096.0000
Family
Annonaceae Juss.; Gen. Pl. [Jussieu] 283 (1789), nom. cons.
125 genera, ca. 2450 spp. Systematics following Nge et al. (2024)
Species listed in database: 2619; hybrids: 8
= Hornschuchiaceae J.Agardh; Theoria Syst. Pl. 65 (1858)
= Monodoraceae J.Agardh; Theoria Syst. Pl. 126 (1858)
096.4782
Genus
Pseuduvaria Miq.; Fl. Ned. Ind. 1: 32 (1858)
Distribution: Malesia, many New Guinea, some tropical Australia
Species listed in database: 61
= Petalolophus Schum.; Nachtr. Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 265 (1905)
= Craibella R.M.K.Saunders et al.; Syst. Bot. 29: 42 (2004)
= Oreomitra Diels; Bot. Jahrb. Syst. 49: 151 (1912)
species :
Pseuduvaria acerosa
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 49 (2006)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria aurantiaca
(Miq.) Merr.; Philipp. J. Sci., C 10: 255 (1915)
Distribution: Moluccas; New Guinea (Irian Jaya)
42 MOL NWG
Synonyms: hidden
= Asimina aurantiaca Zipp. ex Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 25 (1865)
= Orophea aurantiaca Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 25 (1865)
species :
Pseuduvaria beccarii
(Scheff.) J.Sinclair; Gard. Bull. Singapore 15: 7 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Orophea beccarii Scheff.; Ann. Jard. Bot. Buitenzorg 2: 18 (1885)
species :
Pseuduvaria borneensis
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 55 (2006)
Distribution: Borneo (Kalimantan, Sabah, Sarawak)
42 BOR
species :
Pseuduvaria brachyantha
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 59 (2006)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria bruneiensis
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 62 (2006)
Distribution: Brunei
42 BRN
species :
Pseuduvaria calliura
Airy Shaw; Bull. Misc. Inform. Kew 1939: 289 (1939)
Distribution: Borneo
42 BOR
species :
Pseuduvaria cerina
J.Sinclair; Gard. Bull. Singapore 14: 419 (1955)
Distribution: peninsular Malaysia (once from Bukit Bauk, Terengganu)
42 MLY
species :
Pseuduvaria clemensiae
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 68 (2006)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria coriacea
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 71 (2006)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria costata
(Scheff.) J.Sinclair; Gard. Bull. Singapore 15: 7 (1956)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Orophea costata Scheff.; Ann. Jard. Bot. Buitenzorg 2: 19 (1885)
species :
Pseuduvaria cymosa
(J.Sinclair) Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 76 (2006)
Distribution: peninsular Malaysia (Perak, Selangor)
42 MLY
Synonyms: hidden
= Pseuduvaria macrophylla var. cymosa J.Sinclair; Gard. Bull. Singapore 14: 410, Fig. 32 (1955)
species :
Pseuduvaria dielsiana
(Lauterb.) J.Sinclair; Gard. Bull. Singapore 14: 403 (1955)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Goniothalamus dielsianus Lauterb.; Nachtr. Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 266 (1905)
= Orophea dielsiana (Lauterb.) Diels; Bot. Jahrb. Syst. 49: 159 (1912)
species :
Pseuduvaria dolichonema
(Diels) J.Sinclair; Gard. Bull. Singapore 15: 7 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Orophea dolichonema Diels; Bot. Jahrb. Syst. 52: 184 (1915)
species :
Pseuduvaria filipes
(Lauterb. & K.Schum.) J.Sinclair; Gard. Bull. Singapore 15: 7 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Orophea filipes K.Schumach. & Lauterb.; Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 319 (1900)
species :
Pseuduvaria fragrans
Y.C.F.Su, Chaowasku & R.M.K.Saunders; Syst. Bot. 35(1): 31 (2010)
Distribution: Thailand (peninsular Thailand)
41 THA
species :
Pseuduvaria froggattii
(F.Muell.) Jessup; Austrobaileya 2(3): 227 (1986)
Distribution: Australia (Queensland: Melissa Ck, N of Cape Tribulation to Black Mtn, N of Kuranda)
50 AUS
Conservation Status: IUCN R
Synonyms: hidden
= Mitrephora froggattii F.Muell.; Australas. J. Pharm. 2: 3 (1887), et in Bot. Centralbl. 30: 324 (1887)
species :
Pseuduvaria galeata
J.Sinclair; Gard. Bull. Singapore 14: 414 (1955)
Distribution: peninsular Malaysia (?Pahang, Johor)
42 MLY
Conservation Status: IUCN I
species :
Pseuduvaria gardneri
Y.C.F.Su, Chaowasku & R.M.K.Saunders; Syst. Bot. 35(1): 33 (2010)
Distribution: Thailand
41 THA
species :
Pseuduvaria glabrescens
(Jessup) Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 88 (2006)
Distribution: Australia (Queensland: N of Kuranda to the Little Mulgrave R.)
50 AUS
Synonyms: hidden
= Pseuduvaria mulgraveana var. glabrescens Jessup; Austrobaileya 2(4): 312 (1987)
species :
Pseuduvaria glossopetala
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. 35(1): 34 (2010)
Distribution: Thailand (peninsular Thailand); peninsular Malaysia
41 THA 42 MLY
species :
Pseuduvaria grandifolia
(Warb.) J.Sinclair; Gard. Bull. Singapore 15: 7 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Mitrephora grandifolia (Warb.) Diels; Bot. Jahrb. Syst. 49: 155 (1912)
= Stelechocarpus grandifolia Warb.; Bot. Jahrb. Syst. 13: 306 (1891)
species :
Pseuduvaria hyadena
I.M.Turner; Malayan Nat. J. 62(4): 362 (-366, figs. 2-3) (2010)
Distribution: Borneo (Kalimantan)
42 BOR
species :
Pseuduvaria hylandii
Jessup; Austrobaileya 2(4): 308 (1987)
Distribution: Australia (NE-Queensland: upper catchments of the Russell and Mulgrave Rivers)
50 AUS
Conservation Status: IUCN R
species :
Pseuduvaria khaosokensis
Yoosukkee & Chaowasku; Ann. Bot. Fenn. 58(1-3): 52 (2020)
Distribution: Thailand (peninsular Thailand)
41 THA
species :
Pseuduvaria kingiana
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 94 (2006)
Distribution: peninsular Malaysia
42 MLY
species :
Pseuduvaria kwangtungensis
(P.T.Li) Qing L.Wang & B.Xue; PhytoKeys 172: 5 (2021)
Distribution: China (Hainan)
36 CHI
Synonyms: hidden
= Meiogyne kwangtungensis P.T.Li; Acta Phytotax. Sin. 14(1): 104-105, f. 1 (1976)
species :
Pseuduvaria latifolia
(Blume) Bakh.f.; Blumea 12: 61 (1963)
Distribution: Sumatra; Java
42 JAW SUM
Synonyms: hidden
= Bocagea latifolia Blume; Fl. Javae, Annonac. 89 (1830)
= Mitrephora glandulifera Boerl.; Icon. Bogor. [Boerlage] 1: t. 60 (1899)
= Orophea latifolia (Blume) Miq.; Fl. Ned. Ind. 1(2): 30 (1856)
= Pseuduvaria glandulifera (Boerl.) Merr.; Philipp. J. Sci., C 10: 255 (1915)
= Uva hasseltii (Blume) Kuntze; Revis. Gen. Pl. 1: 8 (1891)
= Uvaria hasseltii Blume; Fl. Javae, Annonac. 46, t. XXI (1830)
species :
Pseuduvaria lignocarpa
J.Sinclair; Gard. Bull. Singapore 15: 7 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria livingstoneana
Sukd.Bera & Karthig.; Phytotaxa 718(1): 72 (2025)
Distribution: Andaman Isl.
41 AND
species :
Pseuduvaria luzonensis
(Merr.) Y.C.F.Su & R.M.K.Saunders; Blumea 46(3): 590 (2001)
Distribution: Philippines (Luzon)
42 PHI
Synonyms: hidden
= Orophea luzonensis Merr.; Philipp. J. Sci., C 3: 224 (1908), as ´luzoniensis´
= Orophea maculata Merr.; Publ. Bur. Sci. Gov. Lab. 35: 11 (1906), nom. illeg., non O. maculata Scort. ex King (1892)
= Pseuduvaria grandiflora Merr.; Philipp. J. Sci. 14: 392 (1919)
species :
Pseuduvaria macgregorii
Merr.; Philipp. J. Sci. 30: 392 (1926)
Distribution: Philippines (Samar)
42 PHI
species :
Pseuduvaria macrocarpa
(Burck) Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 107 (2006)
Distribution: Moluccas; New Guinea (Irian Jaya, Papua New Guinea)
42 MOL NWG
Synonyms: hidden
= Meiogyne macrocarpa Burck; Nova Guinea 8: 430 (1911)
= Pseuduvaria versteegii (Diels) Merr.; Philipp. J. Sci., C 10: 255 (1915)
= Mitrephora versteegii Diels; Bot. Jahrb. Syst. 49: 154 (1912)
species :
Pseuduvaria macrophylla
(Oliv.) Merr.; Philipp. J. Sci., C 10: 255 (1915)
Distribution: Thailand (peninsular Thailand); peninsular Malaysia (Kedah, Penang, Kelantan, Terengganu, Perak, Pahang, Selangor, Negeri Sembilan, Melaka); Sumatra; Borneo
41 THA 42 BOR MLY SUM
Synonyms: hidden
= Mitrephora macrophylla Oliv.; Hooker´s Icon. Pl. 16: t. 1562 (1887)
= Pseuduvaria nervosa J.Sinclair; Gard. Bull. Singapore 14: 416 (1955)
species :
Pseuduvaria megalopus
(K.Schum.) Y.C.F.Su & Mols; Syst. Bot. 30(3): 501 (2005)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Petalolophus megalopus K.Schum.; Nachtr. Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 265 (1905)
species :
Pseuduvaria mindorensis
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 117 (2006)
Distribution: Philippines (Mindoro)
42 PHI
species :
Pseuduvaria mollis
(Warb.) J.Sinclair; Gard. Bull. Singapore 15: 9 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Goniothalamus mollis Warb.; Bot. Jahrb. Syst. 13: 306 (1891)
= Mitrephora mollis (Warb.) K.Schumach. & Lauterb.; Fl. Schutzgeb. Südsee [Schumann & Lauterbach] 320 (1900)
= Papualthia bracteata Diels; Bot. Jahrb. Syst. 49: 142 (1913)
= Papualthia mollis (Warb.) Diels; Bot. Jahrb. Syst. 49: 142 (1912)
species :
Pseuduvaria monticola
J.Sinclair; Gard. Bull. Singapore 14: 408 (1955)
Distribution: Thailand (peninsular Thailand); peninsular Malaysia (Pahang, Selangor)
41 THA 42 MLY
species :
Pseuduvaria mulgraveana
Jessup; Austrobaileya 2(4): 310 (1987)
Distribution: Australia (Queensland: N of Kuranda to Miriwinni near Mt Bartle Frere)
50 AUS
Conservation Status: IUCN R
species :
Pseuduvaria multiovulata
(C.E.C.Fisch.) J.Sinclair; Gard. Bull. Singapore 14: 43, 47 (1953)
Distribution: Myanmar (Taninthayi); Thailand (SW- & peninsular Thailand)
41 MYA THA
Synonyms: hidden
= Mitrephora multiovulata C.E.C.Fisch.; Bull. Misc. Inform. Kew 1926: 450 (1926)
species :
Pseuduvaria nova-guineensis
J.Sinclair; Gard. Bull. Singapore 15: 9 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria obliqua
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 131 (2006)
Distribution: Borneo (Sarawak, Kalimantan)
42 BOR
species :
Pseuduvaria oxycarpa
(Boerl. ex Koord.) Y.C.F.Su & R.M.K.Saunders; Blumea 46(3): 591 (2001)
Distribution: Sulawesi
42 SUL
Synonyms: hidden
= Mitrephora oxycarpa Boerl. ex Koord.; Meded. Lands Plantentuin 19: 335 (1898)
species :
Pseuduvaria pamattonis
(Miq.) Y.C.F.Su & R.M.K.Saunders; Blumea 46(3): 591 (2001)
Distribution: Borneo; Brunei; Philippines
42 BOR BRN PHI
Synonyms: hidden
= Orophea pamattonis Miq.; Fl. Ned. Ind. 1(2): 24 (1856)
= Hydnocarpus unonifolia Elmer; Leafl. Philipp. Bot. 5: 1779 (1913)
species :
Pseuduvaria parviflora
(Jovet-Ast) Bân; Tap Chí Sinh Hoc 16: 13 (1994)
Distribution: C-Vietnam
41 VIE
Synonyms: hidden
= Mitrephora parviflora Ast ex Jovet-Ast; Notul. Syst. (Paris) 9: 80 (1940), latine
= Mitrephora parviflora Ast; Fl. Indo-Chine [P.H.Lecomte et al.] Suppl. 1: 102 (1938), gallice
species :
Pseuduvaria parvipetala
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 143 (2006)
Distribution: Sumatra; Borneo (Sarawak, Kalimantan)
42 BOR SUM
species :
Pseuduvaria philippinensis
Merr.; Philipp. J. Sci., C 10: 256 (1915)
Distribution: Palawan; Philippines (Luzon, Samar)
42 PHI PLW
species :
Pseuduvaria phuyensis
(R.M.K.Saunders, Y.C.F.Su & Chalermglin) Y.C.F.Su & R.M.K.Saunders; Syst. Bot. 35(1): 37 (2010)
Distribution: Thailand (SW-Thailand)
41 THA
Synonyms: hidden
= Craibella phuyensis R.M.K.Saunders, Y.C.F.Su & Chalermglin; Syst. Bot. 29: 42 (2004)
species :
Pseuduvaria prainii
(King) Merr.; Philipp. J. Sci., C 10: 255 (1915)
Distribution: Andaman Isl. (South Andaman Isl., Little Andaman Isl.); Nicobar Isl. (Great Nicobar Isl.)
41 AND NCB
Conservation Status: IUCN R
Synonyms: hidden
= Mitrephora prainii King; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 61(2): 88 (1892)
species :
Pseuduvaria pulchella
(Diels) J.Sinclair; Gard. Bull. Singapore 15: 10 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Orophea pulchella Diels; Bot. Jahrb. Syst. 49: 159 (1912)
species :
Pseuduvaria reticulata
(Blume) Miq.; Fl. Ned. Ind. 1(2): 32 (1856)
Distribution: Bangladesh; Andaman Isl.; Nicobar Isl.; Myanmar (Ayeyarwady, Rakhine, Taninthayi); Thailand (peninsular Thailand); peninsular Malaysia; Sumatra; Borneo; Java; Lesser Sunda Isl. (Bali, etc.)
40 BAN 41 AND MYA NCB THA 42 BOR JAW LSI MLY SUM
Synonyms: hidden
= Mitrephora aperta Teijsm. & Binn.; Cat. Hort. Bot. Bogor. [Teijsmann & Binnendijk] 175 (1866)
= Mitrephora chrysocarpa (Miq.) Boerl.; Icon. Bogor. [Boerlage] 1(2): 141 (1899)
= Mitrephora chrysocarpa var. sakumbangensis (Miq.) Ridl.; Sarawak Mus. J. 1(3): 87 (1913)
= Mitrephora diepenhorstii (Teijsm. & Binn.) Teijsm. & Binn.; Natuurk. Tijdschr. Ned.-Indië 27: Errata, opp. p. 1 (1864)
= Mitrephora reticulata (Blume) Hook.f. & Thomson; Fl. Brit. India [J.D.Hooker] 1: 77 (1872)
= Mitrephora reticulata var. acutata Boerl.; Cat. Pl. Phan. [Boerlage] 1: 31 (1899)
= Mitrephora sumatrana (Miq.) Boerl.; Icon. Bogor. [Boerlage] 1(2): 141 (1899)
= Orophea chrysocarpa Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 24 (1865)
= Orophea chrysocarpa var. sakumbangensis Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 24 (1865)
= Orophea diepenhorstii (Teijsm. & Binn.) Scheff.; Flora 52: 302 (1869), et in Tijdschr. Ned.-Indië 31: 10 (1870)
= Orophea reticulata (Blume) Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 23 (1865)
= Orophea sumatrana Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 24 (1865)
= Pseuduvaria diepenhorstii Teijsm. & Binn.; Tijdschr. Ned.-Indië 27: 38 (1864)
= Uvaria reticulata Blume; Fl. Javae, Annonac. 50, t. 24 (1830)
species :
Pseuduvaria rugosa
(Blume) Merr.; Philipp. J. Sci., C 10: 255 (1915)
Distribution: Nicobar Isl. (Central Nicobar Isl.); Myanmar; Thailand (N-, SW-, SE & peninsular Thailand); peninsular Malaysia (Perlis, Penang, Kelantan, Perak, Pahang, Melaka); Sumatra; Borneo; Java; Lesser Sunda Isl.
41 MYA NCB THA 42 BOR JAW LSI MLY SUM
Synonyms: hidden
= Mitrephora micrantha Teijsm. & Binn.; Cat. Hort. Bot. Bogor. [Teijsmann & Binnendijk] 176 (1866)
= Mitrephora rugosa (Blume) Boerl.; Cat. Hort. Bot. Bogor. [Boerlage] 32 (1899), et in Icon. Bogor. [Boerlage] 1(2): 140 (1899)
= Orophea rugosa (Blume) Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 23 (1865)
= Orophea trachycarpa Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 23 (1865)
= Uva rugosa (Blume) Kuntze; Revis. Gen. Pl. 1: 7 (1891)
= Uvaria rugosa (Miq.) Blume; Bijdr. Fl. Ned. Ind. 12 (1825)
species :
Pseuduvaria sessilicarpa
(J.Sinclair) Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 160 (2006)
Distribution: peninsular Malaysia (Johor)
42 MLY
Synonyms: hidden
= Pseuduvaria macrophylla var. sessilicarpa J.Sinclair; Gard. Bull. Singapore 14: 412 (1955)
species :
Pseuduvaria sessilifolia
Sinclair; Gard. Bull. Singapore 15: 10 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria setosa
(King) J.Sinclair; Gard. Bull. Singapore 14: 43 (1953)
Distribution: Thailand (peninsular Thailand); peninsular Malaysia (Kedah, Kelantan, Perak, Pahang, Selangor)
41 THA 42 MLY
Synonyms: hidden
= Orophea setosa King; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 61(2): 80 (1892)
= Pseuduvaria setosa var. major J.Sinclair; Gard. Bull. Singapore 14: 406 (1955)
species :
Pseuduvaria silvestris
(Diels) J.Sinclair; Gard. Bull. Singapore 15: 13 (1956)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Oreomitra bullata Diels; Bot. Jahrb. Syst. 49: 151 (1912)
= Orophea rhytidocarpa Diels; J. Arnold Arbor. 20: 74 (1939)
= Orophea rhytidophylla Diels; Notizbl. Bot. Gart. Berlin-Dahlem 11: 82 (1931)
= Orophea silvestris Diels; Bot. Jahrb. Syst. 49: 160 (1912)
= Pseuduvaria rhytidophylla (Diels) J.Sinclair; Gard. Bull. Singapore 15: 10 (1956)
species :
Pseuduvaria subcordata
Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 170 (2006)
Distribution: New Guinea (Papua New Guinea)
42 NWG
species :
Pseuduvaria taipingensis
J.Sinclair; Gard. Bull. Singapore 14: 406 (1955)
Distribution: peninsular Malaysia (only known from Maxwell´s Hill, Perak)
42 MLY
species :
Pseuduvaria trimera
(Craib) Y.C.F.Su & R.M.K.Saunders; Syst. Bot. Monogr. 79: 174 (2006)
Distribution: China (S-Yunnan); Myanmar; Thailand (N-Thailand); Laos; Vietnam
36 CHI 41 LAO MYA THA VIE
Synonyms: hidden
= Mitrephora trimera Craib; Bull. Misc. Inform. Kew 1913: 65 (1913)
= Pseuduvaria indochinensis Merr.; J. Arnold Arbor. 19(1): 28 (1938)
species :
Pseuduvaria unguiculata
(Elmer) Y.C.F.Su & R.M.K.Saunders; Blumea 46(3): 592 (2001)
Distribution: Philippines (Mindanao, Luzon)
42 PHI
Synonyms: hidden
= Mitrephora ellipanthoides Elmer; Leafl. Philipp. Bot. 5: 1721 (1913)
= Orophea ellipanthoides (Elmer) Merr.; Enum. Philipp. Fl. Pl. 2: 167 (1923)
= Orophea unguiculata Elmer; Leafl. Philipp. Bot. 5: 1724 (1913)
= Pseuduvaria caudata Merr.; Philipp. J. Sci. 17: 251 (1920 publ. 1921)
species :
Pseuduvaria villosa
Jessup; Austrobaileya 2(4): 312 (1987)
Distribution: Australia (Queensland: N Johnstone River to Liverpool Ck)
50 AUS
Conservation Status: IUCN R