518.0000 Family Lamiaceae Martinov; Tekhno-Bot. Slovar 355 (1820), nom. cons., nom. alt.

243 genera, ca. 7800 spp.

Species listed in database: 7809; hybrids: 381

518.9402 Genus Stachys L.; Sp. Pl. [Linnaeus] 2: 580 (1753)

Distribution: temp., subtropical and trop. regions, few in trop. Africa (22), except Malesia, Australia, and New Zealand. The genus is hopelessly polyphyletic and needs splitting in 9 genera (Salmaki et al. 2019); alternatively Sideritis and Prasium would have to be included.

Species listed in database: 375; hybrids: 10

species :

Stachys acerosa Boiss.; Diagn. Pl. Orient. 7: 57 (1846)

Distribution: Iran (SW-Iran)

34 IRN

Remarks: clade 12 Otisia

species :

Stachys aculeolata Hook.f.; J. Proc. Linn. Soc., Bot. 6: 18 (1861)

Distribution: Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; D.R.Congo [Zaire]; Rwanda; Burundi; Ethiopia; Uganda; Kenya; Tanzania

22 NGA 23 BIO BUR CMN RWA ZAI 24 ETH 25 KEN TAN UGA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys aberdarica T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 639 (1923)

= Stachys aculeolata var. afromontana T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 633 (1923)

= Stachys aspericaulis Pax; Bot. Jahrb. Syst. 39: 645 (1907)

= Stachys bambuseti T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 635 (1923)

= Stachys lindblomiana T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 641 (1923)

= Stachys sidamoensis f. neumannii T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 637 (1923)

= Stachys sidamoensis Gürke; Bot. Jahrb. Syst. 36: 126 (1905)

= Stachys subrenifolia T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 644 (1923)

variant :

var. camerunensis T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 629 (1921)

Distribution: Cameroon

23 CMN

variant :

var. inermis Y.B.Harv.; Kew Bull. 51(3): 440 (1996)

Distribution: D.R.Congo [Zaire]; Burundi; Uganda; Tanzania; Malawi

23 BUR ZAI 25 TAN UGA 26 MLW

species :

Stachys adulterina Hemsl.; J. Linn. Soc., Bot. 26: 300 (1890)

Distribution: China (Hubei, Sichuan)

36 CHI

species :

Stachys aegyptiaca Pers.; Syn. Pl. [Persoon] 2: 124 (1806)

Distribution: Egypt (Eastern Desert, NE-Egypt); Israel (S-Negev Desert); Sinai peninsula (C-Sinai, S-Sinai); Jordania (S-Jordania)

20 EGY 34 ISR JOR SIN

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys affinis Fresen.; Mus. Senckenberg. 1: 91 (1833), nom. illeg.

= Stachys boveana Bois; Bull. Mus. Natl. Hist. Nat. 24: 149 (1918)

= Stachys palaestina Vahl; Symb. Bot. [Vahl] 2: 64 (1791), nom. illeg.

= Stachys pauciflora Benth.; Labiat. Gen. Spec. [Bentham] 560 (1834)

species :

Stachys aethiopica L.; Mant. Pl. 1: 82 (1767)

Distribution: Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, N-Cape Prov., W-Cape Prov., E-Cape Prov.)

26 MOZ ZIM 27 SAF

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Betonica capensis Burm.f.; Fl. Ind. [N.L.Burman] Prodr. Fl. Cap. 16 (1768)

= Stachys attenuata Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 351 (1910)

= Stachys capensis C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 530 (1845)

= Stachys fruticetorum Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1083 (1903)

= Stachys harveyi Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 350 (1910)

= Stachys hispidula Hochst.; Flora 28: 66 (1845)

= Stachys pulchella Salisb.; Prodr. Stirp. Chap. Allerton 83 (1796), nom. illeg.

= Stachys serrulata Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 549 (1833)

species :

Stachys affinis Bunge; Enum. Pl. China Bor. 51 (1833)

Name: Artichoke betony (EN), Knollen-Ziest, Japanknolle (DE)

Distribution: Germany [I]; China (Gansu, Hebei, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Xinjiang); Myanmar (Kachin, Shan); USA [I] (New York [I])

11 ger 36 CHI 41 MYA 75 usa

Synonyms: hidden

= Stachys geobombycis var. alba C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 223 (1965)

= Stachys sieboldii Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 112 (1865)

= Stachys sieboldii var. alba (C.Y.Wu & H.W.Li) H.B.Chen; Fl. Fujianica 4: 604 (1989 publ. 1990)

= Stachys sieboldii var. glabrescens C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 222 (1965)

= Stachys sieboldii var. malacotricha Hand.-Mazz.; Acta Horti Gothob. 9: 83 (1934)

= Stachys tubifera Naudin; Rev. Hort. (Paris) 59: 290 (1887), nom. inval.

species :

Stachys agraria Schltdl. & Cham.; Linnaea 5: 100 (1830)

Name: Mousesear

Distribution: Mexico (Aguascalientes, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala; Honduras; Nicaragua; Galapagos [I]

79 MEX 80 GUA HON NIC 83 gal

Synonyms: hidden

= Stachys agraria f. albiflora Benke; Rhodora 35: 45 (1933)

= Stachys camporum M.E.Jones; Contr. W. Bot. 12: 70 (1908)

= Stachys confusa Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 122 (1898)

= Stachys polysegia Briq.; Annuaire Conserv. Jard. Bot. Genève 13/14: 395 (1911)

= Stachys umbrosa Scheele; Linnaea 22: 595 (1849)

species :

Stachys ahmetsavranii Dogu & Bagci; Bangladesh J. Bot. 50(2): 320, figs. 1-2 (2021)

Distribution: Turkey

34 TUR

species :

Stachys ajugoides Benth.; Linnaea 6: 80 (1831)

Name: Bugle hedgenettle

Distribution: USA (California)

76 USA

Synonyms: hidden

= Stachys ajugoides var. cymosa Jeps.; Fl. Calif. [Jepson] 3: 427 (1943), nom. inval.

= Stachys ajugoides var. stricta Jeps.; Fl. W. Calif. [Jepson] 457 (1901)

= Stachys ajugoides var. velutina Jeps.; Fl. W. Calif. [Jepson] 457 (1901)

= Stachys cymosa A.Heller ex Jeps.; Fl. Calif. [Jepson] 3: 427 (1943), pro syn.

= Stachys ramosa A.Heller; Muhlenbergia 1: 116 (1905)

species :

Stachys alabamica B.R.Keener & L.J.Davenp.; J. Bot. Res. Inst. Texas 10(2): 318, figs, 2-4 (2016)

Distribution: USA (Alabama)

78 USA

species :

Stachys albens A.Gray; Proc. Amer. Acad. Arts 7: 387 (1868)

Name: Whitestem hedgenettle

Distribution: USA (California, Nevada [I], Utah [I])

76 USA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys albens var. juliensis Jeps.; Man. Fl. Pl. Calif. [Jepson] 877 (1925)

= Stachys lanuginosa Greene; Pittonia 3: 342 (1898)

= Stachys malacophylla Greene; Pittonia 3: 343 (1898)

= Stachys pycnostachya Torr.; U.S. Expl. Exped., Phan. Pacific N. Amer. 17: 408 (1874)

= Stachys velutina Greene; Erythea 2: 121 (1894), nom. illeg.

species :

Stachys albicaulis Lindl.; Edwards´s Bot. Reg., t. 1558 (1833)

Name: Hierba de Santa María [Chile]

Distribution: Argentina (Mendoza); Chile (Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía)

85 ARG CLE

Synonyms: hidden

= Stachys albicaulis var. candidissima Reiche; Anales Univ. Chile 122: 284 (1908)

= Stachys eriophylla Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 113 (1898)

species :

Stachys albiflora N.E.Br.; Bull. Misc. Inform. Kew 1901: 131 (1901)

Distribution: South Africa (Free State, KwaZulu-Natal)

27 SAF

species :

Stachys albotomentosa Ramamoorthy; Anales Inst. Biol. Univ. Nac. Autón. México, Bot. 54: 158 (1983 publ. 1987)

Distribution: Mexico (Guanajuato, Hidalgo, Queretaro, San Luis Potosi, Tamaulipas)

79 MEX

variant :

var. potosina B.L.Turner; Phytologia 76(5): 393 (1994)

Distribution: Mexico (San Luis Potosi)

79 MEX

species :

Stachys aleurites Boiss. & Heldr. ex Benth.; Prodr. [A.P. de Candolle] 12: 472 (1848)

Distribution: Turkey (SW-Anatolia)

34 TUR

Synonyms: hidden

= Stachys tengwalliana Rech.f.; Oesterr. Bot. Z. 99: 46 (1952)

species :

Stachys alpigena T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 641 (1923)

subspecies :

subsp. alpigena

Distribution: Rwanda; Kenya

23 RWA 25 KEN

Remarks: clade 2 Stachys s. str.

subspecies :

subsp. longipetala Sebsebe; Kew Bull. 48(2): 330 (1993)

Distribution: Ethiopia

24 ETH

species :

Stachys alpina L.; Sp. Pl. [Linnaeus] 2: 581 (1753)

subspecies :

subsp. alpina

Name: Alpen-Ziest (DE), Limestone Woundwort (EN)

Distribution: England (Wales, W-England); Belgium; Luxembourg; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland, Sachsen, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; N- & C-Spain; Andorra; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains of N-Greece); Ukraine

10 GRB 11 AUT BGM CZE GER HUN LIE LUX POL SLO SWI 12 ADO FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB 14 UKR

Remarks: clade 1 Eriostomum, germanica agg.; doubtfully native in England

Synonyms: hidden

= Eriostomum alpinum (L.) Hoffmanns. & Link ex Steud.; Nomencl. Bot. [Steudel], ed. 1, 1: 806 (1821), nom. inval.

= Galeopsis fusca Moench; Methodus [Moench] 397 (1794), nom. illeg.

= Stachys curviflora Tausch; Flora 14: 216 (1831)

= Stachys dinarica De Marb.; Acta Univ. Lund. 27(5): 61 (1892)

= Stachys fusca (Moench) Moench; Methodus [Moench], Suppl. 136 (1802)

= Stachys orientalis L.; Sp. Pl. [Linnaeus] 2: 582 (1753)

= Stachys urticifolia Tausch; Flora 14: 216 (1831)

subspecies :

subsp. dinarica Murb.; Beitr. Fl. Südbosnien [Acta Univ. Lund. 27(5)]: 61 (1891)

Distribution: Croatia; Montenegro; ?Serbia; Kosovo; Bulgaria

13 BUL CRO KOS MNT

subspecies :

subsp. macrophylla (Albov) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 277 (1974)

Distribution: Georgia [Caucasus]; Turkey (N-Anatolia); Iran

33 GEO 34 IRN TUR

Synonyms: hidden

= Stachys macrophylla Albov; Prodr. Fl. Colchic. 202 (1895)

species :

Stachys amanica P.H.Davis; Kew Bull. [6]: 108 (1951)

Distribution: Turkey (S-Anatolia)

34 TUR

Synonyms: hidden

= Stachys debilis Bornm.; Notizbl. Bot. Gart. Berlin-Dahlem 7: 35 (1917), nom. illeg.

species :

Stachys anamurensis Sümbül; Turk. J. Bot. 14(1): 57 (1990)

Distribution: Turkey

34 TUR

species :

Stachys andongensis Hiern; Cat. Afr. Pl. [Hiern] 1: 874 (1900)

Distribution: Angola

26 ANG

species :

Stachys angustifolia M.Bieb.; Fl. Taur.-Caucas. 2: 52 (1808)

Distribution: Croatia; Romania; Bulgaria; European Turkey; Greece (scattered N-Greece, south to Mt. Olimbos, Mt. Athos, Thasos); ?C-European Russia; Ukraine; Crimea

13 BUL CRO GRC ROM TUE 14 KRY UKR

Synonyms: hidden

= Stachys abrotanifolia Boiss. & Heldr.; Fl. Orient. [Boissier] 4(2): 732 (1879)

= Stachys angustifolia subsp. tenuifolia Nyman; Consp. Fl. Eur. 580 (1881)

= Stachys angustifolia var. haynaldii Janka ex Nyman; Consp. Fl. Eur. 580 (1881), nomen

= Stachys tenuifolia Janka; Oesterr. Bot. Z. 22: 180 (1872), nom. illeg.

= Stachys tenuifolia Pall. ex Link; Enum. Hort. Berol. Alt. 2: 109 (1822), nom. illeg.

species :

Stachys anisochila Vis. & Pancic; Mem. Reale Ist. Veneto Sci. 15: 13 (1870)

Distribution: Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; ?North Macedonia; S- & E-Bulgaria

13 BIH BUL KOS MNT SRB

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys anisochila var. sendteri (Beck) Nyman; Consp. Fl. Eur. Suppl. 2: 252 (1890)

= Stachys recta subsp. anisochila (Vis. & Pancic) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)

= Stachys sendteri Beck; Ann. K.K. Naturhist. Hofmus. 2: 144 (1887)

= Stachys sendteri f. versipila H.Lindb.; Öfvers. Finska Vetensk.-Soc. Förh. 48(13): 92 (1906)

= Stachys sendteri var. adenocalyx H.Lindb.; Öfvers. Finska Vetensk.-Soc. Förh. 48(13): 92 (1906)

species :

Stachys annua (L.) L.; Sp. Pl. [Linnaeus], ed. 2: 813 (1763)

subspecies :

subsp. annua

Name: Einjähriger Ziest (DE), Annual Yellow-woundwort (EN)

Distribution: England [I] (England [I], Wales [I]); Denmark [I]; Norway [I]; Sweden [I]; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin [I], Baden-Württemberg, Bayern, Hessen, +Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein [I], Saarland, +Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; N- & NW-Spain; France; Italy; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (scattered mountains of NE-, NW-, C-Greece, rare N-Peloponnisos); S-European Russia; Estonia [I]; Latvia [I]; Lithuania [I]; Belarus; C-European Russia; E-European Russia; N-European Russia; NW-European Russia; Ukraine; Crimea; Siberia (Altai, Krasnoyarsk, Tuva, W-Siberia); Russian Far East [I] (Amur [I], Primorye [I]); Kazakhstan; Uzbekistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia, S-Anatolia, SE-Anatolia, SSW-Anatolia, W-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, W-Iran); Lebanon (C-Lebanon, coastal W-Lebanon); Syria (coastal W-Syria, NW-Syria, W-Syrian Mountains); Canada [I] (New Brunswick [I]); USA [I] (Kansas [I], Massachusetts [I], Maine [I], New Jersey [I], New York [I], Pennsylvania [I], Rhode Island [I], Virginia [I])

10 den grb nor swe 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SMA SRB TUE 14 BLR est KRY lat lit RUS UKR 30 SIB 31 rfe 32 KAZ UZB 33 ARM AZE GEO NCS 34 IRN IRQ LEB SYR TUR 72 cdn 74 usa 75 usa 78 usa

Remarks: clade 12 Otisia

Synonyms: hidden

= Betonica annua L.; Sp. Pl. [Linnaeus] 2: 578 (1753)

= Betonica annua var. glabra L.; Sp. Pl. [Linnaeus] 2: 573 (1753)

= Betonica annua var. hirsuta L.; Sp. Pl. [Linnaeus] 2: 574 (1753)

= Olisia annua (L.) Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 136 (1869)

= Olisia delphinensis Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 136 (1869)

= Prasium stachydium E.H.L.Krause; Deutschl. Fl. [Sturm], ed. 2, Abbild., 11: 115 (1903), nom. illeg.

= Sideritis trinervia Gilib.; Fl. Lit. Inch. 1: 81 (1782), opus utique oppr.

= Stachys adenocalyx K.Koch; Linnaea 21: 691 (1848)

= Stachys annua var. maxima K.Koch; Linnaea 17: 299 (1844)

= Stachys balbisii Link; Enum. Hort. Berol. Alt. 2: 110 (1822)

= Stachys betonica Crantz; Stirp. Austr. Fasc., ed. 2, 2: 264 (1769), nom. illeg.

= Stachys decumbens Willd. ex Benth.; Linnaea 11: 336 (1837), nom. illeg.

= Stachys lazica Boiss.; Fl. Orient. [Boissier] 4(2): 745 (1879)

= Stachys micrantha K.Koch; Linnaea 21: 690 (1849)

= Stachys nervosa Gaterau; Descr. Pl. Montauban 107 (1789)

= Stachys procumbens Willd. ex Benth.; Linnaea 11: 336 (1837)

= Stachys pubescens Ten.; Fl. Napol. 1(Prodr.): XXXIV (1812)

= Stachys pubescens var. grandiflora Regel; Index Sem. (St. Petersburg [Petropolitanus]) 1867: 32 (1868)

subspecies :

subsp. ammophila (Boiss. & C.I.Blanche) Sam.; Ark. Bot., a. s. 5: 376 (1960)

Distribution: Turkey; Lebanon; Syria

34 LEB SYR TUR

Synonyms: hidden

= Stachys annua var. ammophila Boiss. & C.I.Blanche; Diagn. Pl. Orient., Sér. 2, 4: 43 (1859)

subspecies :

subsp. cilicica (Boiss.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 289 (1974)

Distribution: Turkey; ?Lebanon; Syria

34 SYR TUR

Synonyms: hidden

= Stachys cilicica Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 42 (1859)

variant :

var. lycaonica R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 289 (1974)

Distribution: Turkey

34 TUR

species :

Stachys antalyensis Ayasligil & Davis; Notes Roy. Bot. Gard. Edinburgh 42: 72, f. ID-F (1984)

Distribution: Turkey (SW-Anatolia)

34 TUR

species :

Stachys aperta Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 37 (1934)

Distribution: Peru; Bolivia (Chuquisaca, Cochabamba, La Paz)

83 BOL PER

species :

Stachys appalachiana D.B.Poind. & J.B.Nelson; J. Bot. Res. Inst. Texas 5(2): 405 (2011)

Distribution: USA (North Carolina)

78 USA

species :

Stachys arabica Hornem.; Hort. Bot. Hafn. [Hornemann] 2: 554 (1815)

Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon); Syria (NW-Syria, Jbel Druze); Israel (coastal W-Israel, Rift Valley, N-Israel); Jordania (S-Jordania, W-Jordania)

34 ISR JOR LEB SYR TUR

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys macrosperma Boiss.; Diagn. Pl. Orient. 12: 84 (1853)

species :

Stachys arachnoidea Codd; Bothalia 12(2): 182 (1977)

Distribution: South Africa (Limpopo, Mpulamanga); Eswatini [Swaziland]

27 SAF SWZ

species :

Stachys araucana Phil.; Anales Univ. Chile 90: 563 (1895)

Distribution: Chile

85 CLE

Remarks: provisionally accepted name

species :

Stachys arenaria Vahl; Symb. Bot. [Vahl] 2: 64 (1791)

Name: Sand-Ziest (DE)

Distribution: Italy; Sicily; Croatia; Morocco; Algeria; Tunisia

13 CRO ITA SIC 20 ALG MOR TUN

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys arenaria subsp. divaricatidens H.Lindb.; Acta Soc. Sci. Fenn., Ser. B, Opera Biol. 1(2) (Itin. Medit.): 133 (1932)

= Stachys arenaria subsp. mollis (Willd. ex Benth.) F.Gómez; Anales Jard. Bot. Madrid 58(1): 199 (2000)

= Stachys arenaria var. angustifolia (Sennen) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 19 (1979), nom. inval.

= Stachys arenaria var. divaricatidens (H.Lindb.) Maire; Bull. Soc. Hist. Nat. Afrique N. 24: 227 (1933)

= Stachys arenaria var. latifolia Maire; Mém. Soc. Sci. Nat. Maroc 21-22: 15 (1929)

= Stachys arenaria var. maroccana Batt.; Contr. Fl. Atl. 73 (1919)

= Stachys arenaria var. peduncularis (Pau) Maire; Cat. Pl. Maroc 4: 1112 (1941)

= Stachys arenaria var. villosissima Andr.; Ind. Horti Bot. Univ. Budapest 5: 22 (1941)

= Stachys diffusa Schweigg.; Enum. Pl. Hort. Regiom. 69 (1812)

= Stachys hirta Georgi; Beschr. Nation. Russ. Reich. 3(5): 1086 (1800), nom. illeg.

= Stachys macedonica D.Don ex Griseb.; Spic. Fl. Rumel. 2: 143 (1844)

= Stachys mollis Willd. ex Benth.; Linnaea 11: 335 (1837)

= Stachys peduncularis Pau; Cavanillesia 8: 111 (1937)

= Stachys prostrata Lag. ex DC.; Mém. Soc. Phys. Genève 1(2): 455 (1823)

= Stachys pseudoarenaria Sennen; Diagn. Nouv. 199 (1936)

= Stachys pseudoarenaria var. angustifolia Sennen; Diagn. Nouv. 199 (1936)

= Stachys purpurea Poir.; Encycl. [J.Lamarck et al.], Suppl. 5: 227 (1817)

= Stachys rubra Donn.Sm.; Hort. Cantabrig., ed. 4: 136 (1807)

= Stachys scordifolia Willd.; Enum. Pl. Hort. Berol. [Willdenow] 616 (1809)

= Stachys scordioides Desf.; Tabl. École Bot. 219 (1804)

species :

Stachys arenariiformis Rouy; Bull. Soc. Bot. France 37: 166 (1890)

Distribution: Romania; Bulgaria

13 BUL ROM

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys recta subsp. arenariiformis (Rouy) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)

species :

Stachys argillicola Sebsebe; Kew Bull. 48(2): 333 (1993)

subspecies :

subsp. argillicola

Distribution: Ethiopia; Kenya

24 ETH 25 KEN

subspecies :

subsp. barbisina Sebsebe; Kew Bull. 48(2): 334 (1993)

Distribution: Ethiopia; Uganda; Kenya; Tanzania

24 ETH 25 KEN TAN UGA

species :

Stachys aristata Greenm.; Proc. Amer. Acad. Arts 39: 87 (1904)

Distribution: Mexico (Jalisco, Sinaloa)

79 MEX

species :

Stachys arrecta L.H.Bailey; Gentes Herbarum 1: 43 (1920)

Distribution: China (Anhui, Henan, Hubei, Hunan, Jiangsu, Shaanxi, Shanxi, Zhejiang)

36 CHI

species :

Stachys arriagana B.L.Turner; Phytologia 77(4): 350 (1994 publ. 1995)

Distribution: Mexico (San Luis Potosi)

79 MEX

species :

Stachys arvensis (L.) L.; Sp. Pl. [Linnaeus], ed. 2: 814 (1763)

Name: Acker-Ziest (DE); Field Woundwort (EN); Hierba de gato (ES)

Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Sweden; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, +Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Austria; Poland; Slovakia; Portugal; Spain; Gibraltar; Baleares; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Sardinia; Italy; Sicily; Pantelleria; Slovenia; Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Greece (rare lowlands near coast, Kiklades, Kerkyra, Ionian Isl.); Crete; European Russia; Morocco; Algeria; Tunisia; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl., Ilhas Desertas, Porto Santo Isl.); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma); Cape Verde Isl. (Santo Antao Isl., Fogo Isl., Ilha Brava); São Tomé [I]; South Africa [I] (E-Cape Prov. [I]); Ascension Isl. [I]; St. Helena [I]; Mauritius [I]; La Réunion [I]; East Aegaean Isl. (Lesvos, Psara, Ikaria, Samos, Karpathos, Rhodos); Turkey (NW-Anatolia: Bithynia, SSW-Anatolia, SW-Anatolia); Afghanistan (Kandahar); Lebanon (coastal W-Lebanon); Syria (coastal W-Syria); Israel (coastal W-Israel); China (Fujian, Guangdong, Guangxi); Taiwan; India [c]; Sri Lanka [I]; Australia [I] (Western Australia [I], South Australia [I], Queensland [I], New South Wales [I], Victoria [I], Tasmania [I]); Lord Howe Isl. [I]; Norfolk Isl. [I]; New Zealand [I]; New Caledonia [I]; Easter Isl. [I]; Hawaii [I] (Kauai [I], Oahu [I], Molokai [I], Lanai [I], Maui [I], Hawaii Isl. [I]); Canada [I] (British Columbia [I], Nova Scotia [I]); USA [I] (California [I], District of Columbia [I], Louisiana [I], Massachusetts [I], Maine [I], New Jersey [I], New York [I], Oregon [I], Pennsylvania [I], Texas [I], Virginia [I]); Puerto Rico [I]; Guadeloupe [I]; Dominica [I]; Martinique [I]; Colombia [I]; Venezuela [I]; Ecuador [I]; Peru [I]; Bolivia [I]; Brazil [I]; Paraguay [I]; Uruguay [I]; Argentina [I]; Chile [I] (Valparaíso [I], Los Lagos [I]); Juan Fdz. Isl.

10 DEN GRB IRE SWE 11 AUT BGM GER LUX NET POL SLO SWI 12 BAL CIS COR FRA GIB POR SAR SPA 13 ALB CRO GRC ITA KOS KRI MNT NMA PTL SIC SLV SRB 14 RUS 20 ALG MOR TUN 21 AZO CNY CVI MDR 23 sao 27 saf 28 asc sth 29 mau reu 34 AFG EAI ISR LEB SYR TUR 36 CHI 38 TAI 40 ind srl 50 aus lhi nfk 51 nzl 60 nwc 61 eas 63 haw 71 cdn 72 cdn 73 usa 75 usa 76 usa 77 usa 78 usa 81 dom gdl mar pue 82 clm ven 83 bol ecu per 84 bra 85 arg cle JNF par uru

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Cardiaca arvensis (L.) Lam.; Fl. Franç. [Lamarck] 2: 383 (1779)

= Glechoma arvensis L.; Sp. Pl. [Linnaeus] 2: 578 (1753)

= Glechoma belgica L.; Sp. Pl. [Linnaeus] 2: 578 (1753)

= Glechoma marrubiastrum Vill.; Hist. Pl. Dauphiné [Villars] 2: 371 (1787)

= Leonurus arvensis (L.) Durande; Fl. Bourgogne 1: 83 (1782)

= Stachys arvensis f. albiflora H.Lindb.; Acta Soc. Sci. Fenn., Ser. B, Opera Biol. 1(2) (Itin. Medit.): 133 (1932), nom. illeg.

= Stachys arvensis f. stricta (G.Mey.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2416 (1927)

= Stachys arvensis var. albiflora Bolzon; Bull. Soc. Bot. Ital. 1893: 309 (1893)

= Stachys arvensis var. bermudiana Millsp.; Publ. Field Columbian Mus., Bot. Ser. 2: 94 (1900)

= Stachys arvensis var. lamiiformis Domin; Biblioth. Bot. 22(89): 568 (1928)

= Stachys brasiliensis Benth.; Labiat. Gen. Spec. [Bentham] 550 (1834)

= Trixago arvensis (L.) Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 102 (1809)

= Trixago arvensis (L.) Moritzi; Pfl. Schweiz [Moritzi] 312 (1832), isonym

= Trixago colorata C.Presl; Fl. Sicul. [Presl] xxxvii (1826)

= Trixago cordifolia Moench; Methodus [Moench] 398 (1794), nom. illeg.

= Trixago punctata Gilib.; Fl. Lit. Inch. 1: 73 (1782), opus utique oppr.

= Trixella arvensis (L.) Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 135 (1869)

species :

Stachys aspera Michx.; Fl. Bor.-Amer. [Michaux] 2: 5 (1803)

Name: Hyssopleaf hedgenettle

Distribution: USA (Alabama, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kentucky, Maryland, Maine, Missouri, Mississippi, North Carolina, New Jersey, New York, Ohio, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)

74 USA 75 USA 78 USA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys ambigua (A.Gray) Britton; Mem. Torrey Bot. Club 5: 288 (1894), nom. illeg.

= Stachys aspera var. glabrata Benth.; Labiat. Gen. Spec. [Bentham] 543 (1834)

= Stachys aspera var. tenuiflora C.L.Hitchc.; Trans. Acad. Sci. St. Louis 5: 513 (1891)

= Stachys grayana House; Bull. New York State Mus. Nat. Hist., Nos. 233-4: 60 (1920 publ. 1921)

= Stachys hyssopifolia var. ambigua A.Gray; Syn. Fl. N. Amer. 2: 387 (1878)

= Stachys palustris subsp. aspera (Michx.) Derv.-Sok.; Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 80(3): 89 (1975)

= Stachys palustris var. aspera (Michx.) A.Gray; Manual [A.Gray], ed. 2: 317 (1856)

= Stachys tenuifolia var. aspera (Michx.) Fernald

species :

Stachys asperata Hedge; Fl. Madagasc., Fam. 175: 28 (1998)

Distribution: Madagascar

29 MDG

species :

Stachys atherocalyx K.Koch; Linnaea 21: 691 (1849)

Distribution: Germany [I]; Serbia; Kosovo; North Macedonia; Romania; Bulgaria; NE-Greece (S-Thrakia); European Russia; Crimea; ?Northern Caucasus; Georgia [Caucasus]; Armenia; Turkey (NE-Anatolia)

11 ger 13 BUL GRC KOS NMA ROM SRB 14 KRY RUS 33 ARM GEO 34 TUR

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys acanthodonta Klokov; Fl. URSR 9: 648 (1960)

= Stachys linearifolia K.Koch; Linnaea 21: 692 (1849)

= Stachys recta subsp. atherocalyx (K.Koch) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)

= Stachys recta var. sideritidoides Boiss.; Fl. Orient. [Boissier] 4(2): 730 (1879)

= Stachys sideritidoides K.Koch; Linnaea 21: 692 (1849), nom. illeg.

species :

Stachys aucheri Benth.; Prodr. [A.P. de Candolle] 12: 487 (1848)

Distribution: Iran (SW-Iran)

34 IRN

Synonyms: hidden

= Stachys koelzii Rech.f.; Fl. Iranica [Rechinger] 150: 383 (1982)

species :

Stachys aurea Benth.; Prodr. [A.P. de Candolle] 12: 492 (1848)

Distribution: South Africa (N-Cape Prov., W-Cape Prov.)

27 SAF

Remarks: clade 12 Otisia

Synonyms: hidden

= Betonica heraclea L.; Mant. Pl. 1: 83 (1767)

= Phlomis parvifolia Burch.; Trav. S. Africa 1: 225 (1822), nom. illeg.

= Sideritis plumosa Thunb.; Prodr. Pl. Cap. 2: 95 (1800)

= Stachys hantamensis Vatke; Bot. Zeitung (Berlin) 33: 462 (1875)

= Stachys integrifolia Vahl ex Benth.; Labiat. Gen. Spec. [Bentham] 562 (1834), nom. illeg.

= Stachys teres Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 364 (1910)

species :

Stachys babunensis Micevski; Prilozi Oddel. Biol. Med. Nauki, Makedonska Akad. Nauk Umet. 13: 30 (1992)

Distribution: North Macedonia

13 NMA

species :

Stachys bakeri Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 266 (1896)

Distribution: Madagascar

29 MDG

Synonyms: hidden

= Stachys debilis Baker; J. Linn. Soc., Bot. 20: 234 (1884) (non Kunth 1817 nec Bornm. 1917)

species :

Stachys balansae Boiss. & Kotschy; Fl. Orient. [Boissier] 4(2): 722 (1879)

Distribution: Crimea; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, SE-Anatolia); Iraq (NE-Iraq); Iran (N-Iran, Iranian Aserbaijan)

14 KRY 33 ARM GEO 34 IRN IRQ TUR

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Stachys balansae f. latifolia Kusn.; Uchen. Zap. Imp. Yur´evsk. Univ. 6(3): 70 (1898)

= Stachys terekensis Knorring; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 15: 341 (1953)

species :

Stachys balensis Sebsebe; Kew Bull. 48(2): 331 (1993)

Distribution: Ethiopia

24 ETH

species :

Stachys bayburtensis R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 279 (1974)

Distribution: Turkey (NE-Anatolia)

34 TUR

species :

Stachys baytopiorum Kit Tan & Yildiz; Notes Roy. Bot. Gard. Edinburgh 45(3): 450 (1988 publ. 1989)

Distribution: Turkey

34 TUR

species :

Stachys beckeana Dörfl. & Hayek; Denkschr. Kaiserl. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 94: 186 (1918)

Distribution: Bosnia & Hercegovina

13 BIH

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys hayekii K.Malý; Glasn. Zemaljsk. Muz. Bosni Hercegovini 40: 160 (1920)

= Stachys recta subsp. hayekii (K.Malý) Bjelcic; Posebna Izd. Zemaljsk. Muz. Bosne Hercegovine 3: 18 (1974)

species :

Stachys benthamiana Boiss.; Fl. Orient. [Boissier] 4(2): 734 (1879) emend. Rech.f., Bot. Jahrb. Syst. 71: 532 (1941)

Distribution: Iraq (NE-Iraq); Iran (W-Iran)

34 IRN IRQ

Remarks: clade 12 Otisia

species :

Stachys bergii G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116(1): 40 (1989 publ. 1990), nom. nov.

Name: Berg´s hedgenettle

Distribution: USA (California)

76 USA

Synonyms: hidden

= Stachys rigida subsp. lanata Epling; Madroño 4: 270 (1938)

species :

Stachys bhattacharjeeae Ö.Güner; Feddes Repert. 133(4): 258 (2022)

Distribution: Iraq

34 IRQ

species :

Stachys biflora Hook. & Arn.; Bot. Beechey Voy. 155 (1833)

Distribution: Mexico (Chihuahua, Coahuila, Jalisco, Michoacán, Nayarit, Nuevo León, San Luis Potosi, Veracruz)

79 MEX

Synonyms: hidden

= Stachys agraria var. glabrior Benth.; Prodr. [A.P. de Candolle] 12: 479 (1848)

= Stachys grahamii Benth.; Labiat. Gen. Spec. [Bentham] 551 (1834)

= Stachys latipes Greenm.; Proc. Amer. Acad. Arts 39: 87 (1904)

species :

Stachys bigelovii A.Gray; Proc. Amer. Acad. Arts 8: 371 (1872)

Name: Rock hedgenettle

Distribution: USA (Texas); Mexico (Baja California Sur, Chihuahua, Coahuila, Colima, Durango, Jalisco, Michoacán, Nuevo León, San Luis Potosi, Sinaloa, Sonora, Zacatecas)

77 USA 79 MEX

species :

Stachys bithynica Boiss.; Diagn. Pl. Orient. 5: 28 (1844)

Distribution: European Turkey; Crimea; Transcaucasus; Turkey

13 TUE 14 KRY 34 TUR

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Stachys germanica subsp. bithynica (Boiss.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 276 (1974)

= Stachys germanica var. bithynica (Boiss.) Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)

= Stachys germanica var. procera Nábelek; Spisy Prír. Fak. Masarykovy Univ. 70: 61 (1926)

= Stachys pisidica Boiss. & Heldr.; Diagn. Pl. Orient. 12: 75 (1853)

species :

Stachys bizensis Schweinf. ex Penz.; Atti Congr. Bot. Int. Genova [1892] 356 (1893), nomen

Distribution: Eritrea

24 ERI

species :

Stachys bogotensis Kunth; Nov. Gen. Sp. [H.B.K.] 2: 309 (1818)

Distribution: Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, Meta, Nariño, Norte de Santander, Santander)

82 CLM

Synonyms: hidden

= Stachys hians Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 116 (1898)

species :

Stachys bolusii Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 343 (1910)

Distribution: South Africa (W-Cape Prov.)

27 SAF

species :

Stachys bombycina Boiss.; Diagn. Pl. Orient. 12: 79 (1853)

Distribution: Turkey (SSW-Anatolia, SW-Anatolia)

34 TUR

Remarks: clade 5 Distantes

Synonyms: hidden

= Stachys pseudobombycina Kaynak, Daskin & Yilmaz; Nordic J. Bot. 28: 341 (2010)

species :

Stachys boraginoides Schltdl. & Cham.; Linnaea 5: 100 (1830)

Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacán, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Veracruz)

79 MEX

Synonyms: hidden

= Stachys boraginoides var. glandulosa Greenm.; Proc. Amer. Acad. Arts 41: 245 (1905)

species :

Stachys brachiata Bojer ex Benth.; Labiat. Gen. Spec. [Bentham] 547 (1834)

Distribution: Madagascar

29 MDG

Synonyms: hidden

= Stachys madagascariensis Briq.; Bull. Herb. Boissier 2: 137 (1894)

= Stachys madagascariensis var. goudotii Briq.; Bull. Herb. Boissier 2: 140 (1894)

= Stachys rupicola Bojer ex Benth.; Prodr. [A.P. de Candolle] 12: 477 (1848)

species :

Stachys brachyclada de Noé ex Coss.; Ann. Sci. Nat., Bot., Sér. 4, 1: 226 (1854), et in Bull. Soc. Bot. France 2: 583 (1855)

Distribution: E-Spain; Baleares; France; N-Morocco (Rif); Algeria

12 BAL FRA SPA 20 ALG MOR

Synonyms: hidden

= Stachys brachyclada var. immaculata Maire & Wilczek; Bull. Soc. Hist. Nat. Afrique N. 32: 214 (1941)

= Stachys brachyclada var. punctata Maire; Bull. Soc. Hist. Nat. Afrique N. 26: 74 (1936)

= Stachys hirta var. parviflora de Noé ex Bory & Durieu; Expl. Sci. Algérie 1: t. 65 (1848)

species :

Stachys brahuicus (Boiss.) comb.ined.

Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts., SW-Iran); Afghanistan (Balkh, Farah, Herat, Kabul, Kandahar, Orozgan / Daykundi, Samangan); Pakistan (Baluchistan, Kalat)

34 AFG IRN 40 PAK

Remarks: clade 12 Otisia

Synonyms: hidden

= Chamaesphacos afghanicus Aitch. & Hemsl.; Trans. Linn. Soc. London, Bot., Ser. 2, 3: 97 (1886)

= Chamaesphacos brahuicus (Boiss.) Aitch. & Hemsl.; Trans. Linn. Soc. London, Bot., Ser. 2, 3: 97 (1886)

= Thuspeinanta brahuica (Boiss.) Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 229 (1896)

= Tapeinanthus brahuicus Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 29 (1859)

species :

Stachys bridgesii Benth; Labiat. Gen. Spec. [Bentham] 546 (1834)

Name: Hierba de Santa Rosa [Chile]

Distribution: Chile (Coquimbo, Metropolitana de Santiago, O´Higgins, Maule, Los Rios, Los Lagos)

85 CLE

species :

Stachys bullata Benth.; Labiat. Gen. Spec. [Bentham] 547 (1833)

Name: California hedgenettle

Distribution: USA (California)

76 USA

Synonyms: hidden

= Stachys californica Benth.; Prodr. [A.P. de Candolle] 12: 469 (1848)

= Stachys acuminata Greene; Bull. Calif. Acad. Sci. 2(7): 410 (1887)

species :

Stachys burchelliana Launert; Mitt. Bot. Staatssamml. München 7: 301 (1968)

Distribution: Namibia; South Africa (N-Cape Prov.)

27 NAM SAF

Remarks: clade 12 Otisia

Synonyms: hidden

= Phlomis micrantha Burch.; Trav. S. Africa 1: 340 (1822)

= Sideritis rugosa Thunb.; Prodr. Pl. Cap. 2: 95 (1800)

= Stachys burchellii Benth.; Labiat. Gen. Spec. [Bentham] 561 (1834), nom. illeg.

species :

Stachys burgsdorffioides (Benth.) Boiss.; Diagn. Pl. Orient. 12: 85 (1853)

subspecies :

subsp. burgsdorffioides

Distribution: Turkey (E-Anatolia, Inner Anatolia, S-Anatolia, SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); Iraq (NW-Iraq); Syria (NW-Syria)

34 IRQ SYR TUR

Synonyms: hidden

= Stachys satureoides var. burgsdorffioides Benth.; Prodr. [A.P. de Candolle] 12: 481 (1848)

subspecies :

subsp. ladanoides Hand.-Mazz.; Ann. K.K. Naturhist. Hofmus. 27: 417 (1913)

Distribution: Turkey

34 TUR

species :

Stachys buttleri R.R.Mill; Notes Roy. Bot. Gard. Edinburgh 38: 57 (1980)

Distribution: Turkey (SSW-Anatolia)

34 TUR

species :

Stachys byzantina K.Koch; Linnaea 21: 686 (1849)

Name: Wolliger Ziest (DE), Lamb´s-ear, Woolly Hedgenettle (EN)

Distribution: England [I] (England [I], Wales [I], Isle of Man [I], Scotland [I], Isles of Scilly [I]); Ireland [I] (Ireland [I], Northern Ireland [I]); Germany [I]; Slovakia [I]; Hungary [I]; Spain [I]; France [I]; Monaco [c]; Channel Isl. [I]; Slovenia [I]; Croatia [I]; European Turkey; European Russia; Siberia [I] (Altai [I]); ?Northern Caucasus; Georgia [Caucasus]; Turkey (Inner Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan); China [I]; Australia [I] (South Australia [I], Queensland [I]); Lord Howe Isl. [I]; New Zealand [I]; Canada [I] (British Columbia [I], Ontario [I], Québec [I]); USA [I] (California [I], Connecticut [I], Illinois [I], New York [I], Utah [I], Virginia [I], Vermont [I]); Peru [I]

10 grb ire 11 ger hun slo 12 cis fra mco spa 13 cro slv TUE 14 RUS 30 sib 33 GEO 34 IRN TUR 36 chi 50 aus lhi 51 nzl 71 cdn 72 cdn 74 usa 75 usa 76 usa 78 usa 83 per

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Eriostomum lanatum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)

= Stachys germanica subsp. lanata (Jacq.) Bonnier; Fl. Ill. France 16 (1927)

= Stachys lanata Jacq.; Icon. Pl. Rar. [Jacquin] 1: 11 (1781) non Crantz (1769)

= Stachys lanata var. parvifolia K.Koch; Linnaea 21: 686 (1849)

= Stachys olympica Poir.; Encycl. [J.Lamarck et al.], Suppl. 5: 226 (1817)

= Stachys taurica Zefir.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 14: 348 (1951)

species :

Stachys afra E.Mey. ex Benth.; Prodr. [A.P. de Candolle] 12: 495 (1848)

Distribution: South Africa (Limpopo, Gauteng, Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.)

27 SAF

species :

Stachys calcicola Epling; Bull. Torrey Bot. Club 71: 484 (1944)

Distribution: Guatemala; Colombia (Boyacá)

80 GUA 82 CLM

species :

Stachys candida Bory & Chaub.; Expéd. Sci. Morée, Bot. 3(2): 167 (1832)

Distribution: S-Greece (Mt. Taygetos and surroundings)

13 GRC

Remarks: clade 11 Swainsoniana, candida agg.

species :

Stachys canescens Bory & Chaub.; Expéd. Sci. Morée, Bot. 3(2): 167 (1832)

Distribution: Albania; S-Greece (S-Peloponnisos: SW-Messinia, Mani peninsula)

13 ALB GRC

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys messeniaca Boiss.; Prodr. [A.P. de Candolle] 12: 475 (1848)

= Stachys messeniaca var. canescens (Bory & Chaub.) Nyman; Consp. Fl. Eur. 578 (1881)

species :

Stachys carduchorum (R.Bhattacharjee) Rech.f.; Fl. Iranica [Rechinger] 150: 363 (1982)

Distribution: Turkey; Iran

34 IRN TUR

Synonyms: hidden

= Stachys balansae subsp. carduchorum R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 277 (1974)

species :

Stachys caroliniana J.B.Nelson & D.A.Rayner; J. Bot. Res. Inst. Texas 8(2): 431 (2014)

Distribution: USA (South Carolina)

78 USA

species :

Stachys cataonica R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 280 (1974)

Distribution: Turkey (Inner Anatolia)

34 TUR

species :

Stachys chamissonis Benth.; Linnaea 6: 80 (1831)

Name: Coastal hedgenettle

Distribution: USA (California)

76 USA

Synonyms: hidden

= Stachys flaccida Eastw.; Bull. Torrey Bot. Club 29: 80 (1902)

variant :

var. cooleyae (A.Heller) G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116: 41 (1989 publ. 1990)

Name: Cooleys Hedgenettle

Distribution: Alaska; Canada (British Columbia); USA (California, Oregon, Washington State)

70 ASK 71 CDN 73 USA 76 USA

Synonyms: hidden

= Stachys caurina Piper; Proc. Biol. Soc. Washington 32: 42 (1919)

= Stachys confertiflora Piper; Proc. Biol. Soc. Washington 32: 42 (1919)

= Stachys cooleyae A.Heller; Bull. Torrey Bot. Club 26: 590 (1899)

species :

Stachys chasmosericea Ayasligil & Davis; Notes Roy. Bot. Gard. Edinburgh 42: 72, f. 1A-C (1984)

Distribution: Turkey (SW-Anatolia)

34 TUR

species :

Stachys chinensis Bunge ex Benth.; Labiat. Gen. Spec. [Bentham] 544 (1834)

Distribution: Russian Far East (Primorye); China (Gansu, Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi)

31 RFE 36 CHI

Synonyms: hidden

= Stachys aspera subsp. chinensis (Bunge ex Benth.) Krestovsk.; Turczaninowia 7(4): 20 (2004)

= Stachys aspera var. chinensis (Bunge ex Benth.) Maxim.; Bull. Soc. Imp. Naturalistes Moscou 54(1): 45 (1879)

= Stachys baicalensis var. chinensis (Bunge ex Benth.) Kom.; Trudy Imp. S.-Peterburgsk. Bot. Sada 25: 370 (1907)

= Stachys chanetii H.Lév.; Repert. Spec. Nov. Regni Veg. 9: 222 (1911)

= Stachys chinensis var. albiflora C.Y.Li; Fl. Pl. Herb. Chin. Bor.-Or. 7: 256 (1981)

species :

Stachys choruhensis Kit Tan & Sorger; Pl. Syst. Evol. 155(1-4): 99 (1987)

Distribution: NE-Turkey

34 TUR

species :

Stachys chrysantha Boiss. & Heldr.; Diagn. Pl. Orient. 1(7): 56 (1846)

Distribution: S-Greece (NC- to SE-Peloponnisos)

13 GRC

Remarks: clade 11 Swainsoniana, candida agg.

Synonyms: hidden

= Stachys candida subsp. chrysantha (Boiss. & Heldr.) Nyman; Consp. Fl. Eur. 580 (1881)

species :

Stachys circinata L´Hér.; Stirp. Nov. 51 (1786)

Distribution: S-Spain; Gibraltar; Morocco; Algeria; Tunisia

12 GIB SPA 20 ALG MOR TUN

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys circinata f. hozmariensis Pau; Mem. Real Soc. Esp. Hist. Nat. 12: 376 (1924)

= Stachys circinata subsp. numidica (Pomel) Batt.; Fl. Algérie [Battandier et al.] Dicot. 704 (1890)

= Stachys circinata var. zaiana (Emb. & Maire) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 102(11): 1532 (2017)

= Stachys citrina subsp. zaiana Emb. & Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 208 (1932)

= Stachys erioleuca Pomel; Nouv. Mat. Fl. Atl. 305 (1875)

= Stachys latifolia Aiton; Hort. Kew. [W.Aiton] 2: 303 (1789)

= Stachys numidica Pomel; Nouv. Mat. Fl. Atl. 117 (1874)

= Stachys pallida Salisb.; Prodr. Stirp. Chap. Allerton 63 (1796)

= Stachys velutina Willd.; Enum. Pl. [Willdenow], Suppl. 41 (1814)

species :

Stachys citrina Boiss. & Heldr. ex Benth.; Prodr. [A.P. de Candolle] 12: 490 (1848)

subspecies :

subsp. citrina

Distribution: Turkey (Inner Anatolia, S-Anatolia, SSW-Anatolia)

34 TUR

subspecies :

subsp. chamaesideritis (Boiss. & Balansa) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 281 (1974)

Distribution: Turkey

34 TUR

Synonyms: hidden

= Stachys chamaesideritis Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 41 (1859)

species :

Stachys clingmanii Small; Fl. S.E. U.S. [Small] 1032 (1903)

Name: Clingman´s hedgenettle

Distribution: USA (Alabama, Arkansas, District of Columbia, Illinois, Indiana, Kentucky, Maryland, North Carolina, South Carolina, Tennessee, Virginia, Vermont, West Virginia)

74 USA 75 USA 78 USA

species :

Stachys coccinea Ortega; Nov. Pl. Descr. Dec. 20 (1797)

Name: Scarlet hedgenettle

Distribution: USA (Arizona, New Mexico, Texas); Mexico (Aguascalientes, Baja California Sur, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala; Nicaragua

76 USA 77 USA 79 MEX 80 GUA NIC

Synonyms: hidden

= Stachys cardinalis Kunze; Bot. Zeitung (Berlin) 2: 645 (1844)

= Stachys coccinea Jacq.; Pl. Rar. Hort. Schoenbr. 3: 18 (1798), nom. illeg.

= Stachys limitanea A.Nelson; Amer. J. Bot. 25: 115 (1938)

= Stachys oaxacana Fernald; Proc. Amer. Acad. Arts 35: 564 (1900)

species :

Stachys collina Brandegee; Univ. Calif. Publ. Bot. 3: 391 (1909)

Distribution: Mexico (Guerrero, Oaxaca, Puebla)

79 MEX

species :

Stachys comosa Codd; Bothalia 16(1): 51 (1986)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)

27 SAF

species :

Stachys cordata Riddell; W. J. Med. Phys. Sci. 9(36): 582 (1836)

Name: Heartleaf hedgenettle

Distribution: USA (Alabama, Arkansas, Georgia, Illinois, Indiana, Kentucky, Maryland, North Carolina, New Jersey, New York, Ohio, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)

74 USA 75 USA 78 USA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys nuttallii Shuttlew. ex Benth.; Prodr. [A.P. de Candolle] 12: 469 (1848)

= Stachys nuttallii var. occidentalis Benth.; Prodr. [A.P. de Candolle] 12: 469 (1848)

= Stachys palustris var. cordata A.Gray; Manual [A.Gray], ed. 2: 317 (1856)

= Stachys riddellii House; Bull. New York State Mus. Nat. Hist. 233-234: 60 (1920 publ. 1921), nom. illeg.

= Stachys salvioides Small; Fl. S.E. U.S. [Small] 1032 (1903)

= Stachys subcordata Rydb.; Torreya 27: 88 (1927)

species :

Stachys cordifolia Prain; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 59: 310 (1890 publ. 1891)

Distribution: Myanmar

41 MYA

species :

Stachys corsica Pers.; Syn. Pl. [Persoon] 2: 124 (1806)

Distribution: Corsica; Sardinia

12 COR SAR

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Glechoma grandiflora DC.; Fl. Franç. [de Candolle & Lamarck], éd. 3, 3: 538 (1805)

species :

Stachys costaricensis Briq. ex T.Durand & Pittier; Bull. Soc. Roy. Bot. Belgique 30: 240 (1891 publ. 1892)

Distribution: Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Colombia (Antioquia)

80 COS ELS GUA HON NIC PAN 82 CLM

Synonyms: hidden

= Stachys guatemalensis Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 34 (1934)

species :

Stachys crenata Raf.; Fl. Ludov. 42 (1817)

Name: Mousesear

Distribution: USA (Alabama, Arizona, Florida, Kentucky, Louisiana, Mississippi, Texas); Mexico (Coahuila)

76 USA 77 USA 78 USA 79 MEX

species :

Stachys cretica L.; Sp. Pl. [Linnaeus] 2: 581 (1753)

subspecies :

subsp. cretica

Distribution: Germany [I]; Greece (C- & S-Greece); Crete (widespread)

11 ger 13 GRC KRI

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Eriostomum creticum (L.) Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)

= Stachys albereana Neyraut & Debeaux; Rev. Bot. Bull. Mens. 10: 263 (1891)

= Stachys altissima Desf.; Tabl. École Bot., ed. 3 (Cat. Pl. Horti Paris.) 101 (1829)

= Stachys pungens Banks & Sol.; Nat. Hist. Aleppo, ed. 2 [Russell]: 2: 255 (1794)

subspecies :

subsp. anatolica Rech.f.; Ann. Naturhist. Mus. Wien 48: 175 (1937)

Distribution: Turkey

34 TUR

subspecies :

subsp. bulgarica Rech.f.; Ann. Naturhist. Mus. Wien 48: 171 (1937)

Distribution: Bulgaria; European Turkey; Greece (scattered N-Greece); Turkey

13 BUL GRC TUE 34 TUR

Synonyms: hidden

= Stachys thracica Davidov; Spis. Bulg. Akad. Nauk 12: 109 (1915)

subspecies :

subsp. cassia (Boiss.) Rech.f.; Bot. Jahrb. Syst. 71: 534 (1941)

Distribution: Croatia; ?Montenegro; North Macedonia; Albania; Bulgaria; Greece (N-Greece, Thasos, Samothraki, Chalkidiki); Turkey

13 ALB BUL CRO GRC NMA 34 TUR

Synonyms: hidden

= Sideritis cassia Boiss.; Diagn. Pl. Orient. 12: 76 (1853)

= Stachys cassia (Boiss.) Boiss.; Diagn. Pl. Orient. 13: 74 (1854)

= Stachys germanica var. cassia (Boiss.) Briq.; Lab. Alp. Mar. 227 (1893)

subspecies :

subsp. garana (Boiss.) Rech.f.; Ann. Naturhist. Mus. Wien 48: 176 (1937)

Distribution: Turkey; Iraq; Iran

34 IRN IRQ TUR

Synonyms: hidden

= Stachys garana Boiss.; Diagn. Pl. Orient. 12: 76 (1853)

= Stachys cretica var. garana (Boiss.) Boiss.; Fl. Orient. [Boissier] 4(2): 749 (1879)

= Stachys germanica var. garana (Boiss.) Briq.; Lab. Alp. Mar. 223 (1893)

subspecies :

subsp. gulendamiana Akçiçek; Phytotaxa 539(3): 258 (2022)

Distribution: Turkey (W-Anatolia)

34 TUR

subspecies :

subsp. kutahyensis Akçiçek; Turk. J. Bot. 34(2): 132 (2010)

Distribution: Turkey

34 TUR

subspecies :

subsp. lesbiaca Rech.f.; Ann. Naturhist. Mus. Wien 48: 175 (1937)

Distribution: NE-Greece (Samothraki, Limnos); East Aegaean Isl. (Lesvos, Chios, Ikaria); Turkey (W-Anatolia)

13 GRC 34 EAI TUR

subspecies :

subsp. mersinaea (Boiss.) Rech.f.; Ann. Naturhist. Mus. Wien 48: 175 (1937)

Distribution: Turkey

34 TUR

Synonyms: hidden

= Stachys mersinaea Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 37 (1859)

= Stachys germanica var. mersinaea (Boiss.) Briq.; Lab. Alp. Mar. 223 (1893)

= Stachys cretica var. paniculata Boiss.; Fl. Orient. [Boissier] 4(2): 719 (1879)

subspecies :

subsp. salviifolia (Ten.) Rech.f.; Ann. Naturhist. Mus. Wien 48: 170 (1937)

Name: Salbeiblättriger Kretischer Ziest (DE)

Distribution: Germany [I]; France; Italy; Sicily; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (Kerkyra, Lefkada, Epirus, N-Peloponnisos, Chalkidiki, Mt. Falakron)

11 ger 12 FRA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA SIC SLV SMA SRB

Synonyms: hidden

= Eriostomum salviifolium (Ten.) C.Presl; Fl. Sicul. [Presl] 37 (1826)

= Stachys germanica subsp. italica (Briq.) P.Fourn.; Quatre Fl. France 832 (1937)

= Stachys germanica subsp. salviifolia (Ten.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2427 (1927)

= Stachys germanica var. boissieri Briq.; Lab. Alp. Mar. 222 (1893)

= Stachys germanica var. italica Briq.; Lab. Alp. Mar. 224 (1893)

= Stachys salviifolia Ten.; Fl. Napol. 34 (1811)

subspecies :

subsp. smyrnaea Rech.f.; Ann. Naturhist. Mus. Wien 48: 174 (1937)

Name: Smyrna-Ziest (DE); Smyrna Woundwort (EN)

Distribution: European Turkey; East Aegaean Isl. (Chios, Ikaria, Samos, Lipsi, Leros, Kalimnos, Kos, Tilos, Simi, Rhodos); Turkey (widespread); Cyprus

13 TUE 34 CYP EAI TUR

subspecies :

subsp. trapezuntica Rech.f.; Ann. Naturhist. Mus. Wien 48: 172 (1937)

Distribution: NE-Turkey

34 TUR

subspecies :

subsp. vacillans Rech.f.; Bot. Jahrb. Syst. 71: 536 (1941)

Distribution: Turkey; Lebanon; Syria; Israel; Jordania

34 ISR JOR LEB SYR TUR

subspecies :

subsp. velata (Klokov) Greuter & Burdet; Willdenowia 15(1): 78 (1985)

Distribution: Crimea; Georgia [Caucasus]; Armenia

14 KRY 33 ARM GEO

Synonyms: hidden

= Stachys velata Klokov; Fl. URSR 9: 646 (1960)

species :

Stachys cudiensis Firat & Ö.Güner; Phytotaxa 570(1): 45 (2022)

Distribution: Turkey

34 TUR

species :

Stachys cuhacioglui Yildirim & Ö.Güner; Türl. Hab. 4(2): 100 (2023)

Distribution: Turkey

34 TUR

species :

Stachys cuneata Banks ex Benth.; Labiat. Gen. Spec. [Bentham] 560 (1833)

Distribution: South Africa (N-Cape Prov., W-Cape Prov.)

27 SAF

Synonyms: hidden

= Stachys denticulata Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 560 (1834)

species :

Stachys cydni Kotschy ex Gemici & Leblebici; Turk. J. Bot. 22(5): 359 (1998)

Distribution: Turkey

34 TUR

species :

Stachys cymbalaria Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1088 (1903)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)

27 SAF

species :

Stachys darcyana A.Pool; Novon 17(1): 61 (2007)

Distribution: Costa Rica; Panama

80 COS PAN

species :

Stachys debilis Kunth; Nov. Gen. Sp. [H.B.K.] 2: 309 (1818)

Distribution: Colombia (Boyacá, Nariño); Ecuador

82 CLM 83 ECU

Remarks: clade 2 Stachys s. str.

species :

Stachys didymantha Brenan; Mem. New York Bot. Gard. 9: 53 (1954)

Distribution: Malawi

26 MLW

species :

Stachys dinteri Launert; Mitt. Bot. Staatssamml. München 2: 313 (1957)

Distribution: Namibia

27 NAM

species :

Stachys distans Benth.; Prodr. [A.P. de Candolle] 12: 472 (1848)

Distribution: Turkey (SSW-Anatolia); Lebanon (Antilebanon, C-Lebanon, coastal W-Lebanon); Israel (coastal W-Israel, N-Israel)

34 ISR LEB TUR

Remarks: clade 5 Distantes

Synonyms: hidden

= Stachys teucriifolia Boiss. & Kotschy; Diagn. Pl. Orient., Sér. 2, 4: 40 (1859)

variant :

var. cilicica R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33(2): 287 (1974)

Distribution: Turkey; Lebanon; Syria

34 LEB SYR TUR

species :

Stachys diversifolia Boiss.; Diagn. Pl. Orient. 12: 80 (1853)

Distribution: Turkey (S-Anatolia); Lebanon (coastal W-Lebanon); Syria (W-Syrian Mountains)

34 LEB SYR TUR

species :

Stachys dregeana Benth.; Comm. Pl. Afr. Austr. [Meyer] 240 (1838)

Distribution: South Africa (Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho

27 LES SAF

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys dregeana var. lasiocalyx (Schltr.) Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 362 (1910)

= Stachys dregeana var. tenuior Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 362 (1910)

= Stachys foliosa Benth.; Comm. Pl. Afr. Austr. [Meyer] 241 (1838)

= Stachys lasiocalyx Schltr.; J. Bot. 36: 317 (1898)

= Stachys rugosa var. foliosa (Benth.) Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 359 (1910)

species :

Stachys drummondii Benth.; Labiat. Gen. Spec. [Bentham] 551 (1833)

Name: Drummond´s hedgenettle

Distribution: USA (Texas); Mexico (Morelos, Nuevo León, Puebla, San Luis Potosi, Tamaulipas)

77 USA 79 MEX

species :

Stachys durandiana Coss.; Bull. Soc. Bot. France 20: 256 (1873)

Distribution: Morocco

20 MOR

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Sideritis grantii Batt.; Contr. Fl. Atl. 67 (1919)

= Stachys durandiana var. grantii (Batt.) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 42 (1979)

= Stachys grantii (Batt.) Batt.; Bull. Soc. Hist. Nat. Afrique N. 13: 33 (1922)

species :

Stachys duriaei de Noé; Bull. Soc. Bot. France 2: 583 (1855)

Distribution: Algeria; Tunisia

20 ALG TUN

Synonyms: hidden

= Stachys argilacea Pomel; Nouv. Mat. Fl. Atl. 118 (1874)

= Stachys duriaei var. argilacea (Pomel) Batt.; Fl. Algérie [Battandier et al.] Dicot. 707 (1890)

= Stachys duriaei var. ochroleuca de Noé; Bull. Soc. Bot. France 2: 583 (1855)

= Stachys duriaei var. purpurea Batt.; Bull. Soc. Bot. France 33: 356 (1886)

= Stachys ochroleuca (de Noé) Pomel; Nouv. Mat. Fl. Atl. 119 (1874), nom. illeg.

= Stachys pulchra Pomel; Nouv. Mat. Fl. Atl. 119 (1874)

species :

Stachys ehrenbergii Boiss.; Fl. Orient. [Boissier] 4(2): 721 (1879)

Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon)

34 LEB TUR

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Leonurus mollis Ehrenb. ex Boiss.; Fl. Orient. [Boissier] 4(2): 721 (1879)

= Stachys germanica var. ehrenbergii (Boiss.) Briq.; Lab. Alp. Mar. 233 (1893)

species :

Stachys elliptica Kunth; Nov. Gen. Sp. [H.B.K.] 2: 308 (1818)

Distribution: Colombia (Arauca, Boyacá, Cundinamarca, Meta, Norte de Santander, Santander); Ecuador

82 CLM 83 ECU

Synonyms: hidden

= Stachys elliptica var. aristata Wedd.; Chlor. Andina 2: 153 (1860)

= Stachys elliptica var. grandiflora Wedd.; Chlor. Andina 2: 153 (1860)

species :

Stachys eplingii J.B.Nelson; Brittonia 31(4): 492 (1979)

Name: Epling´s hedgenettle

Distribution: USA (Alabama, Arkansas, District of Columbia, Georgia, Kentucky, North Carolina, Oklahoma, South Carolina, Tennessee, Virginia, West Virginia)

74 USA 75 USA 78 USA

species :

Stachys erectiuscula Gürke; Bot. Jahrb. Syst. 28: 315 (1900)

Distribution: South Africa (Mpulamanga, KwaZulu-Natal)

27 SAF

Synonyms: hidden

= Stachys erectiuscula var. natalensis Skan

species :

Stachys eremicola Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 30 (1934), in Repert. Spec. Nov. Regni Veg., Beih. 85: 10 (1935)

Distribution: Chile (Antofagasta, Atacama, Coquimbo)

85 CLE

Synonyms: hidden

= Stachys crenata Phil.; Anales Univ. Chile 90: 562 (1895)

species :

Stachys eriantha Benth.; Labiat. Gen. Spec. [Bentham] 549 (1833)

Distribution: Mexico (Ciudad de Mexico, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tlaxcala, Veracruz); Colombia (Boyacá, Cauca, Cundinamarca, Nariño, Norte de Santander, Santander); Venezuela (Tachira); Ecuador

79 MEX 82 CLM VEN 83 ECU

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys elliptica var. repens Wedd.; Chlor. Andina 2: 153 (1860)

= Stachys repens M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 194 (1844), nom. illeg.

species :

Stachys euadenia P.H.Davis; Kew Bull. [6]: 110 (1951)

Distribution: Turkey (SSW-Anatolia)

34 TUR

species :

Stachys euboica Rech.f.; Bot. Jahrb. Syst. 80: 391 (1961)

Distribution: Greece (S-Evvoia)

13 GRC

Remarks: provisionally accepted name, of uncertain status

species :

Stachys fendleri Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 115 (1898)

Distribution: Colombia (Cesar, Norte de Santander); Venezuela (Aragua, Venezuela Distrito Federal, Merida, Miranda, Tachira, Zulia)

82 CLM VEN

species :

Stachys filifolia Hedge; Fl. Madagasc., Fam. 175: 37 (1998)

Distribution: Madagascar

29 MDG

species :

Stachys flavescens Benth.; Comm. Pl. Afr. Austr. [Meyer] 241 (1838)

Distribution: South Africa (N-Cape Prov., W-Cape Prov.)

27 SAF

Synonyms: hidden

= Stachys gariepina Benth.; Prodr. [A.P. de Candolle] 12: 493 (1848)

species :

Stachys flexuosa Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 352 (1910)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)

27 SAF

species :

Stachys floccosa Benth.; Labiat. Gen. Spec. [Bentham] 739 (1835)

Distribution: Pakistan (Kurram, Chitral, Swat, Hazara); NW-India (Assam, Himachal Pradesh, Jammu & Kashmir, Uttarakhand)

40 IND PAK

Synonyms: hidden

= Stachys sericea Wall. ex Benth.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830), nom. illeg., non Cav. 1801

= Stachys vestita Benth.; Labiat. Gen. Spec. [Bentham] 538 (1833)

species :

Stachys floridana Shuttlew. ex Benth.; Prodr. [A.P. de Candolle] 12: 478 (1848)

Name: Florida hedgenettle

Distribution: USA (Alabama, Arkansas, California, Florida, Georgia, Louisiana, Mississippi, North Carolina, South Carolina, Tennessee, Texas, Virginia)

76 USA 77 USA 78 USA

Remarks: clade 2 Stachys s. str.

species :

Stachys fominii Sosn. ex Grossh.; Fl. Kavkaza [Grossheim] 3: 313 (1932)

Distribution: Azerbaijan

33 AZE

species :

Stachys fontqueri Pau; Mem. Real Soc. Esp. Hist. Nat. 12: 377 (1924)

Distribution: Morocco

20 MOR

species :

Stachys forsythii Hedge; Fl. Madagasc., Fam. 175: 30 (1998)

Distribution: Madagascar

29 MDG

species :

Stachys fragillima Bornm.; Bull. Herb. Boissier 7: 118 (1899)

Distribution: Iraq (NE-Iraq)

34 IRQ

species :

Stachys fruticulosa M.Bieb.; Tabl. Prov. Mer Casp. Append. 1: 15 (1798)

Distribution: Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (NE-Anatolia); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, W-Iran)

33 ARM AZE GEO 34 IRN TUR

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys araxina Kapeller; Zametki Sist. Geogr. Rast. 16: 22 (1951)

= Stachys fruticulosa subsp. grossheimii (Kapeller) Menitsky; Bot. Zhurn. (Moscow & Leningrad) 77(6): 70 (1992)

= Stachys fruticulosa var. leptophylla K.Koch; Linnaea 21: 694 (1849)

= Stachys fruticulosa var. macrocheilos (Boiss.) Boiss.; Fl. Orient. [Boissier] 3: 737 (1879)

= Stachys grossheimii Kapeller; Trudy Tbilissk. Bot. Inst. 12: 223 (1948)

= Stachys macrocheilos Boiss.; Diagn. Pl. Orient. 5: 30 (1844)

= Stachys sosnowskyi Kapeller; Zametki Sist. Geogr. Rast. 16: 20 (1951)

species :

Stachys gaziantepensis Dinc & Dogru; Proc. Natl. Acad. Sci. India, B 86(3): 631-635 (2016)

Distribution: Turkey

34 TUR

species :

Stachys geobombycis C.Y.Wu; Acta Phytotax. Sin. 10: 222 (1965)

Distribution: China (Fujian, Guangdong, Guangxi, Hubei, Hunan, Jiangxi, Zhejiang)

36 CHI

species :

Stachys germanica L.; Sp. Pl. [Linnaeus] 2: 581 (1753)

subspecies :

subsp. germanica

Name: Deutscher Ziest (DE), Downy Woundwort (EN)

Distribution: England (England); Belgium; Germany (Brandenburg, Baden-Württemberg, Bayern, Hessen, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein [I], Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; ?Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Baleares; France; Corsica; Sardinia; Italy; Sicily; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (scattered N- & C-Greece, Kerkyra, Epirus, south to S-Evvoia); C-European Russia; Ukraine; Crimea; Morocco; Canary Isl. (Tenerife); Northern Caucasus; Georgia [Caucasus]; Armenia; Turkey (N-Anatolia, NW-Anatolia: Bithynia, SW-Anatolia); Canada [I] (Ontario [I]); USA [I] (Florida [I], New Jersey [I], Ohio [I], Pennsylvania [I], Rhode Island [I], Tennessee [I], Virginia [I], West Virginia [I])

10 GRB 11 AUT BGM CZE GER HUN POL SLO SWI 12 BAL COR FRA POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SIC SLV SMA SRB TUE 14 KRY RUS UKR 20 MOR 21 CNY 33 ARM GEO NCS 34 TUR 72 cdn 75 usa 78 usa

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Betonica germanica (L.) Walther; Fl. Giessen 453 (1802)

= Eriostomum germanicum (L.) Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)

= Eriostomum lusitanicum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)

= Eriostomum polystachyum (Ten.) C.Presl; Fl. Sicul. [Presl] 37 (1826)

= Leonurus germanica (L.) Gueldenst.; Reise Russland [Gueldenstaedt] 2: 20 (1791)

= Stachys acutifolia Bory & Chaub.; Expéd. Sci. Morée, Bot. 168 (1832), nom. superfl.

= Stachys alba Mill.; Gard. Dict., ed. 8: 4 (1768)

= Stachys argentea Tausch; Flora 14: 217 (1831)

= Stachys biennis Roth; Ann. Bot. [Usteri] 14: 27 (1795)

= Stachys castellana Willk.; Suppl. Prodr. Fl. Hispan. [Willkomm & Lange] 153 (1893)

= Stachys cinerea Benth.; Labiat. Gen. Spec. [Bentham] 536 (1833)

= Stachys cordata Klokov; Fl. URSR 9: 647 (1960), nom. illeg.

= Stachys dasyanthes var. alpina Heldr. & Sart.; Exsicc. (Herb. Norm.) 1851: 347 (1851)

= Stachys elongata Benth.; Labiat. Gen. Spec. [Bentham] 536 (1833)

= Stachys excelsa Benth.; Labiat. Gen. Spec. [Bentham] 536 (1833)

= Stachys fuchsii Bubani; Fl. Pyren. [Bubani] 1: 440 (1897)

= Stachys germanica subsp. cordata Greuter & Burdet; Willdenowia 15: 78 (1985)

= Stachys germanica subsp. cordigera Briq.; Lab. Alp. Mar. 232 (1893)

= Stachys germanica subsp. lusitanica (Hoffmanns. & Link) Cout.; Fl. Portugal [Coutinho] 520 (1913)

= Stachys germanica subsp. polystachya (Ten.) Arcang.; Comp. Fl. Ital. [Arcangeli] 552 (1882)

= Stachys germanica var. alpigena K.Koch; Linnaea 21: 687 (1849)

= Stachys germanica var. alpina (Heldr. & Sart.) Briq.; Lab. Alp. Mar. 228 (1893)

= Stachys germanica var. banatica K.Koch; Linnaea 21: 687 (1849)

= Stachys germanica var. canariensis Font Quer & Svent.; Collect. Bot. (Barcelona) 2: 201 (1949)

= Stachys germanica var. lusitanica (Hoffmanns. & Link) Nyman; Consp. Fl. Eur. Suppl. 2: 251 (1889)

= Stachys germanica var. polystachya (Ten.) Nyman; Consp. Fl. Eur. 577 (1881)

= Stachys germanica var. pubescens Benth.; Edwards´s Bot. Reg. 15: t. 1289 (1829)

= Stachys germanica var. serrata Briq.; Lab. Alp. Mar. 231 (1893)

= Stachys germanica var. sublanata Fleischm. ex Nyman; Consp. Fl. Eur. 577 (1881), nomen

= Stachys germanica var. taurica K.Koch; Linnaea 21: 687 (1849)

= Stachys heraclea Webb ex Ball; J. Linn. Soc., Bot. 16: 624 (1878), nom. inval.

= Stachys heterodonta Zefir.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 14: 349 (1951)

= Stachys heterophylla Moench; Methodus [Moench], Suppl. 135 (1802), nom. illeg.

= Stachys hispanica Mill.; Gard. Dict., ed. 8: 6 (1768)

= Stachys lanata Crantz; Stirp. Austr. Fasc., ed. 2, 2: 257 (1769)

= Stachys lusitanica (Hoffmanns. & Link) Steud.; Nomencl. Bot. [Steudel], ed. 1, 1: 806 (1821)

= Stachys major Garsault; Fig. Pl. Med. t. 569 (1764), Descr. Pl. Anim., 33 (1767), Thell. in Bull. Herb. Boissier, Sér. 2, 8: 906 (1908)

= Stachys nova Sadler ex Rchb.; Fl. Germ. Excurs. 319 (1831)

= Stachys obtusata Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)

= Stachys pannonica Láng ex Ten.; Syll. Pl. Fl. Neapol. 291 (1831)

= Stachys polystachya Ten.; Fl. Napol. 1(Prodr.): XXXIV (1812)

= Stachys tomentosa Gaterau; Descr. Pl. Montauban 107 (1789), nom. illeg.

= Stachys verbascifolia Benth.; Labiat. Gen. Spec. [Bentham] 536 (1834)

subspecies :

subsp. dasyanthes (Raf.) Arcang.; Comp. Fl. Ital., ed. 2 [Arcangeli]: 437 (1894)

Distribution: Italy; Sicily

13 ITA SIC

Synonyms: hidden

= Stachys dasyanthes Raf.; Précis Découv. Somiol. 39 (1814)

= Stachys germanica var. dasyanthes (Raf.) Briq.; Lab. Alp. Mar. 231 (1893)

= Stachys germanica var. orsiniana Briq.; Lab. Alp. Mar. 227 (1893)

subspecies :

subsp. heldreichii (Boiss.) Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 285 (1929)

Distribution: Albania; Greece (NW-, NC-, C-Greece, Peloponnisos); Transcaucasus; Turkey

13 ALB GRC 34 TUR

Synonyms: hidden

= Stachys heldreichii Boiss.; Fl. Orient. [Boissier] 4(2): 721 (1879)

subspecies :

subsp. penicillata (Heldr. & Sart. ex Boiss.) Nyman; Consp. Fl. Eur. 578 (1881)

Distribution: C-Greece (rare Evvoia, Peloponnisos)

13 GRC

Synonyms: hidden

= Stachys germanica var. penicillata (Heldr. & Sart. ex Boiss.) Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)

= Stachys penicillata Heldr. & Sartori ex Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 37 (1859)

subspecies :

subsp. velezensis (Sagorski) Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 286 (1929)

Distribution: Croatia; Bosnia & Hercegovina

13 BIH CRO

Synonyms: hidden

= Stachys reinertii subsp. velezensis Sagorski; Allg. Bot. Z. Syst. 18: 15 (1912)

species :

Stachys gilliesii Benth.; Labiat. Gen. Spec. [Bentham] 740 (1833)

Distribution: Ecuador; Peru; Bolivia (Chuquisaca, Santa Cruz, Tarija); S-Brazil (Paraná, Rio Grande do Sul, Santa Catarina); Paraguay (Pres. Hayes); Uruguay (Artigas, Cerro Largo, Flores Prov., Florida Prov., Lavalleja, Paysandu, Rivera, Salto, Soriano, Treinta y Tres); Argentina (Chaco, Cordoba, Corrientes, Formosa, Jujuy, Mendoza, Tucuman); Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío)

83 BOL ECU PER 84 BRA 85 ARG CLE PAR URU

Synonyms: hidden

= Stachys boliviana Briq.; Bull. Herb. Boissier 4: 871 (1896)

= Stachys galanderi Briq.; Bull. Herb. Boissier 4: 870 (1896)

= Stachys lorentzii Briq.; Bull. Herb. Boissier 4: 869 (1896)

= Stachys micrantha Griseb.; Abh. Königl. Ges. Wiss. Göttingen 24: 275 (1879), nom. illeg.

= Stachys petiolosa Briq.; Bull. Herb. Boissier 4: 868 (1896)

= Stachys tucumanensis Briq.; Bull. Herb. Boissier 4: 872 (1896)

species :

Stachys glandulibracteata Y.B.Harv.; Kew Bull. 51(3): 452 (1996)

Distribution: Tanzania

25 TAN

species :

Stachys glandulosa Hutch. & E.A.Bruce; Bull. Misc. Inform. Kew 1941: 178 (1941)

Distribution: Somalia

24 SOM

species :

Stachys glandulosissima Floden; Phytoneuron 2016-53: 2 (2016)

Distribution: USA (Tennessee)

78 USA

species :

Stachys glechomifolia Nábelek; Spisy Prír. Fak. Masarykovy Univ. 70: 61 (1926)

Distribution: Turkey (SE-Anatolia)

34 TUR

species :

Stachys globosa Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 25 (1934)

Distribution: Mexico (Ciudad de Mexico, Hidalgo, Mexico State, Nayarit, San Luis Potosi, Veracruz)

79 MEX

species :

Stachys glutinosa L.; Sp. Pl. [Linnaeus] 2: 581 (1753)

Distribution: Corsica; Sardinia; Italy (Capraia Isl.)

12 COR SAR 13 ITA

Remarks: clade 11 Swainsoniana

species :

Stachys gossweileri G.Taylor; J. Bot. 69(Suppl. 2): 167 (1931)

Distribution: Angola

26 ANG

species :

Stachys goulimyi Rech.f.; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 94: 25 (1957)

Distribution: NC-Greece (Mt. Askio)

13 GRC

species :

Stachys graciliflora C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 530 (1845)

Distribution: South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Eswatini [Swaziland]

27 SAF SWZ

Synonyms: hidden

= Stachys cooperi Skan; Bull. Misc. Inform. Kew 1909: 420 (1910)

species :

Stachys graeca Boiss. & Heldr.; Diagn. Pl. Orient. 12: 77 (1853)

Distribution: Greece (scattered NC-, C-Greece, Peloponnisos, S-Evvoia, Attiki)

13 GRC

Remarks: clade 1 Eriostomum

Synonyms: hidden

= Stachys acutifolia Link; Linnaea 9: 575 (1835), sensu auct.

= Stachys lusitanica Bory & Chaub.; Nouv. Fl. Pélop. 37 (1838), sensu auct.

species :

Stachys grandidentata Lindl.; Bot. Reg. 13: t. 1080 (1827)

Name: Hierba santa [Chile]

Distribution: Chile (Antofagasta, Atacama, Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Lagos)

85 CLE

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys albicaulis Bertero ex Benth.; Prodr. [A.P. de Candolle] 12: 473 (1848), nom. illeg.

= Stachys closii Phil.; Anales Univ. Chile 90: 561 (1895)

= Stachys elliptica Poepp. ex Benth.; Prodr. [A.P. de Candolle] 12: 473 (1848), nom. illeg.

= Stachys serrata Clos; Fl. Chil. [Gay] 4: 500 (1849)

= Stachys toronjilcillo Phil.; Linnaea 33: 193 (1864)

species :

Stachys grandifolia E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 239 (1838)

Distribution: South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Lesotho; Eswatini [Swaziland]

27 LES SAF SWZ

Remarks: clade 2 Stachys s. str.

species :

Stachys graveolens Nábelek; Spisy Prír. Fak. Masarykovy Univ. 70: 62, t. 7, f. 3 et p. 63, f. 16 (1926)

Distribution: Iraq (NW-Iraq)

34 IRQ

species :

Stachys guyoniana de Noé ex Batt.; Fl. Algérie [Battandier et al.] Dicot. 704 (1890)

Distribution: Algeria

20 ALG

Synonyms: hidden

= Stachys guyoniana de Noé ex Munby; Cat. Pl. Algeria 24 (1890)

species :

Stachys hakkariensis Akçiçek & Firat; Phytotaxa 257(2): 168 (2016)

Distribution: Turkey

34 TUR

species :

Stachys hamata Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 49 (1934)

Distribution: Colombia (Nariño, Quindío, Tolima); Venezuela (Sucre); Ecuador

82 CLM VEN 83 ECU

Synonyms: hidden

= Stachys urticoides Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 12 (1935)

species :

Stachys harkerae J.G.González; Ann. Bot. Fenn. 52(5-6): 345-351 (2015)

Distribution: Mexico (Jalisco, Zacatecas)

79 MEX

species :

Stachys harleyana A.Pool; Novon 17(1): 63 (2007)

Distribution: Mexico (Chiapas)

79 MEX

species :

Stachys hebens Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 35 (1934)

Distribution: Colombia (Cundinamarca, Magdalena, Norte de Santander, Santander)

82 CLM

species :

Stachys heraclea All.; Fl. Pedem. 1: 31 (1785)

Distribution: Spain; France; Corsica; Italy; Sicily; +San Marino

12 COR FRA SPA 13 ITA SIC SMA

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Sideritis heraclea Colebr. ex Benth.; Prodr. [A.P. de Candolle] 12: 447 (1848)

= Stachys barbata Lapeyr.; Hist. Pl. Pyrénées 336 (1813)

= Stachys barbigera Viv.; Fl. Cors. Prodr., App. 2: 4 (1830)

= Stachys betonicifolia Pers.; Syn. Pl. [Persoon] 2: 124 (1806)

= Stachys heraclea subsp. malessana Degen & Hervier; Bull. Acad. Int. Géogr. Bot., Sér. 3, 17: 195 (1907)

= Stachys heraclea var. barbigera (Viv.) Nyman; Consp. Fl. Eur. 578 (1881)

= Stachys phlomoides Willd.; Enum. Pl. Hort. Berol. Suppl. [Willdenow] 41 (1814)

= Stachys valentina Lag.; Varied. Ci. 2(4): 39 (1805)

species :

Stachys herrerae Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 18 (1935), in Repert. Spec. Nov. Regni Veg., Beih. 85: 8 (1935)

Distribution: Peru

83 PER

species :

Stachys herrerana Rzed. & Calderón; Acta Bot. Mex. 3: 1 (1988)

Distribution: Mexico (Hidalgo, Veracruz)

79 MEX

Conservation Status: known only from type

species :

Stachys hildebrandtii Vatke; Linnaea 43: 95 (1880 publ. 1882)

Distribution: Somalia; Kenya; Tanzania

24 SOM 25 KEN TAN

Remarks: clade 12 Otisia

species :

Stachys hintoniorum B.L.Turner; Phytologia 77(4): 358 (1994 publ. 1995)

Distribution: Mexico (Coahuila, Nuevo León)

79 MEX

species :

Stachys hispida Pursh; Fl. Amer. Sept. [Pursh] 2: 407 (1813)

Distribution: Canada (Manitoba, Ontario, Québec); USA (North Dakota, Georgia, Illinois, Indiana, Iowa, Kentucky, Maine, Maryland, Missouri, New York, Ohio, Tennessee, Vermont, Wisconsin)

71 CDN 72 CDN 74 USA 75 USA 78 USA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys palustris var. hispida (Pursh) B.Boivin; Phytologia 22: 381 (1972), nom. illeg.

= Stachys pilosa subsp. hispida (Pursh) Á.Löve & D.Löve; Taxon 31: 354 (1982)

= Stachys tenuifolia var. hispida (Pursh) Fernald; Rhodora 45: 469 (1943)

species :

Stachys hissarica Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 614 (1889)

Distribution: Kazakhstan; Uzbekistan; Tajikistan; Afghanistan; Pakistan (Chitral, Swat, Hazara, Murree)

32 KAZ TZK UZB 34 AFG 40 PAK

Synonyms: hidden

= Stachys pseudofloccosa Knorring; Fl. URSS [Komarov et al., eds.] 21: 653 (1954)

species :

Stachys huber-morathii R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 278 (1974)

Distribution: Turkey (N-Anatolia)

34 TUR

species :

Stachys huetii Boiss.; Fl. Orient. [Boissier] 4(2): 722 (1879)

Distribution: Turkey (E-Anatolia)

34 TUR

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Stachys brantii A.Huet ex Boiss.; Fl. Orient. [Boissier] 4(2): 722 (1879)

= Stachys germanica var. huetii (Boiss.) Briq.; Lab. Alp. Mar. 228 (1893)

species :

Stachys huillensis Hiern; Cat. Afr. Pl. [Hiern] 1: 875 (1898)

Distribution: Angola

26 ANG

species :

Stachys humbertii Hedge; Fl. Madagasc., Fam. 175: 34 (1998)

Distribution: Madagascar

29 MDG

species :

Stachys humifusa Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 547 (1833)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)

27 SAF

Synonyms: hidden

= Stachys subsessilis Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 548 (1834)

= Stachys tenella Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 358 (1910)

species :

Stachys hydrophila Boiss.; Diagn. Pl. Orient. 12: 81 (1853) et Fl. 4: 726 (1879)

Distribution: +Algeria; Lebanon (C-Lebanon, coastal W-Lebanon)

20 ALG 34 LEB

species :

Stachys hyssopifolia Michx.; Fl. Bor.-Amer. [Michaux] 2: 4 (1803)

Name: Hyssopleaf hedgenettle (EN), Ysopblättriger Ziest (DE)

Distribution: Germany [I]; USA (Connecticut, Delaware, Florida, Georgia, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Missouri, North Carolina, New Hampshire, New Jersey, New York, Pennsylvania, Rhode Island, South Carolina, Virginia)

11 ger 74 USA 75 USA 78 USA

Synonyms: hidden

= Stachys atlantica Britton; Man. Fl. N. States [Britton] 792 (1901)

= Stachys hyssopifolia f. oswaldiae Oswald; Phytologia 11: 69 (1964)

= Stachys hyssopifolia var. lythroides (Small) J.B.Nelson; Phytologia 46: 94 (1980)

= Stachys lythroides Small; Fl. S.E. U.S. [Small] 1031 (1903)

species :

Stachys hyssopoides Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 558 (1833)

Distribution: South Africa (Limpopo, North-West Prov., Gauteng, Mpulamanga, Free State, KwaZulu-Natal, N-Cape Prov., E-Cape Prov.); Lesotho

27 LES SAF

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys coerulea Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 558 (1834)

= Stachys macilenta E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 240 (1838)

species :

Stachys iberica M.Bieb.; Fl. Taur.-Caucas. 2: 51 (1808)

subspecies :

subsp. iberica

Distribution: Crimea; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, S-Anatolia, SE-Anatolia, SSW-Anatolia); Iraq (NE-Iraq); Iran (Iranian Aserbaijan); Syria (W-Syrian Mountains)

14 KRY 33 ARM GEO 34 IRN IRQ SYR TUR

Synonyms: hidden

= Stachys congesta d´Urv.; Mém. Soc. Linn. Paris 1: 324 (1822)

= Stachys iberica var. congesta (d´Urv.) K.Koch; Linnaea 21: 693 (1849)

= Stachys caucasica K.Koch; Linnaea 21: 693 (1849)

variant :

var. densipilosa R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 284 (1974)

Distribution: Turkey

34 TUR

variant :

var. georgica (Rech.f.) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 100(6): 585 (2015)

Distribution: Georgia [Caucasus]; Turkey; Iraq; Iran

33 GEO 34 IRN IRQ TUR

Synonyms: hidden

= Stachys iberica subsp. georgica Rech.f.; Feddes Repert. Spec. Nov. Regni Veg. 53: 84 (1944)

subspecies :

subsp. stenostachya (Boiss.) Rech.f.; Feddes Repert. Spec. Nov. Regni Veg. 53: 84 (1944)

Distribution: Crimea; Turkey

14 KRY 34 TUR

Synonyms: hidden

= Stachys euphratica Hayek; Repert. Spec. Nov. Regni Veg. 21: 258 (1925)

= Stachys memorabilis Klokov; Fl. URSR 9: 648 (1960)

= Stachys stenostachya Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 41 (1859)

species :

Stachys ilicifolius (Schrenk) Sennikov; Checkl. Vasc. Pl. Tian-Shan 351 (2021)

Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; Iran (EC-Iran, NE-Iran: Mts., W-Iran); Afghanistan (Badghys, Balkh, Herat, Jawzjan / Sar-e-Pol, Samangan); China (Xinjiang)

32 KAZ TKM TZK UZB 34 AFG IRN 36 CHI

Remarks: clade 12 Otisia

Synonyms: hidden

= Chamaesphacos longiflorus Bornm. & Sint.; Mitt. Thüring. Bot. Vereins, N. F. 18: 51 (1905)

= Chamaesphacos ilicifolius Schrenk ex Fisch. & C.A.Mey.; Enum. Pl. Nov. [Fischer & Meyer] 1: 28 (1841)

species :

Stachys iltisii J.B.Nelson; J. Bot. Res. Inst. Texas 2(2): 761 (2008)

Distribution: USA (Arkansas, Oklahoma)

74 USA 78 USA

species :

Stachys inanis Hausskn. & Bornm.; Repert. Spec. Nov. Regni Veg. 22: 188 (1925)

Distribution: Turkey (E-Anatolia, Inner Anatolia)

34 TUR

species :

Stachys inflata Benth.; Labiat. Gen. Spec. [Bentham] 562 (1834)

subspecies :

subsp. inflata

Distribution: Armenia; Azerbaijan; Turkey (NE-Anatolia); Iraq; Iran (EC-Iran, Iranian Aserbaijan, W-Iran, N-Iran)

33 ARM AZE 34 IRN IRQ TUR

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys inflata subsp. albopannosa Rech.f.; Oesterr. Bot. Z. 99: 47 (1952)

subspecies :

subsp. caucasica (Stschegl.) Takht.; Fl. Erevana 260 (1972)

Distribution: Transcaucasus; Iran

34 IRN

Synonyms: hidden

= Stachys inflata var. caucasica Stschegl.; Bull. Soc. Imp. Naturalistes Moscou 24(2): 474 (1851)

= Stachys schtschegleevii Sosn. ex Grossh.; Fl. Kavkaza [Grossheim] 3: 314 (1932), nom. nov.

species :

Stachys intermedia Aiton; Hort. Kew. [W.Aiton] 2: 301 (1789)

Distribution: Northern Caucasus; Transcaucasus

33 NCS

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Stachys germanica var. intermedia (Aiton) Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)

= Stachys sibirica Link; Enum. Hort. Berol. Alt. 2: 109 (1822)

species :

Stachys ionica Halácsy; Consp. Fl. Graec., Suppl. 85 (1908)

Distribution: W-Greece (Ionian Isl.)

13 GRC

Remarks: clade 11 Swainsoniana

species :

Stachys iraqensis R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 294 (1974)

Distribution: Iraq (NW-Iraq)

34 IRQ

species :

Stachys istanbulensis Ö.Güner; Turk. J. Bot. 46(6): 625 (2022)

Distribution: Turkey

34 TUR

species :

Stachys iva Griseb.; Spic. Fl. Rumel. 2: 143 (1844)

Distribution: North Macedonia; NC-Greece

13 GRC NMA

Synonyms: hidden

= Stachys horvaticii Micevski; Acta Bot. Croat. 28: 452 (1969)

= Stachys iva var. ochroleuca Bornm.; Bot. Jahrb. Syst. 61: 89 (1928)

variant :

var. macedonica (Micevski) Krestovsk.; Novosti Sist. Vyssh. Rast. 48: 121 (2017)

Distribution: North Macedonia

13 NMA

Synonyms: hidden

= Stachys horvaticii var. macedonica Micevski; Acta Bot. Croat. 28: 453 (1969)

species :

Stachys jaimehintonii B.L.Turner; Phytologia 76(5): 395 (1994)

Distribution: Mexico (Michoacán)

79 MEX

species :

Stachys jijigaensis Sebsebe; Kew Bull. 48(2): 337 (1993)

Distribution: Ethiopia

24 ETH

species :

Stachys keerlii Benth.; Labiat. Gen. Spec. [Bentham] 551 (1833)

Distribution: Mexico (Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Tamaulipas, Veracruz)

79 MEX

Synonyms: hidden

= Stachys excelsa M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 194 (1844)

= Stachys inclusa Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 12 (1934)

species :

Stachys kermanshabensis Rech.f.; Pl. SySt. Evol. 134: 290 (1980)

Distribution: Iran (SW-Iran)

34 IRN

Remarks: clade 12 Otisia

species :

Stachys ketenoglui Kaynak, Daskin & Yilmaz; Nordic J. Bot. 27(3): 238 (2009)

Distribution: Turkey

34 TUR

species :

Stachys komarovii Knorring; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 15: 343 (1953)

Distribution: Russian Far East (Primorye)

31 RFE

species :

Stachys kotschyi Boiss. & Hohen.; Diagn. Pl. Orient. 5: 32 (1844)

Distribution: Iraq (NE-Iraq)

34 IRQ

Synonyms: hidden

= Stachys haussknechtii Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 281 (1929)

species :

Stachys kouyangensis (Vaniot) Dunn; Notes Roy. Bot. Gard. Edinburgh 8: 167 (1913)

Distribution: China (Guizhou, Hubei, Sichuan, Yunnan); SE-Tibet; Myanmar; Thailand; Vietnam

36 CHI TIB 41 MYA THA VIE

Synonyms: hidden

= Lamium kouyangense Vaniot; Bull. Acad. Int. Géogr. Bot. 14: 175 (1904)

= Scutellaria kouyangensis (Vaniot) Dunn; Notes Roy. Bot. Gard. Edinburgh 8: 167 (1913)

= Stachys franchetiana H.Lév.; Repert. Spec. Nov. Regni Veg. 9: 246 (1911)

= Stachys kouyangensis var. franchetiana (H.Lév.) C.Y.Wu; Acta Phytotax. Sin. 10: 228 (1965)

= Stachys kouyangensis var. leptodon (Dunn) C.Y.Wu; Acta Phytotax. Sin. 10: 227 (1965)

= Stachys kouyangensis var. tuberculata (Hand.-Mazz.) C.Y.Wu; Acta Phytotax. Sin. 10: 227 (1965)

= Stachys kouyangensis var. villosissima C.Y.Wu; Acta Phytotax. Sin. 10: 228 (1965)

= Stachys leptodon Dunn; Notes Roy. Bot. Gard. Edinburgh 8: 167 (1913)

= Stachys sieboldii var. tuberculata Hand.-Mazz.; Acta Horti Gothob. 13: 348 (1939)

species :

Stachys kulalensis Sebsebe; Kew Bull. 48(1): 115 (1993)

Distribution: Kenya

25 KEN

species :

Stachys kuntzei Gürke; Revis. Gen. Pl. 3(2): 262 (1898)

Distribution: South Africa (Mpulamanga, Free State, KwaZulu-Natal); Lesotho

27 LES SAF

Synonyms: hidden

= Stachys petrogenes Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1085 (1903)

species :

Stachys kurdica Boiss. & Hohen.; Diagn. Pl. Orient. 5: 31 (1844)

subspecies :

subsp. kurdica

Distribution: Turkey (E-Anatolia, SE-Anatolia); Iraq (NE-Iraq); Iran (Iranian Aserbaijan, W-Iran)

34 IRN IRQ TUR

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys ballotiformis Vatke; Bot. Zeitung (Berlin) 33: 448 (1875)

= Stachys benthamiana var. clinopodioides Boiss.; Fl. Orient. [Boissier] 4(2): 735 (1879)

= Stachys plebeia Vatke; Bot. Zeitung (Berlin) 33: 449 (1875)

= Stachys subnuda var. kurdica (Boiss. & Hohen.) Boiss.; Fl. Orient. [Boissier] 4(2): 734 (1879)

variant :

var. brevidens Bornm. ex R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 283 (1974)

Distribution: Turkey; Iraq

34 IRQ TUR

subspecies :

subsp. asterocalyx (Rech.f.) Salmaki; Bot. J. Linn. Soc. 170(4): 597 (2012)

Distribution: Iran (SW-Iran)

34 IRN

Synonyms: hidden

= Stachys asterocalyx Rech.f.; Bot. Jahrb. Syst. 71: 533 (1941)

species :

Stachys lallaniana R.Kr.Singh & Sanjeet Kumar; J. Biodivers. Conservation 7(3)-41: 42 (2023), nom. nov.

Distribution: South Africa

27 SAF

Remarks: provisionally accepted name

Synonyms: hidden

= Stachys malacophylla Skan; Bull. Misc. Inform. Kew 1909(10): 421 (1909), nom. illeg.

species :

Stachys lamarckii Benth.; Labiat. Gen. Spec. [Bentham] 562 (1833)

Distribution: Namibia; South Africa (N-Cape Prov.)

27 NAM SAF

Remarks: clade 12 Otisia

Synonyms: hidden

= Sideritis decumbens Thunb.; Prodr. Pl. Cap. 2: 95 (1800)

= Stachys nutans Benth.; Labiat. Gen. Spec. [Bentham] 561 (1834)

= Stachys rugosa Lam.; Tabl. Encycl. 3: 66 (1823), nom. illeg.

species :

Stachys lamioides Benth.; Pl. Hartw. [Bentham] 244 (1846)

Distribution: Colombia (Antioquia, Caldas, Cauca, Huila, Nariño, Putumayo, Quindío, Risaralda, Tolima, Valle); Ecuador

82 CLM 83 ECU

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys lamioides var. aristulata Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 50 (1934)

species :

Stachys langmaniae Rzed. & Calderón; Acta Bot. Mex. 3: 2 (1988)

Distribution: Mexico (Nuevo León, Tamaulipas)

79 MEX

species :

Stachys lanigera (Bornm.) Rech.f.; Bot. Jahrb. Syst. 71: 528 (1941)

Distribution: Iraq (NE-Iraq); Iran (W-Iran)

34 IRN IRQ

Synonyms: hidden

= Stachys fragillima Bornm. var. lanigera Bornm.; Beih. Bot. Centralbl. 33(2): 189 (1915)

species :

Stachys latidens Small; Man. Fl. N. States [Britton] 793 (1901)

Name: Broadtooth hedgenettle

Distribution: USA (Alabama, District of Columbia, Georgia, Kansas, Maryland, North Carolina, South Carolina, Tennessee, Virginia, West Virginia)

74 USA 75 USA 78 USA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys tenuifolia var. latidens (Small) J.B.Nelson; Phytologia 46: 94 (1980)

species :

Stachys lavandulifolia Vahl; Symb. Bot. [Vahl] 1: 42 (1790)

Distribution: Turkmenistan; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia, SE-Anatolia, SSW-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, W-Iran)

32 TKM 33 ARM GEO 34 IRN IRQ TUR

Remarks: clade 12 Otisia

Synonyms: hidden

= Sideritis calycantha M.Bieb.; Fl. Taur.-Caucas. 3: 393 (1819)

= Stachys boissieri Kapeller; Zametki Syst. Geogr. Rast. 16: 17 (1951)

= Stachys calycantha M.Bieb.; Fl. Taur.-Caucas. 3: 393 (1819)

= Stachys lavandulifolia var. glabrescens R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 283 (1974)

= Stachys tomentosa Bunge; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 21(1): 71 (1873), nom. illeg.

= Stachys zuvandica Rzazade; Fl. Azerbaidzhana 7: 611 (1957)

= Zietenia lavandulifolia (Vahl) Link; Enum. Hort. Berol. Alt. 2: 110 (1822)

= Zietenia orientalis Gled.; Mém. Acad. Roy. Sci. Hist. (Berlin) 1766: 3 (1766)

species :

Stachys laxa Boiss. & Buhse; Nouv. Mém. Soc. Imp. Naturalistes Moscou 12: 179 (1860)

Distribution: Iran (EC-Iran, N-Iran)

34 IRN

Synonyms: hidden

= Stachys demavendica Bornm.; Russk. Bot. Zhurn. 1: 8 (1911)

species :

Stachys leucoglossa Griseb.; Spic. Fl. Rumel. 2: 140 (1844)

subspecies :

subsp. leucoglossa

Distribution: ?Serbia; Kosovo; North Macedonia; Bulgaria; NE-Greece (incl. Mt. Athos)

13 BUL GRC KOS NMA

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys recta var. leucoglossa (Griseb.) Boiss.; Fl. Orient. [Boissier] 4(2): 730 (1879)

= Stachys recta subsp. leucoglossa (Griseb.) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)

subspecies :

subsp. samothracica (Degen) Biel & Kit Tan; Phytol. Balcan. 19(2): 271 (2013)

Distribution: NE-Greece (Samothraki)

13 GRC

Synonyms: hidden

= Stachys patula var. samothracica Degen; Oesterr. Bot. Z. 41: 336 (1891)

= Stachys leucoglossa var. samothracica (Degen) Rech.Fil.; Repert. Spec. Nov. Regni Veg., Beih. 100: 131 (1938)

species :

Stachys libanotica Benth.; Prodr. [A.P. de Candolle] 12: 462 (1848)

Distribution: Turkey (S-Anatolia); Lebanon (Antilebanon, C-Lebanon); Syria (Jbel Druze)

34 LEB SYR TUR

Remarks: libanotica agg.

Synonyms: hidden

= Stachys ciliaris Boiss.; Diagn. Pl. Orient. 12: 78 (1853)

species :

Stachys lindenii Benth.; Prodr. [A.P. de Candolle] 12: 467 (1848)

Distribution: Mexico (Chiapas, Guerrero, Jalisco, Mexico State, Michoacán, Oaxaca, Puebla, Queretaro, Veracruz); Guatemala

79 MEX 80 GUA

Synonyms: hidden

= Stachys boraginoides Benth.; Labiat. Gen. Spec. [Bentham] 540 (1834), nom. illeg.

= Stachys lindenii var. aristulata Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 49 (1934)

species :

Stachys linearis Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 559 (1833)

Distribution: Namibia; South Africa (Free State, N-Cape Prov., W-Cape Prov., E-Cape Prov.)

27 NAM SAF

Synonyms: hidden

= Stachys recurva Gürke; Bull. Herb. Boissier 6: 549 (1898)

= Stachys rosmarinifolia Benth.; Labiat. Gen. Spec. [Bentham] 559 (1833)

= Stachys rosmarinifolia var. burkei Benth.; Prodr. [A.P. de Candolle] 12: 494 (1848)

= Stachys rugosa var. linearis (Burch. ex Benth.) Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 359 (1910)

species :

Stachys longespicata Boiss. & Kotschy; Fl. Orient. [Boissier] 4(2): 725 (1879)

Name: Longspike hedgenettle

Distribution: Turkey (Inner Anatolia, S-Anatolia, SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); ?Iraq (?NE-Iraq); Lebanon (Antilebanon, C-Lebanon); Israel (Rift Valley, N-Israel); USA [I] (Maryland [I])

34 ISR LEB TUR 78 usa

species :

Stachys longiflora Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 39 (1859)

Distribution: Turkey (SSW-Anatolia)

34 TUR

species :

Stachys luchuensis (Kudô) comb.ined.

Distribution: Ryukyu Isl.; Taiwan

38 NNS TAI

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Suzukia luchuensis Kudô; J. Soc. Trop. Agric. 3: 226 (1931)

= Glechoma luchuensis (Kudô) Masam.; Hokuriku J. Bot. 5: 8 (1956)

species :

Stachys lurestanica Jamzad; Fl. Iran [ed. Assadi et al.] 76: 230 (2012)

Distribution: Iran

34 IRN

species :

Stachys lyallii Benth.; Labiat. Gen. Spec. [Bentham] 548 (1834)

Distribution: Madagascar

29 MDG

Synonyms: hidden

= Stachys hildebrandtii Briq.; Bull. Herb. Boissier 2: 138 (1894), nom. illeg.

species :

Stachys macraei Benth.; Linnaea 6: 78 (1831)

Distribution: Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Rios, Los Lagos, Aisén)

85 CLE

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Lamium hirtum Poir.; Encycl. [J.Lamarck et al.], Suppl. 1: 296 (1810)

= Stachys chonotica Hook.f.; Bot. Antarct. Voy. I. (Fl. Antarct.) 336 (1846)

= Stachys quadridentata Bertero ex Benth.; Prodr. [A.P. de Candolle] 12: 473 (1848)

species :

Stachys macrotricha Rech.f. & Goulimy; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 94: 24 (1957)

Distribution: NC-Greece (around Grevena)

13 GRC

Remarks: very rare

Synonyms: hidden

= Stachys heraclea var. macrotricha (Rech.f. & Goulimy) Krestovsk.; Novosti Sist. Vyssh. Rast. 42: 214 (2011)

species :

Stachys manantlanensis B.L.Turner; Phytologia 78(3): 209 (1995)

Distribution: Mexico (Colima, Jalisco)

79 MEX

species :

Stachys mandoniana Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 118 (1898)

Distribution: Bolivia (La Paz)

83 BOL

species :

Stachys marashica Ilçim, Çenet & Dadandý; Ann. Bot. Fenn. 45(2): 151 (2008)

Distribution: Turkey

34 TUR

species :

Stachys mardinensis (Post) R.R.Mill; Notes Roy. Bot. Gard. Edinburgh 38: 57 (1980)

Distribution: Turkey (E-Anatolia, Inner Anatolia, SE-Anatolia); Iraq (NE-Iraq)

34 IRQ TUR

Synonyms: hidden

= Nepeta mardinensis Post; Bull. Herb. Boissier 7: 159 (1899)

species :

Stachys maritima Gouan; Fl. Monsp. [Gouan] 91 (1764)

Distribution: NE-Spain; France; Corsica; Italy; Croatia; ?Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; European Turkey; ?Morocco; Algeria; Tunisia; Georgia [Caucasus]; Turkey (N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia)

12 COR FRA SPA 13 ALB BUL CRO ITA KOS NMA SRB TUE 20 ALG TUN 33 GEO 34 TUR

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys annua subsp. maritima Bonnier & Layens; Tabl. Syn. Pl. Vasc. France 253 (1894)

= Stachys maritima var. pubescens K.Koch; Linnaea 17: 299 (1844)

species :

Stachys marrubiifolia Viv.; Fl. Cors. Prodr., App. 1: 2 (1825)

Distribution: Corsica; W-Italy; Algeria; Tunisia

12 COR 13 ITA 20 ALG TUN

Synonyms: hidden

= Stachys marrubiifoliohirta Batt.; Fl. Algérie Tunisie 268 (1905)

= Stachys poiretii Ten.; Syll. Pl. Fl. Neapol. 538 (1831)

species :

Stachys matthewsii G.P.Fleming, J.B.Nelson & J.F.Towns.; J. Bot. Res. Inst. Texas 5(1): 9 (2011)

Distribution: USA (North Carolina)

78 USA

species :

Stachys megalodonta Hausskn. & Bornm. ex P.H.Davis; Notes Roy. Bot. Gard. Edinburgh 21: 46 (1951)

subspecies :

subsp. megalodonta

Distribution: Turkey (SE-Anatolia); Iraq (NE-Iraq); Iran (Iranian Aserbaijan, W-Iran)

34 IRN IRQ TUR

Remarks: clade 12 Otisia

subspecies :

subsp. mardinensis R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 282 (1974)

Distribution: Turkey

34 TUR

species :

Stachys melampyroides Hand.-Mazz.; Ann. K.K. Naturhist. Hofmus. 27: 415, t. 16 (1913)

Distribution: Turkey (SE-Anatolia); Iraq (NE-Iraq)

34 IRQ TUR

species :

Stachys menthifolia Vis.; Flora 12 (Ergänz. 1): 14 (1829)

Distribution: Croatia; ?Montenegro; Serbia; Kosovo; North Macedonia; Albania; NW-Greece (Mt. Grammos, Mt. Timphi to C-Epirus)

13 ALB CRO GRC KOS NMA SRB

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys grandiflora Host; Fl. Austriac. [Host] 2: 169 (1831)

species :

Stachys menthoides Kotschy & Boiss.; Fl. Orient. [Boissier] 4(2): 727 (1879)

Distribution: Turkey (E-Anatolia, SE-Anatolia)

34 TUR

Remarks: clade 4 Setifolia

species :

Stachys mexicana Benth.; Labiat. Gen. Spec. [Bentham] 541 (1833)

Name: Mexican hedgenettle

Distribution: Alaska; Canada (British Columbia); USA (California, Oregon, Washington State)

70 ASK 71 CDN 73 USA 76 USA

Synonyms: hidden

= Stachys ciliata Douglas ex Benth.; Labiat. Gen. Spec. [Bentham] 539 (1834)

= Stachys ciliata subsp. macrantha Piper; Proc. Biol. Soc. Washington 32: 43 (1919)

= Stachys ciliata var. pubens A.Gray; Syn. Fl. N. Amer. 2: 388 (1878)

= Stachys emersonii Piper; Erythea 6: 31 (1898)

= Stachys pubens (A.Gray) A.Heller; Bull. Torrey Bot. Club 25: 581 (1898)

species :

Stachys mialhesii de Noé; Bull. Soc. Bot. France 2: 584 (1855)

Distribution: Algeria

20 ALG

species :

Stachys micheliana Briq. ex Micheli; Mém. Soc. Phys. Genève 32(10): 7 (1897)

Distribution: Colombia (Antioquia, Arauca, Boyacá, Caldas, Cauca, Chocó, Córdoba, Cundinamarca, Huila, Magdalena, Meta, Nariño, Putumayo, Quindío, Risaralda, Santander, Tolima, Valle); SE-Brazil (São Paulo, Rio de Janeiro); S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul)

82 CLM 84 BRA

Synonyms: hidden

= Stachys mayorii Briq.; Mem. Soc. Neuchâteloise Sci. Nat. 5: 402 (1913)

species :

Stachys milanii Petrovic; Scrin. Fl. Select. 6: 117 (1887)

Distribution: Serbia; Kosovo; S- & E-Bulgaria

13 BUL KOS SRB

Synonyms: hidden

= Stachys spinulosa subsp. milanii (Petrov ex Magnier) Nyman; Consp. Fl. Eur. Suppl. 2: 252 (1890)

species :

Stachys milasensis Ö.Güner; Phytotaxa 566(1): 66 (2022)

Distribution: Turkey

34 TUR

species :

Stachys minor (Boiss.) Akçiçek & Dirmenci; Turk. J. Bot. 36(3): 225 (2012)

Distribution: Turkey

34 TUR

Synonyms: hidden

= Stachys libanotica var. minor Boiss.; Fl. Orient. [Boissier] 4(2): 718 (1879)

species :

Stachys mohinora B.L.Turner; Phytologia 77(4): 360 (1994 publ. 1995)

Distribution: Mexico (Chihuahua)

79 MEX

species :

Stachys mollissima Willd.; Hort. Berol. [Willdenow] 60, t. 60 (1805)

Distribution: S-Albania; NW-Greece (Kerkyra, Paxos, Andipaxos, W-Epirus)

13 ALB GRC

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys decumbens Pers.; Syn. Pl. [Persoon] 2: 123 (1806)

species :

Stachys moorei B.L.Turner; Phytologia 77(4): 360 (1994 publ. 1995)

Distribution: Mexico (Hidalgo, Veracruz)

79 MEX

species :

Stachys mouretii Batt. & Pit.; Contr. Fl. Maroc (Tours) 32 (1918)

Distribution: Morocco

20 MOR

species :

Stachys mucronata Sieber ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 2: 733 (1825)

Distribution: E-Crete; East Aegaean Isl. (Kasos, Karpathos)

13 KRI 34 EAI

Remarks: clade 1 Eriostomum, germanica agg.

species :

Stachys multicaulis Benth.; Prodr. [A.P. de Candolle] 12: 486 (1848)

Distribution: Iraq (NW-Iraq); Iran (W-Iran)

34 IRN IRQ

species :

Stachys munzurdagensis R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 290 (1974)

Distribution: Turkey (E-Anatolia)

34 TUR

species :

Stachys namazdaghensis Yild.; Ot Sist. Bot. Derg. 17(2): 85 (2010)

Distribution: Turkey

34 TUR

species :

Stachys natalensis Hochst.; Flora 28: 65 (1845)

Distribution: Zimbabwe; Mozambique; South Africa (Limpopo, Gauteng, Mpulamanga, KwaZulu-Natal); Eswatini [Swaziland]

26 MOZ ZIM 27 SAF SWZ

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys leptoclada Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1084 (1903)

= Stachys transvaalensis Gürke; Bot. Jahrb. Syst. 28: 316 (1900)

variant :

var. galpinii (Briq.) Codd; Bothalia 12(2): 185 (1977)

Distribution: South Africa (Limpopo, Gauteng, Mpulamanga, KwaZulu-Natal); Eswatini [Swaziland]

27 SAF SWZ

Synonyms: hidden

= Stachys galpinii Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1082 (1903)

= Stachys lupulina Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1082 (1903)

= Stachys parilis N.E.Br.; Bull. Misc. Inform. Kew 1901: 131 (1901)

= Stachys villosissima H.M.L.Forbes; Bothalia 4: 38 (1941)

species :

Stachys nelsonii B.R.Keener & L.J.Davenp.; J. Bot. Res. Inst. Texas 10(2): 315, figs, 1, 3-4 (2016)

Distribution: USA (Alabama)

78 USA

species :

Stachys nemorivaga Briq.; Bot. Jahrb. Syst. 19: 192 (1894)

Distribution: D.R.Congo [Zaire]; Angola; Zambia

23 ZAI 26 ANG ZAM

species :

Stachys nepetifolia Desf.; Tabl. École Bot., ed. 3 (Cat. Pl. Horti Paris.) 58 (1829)

Distribution: Mexico (Ciudad de Mexico, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Oaxaca, Queretaro, San Luis Potosi, Tamaulipas, Tlaxcala, Veracruz, Zacatecas)

79 MEX

Synonyms: hidden

= Nepeta circinnata Willd.; Enum. Pl. Hort. Berol. [Willdenow] 602 (1809)

species :

Stachys nephrophylla Rech.f.; Pl. Syst. Evol. 134: 291 (1980)

Distribution: Iraq (NE-Iraq)

34 IRQ

Remarks: clade 12 Otisia

species :

Stachys neurocalycina Boiss.; Diagn. Pl. Orient. 12: 85 (1853) et Fl. 4: 748 (1879)

Distribution: Lebanon (Antilebanon, C-Lebanon, coastal W-Lebanon); Israel (coastal W-Israel, Rift Valley, N-Israel); Jordania (W-Jordania)

34 ISR JOR LEB

Remarks: clade 12 Otisia

species :

Stachys nigricans Benth.; Comm. Pl. Afr. Austr. [Meyer] 238 (1838)

Distribution: South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Eswatini [Swaziland]

27 SAF SWZ

Remarks: clade 2 Stachys s. str.

species :

Stachys nivea Labill.; Icon. Pl. Syr., Dec. 3: 5, t. 3 (1809)

Distribution: Lebanon (Antilebanon); Syria (C-Syrian Desert); Sinai peninsula (S-Sinai)

34 LEB SIN SYR

species :

Stachys nubilorum Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 48 (1934)

Distribution: Guatemala

80 GUA

Synonyms: hidden

= Stachys glechomoides Epling; Bull. Torrey Bot. Club 71: 484 (1944)

species :

Stachys obliqua Waldst. & Kit.; Descr. Icon. Pl. Hung. 2: 142 (1803 publ. 1804)

Distribution: Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; European Turkey; NE-Greece (Nomos Epirou, Rodopi, Thasos); East Aegaean Isl. (Lesvos); Turkey (Inner Anatolia, NW-Anatolia: Bithynia, W-Anatolia, WN-Anatolia)

13 ALB BUL CRO GRC KOS MNT NMA SRB TUE 34 EAI TUR

Synonyms: hidden

= Stachys heraclea var. lutea Benth.; Prodr. [A.P. de Candolle] 12: 463 (1848)

= Stachys montbretii Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 48 (1836)

= Stachys pauciflora Vis.; Ill. Piant. Nuov. 1: 10 (1840), nom. illeg.

species :

Stachys oblongifolia Wall. ex Benth.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830)

Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan); North Korea; Taiwan; NE-India (Assam, Manipur, Meghalaya, Mizoram, Nagaland, Tripura, West Bengal); Bangladesh; Vietnam; W-Java

36 CHI 38 NKO TAI 40 BAN IND 41 VIE 42 JAW

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys imaii Nakai; Bot. Mag. (Tokyo) 26: 169 (1912)

= Stachys martini Vaniot; Bull. Acad. Int. Géogr. Bot. 14: 187 (1904)

= Stachys modica Hance; J. Bot. 20: 292 (1882)

= Stachys palustris var. imaii (Nakai) Nakai; Bot. Mag. (Tokyo) 34: 48 (1920)

= Stachys subargentea Hayata; Icon. Pl. Formosan. 8: 94 (1919)

= Stachys tashiroi Hayata; Icon. Pl. Formosan. 8: 95 (1919)

variant :

var. leptopoda (Hayata) C.Y.Wu; Acta Phytotax. Sin. 10: 222 (1965)

Distribution: China (Fujian, Guangdong, Guangxi, Sichuan, Yunnan); Taiwan

36 CHI 38 TAI

Synonyms: hidden

= Stachys leptopoda Hayata; Icon. Pl. Formosan. 8: 93 (1919)

= Stachys oblongifolia f. leptopoda (Hayata) Kudô; Mem. Fac. Sci. Taihoku Imp. Univ. 2: 188 (1929)

species :

Stachys obscura Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 43 (1859)

Distribution: Turkey (S-Anatolia, SSW-Anatolia); Lebanon (C-Lebanon, coastal W-Lebanon); Syria (coastal W-Syria, W-Syrian Mountains)

34 LEB SYR TUR

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys arabica var. minor Boiss.; Fl. Orient. [Boissier] 4(2): 747 (1879)

species :

Stachys obtusicrena Boiss.; Diagn. Pl. Orient. 7: 57 (1846)

Distribution: Iran (EC-Iran, SW-Iran)

34 IRN

species :

Stachys obtusifolia MacOwan; Bull. Misc. Inform. Kew 1893: 13 (1893)

Distribution: South Africa (E-Cape Prov.)

27 SAF

Synonyms: hidden

= Stachys obtusifolia var. flanaganii Skan

species :

Stachys ochroleuca Phil.; Linnaea 33: 192 (1864)

Distribution: Chile (Coquimbo, Valparaíso, O´Higgins, Maule, Ñuble, Biobío, Araucanía)

85 CLE

Synonyms: hidden

= Stachys dubia Briq.; Bull. Herb. Boissier 4: 873 (1896)

= Stachys litoralis Phil.; Anales Univ. Chile 90: 560 (1895)

= Stachys meyenii Phil.; Anales Univ. Chile 1873: 520 (1873), nom. illeg.

= Stachys rupestris Phil.; Anales Univ. Chile 90: 562 (1895), nom. illeg.

species :

Stachys ocymastrum (L.) Briq.; Lab. Alp. Mar. 252 (1893)

Name: Basilikum-Ziest (DE); Italian hedgenettle (EN); Alfabega (ES)

Distribution: Portugal; Spain; Gibraltar; Baleares; France; Corsica; Sardinia; Italy; Sicily; Greece (very rare Kerkyra, SW-Peloponnisos); NW-Crete; Morocco; Algeria; Tunisia; Libya; Madeira (Madeira Isl., Porto Santo Isl.); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma); Lebanon (coastal W-Lebanon); USA [I] (New Jersey [I])

12 BAL COR FRA GIB POR SAR SPA 13 GRC ITA SIC 20 ALG LBY MOR TUN 21 CNY MDR 34 LEB 75 usa

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Galeopsis hirsuta L.; Sp. Pl. [Linnaeus] 2: 580 (1753)

= Galeopsis hispanica Mill.; Gard. Dict., ed. 8: no. 6 (1768)

= Sideritis ocymastrum L.; Syst. Nat., ed. 10, 2: 1098 (1759)

= Stachys divaricata Viv.; Fl. Cors. Prodr., App. 3 (1825)

= Stachys foucauldiana Sennen; Diagn. Nouv. 199 (1936)

= Stachys hirta L.; Sp. Pl. [Linnaeus], ed. 2: 813 (1763), nom. illeg.

= Stachys hirta var. hirtula (Pomel) Batt.; Fl. Algérie [Battandier et al.] Dicot. 706 (1890)

= Stachys hirta var. lagascae (Caball.) Sennen & Mauricio; Cat. Fl. Rif Orient. 93 (1933)

= Stachys hirta var. virgata Batt.; Fl. Algérie [Battandier et al.] Dicot. 706 (1888)

= Stachys hirtula Pomel; Nouv. Mat. Fl. Atl. 305 (1874)

= Stachys inscripta Rchb.; Iconogr. Bot. Exot. 1: 28 (1824)

= Stachys lagascae Caball.; Mem. Real Soc. Esp. Hist. Nat. 8: 280 (1915)

= Stachys marrubiastrum Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 266 (1896)

= Stachys ocymastrum f. bicolor Faure & Maire; Bull. Soc. Hist. Nat. Afrique N. 24: 227 (1933)

= Stachys ocymastrum f. lagascae (Caball.) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 35 (1979)

= Stachys ocymastrum subsp. bicolor (Faure & Maire) C.E.Blanco; Actes Simposi Int. Bot. Font Quer 2: 338 (1992)

= Stachys ocymastrum var. bicolor (Faure & Maire) Maire; Cat. Pl. Maroc 3: 639 (1934)

= Stachys ocymastrum var. rifana (Font-Quer & Maire) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 35 (1979)

= Stachys ocymastrum var. violascens Faure & Maire; Bull. Soc. Hist. Nat. Afrique N. 20: 197 (1929)

= Stachys rifana Font Quer & Maire; Cavanillesia 3: 54 (1930)

= Stachys rifana var. foucauldiana (Sennen) Font Quer & Sennen; Diagn. Nouv. 240 (1936)

= Tetrahitum hirtum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 104 (1809)

species :

Stachys oligantha Baker; J. Linn. Soc., Bot. 20: 233 (1884)

Distribution: Madagascar

29 MDG

species :

Stachys oreophila Hedge; Fl. Madagasc., Fam. 175: 31 (1998)

Distribution: Madagascar

29 MDG

species :

Stachys pacifica B.L.Turner; Phytologia 76(5): 396 (1994)

Distribution: Mexico (Chihuahua, Colima, Jalisco, Sinaloa, Sonora)

79 MEX

species :

Stachys palaestina L.; Sp. Pl. [Linnaeus], ed. 2: 1674 (1763)

Distribution: Lebanon (Antilebanon); Israel (coastal W-Israel, N-Israel)

34 ISR LEB

Synonyms: hidden

= Stachys lanata Moench; Methodus [Moench] 397 (1794)

species :

Stachys palustris L.; Sp. Pl. [Linnaeus] 2: 580 (1753)

Name: Sumpf-Ziest (DE), Marsh Woundwort (EN)

Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Italy; Malta; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (scattered N- & C-Greece, Kerkyra); Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Ukraine; Crimea; Siberia (Altai, Krasnoyarsk, Tuva, W-Siberia); Russian Far East [I] (Amur [I], Khabarovsk [I], Primorye [I], Sakhalin [I]); Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Turkey (N-Anatolia, NE-Anatolia); Iran (Iranian Aserbaijan); China (Xinjiang); Mongolia; NW-India (Jammu & Kashmir); Australia [I] (Tasmania [I]); New Zealand [I]; Alaska [I]; Canada [I] (Manitoba [I], New Brunswick [I], Newfoundland [I], Nova Scotia [I], Ontario [I], Prince Edward Isl. [I], Québec [I]); USA [I] (Connecticut [I], District of Columbia [I], Delaware [I], Illinois [I], Massachusetts [I], Maryland [I], Maine [I], Michigan [I], Minnesota [I], New Hampshire [I], New Jersey [I], New York [I], Ohio [I], Pennsylvania [I], Rhode Island [I], Vermont [I], Wisconsin [I])

10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 CIS COR FRA POR SPA 13 ALB BIH BUL CRO GRC ITA KOS MLT MNT NMA ROM SLV SMA SRB TUE 14 BLR EST KRY LAT LIT RUS UKR 30 SIB 31 rfe 32 KAZ KGZ TZK UZB 33 ARM GEO NCS 34 IRN TUR 36 CHI 37 MON 40 IND 50 aus 51 nzl 70 ask 71 cdn 72 cdn 74 usa 75 usa 78 usa

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys aquatica Bubani; Fl. Pyren. [Bubani] 1: 444 (1897)

= Stachys austriaca Heynh.; Alph. Aufz. Gew. 697 (1847), nom. inval.

= Stachys maeotica Postrig.; Bot. Zhurn. (Kiev) 7(2): 67 (1950)

= Stachys palustris f. acuminata (Briq.) Ravarut; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 220 (1961)

= Stachys palustris f. angustifolia Serg.; Fl. Sibir. Occid., ed. 2: 2364 (1937)

= Stachys palustris f. canescens Briq. ex Litard.; Candollea 5: 159 (1932)

= Stachys palustris f. cleoniquei B.Boivin; Naturaliste Canad. 94: 648 (1967)

= Stachys palustris f. eglandulosa Ravarut; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 688 (1961)

= Stachys palustris f. latifolia Serg.; Fl. Sibir. Occid., ed. 2: 2364 (1937), nom. illeg.

= Stachys palustris f. petiolata (Peterm.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2417 (1927)

= Stachys palustris f. segetum (Hagen) C.W.Christ.; Fl. Kiel [J. A. Christiansen et al.] 199 (1922)

= Stachys palustris f. sessilis (Celak.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2417 (1927)

= Stachys palustris f. submersa Glück; Biol. Morphol. Untersuch. Wasser- Sumpfgewächse, Teil 3: 62 (1911)

= Stachys palustris f. virescens Briq. ex Litard.; Candollea 5: 159 (1932)

= Stachys palustris subvar. acuminata (Briq.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2418 (1927)

= Stachys palustris subvar. major (Wirtg.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2418 (1927)

= Stachys palustris subvar. oblongifolia (Briq.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2417 (1927)

= Stachys palustris var. acuminata Briq.; Lab. Alp. Mar. 1: 248 (1891)

= Stachys palustris var. agrestis (F.Aresch.) Serg.; Fl. Sibir. Occid., ed. 2: 2365 (1937)

= Stachys palustris var. angustifolia Benth.; Labiat. Gen. Spec. [Bentham] 542 (1834)

= Stachys palustris var. arenicola Farw.; Amer. Midl. Naturalist 11: 82 (1928)

= Stachys palustris var. bracteata Beck; Fl. Nieder-Österreich 2(2): 1013 (1893)

= Stachys palustris var. ciliata Peterm.; Fl. Lips. Excurs. 447 (1838)

= Stachys palustris var. cinerea T.Durand; Bull. Soc. Roy. Bot. Belgique 20(2): 70 (1881)

= Stachys palustris var. hybrida Benth.; Labiat. Gen. Spec. [Bentham] 542 (1834)

= Stachys palustris var. insularis Briq. ex Litard.; Candollea 5: 159 (1932)

= Stachys palustris var. latifolia Peterm.; Fl. Lips. Excurs. 446 (1838)

= Stachys palustris var. major Wirtg.; Fl. Preuss. Rheinprov. 368 (1857)

= Stachys palustris var. oblongifolia Briq.; Lab. Alp. Mar. 1: 248 (1891)

= Stachys palustris var. petiolata Celak.; Arch. Naturwiss. Landesdurchf. Böhmen 2(2) (Prodr. Fl. Böhmen 2): 358 (1871), nom. illeg.

= Stachys palustris var. petiolata Peterm.; Anal. Pfl.-Schlüss. 349 (1846)

= Stachys palustris var. petiolata T.Durand; Bull. Soc. Roy. Bot. Belgique 18(2): 101 (1879)

= Stachys palustris var. pseudoambigua Erdner; Ber. Naturwiss. Vereins Schwaben Neuburg 39-40: 571 (1911)

= Stachys palustris var. pubescens Schur; Enum. Pl. Transsilv. 539 (1866)

= Stachys palustris var. segetalis Brügger; Naturgesch. Beitr. Umg. Chur [Fl. Curiensis] 75 (1874), nom. inval.

= Stachys palustris var. segetalis Klett & Richt.; Fl. Leipzig 510 (1830)

= Stachys palustris var. segetum (Hagen) Nyman; Consp. Fl. Eur. 578 (1881)

= Stachys palustris var. segetum (Hagen) Rchb.; Fl. Germ. Excurs. 319 (1832)

= Stachys palustris var. sessilis Celak.; Arch. Naturwiss. Landesdurchf. Böhmen 2(2) (Prodr. Fl. Böhmen 2): 358 (1871)

= Stachys palustris var. subcanescens Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849)

= Stachys palustris var. umbrosa Celak.; Arch. Naturwiss. Landesdurchf. Böhmen 2(2) (Prodr. Fl. Böhmen 2): 358 (1871)

= Stachys palustris var. viridifolia Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849)

= Stachys palustris var. vulgaris Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849), nom. illeg.

= Stachys palustris var. vulgaris Peterm.; Anal. Pfl.-Schlüss. 349 (1846)

= Stachys segetum Hagen; Chlor. Boruss. [Hagen] 218 (1819)

= Stachys wolgensis Wilensky; Sborn. Statej Mater. Gosplana A.S.S.R. Nemts. Povolzh. 1: 38 (1926)

species :

Stachys paneiana Mouterde; Saussurea 4: 25 (1973)

Distribution: Israel (N-Israel)

34 ISR

species :

Stachys pannosa Phil.; Anales Univ. Chile 90: 564 (1895)

Distribution: Chile (Antofagasta, Atacama, Coquimbo, Valparaíso, Ñuble, Biobío)

85 CLE

species :

Stachys parolinii Vis.; Mem. Reale Ist. Veneto Sci. 1: 46 (1842 publ. 1843)

Distribution: W-Greece (Lefkada, Kefalonia, N-Peloponnisos)

13 GRC

species :

Stachys parviflora Benth.; Prodr. [A.P. de Candolle] 12: 490 (1848)

Distribution: Tajikistan; Iran (NE-Iran: Mts.); Afghanistan (Badakshan, Badghys, Baghlan, Balkh, Bamyan, Farah, Faryab, Ghazni, Ghorat, Herat, Kabul, Kandahar, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan, Samangan, Takhar); Pakistan (Baluchistan, Quetta, Waziristan, Kurram, N. W. Frontier Prov., Peshawar, Swat, Pakistani Punjab); NW-India (Himachal Pradesh, Jammu & Kashmir, Uttarakhand)

32 TZK 34 AFG IRN 40 IND PAK

Remarks: clade 12 Otisia

Synonyms: hidden

= Phlomidoschema parviflorum (Benth.) Vved.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 9 1: 54 (1941)

species :

Stachys parvifolia M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 193 (1844)

Distribution: Mexico (Chiapas, Chihuahua, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Mexico State, Michoacán, Queretaro, San Luis Potosi, Sonora, Tamaulipas, Tlaxcala, Veracruz)

79 MEX

species :

Stachys paulii Grossh.; Trudy Geobot. Obsl. Pastb. S.S.R. Azerbaijdzhana, Ser. B, 1: 77 (1929)

Distribution: Azerbaijan

33 AZE

Synonyms: hidden

= Stachys zappanioides Woronow; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 1: 221 (1933)

species :

Stachys penanevada B.L.Turner; Phytologia 77(4): 362 (1994 publ. 1995)

Distribution: Mexico (Tamaulipas, Nuevo León)

79 MEX

species :

Stachys peruviana Dombey ex Benth.; Labiat. Gen. Spec. [Bentham] 551 (1833)

Distribution: Peru; Bolivia (Chuquisaca, Cochabamba)

83 BOL PER

species :

Stachys petrokosmos Rech.f.; Svensk Bot. Tidskr. 43: 40 (1949)

Distribution: Turkey (S-Anatolia)

34 TUR

species :

Stachys philippiana Vatke; Bot. Zeitung (Berlin) 33: 443 (1875)

Distribution: Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, Maule, Ñuble, Biobío, Araucanía)

85 CLE

Synonyms: hidden

= Stachys andina Phil.; Anales Univ. Chile 90: 562 (1895), nom. illeg.

= Stachys elliptica Phil.; Linnaea 33: 193 (1864), nom. illeg.

species :

Stachys pilifera Benth.; Prodr. [A.P. de Candolle] 12: 487 (1848)

subspecies :

subsp. pilifera

Distribution: Iran (SW-Iran)

34 IRN

subspecies :

subsp. ixodes (Boiss. & Hausskn. ex Boiss.) Salmaki; Bot. J. Linn. Soc. 170(4): 606 (2012)

Distribution: Iran (SW-Iran)

34 IRN

Synonyms: hidden

= Stachys ixodes Boiss. & Hausskn. ex Boiss.; Fl. Orient. [Boissier] 4(2): 738 (1879)

species :

Stachys pilosa Nutt.; J. Acad. Nat. Sci. Philadelphia 7: 48 (1834)

Name: Hairy hedgenettle

Distribution: Alaska; Canada (Alberta, British Columbia, Manitoba, New Brunswick, Northern Territories, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); USA (Arizona, California, Colorado, Connecticut, Iowa, Idaho, Illinois, Kansas, Massachusetts, Maine, Minnesota, Missouri, Montana, North Dakota, Nebraska, New Hampshire, New Jersey, New Mexico, Nevada, New York, Oklahoma, Oregon, Pennsylvania, South Dakota, Utah, Vermont, Washington State, Wisconsin, Wyoming)

70 ASK CDN 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys ampla Rydb.; Bull. Torrey Bot. Club 36: 683 (1909)

= Stachys asperrima Rydb.; Bull. Torrey Bot. Club 36: 682 (1909)

= Stachys borealis Rydb.; Brittonia 1: 95 (1931)

= Stachys bracteata Greene; Pittonia 3: 342 (1898)

= Stachys homotricha (Fernald) Rydb.; Brittonia 1: 95 (1931)

= Stachys leibergii Rydb.; Bull. Torrey Bot. Club 36: 682 (1909)

= Stachys palustris f. stevensonis B.Boivin; Phytologia 22: 380 (1972)

= Stachys palustris subsp. homotricha (Fernald) Hultén; Kungl. Svenska Vetenskapsakad. Handl., Ser. 4, 13: 152 (1971)

= Stachys palustris subsp. pilosa (Nutt.) Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 63 (1934)

= Stachys palustris var. homotricha Fernald; Rhodora 10: 85 (1908)

= Stachys palustris var. nipigonensis Jenn.; J. Wash. Acad. Sci. 10: 459 (1920)

= Stachys palustris var. phaneropoda Weath. ex Fernald; Rhodora 45: 475 (1943)

= Stachys palustris var. pilosa (Nutt.) Fernald; Rhodora 45: 474 (1943)

= Stachys palustris var. puberula Jenn.; J. Wash. Acad. Sci. 10: (1920)

= Stachys pilosa var. homotricha (Fernald) Mohlenbr.; Phytoneuron 2015-67: 2 (2015)

= Stachys puberula (Jenn.) Rydb.; Brittonia 1: 96 (1931)

= Stachys pustulosa Rydb.; Brittonia 1: 95 (1931)

= Stachys schweinitzii Rydb.; Brittonia 1: 95 (1931)

= Stachys scopulorum Greene; Pittonia 3: 342 (1898)

= Stachys teucriformis Rydb.; Bull. Torrey Bot. Club 31: 640 (1904)

= Stachys velutina Schwein.; Narrat. Exped. St. Peter´s River [Keating] 2: 390 (1824), nom. illeg.

variant :

var. arenicola (Britton) G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116: 46 (1989 publ. 1990)

Distribution: Canada (Manitoba, Ontario, Québec); USA (Arkansas, Connecticut, Iowa, Illinois, Indiana, Kansas, Kentucky, Massachusetts, Maine, Michigan, Minnesota, Missouri, Nebraska, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, South Carolina, South Dakota, Virginia, Vermont, Wisconsin, West Virginia)

71 CDN 72 CDN 74 USA 75 USA 78 USA

Synonyms: hidden

= Stachys arenicola Britton; Man. Fl. N. States [Britton] 792 (1901)

= Stachys brevidens Rydb.; Brittonia 1: 95 (1931), nom. illeg.

species :

Stachys pilosissima M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 194 (1844)

Distribution: Mexico (Chiapas, Guerrero, Jalisco, Mexico State, Michoacán, Oaxaca, Puebla, Tamaulipas, Veracruz)

79 MEX

Synonyms: hidden

= Stachys exilis Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 25 (1934)

= Stachys flaccida Fernald; Proc. Amer. Acad. Arts 35: 563 (1900)

= Stachys vulcanica Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 33 (1934)

species :

Stachys pinardii Boiss.; Diagn. Pl. Orient. 5: 29 (1844)

Distribution: Turkey (SSW-Anatolia)

34 TUR

species :

Stachys pinetorum Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 36 (1859)

Distribution: Turkey (S-Anatolia, SSW-Anatolia); Syria (W-Syrian Mountains)

34 SYR TUR

species :

Stachys pittieri Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 119 (1898)

Distribution: Costa Rica

80 COS

species :

Stachys plumosa Griseb.; Spic. Fl. Rumel. 2: 139 (1844)

subspecies :

subsp. plumosa

Distribution: ?Serbia; Kosovo; North Macedonia; Albania; W-Bulgaria; Greece (NC-Greece to Thessalia, NE-Greece to Thasos)

13 ALB BUL GRC KOS NMA

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys pangaea Phitos; Oesterr. Bot. Z. 113: 272 (1966)

= Stachys recta subsp. plumosa (Griseb.) Nyman; Consp. Fl. Eur. 579 (1881)

= Stachys viridis Boiss. & Heldr.; Diagn. Pl. Orient., Sér. 2, 4: 39 (1859)

subspecies :

subsp. freynii (Hausskn.) Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 290 (1929)

Distribution: North Macedonia; NC-Greece

13 GRC NMA

Synonyms: hidden

= Stachys freynii Hausskn.; Mitt. Geogr. Ges. (Thüringen) Jena 5: 70 (1887)

= Stachys dolopica Formánek

species :

Stachys pringlei Greenm.; Proc. Amer. Acad. Arts 41: 245 (1905)

Distribution: Mexico (Hidalgo, Jalisco, Mexico State, Michoacán, Queretaro, San Luis Potosi, Veracruz)

79 MEX

species :

Stachys pseudohumifusa Sebsebe; Kew Bull. 48(1): 117 (1993)

subspecies :

subsp. pseudohumifusa

Distribution: Cameroon; Bioko Isl. [Fernando Poo]; Tanzania

23 BIO CMN 25 TAN

Conservation Status: IUCN EN (Cameroon)

subspecies :

subsp. minutiflora Y.B.Harv.; Kew Bull. 51(3): 451 (1996)

Distribution: Zambia

26 ZAM

subspecies :

subsp. saxeri Y.B.Harv.; Kew Bull. 51(3): 451 (1996)

Distribution: Nigeria; Cameroon

22 NGA 23 CMN

species :

Stachys pseudonigricans Gürke; Bot. Jahrb. Syst. 28: 315 (1900)

Distribution: Tanzania; Angola; Zambia; Malawi; Zimbabwe

25 TAN 26 ANG MLW ZAM ZIM

species :

Stachys pseudophlomis C.Y.Wu; Acta Phytotax. Sin. 10: 226 (1965)

Distribution: China (Hubei, Sichuan)

36 CHI

species :

Stachys pseudopinardii R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 281 (1974)

Distribution: Turkey (SSW-Anatolia)

34 TUR

species :

Stachys pumila Banks & Sol.; Nat. Hist. Aleppo, ed. 2 [Russell]: 2: 225, t. 10 (1794)

Distribution: Turkey (S-Anatolia, SE-Anatolia); Syria (NW-Syria, W-Syrian Mountains)

34 SYR TUR

Synonyms: hidden

= Stachys bornmuelleri Hand.-Mazz.; Ann. K.K. Naturhist. Hofmus. 27: 414 (1913)

= Stachys floribunda Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 51 (1836)

species :

Stachys pusilla (Wedd.) Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 120 (1898)

Distribution: Colombia (Boyacá, Cundinamarca, Tolima); Ecuador; Peru; Bolivia (La Paz)

82 CLM 83 BOL ECU PER

Synonyms: hidden

= Stachys elliptica var. pusilla Wedd.; Chlor. Andina 2: 153 (1860)

species :

Stachys pycnantha Benth.; Pl. Hartw. [Bentham] 331 (1849)

Name: Shortspike hedgenettle

Distribution: USA (California)

76 USA

species :

Stachys pyramidalis J.K.Morton; J. Linn. Soc., Bot. 58: 273 (1962)

Distribution: Nigeria

22 NGA

species :

Stachys radicans Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 24 (1934), in Repert. Spec. Nov. Regni Veg., Beih. 85: 9 (1935)

Distribution: Mexico (Chiapas, Ciudad de Mexico, Hidalgo, Jalisco, Mexico State, Michoacán, Tlaxcala, Veracruz); Colombia (Boyacá, Cundinamarca, Nariño)

79 MEX 82 CLM

species :

Stachys ramosissima Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 50 (1836)

Distribution: Turkey (E-Anatolia, Inner Anatolia)

34 TUR

Synonyms: hidden

= Stachys ramosissima var. elazigensis R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 291 (1974)

species :

Stachys recta L.; Mant. Pl. 82 (1767)

subspecies :

subsp. recta

Name: Aufrechter Ziest (DE), Stiff hedgenettle (EN)

Distribution: Belgium; Luxembourg; Germany (Brandenburg, +Berlin, Baden-Württemberg, Bayern, Hessen, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein [I], Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; NE-Spain; Andorra; France; Italy; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; ?Greece; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; Ukraine; Northern Caucasus; USA [I] (District of Columbia [I])

11 AUT BGM CZE GER HUN LIE LUX POL SLO SWI 12 ADO FRA SPA 13 ALB BIH BUL CRO ITA KOS MNT NMA ROM SLV SMA SRB 14 BLR LAT LIT RUS UKR 33 NCS 78 usa

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Betonica adscendens Walther; Fl. Giessen 453 (1802), nom. illeg.

= Betonica decumbens Moench; Methodus [Moench] 396 (1794), nom. illeg.

= Betonica hirta L.; Cent. Pl. II: 20 (1756)

= Betonica recta (L.) Baill.; Iconogr. Fl. Franç. t. 208

= Ortostachys recta (L.) Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 136 (1869)

= Prasium stachys E.H.L.Krause; Deutschl. Fl. [Sturm], ed. 2, Abbild., 11: 116 (1903), nom. illeg.

= Sideritis maculata Gilib.; Fl. Lit. Inch. 1: 81 (1782), opus utique oppr.

= Stachys albanica Markgr.; Ber. Deutsch. Bot. Ges. 44: 428 (1926)

= Stachys betonica Scop.; Fl. Carniol., ed. 2, 1: 411 (1771)

= Stachys bufonia Thuill.; Fl. Env. Paris, éd. 2(1): 295 (1799)

= Stachys czernjaevii Des.-Shost.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 8: 152 (1940)

= Stachys decumbens Rchb.; Fl. Germ. Excurs. 318 (1831), nom. illeg.

= Stachys delphinensis Jord.; Arch. Fl. France Allem. [F.W.Schultz] 322 (1847), et in Cat. Jard. Bot. Grenoble 1853: 12 (1853)

= Stachys erecta K.Koch; Linnaea 21: 691 (1849)

= Stachys glabrata Simonk.; Enum. Fl. Transsilv. 450 (1886)

= Stachys hirta (L.) Thell.; Repert. Spec. Nov. Regni Veg. 10: 290 (1912), nom. illeg.

= Stachys krynkensis Kotov; Bot. Zhurn. (Kiev) 1(2): 297 (1940)

= Stachys nitens Janka; Linnaea 30: 597 (1859 publ. 1860)

= Stachys patula Griseb.; Spic. Fl. Rumel. 2: 142 (1844)

= Stachys procumbens Lam.; Fl. Franç. [Lamarck] 2: 385 (1779)

= Stachys recta f. angusta Borbás; Természettud. Közl. 1884: 158 (1884)

= Stachys recta f. angustifolia (Ten.) Bolzon; Bull. Soc. Bot. Ital. 1903: 37 (1903)

= Stachys recta subsp. delphinensis (Jord.) Nyman; Consp. Fl. Eur. 579 (1881)

= Stachys recta subsp. hirta (Ten.) Briq.; Lab. Alp. Mar. 263 (1893)

= Stachys recta subsp. sarajevensis (K.Maly) Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 181 (1924)

= Stachys recta subvar. alpina (Godr.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2411 (1927)

= Stachys recta subvar. glandulifera (A.Kern.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2411 (1927)

= Stachys recta subvar. midzorica Adamovic; Oesterr. Bot. Z. 42: 408 (1899)

= Stachys recta var. alpina Godr.; Fl. France [Grenier] 2: 693 (1853)

= Stachys recta var. angusta (Borbás) Diklic; Fl. Srbije 6: 427 (1974)

= Stachys recta var. angustifolia Godr.; Fl. France [Grenier] 2: 692 (1853), nom. illeg.

= Stachys recta var. angustifolia Ten.; Syll. Pl. Fl. Neapol. 292 (1831)

= Stachys recta var. angustifolia Wender.; Fl. Hassiaca [Wenderoth] 197 (1846)

= Stachys recta var. glabrescens Pancic; Verh. Zool.-Bot. Vereins Wien 6: 543 (1856)

= Stachys recta var. hirta Ten.; Syll. Pl. Fl. Neapol. 292 (1831)

= Stachys recta var. lisae Briq.; Lab. Alp. Mar. 262 (1893)

= Stachys recta var. longiflora Pamp.; Boll. Mus. Republ. San Marino 4: 117 (1920)

= Stachys recta var. luxemburgensis Lefort; Arch. Sect. Sci. Inst. Grand-Ducal Luxemb., n. s. 19: 443 (1950)

= Stachys recta var. major Ten.; Syll. Pl. Fl. Neapol. 292 (1831)

= Stachys recta var. midzorica (Adamovic) Diklic; Fl. Srbije 6: 428 (1974)

= Stachys recta var. nitens (Janka) Chrtek; Folia Geobot. Phytotax. 27: 182 (1992)

= Stachys recta var. ramosissima Griseb.; Arch. Naturgesch. (Berlin) 18: 342 (1852)

= Stachys recta var. ramosissima Pancic; Verh. Zool.-Bot. Vereins Wien 6: 543 (1856), nom. illeg.

= Stachys recta var. stenophylla (Spreng.) Nyman; Consp. Fl. Eur. 579 (1881)

= Stachys sarajevensis K.Malý; Glasn. Zemaljsk. Muz. Bosni Hercegovini 40: 160 (1920)

= Stachys sideritis Vill.; Hist. Pl. Dauphiné [Villars] 2: 375 (1787)

= Stachys spinulosa var. glabrata (Simonk.) Nyman; Consp. Fl. Eur. Suppl. 2: 252 (1890)

= Stachys stenophylla Spreng.; Novi Provent. [Sprengel] 40 (1818)

= Stachys sylvestris Forssk.; Fl. Aegypt.-Arab. xxviii (1775)

= Stachys transsilvanica Schur; Enum. Pl. Transsilv. 539 (1866)

= Syderitis maculata Gilib.; Fl. Lit. Inch. 1: 81 (1782), opus utique oppr.

subspecies :

subsp. baldaccii (Maly) Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 183 (1924)

Distribution: Croatia; ?Montenegro; ?North Macedonia; Albania; Greece

13 ALB CRO GRC

Synonyms: hidden

= Stachys baldaccii (K.Malý) Hand.-Mazz. & Janch.; Oesterr. Bot. Z. 56: 105 (1906)

= Stachys subcrenata f. baldaccii K.Malý; Verh. K.K. Zool.-Bot. Ges. Wien 54: 245 (1904)

subspecies :

subsp. doerfleri (Hayek) Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 167 (1924)

Distribution: Albania

13 ALB

Synonyms: hidden

= Stachys doerfleri Hayek; Oesterr. Bot. Z. 70: 18 (1921)

subspecies :

subsp. labiosa (Bertol.) Briq.; Lab. Alp. Mar. 259 (1893)

Distribution: Germany [I]; Austria; France; Italy; +San Marino; Slovenia; Croatia

11 AUT ger 12 FRA 13 CRO ITA SLV SMA

Synonyms: hidden

= Stachys labiosa Bertol.; Fl. Ital. [Bertoloni] 6: 166 (1844)

= Stachys recta subsp. grandiflora (Caruel) Arcang.; Comp. Fl. Ital. [Arcangeli] 551 (1882)

= Stachys recta var. grandiflora Caruel; Prodr. Fl. Tosc. 519 (1860)

= Stachys recta var. labiosa (Bertol.) Briq.; Lab. Alp. Mar. 259 (1893)

= Stachys subcrenata subsp. labiosa (Bertol.) Nyman; Consp. Fl. Eur. 579 (1881)

= Stachys subcrenata var. labiosa (Bertol.) Vis.; Fl. Dalmat. 2: 208 (1847)

subspecies :

subsp. olympica Stoj. & Jordanov; God. Sofiisk. Univ. Fiz.-Mat. Fak. 34: 205 (1937-1938 publ. 1938)

Distribution: Greece (Mt. Olimbos)

13 GRC

subspecies :

subsp. psammophila (Fiori) Arrigoni; Fl. Analítica Toscana 7: 163 (2020)

Distribution: Italy

13 ITA

Synonyms: hidden

= Stachys recta f. psammophila Fiori; Sched. Fl. Ital. Exs., Ser. 3, 14: 217, no. 2537 (1921)

subspecies :

subsp. rhodopaea (Velen.) Chrtek; Folia Geobot. Phytotax. 27(2): 182 (1992)

Distribution: Bulgaria

13 BUL

Synonyms: hidden

= Stachys fragilis var. rhodopaea Velen.; Fl. Bulg. [Velenovsky] 456 (1891)

subspecies :

subsp. subcrenata (Vis.) Briq.; Lab. Alp. Mar. 257 (1893)

Distribution: Germany [I]; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; European Turkey; Greece (mountains of N-Greece, also C-Greece, Peloponnisos, Thasos); Turkey (NW-Anatolia: Bithynia, SW-Anatolia)

11 ger 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA SLV SRB TUE 34 TUR

Synonyms: hidden

= Stachys fragilis f. serpentini Fiori; Nuovo Giorn. Bot. Ital., n. s. 17(4): 640 (1910)

= Stachys fragilis Vis.; Flora 12(1. Ergänzungsbl.): 14 (1829)

= Stachys glandulifera Dalla Torre & Sarnth.; Fl. Tirol [Dalla Torre & Sarnthein] 6(3): 187 (1912)

= Stachys karstiana (Borbás) Hand.-Mazz.; Oesterr. Bot. Z. 56: 100 (1906)

= Stachys montenegrina K.Malý; Glasn. Zemaljsk. Muz. Bosni Hercegovini 22: (1910)

= Stachys omblae H.Lindb.; Öfvers. Finska Vetensk.-Soc. Förh. 48(13): 93 (1906)

= Stachys petrogena Hand.-Mazz. & Janch.; Oesterr. Bot. Z. 56: 101 (1906)

= Stachys ramosissima Rochel; Bot. Reise Banat 82 (1838), nom. illeg.

= Stachys recta f. eriostachyus (A.Kern.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2412 (1927)

= Stachys recta subsp. ramosissima Nyman; Consp. Fl. Eur. 579 (1881)

= Stachys recta subsp. serpentini (Fiori) Arrigoni; Inform. Bot. Ital. 12: 138 (1980 publ. 1981)

= Stachys recta var. fragilis (Vis.) Boiss.; Fl. Orient. [Boissier] 4(2): 730 (1879)

= Stachys recta var. hyssopifolia Ten.; Syll. Pl. Fl. Neapol. 292 (1831)

= Stachys recta var. karstiana (Borbás) Chrtek; Folia Geobot. Phytotax. 27: 182 (1992), isonym

= Stachys recta var. karstiana (Borbás) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2411 (1927)

= Stachys recta var. subcrenata (Vis.) Nyman; Consp. Fl. Eur. 579 (1881)

= Stachys recta var. visianii Briq.; Lab. Alp. Mar. 259 (1893)

= Stachys subcrenata subsp. fragilis (Vis.) Poldini; Stud. Geobot. 21: 8 (2001)

= Stachys subcrenata var. angustifolia Vis.; Fl. Dalmat. 2: 208 (1847)

= Stachys subcrenata var. fragilis (Vis.) Vis.; Fl. Dalmat. 2: 208 (1847)

= Stachys subcrenata var. karstiana Borbás; Math. Term. Közlem. 14: 402 (1877)

= Stachys subcrenata Vis.; Flora 12(1. Ergänzungsbl.): 15 (1829)

subspecies :

subsp. tenoreana Bornm.; Magyar Bot. Lapok 33: 81 (1934)

Distribution: Italy

13 ITA

Synonyms: hidden

= Stachys tenoreana Bornm.; Magyar Bot. Lapok 33: 81 (1934)

species :

Stachys rehmannii Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 345 (1910)

Distribution: South Africa (Limpopo)

27 SAF

species :

Stachys reptans Hedge; Fl. Madagasc., Fam. 175: 26 (1998)

Distribution: Madagascar

29 MDG

Remarks: clade 2 Stachys s. str.

species :

Stachys reticulata Codd; Bothalia 12(2): 183 (1977)

Distribution: South Africa (Mpulamanga)

27 SAF

species :

Stachys richardiana R.Kr.Singh; Phytotaxa 620(2): 198 (2023)

Distribution: Ethiopia

24 ETH

Synonyms: hidden

= Stachys hypoleuca Hochst. ex A.Rich.; Tent. Fl. Abyss. 2: 197 (1850), nom. illeg.

= Stachys palaestina var. hypoleuca (Hochst. ex A.Rich.) Benth.

species :

Stachys riederi Cham.; Linnaea 6: 570 (1831)

Distribution: European Russia; Siberia (Buryatia, Chita, Irkutsk, Krasnoyarsk, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); China (Heilongjiang, Jilin, Liaoning, Nei Mongol, Shaanxi, Shandong, Shanxi); Mongolia; Japan (Hokkaido)

14 RUS 30 SIB 31 RFE 36 CHI 37 MON 38 JAP

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys aspera subsp. baicalensis (Fisch. ex Benth.) Krestovsk.; Turczaninowia 7(4): 20 (2004)

= Stachys aspera var. baicalensis (Fisch. ex Benth.) Maxim.; Bull. Soc. Imp. Naturalistes Moscou 54(1): 44 (1879)

= Stachys baicalensis Fisch. ex Benth.; Labiat. Gen. Spec. [Bentham] 543 (1834)

= Stachys baicalensis var. hispida (Ledeb.) Nakai; Bot. Mag. (Tokyo) 34: 46 (1920)

= Stachys palustris f. baicalensis (Fisch. ex Benth.) Regel; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 4(4): 119 (1861)

= Stachys palustris var. baicalensis (Fisch. ex Benth.) Turcz.; Bull. Soc. Imp. Naturalistes Moscou 7: 156 (1837)

= Stachys palustris var. hispida Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849)

= Stachys riederi var. hispida (Ledeb.) H.Hara; Bot. Mag. (Tokyo) 41: 144 (1937)

= Stachys rigens Oett.; Trudy Bot. Sada Imp. Yur´evsk Univ. 6: 216 (1906)

variant :

var. hispidula (Regel) H.Hara; Bot. Mag. (Tokyo) 51: 144 (1937)

Distribution: Russian Far East (Kuril Isl.); China (Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol); Mongolia; South Korea; Japan (Hokkaido, Honshu, Shikoku, Kyushu)

31 RFE 36 CHI 37 MON 38 JAP KOR

Synonyms: hidden

= Stachys aspera var. hispidula (Regel) Krestovsk.; Sosud. Rast. Sovet. Dal´nego Vostoka 7: 344 (1995), isonym

= Stachys aspera var. hispidula (Regel) Vorosch.; Florist. Issl. Razn. Raionakh S.S.S.R. [A.K.Skvortsov] 189 (1985)

= Stachys baicalensis f. intermedia Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43: 32 (1921)

= Stachys baicalensis var. angustifolia Honda; Bot. Mag. (Tokyo) 46: 374 (1932)

= Stachys baicalensis var. hispidula (Regel) Nakai; Bot. Mag. (Tokyo) 34: 46 (1920)

= Stachys japonica f. intermedia Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43(8): 32 (1921)

= Stachys palustris f. hispidula Regel; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 4(4): 119 (1861)

= Stachys riederi f. angustifolia (Honda) H.Hara; Bot. Mag. (Tokyo) 51: 144 (1937)

= Stachys riederi var. intermedia (Kudô) Kitam.; Acta Phytotax. Geobot. 17: 10 (1957)

variant :

var. japonica (Miq.) H.Hara; Bot. Mag. (Tokyo) 51: 144 (1937)

Distribution: Russian Far East (Amur); China (Anhui, Fujian, Hebei, Henan, Jiangsu, Jiangxi, Liaoning, Nei Mongol, Shandong, Zhejiang); North Korea; South Korea; Japan (Honshu, Kyushu); Ryukyu Isl.

31 RFE 36 CHI 38 JAP KOR NKO NNS

Synonyms: hidden

= Stachys aspera f. glabrata Nakai; Fl. Kor. 2: 147 (1911), nom. illeg.

= Stachys aspera subsp. japonica (Miq.) Krestovsk.; Turczaninowia 7(4): 21 (2004)

= Stachys aspera var. japonica (Miq.) Maxim.; Bull. Soc. Imp. Naturalistes Moscou 54(1): 45 (1879)

= Stachys aspera var. japonica (Miq.) Vorosch.; Florist. Issl. Razn. Raionakh S.S.S.R. [A.K.Skvortsov] 189 (1985)

= Stachys baicalensis var. japonica (Miq.) Kom.; Trudy Imp. S.-Peterburgsk. Bot. Sada 25: 371 (1907)

= Stachys japonica f. angustifolia Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 197 (1867)

= Stachys japonica f. glabrata Matsum. & Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43(8): 30 (1921)

= Stachys japonica f. villosa Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43: 32 (1921)

= Stachys japonica Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 111 (1865)

= Stachys japonica var. tomentosa F.Z.Li & Z.Y.Sun; Bull. Bot. Res., Harbin 25(2): 134 (2005)

= Stachys japonica var. villosa (Kudô) Ohwi; Fl. Jap. [Ohwi] 1008 (1953)

= Stachys riederi f. harana H.Ohba & S.Akiyama; Bull. Univ. Mus. Univ. Tokyo 41: 101 (2005)

= Stachys riederi var. villosa (Kudô) Kitam.; Acta Phytotax. Geobot. 17: 10 (1957)

species :

Stachys rigida Nutt. ex Benth.; Prodr. [A.P. de Candolle] 12: 472 (1848)

subspecies :

subsp. rigida

Name: Rough hedgenettle

Distribution: USA (California, Nevada, Oregon, Washington State); Mexico (Baja California Norte)

73 USA 76 USA 79 MEX

Synonyms: hidden

= Stachys ajugoides var. rigida (Nutt. ex Benth.) Jeps. & Hoover; Fl. Calif. [Jepson] 3: 426 (1943)

= Stachys littoralis Greene; Pittonia 3: 341 (1898), nom. illeg.

= Stachys prattenii Durand; J. Acad. Nat. Sci. Philadelphia, Ser. 2, 3: 100 (1855)

= Stachys rigida subsp. rivularis Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 60 (1934)

= Stachys rivularis A.Heller; Muhlenbergia 1: 33 (1901), nom. illeg.

= Stachys striata Greene; Pittonia 3: 341 (1898)

= Stachys vestita Howell; Fl. N.W. Amer. [Howell] 558 (1901), non Benth.

subspecies :

subsp. quercetorum (A.Heller) Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 59 (1934)

Distribution: USA (California, Oregon)

73 USA 76 USA

Synonyms: hidden

= Stachys ajugoides var. quercetorum (A.Heller) A.Heller; Fl. Calif. [Jepson] 3(2): 426 (1943)

= Stachys gracilenta A.Heller; Muhlenbergia 2: 319 (1907)

= Stachys nuttallii var. leptostachya Benth.; Pl. Hartw. [Bentham] 331 (1849)

= Stachys quercetorum A.Heller; Muhlenbergia 2: 318 (1907)

= Stachys rigida var. quercetorum (A.Heller) G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116: 47 (1989 publ. 1990)

= Stachys viarum A.Heller; Muhlenbergia 2: 316 (1907)

species :

Stachys riparia A.Pool; Novon 17(1): 64 (2007)

Distribution: Costa Rica

80 COS

species :

Stachys rivularis J.M.Wood & M.S.Evans; J. Bot. 35: 489 (1897)

Distribution: South Africa (KwaZulu-Natal)

27 SAF

Synonyms: hidden

= Stachys schlechteri Gürke; Bot. Jahrb. Syst. 26: 74 (1898)

species :

Stachys rizeensis R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 278 (1974)

Distribution: Turkey (NE-Anatolia)

34 TUR

species :

Stachys rosea (Desf.) Boiss.; Fl. Orient. [Boissier] 4(2): 725 (1879)

Distribution: Algeria

20 ALG

Synonyms: hidden

= Sideritis aegyptiaca Pers.; Syn. Pl. [Persoon] 2: 117 (1806)

= Sideritis rosea Desf.; Ann. Mus. Hist. Nat. 10: 302 (1807)

species :

Stachys rothrockii A.Gray; Proc. Amer. Acad. Arts 12: 82 (1877)

Name: Rothrock´s hedgenettle

Distribution: USA (Arizona, New Mexico, Utah)

76 USA 77 USA

species :

Stachys rotundifolia Moc. & Sessé ex Benth.; Labiat. Gen. Spec. [Bentham] 548 (1833)

Distribution: Mexico (Chiapas, Colima, Hidalgo, Jalisco, Mexico State, Morelos, Oaxaca, Puebla, Queretaro, Veracruz)

79 MEX

species :

Stachys rubella Hedge; Fl. Madagasc., Fam. 175: 33 (1998)

Distribution: Madagascar

29 MDG

species :

Stachys rudatisii Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 347 (1910)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)

27 SAF

species :

Stachys rugosa Aiton; Hort. Kew. [W.Aiton] 2: 303 (1789)

Distribution: South Africa (N-Cape Prov., W-Cape Prov., E-Cape Prov.); Lesotho

27 LES SAF

Synonyms: hidden

= Sideritis pallida Thunb.; Prodr. Pl. Cap. 2: 95 (1800)

= Stachys crenulata Briq.; Bot. Jahrb. Syst. 19: 192 (1894)

= Stachys deserti Benth.; Prodr. [A.P. de Candolle] 12: 494 (1848)

= Stachys jugalis Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 562 (1834)

= Stachys multiflora Benth.; Prodr. [A.P. de Candolle] 12: 492 (1848)

= Stachys rugosa var. longiflora Benth.; Comm. Pl. Afr. Austr. [Meyer] 1(2): 241 (1838)

species :

Stachys rupestris Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 49 (1836)

Distribution: Turkey (S-Anatolia, SSW-Anatolia); Syria (NW-Syria, W-Syrian Mountains)

34 SYR TUR

Remarks: clade 12 Otisia

species :

Stachys salisii Jord. & Fourr.; Brev. Pl. Nov. 2: 92 (1868)

Distribution: Corsica; Sardinia; Italy

12 COR SAR 13 ITA

Synonyms: hidden

= Stachys aimerici Gamisans; Monde Pl. 100: 26 (2005)

= Stachys corsica proles salisii (Jord. & Fourr.) Rouy; Fl. France [Rouy & Foucaud] 11: 313 (1909)

= Stachys corsica var. micrantha Bertol.; Fl. Ital. [Bertoloni] 6(1): 154 (1845)

species :

Stachys sanchezii Rzed. & A.García; Phytologia 56(5): 321 (1984)

Distribution: Mexico (Ciudad de Mexico, Hidalgo, Jalisco, Mexico State, Michoacán, Puebla, Queretaro, Veracruz)

79 MEX

species :

Stachys sandersii B.L.Turner; Phytologia 77(4): 372 (1994 publ. 1995)

Distribution: Mexico (San Luis Potosi)

79 MEX

species :

Stachys saturejoides Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 51 (1836)

Distribution: Turkey (S-Anatolia, SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); Syria (Jazira)

34 SYR TUR

species :

Stachys saxicola Coss. & Balansa; Bull. Soc. Bot. France 20: 257 (1873)

subspecies :

subsp. saxicola

Distribution: Morocco; Algeria

20 ALG MOR

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys saxicola subsp. chelifensis Quézel & Simonn.; Bull. Soc. Hist. Nat. Afrique N. 50: 149 (1959)

subspecies :

subsp. laxa Faure & Maire; Bull. Soc. Hist. Nat. Afrique N. 22: 61 (1931)

Distribution: Morocco

20 MOR

subspecies :

subsp. maweana (Ball) Maire; Cat. Pl. Maroc 638 (1934)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Stachys maweana Ball; J. Linn. Soc., Bot. 16: 626 (1878)

= Stachys saxicola var. maweana (Ball) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 102(11): 1534 (2017)

subspecies :

subsp. villosissima (Ball) Maire; Cat. Pl. Maroc 1113 (1941)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Stachys saxicola var. villosissima Ball; J. Linn. Soc., Bot. 16: 626 (1878)

= Stachys saxicola subsp. platyodon Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 208 (1932)

species :

Stachys scaberula Vatke; Bot. Zeitung (Berlin) 33: 462 (1875)

Distribution: Bhutan; NE-India (Assam)

40 BHU IND

species :

Stachys scabrida Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 349 (1910)

Distribution: South Africa (W-Cape Prov., E-Cape Prov.)

27 SAF

Synonyms: hidden

= Stachys priorii Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 353 (1910)

species :

Stachys schimperi Vatke; Linnaea 37: 330 (1871 publ. 1873)

Distribution: Sudan; Eritrea; Saudi Arabia

24 ERI SUD 35 SAU

Synonyms: hidden

= Stachys palaestina var. schimperi (Vatke) Benth.; Fl. Trop. Afr. [Oliver et al.] 5: 467 (1900)

species :

Stachys semsurensis Firat; Phytotaxa 511(3): 276, figs. 1-3 (2021)

Distribution: Turkey

34 TUR

species :

Stachys serbica Pancic; Fl. Serbiae 564 (1874)

Distribution: Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (scattered NW-Greece)

13 ALB BIH BUL GRC KOS MNT NMA SRB

Synonyms: hidden

= Koevia serbica (Pancic) Krestovsk.; Novosti Sist. Vyssh. Rast. 45: 94 (2014)

species :

Stachys sericantha P.H.Davis; Kew Bull. [6]: 111 (1951)

Distribution: Turkey (SW-Anatolia)

34 TUR

Remarks: libanotica agg.

species :

Stachys sericea Cav.; Descr. Pl. [Cavanilles] 75 (1801)

Distribution: Chile (Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Lagos)

85 CLE

Synonyms: hidden

= Stachys candidissima Phil.; Linnaea 29: 19 (1858)

= Stachys glabrata Phil.; Anales Univ. Chile 90: 559 (1895), nom. illeg.

= Stachys longifolia Bertero ex Steud.; Nomencl. Bot. [Steudel], ed. 2, 2: 628 (1841), nom. inval.

= Stachys sideritidoides Gillies ex Benth.; Labiat. Gen. Spec. [Bentham] 740 (1835)

species :

Stachys sericophylla Halácsy; Consp. Fl. Graec. 2: 519 (1902)

Distribution: S-Albania

13 ALB

Remarks: clade 1 Eriostomum, germanica agg.

species :

Stachys sessilifolia E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 239 (1838)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)

27 SAF

Synonyms: hidden

= Stachys bachmannii Gürke; Bot. Jahrb. Syst. 26: 75 (1898)

species :

Stachys sessilis Gürke; Bot. Jahrb. Syst. 26: 74 (1898)

Distribution: South Africa (Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho

27 LES SAF

species :

Stachys setifera C.A.Mey.; Verz. Pfl. Casp. Meer [C.A. von Meyer] 94 (1831)

subspecies :

subsp. setifera

Distribution: Kazakhstan; Uzbekistan; Tajikistan; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, SE-Anatolia: Mesopotamian Anatolia, SSW-Anatolia, SW-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, E-Iran, N-Iran, Iranian Aserbaijan, W-Iran); Afghanistan (Badakshan, Faryab, Kabul); Lebanon (Antilebanon)

32 KAZ TZK UZB 33 ARM GEO 34 AFG IRN IRQ LEB TUR

Remarks: clade 4 Setifolia

Synonyms: hidden

= Stachys bornmuelleri Gand.; Bull. Soc. Bot. France 65: 68 (1918)

= Stachys daenensis Gand.; Bull. Soc. Bot. France 65: 68 (1918)

= Stachys iranica Rech.f.; Oesterr. Bot. Z. 99: 44 (1952)

= Stachys lycopsiformis K.Koch; Linnaea 21: 692 (1848)

= Stachys repens Schur; Enum. Pl. Transsilv. 538 (1866)

= Stachys setifera subsp. daenensis (Gand.) Rech.f.; Fl. Iranica [Rechinger] 150: 366 (1982)

= Stachys setifera subsp. iranica (Rech.f.) Rech.f.; Fl. Iranica [Rechinger] 150: 366 (1982)

= Stachys shirini Parsa; Kew Bull. [3]: 226 (1948)

subspecies :

subsp. lycia (Gand.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 281 (1974)

Distribution: Turkey

34 TUR

Synonyms: hidden

= Stachys lycia Gand.; Bull. Soc. Bot. France 65: 68, in clavi (1918)

species :

Stachys shikikunensis (Kudô) comb.ined.

Distribution: Taiwan

38 TAI

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Suzukia shikikunensis Kudô; J. Soc. Trop. Agric. 2: 146 (1930)

= Glechoma shikikunensis (Kudô) Masam.; Hokuriku J. Bot. 4: 88 (1955)

species :

Stachys siirtensis Ö.Güner & Akçiçek; Phytotaxa 516(3): 253 (2021)

Distribution: Turkey

34 TUR

species :

Stachys simplex Schltr.; J. Bot. 35: 221 (1897)

Distribution: South Africa (Mpulamanga, Free State, KwaZulu-Natal); Eswatini [Swaziland]

27 SAF SWZ

Synonyms: hidden

= Stachys chrysotrichos Gürke; Bot. Jahrb. Syst. 28: 316 (1900)

= Stachys pascuicola Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1086 (1903)

species :

Stachys sivasica Kit Tan & Yildiz; Notes Roy. Bot. Gard. Edinburgh 45(3): 449 (1988 publ. 1989)

Distribution: Turkey

34 TUR

species :

Stachys sparsipilosa R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 287 (1974)

Distribution: Turkey (Inner Anatolia, S-Anatolia, SSW-Anatolia)

34 TUR

Synonyms: hidden

= Stachys iberica var. pallidiflora Boiss.; Fl. Orient. [Boissier] 4(2): 731 (1879), nom. illeg.

species :

Stachys spathulata Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 559 (1833)

Distribution: Mozambique; Namibia; Botswana; South Africa (Limpopo, North-West Prov., Free State, KwaZulu-Natal, N-Cape Prov.); Madagascar

26 MOZ 27 BOT NAM SAF 29 MDG

Synonyms: hidden

= Stachys karasmontana Dinter; Repert. Spec. Nov. Regni Veg. 17: 203 (1921)

= Stachys minima Gürke; Bull. Herb. Boissier 6: 550 (1898)

= Stachys pachycalymna Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1087 (1903)

species :

Stachys spectabilis Choisy ex DC.; Mém. Soc. Phys. Genève 1: 457 (1823)

Distribution: Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, NE-Anatolia, S-Anatolia, SE-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Israel (Rift Valley, N-Israel)

33 ARM GEO 34 IRN IRQ ISR TUR

Remarks: clade 1 Eriostomum, germanica agg.

Synonyms: hidden

= Stachys elata K.Koch; Linnaea 21: 687 (1848)

= Stachys germanica subsp. spectabilis (Choisy ex DC.) Briq.; Lab. Alp. Mar. 234 (1893)

= Stachys germanica var. elata (K.Koch) Briq.; Lab. Alp. Mar. 235 (1893)

= Stachys germanica var. hypoleuca (K.Koch) Briq.; Lab. Alp. Mar. 235 (1893)

= Stachys germanica var. spectabilis (Choisy ex DC.) Briq.; Lab. Alp. Mar. 234 (1893)

= Stachys hypoleuca K.Koch; Linnaea 21: 688 (1849)

= Stachys rosea Hohen.; Bull. Soc. Imp. Naturalistes Moscou 11: 300 (1838), no diagnostic descr.

= Stachys spectabiliformis Kapeller; Zametki Sist. Geogr. Rast. 16: 14 (1951)

= Stachys spectabilis subsp. elata (K.Koch) Krestovsk.; Novosti Sist. Vyssh. Rast. 33: 203 (2001)

species :

Stachys speluncarum Contandr. & Quézel; Bull. Soc. Bot. France 123(7-8): 426 (1976)

Distribution: Turkey

34 TUR

species :

Stachys sphaerodonta Baker; J. Linn. Soc., Bot. 20: 233 (1884)

Distribution: Madagascar

29 MDG

species :

Stachys spinosa L.; Sp. Pl. [Linnaeus] 2: 581 (1753)

Distribution: Crete (widespread); East Aegaean Isl. (Karpathos)

13 KRI 34 EAI

Remarks: clade 11 Swainsoniana

species :

Stachys spinulosa Sm.; Fl. Graec. Prodr. [Sibthorp] 1: 410 (1809)

Name: Dorniger Ziest (DE); Spiny Woundwort (EN)

Distribution: Croatia; ?Montenegro; ?Serbia; Kosovo; North Macedonia; Albania; Greece (C- & S-Greece, Kerkyra, Ionian Isl., Kithira, Sporades, Skiros, Evvoia, Andros, Kiklades: Amorgos, Astipaleia); W-Crete; East Aegaean Isl. (Lesvos, Ikaria, Samos, Kalimnos, Kos, Simi, Rhodos, Kastellorhizo Isl.); Turkey (NW-Anatolia: Bithynia, SSW-Anatolia, SW-Anatolia, W-Anatolia)

13 ALB CRO GRC KOS KRI NMA 34 EAI TUR

Synonyms: hidden

= Galeopsis orientalis Mill.; Gard. Dict., ed. 8: no. 5 (1768)

= Stachys betonicifolia Desf.; Ann. Mus. Hist. Nat. 10: 304 (1807), nom. illeg.

= Stachys orientalls (Mill.) Vahl; Symb. Bot. [Vahl] 2: 64 (1791), nom. illeg.

species :

Stachys splendens Wall. ex Benth.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830)

Distribution: Tibet; Pakistan (Hazara, Murree, Astor); Nepal; Bhutan; N-India (Arunachal Pradesh, Assam, Himachal Pradesh, Jammu & Kashmir, Nagaland, Sikkim, Uttarakhand); Maldives [I]; Bangladesh

36 TIB 40 BAN BHU IND mdv NEP PAK

Synonyms: hidden

= Stachys bikramii Aswal & Mehrotra; J. Econ. Taxon. Bot. 4: 1028 (1983), nom. nov. superfl.

= Stachys emodi Hedge; Fl. Pakistan 192: 185 (1990) (nom. nov.)

= Stachys melissifolia Benth.; Labiat. Gen. Spec. [Bentham] 538 (1834)

= Stachys splendens Wall.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830), nom. inval.

species :

Stachys spreitzenhoferi Heldr.; Oesterr. Bot. Z. 30. 344 (1880)

subspecies :

subsp. spreitzenhoferi

Distribution: S-Greece (Malea peninsula, Kithira)

13 GRC

Remarks: clade 11 Swainsoniana, candida agg.

subspecies :

subsp. virella D.Perss.; Biosyst. Stachys 38 (1981)

Distribution: S-Greece (Malea peninsula)

13 GRC

Synonyms: hidden

= Stachys spreitzenhoferi var. virella (D.Perss.) Krestovsk.; Novosti Sist. Vyssh. Rast. 48: 120 (2017)

species :

Stachys sprucei Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 117 (1898)

Distribution: Ecuador

83 ECU

species :

Stachys spruneri Boiss.; Prodr. [A.P. de Candolle] 12: 488 (1848)

Distribution: C-Greece (Attiki)

13 GRC

Remarks: clade 11 Swainsoniana, swainsonii agg.

species :

Stachys stebbinsii G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116(1): 48 (1989 publ. 1990)

Name: Stebbins´ hedgenettle

Distribution: USA (California)

76 USA

species :

Stachys stricta Greene; Erythea 2: 122 (1894)

Name: Sonoma hedgenettle

Distribution: USA (California)

76 USA

Synonyms: hidden

= Stachys ingrata Greene; Pittonia 3: 341 (1898)

species :

Stachys strictiflora C.Y.Wu; Acta Phytotax. Sin. 10: 220 (1965)

Distribution: China (Yunnan)

36 CHI

variant :

var. latidens C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 221 (1965)

Distribution: China (Yunnan)

36 CHI

species :

Stachys subaphylla Rech.f.; Pl. Syst. Evol. 134: 288 (1980)

Distribution: Iran (N-Iran)

34 IRN

Remarks: clade 12 Otisia

species :

Stachys sublobata Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 354 (1910)

Distribution: South Africa (W-Cape Prov.)

27 SAF

species :

Stachys subnuda Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 50 (1836)

Distribution: Turkey (E-Anatolia, SE-Anatolia)

34 TUR

species :

Stachys swainsonii Benth.; Labiat. Gen. Spec. [Bentham] 535 (1834)

subspecies :

subsp. swainsonii

Distribution: C-Greece (Mt. Giona to Mt. Parnassos)

13 GRC

Remarks: clade 11 Swainsoniana, swainsonii agg.

Synonyms: hidden

= Prasium hirsutum Poir.; Encycl. [J.Lamarck et al.] 5: 611 (1804)

= Prasium minus Vahl ex Walp.; Repert. Bot. Syst. [Walpers] 3: 819 (1845)

subspecies :

subsp. argolica (Boiss.) Phitos & Damboldt; Ber. Deutsch. Bot. Ges. 82: 600 (1969)

Distribution: S-Greece (NE-Peloponnisos)

13 GRC

Synonyms: hidden

= Stachys argolica Boiss.; Diagn. Pl. Orient. 12: 83 (1853)

subspecies :

subsp. scyronica (Boiss.) Phitos & Damboldt; Ber. Deutsch. Bot. Ges. 82: 600 (1969)

Distribution: C-Greece (around Korinth)

13 GRC

Synonyms: hidden

= Stachys scyronica Boiss.; Diagn. Pl. Orient. 5: 31 (1844)

= Stachys swainsonii var. scyronica (Boiss.) Nyman; Consp. Fl. Eur. 580 (1881)

species :

Stachys sylvatica L.; Sp. Pl. [Linnaeus] 2: 580 (1753)

Name: Wald-Ziest (DE), Hedge Woundwort (EN)

Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Italy; Sicily; Malta; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (mountains of N-, NE-Greece, south to Mt. Pelion); Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Ukraine; Crimea; Madeira (Madeira Isl.); Siberia (Altai, Krasnoyarsk, W-Siberia); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (E-Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia); Iran (N-Iran, Iranian Aserbaijan); China (Xinjiang); China [I]; NW-India (Himachal Pradesh, Jammu & Kashmir); New Zealand [I]; Canada [I] (Ontario [I]); USA [I] (New Jersey [I], New York [I], Ohio [I], Pennsylvania [I])

10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS COR FRA POR SPA 13 ALB BIH BUL CRO GRC ITA KOS MLT MNT NMA ROM SIC SLV SMA SRB TUE 14 BLR EST KRY LAT LIT RUS UKR 21 MDR 30 SIB 32 KAZ KGZ TKM UZB 33 ARM AZE GEO NCS 34 IRN TUR 36 CHI 40 IND 51 nzl 72 cdn 75 usa

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys canariensis Jacq.; Icon. Pl. Rar. [Jacquin] 1: 11, t. 108 (1787)

= Stachys cordata Gilib.; Fl. Lit. Inch. 1: 80 (1782), opus utique oppr.

= Stachys cordatifolia Gilib.; Exerc. Phyt. 1: 91 (1792), opus utique oppr.

= Stachys foetida Gueldenst. ex Ledeb.; Fl. Ross. [Ledebour] 3: 413 (1849)

= Stachys glaucescens Muss.-Puschk. ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 2: 736 (1825)

= Stachys sylvatica f. bracteata (Beck) L.Gross; Mitt. Bad. Bot. Vereins 210/211: 82 (1906)

= Stachys sylvatica var. minus Gray; Nat. Arr. Brit. Pl. 2: 372 (1821 publ. 1822)

= Stachys sylvatica unranked mollissima Schultheiss; Abh. Naturhist. Ges. Nürnberg 13, Beil. 1 [Phan.-Fl. Nürnberg-Erlangen]: 633 (1900)

= Stachys sylvatica var. tomentosa Celak.; Arch. Naturwiss. Landesdurchf. Böhmen 4(3) (Prodr. Fl. Böhmen 4): 840 (1881)

= Stachys trapezuntea Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 38 (1859)

species :

Stachys taliensis C.Y.Wu; Acta Phytotax. Sin. 10: 225 (1965)

Distribution: China (Yunnan)

36 CHI

species :

Stachys talyschensis Kapeller; Zametki Syst. Geogr. Rast. 15: 48 (1949)

Distribution: +Azerbaijan

33 AZE

species :

Stachys tamaulipana B.L.Turner; Phytologia 95(1): 28 (2013)

Distribution: Mexico (Nuevo León, Tamaulipas)

79 MEX

species :

Stachys tenerrima Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 32 (1934)

Distribution: Mexico (Baja California Sur)

79 MEX

species :

Stachys tenuifolia Willd.; Sp. Pl., ed. 4 [Willdenow] 3: 100 (1800)

Name: Smooth hedgenettle

Distribution: Canada (Manitoba, New Brunswick, Ontario, Québec); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, South Dakota, Tennessee, Texas, Virginia, Vermont, Wisconsin, West Virginia)

71 CDN 72 CDN 74 USA 75 USA 77 USA 78 USA

Remarks: clade 2 Stachys s. str.

Synonyms: hidden

= Stachys cincinnatensis Kuntze; Revis. Gen. Pl. 2: 531 (1891)

= Stachys glabra Riddell; W. J. Med. Phys. Sci. 9(36): 583 (1836)

= Stachys palustris var. glabra (Riddell) A.Gray; Manual [A.Gray], ed. 2: 317 (1856)

= Stachys palustris var. macrocalyx Jenn.; J. Wash. Acad. Sci. 10: 458 (1920)

= Stachys tenuifolia var. aspera Fernald; Rhodora 10: 85 (1908)

= Stachys tenuifolia var. perlonga Fernald; Rhodora 45: 466 (1943)

= Stachys tenuifolia var. platyphylla Fernald; Rhodora 45: 468 (1943)

species :

Stachys tetragona Boiss. & Heldr.; Fl. Orient. [Boissier] 4(2): 736 (1879)

Distribution: C-Greece (Sporades, N-Evvoia)

13 GRC

Remarks: clade 11 Swainsoniana

species :

Stachys thirkei K.Koch; Linnaea 21: 685 (1849)

Name: Thirke-Ziest (DE)

Distribution: Germany [I]; S-Italy; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Bulgaria; European Turkey; Greece (very rare C-, NE-Greece, N-Peloponnisos); East Aegaean Isl. (Chios); Turkey (N-Anatolia, NW-Anatolia: Bithynia, W-Anatolia, WN-Anatolia)

11 ger 13 BIH BUL CRO GRC ITA KOS MNT NMA SRB TUE 34 EAI TUR

Remarks: germanica agg.

Synonyms: hidden

= Stachys janiana Ces., Pass. & Gibelli; Comp. Fl. Ital. [Cesati] 318 (1874)

= Stachys thirkei var. condensata Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)

species :

Stachys thunbergii Benth.; Labiat. Gen. Spec. [Bentham] 540 (1833)

Distribution: South Africa (W-Cape Prov., E-Cape Prov.)

27 SAF

Synonyms: hidden

= Galeopsis hispida Thunb.; Prodr. Pl. Cap. 2: 96 (1800)

= Stachys hispida (Thunb.) Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 263 (1896), nom. illeg.

species :

Stachys thuspeinantha Sennikov; Checkl. Vasc. Pl. Tian-Shan 351 (2021), nom. nov.

Distribution: Kazakhstan; Turkmenistan; Iraq (W-Iraq: Desert, NW-Iraq); Iran (EC-Iran, NE-Iran: Mts., Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Farah, Herat); Syria (C-Syrian Desert); Jordania (S-Jordania, W-Jordania); Saudi Arabia (N-Saudi Arabia)

32 KAZ TKM 34 AFG IRN IRQ JOR SYR 35 SAU

Remarks: clade 12 Otisia

Synonyms: hidden

= Stachys musili Velen.; Sitzungsber. Königl. Böhm. Ges. Wiss. Prag, Math.-Naturwiss. Cl. (Vestn. Král. Ceské Spolecn. Nauk., Tr. Mat.-Prír.) 1911: 5 (1911 publ. 1912)

= Tapeinanthus persicus Boiss.; Prodr. [A.P. de Candolle] 12: 436 (1848)

= Thuspeinanta persica (Boiss.) Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 229 (1896)

species :

Stachys tlaxiacana B.L.Turner; Phytologia 95(1): 28 (2013)

Distribution: Mexico (Oaxaca)

79 MEX

species :

Stachys tmolea Boiss.; Diagn. Pl. Orient. 5: 29 (1844)

Distribution: Turkey (Inner Anatolia, NW-Anatolia: Bithynia, W-Anatolia, WN-Anatolia)

34 TUR

Remarks: germanica agg.

species :

Stachys torresii B.L.Turner; Phytologia 76(5): 397 (1994)

Distribution: Mexico (Oaxaca)

79 MEX

species :

Stachys tournefortii Poir.; Encycl. [J.Lamarck et al.] 13: 227 (1817)

Distribution: NW-Crete; Libya

20 LBY

Remarks: germanica agg.

Synonyms: hidden

= Stachys sieberi K.Koch; Linnaea 21: 685 (1849)

species :

Stachys trichophylla Baker; J. Linn. Soc., Bot. 25: 342 (1890)

Distribution: Madagascar

29 MDG

species :

Stachys trinervis Aitch. & Hemsl.; Trans. Linn. Soc. London, Bot., Ser. 2, 3: 9, t. 43 (1888)

Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran); Afghanistan (Badghys, Farah, Faryab, Herat)

32 TKM 34 AFG IRN

Remarks: clade 12 Otisia

species :

Stachys truncata Kunze ex Benth.; Labiat. Gen. Spec. [Bentham] 550 (1834)

Distribution: Chile (Coquimbo, Valparaíso, Maule, Aisén)

85 CLE

species :

Stachys tubulosa MacOwan; Bull. Misc. Inform. Kew 1893: 13 (1893)

Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.); Eswatini [Swaziland]

27 SAF SWZ

Synonyms: hidden

= Lamiostachys tubulosa (MacOwan) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 91(8): 1258 (2006)

= Stachys dolichodeira Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1081 (1903)

species :

Stachys tundjeliensis Kit Tan & Sorger; Pl. Syst. Evol. 155(1-4): 99 (1987)

Distribution: Turkey

34 TUR

species :

Stachys turcomanica Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 463 (1886)

Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran)

32 TKM 34 IRN

Remarks: clade 12 Otisia

species :

Stachys turkestanica (Regel) Popov ex Knorring; Fl. URSS [Komarov et al., eds.] 21: 209 (1954)

Distribution: Uzbekistan; Kyrgyzstan; Tajikistan

32 KGZ TZK UZB

Synonyms: hidden

= Stachys alpina var. turkestanica Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada, prepr. 6: 366 (1879)

= Stachys hissarica subsp. turkestanica (Regel) Derv.-Sok.; Novosti Sist. Vyssh. Rast. 14: 205 (1977)

= Stachys hissarica var. turkestanica (Regel) Krestovsk.; Novosti Sist. Vyssh. Rast. 42: 206 (2011)

= Stachys tschatkalensis Knorring; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 15: 344 (1953)

species :

Stachys turneri Rzed. & Calderón; Acta Bot. Mex. 32: 4 (1995)

Distribution: Mexico (Guanajuato)

79 MEX

species :

Stachys tymphaea Hausskn.; Mitt. Geogr. Ges. (Thüringen) Jena 5: 70 (1886)

Distribution: C- & S-Italy; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (mountains of NE-, NC-, NW- to C-Greece, south to Mt. Parnassos)

13 ALB BUL GRC ITA KOS NMA SRB

Remarks: germanica agg.

Synonyms: hidden

= Stachys alpina var. reinertii (Heldr.) Stoj.; Fl. Reipubl. Popularis Bulg. (Fl. Narodna Republ. Bulg.) 9: 914 (1967)

= Stachys germanica subsp. tymphaea (Hausskn.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 276 (1974)

= Stachys germanica var. reinertii (Heldr.) Nyman; Consp. Fl. Eur. Suppl. 2: 251 (1889)

= Stachys reinertii Heldr.; Exsicc. Herb. Graecum Normale 1857: 743 (1857)

species :

Stachys tysonii Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 357 (1910)

Distribution: South Africa (Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho

27 LES SAF

Synonyms: hidden

= Stachys obtusifolia var. angustifolia Skan

species :

Stachys uniflora A.Pool; Novon 17(1): 64 (2007)

Distribution: Costa Rica; Panama

80 COS PAN

species :

Stachys venezuelana Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 114 (1898)

Distribution: Venezuela (Merida, Trujillo); Ecuador

82 VEN 83 ECU

species :

Stachys venulosa Greene; Pittonia 1: 157 (1888)

Distribution: Mexico (Durango, etc.)

79 MEX

species :

Stachys veroniciformis Rech.f.; Pl. Syst. Evol. 134: 289 (1980)

Distribution: Iran (SW-Iran)

34 IRN

species :

Stachys virgata Bory & Chaub.; Expéd. Sci. Morée, Bot. 3(2): 166 (1832)

Distribution: S-Greece (E-Peloponnisos)

13 GRC

Synonyms: hidden

= Stachys zuccarinii Benth.; Labiat. Gen. Spec. [Bentham] 535 (1833)

species :

Stachys viscosa Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 49 (1836)

Distribution: Georgia [Caucasus]; Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia); Iran (Iranian Aserbaijan)

33 AZE GEO 34 IRN TUR

Synonyms: hidden

= Stachys laetivirens Kotschy & Boiss. ex Rech.f.; Bot. Jahrb. Syst. 71: 529 (1941)

= Stachys odontophylla Freyn; Oesterr. Bot. Z. 41: 59 (1891)

species :

Stachys viticina Boiss.; Diagn. Pl. Orient. 12: 77 (1853) et Fl. 4: 724 (1879)

Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon, coastal W-Lebanon); Syria (W-Syrian Mountains); Israel (coastal W-Israel, Rift Valley, N-Israel)

34 ISR LEB SYR TUR

species :

Stachys vulnerabilis Rzed. & Calderón; Acta Bot. Mex. 3: 3 (1988)

Distribution: Mexico (Coahuila, Nuevo León)

79 MEX

species :

Stachys vuralii Yildiz, Dirmenci & Akçiçek; Ann. Bot. Fenn. 48(5): 403 (2011)

Distribution: Croatia; Azerbaijan; Turkey

13 CRO 33 AZE 34 TUR

species :

Stachys willemsei Kit Tan & Hedge; Notes Roy. Bot. Gard. Edinburgh 45(3): 451 (1988 publ. 1989)

Distribution: Turkey

34 TUR

species :

Stachys woronowii (Schischk. ex Grossh.) R.R.Mill; Fl. Turkey 7: 253 (1982)

Distribution: Armenia; Turkey (E-Anatolia, Inner Anatolia, NE-Anatolia, SSW-Anatolia); Iran (Iranian Aserbaijan); Lebanon (Antilebanon, C-Lebanon); China (Xinjiang)

33 ARM 34 IRN LEB TUR 36 CHI

Synonyms: hidden

= Sideritis balansae Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 35 (1859) et Fl. 4: 707 (1879) (non Stachys balansae Boiss. & Kotschy (1879))

= Sideritis woronowii Schischk. ex Grossh.; Fl. Kavkaza [Grossheim] 3: 290 (1932)

= Stachys pseudosideritis R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33(2): 291 (1974) (nom. nov.)

species :

Stachys xanthantha C.Y.Wu; Acta Phytotax. Sin. 10: 224 (1965)

Distribution: China (Sichuan)

36 CHI

Synonyms: hidden

= Stachys xanthantha var. gracilis C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 225 (1965)

species :

Stachys yemenensis Hedge; Notes Roy. Bot. Gard. Edinburgh 40: 71, f. 4 (1982)

Distribution: Saudi Arabia (Asir); Yemen (N-Inner Yemen, W-Yemen)

35 SAU YEM

Remarks: clade 12 Otisia

species :

Stachys yildirimlii Dinç; Ann. Bot. Fenn. 43(2): 143 (2006)

Distribution: Turkey

34 TUR

species :

Stachys yingzuijieensis L.Wu & Y.P.Chen; PhytoKeys 236: 127 (2023)

Distribution: China (Hunan)

36 CHI

species :

Stachys zepcensis Formánek; Oesterr. Bot. Z. 38: 383 (1888)

Distribution: Bosnia & Hercegovina; Serbia

13 BIH SRB

Remarks: clade 11 Swainsoniana

Synonyms: hidden

= Stachys anisochila subsp. zepcensis (Formánek) Vandas; Reliq. Formán. [Vandas] 469 (1909)

= Stachys labiosa var. zepcensis (Formánek) K.Malý; Verh. K.K. Zool.-Bot. Ges. Wien 54: 246 (1904)

= Stachys recta var. chrysophaea Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 183 (1924)

species :

Stachys zeyheri Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 363 (1910)

Distribution: South Africa (W-Cape Prov.)

27 SAF

species :

Stachys zoharyana Eig; Palestine J. Bot., Jerusalem Ser. 4: 172, f. 1 (1948)

Distribution: Israel (coastal W-Israel, N-Israel)

34 ISR

species :

Stachys ×ambigua Sm.; Engl. Bot. [Smith & Sowerby] 30: t. 2089 (1809)

Name: Hybrid Woundwort (EN)

Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Belgium; Germany (Baden-Württemberg, Bayern, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Sachsen, Sachsen-Anhalt, Thüringen); France; Czech Republic; Slovakia; Channel Isl. (Alderney, Guernsey, Jersey); Turkey (E-Anatolia); etc.

10 GRB IRE 11 BGM CZE GER SLO 12 CIS FRA 34 TUR

Remarks: clade 2 Stachys s. str., Stachys palustris x sylvatica

Synonyms: hidden

= Stachys palustris subsp. ambigua (Sm.) Arcang.; Comp. Fl. Ital. [Arcangeli] 552 (1882)

= Stachys palustris subsp. ambigua (Sm.) Hartm.; Sv. Norsk Excurs.-Fl. 85 (1846)

= Stachys palustris var. ambigua (Sm.) Bluff & Fingerh.; Comp. Fl. German. [Bluff & Fingerhuth] 1: 31 (1825)

= Stachys palustris var. ambigua (Sm.) Hook.; Fl. Bor.-Amer. [Hooker] 2: 116 (1838)

= Stachys palustris ×sylvatica var. wirtgenii (F.W.Schultz) Bornm.; Beih. Bot. Centralbl. 37(2): 315 (1919)

= Stachys palustri-sylvatica Schiede; De Pl. Hybr. Spont. Nat. 42 (1825)

= Stachys ×ambigua f. ellipticus Clos; Bull. Soc. Bot. France 36: 71 (1889)

= Stachys ×ambigua f. longepetiolatus Clos; Bull. Soc. Bot. France 36: 71 (1889)

= Stachys ×ambigua f. petiolatus Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2435 (1927)

= Stachys ×ambigua var. elliptica Wallr.; Scholion zu Hampe´s Prodr. Fl. Hercyn.: 222 (1840), nom. inval.

= Stachys ×ambigua var. lanceolata Wallr.; Scholion zu Hampe´s Prodr. Fl. Hercyn.: 222 (1840)

= Stachys ×ambigua var. ovatifolia Wallr.; Scholion zu Hampe´s Prodr. Fl. Hercyn.: 222 (1840), nom. inval.

= Stachys ×wirtgenii F.W.Schultz; Arch. Fl. 1(2): 117 (1861), nom. inval.

species :

Stachys ×bodeana Bunge; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 21(1): 71 (1873), nom. nov.

Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)

34 IRN

Remarks: Stachys inflata x lavandulifolia.

Synonyms: hidden

= Stachys lavandulifolia var. brachyodon Boiss.; Fl. Orient. [Boissier] 4: 743 (1873)

= Stachys ×tomentosa Benth.; Prodr. [A.P. de Candolle] 12: 489 (1848), nom. illeg.

= Stachys ×tomentosa Benth. ex Rech.f.; Fl. Iranica [Rechinger] 150: 395, t. 360 (1982)

species :

Stachys ×burrii P.H.Davis; Kew Bull. [6]: 109 (1951)

Distribution: Turkey (SW-Anatolia)

34 TUR

Remarks: clade 1 Eriostomum, germanica agg.; Stachys cretica ssp. mersinaea x Stachys sericantha

species :

Stachys ×cryptadenia Rech.f.; Bot. Jahrb. Syst. 71: 538 (1941)

Distribution: Croatia (Dalmatia)

13 CRO

Remarks: clade 1 Eriostomum, germanica agg.; Stachys cretica ssp. salviifolia x thrikei

species :

Stachys ×digenea Legué; Bull. Soc. Bot. France 40: 213 (1893)

Distribution: Czech Republic; Slovakia; etc.

11 CZE SLO

Remarks: clade 1 Eriostomum, germanica agg.; Stachys alpina x germanica

species :

Stachys ×leucomalla Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 49: 271 (1940)

Distribution: Iran (N-Iran)

34 IRN

Remarks: Stachys inflata x laxa

species :

Stachys ×medebachensis Feld & O.Koenen; Jahresber. Westfäl. Prov.-Vereins Wiss. 41: 183 (1913)

Distribution: Germany (Bayern, Nordrhein-Westfalen); Switzerland; etc.

11 GER SWI

Remarks: clade 1 Eriostomum, germanica agg.; Stachys alpina x sylvatica

Synonyms: hidden

= Stachys ×hybrida Brügger; Jahresber. Naturforsch. Ges. Graubünden, N. F. 23-24: 102 (1880), nom. inval.

= Stachys ×medebachensis f. superalpina Feld & O.Koenen; Jahresber. Westfäl. Prov.-Vereins Wiss. 41: 183 (1913), nom. inval.

= Stachys ×medebachensis f. supersilatica Feld & O.Koenen; Jahresber. Westfäl. Prov.-Vereins Wiss. 41: 183 (1913), nom. inval.

species :

Stachys ×mirabilis Rouy; Fl. France [Rouy & Foucaud] 11: 310 (1909)

Distribution: France; etc.

12 FRA

Remarks: clade 1 Eriostomum, germanica agg.; Stachys germanica x palustris

species :

Stachys ×paraplesia Rech.f.; Fl. Iranica [Rechinger] 150: 395, t. 361 (1982)

Distribution: Iran (EC-Iran, N-Iran)

34 IRN

Remarks: Stachys inflata x turcomanica

species :

Stachys ×sintenisii Gand.; Bull. Soc. Bot. France 65: 68 (1918)

Distribution: Turkmenistan; NE-Iran (EC-Iran, NE-Iran: Mts.)

32 TKM 34 IRN

Remarks: Stachys lavandulifolia x turcomanica

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A complete, synonymic checklist of the Higher Plants of the World

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A complete, synonymic checklist of the Higher Plants of the World

Version 25.07 (July 5th, 2025)

Covering 356,783 accepted species,
8,788 hybrids, 48,065 accepted infraspecific taxa and 1,003,637 synonyms,
**in total 1,417,272* names**

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

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How to Cite:

Realization

Search

+ Family

Filter

Order

Family (26)

Genus (224)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 25.07 (July 5th, 2025)

Covering 356,783 accepted species,8,788 hybrids, 48,065 accepted infraspecific taxa and 1,003,637 synonyms,in total 1,417,272* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 356,783 accepted species,
8,788 hybrids, 48,065 accepted infraspecific taxa and 1,003,637 synonyms,
**in total 1,417,272* names**

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]