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Ver. 19.4
June 30th, 2024

203.0000 Family Crassulaceae J.St.-Hil.; Expos. Fam. Nat. 2: 123 (1805), nom. cons.
35 genera, ca. 1500 spp.
Species listed in database: 1622; hybrids: 159
= Cotyledonaceae Martinov; Tekhno-Bot. Slovar: 168 (1820)
= Rhodiolaceae Martinov; Tekhno-Bot. Slovar: 546 (1820)
= Sedaceae Roussel; Fl. Calvados, ed. 2: 291 (1806)
= Sempervivaceae Juss.; Gen. Pl.: 207 (1789)
= Tillaeaceae Martinov; Tekhno-Bot. Slovar: 635 (1820)
203.3106 Genus Umbilicus DC.; Bull. Soc. Philom. 49 (1801)
Distribution: Europe, Mediterranean, montane tropical Africa frorn Cameroon to Ethiopia, Somalia and Tanzania, W-Asia to Caucasus (1) and Iran
Species listed in database: 14
= Umbilicaria Pers.; Syn. 1: 510 (1805)
= Chiastophyllum (Ledeb.) Stapf; Index Londin. 2: 176, 316 (1930), A. Berger in Engl. & Prantl, Nat. Pflanzenfam. ed. 2, 18a: 418 (1930)
= Cotyliphyllum Link; Handbuch 2: 23 (1831)
species :
Umbilicus albidoopacus Carlström; Willdenowia 14(1): 22 (1984)
Name: Rhodischer Venusnabel (DE); Rhodian Navelwort (EN)
Distribution: East Aegaean Isl. (Rhodos, Chalki)
34 EAI
species :
Umbilicus botryoides Hochst. ex A.Rich.; Tent. Fl. Abyss. 1: 308 (1848)
Distribution: Egypt (SE-Egypt); Cameroon; E-D.R.Congo [Zaire]; Sudan; Ethiopia; Eritrea; Somalia; Uganda; Kenya; Tanzania; Saudi Arabia (Asir)
20 EGY 23 CMN ZAI 24 ERI ETH SOM SUD 25 KEN TAN UGA 35 SAU
species :
Umbilicus chloranthus Heldr. & Sart. ex Boiss.; Fl. Or. 2: 768 (1872)
Name: Grünes Nabelkraut (DE); Green Venus´ Navel (EN)
Distribution: SE-Italy; Croatia; Albania; Greece (lowlands of C- & S-Greece, Kerkyra, Ionian Isl., Kithira, Sporades, Sifnos, Evvoia, Amorgos); East Aegaean Isl. (Ikaria, Simi, Rhodos); Turkey (W-Anatolia)
13 ALB CRO GRC ITA 34 EAI TUR
Synonyms: hidden
= Cotyledon chlorantha (Heldr. & Sart. ex Boiss.) Halácsy; Consp. Fl. Graec. 1: 578 (1900)
species :
Umbilicus heylandianus Webb & Berthel.; Phyt. Canar. 1: 176 (1840)
Name: Heylands Nabelkraut (DE); Sombrerillo de cumbre (ES)
Distribution: Portugal; N-, W- & C-Spain; Morocco; Canary Isl. (La Palma Isl., Gran Canaria)
12 POR 20 MOR 21 CNY
Synonyms: hidden
= Cotyledon coutinhoi (Mariz) Cout.; Fl. Portug., ed. 2, Palhinha, 336 (1939), in clavi
= Cotyledon praealta (Brot.) Samp.; Bull. Assoc. Pyren. Echange Pl. 19: 4 (1909)
= Cotyledon praealta var. coutinhoi (Mariz) Samp.; (1947)
= Cotyledon strangulata Font Quer; Cavanillesia 1: 71 (1928)
= Cotyledon umbilicus-veneris var. praealta (Brot.) Pérez-Lara; (1971)
= Grammanthes heylandiana (Webb & Berthel.) Webb ex Christ; Bot. Jahrb. Syst. 9: 162 (1887)
= Umbilicus citrinus Wolley-Dod; J. Bot. 52: 12 (1914)
= Umbilicus coutinhoi Mariz; Bol. Soc. Broter. 20: 188 (1905)
= Umbilicus heylandianus var. coutinhoi (Mariz) A.Fern.; Anuar. Soc. Brot. 45: 115 (1979)
= Umbilicus pendulinus var. praealtus Brot.; (1804)
= Umbilicus praealtus (Brot.) Mariz; Bol. Soc. Broter. 20: 185 (1905)
= Umbilicus strangulatus (Font Quer) A.Berger; Engl. & Prantl, Nat. Pflanzenfam. ed. 2, 18a: 417 (1930)
species :
Umbilicus horizontalis (Guss.) DC.; Prodr. 3: 400 (1828)
Name: Waagerechtes Nabelkraut (DE); Horizontal Navelwort (EN)
Distribution: Spain; Gibraltar; Baleares; ?Sardinia; Italy; Sicily; Pantelleria; Isole Pelagie (Lampedusa); San Marino; Croatia; Bosnia & Hercegovina; Montenegro; Albania; Bulgaria; Greece (widespread lowlands, incl. most islands); Crete (widespread); Morocco; Algeria; Tunisia; Libya; Egypt (NW-coastal Egypt); Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl., Porto Santo Isl.); Islas Selvagens; Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); Sudan; N-Somalia; Soqotra; East Aegaean Isl. (widespread, incl. Rhodos, Kastellorhizo Isl.); Turkey (SSW-Anatolia, SW-Anatolia, W-Anatolia); Iran (W-Iran); Cyprus (E-Cyprus, N-Cyprus); Lebanon (C-Lebanon); Saudi Arabia (Asir); United Arab Emirates; Oman
12 BAL GIB SPA 13 ALB BIH BUL CRO GRC ITA KRI MNT PEL PTL SIC SMA 20 ALG EGY LBY MOR TUN 21 AZO CNY MDR SEL 24 SOC SOM SUD 34 CYP EAI IRN LEB TUR 35 OMA SAU UAE
Synonyms: hidden
= Cotyledon gaditana (Boiss.) Pau; Visita Bot. Riff, 2 (Ann. Acad. Polytechn. Porto 6: (1911)
= Cotyledon horizontalis Guss.; Ind. Sem. Hort. Boccad. (1826) 4
= Cotyledon peltata J.C.Wendl.; Bot. Beob.: 49 (1798)
= Cotyledon umbilicus-veneris f. amphitropa (Batt.) Maire; (1977)
= Cotyledon umbilicus-veneris f. parviflora Maire; (1977)
= Cotyledon umbilicus-veneris f. suberecta Maire; (1934)
= Cotyledon umbilicus-veneris subsp. gaditanus (Boiss.) Emb. & Maire; Cat. Pl. Maroc 4: 1019 (1941)
= Cotyledon umbilicus-veneris subsp. horizontalis (Guss.) Batt.; Fl. Algèrie [Batt. & Trabut]: 329 (1889)
= Cotyledon umbilicus-veneris var. amphitropa Batt.; Bull. Soc. Bot. France 28: 227 (1881)
= Cotyledon umbilicus-veneris var. gaditana (Boiss.) Batt.; Fl. Algerie [Batt. & Trabut]: 329 (1889)
= Cotyledon umbilicus-veneris var. horizontalis (Guss.) Menezes
= Cotyledon umbilicus-veneris var. suberecta Maire; Bull. Soc. Hist. Nat. Afrique N. 25: 301 (1934)
= Cotyliphyllum horizontale (Guss.) Link; Handbuch 2: 23 (1831)
= Umbilicus gaditanus Boiss.; Diagn. Pl. Or. Nov. 1(6): 58 (1846)
= Umbilicus gaditanus var. giganteus Batt.; (1885)
= Umbilicus giganteus (Batt.) Batt.; Fl. Algérie [Batt. & Trabut] Tunisie: 133 (1905)
= Umbilicus horizontalis subsp. gaditanus (Boiss.) Losa & Rivas Goday; Arch. Inst. Aclim. Cons. Super. Invest. Ci. 13(2): 232 (1974)
= Umbilicus maroccanus Gand.; Bull. Soc. Bot. France 54: 78 (1907)
= Umbilicus rodriguezii Gand.; Bull. Soc. Bot. France 47: 133 (1900)
= Umbilicus rupestris subsp. gaditanus (Boiss.) O.Bolòs, X.Font & X.Pons; Atlas Corol. Fl. Vasc. Països Catalans 4: no. 583 (1994)
= Umbilicus rupestris subsp. horizontalis (Guss.) O.Bolòs & Vigo; But. Inst. Catalana Hist. Nat., 38 Bot. 1: 66 (1974)
= Umbilicus vulgaris subsp. gaditanus (Boiss.) Batt. & Trab.; (1904)
variant :
var. intermedius (Boiss.) Chamb.; Fl. Turkey & E Aegaean Isl. 4: 213 (1972)
Distribution: NE-Greece; Libya; Egypt (Eastern Desert, NE-Egypt); East Aegaean Isl.; Turkey (S-Anatolia, SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia, SSW-Anatolia); Iraq (NE-Iraq, NW-Iraq); Iran (EC-Iran, N-Iran, S-Iran, W-Iran); Lebanon (Antilebanon, C-Lebanon, coastal W-Lebanon); Syria (coastal W-Syria, C-Syrian Desert, Jazira, NW-Syria, Jbel Druze, W-Syrian Mountains); Israel (coastal W-Israel, Rift Valley, Negev Desert, N-Israel, Judean Desert); Sinai peninsula (C-Sinai, N-Sinai, S-Sinai); Jordania (S-Jordania, W-Jordania); Saudi Arabia (Asir)
13 GRC 20 EGY LBY 34 EAI IRN IRQ ISR JOR LEB SIN SYR TUR 35 SAU
Synonyms: hidden
= Cotyledon intermedia (Boiss.) Bornm.; Bot. Jahrb. Syst. 33(3): 427 (1903)
= Cotyledon umbilicus-veneris subsp. intermedia (Boiss.) Maire & Weiller; (1939)
= Cotyledon umbilicus-veneris var. intermedia (Boiss.) Maire; (1977)
= Umbilicus intermedius Boiss.; Fl. Or. 2: 769 (1872)
= Umbilicus lineolatus Boiss.; Fl. Orient. 2: 769 (1872)
= Umbilicus pendulinus var. intermedius (Boiss.) Post; (1896)
species :
Umbilicus luteus (Huds.) Webb & Berthel.; Phyt. Canar. 1: 177 (1840)
Distribution: S-Italy; Kosovo; North Macedonia; Albania; Bulgaria; Greece (scattered on mainland, Kiklades: Tinos, Amorgos); Crete (Mt. Ida, Mt. Dikti); Morocco; Algeria; East Aegaean Isl. (Lesvos, Chios, Ikaria, Samos); Turkey (E-Anatolia, Inner Anatolia, NW-Anatolia: Bithynia, S-Anatolia, SE-Anatolia, SSW-Anatolia, SW-Anatolia, W-Anatolia); Lebanon (C-Lebanon); Syria (W-Syrian Mountains)
13 ALB BUL GRC ITA KOS KRI NMA 20 ALG MOR 34 EAI LEB SYR TUR
Synonyms: hidden
= Cotyledon altera Thell.; Bull. Herb. Boissier Ser. II. 8: 908 (1908)
= Cotyledon erecta D.Dietr.; Syn. Pl. 2: 1626 (1840)
= Cotyledon font-queri (Maire & Sennen) Sennen & Mauricio; Cat. Fl. Rif Orient.: 44 (1933)
= Cotyledon gaditana subsp. fontqueri (Maire & Sennen) Sennen; Diagn. Nouv. Pl. Espagne & Maroc: 155 ´1936´ (1951)
= Cotyledon lassithiensis (Gand.) Hayek; Repert. Spec. Nov. Regni Veg. Beih. 30: 632 (1925)
= Cotyledon lusitanica Lam.; Encyc. 2: 140 (1786)
= Cotyledon lutea Huds.; Fl. Angl. ed. II. 194 (1762)
= Cotyledon repens (L.) Grande; Bull. Ort. Bot. Napoli 3: 97 (1913)
= Cotyledon umbilicus-veneris f. gigantea Batt.; (1889)
= Cotyledon umbilicus-veneris f. purpurea Maire; (1938)
= Cotyledon umbilicus-veneris Houtt.; Nat. Hist. 2(5): 119 (1775)
= Cotyledon umbilicus-veneris var. font-queri Maire & Sennen; Bull. Soc. Hist. Nat. Afrique N. 24: 213 (1933)
= Cotyledon umbilicus-veneris var. gigantea (Batt.) Maire; Fl. Afrique N. 14: 274 (1977)
= Cotyledon umbilicus-veneris var. pomelii Maire; Bull. Soc. Hist. Nat. Afrique N. 27: 224 (1936)
= Cotyledon umbilicus-veneris var. purpurea (Maire) Maire; Bull. Soc. Hist. Nat. Afrique N. 29: 418 (1936)
= Orostachys umbilicus Schrank & Mart.; Hort. Reg. Monac.: 132 (1829)
= Umbilicus erectus DC.; Lam. & A. P. de Candolle, Fl. Franç., éd. 3 4: 384 (1805)
= Umbilicus erectus var. lassithiensis (Gand.) Stoj.
= Umbilicus lassithiensis Gand.; Fl. Cret. 40, No. 720 (1916), et in Bull. Soc. Bot. France 63: 232 (1919)
species :
Umbilicus mirus (Pamp.) Greuter; Willdenowia 11(2): 277 (1981)
Distribution: Libya (Cyrenaica)
20 LBY
Remarks: possibly to U. parviflorus
Synonyms: hidden
= Sedum mirum Pamp.; Archiv. Bot., Forli 12: (n. s. ii.) 28 (1936)
species :
Umbilicus oppositifolius Ledeb.; Fl. Ross. 2: 176 (1844)
Name: Kaukasus-Walddickblatt (DE)
Distribution: Germany [I]; Northern Caucasus; Georgia [Caucasus]; Armenia
11 ger 33 ARM GEO NCS
Synonyms: hidden
= Chiastophyllum oppositifolium (Ledeb.) A.Berger; Engl. & Prantl, Nat. Pflanzenfam. ed. 2, 18a: 419 (1930)
= Cotyledon oppositifolia Ledeb. ex Nordm.; Bull, Sci. Acad. Petersb. II. 813 (1837)
= Sedum oppositifolium (Ledeb.) Raym.-Hamet; Candollea 4: 43 (1929)
species :
Umbilicus paniculiformis Wickens; Kew Bull. 33(3): 421 (1979)
Distribution: Sudan
24 SUD
Synonyms: hidden
= Cotyledon arabicum Schweinf. ex Täckh.; (1970)
species :
Umbilicus parviflorus (Desf.) DC.; Prodr. 3: 400 (1828)
Name: Kleinblütiges Nabelkraut (DE); Small-flowered Navelwort (EN)
Distribution: Croatia; Greece (Kiklades: Naxos, Amorgos, Milos); Crete (widespread); East Aegaean Isl. (Chios, Ikaria, Samos, Kos, Kasos, Karpathos, Rhodos)
13 CRO GRC KRI 34 EAI
Synonyms: hidden
= Cotyledon parviflora Desf.; Ann. Mus. Hist. Nat. Paris 11: 444 (1808)
= Umbilicus sprunerianus Boiss.; Diagn. Pl. Orient. ser. 1, 6: 57 (1846)
species :
Umbilicus patens Pomel; Nouv. Mat. Fl. Atl. 324 (1876)
Distribution: Morocco; Algeria; Tunisia; Libya
20 ALG LBY MOR TUN
Synonyms: hidden
= Cotyledon horizontalis var. micrantha Pamp.; Nuovo Giorn. Bot. Ital. 24: 137 (1917)
= Cotyledon patens (Pomel) Batt.; Fl. Algérie [Batt. & Trabut], Dicot.: 329 (1889)
= Cotyledon umbilicus-veneris subsp. patens (Pomel) Batt.; Fl. Algèrie [Batt. & Trabut]: 329 (1889)
= Cotyledon umbilicus-veneris var. micrantha (Pamp.) Maire; Fl. Afrique N. 14: 275 (1977)
= Cotyledon umbilicus-veneris var. subhorizontalis Maire & Weiller; Bull. Soc. Hist. Nat. Afrique N. 30: 277 (1939),
= Umbilicus micranthus Pomel; Nouv. Mat. Fl. Atl. 325 (1876)
species :
Umbilicus rupestris (Salisb.) Dandy; Riddelsd., Hedley & Price, Fl. Gloucestershire 611 (1948)
Name: Echter Venusnabel (DE)
Distribution: England; Scotland; Ireland; Germany [I]; Portugal; Spain; Andorra; Gibraltar; Baleares; France; Monaco; Channel Isl.; Corsica; Sardinia; Italy; Sicily; Pantelleria; Isole Pelagie (Lampedusa); San Marino; ?Slovenia; Croatia; Bosnia & Hercegovina; Albania; ?Bulgaria; Greece (widespread on mainland, incl. most islands); Crete (incl. Gavdos); Morocco; Algeria; Tunisia; Libya; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl., Ilhas Desertas, Porto Santo Isl.); East Aegaean Isl. (Lesvos, Chios, Ikaria, Fourni, Samos, Kos, Chios, Rhodos); Turkey (Inner Anatolia, SSW-Anatolia, W-Anatolia, WN-Anatolia); Cyprus (C-Mountains, E-Cyprus, N-Cyprus, W-Cyprus); Lebanon (C-Lebanon); Sinai peninsula (S-Sinai); Saudi Arabia (Hejaz, Midyan); S-India [I] (Karnataka [I], Tamil Nadu [I])
10 GRB IRE 11 ger 12 ADO BAL CIS COR FRA GIB MCO POR SAR SPA 13 ALB BIH CRO GRC ITA KRI PEL PTL SIC SMA 20 ALG LBY MOR TUN 21 AZO MDR 34 CYP EAI LEB SIN TUR 35 SAU 40 ind
Synonyms: hidden
= Cotyledon neglecta Cout.; Fl. Portug., ed. 2, Palhinha, 336 (1939), in adnot.
= Cotyledon ombilicus Lam.; Encycl. 2: 140 (1786)
= Cotyledon pendulina Batt.; Fl. Algérie [Batt. & Trabut], Dicot.: 329 (1889)
= Cotyledon rupestris Salisb.; Prodr. Stirp. Chap. Allerton: 307 (1796)
= Cotyledon tuberosa (L.) Halácsy; Consp. Fl. Graec. 1: 577 (1901)
= Cotyledon umbilicata Lam.; Fl. Franç. 3: 59 (1779)
= Cotyledon umbilicifolia Stokes; Bot. Mat. Med. 2: 545 (1812)
= Cotyledon umbilicus-veneris f. deflexa (Pomel) Batt.; (1889)
= Cotyledon umbilicus-veneris L.; Sp. Pl. 1: 429 (1753)
= Cotyledon umbilicus-veneris subsp. pendulina (DC.) Batt.; Fl. Algerie [Batt. & Trabut]: 329 (1889)
= Cotyledon umbilicus-veneris var. deflexa (Pomel) Maire; Cat. Pl. Maroc 2: 326 (1932)
= Cotyledon umbilicus-veneris var. patula (Pomel) Maire; Fl. Afrique N. 14: 277 (1977)
= Cotyledon umbilicus-veneris var. tuberosa L.; Sp. Pl. 1: 429 (1753)
= Cotyliphyllum erectum Link; Handbuch 2: 23 (1831)
= Cotyliphyllum umbilicus Link; Handbuch 2: 23 (1831)
= Umbilicus aetneus Tornab.; Fl. Sicul.: 249 (1887)
= Umbilicus deflexus Pomel; Nouv. Mat. Fl. Atl. 324 (1876)
= Umbilicus neglectus (Cout.) Rothm. & P.Silva; Agron. Lusit. 2: 88 (1940)
= Umbilicus patulus Pomel; Nouv. Mat. Fl. Atl. 323 (1876)
= Umbilicus pendulinus (DC.) DC.; Pl. Hist. Succ. t. 162 (1829)
= Umbilicus pendulinus var. truncatus Wolley-Dod; (1914)
= Umbilicus pendulinus var. velenovskyi Rohlena ex H.Jacobsen; (1954)
= Umbilicus rupestris var. truncatus (Wolley-Dod) G.D.Rowley; (1956)
= Umbilicus rupestris var. velenovskyi (Rohlena ex H.Jacobsen) G.D.Rowley; (1956)
= Umbilicus simplex K.Koch; Linnaea 19: 41 (1846)
= Umbilicus umbilicatus (Lam.) Breistr.; Bull. Soc. Bot. France, 121 (1-2): 56, 64 (1974)
= Umbilicus vulgaris Batt. & Trab.; Fl. Alger. Tunis, 133 (1905)
species :
Umbilicus schmidtii Bolle; Bonplandia (Hannover) 7: 245 (1859)
Distribution: Cape Verde Isl. (Santo Antao Isl., Ilha de Sao Nicolau, Ilha de Sao Tiago, Fogo Isl.)
21 CVI
Synonyms: hidden
= Umbilicus horizontalis J.A.Schmidt; Beitr. Fl. Cap. Verd. Ins. 258 (1852) (nom. illeg.)
species :
Umbilicus tropaeolifolius Boiss.; Diagn. ser. 1, 3: 14 (1843)
Distribution: NE-Somalia; Turkey (SE-Anatolia, W-Anatolia); Iraq (NE-Iraq, NW-Iraq); Iran (S-Iran, W-Iran); Syria (Jazira)
24 SOM 34 IRN IRQ SYR TUR
Synonyms: hidden
= Cotyledon tropaeolifolia (Boiss.) Bornm.; Beih. Bot. Centralbl. 32: II. 389 (1914)
= Umbilicus oxypetalus Boiss.; Diagn. Ser. I. 3: 15 (1843)
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A complete, synonymic checklist of the Higher Plants of the World


Version 19.4 (June 30th, 2024)

Covering 355,230 accepted species,
8,675 hybrids, 47,796 accepted infraspecific taxa and 997,888 synonyms,
in total 1,409,589* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life