231.0000
Family
Zygophyllaceae R.Br.; Voy. Terra Austral. 2: 545 (1814), nom. cons.
26 genera, ca. 280 spp.
Species listed in database: 290; hybrids: 1
= Agialidaceae Wettst.; Handb. Syst. Bot., ed. 2, 2: 620 (1911)
= Balanitaceae M.Roem.; Fam. Nat. Syn. Monogr. 1: 26 (1846), nom. cons.
= Tribulaceae Trautv.; Estestv. Istor. Gub. Kievsk. Uchebn. Okr. Bot. Sist. 28 (1853)
231.5005
Genus
Zygophyllum L.; Sp. Pl. [Linnaeus] 1: 385 (1753) s. str.
Distribution: E-Mediterranean (1), mostly W-Asia to China, few Africa (?). A few African species still need transfer into Tetraena or Roepera.
Species listed in database: 52
= Agrophyllum Neck.; Elem. Bot. [Necker] 2: 227 (1790)
= Fabago Mill.; Gard. Dict. Abr., ed. 4: s. p. (1754)
= Halimiphyllum (Engl.) Boriss.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18: 144 (1957)
= Sarcozygium Bunge; Linnaea 17: 7, t. 1 (1843)
species :
Zygophyllum atriplicoides
Fisch. & C.A.Mey.; Index Sem. (St. Petersburg [Petropolitanus]) 1: 41 (1835)
subspecies :
subsp. atriplicoides
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Armenia; Iraq (NW-Iraq, W-Iraq: Desert); Iran (EC-Iran, E-Iran, NE-Iran: Mts., S-Iran, W-Iran); Afghanistan (Badakshan, Bamyan, Badghys, Farah, Herat, Samangan, Zabul); Syria (C-Syrian Desert, Jazira, NW-Syria); Pakistan
32 KGZ TKM TZK UZB 33 ARM 34 AFG IRN IRQ SYR 40 PAK
Synonyms: hidden
= Halimiphyllum atriplicoides (Fisch. & C.A.Mey.) Boriss.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18: 150 (1957)
= Halimiphyllum megacarpum (Boriss.) Boriss.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18: 152 (1957)
= Zygophyllum atriplicoides var. megacarpum (Boriss.) Akhiani; Iran. J. Bot. 5(1): 42 (1991)
= Zygophyllum megacarpum Boriss.; Fl. URSS [Komarov et al., eds.] 14: 730 (1949)
subspecies :
subsp. eurypterum
(Boiss. & Buhse) Popov; Byull. Sredne-Aziatsk. Gosud. Univ. 12: 119 (1926)
Distribution: Turkmenistan; Iraq (NE-Iraq); Iran (E-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Balkh, Bamyan, Farah, Ghorat, Helmand, Herat, Parwan, Samangan, Zabul); Lebanon; Pakistan (Baluchistan, Quetta)
32 TKM 34 AFG IRN IRQ LEB 40 PAK
Synonyms: hidden
= Halimiphyllum eurypterum (Boiss. & Buhse) Boriss.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18: 146 (1957)
= Zygophyllum eurypterum Boiss. & Buhse; Nouv. Mém. Soc. Imp. Naturalistes Moscou 12: 49 (1860)
species :
Zygophyllum balchaschense
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 725 (1949)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum betpakdalense
Golosk. & Semiotr.; Bull. Acad. Sci. Kazakhsk. S.S.R., Bot. & Soil Sci. Ser. (7): 74 (1960)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum bogdashanicum
Chang Y.Yang & J.H.Fan; Fl. Xinjiangensis 3: 624 (2011)
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
species :
Zygophyllum borissovae
Beier & Thulin; Pl. Syst. Evol. 240(1-4): 33 (2003), nom. nov.
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Zygophyllum microcarpum Boriss.; Fl. URSS [Komarov et al., eds.] 14: 724 (1949)
species :
Zygophyllum brachypterum
Kar. & Kir.; Bull. Soc. Imp. Naturalistes Moscou 14: 397 (1841)
Distribution: Kazakhstan; C-Iran; China (W-Gansu, Xinjiang); Mongolia
32 KAZ 34 IRN 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum fabago subsp. brachypterum (Kar. & Kir.) Popov; (1925)
species :
Zygophyllum cuspidatum
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 726 (1949)
Distribution: Kazakhstan; China (Xinjiang)
32 KAZ 36 CHI
species :
Zygophyllum darvasicum
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 729 (1949)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Halimiphyllum darvasicum (Boriss.) Boriss.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18: 150 (1957)
species :
Zygophyllum dregeanum
Sond.; Fl. Cap. [Harvey & auct. succ.] 1: 365 (1860)
Distribution: South Africa (N-Cape Prov.)
27 SAF
Remarks: belongs in another genus
Synonyms: hidden
= Zygophyllum densiflorum Schinz; Verh. Bot. Vereins Prov. Brandenburg 31: 181 (1896)
species :
Zygophyllum ebinricum
Chang Y.Yang; Fl. Xinjiangensis 3: 65 (2011)
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
species :
Zygophyllum eichwaldii
C.A.Mey.; Pl. Nov. Casp.-Caucasus [Eichwald] 15 (1831)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
Synonyms: hidden
= Zygophyllum coccineum Lepech.; Tagebuch Reise Russ. 1: 310 (1774)
species :
Zygophyllum fabago
L.; Sp. Pl. [Linnaeus] 1: 385 (1753)
Name: Bohnen-Jochblatt (DE), Syrian beancaper (EN)
Distribution: Germany [I]; Spain; Baleares; France [I]; Sardinia [I]; Romania; E-European Russia; Ukraine; Crimea; Egypt (Eastern Desert); Kazakhstan; Turkmenistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, S-Anatolia, SE-Anatolia); Iraq (NW-Iraq, SE-Iraq); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Badghys, Bamyan, Faryab, Ghazni, Herat, Kabul, Parwan, Qunduz, Samangan); Lebanon (Antilebanon); Syria (C-Syrian Desert, Jazira, NW-Syria, W-Syrian Mountains); Israel (N-Israel); Sinai peninsula (N-Sinai); Jordania (W-Jordania); China (NW-Gansu, W-Nei Mongol, Qinghai, Xinjiang); Pakistan (Baluchistan, Quetta); USA [I] (California [I], Colorado [I], Idaho [I], Kansas [I], Montana [I], New Mexico [I], Nevada [I], New York [I], Pennsylvania [I], Texas [I], Washington State [I])
11 ger 12 BAL fra sar SPA 13 ROM 14 KRY RUS UKR 20 EGY 32 KAZ TKM 33 ARM AZE GEO NCS 34 AFG IRN IRQ ISR JOR LEB SIN SYR TUR 36 CHI 40 PAK 73 usa 74 usa 75 usa 76 usa 77 usa
Synonyms: hidden
= Fabago alata Moench; Methodus [Moench], Suppl. 90 (1802)
= Fabago major Sweet; Brit. Fl. Gard. [Sweet] 6: t. 226 (1834)
= Zygophyllum fabagineum St.-Lag.; Ann. Soc. Bot. Lyon 7: 137 (1880)
= Zygophyllum fabago subsp. dolichocarpum Popov ex Hadidi; Fl. Iranica [Rechinger] 98: 27 (1972)
= Zygophyllum fabago subsp. orientale Boriss. ex Hadidi; Fl. Iranica [Rechinger] 98: 27 (1972)
= Zygophyllum fabago var. sayramense Chang Y.Yang; Fl. Xinjiangensis 3: 623 (2011)
= Zygophyllum micronatum Opiz; Naturalientausch 6: 42 (1824)
= Zygophyllum typicum (Popov) C.Regel; Candollea 16: 232 (1958)
species :
Zygophyllum fabagoides
Popov; Byull. Sredne-Aziatsk. Gosud. Univ. 11: 113 (1925)
Distribution: Kazakhstan; China (Gansu, Xinjiang)
32 KAZ 36 CHI
species :
Zygophyllum fuhaiense
Chang Y.Yang & Y.Feng; Fl. Xinjiangensis 3: 624 (2011)
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
species :
Zygophyllum furcatum
C.A.Mey.; Fl. Altaic. [Ledebour] 2: 106 (1830)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum gobicum
Maxim.; Enum. Pl. Mongolia 125 (1889)
Distribution: Kazakhstan; China (N-Gansu, W-Nei Mongol, Qinghai, E-Xinjiang); Mongolia
32 KAZ 36 CHI 37 MON
species :
Zygophyllum gontscharovii
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 729 (1949)
Distribution: Turkmenistan; Tajikistan; Iran; Afghanistan (Bamyan, Farah, Samangan); Pakistan
32 TKM TZK 34 AFG IRN 40 PAK
Synonyms: hidden
= Halimiphyllum gontscharovii (Boriss.) Boriss.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18: 148 (1957)
= Zygophyllum eurypterum subsp. gontscharovii (Boriss.) Hadidi; Fl. Iranica [Rechinger] 98: 24 (1972)
species :
Zygophyllum hamicum
J.H.Huang ex Chang Y.Yang & J.H.Fan; Fl. Xinjiangensis 3: 622 (2011)
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
species :
Zygophyllum heterocladum
Rech.f. & Patzak; Biol. Skr. 13(4) [Symb. Afghan. 5]: 15 (1963)
Distribution: Afghanistan (Badakshan)
34 AFG
Synonyms: hidden
= Halimiphyllum heterocladum (Rech.f. & Patzak) Adylov; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzbeksk. S.S.R. 20: 6 (1982)
species :
Zygophyllum iliense
Popov; Byull. Sredne-Aziatsk. Gosud. Univ. 12: 112 (1926)
Distribution: Kazakhstan; China (Gansu, W-Nei Mongol, Xinjiang)
32 KAZ 36 CHI
species :
Zygophyllum jaxarticum
Popov; Fl. Uzbekist. 4: 58 (1959)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; China (Xinjiang)
32 KAZ TKM TZK UZB 36 CHI
Synonyms: hidden
= Zygophyllum dielsianum (Popov) Popov; Trudy Sredne-Aziatsk. Gosud Univ., Ser. 8b, Bot. 3: 75 (1928) non Schltr.
= Zygophyllum miniatum subsp. dielsianum Popov
species :
Zygophyllum junggaricum
Chang Y.Yang & J.H.Fan; Fl. Xinjiangensis 3: 623 (2011)
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
species :
Zygophyllum kansuense
Y.X.Liou; Acta Phytotax. Sin. 18(4): 484 (1980)
Distribution: China (Gansu)
36 CHI
species :
Zygophyllum karatavicum
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 726 (1949)
Distribution: Kazakhstan (Karatau)
32 KAZ
species :
Zygophyllum kaschgaricum
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 728 (1949)
Distribution: ?Kazakhstan; China (Xinjiang); Mongolia
36 CHI 37 MON
Synonyms: hidden
= Sarcozygium kaschgaricum (Boriss.) Y.X.Liou; Fl. Reipubl. Popularis Sin. 43(1): 142 (1998)
species :
Zygophyllum kegense
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 727 (1949)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Zygophyllum kopalense
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 726 (1949)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum lehmannianum
Bunge; Arbeiten Naturf. Vereins Riga 1(2): 202 (1847)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan
32 KAZ TKM TZK UZB
Synonyms: hidden
= Zygophyllum karelini Fisch. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou 11: 149 (1839), nomen
species :
Zygophyllum loczyi
Kanitz; Növenyt. Gyujtesek Eredm. Grof Szechenyi Bela Keletazsiai Utjabol 13 (1891)
Distribution: ?Kazakhstan; China (Gansu, W-Nei Mongol, Qinghai, Xinjiang)
36 CHI
species :
Zygophyllum macropodum
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 161, 724 (1949)
Distribution: Kazakhstan; China (Xinjiang)
32 KAZ 36 CHI
species :
Zygophyllum melongena
Bunge; Fl. Altaic. [Ledebour] 2: 104 (1830)
Distribution: Siberia (Altai); Mongolia
30 SIB 37 MON
species :
Zygophyllum miniatum
Cham. & Schltdl.; Linnaea 5: 49 (1830)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Afghanistan (Badghys, Bamyan, Kabul)
32 KAZ KGZ TKM TZK UZB 34 AFG
variant :
var. englerianum
Popov; (1925)
Distribution: Turkmenistan; Uzbekistan (Samarkand, Aralo-Caspia); Afghanistan
32 TKM UZB 34 AFG
Synonyms: hidden
= Zygophyllum englerianum (Popov) Popov; Fl. Uzbekist. 4: 57 (1959)
species :
Zygophyllum mucronatum
Maxim.; Bull. Acad. Imp. Sci. St.-Pétersbourg 27: 438 (1881)
Distribution: China (Gansu, W-Nei Mongol, Ningxia, Qinghai, Xinjiang); Mongolia
36 CHI 37 MON
species :
Zygophyllum neglectum
Grubov; Opred. Sosud. Rast. Mongolii (Key Vasc. Pl. Mongolia): 176 (1982)
Distribution: Mongolia
37 MON
species :
Zygophyllum obliquum
Popov; Byull. Sredne-Aziatsk. Gosud. Univ. 11: 113 (1925)
Distribution: Kazakhstan; Kyrgyzstan; Tajikistan; Afghanistan (Wakhan); China (Gansu, Xinjiang)
32 KAZ KGZ TZK 34 AFG 36 CHI
species :
Zygophyllum ovigerum
Fisch. & C.A.Mey. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou 11: 142 (1839), nomen, et Fl. Orient. [Boissier] 1: 914 (1867)
Distribution: SE-European Russia; W-Kazakhstan; Turkmenistan; Uzbekistan
14 RUS 32 KAZ TKM UZB
species :
Zygophyllum oxianum
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 723 (1949)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; Iran (NE-Iran: Mts., N-Iran); Afghanistan (Faryab, Samangan)
32 KAZ TKM TZK UZB 34 AFG IRN
Synonyms: hidden
= Zygophyllum fabago var. oxianum (Boriss.) Kitam.
species :
Zygophyllum oxycarpum
Popov; Byull. Sredne-Aziatsk. Gosud. Univ. 12: 112 (1926), et in Bull. Univ. As. Centr. Suppl.,12
Distribution: Kazakhstan; China (Xinjiang)
32 KAZ 36 CHI
Synonyms: hidden
= Zygophyllum budunense Semiotr.; Bot. Mater. Gerb. Bot. Inst. Bot. Akad. Nauk Kazakhsk. S.S.R. 3: 19 (1965)
species :
Zygophyllum pamiricum
Grubov; Novosti Sist. Vyssh. Rast. 31: 175 (1998)
Distribution: China (Xinjiang)
36 CHI
species :
Zygophyllum pinnatum
Cham. & Schltdl.; Linnaea 5: 48 (1830)
Distribution: C-European Russia; E-European Russia (incl. S-Urals); Siberia (Altai, Krasnoyarsk); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Iran (N-Iran); China (Xinjiang); Mongolia
14 RUS 30 SIB 32 KAZ KGZ TKM TZK UZB 34 IRN 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum chakassicum (Peschkova) A.L.Ebel; Botanica Pacifica 12(2): 129 (2023)
= Zygophyllum macropterum C.A.Mey.; Fl. Altaic. [Ledebour] 2: 102 (1830)
= Zygophyllum pinnatum subsp. chakassicum Peschkova; Fl. Sibir. 10: 32 (1996)
= Zygophyllum sieversianum Stephan ex Ledeb.; Fl. Ross. [Ledebour] 1(3): 484 (1843)
species :
Zygophyllum potaninii
Maxim.; Bull. Acad. Imp. Sci. St.-Pétersbourg 27: 488 (1881)
Distribution: Kazakhstan; China (Gansu, W-Nei Mongol, Xinjiang); Mongolia
32 KAZ 36 CHI 37 MON
species :
Zygophyllum pterocarpum
Bunge; Fl. Altaic. [Ledebour] 2: 103 (1830)
Distribution: Siberia (Altai, Tuva); Kazakhstan; China (Gansu, W-Nei Mongol, Xinjiang); Mongolia
30 SIB 32 KAZ 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum altaicum Stephan ex Ledeb.; Fl. Ross. [Ledebour] 1: 484 (1843)
= Zygophyllum pterocarpum subsp. tuvinicum Peschkova; Fl. Sibir. 10: 33 (1996)
= Zygophyllum pterocarpum var. microcarpum Y.X.Liou; Acta Phytotax. Sin. 18(4): 485 (1980)
= Zygophyllum tuvinicum (Peschkova) A.L.Ebel; Botanica Pacifica 12(2): 129 (2023)
species :
Zygophyllum rosowii
Bunge; Linnaea 17: 5 (1843)
Distribution: Kazakhstan; Kyrgyzstan; Tajikistan; China (Gansu, Xinjiang); Mongolia
32 KAZ KGZ TZK 36 CHI 37 MON
variant :
var. latifolium
(Schrenk) Popov; Byull. Sredne-Aziatsk. Gosud. Univ. 11: 118 (1925)
Distribution: Kazakhstan; China (Gansu, Nei Mongol, Xinjiang); Mongolia
32 KAZ 36 CHI 37 MON
Synonyms: hidden
= Zygophyllum latifolium Schrenk; Bull. Cl. Phys.-Math. Acad. Imp. Sci. St.-Pétersbourg 2: 198 (1844)
species :
Zygophyllum sinkiangense
Y.X.Liou; Acta Phytotax. Sin. 18: 484 (1980)
Distribution: China (Xinjiang)
36 CHI
species :
Zygophyllum stenopterum
Schrenk; Bull. Cl. Phys.-Math. Acad. Imp. Sci. St.-Pétersbourg 3: 308 (1845)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum subtrijugum
C.A.Mey.; Fl. Altaic. [Ledebour] 2: 105 (1830)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum taldykurganicum
Boriss.; Fl. URSS [Komarov et al., eds.] 14: 727 (1949)
Distribution: Kazakhstan
32 KAZ
species :
Zygophyllum trialatum
Blatt. & Hallb.; J. Indian Bot. 1: 90 (1919)
Distribution: Pakistan (Baluchistan)
40 PAK
Remarks: provisionally accepted name
species :
Zygophyllum turcomanicum
Fisch. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou 11: 149 (1839), nomen, et Fl. Orient. [Boissier] 1: 914 (1867)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan
32 KAZ TKM UZB
species :
Zygophyllum uqturpanicum
Chang Y.Yang & Z.J.Li; Fl. Xinjiangensis 3: 622 (2011)
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
species :
Zygophyllum xanthoxylum
(Bunge) Maxim.; Fl. Tangut. 103 (1889)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang, Qinghai, Nei Mongol, NC-China); Mongolia
32 KGZ TZK UZB 36 CHI 37 MON
Synonyms: hidden
= Sarcozygium xanthoxylum Bunge; Linnaea 17: 8 (1843)
= Zygophyllum xanthoxylum (Bunge) Engl.; Nat. Pflanzenfam. [Engler & Prantl] III, 4: (1890) 81, isonym
= Zygophyllum zanthoxylum ´Engl.´ ex Dippel; Handb. Laubholzk. 2: 358 (1893)
= Zygophyllum ferganense (Drobow) Boriss.; Fl. URSS [Komarov et al., eds.] 14: 184 (1949)
= Zygophyllum xanthoxylon var. ferganense Drobow
species :
Zygophyllum yiwuense
Chang Y.Yang & J.H.Fan; Fl. Xinjiangensis 3: 625 (2011)
Distribution: China (Xinjiang)
36 CHI
Remarks: provisionally accepted name
