eleonorae) (Paulus 2014)
Synonyms: hidden
= Ophrys lojaconoi P.Delforge, Nat. Belg. 76 (3): 289 (1995)
species :
Ophrys isaura Renz & Taubenheim; Orchidee 31 (6): 240 (1980)
Name: Isaurische Ragwurz
Distribution: Turkey (Antalya, Icel)
34 TUR
Remarks: holosericea agg., oestrifera-group
Conservation Status: IUCN E, extremely threatened
Synonyms: hidden
= Ophrys scolopax ssp. isaura (Renz & Taubenheim) H.A.Pedersen & P.J.Cribb, Field Guide Orchids Europe Medit. 303 (2019)
species :
Ophrys kallaikia C.E.Hermos.; Flora Montiber. 71: 134 (2018)
Distribution: Spain
12 SPA
Remarks: provisionally accepted name
species :
Ophrys kassandra Presser, Z.Antonop. & Tenschert; Ber. Arbeitskreis. Heimische Orchid. 35(1): 180 (2018)
Distribution: Greece
13 GRC
Remarks: provisionally accepted name
species :
Ophrys konyana Kreutz & Ruedi Peter; J. Eur. Orch. 39: 71-78 (2007)
Distribution: SW-Turkey (Yeniköy, east of Antalya)
34 TUR
Remarks: reinholdii agg., hybridogenous origin Ophrys lucis x straussii
species :
Ophrys kopetdagensis K.P.Popov & Neshat.; Izv. Akad. Nauk Turkm. SSR, Biol. Nauk, 1982 (4): 17 (1982)
Distribution: Turkmenistan (Kopet Dag)
32 TKM
Remarks: holosericea agg., tetraloniae-group
Conservation Status: extremely rare, known only from one population
species :
Ophrys kotschyi H.Fleischm. & Soó; Notizbl. Bot. Gart. Berlin 9: 908 (1926)
Name: Kotschys Ragwurz
Distribution: Cyprus
34 CYP
Remarks: holosericea agg., oestrifera-group
Conservation Status: IUCN V
Synonyms: hidden
= Ophrys cypria Renz, Fedde, Repert. 27: 202 (1929)
= Ophrys sintenisii ssp. kotschyi (H.Fleischm. & Soó) Soó, (1929)
species :
Ophrys levantina Gölz & H.R.Reinhard; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 17 (3): 461 (1985)
subspecies :
subsp. levantina
Distribution: S-Turkey; Cyprus
34 CYP TUR
Remarks: holosericea agg., bornmuelleri-group, pollinator Eucera sp.
subspecies :
subsp. grandiflora (H.Fleischm. & Soó) Kreutz; Jour. Eur. Orch. 38(1): 127 (2006)
Distribution: S-Turkey; NW-Syria; Lebanon
34 LEB SYR TUR
Remarks: holosericea agg., bornmuelleri-group
Synonyms: hidden
= Ophrys bornmuelleri f. grandiflora H.Fleischm. & Soó, Repert. Spec. Nov. Regni Veg. 24: 26 (1927)
= Ophrys bornmuelleri ssp. grandiflora (H.Fleischm. & Soó) Renz & Taubenheim, Notes Roy. Bot. Gard. Edinb. 41 (2): 276 (1983)
= Ophrys fuciflora ssp. bornmuelleri var.grandiflora (H.Fleischm. & Soó) B. & E.Willing, Orchidee 26 (2): 78 (1975)
= Ophrys holosericea ssp. grandiflora (H.Fleischm. & Soó) Faurh., J. Eur. Orchideen 35(4): 745 (2003)
= Ophrys holosericea ssp. bornmuelleri var.grandiflora (H.Fleischm. & Soó) Landwehr, Wilde Orch. Eur. 2: 472 (1977) [nom. nud.]
= Ophrys fuciflora ssp. grandiflora (H.Fleischm. & Soó) Faurh., Ber. Arbeitskreis. Heimische Orchid. 22(2): 7 (2006)
species :
Ophrys lunulata Parl.; Giorn. Sc. Lett. Sicil. 62: 4 (1838)
Name: Halbmond-Ragwurz
Distribution: Sicily (incl. Isole Lipari); Malta; ?Croatia
13 MLT SIC
Remarks: sphegodes agg., lunulata subagg., pollinator Osmia kohli
Conservation Status: IUCN Ex (Sardegna, Malta), V (Sicilia)
Synonyms: hidden
= Arachnites lunulata (Parl.) Tod., Orch. Sicul. 77 (1842)
= Ophrys aranifera ssp. lunulata (Parl.) E.G.Camus, Bergon & A.Camus, Monogr. Orchid. 289 (1908)
= Ophrys sphegodes ssp. lunulata (Parl.) H.Sund., Eur. Medit. Orchid., ed. 3: 39 (1980)
species :
Ophrys lutea (Gouan) Cav.; Icon. Descr. 2: 46 (1793)
subspecies :
subsp. lutea
Name: Gelbe Ragwurz
Distribution: Malta (incl. Gozo Isl.); Sicily; Italy; +Corsica; Sardinia; Baleares; Spain; Gibraltar; Portugal; France; Algeria
12 BAL COR FRA GIB POR SAR SPA 13 ITA MLT SIC 20 ALG
Remarks: fusca agg., lutea-group, pollinator Andrena cinerea (Sardegna), Andrena senecionis
Synonyms: hidden
= Arachnites lutea (Cav.) Tod., Orch. Sicul. 95 (1842)
= Ophrys glabra Pers., Syn. 2: 510 (1807)
= Ophrys speculum Steud., Nom. ed. II, 2: 219 (1841)
= Ophrys vespifera Brot., Fl. Lusit. 1: 24 (1800 publ. 1804)
= Ophrys vespifera Willd., Sp. Pl. 4: 65 (1805) [nom. superfl.]
= Ophrys lutea Biv., Sicul. Pl. Cent. 2: 41 (1807) [nom. illeg.]
= Ophrys insectifera var. lutea Gouan, Fl. Monsp. 299 (1765)
= Ophrys lutea var. speculissima F.M.Vázquez, J. Eur. Orch. 40(4): 706 (2008)
= Ophrys lutea f. speculissima (F.M.Vázquez) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
subspecies :
subsp. corsica (Soleirol ex G.Foelsche & W.Foelsche) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 176 (2007)
Distribution: Corsica; Sardinia; ?Malta; ?S-France
12 COR SAR
Remarks: fusca agg., lutea-group, pollinator Andrena cinerea (Paulus 2014)
Synonyms: hidden
= Ophrys corsica Soleirol ex G. & W.Foelsche, J. Eur. Orch. 34(4): 823-885 (2002)
= Ophrys phryganae var. corsica (Soleirol ex G. & W.Foelsche) Hennecke, GIROS Orch. Spont. Eur. 60(2): 270 (2017)
subspecies :
subsp. hellenica (Devillers & Devillers-Tersch.) comb.ined.
Distribution: Greece (Kefallonia, Attika, Epirus, Ionian Isl.); S-Italy; Sicily; N-Algeria
13 GRC ITA SIC 20 ALG
Remarks: fusca agg., lutea-group, pollinator Andrena nigroolivacea (Paulus 2014)
Synonyms: hidden
= Ophrys hellenica Devillers & Devillers-Tersch., Naturalistes Belges 94(Orchid. 26): 158 (2013)
= Ophrys phryganae var. hellenica (Devillers & Devillers-Tersch.) Hennecke, GIROS Orch. Spont. Eur. 60(2): 270 (2017)
subspecies :
subsp. melena Renz; Fedde, Repert. 25: 264 (1928) [pro hybr.]
Distribution: Albania; Croatia; Greece (Evvoia, Peloponnisos); East Aegaean Isl.
13 ALB CRO GRC 34 EAI
Remarks: fusca agg., lutea-group
Synonyms: hidden
= Ophrys melena (Renz) Paulus & Gack, Israel J. Bot. 39(1-2): 79 (1990) [nom. inval.]
= Ophrys galilaea ssp. melena (Renz) Del Prete, Webbia 38: 213 (1984)
= Ophrys murbeckii ssp. melena (Renz) Soó, Acta Bot. Acad. Sci. Hung. 5: 440 (1959)
= Ophrys lutea var. melena (Renz) E.Nelson, Monogr. Ophrys 216 (1962) [nom. inval.]
subspecies :
subsp. phryganae (Devillers-Tersch. & Devillers) Melki; Cah. Soc. Fr. Orch. 6 (14e Colloque Peris 1999): 51 (2000)
Name: Phrygana-Ragwurz (DE); Phrygana Bee Orchid (EN)
Distribution: Crete (sensu stricto); Greece; S-Italy; East Aegaean Isl. (Karpathos, Rhodos); ?Croatia
13 GRC ITA KRI 34 EAI
Remarks: fusca agg., lutea-group. Pollinator Andrena panurgimorpha (Paulus 2014) for the typical form; Andrena humilis, A. clypella, A. tadauchii etc. for the sensu-lato-forms
Synonyms: hidden
= Ophrys phryganae Devillers-Tersch. & Devillers, Nat. Belg. 72 (3): 100 (1991)
= Ophrys heracleotica Gavalas, Kreutz & Z.Antonop., Ber. Arbeitskr. Heim. Orch. 27(2): 266-281 (2010)
subspecies :
subsp. penelopeae (Paulus) comb.ined.
Distribution: Greece (Peloponnes, W-Greece, Ionian Isl.); ?East Aegaean Isl. (?Naxos)
13 GRC
Synonyms: hidden
= Ophrys penelopeae Paulus, J. Eur. Orch. 46(2): 293 (2014)
= Ophrys sicula var. penelopeae (Paulus) Hennecke, GIROS Orch. Spont. Eur. 61(1): 208 (2018)
subspecies :
subsp. praemelena (S.Hertel & Presser) comb.ined.
Distribution: NW-Greece (Ionian Isl.: Korfu)
13 GRC
Remarks: fusca agg., lutea-group
Synonyms: hidden
= Ophrys praemelena S.Hertel & Presser, Ber. Arbeitskreis. Heimische Orchid. 27(1): 169 (2010)
subspecies :
subsp. quarteirae Kreutz, M.R.Lowe & Wucherpf.; Jour. Eur. Orch. 39(3/4): 627 (2007)
Distribution: SC-Spain; S-Spain (Algarve); S-Portugal
12 POR SPA
Remarks: fusca agg., lutea-group, pollinator Andrena cf. vetula (Paulus 2014)
Synonyms: hidden
= Ophrys lutea var. quarteirae (Kreutz, M.R.Lowe & Wucherpf.) F.M.Vázquez, J. Eur. Orch. 40(4): 711 (2008)
= Ophrys murbeckii ssp. quarteirae (Kreutz, M.R.Lowe & Wucherpf.) M.R.Lowe, J. Eur. Orch. 43(3): 462 (2011)
= Ophrys quarteirae (Kreutz, M.R.Lowe & Wucherpf.) Devillers & Devillers-Tersch., Naturalistes Belges 94(Orchid. 26): 159 (2013)
subspecies :
subsp. pseudomelena (Turco, Medagli & D´Emerico) comb.ined.
Distribution: Italy (Puglia, Gargano to S-Italy)
13 ITA
Remarks: fusca agg., lutea-group, pollinator Andrena vulpecula (Paulus 2014)
Synonyms: hidden
= Ophrys pseudomelena Turco, Medagli & D´Emerico, GIROS Notiz. 2: 8 (2012)
species :
Ophrys mammosa Desf.; Ann. Mus. Par. 10: 222. t. 12 (1807)
subspecies :
subsp. mammosa
Name: Busen-Ragwurz (DE); Bosom Orchid (EN)
Distribution: Greece (widespread, incl. Kyklades); Crete; East Aegaean Isl. (Samos, Thasos, Paros, Rhodos, Pserimos, etc.); Turkey
13 GRC KRI 34 EAI TUR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys aranifera ssp. mammosa (Desf.) Soó, Notizbl. Bot. Gart. Berlin-Dahlem 9(88): 907 (1926)
= Ophrys doerfleri H.Fleischm., Österr. Bot. Zeitschr. 74: 185 (1925)
= Ophrys mammosa f. falsomammosa (B. & H.Baumann) P.Delforge, Naturalistes Belges 89(Orchid. 21): 251 (2008)
= Ophrys mammosa f. parnassica Vierh., Verh. Zool.-Bot. Ges. Wien 69: 297 (1919) [nom. nud.]
= Ophrys mammosa ssp. falsomammosa B. & H.Baumann, Jour. Eur. Orch. 37(3): 725 (2005)
= Ophrys sphegodes ssp. parnassica (Vierh.) Soó ex J.J.Wood, Orchidee 31 (6): 233 (1980)
= Ophrys sphegodes ssp. mammosa (Desf.) Soó ex E.Nelson, Gestaltwandel 184 (1962)
= Ophrys sphegodes ssp. parnassica (Vierh.) Soó, Acta Bot. Acad. Sci. Hung. 5: 444 (1959)
subspecies :
subsp. ciliciana Kreutz; Ber. Arbeitskreis. Heimische Orchid. 30(2): 275 (2013)
Distribution: Turkey
34 TUR
Synonyms: hidden
= Ophrys ciliciana (Kreutz) P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015)
subspecies :
subsp. cyclocheila (Aver.) Baumann et al.; Jour. Eur. Orch. 35(1): 172 (2003)
Distribution: Azerbaijan; Northern Caucasus; Crimea; Georgia [Caucasus]
14 KRY 33 AZE GEO NCS
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys sphegodes ssp. taurica (Agg.) Soó, Acta Bot. Acad. Sci. Hung. 16(3/4): 383 (1970)
= Ophrys mammosa ssp. taurica (Agg.) Soó, Repert. Spec. Nov. Regni Veg. 85: 446 (1974)
= Ophrys aranifera ssp. taurica (Aggeenko) K.Richt., Pl. Eur. 1: 263 (1890)
= Ophrys aranifera var. taurica Aggeenko, Schrift. St. Peters. Naturf. Ges. 291 (1886-87)
= Ophrys taurica (Agg.) Nevski, Komarov, Fl. URSS 4: 728 (1935)
= Ophrys caucasica ssp. cyclocheila Aver., Bot. Zhurn.79 (10): 124 (1994), et Ber. Arbeitskr. Heim. Orch. 11 (2): 36 (1994 publ. 1995)
= Ophrys cyclocheila (Aver.) P.Delforge, Naturalistes Belges 85 (Orchid. 17): 252 (2004)
= Ophrys cyclocheila Nevski ex Nevski, Fl. URSS 4: 728 (1935) [nom. inval., pro syn.
= Ophrys caucasica f. cyclocheila Nevski, Fl. URSS 4: 728 (1935) [nom. inval., descr. ross.]
= Ophrys araneifera var. cyclocheila (Nevski) Soó, Monogr. Iconogr. Orchid. Eur. 2: 388 (1938) [nom. inval.]
= Ophrys caucasica var. cyclocheila Nevski ex Grossh., Fl. Kavk. 2: 231 (1940) [nom. inval.]
= Ophrys sphegodes f. cyclocheila (Nevski) Soó, Ann. Univ. Scient. Budapest 11: 172 (1969) [nom. inval.]
subspecies :
subsp. epirotica (Renz) H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 725 (2005)
Distribution: NW-Greece; W-Greece; Albania
13 ALB GRC
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys aranifera f. epirotica Renz, Repert. Spec. Nov. Regni Veg. 25: 248 (1928)
= Ophrys mammosa ssp. epirotica (Renz) Gölz & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15 (2): 178 (1983)
= Ophrys epirotica (Renz) Devillers-Tersch. & Devillers, Nat. Belg. 72 (3): 101 (1991)
= Ophrys mammosa ssp. macedonica (H.Fleischm. ex Soó) Kreutz, Kompend. Eur. Orchid. 107 (2004)
= Ophrys aranifera ssp. ×macedonica H. Fleischm. ex Soó, Repert. Spec. Nov. Regni Veg. 26: 279 (1929)
= Ophrys macedonica (H.Fleischm. ex Soó) Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 381 (1994) [pro hybr.]
= Ophrys mammosa ssp. grammosa B. & E.Willing, (1985)
= Ophrys ×macedonica (H.Fleischm. ex Soó) Soó ex Soó, Fedde, Repert. 24: 27 (1927)
subspecies :
subsp. taigetica (Presser & S.Hertel) comb.ined.
Distribution: S-Greece (Peloponnes: Taigetos)
13 GRC
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys taigetica Presser & S.Hertel, Ber. Arbeitskreis. Heimische Orchid. 27(1): 160 (2010)
subspecies :
subsp. zeusii (M.Hirth) comb.ined.
Distribution: Macedonia; N-Greece; Albania; ?Croatia
13 ALB GRC NMA
Remarks: mammosa agg., possibly identical with ssp. macedonica/epirotica. Pollinator Andrena paucisquama (Paulus 2014)
Synonyms: hidden
= Ophrys sphegodes ssp. zeusii (M.Hirth) Kreutz, Kompend. Eur. Orchid. 117 (2004)
= Ophrys zeusii Hirth, Jahresber. Naturwiss. Ver. Wuppertal 55: 163-188 (2002)
subspecies :
subsp. gortynia (H.Baumann & Künkele) H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 726 (2005)
Distribution: Crete; Greece (Kyklades: Paros etc.); East Aegaean Isl. (Naxos etc.)
13 GRC KRI 34 EAI
Remarks: mammosa agg. Pollinator Colletes albomaculatus
Synonyms: hidden
= Ophrys gortynia (Baumann & Künkele) Paulus, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (4): 855 (1988)
= Ophrys sphegodes ssp. gortynia H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 377 (1986)
subspecies :
subsp. hansreinhardii (M.Hirth) comb.ined.
Distribution: Albania; Macedonia; NW-Greece
13 ALB GRC NMA
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys hansreinhardii M.Hirth, Jour. Eur. Orch. 39(3/4): 474 (2007)
subspecies :
subsp. kojurensis (Gölz) comb.ined.
Distribution: Iran
34 IRN
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys kojurensis Gölz, J. Eur. Orch. 38(1): 86 (2006)
subspecies :
subsp. parviflora Kreutz & H.Heitz; Ber. Arbeitskreis. Heimische Orchid. 29(2): 149 (2013)
Distribution: Greece
13 GRC
Synonyms: hidden
= Ophrys chaonica P.Delforge, Naturalistes Belges 96(Orchid. 28): 17 (2015), nom. nov.
subspecies :
subsp. posteria B. & H.Baumann; Jour. Eur. Orch. 37(3): 725 (2005)
Distribution: Cyprus; ?SW-Turkey
34 CYP
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys posteria (B. & H.Baumann) Devillers & Devillers-Tersch., Naturalistes Belges 93(Orchid. 25): 162 (2012)
subspecies :
subsp. serotina B. & E.Willing; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 17 (4): 525 (1985)
Distribution: NW-Greece; Albania
13 ALB GRC
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys hystera Kreutz & Ruedi Peter, J. Eur. Orchid. 30 (1): 128 (prepr. 30 Sept. 1997) [nom. nov. superfl.]
= Ophrys mammosa ssp. leucophthalma (Devillers-Tersch. & Devillers) Kreutz, Kompend. Eur. Orchid. 107 (2004)
= Ophrys leucophthalma Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 380 (1994) [fide Kretzschmar 2002]
subspecies :
subsp. parviflora Kreutz & H.Heitz; Ber. Arbeitskreis. Heimische Orchid. 29(2): 149 (2013)
Distribution: Greece
13 GRC
subspecies :
subsp. ustulata G. & W.Thiele; Jour. Eur. Orch. 39(2): 279 (2007)
Distribution: NW-Greece (Ioannina)
13 GRC
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys ustulata (G.Thiele & W.Thiele) P.Delforge, Naturalistes Belges 96(Orchid. 28): 19 (2015)
subspecies :
subsp. mavromata (A.Alibertis & Triantafyll.) comb.ined.
Distribution: Greece
13 GRC
Remarks: mammosa agg., status unverified
Synonyms: hidden
= Ophrys mavromata A.Alibertis & Triantafyll., J. Eur. Orch. 47(2-4): 288 (2015)
subspecies :
subsp. albanica (Presser) comb.ined.
Distribution: Albania
13 ALB
Remarks: mammosa agg., status unverified
Synonyms: hidden
= Ophrys albanica Presser, Ber. Arbeitskreis. Heimische Orchid. 32(2): 122 (2016)
subspecies :
subsp. prespaensis (M.Hirth & Paulus) comb.ined.
Distribution: Greece
13 GRC
Synonyms: hidden
= Ophrys prespaensis M.Hirth & Paulus, J. Eur. Orch. 49(2): 284 (2017)
subspecies :
subsp. willingii (Paulus & M.Hirth) comb.ined.
Distribution: Greece
13 GRC
Synonyms: hidden
= Ophrys willingii Paulus & M.Hirth, J. Eur. Orch. 49(2): 270 (2017)
species :
Ophrys mavrochila S.Hertel & Presser; Ber. Arbeitskreis. Heimische Orchid. 35(1): 141 (2018)
Distribution: Greece
13 GRC
Remarks: provisionally accepted name
species :
Ophrys oestrifera M.Bieb.; Fl. Taur. Caucasus 2: 369 (1808)
subspecies :
subsp. oestrifera
Name: Bremsen-Ragwurz
Distribution: Crimea
14 KRY
Remarks: holosericea agg., oestrifera-group. In strict sense endemic to Crimea
Synonyms: hidden
= Ophrys scolopax ssp. oestrifera (M.B.) Soó, Acta Bot. Acad. Sci. Hung. 16 (3-4): 386 (1970 publ. 1971)
= Ophrys muscaria Pall., Reise Suedl. Statth. Russ. Reich. 2: 172 [non Scop., nec Lam.]
= Ophrys proteus Radozytski, (1842)
subspecies :
subsp. minuscula (G.Thiele & W.Thiele) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 28(2): 268 (2011)
Distribution: Greece (widespread, incl. Kyklades)
13 GRC
Remarks: holosericea agg., oestrifera-group. The most common taxon of the oestrifera-group in Greece. If oestrifera is taken sensu lato, this would be a synonym. Pollinator Eucera euroa (Paulus 2014)
Synonyms: hidden
= Ophrys cornuta var. minuscula G.Thiele & W.Thiele, J. Eur. Orchideen 34(3): 646 (2002)
= Ophrys cerastes Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid. 17): 234 (2004)
= Ophrys cerastes var. minuscula (G.Thiele & W.Thiele) Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid. 17): 252 (2004)
= Ophrys minuscula (G.Thiele & W.Thiele) Presser & S.Hertel, Ber. Arbeitskreis. Heimische Orchid. 27(1): 185 (2010), nom. inval., et Ber. Arbeitskreis. Heimische Orchid. 29 (1): 245 (2012)
subspecies :
subsp. stavri Kalog., Delipetrou & A.Alibertis; J. Eur. Orch. 44(1): 66 (2012)
Distribution: S-Greece (Peloponnisos, Attiki)
13 GRC
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys stavri (Kalog., Delipetrou & A.Alibertis) P.Delforge, Naturalistes Belges 95: 226 (2014)
subspecies :
subsp. lemnosiana B. & H.Baumann; Jour. Eur. Orch. 37(3): 727 (2005)
Distribution: Greece (Limnos)
13 EAI GRC
Remarks: holosericea agg., oestrifera-group
subspecies :
subsp. nestoris (A.Alibertis & Brütsch) comb.ined.
Distribution: Greece
13 GRC
Remarks: holosericea agg., oestrifera-group. Status uncertain
Synonyms: hidden
= Ophrys scolopax ssp. nestoris A.Alibertis & Brütsch, Orchidophile (Asnières) 191: 285 (2011)
= Ophrys nestoris (A.Alibertis & Brütsch) Presser & S.Hertel, Ber. Arbeitskreis. Heimische Orchid. 35(1): 149 (2018)
subspecies :
subsp. bicornis (Sadler) Kreutz; Kompend. Eur. Orchid. 108 (2004)
Distribution: N-Greece; Hungary; Serbia; Bosnia & Hercegovina; Romania; Bulgaria; ?Russia
11 HUN 13 BIH BUL GRC ROM SRB
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys scolopax var. banatica (Rchb.fil. ex Pantu) Soó, Acta Bot. Acad. Sci. Hung. 16 (3-4): 367 (1970 publ. 1971)
= Ophrys bicornis Sadler, Nendtv., Pl. Quinq. Eccl. (1836) 35
= Ophrys scolopax var. bicornis (Sadler) Nyman
subspecies :
subsp. orphanidea (Saliaris & P.Delforge) comb.ined.
Distribution: East Aegaean Isl. (Chios, Lesvos)
34 EAI
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys orphanidea Saliaris & P.Delforge, Naturalistes Belges 88(Orchid. 20): 177 (2007)
subspecies :
subsp. rhodostephane (Devillers & Devillers-Tersch.) Kreutz; Kompend. Eur. Orchid. 109 (2004)
Distribution: Croatia (Zadar south, Dalmatia); Montenegro; ?Albania; ?Greece (?Corfu)
13 CRO MNT
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys rhodostephane Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid.17): 232 (2004)
subspecies :
subsp. grandiflora (B.Willing & E.Willing) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 177 (2007)
Distribution: NW-Greece (incl. Ionian Isl.); Turkey
13 GRC 34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys cornuta f. crassicornis Renz, Repert. Spec. Nov. Regni Veg. 25: 259 (1928)
= Ophrys crassicornis (Renz) Devillers-Tersch. & Devillers, Naturalistes Belges 85 (Orchid. 17): 234 (2004)
= Ophrys oestrifera var. grandiflora B. & E.Willing, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 17 (4): 528 (1985)
= Ophrys scolopax f. crassicornis (Renz) Soó, Acta Bot. Acad. Sci. Hung. 16 (3-4): 367 (1970 publ. 1971)
= Ophrys scolopax var. grandiflora (B.Willing & E.Willing) Biel, Ber. Arbeitskr. Heim. Orch. 17 (2): 61 (2000)
= Ophrys sepioides Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid. 17): 233 (2004)
= Ophrys dicipulus Valahas, African Plant Sci. 7: 305 (2013)
subspecies :
subsp. leptomera (P.Delforge) Kreutz; Kompend. Eur. Orchid. 109 (2004)
Distribution: Greece (Evvoia, Attica, Peloponnisos, Kyklades: Paros, Naxos)
13 GRC
Remarks: holosericea agg., oestrifera-group. Pollinator Eucera helvola
Synonyms: hidden
= Ophrys leptomera P.Delforge, Nat. Belg. 81 (3): 192 (2000)
subspecies :
subsp. cornuta (Stev.) K.Richt.; Pl. Eur. 1: 264 (1890) [nom. illeg.]
Distribution: Northern Caucasus; Armenia; Turkey
33 ARM NCS 34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys orientalis ssp. cornuta (Stev.) Soó, Monogr. Iconogr. Orchid. Eur. 2: 65, 448 (1931) [nom. illeg.]
= Ophrys cornuta Stev. ex M.Bieb., Fl. Taur. Caucasus 1: 370 (1809), in obs., et Mem. Soc. Nat. Mosc. 2: 175 (1809)
= Ophrys fuciflora ssp. cornuta (Stev.) H.Sund., Europ. Medit. Orchid., ed. 3: 39 (1980)
= Ophrys holosericea ssp. cornuta (Stev.) H.Sund., Taxon 24 (5-6): 625 (1975)
= Ophrys scolopax ssp. cornuta (Stev.) E.G.Camus, Monogr. Orch. Eur. 270 (1908)
= Ophrys scolopax ssp. cornuta Stev., (1819)
subspecies :
subsp. bremifera (Stev.) K.Richt.; Pl. Eur. 1: 264 (1890)
Distribution: East Aegaean Isl. (Nisyros, Tilos, etc.); Turkey; S-Azerbaijan (Talysh); Iran
33 AZE 34 EAI IRN TUR
Remarks: holosericea agg., oestrifera-group. Status of name and taxon disputed. Delforge believes it to be a Caucasian hybrid oestrifera ssp. abchasica x apifera. Other authors consider it as a mere variety of ssp. cornuta
Synonyms: hidden
= Ophrys bremifera Stev. ex M.Bieb., Fl. Taur. Caucasus 1: 370, in obs., et Mem. Soc. Nat. Mosc. 2: 174 (1809)
= Ophrys scolopax ssp. bremifera (Stev.) Biel, Ber. Arbeitskr. Heim. Orch. 16 (1): 54 (1999)
subspecies :
subsp. mycenensis (S.Hertel & Paulus) comb.ined.
Distribution: Greece (EC-Greece, N- & E-Peloponnisos, Evvoia, Skiathos)
13 GRC
Remarks: holosericea agg., oestrifera-group. Pollinator Chalicodoma manicata.
Synonyms: hidden
= Ophrys mycenensis S.Hertel & Paulus, J. Eur. Orch. 42(3-4): 457 (2010)
subspecies :
subsp. cornutula (Paulus) Kreutz; Kompend. Eur. Orchid. 109 (2004)
Name: Kleine Gehörnte Ragwurz (DE); Small Horned Bee Orchid (EN)
Distribution: East Aegaean Isl. (S-Rhodos)
34 EAI
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys cornutula Paulus, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 38-86 (2001)
subspecies :
subsp. chalkae (M.Hirth & H.Spaeth) comb.ined.
Name: Chalki-Ragwurz (DE); Chalki Horned Bee Orchid (EN)
Distribution: East Aegaean Isl. (Chalki)
34 EAI
Remarks: holosericea agg., oestrifera-group. Status unverified
Synonyms: hidden
= Ophrys chalkae M.Hirth & H.Spaeth, J. Eur. Orch. 42(3-4): 573 (2010)
subspecies :
subsp. tili (M.Hirth & H.Spaeth) comb.ined.
Distribution: East Aegaean Isl. (Tilos)
34 EAI
Remarks: holosericea agg., oestrifera-group, Status unverified
Synonyms: hidden
= Ophrys tili M.Hirth & H.Spaeth, J. Eur. Orch. 42(3-4): 574 (2010)
subspecies :
subsp. ellinicaea (Kreutz & Tenschert) comb.ined.
Distribution: East Aegaean Isl. (Kos)
34 EAI
Synonyms: hidden
= Ophrys ellinicaea Kreutz & Tenschert, Ber. Arbeitskreis. Heimische Orchid. 28(2): 295 (2011)
subspecies :
subsp. ceto (Devillers, Devillers-Tersch. & P.Delforge) comb.ined.
Distribution: Greece (Kyklades: Naxos, Paros)
13 GRC
Remarks: holosericea agg., oestrifera-group. Pollinator Eucera euroa
Synonyms: hidden
= Ophrys ceto Devillers, Devillers-Tersch. & P.Delforge, Naturalistes Belges 85 (Orchid. 17): 234 (2004)
subspecies :
subsp. minutula (Gölz & H.R.Reinhard) Kreutz; Kompend. Eur. Orchid. 109 (2004)
Distribution: East Aegaean Isl. (Lesvos, Samos, Chios); W-Turkey
34 EAI TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys scolopax ssp. minutula (Gölz & H.R.Reinhard) Biel, Ber. Arbeitskr. Heim. Orch. 16 (1): 54 (1999)
= Ophrys minutula Gölz & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 21 (4): 1055 (1989)
= Ophrys scolopax var. minutula (Gölz & H.R.Reinhard) H.A.Pedersen & Faurh., Orchidee 48 (6): 235 (1997)
subspecies :
subsp. hygrophila (Gügel, Kreutz, D. & U.Rückbr.) Kreutz; Kompend. Eur. Orchid. 109 (2004)
Distribution: SW-Turkey (Antalya, Konya)
34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys hygrophila Gügel, Kreutz, D. & U.Rückbr., J. Eur. Orchid. 29 (4): 671 (1997)
subspecies :
subsp. phrygia (H.Fleischm. & Bornm.) H.Baumann & Lorenz; Jour. Eur. Orch. 37(3): 727 (2005)
Distribution: S-Turkey (Antalya); East Aegaean Isl. (Chios)
34 EAI TUR
Remarks: holosericea agg., oestrifera-group
Conservation Status: IUCN E
Synonyms: hidden
= Ophrys phrygia H.Fleischm. & Bornm., Ann. Naturhist. Mus. Wien 36: 9 (1923)
= Ophrys scolopax ssp. phrygia (H.Fleischm. & Bornm.) H.A.Pedersen & P.J.Cribb, Field Guide Orchids Europe Medit. 304 (2019)
subspecies :
subsp. kreutzii (W.Hahn, R.Wegener & J.Mast) comb.ined.
Distribution: S-Turkey (Karaman prov.)
34 TUR
Remarks: holosericea agg., oestrifera-group, near ssp. phrygia. Pollinators Eucera nigripes, Eucera squamosa
Synonyms: hidden
= Ophrys kreutzii W.Hahn, R.Wegener & J.Mast, Ber. Arbeitskreis. Heimische Orchid. 25(2): 168 (2008)
subspecies :
subsp. abchasica Kümpel; Ber. Arbeitskr. Heim. Orch. 5 (1­2): 25 (1988)
Distribution: NE-Turkey; Georgia [Caucasus] (Abkhazia, Shiga Kartli, Kartli); Azerbaijan
33 AZE GEO 34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys abchasica (Kümpel) P.Delforge, Nat. Belg. 72 (3): 100 (1991)
subspecies :
subsp. ulupinara (W.Hahn, Passin & R.Wegener) comb.ined.
Distribution: S-Turkey (Antalya)
34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys ulupinara W.Hahn, Passin & R.Wegener, Ber. Arbeitskreis. Heimische Orchid. 25(2): 176 (2008)
subspecies :
subsp. masticorum (P.Delforge & Saliaris) comb.ined.
Distribution: East Aegaean Isl. (Chios, ?Lesvos)
34 EAI
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys masticorum P.Delforge & Saliaris, Naturalistes Belges 88(Orchid. 20): 177 (2007)
subspecies :
subsp. akcakarae Kreutz; Jour. Eur. Orch. 38(1): 161 (2006)
Distribution: SE-Turkey (Anatolia: Diyarbakir, Siirt, Sirnak)
34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys aramaeorum f. akcakarae (Kreutz) P.Delforge, Naturalistes Belges 87(Orchid. 19): 260 (2006)
= Ophrys akcakarae (Kreutz) Kreutz, J. Eur. Orch. 46(2): 455 (2014)
subspecies :
subsp. akcaliensis Perschke; J. Eur. Orch. 40(2): 237 (2008)
Distribution: SE-Turkey (Cilicia)
34 TUR
Remarks: holosericea agg., oestrifera-group
subspecies :
subsp. karadenizensis (M.Schönfelder & H.Schönfelder) Kreutz; Kompend. Eur. Orchid. 109 (2004)
Distribution: N-Turkey
34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys karadenizensis M.Schönfelder & H.Schönfelder, Ber. Arbeitskr. Heim. Orch. 14 (2): 18, 17 (1997 publ. 1998)
subspecies :
subsp. minutiflora J.Essink, L.Essink, Gügel, Kreutz & Zaiss; Ber. Arbeitskreis. Heimische Orchid. 26(2): 53 (2009)
Distribution: Turkey
34 TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys minutiflora (J.Essink, L.Essink, Gügel, Kreutz & Zaiss) P.Delforge, Naturalistes Belges 96(Orchid. 28): 18 (2015)
subspecies :
subsp. elbursiana Kreutz; Ber. Arbeitskreis. Heimische Orchid. 27(2): 227 (2010 publ. 2011)
Distribution: N-Iran
34 IRN
Remarks: holosericea agg., oestrifera-group
subspecies :
subsp. montis-gargani Van de Vijver & W.Van Looken; Journ. Eur. Orch. 42(1): 167-180 (2010)
Distribution: C-Italy (Monte Gargano)
13 ITA
Remarks: holosericea agg., oestrifera-group
subspecies :
subsp. mattinatae (Medagli, A.Rossini, Quitadamo, D´Emerico & Turco) Kreutz; GIROS Orch. Spont. Eur. 58: 6 (2015)
Distribution: C-Italy (Monte Gargano)
13 ITA
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys mattinatae Medagli, A.Rossini, Quitadamo, D´Emerico & Turco, GIROS Notiz. 51: 103 (2012)
subspecies :
subsp. zinsmeisteri (A.Fuchs) Kreutz; Kompend. Eur. Orchid. 110 (2004)
Distribution: SE-Austria; Croatia; ?Slovenia
11 AUT 13 CRO
Remarks: holosericea agg., oestrifera-group. Pollinator Eucera longicornis (Paulus 2014)
Synonyms: hidden
= Ophrys ×zinsmeisteri A.Fuchs, Repert. Spec. Nov. Regni Veg., Beih. 51: 132 (1928)
= Ophrys oestrifera ssp. balcanica Soó ex Hayek, Repert. Spec. Nov. Regni Veg., Sonderbeih. 30(3): 378 (1933)
= Ophrys balcanica Soó, Fedde, Repert. 26: 279 (1929)
= Ophrys kvarneri Perko & Kerschb., Ber. Arbeitskreis. Heimische Orchid. 20(1): 46 (2003)
species :
Ophrys omegaifera H.Fleischm.; Österr. Bot. Zeitschr. 74: 184 (1925)
subspecies :
subsp. omegaifera
Name: Omega-Ragwurz (DE); Omega Bee Orchid (EN)
Distribution: East Aegaean Isl. (widespread, including Rhodos, Karpathos, Chalki, Kastellorizo, Pserimos, Tilos); Crete; Greece (Kyklades); SW-Turkey; Jordania
13 GRC KRI 34 EAI JOR TUR
Remarks: fusca agg., omegaifera-group, Pollinator Anthophora dalmatica (atroalba ssp. agamoides)
Synonyms: hidden
= Ophrys fusca ssp. omegaifera (H.Fleischm.) E.Nelson, Gestaltwandel 209 (1962)
= Ophrys lutea ssp. omegaifera (H.Fleischm.) Soó, Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 906 (1926)
subspecies :
subsp. polycratis (P.Delforge) comb.ined.
Distribution: East Aegaean Isl. (Samos, Chios)
34 EAI
Remarks: fusca agg., omegaifera-group
Synonyms: hidden
= Ophrys polycratis P.Delforge, Naturalistes Belges 89(Orchid. 21): 205 (2008)
subspecies :
subsp. dyris (Maire) Del Prete; Webbia 38: 213 (1984)
Name: Marokkanische Ragwurz
Distribution: Baleares (Mallorca, Ibiza); Spain; S-Portugal; Morocco; Algeria
12 BAL POR SPA 20 ALG MOR
Remarks: fusca agg., omegaifera-group, Pollinator Anthophora atroalba, Anthophora balearica
Synonyms: hidden
= Ophrys atlantica ssp. dyris (Maire) G.Keller, Monogr. Icon. Orch. Eur. 2: 403 (1969)
= Ophrys fusca ssp. dyris (Maire) Soó, Monogr. Iconogr. Orch. Eur. 2: 312 (1935)
= Ophrys dyris Maire, Bull. Soc. Hist. Nat. Afr. Nord 22: 65 (1931)
= Ophrys dyris f. albina De Bellard, Calzado, Hervás, J.C.Huertas, Reyes Carr. & Ruíz Cano, Micobot. Jaén 14(1): [24] (2019)
= Ophrys omegaifera var. dyris (Maire) Soó, Acta Bot. Acad. Sci. Hung. 25 (3-4): 361 (1979 publ. 1980)
= Ophrys fusca var. dyris (Maire) O.Bolòs & Vigo, Fl. Països Catalans 4: 659 (2001)
= Ophrys omegaifera ssp. dyris var.maculata F.M.Vázquez, Jour. Eur. Orch. 40(4), 705 (2008)
subspecies :
subsp. lenae (Lowe & D.Tyteca) comb.ined.
Distribution: Portugal
12 POR
Remarks: fusca agg., omegaifera-group, probably hybridogenous origin O. fusca x O. dyris
Synonyms: hidden
= Ophrys lenae Lowe & D.Tyteca, J. Eur. Orch. 44(1): 213 (2012)
subspecies :
subsp. algarvensis (D.Tyteca, Benito & M.Walravens) Kreutz; Kompend. Eur. Orchid. 110 (2004)
Distribution: S-Portugal; S-Spain
12 POR SPA
Remarks: fusca agg., omegaifera-group, pollinator Colletes albomaculatus
Synonyms: hidden
= Ophrys algarvensis D.Tyteca, Ayuso & Walravens, Jour. Eur. Orch. 35(1): 57-78 (2003)
subspecies :
subsp. fleischmannii (Hayek) Del Prete; Webbia 38: 213 (1984)
Distribution: Crete
13 KRI
Remarks: fusca agg., omegaifera-group, pollinator Anthophora sicheli. Records from Kyklades refer to ssp. israelitica
Synonyms: hidden
= Ophrys funerea ssp. fleischmannii Soó, Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 905 (1926)
= Ophrys fusca ssp. fleischmannii (Hayek) Soó, Notizbl. Bot. Gart. Berlin-Dahlem 9: 905 (1926)
= Ophrys heldreichii H.Fleischm., Österr. Bot. Zeitschr. 74: 182 (1925) [non O. heldreichii Schltr.]
= Ophrys lutea var. fleischmannii (Hayek) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 379 (1973)
= Ophrys fleischmannii Hayek, Fedde, Repert. 22: 388 (1926)
= Ophrys omegaifera var. fleischmannii (Hayek) Soó, Acta Bot. Acad. Sci. Hung. 25 (3-4): 361 (1979 publ. 1980)
subspecies :
subsp. basilissa (C.Alibertis, A.Alibertis & H.R.Reinhard) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 196 (2002)
Name: Große Omega-Ragwurz (DE); Large Omega Bee Orchid (EN)
Distribution: Crete; Greece (Kyklades: Paros, Herakleia, Patmos, Kalymnos, Neros); East Aegaean Isl. (Chios, Kos, Rhodos)
13 GRC KRI 34 EAI
Remarks: fusca agg., omegaifera-group, pollinator Anthophora subterranea (nigrocincta)
Synonyms: hidden
= Ophrys basilissa C.Alibertis, A.Alibertis & H.R.Reinhard, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 22 (1): 201 (1990)
= Ophrys omegaifera var. basilissa (C.Alibertis, A.Alibertis & H.R.Reinhard) Faurh., Orchidee (Hamburg) 53(3): 345 (2002)
subspecies :
subsp. scyria (P.Delforge & Onckelinx) comb.ined.
Distribution: Greece (Skyros Isl.)
13 GRC
Synonyms: hidden
= Ophrys scyria P.Delforge & Onckelinx, Naturalistes Belges 94(Orchid. 26): 238 (2013)
subspecies :
subsp. sitiaca (Paulus, C. & A.Alibertis) Kreutz; Kompend. Eur. Orchid. 111 (2004)
Name: Sitia-Ragwurz (DE); Sitia Bee Orchid (EN)
Distribution: Crete; Greece (Kyklades: Paros, Herakleia, Chios); East Aegaean Isl. (Rhodos, Samos, Phournoi, Arkoi, Patmos, Leros, Kalymnos, Kastellorizo); SW-Turkey
13 GRC KRI 34 EAI TUR
Remarks: fusca agg., omegaifera-group, pollinator Andrena (Melandrena) nigroaenea. Very variable taxon, probably of hybridogenous origin. Belongs possibly in the fusca s. str. group (Paulus 2014)
Synonyms: hidden
= Ophrys israelitica ssp. sitiaca (Paulus, C. & A.Alibertis) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 725 (2005)
= Ophrys sitiaca Paulus, C. & A.Alibertis, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (4): 839 (1988)
subspecies :
subsp. apollonae (Paulus & M.Hirth) comb.ined.
Name: Apollona-Ragwurz (DE); Apollona Bee Orchid (EN)
Distribution: East Aegaean Isl. (Samos, Chios, Nisyros, Rhodos)
34 EAI
Remarks: fusca agg., omegaifera-group, pollinator Anthophora nigriceps. Very early flowering
Synonyms: hidden
= Ophrys apollonae Paulus & M.Hirth, Journ. Eur. Orch. 41(3/4): 647 (2009)
subspecies :
subsp. hayekii (H.Fleischm. & Soó) Kreutz; Kompend. Eur. Orchid. 111 (2004)
Distribution: Sicily; Algeria; +Tunisia
13 SIC 20 ALG TUN
Remarks: fusca agg., omegaifera-group
Synonyms: hidden
= Ophrys atlantica ssp. hayekii (H.Fleischm. & Soó) Soó, Acta Bot. Acad. Sci. Hung. 5: 440 (1959)
= Ophrys fusca ssp. hayekii H.Fleischm. & Soó, Repert. Spec. Nov. Regni Veg. 24: 26 (1927)
= Ophrys hayekii H.Fleischm. & Soó, Repert. Spec. Nov. Regni Veg. 24: 26 (1927 publ. 1928) [nom. nud.]
= Ophrys mirabilis Geniez & Melki, L´Orchidophile 98: 164, 162 (1991)
species :
Ophrys oxyrrhynchos Tod.; Imparziale Giorn. Sc. Lett. Art. (1840) 74
subspecies :
subsp. oxyrrhynchos
Name: Schnabel-Ragwurz
Distribution: Sicily; Malta; S-Italy
13 ITA MLT SIC
Remarks: holosericea agg.
Synonyms: hidden
= Arachnites oxyrrhynchos (Tod.) Tod., Orchid. Sic. 81 (1842) [nom. illeg.]
= Ophrys arachnites ssp. oxyrrhynchos (Tod.) Arcang., Comp. Fl. Ital. ed. 1: 661 (1882)
= Ophrys fuciflora ssp. oxyrrhynchos (Tod.) Soó, Fedde, Repert. 24: 26 (1927)
= Ophrys holosericea ssp. oxyrrhynchos (Tod.) H.Sund., Taxon 24 (5-6): 625 (1975)
= Ophrys nicotrae Zodda, Malpighia 14: 183 (1900)
subspecies :
subsp. biancae (Tod.) Faurh. & H.A.Pedersen; Orchidee (Hamburg) 53(3): 345 (2002)
Name: Biancas Ragwurz
Distribution: Sicily
13 SIC
Remarks: holosericea agg., pollinator Eucera euroa. Hybridogenous origin O. oxyrrhynchos x tenthredinifera
Synonyms: hidden
= Arachnites biancae Tod.1842
= Ophrys biancae Macch., Nuov. Giorn. Bot. Ital. 13: 315 (1881)
= Ophrys discors Bianca, Tod., Orch. Sicul. 84 (1842)
= Ophrys di-stefani Lojac., Fl. Sic. 3: 38 (1909)
= Ophrys ×anapi O. & E.Danesch, Orchid. Eur. Ophrys-Hybr. 227, 137 (1972)
= Ophrys holosericea ssp. biancae (Tod.) Galesi, Cristaudo & Maugeri, J. Eur. Orchideen 36(2): 506 (2004)
= Ophrys fuciflora ssp. biancae (Tod.) Faurh., Ber. Arbeitskreis. Heimische Orchid. 22(2): 7 (2006)
subspecies :
subsp. celiensis (O. & E.Danesch) Del Prete; Webbia 37 (2): 252 (1984)
Distribution: S-Italy
13 ITA
Remarks: holosericea agg., pollinator Eucera graeca. Hybridogen, origin O. apulica x oxyrrhynchos
Synonyms: hidden
= Ophrys holosericea ssp. celiensis (O. & E.Danesch) O. & E.Danesch, Pl. Syst. Evol. 124 (2): 82 (1975)
= Ophrys celiensis (O. & E.Danesch) P.Delforge, Nat. Belg. 81 (4): 397 (2000)
= Ophrys fuciflora ssp. celiensis O. & E.Danesch, Die Orchidee 21(1): 21 (1970)
subspecies :
subsp. lacaitae (Lojac.) Del Prete; Webbia 37 (2): 252 (1984)
Name: Lacaitas Ragwurz
Distribution: Sicily; S-Italy; Croatia (Vis)
13 CRO ITA SIC
Remarks: holosericea agg., pollinator Eucera eucnemidea. Frequently included in O. holosericea
Synonyms: hidden
= Ophrys fuciflora ssp. lacaitae (Lojac.) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 382 (1973)
= Ophrys fuciflora ssp. lacaitae (Lojac.) H.Sund., Europ. Medit. Orchid., ed. 3: 39 (1980)
= Ophrys holosericea ssp. lacaitae (Lojac.) W.Rossi, Inform. Bot. Ital. 13: 201 (1983)
= Ophrys lacaitae Lojac., Fl. Sic. 3: 41 (1909)
subspecies :
subsp. ingrassiae Dura, Turco, Gennaio & Medagli; GIROS Notiz. 46: 37 (-39) (2011)
Distribution: Italy (Apulia)
13 ITA
Remarks: holosericea agg.
species :
Ophrys pallida Raf.; Caratt. 87 (1810)
Name: Bleiche Ragwurz
Distribution: W-Sicily; Malta; Tunisia; E-Algeria
13 MLT SIC 20 ALG TUN
Remarks: fusca agg. Pollinator Andrena orbitalis. A morphologically very distinct taxon which probably deserved rank of valid species.
Synonyms: hidden
= Arachnites pallida (Raf.) Tod., Orch. Sicul. 100 (1842)
= Ophrys fusca ssp. pallida (Raf.) E.G.Camus, Icon. Orch. Eur. 293 (1842)
= Ophrys insectifera var. rosea Desf., (1799)
= Ophrys rosea (Desf.) G.Samp., Terc. Apend. Lista Espec. Herb. Portug. 5 (1914)
= Ophrys rosea Desf. ex Boiss., Fl. Orient. 5: 77 (1884)
= Ophrys fusca ssp. pectus (Mutel) Kreutz, Eurorchis 17: 108 (2005)
= Ophrys pectus Mutel, Ann. Sc. Nat. Ser. 2, 3: 242 (1835)
species :
Ophrys panattensis Scrugli, Cogoni & A.Pessei; Orchidee 43 (5): 225 (1992), pro hybr.
Distribution: Sardinia
12 SAR
Remarks: sphegodes agg., lunulata subagg., pollinator Osmia bicornis (rufa). Hybridogenous origin Ophrys morisii x scolopax
species :
Ophrys promontorii O. & E.Danesch; Orchidee 22 (6): 258 (1971)
Distribution: EC-Italy (Mte. Gargano)
13 ITA
Remarks: sphegodes agg., lunulata subagg., pollinator Osmia mustelina
species :
Ophrys rasinii Soca; J. Eur. Orch. 49(2): 379 (2017)
Distribution: Italy
13 ITA
Remarks: provisionally accepted name
species :
Ophrys regis-ferdinandii (Acht. & Kellerer ex Kuzmanov) Buttler; Jahresber. Naturwiss. Ver. Wuppertal 36: 47 (1983)
Name: König-Ferdinand-Ragwurz (DE); King Ferdinands Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, Symi, Tilos, Samos, Chios, Chalki, Pserimos); SW-Turkey
34 EAI TUR
Remarks: ciliata agg.
Synonyms: hidden
= Ophrys speculum f. regis ferdinandii Acht. & Kellerer ex Renz, Denkschr. Akad. Wiss. Wien, Math. Nat. Cl. 105, 1: 819 (1943)
= Ophrys speculum var. regis-ferdinandii (Acht. & Kellerer ex Renz) Soó, Acta Bot. Acad. Sci. Hung. 5: 439 (1959)
= Ophrys speculum ssp. regis-ferdinandii Acht. & Kellerer, Izv. Carsk. Prirod. Inst. Sofija 12: 206 (1939) [nom. inval.]
= Ophrys speculum ssp. regis-ferdinandii-coburgii (Acht. & Kellerer ex Renz) Acht. & Kellerer ex Kuzmanov, Fragm. Flor. Geobot. 17(2): 194 (1971)
= Ophrys vernixia ssp. regis-ferdinandii (Acht. & Kellerer ex Kuzmanov) Renz & Taubenheim, Notes Roy. Bot. Gard. Edinb. 41 (2): 277 (1983)
= Ophrys ciliata ssp. regis-ferdinandii (Acht. & Kellerer ex Renz) H.Baumann, Künkele & R.Lorenz, J. Eur. Orchideen 36(2): 572 (2004)
species :
Ophrys reinholdii Spruner ex H.Fleischm.; Österr. Bot. Zeitschr. 57: 5, 74 (1908)
subspecies :
subsp. reinholdii
Name: Reinholds Ragwurz (DE); Reinholds Bee Orchid (EN)
Distribution: Macedonia; Albania; W-Greece; S-Greece (incl. Kyklades); NW-Greece (Ionian Isl.: Korfu); East Aegaean Isl. (Samos, Naxos, Lesvos, Nisyros, Tilos, etc., Rhodos); SW-Turkey; Iraq
13 ALB GRC NMA 34 EAI IRQ TUR
Remarks: reinholdii agg., pollinator Eupavlovskia obscura
Synonyms: hidden
= Ophrys mimnolea O.Schwarz, Fedde, Repert. 36: 78 (1934)
= Ophrys naxensis Rech., Ann. Naturhist. Mus. Wien 43: 333 (1929) [nomen]
= Ophrys reinholdii f. albovirescens B. & E.Willing, L´Orchidophile 72: 1099 (1986)
subspecies :
subsp. reinhardiorum (Paulus) comb.ined.
Distribution: NE-Greece; N-Greece (Thessalia)
13 GRC
Remarks: reinholdii agg., pollinator Eupavlovskia funeraria
Synonyms: hidden
= Ophrys reinhardiorum Paulus, Ber. Arbeitskreis. Heimische Orchid. 25(1): 19 (2008)
species :
Ophrys schulzei Bornm. & H.Fleischm.; Mitt. Thüring. Bot. Ver. n. s. 28: 60 (1911)
Name: Schultzes Ragwurz
Distribution: Syria; Northern Caucasus; SE-Turkey (Diyarbakir, Hatay); Iran; Iraq; Lebanon
33 NCS 34 IRN IRQ LEB SYR TUR
Remarks: holosericea agg., oestrifera-group
Synonyms: hidden
= Ophrys luristanica Renz, Orchidee 24 (2): 47 (1973)
= Ophrys schulzei ssp. kurdica Fleischm., [nom. inval.]
species :
Ophrys scolopax Cav.; Icon. Descr. 2: 46 (1793)
subspecies :
subsp. scolopax
Name: Schnepfen-Ragwurz
Distribution: Spain; France; Portugal; Baleares; Algeria
12 BAL FRA POR SPA 20 ALG
Remarks: holosericea agg., scolopax s. str.-group, pollinator Eucera nigrescens and E. interrupta
Synonyms: hidden
= Ophrys asilifera Vayr., Anal. Soc. Esp. Hist. Nat. 9: 98 (1880)
= Ophrys aurita Brot., Phyt. Lusit. 2: 34, t. 91. f. II. (1827)
= Ophrys bullata Cirillo ex Steud., Nom. ed. II, 2: 219 (1841)
= Ophrys fuciflora ssp. scolopax (Cav.) H.Sund., Europ. Medit. Orchid., ed. 3: 39 (1980)
= Ophrys holosericea ssp. scolopax (Cav.) H.Sund., Taxon 24 (5-6): 625 (1975)
= Ophrys nouletii E.G.Camus, (1893)
= Ophrys sancti-martinii Duffort, (1931)
= Ophrys scolopax f. carbajoi (F.M.Vázquez) Hervás, De Bellard, Calzado, J.C.Huertas, Reyes Carr. & Ruíz Cano, Micobot. Jaén 14(1): [26] (2019)
= Ophrys scolopax f. chlorosepala Thell., Allg. Bot. Z. Syst. 20: 61 (1914)
= Ophrys scolopax f. minuta De Bellard, Calzado, Hervás, J.C.Huertas, Reyes Carr. & Ruíz Cano, Micobot. Jaén 14(1): [26] (2019)
= Ophrys scolopax var. chlorosepala (Thell.) Hervás, De Bellard, Calzado, J.C.Huertas, Reyes Carr. & Ruíz Cano, Micobot. Jaén 14(1): [26] (2019)
= Ophrys scolopax var. sepalina F.M.Vázquez, Folia Bot. Extremadur. 3: 306 (2009)
subspecies :
subsp. apiformis (Desf.) Maire & Weiller; Fl. Afr. Nord 6: 260 (1959)
Distribution: Pantelleria Isl.; Algeria; Morocco; Tunisia; ?Sardinia; ?S-Spain; Sicily
13 SIC 20 ALG MOR TUN
Remarks: holosericea agg., scolopax s. str.-group
Synonyms: hidden
= Ophrys sphegifera Willd., Sp. Pl. 4: 65 (1805)
= Ophrys picta var. sphegifera (Willd.) F.M.Vázquez, Folia Bot. Extremadur. 3: 310 (2009)
= Ophrys apiformis (Desf.) Steud., Nom. ed. I, 567 (1821)
= Ophrys fuciflora ssp. apiformis (Desf.) H.Sund., Europ. Medit. Orchid., ed. 3: 39 (1980)
= Ophrys holosericea ssp. apiformis (Desf.) H.Sund., Taxon 24 (5-6): 625 (1975)
= Ophrys insectifera var. apiformis Desf., Fl. Atlant. 2(8): 321 (1799)
= Ophrys ouritensis Guitt., Bull. Soc. Bot. France 264 (1963)
subspecies :
subsp. picta (Link) Kreutz; Kompend. Eur. Orchid. 114 (2004)
Distribution: Spain; Portugal; France; Sardinia; ?Algeria; ?Morocco; ?Tunisia
12 FRA POR SAR SPA
Remarks: holosericea agg., scolopax s. str.-group
Synonyms: hidden
= Ophrys picta Link, Schrad., Jour. für die Bot. 2: 325 (1799)
= Ophrys corniculata Brot., Phyt. Lusit. 2: 38, t. 93 (1827)
= Ophrys scolopax var. picta (Link) Rchb.fil., Icon. Fl. Germ. Helv. 13/14: 98-101 (1851)
subspecies :
subsp. philippei (Gren.) H.A.Pedersen & P.J.Cribb; Field Guide Orchids Europe Medit. 304 (2019)
Distribution: France (Toulon)
12 FRA
Remarks: holosericea agg., tetraloniae-group. Systematic position uncertain, a very distinctive taxon
Conservation Status: Extremely rare, only a few small populations known
Synonyms: hidden
= Ophrys ×philippei Gren., Mem. Soc. Emul. Doubs, ser. 3, 4: 401 (1860), pro hybr.
= Ophrys ×philippei nm.calenae O. & E.Danesch, Orchid. Eur. Ophrys-Hybr. 228, 97 (1972)
= Ophrys ×philippei nm.maurensis O. & E.Danesch, Orchid. Eur. Ophrys-Hybr. 228, 94 (1972)
= Ophrys insectifera ssp. philippei (Gren.) Moggr. & Rchb.fil., Nova Acta Acad. Leop.-Carol. 35: 11 & 12 (1865)
= Ophrys oestrifera ssp. philippei (Gren.) K.Richt., Pl. Eur. 1: 264 (1890)
subspecies :
subsp. corbariensis (J.Samuel & J.-M.Lewin) Kreutz; Eurorchis 17: 109 (2005)
Distribution: France (Aude, E-Corbieres)
12 FRA
Remarks: holosericea agg., scolopax s. str.-group
Synonyms: hidden
= Ophrys corbariensis J.Samuel & J.-M.Lewin, Orchidophile (Asnieres) 154: 254, 251 (2002)
subspecies :
subsp. conradiae (Melki & Deschâtres) H.Baumann, Giotta, R.Lorenz, Künkele & Piccitto; J. Eur. Orchid. 27 (2): 220 (1995)
Distribution: Corsica; Sardinia; ?Tunisia
12 COR SAR
Remarks: holosericea agg., scolopax s. str.-group. Possibly a ssp. of O. holosericea in the tetraloniae-group
Synonyms: hidden
= Ophrys conradiae Melki & Deschâtres, L´Orchidophile 107: 104 (1993)
= Ophrys scolopax ssp. sardoa H.Baumann, Giotta, Künkele, R.Lorenz & Piccitto, J. Eur. Orchid. 27 (2): 222 (1995)
= Ophrys sardoa (H.Baumann, Giotta, R.Lorenz, Künkele & Piccitto) Paulus & Gack, J. Eur. Orchid. 31 (2): 386 (1999)
subspecies :
subsp. santonica (J.M.Mathé & Melki) R.Engel & Quentin; L´Orchidophile (Asnieres) 27 (124): 205 (1996)
Distribution: France (Charente-Maritime, ?Languedoc-Roussillon, ?Provence); N-Spain (Catalunya, Soria, Navarra, Huesca)
12 FRA SPA
Remarks: holosericea agg., scolopax s. str.-group. Possibly a ssp. of O. holosericea in the tetraloniae-group
Synonyms: hidden
= Ophrys santonica J.M.Mathé & Melki, L´Orchidophile 113: 159 (1994) [fide Delforge 2000]
= Ophrys aestivalis J.M.Mathé & Melki, L´Orchidophile 112: 124, 120 (1994) [homonym]
= Ophrys juliana Kerguélen, Bull. Ass. Inform. Bot. 1: 25 (1994)
subspecies :
subsp. vetula (Risso) Kreutz; Kompend. Eur. Orchid. 114 (2004)
Distribution: ?France; Spain; ?NW-Italy; ?Portugal
12 SPA
Remarks: holosericea agg., scolopax s. str.-group. Probably a ssp. of O. holosericea in the tetraloniae-group
Synonyms: hidden
= Ophrys vetula Risso, Fl. Nice 464 (1844)
= Ophrys scolopax var. atropos Barla, Fl. Illus. Nice, Orchid. 71 (1868)
= Ophrys atropos Barla, Icon. Orch. t. 59 (1868)
species :
Ophrys scontusiana Licheri & Biagioli; GIROS Orch. Spont. Eur. 62(2): 387 (2019)
Distribution: Sardinia
12 SAR
Remarks: provisionally accepted name
species :
Ophrys sicula Tineo; Pl. Rar. Sicil. 13 (1846)
Name: Sizilianische Ragwurz (DE); Sicilian Bee Orchid (EN)
Distribution: Albania; Croatia; Crete (incl. Gavdos Isl.); Greece (incl. Kyklades); East Aegaean Isl. (including Rhodos, Karpathos, Kasos, Chalki, Kastellorizo, Pserimos, Tilos); Turkey; Cyprus; Israel; Syria; Lebanon; Sicily; Corsica; Malta (incl. Gozo, Comino Isl.); European Turkey
12 COR 13 ALB CRO GRC KRI MLT SIC TUE 34 CYP EAI ISR LEB SYR TUR
Remarks: fusca agg., lutea-group, pollinator Andrena hesperia
Synonyms: hidden
= Arachnites lutea var. minor Tod., Orchid. Sicul.: 97, tab. 2 (1842)
= Ophrys galilaea H.Fleischm. & Bornm., Ann. Naturhist. Mus. Wien 36: 12 (1923)
= Ophrys lutea ssp. galilaea (H.Fleischm. & Bornm.) Soó, Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 906 (1926)
= Ophrys lutea ssp. minor (Guss.) O. & E.Danesch ex Gölz & H.R.Reinhard, Orchidee 29 (3): 104 (1978), with basionym ref.
= Ophrys lutea ssp. minor (Tod.) O. & E.Danesch, Pl. Syst. Evol. 124 (2): 82 (1975)
= Ophrys lutea ssp. sicula (Tineo) Soldano, Atti Soc. Ital. Sci. Nat. Mus. Civ. Stor. Nat. Milano, 133 (10): 115 (1992 publ. 1993)
= Ophrys lutea var. galilaea (H.Fleischm. & Bornm.) Soó, Acta Bot. Acad. Sci. Hung. 16 (3-4): 390 (1970 publ. 1971) [without basionym ref.]
= Ophrys lutea var. galilaea (H.Fleischm. & Bornm.) Hennecke, Ber. Arbeitskreis. Heimische Orchid. 33(1): 164 (2016)
= Ophrys lutea var. minor (Tod.) Raynaud, Orchid. Maroc 16 (1985)
= Ophrys lutea var. minor (Tod.) Guss., Fl. Sicul. Syn. 2: 550 (1844)
= Ophrys minor ssp. galilaea (H.Fleischm. & Bornm.) Paulus & Gack, Jahresber. Naturwiss. Ver. Wuppertal 39: 66 (1986)
= Ophrys murbeckii Soó, [nec H. Fleischm.]
= Ophrys sicula ssp. galilaea (H.Fleischm. & Bornm.) Paulus & Gack, Jahresber. Naturwiss. Ver. Wuppertal 43: 106 (1990)
= Ophrys sicula var. transadriatica Devillers & Devillers-Tersch., Naturalistes Belges 94(Orchid. 26): 159 (2013)
species :
Ophrys sphegodes Mill.; Gard. Dict. ed. 8, n. 8 (1768)
subspecies :
subsp. sphegodes
Name: Spinnen-Ragwurz
Distribution: Albania; Austria; Belgium; Luxembourg; Baleares; England; Channel Isl.; Bulgaria; Corsica; Slovakia; France; Germany; Switzerland; Spain; Hungary; Italy; Luxembourg; Serbia & Kosovo; Montenegro; Macedonia; Bosnia & Hercegovina; Slovenia; Croatia; Portugal; Romania; Sardinia; Sicily
10 GRB 11 AUT BGM GER HUN LUX SLO SWI 12 BAL CIS COR FRA POR SAR SPA 13 ALB BIH BUL CRO ITA MNT NMA ROM SIC SLV SRB
Remarks: sphegodes agg., pollinator Anthophora atriceps, A. nigroaenea, A. limata
Synonyms: hidden
= Arachnites aranifera (Huds.) Bubani, Fl. Pyren. 4: 48 (1901)
= Myodium araniferum (Huds.) Salisb., Trans. Hort. Soc. 1: 289 (1812)
= Ophrys arachnitis (L.) Mill., Gard. dict. ed. 8, no. 7 (1768)
= Ophrys araneifera var. ruppertii (A.Fuchs) Soó, Feddes Repert. 24: 27 (1927)
= Ophrys aranifera Huds., Fl. Angl. ed. II, 2: 392 (1778)
= Ophrys aranifera ssp. fuchsii (Zimm.) Soó, Feddes Repert. 24: 27 (1927)
= Ophrys aranifera ssp. fucifera Watson, London Cat. Brit. Pl. 21 (1867)
= Ophrys aranifera ssp. gigantea (A.Fuchs) Soó, Repert. Spec. Nov. Regni Veg. 26: 279 (1929)
= Ophrys aranifera ssp. moesziana Soó, Repert. Spec. Nov. Regni Veg. 24: 35 (1927)
= Ophrys aranifera ssp. vierhapperi (Soó) Soó, Monogr. Iconogr. Orchid. Eur. 2: 313, 388 (1935)
= Ophrys aranifera var. fuchsii (Zimm.) Schltr., Monogr. Icon. Orch. Eur. 1: 108 (1926)
= Ophrys aranifera var. vierhapperi Soó, Repert. Spec. Nov. Regni Veg. 24: 35 (1927)
= Ophrys crucigera Jacq., Icon. pl. Rar. 1: 18, t. 185 (1784)
= Ophrys fuchsii W.Zimm., Mitt. Bayer. Bot. Ges. 3: 388 (1917)
= Ophrys fucifera Curtis, Fl. Lond. fasc. 6: t. 67 (1794)
= Ophrys fucifera Sm., Engl. Bot. t. 2649 (1830)
= Ophrys galeopsidea Lag. ex Colmeiro, Enum. Pl. Penins. Hispano-Lusit. 5: 42 (1889)
= Ophrys insectifera ssp. aranifera (Huds.) Moggr. & Rchb.fil., Nova Acta Acad. Leop.-Carol. 35: 9 (1869)
= Ophrys insectifera var. arachnites L., Sp. pl. 2: 949 (1753)
= Ophrys lutea ssp. pseudospeculum (DC.) Kerguélen, Index Synonym. Fl. France (Coll. Patrim. Nat., 8): xv (1993)
= Ophrys mammosa Desf., Ann. Mus. Par. 10: 222, t. 12 (1807)
= Ophrys mammosa ssp. vierhapperi (Soó) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 380 (1973)
= Ophrys pseudaranifera Murr, (1898)
= Ophrys pseudospeculum DC., Fl. Fr. Suppl. 332 (1815)
= Ophrys sphegodes ssp. aranifera (Huds.) O.Schwarz, Mitt. Thüring. Bot. Ges. 1: 92 (1949) [nom. illeg.]
= Ophrys sphegodes ssp. fuchsii (Zimm.) Soó, Acta Bot. Acad. Sci. Hung. 16: 382 (1970)
= Ophrys sphegodes ssp. gigantea (A.Fuchs) O.Schwarz, Mitt. Thüring. Bot. Ges. 1: 92 (1949)
= Ophrys sphegodes ssp. moesziana (Soó) Soó, Acta Bot. Acad. Sci. Hung. 16(3/4): 390 (1970 publ. 1971)
= Ophrys sphegodes ssp. vierhapperi (Soó) Soó, Acta Bot. Acad. Sci. Hung.18: 380 (1973)
= Ophrys sphegodes var. gigantea A.Fuchs, Mitt. Bayer. Bot. Ges. 3: 280 (1917)
= Ophrys sphegodes var. subaesculapiana Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 282 (1986)
= Ophrys sphegodes var. subspruneriana Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 282 (1986)
= Ophrys sphegodes var. subtommasiniana Balayer, Bull. Soc. Bot. Fr., Lett. Bot. 133 (3): 282 (1986)
= Ophrys vindelica W.Zimm. ex A.Fuchs, Repert. Spec. Nov. Regni Veg., Beih. 51: 136 (1928)
= Ophrys ×ruppertii A.Fuchs, Ber. Bayer. Bot. Ges. 16: 76 (1917), et Repert. Spec. Nov. Regni Veg., Beih. 51: 134 (1928)
= Ophrys ×todaroana Macch., Nuov. Giorn. Bot. Ital. 13: 314 (1881)
subspecies :
subsp. liburnica (Devillers & Devillers-Tersch.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 180 (2007)
Distribution: Croatia; Montenegro
13 CRO MNT
Remarks: sphegodes agg., pollinator Andrena nigroaenea (Paulus 2014). Probably only a variant of the nominotypical subspecies.
Synonyms: hidden
= Ophrys liburnica Devillers & Devillers-Tersch., Naturalistes Belges 85 (Orchid. 17): 147 (2004)
subspecies :
subsp. grammica (B.Willing & E.Willing) Kreutz; Kompend. Eur. Orchid. 116 (2004)
Distribution: N-Greece; W-Greece; Macedonia; Albania; S-Croatia; Crete
13 ALB CRO GRC KRI NMA
Remarks: sphegodes agg., pollinator Andrena nigroaenea (Paulus 2014)
Synonyms: hidden
= Ophrys grammica (B.Willing & E.Willing) Devillers-Tersch. & Devillers, Nat. Belg. 72 (3): 101 (1991)
= Ophrys mammosa ssp. grammica B. & E.Willing, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 17 (4): 523 (1985)
= Ophrys aranifera f. oodicheila Renz, Repert. Spec. Nov. Regni Veg. 25: 248 (1928)
= Ophrys sphegodes ssp. oodicheila (Renz) Riech., Jour. Eur. Orch. 36(2): 533 (2004)
= Ophrys cretensis ssp. samica A.Alibertis, J. Eur. Orch. 43(4): 753-758 (2011)
subspecies :
subsp. herae (M.Hirth & H.Spaeth) Kreutz; Kompend. Eur. Orchid. 116 (2004)
Distribution: East Aegaean Isl. (Samos); N-Turkey; E-Greece; Albania; Croatia
13 ALB CRO GRC 34 EAI TUR
Remarks: sphegodes agg., pollinator Andrena thoracica
Synonyms: hidden
= Ophrys herae M.Hirth & H.Spaeth, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 24 (1): 6, 3 (1992)
= Ophrys ×pseudomammosa Renz, Fedde, Repert. 25: 262 (1928)
subspecies :
subsp. janrenzii (M.Hirth) Kreutz; Orchidee (Hamburg) 57(1): 101 (2006)
Distribution: Greece (Korfu)
13 GRC
Remarks: sphegodes agg., pollinator Andrena thoracica
Synonyms: hidden
= Ophrys herae f. janrenzii (M.Hirth) P.Delforge, Naturalistes Belges 89(Orchid. 21): 251 (2008)
= Ophrys janrenzii M.Hirth, Jahresber. Naturwiss. Vereins Wuppertal 55: 171 (2002)
= Ophrys mammosa ssp. janrenzii (M.Hirth) Kreutz, Ber. Arbeitskreis. Heimische Orchid. 24(1): 177 (2007)
= Ophrys herae ssp. janrenzii (M.Hirth) M.Hirth, J. Eur. Orch. 37(1): 176 (2005)
subspecies :
subsp. osmaniaca (Kreutz) comb.ined.
Distribution: Turkey
34 TUR
Synonyms: hidden
= Ophrys herae ssp. osmaniaca Kreutz, Ber. Arbeitskreis. Heimische Orchid. 28(2): 279 (2011)
= Ophrys osmaniaca (Kreutz) P.Delforge, Naturalistes Belges 96(Orchid. 28): 18 (2015)
subspecies :
subsp. caucasica (Woronow ex Grossh.) Soó; Ann. Univ. Sci. Budap., Sect. Biol. 11: 72 (1969)
Distribution: Georgia [Caucasus] (Abkhazia); Armenia; NE-Turkey
33 ARM GEO 34 TUR
Remarks: sphegodes agg.
Conservation Status: IUCN I
Synonyms: hidden
= Ophrys aranifera ssp. caucasica (Woronow) Soó, Monogr. Iconogr. Orchid. Eur. 2: 388 (1938)
= Ophrys caucasica Woronow, Act. Hort. Tiflis 8: III. 70 (1908) ex Grossh., Fl. Kavkaza 1: 261 (1928)
= Ophrys mammosa ssp. caucasica (Woronow ex Grossh.) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 383 (1973)
subspecies :
subsp. classica (Devillers-Tersch. & Devillers) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 179 (2007)
Distribution: C-Italy; ?Croatia
13 ITA
Remarks: sphegodes agg., possibly also to O. exaltata / arachnitiformis
Synonyms: hidden
= Ophrys classica Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 351 (2000)
subspecies :
subsp. kojurensis (Gölz) comb.ined.
Distribution: N-Iran
34 IRN
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys kojurensis Gölz, Jour. Eur. Orch. 38(1): 85 (2006)
subspecies :
subsp. knossia (A.Alibertis) comb.ined.
Distribution: Crete
13 KRI
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys grammica ssp. knossia A.Alibertis, Orchidophile (Asnières) 191: 285 (fig. 1) (2011)
= Ophrys knossia (A.Alibertis) P.Delforge, Naturalistes Belges 93(Orchid. 25): 242 (2012)
subspecies :
subsp. grassoana Cristaudo et al.; Journ. Eur. Orch. 42(1): 181-200 (2010)
Distribution: Sicily (Mt. Etna)
13 SIC
Remarks: sphegodes agg.
subspecies :
subsp. catalcana Kreutz; Ber. Arbeitskreis. Heimische Orchid. 27(2): 231 (2010 publ. 2011)
Distribution: European Turkey
13 TUE
Remarks: sphegodes agg.
subspecies :
subsp. jeanpertii (E.G.Camus) Del Prete & Conte; Webbia 34 (2): 560 (1980) [nothossp.]
Distribution: France; Austria
11 AUT 12 FRA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys speculum ssp. jeanpertii (E.G.Camus) Del Prete & Conte, Webbia 34 (2): 560 (1980)
= Ophrys sphegodes nssp. jeanpertii (E.G.Camus) J.Duvign., Nouv. Fl. Belgique, Luxembourg, N. France & Rég. vois. (ed. 3): 922 (1983)
= Ophrys ×jeanpertii E.G.Camus, Bull. Soc. Bot. Fr. 38: 40, 41 (1891)
= Ophrys ×paulii A.Fuchs, Repert. Spec. Nov. Regni Veg., Beih. 51: 132 (1928)
subspecies :
subsp. massiliensis (Viglione & Véla) Kreutz; Kompend. Eur. Orchid. 116 (2004)
Distribution: SE-France; NW-Italy
12 FRA 13 ITA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys massiliensis Viglione & Véla, L´Orchidophile 135: 13 (1999)
= Ophrys aranifera ssp. massiliensis (Viglione & Véla) Véla, Candollea 62(1): 120 (2007)
subspecies :
subsp. melitensis Salk.; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 24 (4): 633, 640 (1992)
Distribution: Malta (incl. Gozo, Comino Isl.)
13 MLT
Remarks: sphegodes agg., pollinator Chalicodoma sicula
Synonyms: hidden
= Ophrys melitensis (Salk.) Devillers-Tersch. & Devillers, Nat. Belg. 75 ("Orchidees" no. 7, suppl.): 380 (1994)
subspecies :
subsp. montenegrina H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (3): 623 (1988)
Distribution: Montenegro; S-Croatia (incl. Dalmatia)
13 CRO MNT
Remarks: sphegodes agg. Possibly the same as O. herae ssp. janrenzii (Paulus 2014)
Synonyms: hidden
= Ophrys montenegrina (H.Baumann & Künkele) Devillers-Tersch. & Devillers, Nat. Belg. 72 (3): 101 (1991)
subspecies :
subsp. nicolae (Q.G.Manni) comb.ined.
Distribution: S-Italy (Apulia)
13 ITA
Synonyms: hidden
= Ophrys nicolae Q.G.Manni, J. Eur. Orch. 48(2-4): 288 (2016)
subspecies :
subsp. metaurensis (Klaver & Kreutz) comb.ined.
Distribution: Italy (Pesaro, Urbino)
13 ITA
Synonyms: hidden
= Ophrys metaurensis Klaver & Kreutz, GIROS Orch. Spont. Eur. 63(1): 139 (2020)
subspecies :
subsp. panormitana (Tod.) E.Nelson; Monogr. Ophrys 194 (1962)
Distribution: Sicily
13 SIC
Remarks: sphegodes agg., pollinator Andrena thoracica, A. sabulosa
Synonyms: hidden
= Arachnites fuciflora var. panormitana Tod., Orchid. Sicul. 75 (1842)
= Ophrys panormitana (Tod.) Soó, Acta Bot. Acad. Sci. Hung. 18 (3-4): 384 (1973)
= Ophrys sphegodes ssp. panormitana (Tod.) Kreutz, Kompend. Eur. Orchid. 116 (2004), isonym
= Ophrys sphegodes ssp. sicula E.Nelson ex Soó, Journ. Linn. Soc. 76: 367 (1978)
= Ophrys spruneri ssp. panormitana (Tod.) Soó, Acta Bot. Acad. Sci. Hung. 16: 383 (1970)
= Ophrys exaltata ssp. panormitana (Tod.) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 720 (2005)
= Ophrys aranifera ssp. panormitana (Tod.) Véla & R.Martin, Étude Cartogr. Prélim. Orchid. Sicile 91 (2015)
subspecies :
subsp. praecox Corrias; Boll. Soc. Sarda Sci. Nat., 22: 325 (1983)
Distribution: Sardinia; Corsica; ?Croatia
12 COR SAR
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys aranifera ssp. praecox (Corrias) Véla, Orchidophile (Asnières) 176: 38 (2008)
= Ophrys panormitana ssp. praecox (Corrias) Paulus & Gack, Jahresber. Naturwiss. Ver. Wuppertal 48: 204 (1995)
= Ophrys praecox (Corrias) Devillers-Tersch. & Devillers, Nat. Belg. 69 (2): 100 (1988)
= Ophrys panormitana var. praecox (Corrias) P.Delforge, Nat. Belg. 81 (4): 398 (2000)
subspecies :
subsp. tarquinia (P.Delforge) Kreutz; Kompend. Eur. Orchid. 117 (2004)
Distribution: Italy (Tuscany)
13 ITA
Remarks: sphegodes agg., pollinator Andrena tibialis. Possibly to O. exaltata / arachnitiformis
Synonyms: hidden
= Ophrys tarquinia P.Delforge, Nat. Belg. 81 (3): 84 (2000)
subspecies :
subsp. provincialis H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (3): 617 (1988)
Distribution: SE-France; ?NW-Italy
12 FRA
Remarks: sphegodes agg., provincialis-group, pollinator Anthophora atriceps
Synonyms: hidden
= Ophrys provincialis (Baumann & Künkele) Paulus, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20 (4): 855 (1988)
= Ophrys sphegodes ssp. provincialis Nelson, Monogr. Ophrys 197 (1962) [nom. inval.]
= Ophrys sphegodes var. provincialis (H.Baumann & Künkele) P.J.Cribb, Field Guide Orchids Europe Medit. 241 (2019)
subspecies :
subsp. argensonensis (J.-C.Guérin & A.Merlet) Kreutz; Kompend. Eur. Orchid. 115 (2004)
Distribution: France (Charente-Maritime)
12 FRA
Remarks: sphegodes agg., provincialis-group
Synonyms: hidden
= Ophrys argensonensis Guérin & Merlet, L´Orchidophile (133) 169-173 (1998)
subspecies :
subsp. suboccidentalis (J.-P.Ring, J.-C.Querré & Y.Wilcox) comb.ined.
Distribution: France
12 FRA
Remarks: sphegodes agg.
Synonyms: hidden
= Ophrys suboccidentalis J.-P.Ring, J.-C.Querré & Y.Wilcox, Orchidophile (Asnières) 215: 385 (2017)
subspecies :
subsp. taurica (Aggeenko) Soó ex Niketic & Djordjevic; Bull. Nat. Hist. Mus. Belgrade 11: 106 (2018)
Distribution:
Synonyms: hidden
= Ophrys aranifera var. taurica Aggeenko, Bot. Zap. 1(2): 291 (1887)
species :
Ophrys spruneri Nyman; Consp. 698 (1882)
subspecies :
subsp. spruneri
Name: Spruners Ragwurz
Distribution: Greece (except North, but incl. Ionian Isl.); Crete; East Aegaean Isl. (Chios, Ikaria)
13 GRC KRI 34 EAI
Remarks: mammosa agg., pollinator Xylocopa iris
Synonyms: hidden
= Ophrys corinthiaca Hausskn., Mitt. Thüring. Bot. Ver. n. s. 13-14: 25 (1899)
= Ophrys delphica H.Fleischm. ex Hayek, Fedde, Repert. 22: 389 (1926)
= Ophrys ferrum-equinum ssp. spruneri (Nyman) E.G.Camus, Bergon & A.Camus, Monogr. orchid. 274 (1908)
= Ophrys galactostictos Heldr. & Sart ex Boiss., Fl. Orient. 5: 79 (1884)
= Ophrys hiulca Spruner ex Rchb.fil., Icon. Fl. Germ. 13: 93, t. 101. f. 2 (1851)
= Ophrys reinholdi Spruner ex Boiss., Fl. Orient. 5: 79 (1884)
= Ophrys sphaciotica H.Fleischm., Österr. Bot. Zeitschr. 74: 186 (1925)
= Ophrys sphegodes ssp. spruneri (Nyman) E.Nelson, Monogr. Ophrys 183 (1962) [nom. inval.]
subspecies :
subsp. grigoriana (G. & H.Kretzschmar) H.Kretzschmar; Orchideen auf Kreta, Kasos, Karpathos 236 (2002)
Distribution: Crete
13 KRI
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys grigoriana G. & H.Kretzschmar, Ber. Arbeitskr. Heim. Orch. 12 (1): 56 (1995)
species :
Ophrys straussii H.Fleischm. & Bornm.; Ann. Naturhist. Mus. Wien 36: 13 (1923)
subspecies :
subsp. straussii
Distribution: Turkey (S-Anatolia, SE-Anatolia); N-Iraq; Iran
34 IRN IRQ TUR
Remarks: reinholdii agg.
Conservation Status: IUCN E
Synonyms: hidden
= Ophrys straussii H.Fleischm., Beih. Bot. Centralbl. 33(2): 204 (1915)
= Ophrys reinholdii ssp. straussii (H.Fleischm. & Bornm.) E.Nelson, Monogr. Ophrys 149 (1962)
variant :
var. leucotaenia (Renz & Taubenheim) R.Peter; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 24 (4): 575 (1992)
Distribution: Turkey (S-Anatolia, SE-Anatolia); Iraq; Iran
34 IRN IRQ TUR
Remarks: reinholdii agg.
Synonyms: hidden
= Ophrys reinholdii ssp. leucotaenia Renz & Taubenheim, Orchidee 31 (6): 238 (1980)
= Ophrys straussii f. leucotaenia (Renz & Taubenheim) P.Delforge, Naturalistes Belges 90(Orchid.22): 327 (2009)
subspecies :
subsp. antiochiana (H.Baumann & Künkele) Kreutz; Kompend. Eur. Orchid. 117 (2004)
Distribution: S-Turkey (Ziyaret Mts.); Syria
34 SYR TUR
Remarks: reinholdii agg., hybridogenous origin between O. straussii x O. transhyrcana ssp. amanensis
Synonyms: hidden
= Ophrys ×antiochiana H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 527 (1986) [pro hybr.
= Ophrys reinholdii ssp. antiochiana (H.Baumann & Künkele) H.Baumann & Lorenz, Jour. Eur. Orch. 37(3): 728 (2005)
species :
Ophrys subfusca (Rchb.fil.) Hausskn.; Mitt. Thüring. Bot. Vereins, N. F.13/14: 25 (1899)
subspecies :
subsp. subfusca
Distribution: N-Tunisia; NE-Algeria
20 ALG TUN
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys bilunulata ssp. subfusca (Rchb.f.) Paulus, J. Eur. Orch. 52(2-4): 372 (2020)
= Ophrys fusca ssp. numida (Devillers-Tersch. & Devillers) Kreutz, Orchidee (Hamburg) 57(1): 103 (2006) (fide Rebbas & Vela 2013)
= Ophrys fusca ssp. subfusca (Rchb.fil.) E.G. & A.Camus, Iconogr. orchid. Europe, Text(1): 292 (1928)
= Ophrys lutea ssp. murbeckii (H.Fleischm.) Soó, Fedde, Repert. 24: 25 (1927)
= Ophrys lutea ssp. numida (Devillers-Tersch. & Devillers) Kreutz, Kompend. Eur. Orchid. 106 (2004) (fide Rebbas & Vela 2013)
= Ophrys lutea ssp. subfusca (Rchb.fil.) Murb., Acta Reg. Soc. Physiogr. Lund 10: 21 (1899)
= Ophrys lutea var. subfusca Rchb.fil., Icon. Fl. Germ. Helv. 13/14: 76 (1851)
= Ophrys numida Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 297 (2000) (fide Rebbas & Vela 2013)
= Orchis subfusca Murb., Contrib. Fl. Nord-Ouest Afr. Ser. I(3): 21 (1899)
subspecies :
subsp. battandieri (E.G.Camus) Kreutz; Orchidee (Hamburg) 57(1): 102 (2006)
Distribution: Algeria; Tunisia; Morocco
20 ALG MOR TUN
Remarks: fusca agg., subfusca-group, pollinator Andrena vetula (Paulus 2014)
Synonyms: hidden
= Ophrys ×battandieri E.G.Camus, Monogr. Orch. 307 (1908)
= Ophrys murbeckii H.Fleischm., Österr. Bot. Zeitschr. 74: 183 (1925)
= Ophrys galilaea ssp. murbeckii (H.Fleischm.) Del Prete, Webbia 38: 213 (1984)
= Ophrys ×gauthieri Lièvre, Bull. Soc. Hist. Nat. Afr. Nord 13: 196 (1922)
= Ophrys ×lucronii Benito, Estud. Mus. Cienc. Nat. Álava 18-19: 103 (2004)
= Ophrys ×gauthieri nssp. lucronii (Benito) F.M.Vázquez, Folia Bot. Extremadur. 3: 267 (2009)
= Ophrys ×pouyannei Maire, Bull. Soc. Hist. Nat. Afr. Nord 22: 315 (1931)
= Ophrys ×gauthieri nssp. fenarolii (Ferlan) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 465 (1986)
= Ophrys ×subfusca nssp. fenarolii (Ferlan) Del Prete, Webbia 37 (2): 255 (1984)
= Ophrys ×fenarolii Ferlan, Phyton 6: 214 (1956)
= Ophrys lutea ssp. battandieri (E.G.Camus) Kreutz, Kompend. Eur. Orchid. 105 (2004)
subspecies :
subsp. carpitana (M.R.Lowe, Gügel & Kreutz) comb.ined.
Distribution: Tunisia (Cap Bon)
20 TUN
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys carpitana M.R.Lowe, Gügel & Kreutz, Jour. Eur. Orch. 39(3/4): 638 (2007)
= Ophrys pectus auct.et sensu Delforge, nec Mutel
subspecies :
subsp. aspea (Devillers-Tersch. & Devillers) Kreutz; Orchidee (Hamburg) 57(1): 102 (2006)
Distribution: Tunisia (Cap Bon); Libya
20 LBY TUN
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys aspea Devillers-Tersch. & Devillers, Nat. Belg. 81 (3): 297 (2000)
= Ophrys lutea ssp. aspea (Devillers-Tersch. & Devillers) Faurh., Ber. Arbeitskreis. Heimische Orchid. 20(1): 83 (2003), comb. inval.
= Ophrys lutea ssp. aspea (Devillers-Tersch. & Devillers) Faurh., J. Eur. Orch. 41(1): 194 (2009)
subspecies :
subsp. lucentina (P.Delforge) Kreutz; Orchidee (Hamburg) 57(1): 103 (2006)
Distribution: SW-France; Spain (Alicante)
12 FRA SPA
Remarks: fusca agg., subfusca-group. Pollinator Andrena vulpecula
Synonyms: hidden
= Ophrys lucentina P.Delforge, Nat. Belg. 80 (3): 253 (1999)
= Ophrys fusca ssp. lucentina (P.Delforge) Kreutz, Kompend. Eur. Orchid. 94 (2004)
subspecies :
subsp. laurensis (Geniez & Melki) Kreutz; Orchidee (Hamburg) 57(1): 102 (2006)
Distribution: E-Sicily (Mte. Lauro)
13 SIC
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys laurensis Geniez & Melki, Actes Coll. S. F. O. 12: 14-15 (1991 publ. 1992)
= Ophrys lutea ssp. laurensis (Geniez & Melki) Kreutz, Kompend. Eur. Orchid. 105 (2004)
= Ophrys sicula var. laurensis (Geniez & Melki) Hennecke, GIROS Orch. Spont. Eur. 61(1): 200 (2018)
subspecies :
subsp. flammeola (P.Delforge) Kreutz; Orchidee (Hamburg) 57(1): 102 (2006)
Distribution: Sicily
13 SIC
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys flammeola P.Delforge, Nat. Belg. 81 (3): 256 (2000)
= Ophrys fusca ssp. flammeola (P.Delforge) Kreutz, Kompend. Eur. Orchid. 94 (2004)
= Ophrys sicula var. flammeola (P.Delforge) Hennecke, GIROS Orch. Spont. Eur. 61(1): 200 (2018)
subspecies :
subsp. archimedea (P.Delforge & M.Walravens) Kreutz; Orchidee (Hamburg) 57(1): 101 (2006)
Distribution: Sicily
13 SIC
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys archimedea P.Delforge & M.Walravens, Nat. Belg. 81 (3): 256 (2000)
= Ophrys lutea ssp. archimedea (P.Delforge & M.Walravens) Kreutz, Kompend. Eur. Orchid. 105 (2004)
= Ophrys sicula var. archimedea (P.Delforge & M.Walravens) Hennecke, GIROS Orch. Spont. Eur. 61(1): 199 (2018)
subspecies :
subsp. liveranii Orrù & M.P.Grasso; GIROS Notiz. 30: 9 (2005)
Distribution: C-Sardinia
12 SAR
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys lepida S. & J.-M.Moingeon, L´Orchidophile 166: 173-178 (2005)
= Ophrys subfusca ssp. lepida (S.Moingeon & J.-M.Moingeon) Kreutz, GIROS Orch. Spont. Eur. 58: 7 (2015)
= Ophrys sicula var. lepida (S.Moingeon & J.-M.Moingeon) Hennecke, GIROS Orch. Spont. Eur. 61(1): 200 (2018)
= Ophrys sicula var. liveranii (Orrù & M.P.Grasso) Hennecke, GIROS Orch. Spont. Eur. 61(1): 200 (2018)
subspecies :
subsp. pintoi (Lowe & D.Tyteca) comb.ined.
Distribution: Portugal
12 POR
Remarks: fusca agg., subfusca-group
Synonyms: hidden
= Ophrys pintoi Lowe & D.Tyteca, J. Eur. Orch. 44(1): 211 (2012)
subspecies :
subsp. blitopertha (Paulus & Gack) Kreutz; Orchidee (Hamburg) 57(1): 102 (2006)
Name: Käfer-Ragwurz (DE); Beetle Orchid (EN)
Distribution: East Aegaean Isl. (Lesvos, Samos, Naxos, Chios, Rhodos, Pserimos, Tilos); SW-Turkey; Greece (Kyklades)
13 GRC 34 EAI TUR
Remarks: fusca agg., blithopertha-group. Pollinator Blitopertha lineolata (Coleoptera)
Synonyms: hidden
= Ophrys blitopertha Paulus & Gack, J. Eur. Orchid. 30 (1): 181 (1998)
= Ophrys fusca ssp. blitopertha (Paulus & Gack) Faurh. & H.A.Pedersen, Orchidee (Hamburg) 53(3): 345 (2002)
subspecies :
subsp. persephonae (Paulus) Kreutz; Orchidee (Hamburg) 57(1): 103 (2006)
Name: Persephone-Ragwurz (DE); Persephone Bee Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, ?Chios, ?Samos); SW-Turkey
34 EAI TUR
Remarks: fusca agg., blithopertha-group. Status uncertain, possibly a synonym of ssp. blitopertha
Synonyms: hidden
= Ophrys persephonae Paulus, Ber. Arbeitskrs. Heim. Orchid. 18 (1): 38-86 (2001)
= Ophrys fusca ssp. persephonae (Paulus) Kreutz, Eurorchis 17: 108 (2005)
subspecies :
subsp. urteae (Paulus) comb.ined.
Distribution: Turkey (Antalya)
34 TUR
Remarks: fusca agg., blithopertha-group, pollinator Blitopertha nigripennis (Coleoptera)
Synonyms: hidden
= Ophrys urteae Paulus, Ber. Arbeitskreis. Heimische Orchid. 26(2): 10 (2009)
= Ophrys sicula var. urteae (Paulus) Hennecke, GIROS Orch. Spont. Eur. 61(1): 208 (2018)
species :
Ophrys tarentina Gölz & H.R.Reinhard; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 14 (1): 25 (1982)
Name: Tarenter Ragwurz
Distribution: S-Italy
13 ITA
Remarks: sphegodes agg., lunulata subagg., pollinator Osmia tricornis
species :
Ophrys tenthredinifera Willd.; Sp. Pl. 4: 67 (1805)
subspecies :
subsp. tenthredinifera
Name: Wespen-Ragwurz
Distribution: S-France; Spain; Gibraltar; Portugal; Baleares; Tunisia; Algeria; Morocco; Crete (incl. Gavdos Isl.); Croatia; Greece (Skyros, etc.); East Aegaean Isl. (Karpathos, Kasos, Rhodos, Chios); Turkey; Cyprus; ?Libya (?Cyrenaica)
12 BAL FRA GIB POR SPA 13 CRO GRC KRI 20 ALG MOR TUN 34 CYP EAI TUR
Remarks: tenthredinifera agg., pollinator Eucera nigrilabris and other Eucera. According to Hirth & Paulus (2012) not distinguishable from the eastern ssp. villosa
Synonyms: hidden
= Arachnites tenthredinifera (Willd.) Tod., Orch. Sicul. 85 (1842)
= Ophrys arachnites Link, Schrad., Jour. für die Bot. 2: 325 (1799)
= Ophrys dictynnae P.Delforge, Natural. belges 86 (Orchid. 18): 95-144 (2005)
= Ophrys hookeri Orph. ex Soó, Bot. Ark. 23: 25 (1928)
= Ophrys limbata Link, Handb. 1: 247 (1829)
= Ophrys lychnitis Paulus & Hirth, J. Eur. Orch. 48(2-4): 374 (2016)
= Ophrys tabanifera Sieber ex Nyman, Consp. 697 (1882)
= Ophrys tenthredinifera f. lutescens (Batt.) Raynaud, Orchid. Maroc 26 (1985)
= Ophrys tenthredinifera ssp. dictynnae (P.Delforge) Kreutz, Orchidee (Hamburg) 57(1): 103 (2006)
= Ophrys tenthredinifera ssp. villosa (Desf.) H.Baumann & Künkele, Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 18 (3): 384 (1986)
= Ophrys villosa Desf., Ann. Mus. Par. 10: 225, t. 14 (1807)
variant :
var. expansa Lumare & Medagli; GIROS Orch. Spont. Eur. 61(1): 30 (2018)
Distribution: Italy
13 ITA
variant :
var. glabrescens F.M.Vázquez; Folia Bot. Extremadur. 3: 320 (2009)
Distribution: Spain
12 SPA
variant :
var. riphaea (F.M.Vázquez) comb.ined.
Distribution: Morocco
20 MOR
Synonyms: hidden
= Ophrys neglecta var. riphaea F.M.Vázquez, J. Eur. Orch. 44(3): 588 (2012)
= Ophrys riphaea (F.M.Vázquez) P.Delforge, Naturalistes Belges 94(Orchid. 26): 296 (2013)
variant :
var. korae (Hirth & Paulus) comb.ined.
Name: Kora-Ragwurz (DE); Kora Sawfly Orchid (EN)
Distribution: East Aegaean Isl. (Rhodos, Samos, ?Lesvos)
34 EAI
Remarks: tenthredinifera agg. Probably only a color variant of ssp. tenthredinifera (villosa)
Synonyms: hidden
= Ophrys korae Hirth & Paulus, Journ. Eur. Orch. 44(3): 646 (2012)
subspecies :
subsp. spectabilis Kreutz & Zelesny; Ber. Arbeitskreis. Heimische Orchid. 24(1): 116 (2007)
Distribution: Baleares (Mallorca)
12 BAL
Remarks: tenthredinifera agg.
Synonyms: hidden
= Ophrys spectabilis (Kreutz & Zelesny) Paulus, J. Eur. Orch. 43(1): 38 (2011)
subspecies :
subsp. ficalhoana (J.A.Guim.) M.R.Lowe & D.Tyteca; J. Eur. Orchid. 32 (2): 297 (2000)
Distribution: Portugal; Spain; SW-France; ?Morocco; ?W-Algeria
12 FRA POR SPA
Remarks: tenthredinifera agg., pollinator Eucera sp.
Synonyms: hidden
= Ophrys ficalhoana J.A.Guim., Bol. Soc. Brot. 5: 39 (1887)
= Ophrys tenthredinifera var. ronda Schltr., [nom. nud.]
= Ophrys tenthredinifera f. mariana Rivas Goday & Bellot, (1945)
= Ophrys tenthredinifera var. ficalhoana (J.A.Guim.) F.M.Vázquez, Folia Bot. Extremadur. 3: 316 (2009)
= Ophrys ficalhoana (J.A.Guim.) Wucherpf., Ber. Arbeitskreis. Heimische Orchid. 33(1): 113 (2016), nom. illeg. superfl.
variant :
var. praecox Rchb.fil. ex E.G.Camus, Bergon & A.Camus; Monogr. Orchid. 262 (1908)
Distribution: W-Spain; Portugal
12 POR SPA
Remarks: tenthredinifera agg.
Synonyms: hidden
= Ophrys tenthredinifera ssp. guimaraesii D.Tyteca, J. Eur. Orchideen 32 (2): 297 (2000)
= Ophrys tenthredinifera ssp. praecox D.Tyteca, L´Orchidophile (Asnieres) 16(66): 810 [nom. illeg.]
= Ophrys praecox D.Tyteca, [nom. illeg.]
= Ophrys tenthredinifera subvar. praecox J.A.Guim., [nom. illeg.]
= Ophrys tenthredinifera var. guimaraesii (D.Tyteca) Kreutz, Kompend. Eur. Orchid. 118 (2004)
subspecies :
subsp. neglecta (Parl.) E.G.Camus; Monogr. Orchid. (1908)
Distribution: C-Italy; S-Italy; Sardinia; ?S-Corsica
12 SAR 13 ITA
Remarks: tenthredinifera agg., pollinator Eucera clypeata and E. oraniensis
Synonyms: hidden
= Ophrys rosea ssp. neglecta (Parl.) Guadagno, (1931)
= Ophrys neglecta Parl., Fl. Ital. 3: 548 (1860)
= Ophrys rosea Grande, Nuov. Giorn. Bot. Ital., n. s. 27: 238 (1920)
subspecies :
subsp. aprilia (Devillers & Devillers-Tersch.) Kreutz; Kompend. Eur. Orchid. 118 (2004)
Distribution: S-Corsica; ?Sardinia
12 COR
Remarks: tenthredinifera agg.
Synonyms: hidden
= Ophrys aprilia Devillers & Devillers-Tersch., Jour. Eur. Orch. 35(1): 152 (2003)
subspecies :
subsp. grandiflora (Ten.) Kreutz; Kompend. Eur. Orchid. 118 (2004)
Distribution: Sicily; S-Italy; Tunisia; Algeria
13 ITA SIC 20 ALG TUN
Remarks: tenthredinifera agg.
Synonyms: hidden
= Ophrys grandiflora Ten., Cat. Pl. Hort. Neap. (1819) App. 2: 83, Fl. Nap. 2: 309
= Ophrys tenoreana Lindl., Bot. Reg. t. 1093 (1827)
subspecies :
subsp. leochroma (P.Delforge) Kreutz; Orchidee (Hamburg) 57(1): 103 (2006)
Name: Löwenmähnen-Ragwurz (DE); Lion-mane Sawfly Orchid (EN)
Distribution: Crete; Greece (Euboia, Attika, Peloponnes, Ionian Isl.: Kefallenia, Zakynthos, Kyklades: Kea, Astypaleia, Amorgos, Milos, Kimolos); East Aegaean Isl. (Samos, Rhodos, Chalki, Tilos); W-Turkey
13 GRC KRI 34 EAI TUR
Remarks: tenthredinifera agg., pollinator Eucera kullenbergi. Status uncertain, possibly a color variant of ssp. villosa
Synonyms: hidden
= Ophrys leochroma P.Delforge, Natural. belges 86 (Orchid. 18): 95-144 (2005)
subspecies :
subsp. lycomedis (P.Delforge) comb.ined.
Distribution: Greece (Kyklades: Skyros)
13 GRC
Remarks: tenthredinifera agg., pollinator Eucera caspica
Synonyms: hidden
= Ophrys lycomedis P.Delforge, Naturalistes Belges 94(Orchid. 26): 296 (2013)
subspecies :
subsp. ulyssea (P.Delforge) Kreutz; Orchidee (Hamburg) 57(1): 103 (2006)
Distribution: NW-Greece (Ionian Isl.: Corfu, Lefkada, Kefallenia, Zakynthos, Kythera); SW-Greece; W-Greece;
13 GRC
Remarks: tenthredinifera agg., pollinator Eucera bidentata
Synonyms: hidden
= Ophrys ulyssea P.Delforge, Natural. belges 86 (Orchid. 18): 95-144 (2005)
= Ophrys amphidami P.Delforge, Naturalistes Belges 94(Orchid. 26): 295 (2013) [fide Hirth & Paulus 2016]
subspecies :
subsp. sanctae-marcellae Saliaris, A.Saliaris & A.Alibertis; J. Eur. Orch. 43(3): 605 (2011)
Distribution: East Aegaean Isl.
34 EAI
Synonyms: hidden
= Ophrys leochroma var. sanctae-marcellae (Saliaris, A.Saliaris & A.Alibertis) P.Delforge, Naturalistes Belges 93(Orchid. 25): 243 (2012)
= Ophrys villosa var. sanctae-marcellae (Saliaris, A.Saliaris & A.Alibertis) P.Delforge, Naturalistes Belges 94(Orchid. 26): 296 (2013)
subspecies :
subsp. tingurtiae Kreutz, Miara, Ait-Hamm. & Rebbas; Ber. Arbeitskreis. Heimische Orchid. 31(2): 147 (2015)
Distribution: Algeria
20 ALG
species :
Ophrys transhyrcana Czerniak.; Not. Syst. Herb. Hort. Petrop. 4: 1 (1923)
subspecies :
subsp. transhyrcana
Distribution: Crete (Gavdos Isl.); Turkey; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkmenistan
13 KRI 32 TKM 33 ARM AZE GEO NCS 34 TUR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys adonidis A.Camus & Gomb., Notul. Syst. (Paris) 14: 104 (1950)
= Ophrys aranifera ssp. transhyrcana (Czerniak.) Soó, Repert. Spec. Nov. Regni Veg., Sonderbeih. A2: 49 (1931)
= Ophrys mammosa ssp. transhyrcana (Czerniak.) Paulus & Gack, Jahresber. Naturwiss. Ver. Wuppertal 39: 68 (1986)
= Ophrys sphegodes ssp. transhyrcana (Czerniak.) Soó, Acta Bot. Acad. Sci. Hung. 5: 444 (1970)
= Ophrys sphegodes ssp. transhyrcana (Czerniak.) Buttler, Willdenowia 16 (1): 115 (1986)
= Ophrys sphegodes var. transhyrcana (Czerniak.) P.J.Cribb, Field Guide Orchids Europe Medit. 259 (2019)
subspecies :
subsp. morio (Paulus & Kreutz) Kreutz; Kompend. Eur. Orchid. 119 (2004)
Distribution: Cyprus
34 CYP
Remarks: mammosa agg., pollinator Andrena morio
Synonyms: hidden
= Ophrys morio Paulus & Kreutz, Orchid. Zypern 415 (2004)
subspecies :
subsp. mouterdeana (B. & H.Baumann) Kreutz; Eurorchis 17: 110 (2005)
Distribution: Lebanon; Israel; Jordania; Syria
34 ISR JOR LEB SYR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys mammosa ssp. mouterdeana B. & H.Baumann, Jour. Eur. Orch. 37(2): 263, 264 (2005)
= Ophrys mouterdeana (B. & H.Baumann) P.Delforge, Naturalistes Belges 96(Orchid. 28): 18 (2015)
= Ophrys doerfleri ssp. mouterdeana (B. & H.Baumann) Paulus & M.Hirth, J. Eur. Orch. 49(2): 253 (2017)
subspecies :
subsp. hittitica (Kreutz & Ruedi Peter) comb.ined.
Distribution: S-Turkey (Hatay, Antakya); NW-Syria
34 SYR TUR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys hittitica Kreutz & Ruedi Peter, J. Eur. Orchid. 30 (1): 128 (prepr. 30 Sept. 1997)
subspecies :
subsp. paphlagonica Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 119 (2007)
Distribution: N-Turkey
34 TUR
Remarks: mammosa agg.
Synonyms: hidden
= Ophrys paphlagonica (Kreutz) P.Delforge, Naturalistes Belges 96(Orchid. 28): 18 (2015)
subspecies :