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492.0000 Family Rubiaceae Juss.; Gen. Pl. [Jussieu] 196 (1789), nom. cons.
589 genera, ca. 14.200 spp. mostly tropics, comparably few temperate. Phylogeny: Razafimandimbison & Rydin (2024)
Species listed in database: 14370; hybrids: 47
= Aparinaceae Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 2: 3, 38 (1813-1829)
= Asperulaceae Cham. ex Spenn.; Handb. Angew. Bot. 2: 493 (1835)
= Catesbaeaceae Martinov; Tekhno-Bot. Slovar 111 (1820)
= Cephalanthaceae Raf.; Ann. Gén. Sci. Phys. 6: 86 (1820)
= Cinchonaceae Batsch; Tab. Affin. Regni Veg. 234 (1802)
= Coffeaceae Batsch; Tab. Affin. Regni Veg. 233 (1802)
= Coutareaceae Martinov; Tekhno-Bot. Slovar 168 (1820)
= Dialypetalanthaceae Rizzini & Occhioni; Lilloa 17: 253 (1948), nom. cons.
= Galiaceae Lindl.; Intr. Nat. Syst. Bot., ed. 2: 249 (1836)
= Gardeniaceae Dumort.; Anal. Fam. Pl. 29, 32 (1829)
= Guettardaceae Batsch; Tab. Affin. Regni Veg. 235 (1802)
= Hameliaceae Mart.; Consp. Regn. Veg. [Martius] 31 (1835)
= Hedyotidaceae Dumort.; Commentat. Bot. [Dumortier] 57 (1822)
= Henriqueziaceae Bremek.; Acta Bot. Neerl. 6 (Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 141): 371 (1957)
= Houstoniaceae Raf.; Good Book 21 (1840)
= Hydrophylacaceae Martinov; Tekhno-Bot. Slovar 318 (1820)
= Lippayaceae Meisn.; Pl. Vasc. Gen. [Meisner], Tab. Diagn. 156, Comm. 112 (1838)
= Lygodisodeaceae Bartl.; Ord. Nat. Pl. 123, 207 (1830)
= Naucleaceae Wernham; New Phytol. 11: 225 (1912)
= Nonateliaceae Martinov; Tekhno-Bot. Slovar 420 (1820)
= Operculariaceae Juss. ex Perleb; Vers. Arzneikr. Pfl. 207 (1818)
= Pagamaeaceae Martinov; Tekhno-Bot. Slovar 447 (1820)
= Psychotriaceae F.Rudolphi; Syst. Orb. Veg. 49 (1830)
= Randiaceae Martinov; Tekhno-Bot. Slovar 534 (1820)
= Sabiceaceae Martinov; Tekhno-Bot. Slovar 555 (1820)
= Spermacoceaceae Bercht. & J.Presl; Přir. Rostlin 256 (1820)
= Theligonaceae Dumort.; Anal. Fam. Pl. 15, 17 (1829), nom. cons.
492.2301 Genus Pagamea Aubl.; Hist. Pl. Guiane 1: 112, t. 44 (1775)
Distribution: Tropical S-America
Species listed in database: 30
= Aphanocarpus Steyerm.; Mem. New York Bot. Gard. 12(3): 263 (1965)
= Coryphothamnus Steyerm.; Mem. New York Bot. Gard. 12(3): 264 (1965)
= Pagameopsis Steyerm.; Mem. New York Bot. Gard. 12(3): 267 (1965)
species :
Pagamea acrensis Steyerm.; Brittonia 33: 385 (1981)
Distribution: N-Brazil (Amazonas, Acre)
84 BRA
species :
Pagamea anisophylla Standl. & Steyerm.; Fieldiana, Bot. 28: 584 (1953)
Distribution: Venezuela (Amazonas)
82 VEN
species :
Pagamea aracaensis B.M.Boom; Brittonia 41: 129 (1989)
Distribution: N-Brazil (Amazonas)
84 BRA
species :
Pagamea auyantepuiensis Steyerm.; Fieldiana, Bot. 228: 584 (1953)
Distribution: Venezuela (Bolívar: Auyan-tepui)
82 VEN
Synonyms: hidden
= Coryphothamnus auyantepuiensis (Steyerm.) Steyerm.; Mem. New York Bot. Gard. 12: 264 (1965)
species :
Pagamea capitata Benth.; J. Proc. Linn. Soc., Bot. 1: 85, 109 (1857)
subspecies :
subsp. capitata
Distribution: Venezuela (Bolívar); Guyana; Surinam
82 GUY SUR VEN
Synonyms: hidden
= Pagamea brevipedunculata Steyerm.; Mem. New York Bot. Gard. 12: 283 (1965)
subspecies :
subsp. caudata (Huber) Steyerm.; Mem. New York Bot. Gard. 12: 280 (1965)
Distribution: N-Brazil (Pará)
84 BRA
Synonyms: hidden
= Pagamea caudata Huber; Bull. Soc. Bot. Genève, Sér. 2, 6: 211 (1914)
subspecies :
subsp. conferta (Standl.) Steyerm.; Mem. New York Bot. Gard. 12: 281 (1965)
Distribution: Venezuela (Amazonas, Bolívar)
82 VEN
Synonyms: hidden
= Pagamea capitata f. breviloba Steyerm.; Mem. New York Bot. Gard. 12: 281 (1965)
= Pagamea conferta Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 7: 419 (1931)
subspecies :
subsp. thibaudiifolia (Wernham) Steyerm.; Mem. New York Bot. Gard. 12: 280 (1965)
Distribution: Guyana; Surinam
82 GUY SUR
Synonyms: hidden
= Cephaelis thibaudiifolia Wernham; J. Bot. 52: 313 (1914)
species :
Pagamea coriacea Spruce ex Benth.; J. Proc. Linn. Soc., Bot. 1: 110 (1857)
Distribution: Colombia (Amazonas, Caquetá, Guainía, Vaupés); Venezuela (Amazonas); Guyana; Peru; N-Brazil (Pará, Amazonas, Rondônia)
82 CLM GUY VEN 83 PER 84 BRA
Synonyms: hidden
= Pagamea coriacea var. acuta Steyerm.; Mem. New York Bot. Gard. 12(3): 273 (1965)
= Pagamea coriacea var. pubescens Steyerm.; Mem. New York Bot. Gard. 12: 273 (1965)
species :
Pagamea diceras Steyerm.; Mem. New York Bot. Gard. 12: 281 (1965)
Distribution: Venezuela (Amazonas)
82 VEN
species :
Pagamea duckei Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 17. 278 (1937)
Distribution: N-Brazil (Amazonas, Acre)
84 BRA
species :
Pagamea dudleyi Steyerm.; Phytologia 31: 226 (1975)
Distribution: Ecuador; Peru
83 ECU PER
species :
Pagamea duidana Standl. & Steyerm.; Fieldiana, Bot. 28: 586 (1953)
Distribution: Venezuela (Amazonas: Cerro Duida)
82 VEN
species :
Pagamea garryoides Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 7: 420 (1931)
Distribution: Venezuela (Amazonas, Bolívar)
82 VEN
Synonyms: hidden
= Pagameopsis garryoides (Standl.) Steyerm.; Mem. New York Bot. Gard. 12: 267 (1965)
species :
Pagamea guianensis Aubl.; Hist. Pl. Guiane 113 (1775)
Distribution: Venezuela (Amazonas, Apure); Guyana; Surinam; French Guiana; Bolivia (La Paz, Santa Cruz); N-Brazil (Roraima, Amapá, Pará, Amazonas, Rondônia); NE-Brazil (Maranhão, Bahia); WC-Brazil (Mato Grosso)
82 FRG GUY SUR VEN 83 BOL 84 BRA
Synonyms: hidden
= Pagamea guianensis var. angustifolia Progel; Fl. Bras. [Martius] 6(1): 285 (1868)
= Pagamea guianensis var. parviflora Spruce ex Progel; Fl. Bras. [Martius] 6(1): 285 (1868)
variant :
var. macrocarpa Steyerm.; Mem. New York Bot. Gard. 12: 278 (1965)
Distribution: Colombia (Amazonas, Caquetá, Guainía, Vaupés, Vichada); N-Brazil (Amazonas)
82 CLM 84 BRA
species :
Pagamea harleyi Steyerm.; Kew Bull. 38: 317 (1983)
Distribution: NE-Brazil (Bahia)
84 BRA
species :
Pagamea hirsuta Spruce ex Benth.; J. Proc. Linn. Soc., Bot. 1: 111 (1857)
Distribution: Colombia (Amazonas, Guainía, Vaupés); Venezuela (Amazonas)
82 CLM VEN
species :
Pagamea jauaensis Steyerm.; Mem. New York Bot. Gard. 23: 883 (1972)
Distribution: Venezuela (Amazonas, Bolívar)
82 VEN
species :
Pagamea macrophylla Spruce ex Benth.; J. Proc. Linn. Soc., Bot. 1: 110 (1857)
Distribution: Colombia (Vaupés); N-Brazil (Amazonas)
82 CLM 84 BRA
species :
Pagamea magniflora Steyerm.; Mem. New York Bot. Gard. 12: 285 (1965)
Distribution: Venezuela (Bolívar)
82 VEN
species :
Pagamea maguirei (Steyerm.) comb.ined.
Distribution: Venezuela (Amazonas, Bolívar); N-Brazil (Amazonas)
82 VEN 84 BRA
Synonyms: hidden
= Pagameopsis maguirei Steyerm.; Mem. New York Bot. Gard. 12: 267 (1965)
= Pagameopsis maguirei subsp. neblinensis Steyerm.; Ann. Missouri Bot. Gard. 74: 108 (1987)
= Pagameopsis maguirei subsp. pusillus Steyerm.; Ann. Missouri Bot. Gard. 74: 106 (1987)
= Pagameopsis maguirei var. angustifolius Steyerm.; Mem. New York Bot. Gard. 12(3): 270 (1965)
= Pagameopsis maguirei var. glabrus Steyerm.; Ann. Missouri Bot. Gard. 74: 108 (1987)
= Pagameopsis maguirei var. pirapucuensis Steyerm.; Ann. Missouri Bot. Gard. 74: 108 (1987)
species :
Pagamea montana Gleason & Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 7: 421 (1931)
Distribution: Venezuela (Amazonas, Bolívar); N-Brazil (Amazonas)
82 VEN 84 BRA
species :
Pagamea pauciflora Standl. & Steyerm.; Fieldiana, Bot. 28: 588 (1953)
Distribution: Venezuela (Bolívar)
82 VEN
species :
Pagamea pilosa (Standl.) Steyerm.; Mem. New York Bot. Gard. 12: 276 (1965)
Distribution: N-Brazil (Amazonas)
84 BRA
Synonyms: hidden
= Pagamea guianensis var. pilosa Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 17: 279 (1937)
species :
Pagamea plicata Spruce ex Benth.; J. Proc. Linn. Soc., Bot. 1: 109 (1857)
Distribution: Colombia (Amazonas, Caquetá, Guainía, Vaupés); Venezuela (Amazonas, Bolívar); N-Brazil (Amazonas); WC-Brazil (Mato Grosso)
82 CLM VEN 84 BRA
Synonyms: hidden
= Pagamea plicata var. multinervia Steyerm.; Mem. New York Bot. Gard. 12: 274 (1965)
variant :
var. glabrescens Benth.; J. Proc. Linn. Soc., Bot. 1: 109 (1857)
Distribution: NE-Brazil (Piauí, Pernambuco)
84 BRA
species :
Pagamea plicatiformis Steyerm.; Mem. New York Bot. Gard. 12: 274 (1965)
Distribution: Venezuela (Amazonas); N-Brazil (Amazonas)
82 VEN 84 BRA
species :
Pagamea puberula Steyerm.; Mem. New York Bot. Gard. 12: 276 (1965)
Distribution: N-Brazil (Pará, Amazonas)
84 BRA
species :
Pagamea sessiliflora Spruce ex Benth.; J. Proc. Linn. Soc., Bot. 1: 110 (1857)
Distribution: Venezuela (Amazonas)
82 VEN
species :
Pagamea spruceana Vicent. & E.M.B.Prata; Phytotaxa 269(3): 187 (2016)
Distribution: Brazil (Amazonas)
84 BRA
species :
Pagamea standleyana Steyerm.; Fieldiana, Bot. 28: 589 (1953)
Distribution: Venezuela (Bolívar)
82 VEN
Synonyms: hidden
= Pagamea reducta Steyerm.; Mem. New York Bot. Gard. 12: 283 (1965)
= Pagamea uniflora Steyerm.; Mem. New York Bot. Gard. 12: 284 (1965)
species :
Pagamea steyermarkii Standl.; Fieldiana, Bot. 228: 589 (1953)
Distribution: Venezuela (Bolívar)
82 VEN
Synonyms: hidden
= Aphanocarpus steyermarkii f. elongatus Steyerm.; Ann. Missouri Bot. Gard. 71(1): 331 (1984)
= Aphanocarpus steyermarkii f. glabrior Steyerm.; Ann. Missouri Bot. Gard. 71(1): 330 (1984)
= Aphanocarpus steyermarkii (Standl.) Steyerm.; Mem. New York Bot. Gard. 12: 264 (1965)
species :
Pagamea thyrsiflora Spruce ex Benth.; J. Proc. Linn. Soc., Bot. 1: 110 (1857)
Distribution: Colombia (Amazonas, Caquetá, Guainía, Vaupés); Venezuela (Amazonas, Bolívar); Guyana; Ecuador; N-Brazil (Pará, Rondônia)
82 CLM GUY VEN 83 ECU 84 BRA
species :
Pagamea velutina Steyerm.; Mem. New York Bot. Gard. 12: 275 (1965)
Distribution: Venezuela (Amazonas); N-Brazil (Amazonas)
82 VEN 84 BRA
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.08 (August 2nd, 2026)

Covering 360,349 accepted species,
8,927 hybrids, 47,544 accepted infraspecific taxa and 1,011,667 synonyms,
in total 1,428,487* names

[*about 20,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life