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041.0000 Family Polypodiaceae J. & C.Presl; Delic. Prag. 159 (1822)
67 genera, ca. 1700 spp. Phylogeny: Wei & Zhang 2022
Species listed in database: 1750; hybrids: 50
= Platyceriaceae Ching; Acta Phytotax. Sin. 16(3): 18 (1978)
= Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
= Grammitidaceae Newman; Hist. Brit. Ferns 7 (1840) (Grammitaceae auct., missp.)
= Gymnogrammitidaceae Ching; Acta Phytotax. Sin. 11: 12 (1966)
= Loxogrammaceae Ching ex Pic.Serm.; Webbia 29: 11 (1975)
= Platyceriaceae Ching; Sunyatsenia 5: ? (1940)
= Pleurosoriopsidaceae Sa.Kurata & Ikebe ex Ching; Acta Phytotax. Sin. 16(4): 17 (1978)
041.9001 Genus Terpsichore A.R.Sm.; Novon 3: 478-489 (1993)
Distribution: Neotropics, 1 to trop. Africa. Checklist revised for this database by Parris (2020)
Species listed in database: 16
species :
Terpsichore alfaroi (Donn.Sm.) A.R.Sm.; Novon 3: 485 (1993)
Distribution: Guatemala; Costa Rica; Colombia (Cundinamarca, Santander); Venezuela (Aragua); ?Brazil (?Rio de Janeiro)
80 COS GUA 82 CLM VEN
Synonyms: hidden
= Alansmia alfaroi (Donn.Sm.) Moguel & M.Kessler; Brittonia 63(2): 239 (2011)
= Ctenopteris alfaroi (Donn.Sm.) Copel.; Philipp. J. Sci. 84: 433 (1956)
= Grammitis alfaroi (Donn.Sm.) C.V.Morton; Contr. U.S. Natl. Herb. 38: 103 (1967)
= Polypodium alfaroi Donn.Sm.; Bot. Gaz. 33: 262 (1902)
= Polypodium oligosorum Moritz ex Mett.; Linnaea 36: 132 (1869), nom. illeg., non Klotzsch (1847)
species :
Terpsichore alsophilicola (Christ) A.R.Sm.; Novon 3(4): 485 (1993)
Distribution: Costa Rica; Panama; Colombia (Chocó); Ecuador
80 COS PAN 82 CLM 83 ECU
Synonyms: hidden
= Ctenopteris alsophilicola (Christ) Copel.; Philipp. J. Sci. 84: 423 (1956)
= Grammitis alsophilicola (Christ) F.Seym.; Phytologia 31: 176 (1975)
= Polypodium alsophilicola Christ; Bull. Soc. Bot. Genève, Sér. 2, 1: 219 (1909)
species :
Terpsichore asplenifolia (L.) A.R.Sm.; Novon 3(4): 485 (1993)
Distribution: Mexico (Chiapas, Guerrero, Michoacan, Oaxaca, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Nevis; St. Kitts; Montserrat; Guadeloupe; Dominica; Martinique; St. Lucia; St. Vincent; Grenada; Trinidad; Colombia (Antioquia, Cesar, Cundinamarca, Huila, Nariño); Venezuela (Amazonas, Anzoategui, Aragua, Bolívar, Carabobo, Cojedes, Venezuela Distrito Federal, Guarico, Lara, Merida, Miranda, Portuguesa, Sucre, Tachira, Trujillo, Yaracuy); Guyana; Ecuador; Peru; Bolivia (Cochabamba, La Paz); Brazil (Amazonas, Bahia)
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 CUB DOM GDL GRE HIS JAM MAR MTS PUE STK STL STV TRT 82 CLM GUY VEN 83 BOL ECU PER 84 BRA
Synonyms: hidden
= Ctenopteris asplenifolia (L.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.]: 219 (1947)
= Grammitis asplenifolia (L.) Proctor; Brit. Fern Gaz. 9: 76 (1962)
= Phegopteris asplenifolia (L.) Fée; Mém. Foug. 5 (Gen. Filic.) 243 (1852)
= Polypodium asplenifolium L.; Sp. Pl. [Linnaeus] 2: 1084 (1753)
= Polypodium laxifrons Liebm.; Kongel. Danske Vidensk. Selsk. Skr., Naturvidensk. Math. Afd., Ser. 5, 1: 204 (1899)
= Polypodium laxifrons var. lividum Kuhn; Bot. Jahrb. Syst. 24(1): 126 (1897)
= Polypodium suspensum var. asplenifolium (L.) Krug; Bot. Jahrb. Syst. 24(1): 127 (1897)
species :
Terpsichore atroviridis (Copel.) A.R.Sm.; Novon 3(4): 486 (1993)
Distribution: Nicaragua; Costa Rica; Panama
80 COS NIC PAN
Synonyms: hidden
= Ctenopteris atroviridis Copel.; Philipp. J. Sci. 84: 461 (1956)
= Grammitis atroviridis (Copel.) F.Seym.; Phytologia 31: 176 (1975)
species :
Terpsichore bipinnata (Stolze) A.R.Sm.; Novon 5(1): 21 (1995)
Distribution: Peru
83 PER
Synonyms: hidden
= Grammitis bipinnata Stolze; Fieldiana, Bot. 32: 92 (1993)
species :
Terpsichore chrysleri (Copel.) A.R.Sm.; Novon 3: 486 (1993)
Distribution: Costa Rica; Panama; Jamaica; Hispaniola (Haiti, Dominican Republic); Colombia (Antioquia, Boyacá, Cundinamarca, Huila, Magdalena); Venezuela (Aragua, Venezuela Distrito Federal, Lara, Merida, Tachira); Ecuador; Bolivia (Cochabamba, La Paz); Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, São Paulo)
80 COS PAN 81 HIS JAM 82 CLM VEN 83 BOL ECU 84 BRA
Synonyms: hidden
= Ctenopteris chrysleri Copel.; Philipp. J. Sci. 84: 448 (1956)
= Grammitis chrysleri (Copel.) Proctor; Brit. Fern Gaz. 9: 218 (1965)
= Polypodium arcuatum Moritz ex Mett.; Abh. Senckenberg. Naturf. Ges. 2(1) [Polypodium]: 56 (1856), nom. illeg.
= Polypodium chrysleri (Copel.) Proctor; Bull. Inst. Jamaica, Sci. Ser. 5: 46 (1953)
= Polypodium repandum Fée; Crypt. Vasc. Brésil 1: 87, t. 29, f. 1 (1869)
species :
Terpsichore dolorensis (Hieron.) A.R.Sm.; Novon 3(4): 486 (1993)
Distribution: Colombia (Antioquia)
82 CLM
Remarks: status uncertain, possibly a synonym of T. asplenifolia
Synonyms: hidden
= Ctenopteris dolorensis (Hieron.) Copel.; Philipp. J. Sci. 84: 449 (1956)
= Grammitis dolorensis (Hieron.) Lellinger; Proc. Biol. Soc. Washington 89(61): 714 (1977)
= Polypodium dolorense Hieron.; Bot. Jahrb. Syst. 34: 512 (1904)
species :
Terpsichore eggersii (Baker ex Hook.) A.R.Sm.; Novon 3(4): 486 (1993)
Distribution: St. Kitts; Montserrat; Guadeloupe; Dominica; Martinique; St. Vincent; Grenada
81 DOM GDL GRE MAR MTS STK STV
Synonyms: hidden
= Grammitis eggersii (Baker) Proctor; Brit. Fern Gaz. 9: 77 (1962)
= Polypodium eggersii Baker; Hooker´s Icon. Pl. t. 1671 (1886)
= Polypodium tenuiculum var. husnotii Krug; Bot. Jahrb. Syst. 24(1): 127 (1897)
species :
Terpsichore exornans (Maxon) A.R.Sm.; Novon 3(4): 486 (1993)
Distribution: Jamaica
81 JAM
Synonyms: hidden
= Ctenopteris exornans (Maxon) Copel.; Philipp. J. Sci. 84: 446 (1956)
= Grammitis exornans (Maxon) Proctor; Brit. Fern Gaz. 9: 219 (1965)
= Polypodium exornans Maxon; Amer. Fern J. 18: 47 (1928)
species :
Terpsichore flexuosa (Maxon) A.R.Sm.; Novon 5(1): 21 (1995)
Distribution: Cuba
81 CUB
Synonyms: hidden
= Ctenopteris flexuosa (Maxon) Copel.; Philipp. J. Sci. 84: 434 (1956)
= Grammitis maxoniana Lellinger; Amer. Fern J. 74(2): 58 (1984), nom. nov.
= Grammitis terpsichore Christenh.; Global Fl. 4: 49 (2018), nom. nov.
= Lellingeria flexuosa (Maxon) A.R.Sm. & R.C.Moran; Amer. Fern J. 81(3): 84 (1991)
= Polypodium flexuosum Maxon; Contr. U.S. Natl. Herb. 17: 597, t. 42 (1916) (non Grammitis flexuosa Humb. & Bonpl. 1816)
species :
Terpsichore hanekeana (Proctor) A.R.Sm.; Novon 3(4): 486 (1993)
Distribution: Puerto Rico
81 PUE
Synonyms: hidden
= Grammitis hanekeana Proctor; Mem. New York Bot. Gard. 53: 341 (1989)
species :
Terpsichore jenmanii (Underw. ex Maxon) A.R.Sm.; Novon 3(4): 487 (1993)
Distribution: Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic)
81 CUB HIS JAM
Synonyms: hidden
= Ctenopteris jenmanii (Underw. ex Maxon) Copel.; Philipp. J. Sci. 84: 442 (1956)
= Grammitis jenmanii (Underw. ex Maxon) Proctor; Brit. Fern Gaz. 9: 219 (1965)
= Polypodium jenmanii Underw.; Contr. U.S. Natl. Herb. 16: 62 (1912)
= Polypodium lasiolepis Jenman; Bull. Bot. Dept., Jamaica 4: 118 (1897), nom. illeg., non Mett. (1869)
species :
Terpsichore lehmanniana (Hieron.) A.R.Sm.; Novon 3(4): 487 (1993)
Distribution: Guatemala; Belize; ?El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Colombia (Antioquia); Ecuador
80 BLZ COS GUA HON NIC PAN 82 CLM 83 ECU
Synonyms: hidden
= Ctenopteris lehmanniana (Hieron.) Copel.; Philipp. J. Sci. 84: 448 (1956)
= Grammitis lehmanniana (Hieron.) C.V.Morton; Contr. U.S. Natl. Herb. 38: 104 (1967)
= Polypodium lehmannianum Hieron.; Bot. Jahrb. Syst. 34: 513 (1904) (non Polypodium lehmanni Mett. (1857))
= Polypodium pastoense C.Chr.; Index Filic. 551 (1906) (nom. illeg. superfl.)
= Polypodium sublongipes Christ; Bull. Soc. Bot. Genève, Sér. 2, 1: 218 (1909)
species :
Terpsichore liogieri (Proctor) A.R.Sm.; Novon 3(4): 487 (1993)
Distribution: Puerto Rico
81 PUE
Synonyms: hidden
= Grammitis liogieri Proctor; Mem. New York Bot. Gard. 53: 341 (1989)
species :
Terpsichore paulistana (Brade & Rosenst.) A.R.Sm.; Novon 3(4): 488 (1993)
Distribution: Brazil (Rio de Janeiro)
84 BRA
Synonyms: hidden
= Grammitis paulistana (Brade & Rosenst.) Christenh.; Global Fl. 4: 47 (2018)
= Polypodium paulistanum Brade & Rosenst. ex Brade; Arch. Inst. Biol. (São Paulo) 2: 3 (1935)
species :
Terpsichore staheliana (Posth.) A.R.Sm.; Novon 3(4): 488 (1993)
Distribution: Costa Rica; Panama; Colombia (Chocó); Venezuela (Bolívar); Surinam; French Guiana
80 COS PAN 82 CLM FRG SUR VEN
Synonyms: hidden
= Ctenopteris staheliana (Posth.) K.U.Kramer; Amer. Fern J. 64: 114 (1974)
= Grammitis staheliana (Posth.) Lellinger; Proc. Biol. Soc. Washington 89(61): 716 (1977)
= Polypodium stahelianum Posth.; Recueil Trav. Bot. Néerl. 23: 401, f. 2 (1924)
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A complete, synonymic checklist of the Higher Plants of the World

Version 26.02 (Feb. 1st, 2026)

Covering 358,774 accepted species,
8,847 hybrids, 48,003 accepted infraspecific taxa and 1,006,442 synonyms,
in total 1,422,066* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

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