293.0000
Family
Brassicaceae Burnett; Outlines Bot. [Burnett] 854, 1093, 1123 (1835), nom. cons.
339 genera, ca. 4050 spp. Updated phylogeny see Hendriks et al. (2022) and German et al. (2023)
Species listed in database: 4089; hybrids: 111
= Arabidaceae Döll; Rhein. Fl. [Döll] 573 (1843)
= Cruciferae Juss.; Gen. Pl. [Jussieu] 237 (1789), nom. cons.
= Drabaceae Martinov; Tekhno-Bot. Slovar 215 (1820)
= Erysimaceae Martinov; Tekhno-Bot. Slovar 238 (1820)
= Isatidaceae Döll; Fl. Baden [Döll] 3: 1310 (1861)
= Raphanaceae Horan.; Char. Ess. Fam. 169 (1847)
= Schizopetalaceae A.Juss.; Dict. Univ. Hist. Nat. 11: 419 (1848)
= Sisymbriaceae Martinov; Tekhno-Bot. Slovar 583 (1820)
= Stanleyaceae Nutt.; J. Acad. Nat. Sci. Philadelphia 7: 85 (1834)
= Thlaspiaceae Martinov; Tekhno-Bot. Slovar 633 (1820)
293.3901
Genus
Physaria (Nutt. ex Torr. & A.Gray) A.Gray; Gen. Amer. Bor. 1: 162 (1848)
Distribution: North America (88) to C-Mexico; NE-European Russia, S-South America
Species listed in database: 110
= Cistocarpium Spach; Hist. Nat. Vég. [Spach] 6: 471 (1838)
= Coulterina Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Discovium Raf.; J. Phys. Chim. Hist. Nat. Arts 89: 96 (1819)
= Lesquerella S.Watson; Proc. Amer. Acad. Arts 23: 250 (1888)
= Vesicaria Tourn. ex Adans.; Fam. Pl. [Adanson] 2: 429 (1763)
species :
Physaria acutifolia
Rydb.; Bull. Torrey Bot. Club 279 (1901)
Name: Sharpleaf twinpod
Distribution: USA (Arizona, Colorado, Idaho, Montana, Nevada, New Mexico, South Dakota, Utah, Wyoming)
73 USA 74 USA 76 USA 77 USA
Synonyms: hidden
= Physaria acutifolia var. stylosa (Rollins) S.L.Welsh; Great Basin Naturalist 46(2): 255 (1986)
= Physaria australis (Payson) Rollins; Rhodora 41: 408 (1939)
= Physaria didymocarpa var. australis Payson; Ann. Missouri Bot. Gard. 5: 144 (1918)
= Physaria stylosa Rollins; Contr. Gray Herb. 214: 14 (1984)
species :
Physaria alpestris
Suksd.; W. Amer. Sci. 15: 58 (1906)
Name: Washington twinpod
Distribution: USA (Washington State)
73 USA
Synonyms: hidden
= Lesquerella alpestris (Suksd.) Mulligan; Canad. J. Bot. 46: 528 (1968)
species :
Physaria alpina
Rollins; Brittonia 33(3): 339 (1981)
Name: Avery Peak twinpod
Distribution: USA (WC-Colorado)
73 USA
species :
Physaria angustifolia
(Nutt.) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
Name: Threadleaf bladderpod
Distribution: USA (Oklahoma, N-Texas)
74 USA 77 USA
Synonyms: hidden
= Lesquerella angustifolia (Nutt.) S.Watson; Proc. Amer. Acad. Arts 23: 253 (1888)
= Lesquerella longifolia Cory; Field & Lab. 18: 96 (1950)
= Vesicaria angustifolia Nutt.; Fl. N. Amer. [Torrey & A.Gray] 1: 101 (1838)
species :
Physaria arctica
(Wormsk. ex Hornem.) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
Name: Arctic bladderpod
Distribution: Russian Far East (Magadan); Alaska; Greenland; Canada (Alberta, British Columbia, Manitoba, Newfoundland, Labrador, N. W. T., Nunavut, Quebec, Yukon)
31 RFE 70 ASK CDN GNL 71 CDN 72 CDN
Synonyms: hidden
= Alyssum arcticum Wormsk. ex Hornem.; Fl. Dan. [Oeder et al.] 9: 3, t. 1520 (1816)
= Alyssum arcticum Wormsk. ex Vahl; Fl. Dan. [Oeder et al.] t. 1520 (1818)
= Lesquerella arctica (Wormsk. ex Hornem.) S.Watson; Proc. Amer. Acad. Arts 23: 255 (1888)
= Lesquerella arctica subsp. purshii (S.Watson) A.E.Porsild; Natl. Mus. Canada Bull. 121: 191 (1951)
= Lesquerella arctica var. minor Tatew. & Kobayashi; J. Fac. Agric. Hokkaido Imp. Univ. 36: 49 (1934)
= Lesquerella arctica var. purshii S.Watson; Proc. Amer. Acad. Arts 23: 254 (1888)
= Lesquerella arctica var. scammaniae Rollins; Amer. J. Bot. 26: 421 (1939) (´scammanae´)
= Lesquerella beringiana Tatew. & Kobay.; J. Fac. Agric. Hokkaido Imp. Univ. 36: 49 (1934)
= Lesquerella purshii (S.Watson) Fernald; Rhodora 35: 267 (1933)
= Vesicaria arctica (Wormsk. ex Hornem.) Richardson; Narr. Journey Polar Sea [Franklin] App. 743, et App. ed. 2: 26 (1823)
= Vesicaria arctica var. leiocarpa Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 5: 19 (1877)
= Vesicaria leiocarpa (Trautv.) N.Busch; Fl. URSS [Komarov et al., eds.] 8: 362 (1939)
species :
Physaria arenosa
(Richardson) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
subspecies :
subsp. arenosa
Name: Great Plains bladderpod
Distribution: Canada (Alberta, Manitoba, Saskatchewan); USA (Colorado, Montana, Nebraska, North Dakota, South Dakota, Wyoming)
71 CDN 73 USA 74 USA
Synonyms: hidden
= Lesquerella arenosa (Richardson) Rydb.; Bull. Torrey Bot. Club 236 (1902)
= Lesquerella argentea var. arenosa (Richardson) Rydb.; Contr. U.S. Natl. Herb. 3: 485 (1896)
= Lesquerella ludoviciana var. arenosa (Richardson) S.Watson; Proc. Amer. Acad. Arts 23: 252 (1888)
= Lesquerella lunellii A.Nelson; Bot. Gaz. 42: 49 (1906)
= Lesquerella lunellii var. lutea A.Nelson; Bot. Gaz. 54: 149 (1912)
= Lesquerella macounii Greene; Pittonia 4: 301 (1901)
= Lesquerella rosea Greene; Pittonia 4: 301 (1901)
= Lesquerella versicolor Greene; Pittonia 4: 301 (1901)
= Vesicaria arenosa Richardson; Narr. Journey Polar Sea [Franklin] App. 743, et App. ed. 2: 26 (1823)
subspecies :
subsp. argillosa
(Rollins & E: A.Shaw) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
Distribution: USA (Colorado, Nebraska, South Dakota, Wyoming)
73 USA 74 USA
Synonyms: hidden
= Lesquerella arenosa var. argillosa Rollins & E: A.Shaw; Gen. Lesquerella 178 (1973)
= Physaria arenosa var. argillosa (Rollins & E.A.Shaw) B.L.Turner; Phytologia 86(3): 203 (2004)
species :
Physaria argentea
(Schauer) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
Distribution: Mexico (Chihuahua, Ciudad de Mexico, Guanajuato, Hidalgo, Mexico State, Puebla, Queretaro, Veracruz, Zacatecas)
79 MEX
Synonyms: hidden
= Alyssum schauerianum Kuntze; Revis. Gen. Pl. 1: 16 (1891)
= Lesquerella argentea (S.Schauer) S.Watson; Proc. Amer. Acad. Arts 23: 252 (1888)
= Lesquerella schaueriana (Kuntze) Payson; Ann. Missouri Bot. Gard. 8: 139 (1921 publ. 1922)
= Vesicaria argentea S.Schauer; Linnaea 20: 720 (1847)
species :
Physaria argyraea
(A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
subspecies :
subsp. argyraea
Name: Silver bladderpod
Distribution: USA (S-Texas); NE-Mexico (Coahuila, Nuevo León)
77 USA 79 MEX
Synonyms: hidden
= Alyssum argyreum (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella argyraea (A.Gray) S.Watson; Proc. Amer. Acad. Arts 23: 251 (1888)
= Vesicaria argyraea A.Gray; Boston J. Nat. Hist. 6: 146 (1850)
subspecies :
subsp. diffusa
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Guanajuato, Jalisco, Nuevo Leon, Queretaro, San Luis Potosi, Tamaulipas, Zacatecas)
79 MEX
Synonyms: hidden
= Lesquerella argyraea subsp. diffusa (Rollins) Rollins & E.A.Shaw; Gen. Lesquerella 83 (1973)
= Lesquerella argyraea var. diffusa (Rollins) Rollins; Crucifer. Continental N. Amer. 607 (1993)
= Lesquerella diffusa Rollins; J. Arnold Arbor. 21: 395 (1940)
= Physaria argyraea var. diffusa (Rollins) B.L.Turner; Phytologia 86(3): 203 (2004)
species :
Physaria arizonica
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 321 (2002)
Name: Arizona bladderpod
Distribution: USA (Arizona, Utah)
76 USA
Synonyms: hidden
= Lesquerella arizonica S.Watson; Proc. Amer. Acad. Arts 23: 254 (1888)
= Lesquerella arizonica var. nudicaulis Payson; Ann. Missouri Bot. Gard. 8: 208, fig. 25 (1922)
= Physaria arizonica var. andrusensis N.D.Atwood, S.L.Welsh & L.C.Higgins; Rhodora 109(940): 402 (-405
species :
Physaria aurea
(Wooton) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: Golden bladderpod
Distribution: USA (SC-New Mexico: Jicarilla, Sacramento)
76 USA 77 USA
Synonyms: hidden
= Lesquerella aurea Wooton; Bull. Torrey Bot. Club 260 (1898)
species :
Physaria bellii
G.A.Mulligan; Canad. J. Bot. 44: 1662 (1966)
Name: Front Range twinpod
Distribution: USA (N-Colorado)
73 USA
species :
Physaria berlandieri
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Distribution: Mexico (Coahuila, Nuevo Leon, San Luis Potosi, Tamaulipas)
79 MEX
Synonyms: hidden
= Lesquerella berlandieri S.Watson; Proc. Amer. Acad. Arts 23: 252 (1888)
species :
Physaria brassicoides
Rydb.; Bull. Torrey Bot. Club 237 (1902)
Name: Double twinpod
Distribution: USA (Colorado, Montana, Nebraska, North Dakota, South Dakota, Wyoming)
73 USA 74 USA
species :
Physaria calcicola
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: Rocky Mountain bladderpod
Distribution: USA (SE-Colorado, NE-New Mexico)
73 USA 77 USA
Synonyms: hidden
= Lesquerella calcicola Rollins; Amer. J. Bot. 26: 419 (1939)
species :
Physaria calderi
(G.A.Mulligan & A.E.Porsild) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: Calder´s bladderpod
Distribution: Alaska (Ogilvie, Richardson Mts.); Canada (N. W. T., Yukon)
70 ASK CDN
Synonyms: hidden
= Lesquerella arctica subsp. calderi (G.A.Mulligan & A.E.Porsild) Hultén; Bot. Not. 126: 488 (1973)
= Lesquerella arctica var. calderi (G.A.Mulligan & A.E.Porsild) Welsh
= Lesquerella calderi G.A.Mulligan & A.E.Porsild; Canad. J. Bot. 47: 215 (1969)
species :
Physaria carinata
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
subspecies :
subsp. carinata
Name: Idaho bladderpod
Distribution: USA (W-Wyoming, EC-Idaho, Montana)
73 USA
Synonyms: hidden
= Lesquerella carinata Rollins; Contr. Gray Herb. 171: 42 (1950)
= Lesquerella carinata var. languida Rollins; Crucifer. Continental N. Amer. 611 (1993)
subspecies :
subsp. paysonii
(Rollins) O´Kane; Novon 17: 379 (2007)
Name: Payson´s bladderpod
Distribution: USA (W-Wyoming, E-Idaho, W-Montana)
73 USA
Synonyms: hidden
= Lesquerella paysonii Rollins; Contr. Gray Herb. 171: 44 (1950)
= Physaria paysonii (Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
subspecies :
subsp. pulchella
(Rollins) O´Kane; Novon 17: 379 (2007)
Name: Beautiful bladderpod
Distribution: USA (Idaho, Montana)
73 USA
Synonyms: hidden
= Lesquerella pulchella Rollins; Novon 5(1): 72 (1995)
= Physaria pulchella (Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
species :
Physaria chambersii
Rollins; Rhodora 41: 403 (1939)
Name: Chambers´ twinpod
Distribution: USA (Arizona, California, Idaho, Nevada, Oregon, Utah)
73 USA 76 USA
Synonyms: hidden
= Physaria acutifolia var. canaani S.L.Welsh; Utah Fl., ed. 3: 287 (2003)
= Physaria chambersii var. purpurea S.L.Welsh & Reveal; Great Basin Naturalist 37: 345 (1977 publ. 1978)
= Physaria chambersii var. sobolifera S.L.Welsh; Great Basin Naturalist 46(2): 255 (1986)
species :
Physaria cinerea
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: Basin bladderpod
Distribution: USA (C-Arizona)
76 USA
Synonyms: hidden
= Lesquerella cinerea S.Watson; Proc. Amer. Acad. Arts 23: 255 (1888)
species :
Physaria condensata
Rollins; Rhodora 41: 407 (1939)
Name: Tufted twinpod
Distribution: USA (W-Wyoming)
73 USA
species :
Physaria congesta
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: Dudley Bluffs bladderpod
Distribution: USA (Colorado: Rio Blanco Co.)
73 USA
Synonyms: hidden
= Lesquerella congesta Rollins; Contr. Gray Herb. 214: 8 (1984)
species :
Physaria cordiformis
Rollins; Contr. Gray Herb. 171: 47 (1950)
Name: Wassuk Range bladderpod
Distribution: USA (California, Idaho, Nevada, Utah)
73 USA 76 USA
Synonyms: hidden
= Lesquerella cordiformis (Rollins) Rollins & E.A.Shaw; Gen. Lesquerella 248 (1973)
= Lesquerella kingii var. cordiformis (Rollins) Maguire & A.H.Holmgren; Madroño 11: 183 (1951)
= Lesquerella kingii var. nevadensis Maguire & A.H.Holmgren; Madroño 11: 182 (1951)
species :
Physaria crassistigma
O´Kane & Al-Shehbaz; Novon 14(2): 200 (2004)
Distribution: Argentina (Mendoza)
85 ARG
species :
Physaria curvipes
(A.Nelson) Grady & O´Kane; Novon 17: 183 (2007)
Distribution: USA (Montana, Wyoming: Big Horn Mts.)
73 USA
Synonyms: hidden
= Lesquerella curvipes A.Nelson; Bull. Torrey Bot. Club 205 (1898)
species :
Physaria densiflora
(A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: Denseflower bladderpod
Distribution: USA (SC-Texas); Mexico (Tamaulipas)
77 USA 79 MEX
Synonyms: hidden
= Alyssum densiflorum (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella densiflora (A.Gray) S.Watson; Proc. Amer. Acad. Arts 23: 251 (1888)
= Vesicaria densiflora A.Gray; Boston J. Nat. Hist. 6: 145 (1850)
species :
Physaria didymocarpa
(Hook.) A.Gray; Gen. Amer. Bor. 1: 162 (1848)
subspecies :
subsp. didymocarpa
Name: Common twinpod
Distribution: Canada (Alberta, British Columbia, Idaho, Montana, Washington State, Wyoming)
71 CDN 73 USA
Synonyms: hidden
= Coulterina didymocarpa (Hook.) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Physaria didymocarpa var. contractoreplum Kuntze; Revis. Gen. Pl. 1: 35 (1891)
= Physaria macrantha Blank.; Sci. Stud. Montana Coll. Agric., Bot. 1: 60 (1905)
= Vesicaria didymocarpa Hook.; Fl. Bor.-Amer. [Hooker] 1: 49 (1830)
subspecies :
subsp. lanata
(A.Nelson) O´Kane; Novon 17: 379 (2007)
Distribution: USA (N-Wyoming, Montana)
73 USA
Synonyms: hidden
= Physaria didymocarpa var. lanata A.Nelson; Bull. Torrey Bot. Club 31: 241 (1901)
= Physaria lanata (A.Nelson) Rydb.; Bull. Torrey Bot. Club 39: 322 (1912)
subspecies :
subsp. lyrata
(C.L.Hitchc.) O´Kane; Novon 17: 379 (2007)
Name: Idaho twinpod
Distribution: USA (E-Idaho: Salmon River drainage)
73 USA
Synonyms: hidden
= Physaria didymocarpa var. lyrata C.L.Hitchc.; Univ. Wash. Publ. Biol. 17(2): 530 (1964)
species :
Physaria dornii
Lichvar; Brittonia 35(2): 151 (1983)
Name: Tunp Range twinpod
Distribution: USA (W-Wyoming)
73 USA
species :
Physaria douglasii
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
subspecies :
subsp. douglasii
Name: Douglas´ bladderpod
Distribution: Canada (British Columbia); USA (Idaho, Montana, Oregon, Washington)
71 CDN 73 USA
Synonyms: hidden
= Lesquerella douglasii S.Watson; Proc. Amer. Acad. Arts 23: 255 (1888)
subspecies :
subsp. tuplashensis
(Rollins, K.A.Beck & Caplow) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: White Bluffs bladderpod
Distribution: USA (Washington: White Bluffs of the Columbia River)
73 USA
Synonyms: hidden
= Lesquerella tuplashensis Rollins, K.A.Beck & Caplow; Rhodora 97(891): 203 (1995 publ. 1996)
species :
Physaria eburniflora
Rollins; Brittonia 33(3): 333 (1981)
Name: Devils Gate twinpod
Distribution: USA (C-Wyoming)
73 USA
species :
Physaria engelmannii
(A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 322 (2002)
Name: Engelmann´s bladderpod
Distribution: USA (Texas, Oklahoma)
74 USA 77 USA
Synonyms: hidden
= Alyssum engelmannii (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella engelmannii (A.Gray) S.Watson; Proc. Amer. Acad. Arts 23: 254 (1888)
= Vesicaria engelmannii A.Gray; Gen. Amer. Bor. 1: 162, t. 70 (1848)
= Vesicaria engelmannii var. elatior A.Gray; Boston J. Nat. Hist. 6: 145 (1850)
= Vesicaria pulchella Kunth & C.D.Bouché; Index Sem. (Berlin) 1848: 15 (1848)
species :
Physaria eriocarpa
Grady & O´Kane; Novon 17: 184, fig. 3 (2007)
Distribution: USA (Montana: Sheep Mountain)
73 USA
species :
Physaria fallax
K.A.Mathis & O´Kane; Phytoneuron 2024-61: 7 (2024)
Distribution: USA (N- & E-Arizona, SW-New Mexico, S-Utah)
76 USA 77 USA
species :
Physaria fendleri
(A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Fendler´s bladderpod
Distribution: USA (Arizona, Colorado, Kansas, New Mexico, Oklahoma, Texas, Utah); Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Hidalgo, Nayarit, Nuevo Leon, Queretaro, San Luis Potosi, Sonora, Zacatecas)
73 USA 74 USA 76 USA 77 USA 79 MEX
Synonyms: hidden
= Alyssum fendleri (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Alyssum stenophyllum (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella fendleri (A.Gray) S.Watson; Proc. Amer. Acad. Arts 23: 255 (1888)
= Lesquerella foliacea Greene; Pittonia 5: 134 (1903)
= Lesquerella praecox Wooton & Standl.; Contr. U.S. Natl. Herb. 16: 126 (1913)
= Lesquerella stenophylla (A.Gray) Rydb.; Bull. Torrey Bot. Club 33: 142 (1906)
= Vesicaria fendleri A.Gray; Mem. Amer. Acad. Arts, n. s. 4: 9 (1849)
= Vesicaria stenophylla A.Gray; Boston J. Nat. Hist. 6: 149 (1850)
= Vesicaria stenophylla var. diffusa A.Gray; Smithsonian Contr. Bot. 5 (Art. 6): 13 (1853)
= Vesicaria stenophylla var. humilis A.Gray; Smithsonian Contr. Bot. 5 (Art. 6): 13 (1853)
= Vesicaria stenophylla var. procera A.Gray; Smithsonian Contr. Bot. 5 (Art. 6): 13 (1853)
species :
Physaria filiformis
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Limestoneglade bladderpod
Distribution: USA (Alabama, Arkansas, Missouri)
75 USA 78 USA
Synonyms: hidden
= Lesquerella filiformis Rollins; Rhodora 58: 201 (1956)
species :
Physaria floribunda
Rydb.; Bull. Torrey Bot. Club 279 (1901)
subspecies :
subsp. floribunda
Name: Pointtip twinpod
Distribution: USA (Colorado, New Mexico)
73 USA 77 USA
subspecies :
subsp. osterhoutii
(Payson) O´Kane; Novon 17: 379 (2007)
Distribution: USA (NC-Colorado)
73 USA
Synonyms: hidden
= Physaria floribunda var. osterhoutii (Payson) Rollins; Crucifer. Continental N. Amer. 707 (1993)
= Physaria osterhoutii Payson; Ann. Missouri Bot. Gard. 5: 146 (1918)
species :
Physaria fremontii
(Rollins & E.A.Shaw) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Fremont´s bladderpod
Distribution: USA (Wyoming: Wind River Mts.)
73 USA
Synonyms: hidden
= Lesquerella fremontii Rollins & E.A.Shaw; Gen. Lesquerella 228 (1973)
species :
Physaria garrettii
(Payson) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Garrett´s bladderpod
Distribution: USA (Utah: Wasatch Mts.)
76 USA
Synonyms: hidden
= Lesquerella garrettii Payson; Ann. Missouri Bot. Gard. 8: 213 (1921 publ. 1922)
species :
Physaria geyeri
(Hook.) A.Gray; Gen. Amer. Bor. 1: 162 (1848)
subspecies :
subsp. geyeri
Name: Geyer´s twinpod
Distribution: USA (Montana, Oregon, Washington)
73 USA
Synonyms: hidden
= Coulterina geyeri (Hook.) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella geyeri (Hook.) Mulligan; Canad. J. Bot. 46: 528 (1968)
= Vesicaria geyeri Hook.; J. Bot. [Hooker] 6: 70 (1847)
subspecies :
subsp. purpurea
(Rollins) O´Kane; Novon 17: 380 (2007)
Name: Purple twinpod
Distribution: USA (C-Idaho)
73 USA
Synonyms: hidden
= Physaria geyeri var. purpurea Rollins; Rhodora 41: 401 (1939)
species :
Physaria globosa
(Desv.) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Globe bladderpod
Distribution: USA (NC-Kentucky to C-Tennessee, Indiana)
75 USA 78 USA
Synonyms: hidden
= Alyssum globosum (Desv.) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Alyssum globosum (Desv.) Steud.; Nomencl. Bot. [Steudel], ed. 1, 1: 34 (1821)
= Alyssum shortii (Torr. & A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella globosa (Desv.) S.Watson; Proc. Amer. Acad. Arts 23: 252 (1888)
= Vesicaria globosa Desv.; J. Bot. Agric. 3: 171, 184 (1815)
= Vesicaria shortii Torr. & A.Gray; Fl. N. Amer. [Torrey & A.Gray] 1(1): 102 (1838)
species :
Physaria gooddingii
(Rollins & E.A.Shaw) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Goodding´s bladderpod
Distribution: USA (Arizona: Greenlee Co., New Mexico: Catron, Sierra, western Socorro Co.)
76 USA 77 USA
Synonyms: hidden
= Lesquerella gooddingii Rollins & E.A.Shaw; Gen. Lesquerella 164 (1973)
species :
Physaria gordonii
(A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Gordon´s bladderpod
Distribution: USA (Arizona, Kansas, New Mexico, Oklahoma, Texas, Virginia [I]); Mexico (Aguascalientes, Chihuahua, Sonora)
74 USA 76 USA 77 USA 78 USA 79 MEX
Synonyms: hidden
= Alyssum gordonii (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella gordonii (A.Gray) S.Watson; Proc. Amer. Acad. Arts 23: 253 (1888)
= Lesquerella gordonii var. densifolia Rollins; Crucifer. Continental N. Amer. 621 (1993)
= Physaria gordonii subsp. densifolia (Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
= Physaria gordonii var. densifolia (Rollins) B.L.Turner; Phytologia 86(3): 203 (2004)
= Vesicaria gordonii A.Gray; Boston J. Nat. Hist. 6: 149 (1850)
species :
Physaria gracilis
(Hook.) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
subspecies :
subsp. gracilis
Name: Spreading bladderpod
Distribution: USA (Alabama, Illinois [I], Iowa [I], Louisiana, Mississippi, Missouri [I], Oklahoma, W-Tennessee [I], Texas)
74 USA 75 USA 77 USA 78 USA
Synonyms: hidden
= Alyssum gracile (Hook.) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella gracilis (Hook.) S.Watson; Proc. Amer. Acad. Arts 23: 253 (1888)
= Lesquerella polyantha (Schltdl.) Small; (1903)
= Vesicaria gracilis Hook.; Bot. Mag. t. 3533 (1836)
= Vesicaria polyantha Schltdl.; Bot. Zeitung (Berlin) 11: 620 (1853)
subspecies :
subsp. nuttallii
(Torr. & A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Name: Nuttall´s bladderpod
Distribution: USA (Kansas, Oklahoma, Texas)
74 USA 77 USA
Synonyms: hidden
= Alyssum nuttallii (Torr. & A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Alyssum repandum (Nutt. ex Torr. & A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella gracilis subsp. nuttallii (Torr. & A.Gray) Rollins & E.A.Shaw; Gen. Lesquerella 53 (1973)
= Lesquerella gracilis var. repanda (Nutt.) Payson; Ann. Missouri Bot. Gard. 8: 186 (1922)
= Lesquerella nuttallii (Torr. & A.Gray) S.Watson; Proc. Amer. Acad. Arts 22: 252 (1888)
= Lesquerella repanda (Nutt.) S.Watson; Proc. Amer. Acad. Arts 22: 254 (1888)
= Physaria acutifolia var. repanda (Rollins) S.L.Welsh; Utah Fl., ed. 3: 287 (2003)
= Physaria gracilis var. repanda (Nutt. ex Torr. & A.Gray) B.L.Turner; Phytologia 86(3): 204 (2004)
= Physaria repanda Rollins; Contr. Gray Herb. 214: 11 (1984)
= Vesicaria nuttallii Torr. & A.Gray; Fl. N. Amer. [Torrey & A.Gray] 1: 101 (1838)
= Vesicaria repanda Nutt.; Fl. N. Amer. [Torrey & A.Gray] 1: 101 (1838)
species :
Physaria grahamii
Morton ex E.H.Graham; Ann. Carnegie Mus. 26: (Bot. Stud. Uinta Basin, Utah & Colorado) 220 (1937)
Name: Graham´s twinpod
Distribution: USA (Utah)
76 USA
Synonyms: hidden
= Physaria acutifolia var. purpurea S.L.Welsh & Reveal; Great Basin Naturalist 37(3): 345 (1977 publ. 1978)
species :
Physaria hemiphysaria
(Maguire) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
subspecies :
subsp. hemiphysaria
Name: Intermountain bladderpod
Distribution: USA (Utah: Wasatch Plateau)
76 USA
Synonyms: hidden
= Lesquerella hemiphysaria Maguire; Amer. Midl. Naturalist 27: 466 (1942)
subspecies :
subsp. lucens
(S.L.Welsh & Reveal) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
Distribution: USA (Utah: West Tavaputs Plateau)
76 USA
Synonyms: hidden
= Lesquerella hemiphysaria var. lucens S.L.Welsh & Reveal; Great Basin Naturalist 37(3): 338 (1977 publ. 1978)
= Physaria hemiphysaria var. lucens (S.L.Welsh & Reveal) S.L.Welsh; Brittonia 56(3): 247 (2004)
species :
Physaria hitchcockii
(Munz) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
subspecies :
subsp. hitchcockii
Name: Hitchcock´s bladderpod
Distribution: USA (Nevada: Sheep Range, Spring Mts., Utah: Table Cliff Plateau)
76 USA
Synonyms: hidden
= Lesquerella hitchcockii Munz; Bull. Torrey Bot. Club 56: 163 (1929)
subspecies :
subsp. confluens
(Maguire & A.H.Holmgren) O´Kane; Novon 17: 380 (2007)
Distribution: USA (Nevada: Grant, Quinn Canyon, Schell Creek ranges)
76 USA
Synonyms: hidden
= Lesquerella confluens (Maguire & A.H.Holmgren) Reveal; Great Basin Naturalist 30: 96 (1970)
= Lesquerella hitchcockii subsp. confluens Maguire & A.H.Holmgren; Madroño 11: 174 (1951)
= Physaria hitchcockii var. confluens (Maguire & A.H.Holmgren) N.H.Holmgren; Brittonia 56: 247 (2004)
subspecies :
subsp. rubicundula
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
Name: Tum bladderpod
Distribution: USA (Utah: Aquarius, Markagunt, Paunsaugunt plateaus)
76 USA
Synonyms: hidden
= Lesquerella hitchcockii subsp. rubicundula (Rollins) Maguire & A.H.Holmgren; Madroño 11: 175 (1951)
= Lesquerella rubicundula Rollins; Contr. Dudley Herb. 3: 178 (1941)
= Physaria hitchcockii var. rubicundula (Rollins) B.L.Turner; Phytologia 86(3): 204 (2004)
= Physaria rubicundula (Rollins) S.L.Welsh; Utah Fl., ed. 3: 291 (2003)
species :
Physaria humilis
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
Name: St. Marys Peak bladderpod
Distribution: USA (SW-Montana: peaks of the Bitterroot Mts.)
73 USA
Synonyms: hidden
= Lesquerella humilis Rollins; Contr. Gray Herb. 214: 9 (1984)
species :
Physaria inflata
(Rollins & E.A.Shaw) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
Distribution: Mexico (Nuevo Leon, Coahuila)
79 MEX
Synonyms: hidden
= Lesquerella inflata Rollins & E.A.Shaw; Gen. Lesquerella 150 (1973)
species :
Physaria integrifolia
(Rollins) Lichvar; Madroño 31(4): 203 (1984)
Name: Snake River twinpod
Distribution: USA (Montana, W-Wyoming, Idaho)
73 USA
Synonyms: hidden
= Physaria didymocarpa var. integrifolia Rollins; Rhodora 41: 407 (1939)
= Physaria integrifolia var. monticola Lichvar; Madroño 331(4): 204 (1984)
species :
Physaria intermedia
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
Name: Mid bladderpod
Distribution: USA (NC-New Mexico)
Error
Remarks: many specimens formerly treated under this name are now segregated as P. fallax
Synonyms: hidden
= Lesquerella alpina var. intermedia S.Watson; Proc. Amer. Acad. Arts 23: 251 (1888)
= Lesquerella intermedia (S.Watson) A.Heller; Pl. World 1: 22 (1897)
species :
Physaria johnstonii
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
Distribution: Mexico (Coahuila)
79 MEX
Synonyms: hidden
= Lesquerella johnstonii Rollins; Contr. Dudley Herb. 3: 179 (1941)
species :
Physaria kingii
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
subspecies :
subsp. kingii
Name: King bladderpod
Distribution: USA (California, Idaho, Nevada, Oregon)
73 USA 76 USA
Synonyms: hidden
= Lesquerella kingii (S.Watson) S.Watson; Proc. Amer. Acad. Arts 23: 251 (1888)
= Vesicaria kingii S.Watson; Proc. Amer. Acad. Arts 20: 353 (1885)
subspecies :
subsp. bernardina
(Munz) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
Distribution: USA (California: Bear Lake in the San Bernardino Mts.)
76 USA
Synonyms: hidden
= Lesquerella bernardina Munz; Bull. S. Calif. Acad. Sci. 31: 62 (1932)
= Lesquerella kingii subsp. bernardina (Munz) Munz; Aliso 4: 91 (1958)
= Physaria kingii var. bernardina (Munz) B.L.Turner; Phytologia 86(3): 204 (2004)
subspecies :
subsp. cobrensis
(Rollins & E.A.Shaw) O´Kane & Al-Shehbaz; Novon 12(3): 324 (2002)
Distribution: USA (Idaho, Nevada, Oregon)
73 USA 76 USA
Synonyms: hidden
= Lesquerella kingii var. cobrensis Rollins & E.A.Shaw; Gen. Lesquerella 255 (1973)
= Physaria cobrensis (Rollins & E.A.Shaw) N.H.Holmgren; Brittonia 56(3): 247 (2004)
= Physaria kingii var. cobrensis (Rollins & E.A.Shaw) B.L.Turner; Phytologia 86(3): 205 (2004)
subspecies :
subsp. diversifolia
(Greene) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Distribution: USA (Oregon: Elkhorn, Wallowa Mts.)
73 USA
Synonyms: hidden
= Lesquerella diversifolia Greene; Pittonia 4: 309 (1901)
= Lesquerella kingii subsp. diversifolia (Greene) Rollins & E.A.Shaw; Gen. Lesquerella 255 (1973)
= Lesquerella kingii var. sherwoodii (M.Peck) C.L.Hitchc.; Vasc. Pl. Pacif. N. W. [C.L. Hitchcock et al.] 2: 523 (1964)
= Lesquerella occidentalis subsp. diversifolia (Greene) Maguire & A.H.Holmgren; Madroño 11: 178 (1951)
= Lesquerella occidentalis var. diversifolia (Greene) C.L.Hitchc.; Vasc. Pl. Pacif. N. W. [C.L. Hitchcock et al.] 2: 525 (1964)
= Lesquerella sherwoodii M.Peck; Proc. Biol. Soc. Washington 47: 186 (1934)
= Physaria kingii var. diversifolia (Greene) B.L.Turner; Phytologia 86(3): 205 (2004)
subspecies :
subsp. kaibabensis
(Rollins) O´Kane; Novon 17: 380 (2007)
Name: Kaibab bladderpod
Distribution: USA (Arizona: Kaibab Plateau)
76 USA
Synonyms: hidden
= Lesquerella kaibabensis Rollins; Contr. Gray Herb. 211: 110 (1982)
= Physaria kaibabensis (Rollins) N.H.Holmgren; Brittonia 56(3): 247 (2004)
subspecies :
subsp. latifolia
(A.Nelson) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Ward´s bladderpod
Distribution: USA (Arizona, California, Nevada, Utah); Mexico (Baja California Norte)
76 USA 79 MEX
Synonyms: hidden
= Lesquerella barnebyi Maguire; Leafl. W. Bot. 6: 73 (1950)
= Lesquerella kingii subsp. latifolia (A.Nelson) Rollins & E: A.Shaw; Gen. Lesquerella 257 (1973)
= Lesquerella kingii var. parvifolia (Maguire & A.H.Holmgren) S.L.Welsh & Reveal; Great Basin Naturalist 37: 339 ((1978 publ. 1977))
= Lesquerella latifolia A.Nelson; Bot. Gaz. 42: 49 (1906)
= Lesquerella occidentalis var. parvifolia Maguire & A.H.Holmgren; Madroño 11: 179 (1951)
= Lesquerella wardii S.Watson; Proc. Amer. Acad. Arts 23: 252 (1888)
= Physaria kingii var. parvifolia (Maguire & A.H.Holmgren) S.L.Welsh; Utah Fl., ed. 3: 289 (2003)
= Physaria wardii (S.Watson) O´Kane & Al-Shehbaz; Novon 12: 328 (2002)
subspecies :
subsp. utahensis
(Rydb.) O´Kane; Novon 17: 380 (2007)
Name: Utah bladderpod
Distribution: USA (Utah: Uinta, Wasatch Mts.)
76 USA
Synonyms: hidden
= Lesquerella utahensis Rydb.; Bull. Torrey Bot. Club 252 (1903)
= Physaria utahensis (Rydb.) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
species :
Physaria klausii
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Rogers Pass bladderpod
Distribution: USA (Montana)
73 USA
Synonyms: hidden
= Lesquerella klausii Rollins; Contr. Gray Herb. 214: 10 (1984)
species :
Physaria lata
(Wooton & Standl.) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Lincoln County bladderpod
Distribution: USA (New Mexico: Lincoln Co.)
76 USA 77 USA
Synonyms: hidden
= Lesquerella lata Wooton & Standl.; Contr. U.S. Natl. Herb. 16: 126 (1913)
species :
Physaria lateralis
O´Kane & Al-Shehbaz; Novon 14(2): 198 (2004), nom. nov.
Distribution: Argentina (Cordoba, Mendoza, Neuquen, San Luis)
85 ARG
Synonyms: hidden
= Lesquerella mendocina var. microcarpa O.E.Schulz; Notizbl. Bot. Gart. Berlin-Dahlem 12: 41 (1934)
species :
Physaria lepidota
Rollins; Brittonia 33(3): 335 (1981)
subspecies :
subsp. lepidota
Name: Kane County twinpod
Distribution: USA (C-Utah)
76 USA
subspecies :
subsp. membranacea
(Rollins) O´Kane; Novon 17: 531 (2007)
Distribution: USA (SC-Utah)
76 USA
Synonyms: hidden
= Physaria chambersii var. membranacea Rollins; Rhodora 41: 405, plate 556, figs. 17, 18 (1939)
= Physaria lepidota var. membranacea (Rollins) Rollins; Brittonia 33(3): 338 (1981)
species :
Physaria lesicae
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Pryor Mountains bladderpod
Distribution: USA (Montana)
73 USA
Synonyms: hidden
= Lesquerella lesicae Rollins; Novon 5(1): 71 (1995)
species :
Physaria lindheimeri
(A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Lindheimer´s bladderpod
Distribution: USA (S-Texas); Mexico (Tamaulipas)
77 USA 79 MEX
Synonyms: hidden
= Alyssum lindheimeri (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella gracilis var. pilosa Lundell; Wrightia 5(9): 331 (1977)
= Lesquerella lindheimeri (A.Gray) S.Watson; Proc. Amer. Acad. Arts 23: 253 (1888)
= Vesicaria lindheimeri A.Gray; Boston J. Nat. Hist. 6: 145 (1850)
species :
Physaria ludoviciana
(Nutt.) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Foothill bladderpod
Distribution: USA (Arizona, California, Colorado, Illinois, Iowa, Kansas, Minnesota, Montana, Nebraska, Nevada, New Mexico, North Dakota, Oklahoma, South Dakota, Utah, Wisconsin, Wyoming)
73 USA 74 USA 75 USA 76 USA 77 USA
Synonyms: hidden
= Alyssum ludovicianum Nutt.; Gen. N. Amer. Pl. [Nuttall] 2: 63 (1818), not A. argenteum Vitman 1790
= Lesquerella argentea (Pursh) MacMill.; Metasp. Minnesota Valley 203 (1892) (non (Schauer) S. Watson 1888)
= Lesquerella ludoviciana (Nutt.) S.Watson; Proc. Amer. Acad. Arts 23: 252 (1888)
= Myagrum argenteum Pursh; Fl. Amer. Sept. [Pursh] 2: 434 (1814)
= Physaria argentea MacMill.; Metasp. Minnesota Valley 263 (1892)
= Vesicaria ludoviciana (Nutt.) DC.; Syst. Nat. [Candolle] 2: 297 (1821)
species :
Physaria macrocarpa
(A.Nelson) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Largefruit bladderpod
Distribution: USA (Wyoming: Sweetwater Co.)
73 USA
Synonyms: hidden
= Lesquerella macrocarpa A.Nelson; Bot. Gaz. 34: 366 (1902)
species :
Physaria mcvaughiana
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: McVaugh´s bladderpod
Distribution: USA (Texas); Mexico (Chihuahua, Coahuila)
77 USA 79 MEX
Synonyms: hidden
= Lesquerella mcvaughiana Rollins; Contr. Gray Herb. 171: 44 (1950)
species :
Physaria mendocina
(Phil.) O´Kane & Al-Shehbaz; Novon 14(2): 197 (2004)
Distribution: Uruguay (Paysandu); Argentina (Buenos Aires, Chubut, Cordoba, La Pampa, Mendoza, Neuquen, Rio Negro, San Luis)
85 ARG URU
Synonyms: hidden
= Alyssum mendocinum (Phil.) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella mendocina (Phil.) Kurtz; Revista Mus. La Plata 5: 284 (1883)
= Lesquerella mendocina f. nana O.E.Schulz; Notizbl. Bot. Gart. Berlin-Dahlem 11: 875 (1933)
= Lesquerella montevidensis (Eichler) S.Watson; Proc. Amer. Acad. Arts 22: 251 (1888)
= Vesicaria arctica Barnéoud; Fl. Chil. [Gay] 1(2): 161 (1846), non (Wormsk. ex Hornem.) Richardson
= Vesicaria mendocina Phil.; Linnaea 33: 12 (1864 publ. 1865)
= Vesicaria montevidensis Eichler; Fl. Bras. [Martius] 13(1): 302 (1865)
species :
Physaria mexicana
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Distribution: Mexico (Coahuila, Nuevo Leon)
79 MEX
Synonyms: hidden
= Lesquerella mexicana Rollins; Bol. Soc. Bot. México 23: 45 (1958)
species :
Physaria mirandiana
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Distribution: Mexico (Coahuila, Nuevo Leon, San Luis Potosi, Tamaulipas, Zacatecas)
79 MEX
Synonyms: hidden
= Lesquerella mirandiana Rollins; Bol. Soc. Bot. México 23: 43 (1958)
species :
Physaria montana
(A.Gray) Greene; Fl. Francisc. 2: 249 (1891)
Name: Mountain bladderpod
Distribution: USA (Arizona, Colorado, Nebraska, New Mexico, South Dakota, Wyoming)
73 USA 74 USA 76 USA 77 USA
Synonyms: hidden
= Lesquerella montana (A.Gray) S.Watson; Proc. Amer. Acad. Arts 22: 251 (1888)
= Lesquerella montana var. suffruticosa Payson; Ann. Missouri Bot. Gard. 8: 200 (1921 publ. 1922)
= Lesquerella rosulata A.Nelson; Bull. Torrey Bot. Club 25: 205 (1898)
= Lesquerella shearis Rydb.; Bull. Torrey Bot. Club 29: 237 (1902)
= Vesicaria montana A.Gray; Proc. Acad. Nat. Sci. Philadelphia 15: 58 (1863 publ. 1864)
species :
Physaria multiceps
(Maguire) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Manyhead bladderpod
Distribution: USA (Idaho, Utah, Wyoming)
73 USA 76 USA
Synonyms: hidden
= Lesquerella multiceps Maguire; Amer. Midl. Naturalist 27: 465 (1942)
species :
Physaria navajoensis
(O´Kane) O´Kane & Al-Shehbaz; Novon 12(3): 325 (2002)
Name: Navajo bladderpod
Distribution: USA (New Mexico, Arizona)
76 USA 77 USA
Synonyms: hidden
= Lesquerella navajoensis O´Kane; Madroño 46(2): 88 (1999)
species :
Physaria neeseae
S.L.Welsh & N.D.Atwood; Utah Fl., ed. 4: 335 (2008)
Distribution: USA (Utah)
76 USA
species :
Physaria nelsonii
O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002), nom. nov.
Name: Dense bladderpod
Distribution: USA (Utah, Wyoming)
73 USA 76 USA
Synonyms: hidden
= Lesquerella alpina subsp. condensata (A.Nelson) Rollins & E.A.Shaw; Gen. Lesquerella 191 (1973)
= Lesquerella alpina var. condensata (A.Nelson) C.L.Hitchc.; Vasc. Pl. Pacif. N. W. [C.L. Hitchcock et al.] 2: 522 (1964)
= Lesquerella condensata A.Nelson; Bull. Torrey Bot. Club 238 (1899), not Physaria condensata Rollins 1939
= Physaria reediana var. condensata (A.Nelson) S.L.Welsh; Utah Fl., ed. 3: 291 (2003)
species :
Physaria newberryi
A.Gray; Rep. Colorado R. [Ives] 4(Bot.): 6 (1860)
subspecies :
subsp. newberryi
Name: Newberry´s twinpod
Distribution: USA (Arizona, Nevada, New Mexico, Utah)
76 USA 77 USA
Synonyms: hidden
= Coulterina newberryi (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Physaria didymocarpa var. newberryi (A.Gray) M.E.Jones; Proc. Calif. Acad. Sci., Ser. 2, 5: 624 (1895)
= Physaria newberryi var. racemosa Rollins; Brittonia 33(3): 339 (1981)
= Vesicaria newberryi A.Gray; Rep. Colorado R. [Ives], 4(Bot.): 6 (1861)
subspecies :
subsp. yesicola
(Sivinski) O´Kane; Novon 17: 381 (2007)
Distribution: USA (New Mexico)
77 USA
Synonyms: hidden
= Physaria newberryi var. yesicola Sivinski; Sida 18(3): 673 (1999)
species :
Physaria obcordata
Rollins; J. Arnold Arbor. 64(4): 495 (1983)
Name: Dudley Bluffs twinpod
Distribution: USA (NW-Colorado: Thirteenmile Creek Tongue, the Parachute Creek Member of the Green River Formation)
73 USA
species :
Physaria obdeltata
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Name: Middle Butte bladderpod
Distribution: USA (Idaho)
73 USA
Synonyms: hidden
= Lesquerella obdeltata Rollins; Crucifer. Continental N. Amer. 640 (1993)
species :
Physaria occidentalis
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
subspecies :
subsp. occidentalis
Name: Western bladderpod
Distribution: USA (California, Idaho, Nevada, Oregon, Washington)
73 USA 76 USA
Synonyms: hidden
= Lesquerella cusickii M.E.Jones; Contr. W. Bot. 12: 2 (1908)
= Lesquerella occidentalis (S.Watson) S.Watson; Proc. Amer. Acad. Arts 22: 251 (1888)
= Lesquerella occidentalis subsp. cusickii (M.E.Jones) Maguire & A.H.Holmgren; Madroño 11: 179 (1951)
= Lesquerella occidentalis var. cusickii (M.E.Jones) C.L.Hitchc.; Vasc. Pl. Pacif. N. W. [C.L. Hitchcock et al.] 2: 525 (1964)
= Vesicaria occidentalis S.Watson; Proc. Amer. Acad. Arts 20: 353 (1885)
subspecies :
subsp. cinerascens
(Maguire & Holmgren) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Name: Tunnel Springs Mountain bladderpod
Distribution: USA (Nevada, NW-Utah)
76 USA
Synonyms: hidden
= Lesquerella goodrichii Rollins; J. Arnold Arbor. 64(4): 503 (1983)
= Lesquerella occidentalis subsp. cinerascens (Maguire & A.H.Holmgren) Rollins & E.A.Shaw; Gen. Lesquerella 225 (1973)
= Lesquerella occidentalis var. cinerascens Maguire & A.H.Holmgren; Madroño 11: 178 (1951)
= Physaria goodrichii (Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 323 (2002)
= Physaria occidentalis var. cinerascens (Maguire & A.H.Holmgren) S.L.Welsh; Utah Fl., ed. 3: 290 (2003)
= Physaria occidentalis var. goodrichii (Rollins) S.L.Welsh; Utah Fl., ed. 3: 290 (2003)
species :
Physaria okanensis
Al-Shehbaz & Prina; Novon 19(3): 284 (2009)
Distribution: Argentina (Catamarca)
85 ARG
species :
Physaria oregona
S.Watson; Proc. Amer. Acad. Arts 17: 363 (1882)
Name: Oregon twinpod
Distribution: USA (W-Idaho, E-Oregon, Washington State)
73 USA
Synonyms: hidden
= Coulterina oregona (S.Watson) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella oregona (S.Watson) Mulligan; Canad. J. Bot. 46: 528 (1968)
species :
Physaria ovalifolia
(Rydb.) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
subspecies :
subsp. ovalifolia
Name: Roundleaf bladderpod
Distribution: USA (Colorado, Kansas, Nebraska, New Mexico, Oklahoma, Texas)
73 USA 74 USA 77 USA
Synonyms: hidden
= Lesquerella engelmannii subsp. ovalifolia (Rydb.) Clarke; Brittonia 27(3): 271 (1975)
= Lesquerella ovalifolia Rydb.; Ill. Fl. N. U.S. [Britton & Brown] 2: 137 (1897)
= Lesquerella ovata Greene; Pittonia 4: 308 (1901)
subspecies :
subsp. alba
(Goodman) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Distribution: USA (NC-Kansas, S-Oklahoma)
74 USA
Synonyms: hidden
= Lesquerella engelmannii subsp. alba (Goodman) C.Clark; Brittonia 27: 270 (1975)
= Lesquerella ovalifolia subsp. alba (Goodman) Rollins & E: A.Shaw; Gen. Lesquerella 127 (1973)
= Lesquerella ovalifolia var. alba Goodman; Rhodora 38: 239 (1936)
= Physaria ovalifolia var. alba (Goodman) B.L.Turner; Phytologia 86: 205 (2004)
species :
Physaria pachyphylla
O´Kane & Grady; Novon 17: 187, fig. 4 (2007)
Distribution: USA (S-Montana: Pryor Mountain Desert)
73 USA
species :
Physaria pallida
(Torr. & A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Name: White bladderpod
Distribution: USA (Texas: Weches Formation in San Augustine Co.)
77 USA
Conservation Status: endangered
Synonyms: hidden
= Alyssum pallidum (Torr. & A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella pallida (Torr. & A.Gray) S.Watson; Proc. Amer. Acad. Arts 22: 253 (1888)
= Vesicaria grandiflora var. pallida Torr. & A.Gray; Fl. N. Amer. [Torrey & A.Gray] 1: 101 (1838)
= Vesicaria pallida (Torr. & A.Gray) Torr. & A.Gray; Fl. N. Amer. [Torrey & A.Gray] 1: 668 (1840)
species :
Physaria palmeri
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Distribution: Mexico (Baja California Norte, Sonora)
79 MEX
Synonyms: hidden
= Lesquerella palmeri Watson; Proc. Amer. Acad. Arts 22: 252 (1888)
species :
Physaria parviflora
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Name: Piceance bladderpod
Distribution: USA (Colorado: Parachute Creek Member of the Green River Formation, Rio Blanco Co.)
73 USA
Synonyms: hidden
= Lesquerella parviflora Rollins; J. Arnold Arbor. 64(4): 506 (1983)
species :
Physaria parvula
(Greene) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Name: Pygmy bladderpod
Distribution: USA (NC-Colorado to NE-Utah, and, SW-Wyoming)
73 USA
Synonyms: hidden
= Lesquerella alpina subsp. parvula (Greene) Rollins & E: A.Shaw; Genus Lesquerella N. America 189 (1973)
= Lesquerella alpina var. parvula (Greene) S.L.Welsh & Reveal; Great Basin Naturalist 37: 337 (1977 publ. 1978)
= Lesquerella parvula Greene; Pittonia 4: 308 (1901)
species :
Physaria pendula
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 326 (2002)
Name: Snake Range bladderpod
Distribution: USA (Nevada)
76 USA
Synonyms: hidden
= Lesquerella pendula Rollins; Crucifer. Continental N. Amer. 647 (1993)
species :
Physaria peninsularis
(Wiggins) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
Distribution: Mexico (Baja California Norte)
79 MEX
Synonyms: hidden
= Lesquerella peninsularis Wiggins; Contr. Dudley Herb. 1: 166 (1933)
species :
Physaria pinetorum
(Wooton & Standl.) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
subspecies :
subsp. pinetorum
Name: White Mountain bladderpod
Distribution: USA (Arizona, New Mexico)
76 USA 77 USA
Synonyms: hidden
= Lesquerella pinetorum Wooton & Standl.; Contr. U.S. Natl. Herb. 16: 126 (1913)
subspecies :
subsp. iveyana
(O´Kane, K.N.Sm. & K.A.Arp) O´Kane; Phytoneuron 2022-18: 1-2 (2022)
Distribution: USA (New Mexico: Sandia Mts.)
77 USA
Synonyms: hidden
= Physaria iveyana O´Kane, K.N.Sm. & K.A.Arp; Phytoneuron 2012-53: 1 (2012)
species :
Physaria prostrata
(A.Nelson) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
Name: Low bladderpod
Distribution: USA (SW-Wyoming, Idaho, N-Utah)
73 USA 76 USA
Synonyms: hidden
= Lesquerella prostrata A.Nelson; Bull. Torrey Bot. Club 124 (1899)
species :
Physaria pruinosa
(Greene) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
Name: Pagosa Springs bladderpod
Distribution: USA (Colorado: Archuleta Co., N-New Mexico)
73 USA 77 USA
Synonyms: hidden
= Lesquerella pruinosa Greene; Pittonia 4: 307 (1901)
species :
Physaria pueblensis
(Payson) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
Distribution: Mexico (Oaxaca, Puebla)
79 MEX
Synonyms: hidden
= Lesquerella pueblensis Payson; Ann. Missouri Bot. Gard. 8: 169 (1921 publ. 1922)
species :
Physaria pulvinata
O´Kane & Reveal; Brittonia 58: 74, fig. 1 (2006)
Distribution: USA (Colorado: Dolores, San Miguel Counties)
73 USA 76 USA
species :
Physaria purpurea
(A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
Name: Rose bladderpod
Distribution: USA (Arizona, New Mexico, Texas); Mexico (Chihuahua, Coahuila, Nuevo Leon, San Luis Potosi, Sonora, Veracruz, Zacatecas)
76 USA 77 USA 79 MEX
Synonyms: hidden
= Alyssum purpureum (A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891), nom. illeg.
= Lesquerella purpurea (A.Gray) S.Watson; Proc. Amer. Acad. Arts 22: 253 (1888)
= Lesquerella purpurea subsp. foliosa (Rollins) Rollins & E.A.Shaw; Gen. Lesquerella 114 (1973)
= Lesquerella purpurea subsp. foliosa (Rollins) Rollins & E: A.Shaw; Gen. Lesquerella 114 (1973)
= Lesquerella purpurea var. foliosa Rollins; Contr. Dudley Herb. 3: 180 (1941)
= Physaria purpurea var. foliosa (Rollins) B.L.Turner; Phytologia 86(3): 206 (2004)
= Vesicaria purpurea A.Gray; Smithsonian Contr. Knowl. 5(6): 14 (1853)
= Vesicaria purpurea var. albiflora A.Gray; Rep. Colorado R. [Ives] 4(Bot.): 4: 6 (1861)
species :
Physaria pycnantha
Grady & O´Kane; Novon 17: 188, fig. 5 (2007)
Distribution: USA (Idaho, Montana)
73 USA
species :
Physaria pygmaea
O´Kane & Al-Shehbaz; Novon 14(2): 203 (2004)
Distribution: Argentina (Jujuy, La Rioja, Tucuman)
85 ARG
species :
Physaria rectipes
(Wooton & Standl.) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
Name: Straight bladderpod
Distribution: USA (NW-New Mexico, SW-Colorado, Utah, Arizona)
73 USA 76 USA 77 USA
Synonyms: hidden
= Lesquerella rectipes Wooton & Standl.; Contr. U.S. Natl. Herb. 16: 127 (1913)
species :
Physaria recurvata
(Engelm. ex A.Gray) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
Name: Gaslight bladderpod
Distribution: USA (SC-Texas: Edwards Plateau)
77 USA
Synonyms: hidden
= Alyssum recurvatum (Engelm. ex A.Gray) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella recurvata (Engelm. ex A.Gray) S.Watson; Proc. Amer. Acad. Arts 22: 253 (1888)
= Vesicaria recurvata Engelm.; Boston J. Nat. Hist. 6: 147 (1850)
species :
Physaria reediana
O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002), nom. nov.
Name: Alpine bladderpod
Distribution: USA (Colorado, Nebraska, Wyoming)
73 USA 74 USA
Synonyms: hidden
= Alyssum alpinum (Nutt.) Kuntze; Revis. Gen. Pl. 2: 931 (1891)
= Lesquerella alpina (Nutt. ex Torr. & A.Gray) S.Watson; Proc. Amer. Acad. Arts 23: 251 (1888)
= Lesquerella alpina var. laevis (Payson) C.L.Hitchc.; Vasc. Pl. Pacif. N. W. [C.L. Hitchcock et al.] 2: 522 (1964)
= Lesquerella condensata var. laevis Payson; Ann. Missouri Bot. Gard. 8: 212 (1921 publ. 1922)
= Vesicaria alpina Nutt.; Fl. N. Amer. [Torrey & A.Gray] 1: 102 (1838), not Physaria alpina Rollins 1981
species :
Physaria rollinsii
G.A.Mulligan; Canad. J. Bot. 44: 1663 (1966)
Name: Rollins´ twinpod
Distribution: USA (Colorado)
73 USA
species :
Physaria rosei
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
subspecies :
subsp. rosei
Distribution: Mexico (Ciudad de Mexico, Hidalgo, Mexico State, Puebla, Veracruz)
79 MEX
Synonyms: hidden
= Lesquerella argentea subsp. lepidota (Rose) Rollins & E.A.Shaw; Gen. Lesquerella 159 (1973)
= Lesquerella lepidota (Rose) M.E.Jones; Contr. W. Bot. 18: 36 (1933-5)
= Lesquerella rosei Rollins; Contr. Gray Herb. 214: 23 (1984)
= Synthlipsis lepidota Rose; Contr. U.S. Natl. Herb. 8: 294 (1905)
subspecies :
subsp. perotensis
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Distribution: Mexico (Hidalgo to Veracruz)
79 MEX
Synonyms: hidden
= Lesquerella rosei var. perotensis Rollins; Contr. Gray Herb. 214: 23 (1984)
= Physaria rosei var. perotensis (Rollins) B.L.Turner; Phytologia 86(3): 206 (2004)
species :
Physaria saximontana
Rollins; Contr. Gray Herb. 214: 13 (1984)
subspecies :
subsp. saximontana
Name: Fremont County twinpod
Distribution: USA (C-Wyoming)
73 USA
subspecies :
subsp. dentata
(Rollins) O´Kane; Novon 17: 381 (2007)
Distribution: USA (Montana)
73 USA
Synonyms: hidden
= Physaria saximontana var. dentata Rollins; Contr. Gray Herb. 214: 14 (1984)
species :
Physaria schaffneri
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Distribution: Mexico (Coahuila, Durango, Zacatecas, Aguascalientes, San Luis Potosi, Guanajuato, Queretaro)
79 MEX
Synonyms: hidden
= Lesquerella schaffneri (S.Watson) S.Watson; Proc. Amer. Acad. Arts 22: 254 (1888)
= Vesicaria schaffneri S.Watson; Proc. Amer. Acad. Arts 17: 320 (1882)
species :
Physaria scrotiformis
O´Kane; Novon 17: 376, fig. 1 (2007)
Distribution: USA (Colorado: La Plata Co.)
73 USA
species :
Physaria sessilis
(S.Watson) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Name: Sessile bladderpod
Distribution: USA (SC-Texas)
77 USA
Synonyms: hidden
= Lesquerella gracilis var. sessilis S.Watson; Proc. Amer. Acad. Arts 23: 253 (1888)
= Lesquerella sessilis (S.Watson) Small; Fl. S.E. U.S. [Small] 471 (1903)
species :
Physaria sinuosa
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Distribution: Mexico (Puebla)
79 MEX
Synonyms: hidden
= Lesquerella sinuosa Rollins; Contr. Gray Herb. 214: 23 (1984)
species :
Physaria spatulata
(Rydb.) Grady & O´Kane; Novon 17: 190 (2007)
Distribution: Canada (Alberta, Saskatchewan); USA (Montana, Nebraska, North Dakota, South Dakota, Wyoming)
71 CDN 73 USA 74 USA
Synonyms: hidden
= Lesquerella alpina var. spatulata (Rydb.) Payson; Ann. Missouri Bot. Gard. 8: 210 (1922)
= Lesquerella nodosa Greene; Pittonia 4: 309 (1901)
= Lesquerella spatulata Rydb.; Contr. U.S. Natl. Herb. 3: 486 (1896)
= Physaria reediana subsp. spatulata (Rydb.) O´Kane & Al-Shehbaz; Novon 12(3): 327 (2002)
= Physaria reediana var. spatulata (Rydb.) B.L.Turner; Phytologia 86(3): 206 (2004)
species :
Physaria subumbellata
(Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Name: Parasol bladderpod
Distribution: USA (NW-Colorado, Utah, Wyoming)
73 USA 76 USA
Synonyms: hidden
= Lesquerella subumbellata Rollins; Amer. J. Bot. 26: 420 (1939)
species :
Physaria tenella
(A.Nelson) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Name: Moapa bladderpod
Distribution: USA (Arizona, California, Nevada, Utah); Mexico (Sonora)
76 USA 79 MEX
Synonyms: hidden
= Lesquerella gordonii var. sessilis S.Watson; Proc. Amer. Acad. Arts 23: 253 (1888)
= Lesquerella tenella A.Nelson; Bot. Gaz. 47: 426 (1909)
species :
Physaria thamnophila
(Rollins & E.A.Shaw) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
subspecies :
subsp. thamnophila
Name: Zapata bladderpod
Distribution: USA (Texas: Zapata Co., Starr Co.)
77 USA
Synonyms: hidden
= Lesquerella thamnophila Rollins & E.A.Shaw; Gen. Lesquerella 86 (1973)
subspecies :
subsp. loretensis
Sedio, R.J.Williams, D.M.Price, C.F.Best, A.Contr., Manhart & Pepper; J. Bot. Res. Inst. Texas 17(2): 422 (2023)
Distribution: Mexico (Tamaulipas)
79 MEX
species :
Physaria tumulosa
(Barneby) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Distribution: USA (Utah: Winsor Member of the Carmel Formation)
76 USA
Synonyms: hidden
= Lesquerella hitchcockii subsp. tumulosa Barneby; Leafl. W. Bot. 10: 313 (1966)
= Lesquerella tumulosa (Barneby) Reveal; Great Basin Naturalist 30: 97 (1970)
= Physaria rubicundula var. tumulosa (Barneby) S.L.Welsh; Utah Fl., ed. 3: 291 (2003)
species :
Physaria urbaniana
(Muschl.) O´Kane & Al-Shehbaz; Novon 14(2): 202 (2004)
Distribution: Bolivia (Tarija); Argentina (Catamarca, Jujuy, La Rioja, Mendoza, Salta, Tucuman)
83 BOL 85 ARG
Synonyms: hidden
= Alyssum urbanianum Muschl.; Bot. Jahrb. Syst. 40: 274 (1908)
= Alyssum bolivianum Muschl.; Bot. Jahrb. Syst. 40: 275 (1908)
species :
Physaria valida
(Greene) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Name: Strong bladderpod
Distribution: USA (SC-New Mexico, W-Texas)
77 USA
Synonyms: hidden
= Lesquerella valida Greene; Pittonia 4: 68 (1899)
= Lesquerella lepidota Cory; Rhodora 32: 110 (1930)
species :
Physaria vicina
(J.L.Anderson, Reveal & Rollins) O´Kane & Al-Shehbaz; Novon 12(3): 328 (2002)
Name: Uncompaghre bladderpod
Distribution: USA (Colorado)
73 USA
Synonyms: hidden
= Lesquerella vicina J.L.Anderson, Reveal & Rollins; Novon 7(1): 9 (1997)
species :
Physaria vigana
B.L.Turner; Phytologia 89(1): 75 (2007)
Distribution: Mexico (Coahuila, Nuevo Leon)
79 MEX
species :
Physaria vitulifera
Rydb.; Bull. Torrey Bot. Club 278 (1901)
Name: Roundtip twinpod
Distribution: USA (C-Colorado)
73 USA
species :
Physaria wyndii
(Rollins & E.A.Shaw) O´Kane & Al-Shehbaz; Novon 12(3): 329 (2002)
Distribution: Mexico (NW-Coahuila)
79 MEX
Synonyms: hidden
= Lesquerella wyndii Rollins & E.A.Shaw; Gen. Lesquerella 154 (1973)
