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044.0000 Family Thelypteridaceae Ching ex Pic.Serm.; Webbia 24: 711 (1970)
5-34 genera (depending on genus concept), 1100-1200 spp. New genus concept adopted here from Fawcett & Smith (2021), based on molecular results.
Species listed in database: 1180; hybrids: 38
044.2001 Genus Thelypteris Schmidel; Icon. Pl., ed. Keller: 3, 45, t. 11, text (1763), nom. cons.
Distribution: 1 sp. north-temperate, 1 sp. south-temperate
Species listed in database: 2
= Dryopteris sect.Asterochlaena C.Chr.; Biol. Arbejder Tilegnede Eug. Warming 84 (1911)
species :
Thelypteris confluens (Thunb.) C.V.Morton; Contr. U.S. Natl. Herb. 38: 71 (1967)
Distribution: Nigeria; Congo [Brazzaville]; D.R.Congo [Zaire]; Rwanda; Burundi; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; Namibia; Botswana; South Africa (widespread); Lesotho; Eswatini [Swaziland]; Madagascar; China (S-Yunnan); S-India (?Odisha, Tamil Nadu); Sri Lanka; Myanmar; N-Thailand; Laos; Vietnam; N-Sumatra; New Guinea (Irian Jaya, Papua New Guinea); Australia (SE-Queensland, NE-Victoria); New Zealand (North Isl.); Argentina (Buenos Aires, Chaco)
22 NGA 23 BUR CON RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 BOT LES NAM SAF SWZ 29 MDG 36 CHI 40 IND SRL 41 LAO MYA THA VIE 42 NWG SUM 50 AUS 51 NZL 85 ARG
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium dregei Fée; Mém. Foug. 5 (Gen. Filic.) 292 (1852)
= Aspidium squamigerum (Schltdl.) Fée; Mém. Foug. 8 (Icon. Esp. Nouv.): 104 (1857)
= Aspidium squamulosum Kaulf.; Adumbr. Pl. 1: 23 (1825), sphalm.
= Aspidium thelypteris var. squamigerum Schltdl.; Adumbr. Pl. 23, t. 11 (1825)
= Athyrium cyclolepis C.Chr. & Tardieu; Bull. Mus. Natl. Hist. Nat., Sér. 2, 6: 109, f. 3-4 (1934)
= Dryopteris cabrerae Weath.; Bol. Soc. Argent. Bot. 3: 31, f. 1-5 (1949)
= Dryopteris thelypteris subsp. squamigera (Schltdl.) Bonap.; Notes Ptéridol. 15: 17 (1924)
= Dryopteris thelypteris var. squamigera (Schltdl.) C.Chr.; Index Filic. 297 (1905)
= Lastrea cabrerae (Weath.) Capurro; Anales Comis. Invest. Ci., Buenos Aires 2: 139 (1961)
= Lastrea fairbankii Bedd.; Ferns Brit. India, t. 254 (1867)
= Lastrea invisa J.B.Armstr.; Trans. & Proc. New Zealand Inst. 13: 363 (1882)
= Lastrea squamulosa (Kaulf.) C.Presl; Tent. Pterid. 76 (1836)
= Lastrea thelypteris var. squamigera (Schltdl.) Bedd.; Ferns Brit. India Suppl. 54 (1892)
= Nephrodium squamigerum (Schltdl.) Hook. & Arn.; Bot. Beechey Voy. 106 (1832)
= Nephrodium squamulosum (Kaulf.) Hook.f.; Bot. Antarct. Voy. II. (Fl. Nov.-Zel.) 1: 39 (1855), sphalm.
= Nephrodium thelypteris var. squamigerum (Schltdl.) Hook.; Sp. Fil. [W.J.Hooker] 4: 88 (1862) (´squamulosum´ errore)
= Pteris confluens Thunb.; Prodr. Pl. Cap. 2: 171 (1800)
= Thelypteris cabrerae (Weath.) Abbiatti; Revista Mus. La Plata, Secc. Bot. 9: 19 (1958)
= Thelypteris cyclolepis (C.Chr. & Tardieu) Tagawa; J. Jap. Bot. 26: 20 (1951)
= Thelypteris fairbankii (Bedd.) Y.X.Lin, K.Iwats. & M.G.Gilbert; Fl. China 2-3: 323 (2013)
= Thelypteris palustris var. squamigera (Schltdl.) Weath.; Contr. Gray Herb. 73: 40 (1924)
= Thelypteris squamigera (Schltdl.) Ching; Bull. Fan Mem. Inst. Biol., Bot. 6: 329 (1936) (err. typ. squamulosa)
= Thelypteris thelypteroides subsp. squamigera (Schltdl.) Soó; Acta Bot. Acad. Sci. Hung. 23(3-4): 391 (1977 publ. 1978)
species :
Thelypteris palustris Schott; Gen. Fil. [Schott] ad t. 10 (1834)
subspecies :
subsp. palustris
Name: Sumpffarn (DE), Marsh Fern (EN)
Distribution: England (England, Wales, S-Scotland); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; France; Corsica; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (scattered N- and C-Greece, Peloponnisos, Samothraki); Estonia; Latvia; Lithuania; Belarus; C-European Russia; N-European Russia; W-European Russia; Moldova; Ukraine; Crimea; Morocco; Algeria; Sudan; Siberia (Altai, Buryatia, Irkutsk, Krasnoyarsk, W-Siberia); Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (Anatolia); Lebanon; Iraq; ?Syria; Iran; Israel; Palestine; China (Hebei, Heilongjiang, Henan, Jilin, Nei Mongol, Shandong, Sichuan, Xinjiang); North Korea; South Korea (incl. Cheju Isl.); Japan; India (Himachal Pradesh, Jammu & Kashmir)
10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 COR FRA POR SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB 14 BLR EST KRY LAT LIT MLD RUS UKR 20 ALG MOR 24 SUD 30 SIB 32 KAZ KGZ TZK UZB 33 ARM AZE GEO NCS 34 IRN IRQ ISR LEB PAL TUR 36 CHI 38 JAP KOR NKO 40 IND
Remarks: subgenus Thelypteris s. str.
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Acrostichum thelypteris L.; Sp. Pl. [Linnaeus] 2: 1071 (1753) (syn. subst.)
= Aspidium palustre Gray; Nat. Arr. Brit. Pl. 2: 9 (1821), nom. superfl.
= Aspidium thelypteris (L.) Sw.; J. Bot. [Schrader] 1800(2): 40 (1802)
= Aspidium thelypteris f. incisum (Asch.) Werner Christ.; Fl. Kiel [J.A.Christiansen et al.] 305 (1922)
= Aspidium thelypteris f. rogoetzianum (Bolle) Werner Christ.; Fl. Kiel [J.A.Christiansen et al.] 305 (1922)
= Aspidium thelypteris var. incisum Asch.; Fl. Brandenburg 1: 922 (1864)
= Aspidium thelypteris var. rogoetzianum Bolle; Verh. Bot. Vereins Prov. Brandenburg 1: 73 (1859)
= Athyrium thelypteris (L.) Spreng.; Anleit. Kenntn. Gew. 3: 134 (1804)
= Dryopteris thelypteris (L.) A.Gray; Manual (Gray), ed. 1: 630 (1848)
= Dryopteris thelypteris var. koreana Nakai; Bot. Mag. (Tokyo) 45: 97 (1931)
= Filix thelypteris (L.) Farw.; Rep. (Annual) Michigan Acad. Sci. 6: 200 (1904)
= Filix-mas thelypteris (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Hemestheum thelypteris (L.) Newman; Phytologist 4: app. XXII (1851)
= Hemestheum thelypteris var. incisum (Asch.) Rouy; Fl. France [Rouy & Foucaud] 14: 402 (1913)
= Lastrea palustris (Schott) Bubani; Fl. Pyren. [Bubani] 4: 432 (1901), nom. superfl.
= Lastrea palustris (Schott) J.Sm.; Cult. Ferns 56 (1857), nom. superfl.
= Lastrea thelypteris (L.) Bory; Dict. Class. Hist. Nat. [Bory] 9: 233 (1826)
= Lastrea thelypteris (L.) C.Presl; Tent. Pterid. 76 (1836), isonym
= Nephrodium thelypteris (L.) Strempel; Fil. Berol. Syn. 32 (1822)
= Nephrodium thelypteris var. punctatum Coss.; Notes Pl. Crit. 1: 20 (1848)
= Polypodium palustre Salisb.; Prodr. Stirp. Chap. Allerton 403 (1796), nom. illeg., non Burm.f. (1768)
= Polypodium pterioides Lam.; Fl. Franç. [Lamarck] 1: 18 (1779), nom. illeg. superfl.
= Polypodium thelypteris (L.) Weis; Pl. Crypt. Fl. Gott. 307 (1770)
= Polystichum convolutum Dulac; Fl. Hautes-Pyrenées 32 (1867)
= Polystichum thelypteris (L.) Roth; Tent. Fl. Germ. 3: 77 (1799)
= Tectaria pterioides (Lam.) Lag.; Anales Ci. Nat. 5: 144 (1802), nom. illeg.
= Thelypteris palustris f. glabra H.Itô; Nov. Fl. Jap. [Nakai & Honda] 4: 127 (1939)
= Thelypteris thelypterioides subsp. glabra Holub; Taxon 21: 332 (1972)
= Thelypteris thelypteris (L.) Nieuwl.; Midl. Naturalist (Notre Dame) 1: 226 (1910), nom. inval.
subspecies :
subsp. pubescens (G.Lawson) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 280-281 (1997)
Name: Eastern marsh fern
Distribution: China (N-Jiangsu, Heilongjiang, Jilin, Shandong); Japan; Canada (Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon; USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, South Dakota, Tennessee, Texas, Virginia, Vermont, Wisconsin, West Virginia); +Bermuda; Mexico (Ciudad de Mexico, Michoacan); Cuba; ?Peru
36 CHI 38 JAP 71 CDN 72 CDN SPM 74 USA 75 USA 77 USA 78 USA 79 MEX 81 BER CUB
Remarks: subgenus Thelypteris s. str.
Synonyms: hidden
= Aspidium thelypteris f. pufferae (A.A.Eaton) B.L.Rob.; Rhodora 9: 83 (1907)
= Dryopteris thelypteris f. haleana (Fernald) C.F.Reed; Castanea 16: 71 (1951)
= Dryopteris thelypteris f. linearis (Farw.) M.Broun; Index No. Amer. Ferns: 82 (1938)
= Dryopteris thelypteris f. pufferae (A.A.Eaton) A.Prince ex Weath.; Amer. Fern J. 26: 95 (1936)
= Dryopteris thelypteris f. suaveolens (Clute) A.Prince ex Weath.; Amer. Fern J. 26: 95 (1936)
= Dryopteris thelypteris subsp. pubescens (G.Lawson) Hultén; Kungl. Svenska Vetenskapsakad. Handl., Ser. 4, 8(5): 180 (1962)
= Dryopteris thelypteris var. haleana (Fernald) M.Broun ex Weath.; Amer. Fern J. 26: 95 (1936)
= Dryopteris thelypteris var. pubescens (G.Lawson) A.Prince; Amer. Fern J. 26: 95 (1836)
= Dryopteris thelypteris var. pubescens (G.Lawson) Nakai; Bot. Mag. (Tokyo) 45: 97 (1931), isonym
= Dryopteris tremula Christ; Notul. Syst. (Paris) 1: 234 (1910)
= Filix thelypteris f. frondosa Farw.; Pap. Michigan Acad. Sci. 2: 14 (1923)
= Filix thelypteris var. linearis Farw.; Pap. Michigan Acad. Sci. 2: 14 (1923)
= Filix-mas thelypteris subvar. frondosa (Farw.) Farw.; Amer. Midl. Naturalist 12: 258 (1931)
= Filix-mas thelypteris f. frondosa (Farw.) Farw.; Pap. Michigan Acad. Sci. 2: 14 (1923)
= Filix-mas thelypteris var. linearis (Farw.) Farw.; Amer. Midl. Naturalist 12: 258 (1931)
= Filix-mas thelypteris var. pubescens (G.Lawson) Farw.; Amer. Midl. Naturalist 12: 258 (1931)
= Lastrea thelypteris var. pubescens G.Lawson; Edinburgh New Philos. J., Ser. 2, 19: 277 (1864)
= Nephrodium thelypteris f. pufferae A.A.Eaton; Fern Bull. 10: 78 (1902)
= Nephrodium thelypteris f. suaveolens Clute; Fern Bull. 18: 87 (1910)
= Thelypteris confluens var. pubescens (G.Lawson) J.S.Pringle; Tech. Bull. Roy. Bot. Gards. Hamilton no. 4: 42 (1969)
= Thelypteris palustris f. frondosa (Farw.) Gruber; Amer. Fern J. 30: 91 (1940)
= Thelypteris palustris f. linearis (Farw.) C.F.Reed; Phytologia 17: 301 (1968)
= Thelypteris palustris f. pubescens (G.Lawson) Clute; Our Ferns 152 (1901)
= Thelypteris palustris f. pubescens (G.Lawson) H.Itô; (1939)
= Thelypteris palustris f. pufferae (A.A.Eaton) C.F.Reed; Phytologia 17: 301 (1968), isonym
= Thelypteris palustris f. pufferae (A.A.Eaton) L.B.Sm.; Rhodora 30: 16 (1928)
= Thelypteris palustris f. serratipinnula Gruber; Amer. Fern J. 30: 91 (1940)
= Thelypteris palustris f. suaveolens (Clute) C.F.Reed; Phytologia 17: 301 (1968), isonym
= Thelypteris palustris f. suaveolens (Clute) Fernald; Rhodora 23: 165 (1921)
= Thelypteris palustris var. haleana Fernald; Rhodora 31: 34, fig. 5, 6 (1929)
= Thelypteris palustris var. pubescens (G.Lawson) Fernald; Rhodora 31: 34 (1929)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.04 (April 17th, 2025)

Covering 356,375 accepted species,
8,782 hybrids, 48,090 accepted infraspecific taxa and 1,002,897 synonyms,
in total 1,416,144* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

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With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life