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Ver. 14.9
February 4th, 2023

034.0000 Family Dryopteridaceae Herter; Rev. Südam. Bot. 9: 15 (1949), nom. cons.
26 genera, ca. 2100 spp.
Species listed in database: 2162; hybrids: 182
= Aspidiaceae Burnett; Outl. Bot.: 324, 1156 (1835), nom. illeg.
= Aspidiaceae Mett. ex A.B.Frank; Leunis, Syn. Pflanzenk. ed. 2. 3: 1469 (1877), nom. illeg.
= Bolbitidaceae (Pic.Serm.) Ching; Acta Phytotax. Sin. 16: 15 (1978)
= Elaphoglossaceae Pic.Serm.; Webbia 23: 209 (1968)
= Filicaceae Juss.; Gen. Pl.: 14. (1789), as ´Filices´, nom. illeg., rej.
= Hypodematiaceae Ching; Acta Phytotax. Sin. 13: 96 (1975)
= Peranemataceae (C.Presl) Ching; Sunyatsenia 5: 246 (1940), nom. rej.
= Peranematoideae (C.Presl) Fraser-Jenk.; subfamily
034.3006 Genus Dryopteris Adans.; Fam. Pl. 2: 20 (1763), nom. cons., sensu lato
Distribution: Northern temperate zone (over 150 spp. in China and Taiwan), tropical mountains, only 22 New World. Inclusion of all the satellite genera Acrorumohra, Adenoderris, Peranema, Diacalpe, Dryopsis, Nothoperanema etc. fide Zhang (2012) and others, based on molecular results. Fraser-Jenkins et al. (2018) argue for keeping some genera distinct, like Dryopsis, Stenolepia and Nothoperanema.
Species listed in database: 346; hybrids: 81
= Acrophorus C.Presl; Tent. Pterid. 93 (1836) (fide Zhang (2012))
= Acrorumohra (H.Itô) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 101 (1939) (fide Zhang (2012))
= Adenoderris J.Sm.; Hist. Fil. 222 (1875)
= Anopodium J.Sm.; Cat. cult. ferns 16 (1857)
= Arsenopteris Webb & Berthel.; Hist. nat. Canar. 82, pars 3: 437 (1847)
= Arthrobotrys Wall.; Num. List no. 395 (1828)
= Catapodium J.Sm.; Cat. cult. ferns 16 (1857)
= Ctenitis subgen.Dryopsis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 280 (1938)
= Desmopodium J.Sm. ex T.Moore; Ind. 308 (1861)
= Diacalpe Blume; Enum. Pl. Jav. 241 (1828) (fide Zhang (2012))
= Dichasium (A.Braun) Fée; Mém. Foug. 5 (Gen. Fil.):302 (1852)
= Diclisodon T.Moore; Index Fil. (T. Moore) 95 (1857)
= Dryopsis Holttum & Edwards; Kew Bull. 41: 179 (1986) (fide Zhang (2012))
= Dryopteris sect.Dryopsis (Holttum & P.J.Edwards) Li Bing Zhang; Phytotaxa 71: 18 (2012)
= Dryopteris subgen.Lophodium (Newman) Tzvelev; Novosti Sist. Vyssh. Rast. 35: 10 (2003)
= Filix Hill; Family Herbal 171 (1755)
= Filix-Mas Hill; Brit. Herb. 527 (1756)
= Gleichenia Neck.; Element. Bot. 3: 314 (1790)
= Gymnothalamium Zenker ms. ex Kunze; Linn. 24: 287 (1851)
= Lophodium Newman; Phytol. 4: 371, app. XVI (1851)
= Megalastrum J.Sm.; Hist. Fil. 216 (1875)
= Microdictyon Fée; Mém. Foug. 11 (Hist. Foug. Ant.): 66 (1866)
= Monogonia C.Presl; Tent. Pterid. 146 (1836)
= Monophlebia T.Moore; Index Fil. (T. Moore) 89 (1857)
= Nematopera Kunze; Bot. Zeit. (Berlin) 797 (1846) (corr. nom.)
= Nephrodium Rich.; Marthe, Cat. Pl. Jard. Méd. Paris 120 (1801)
= Nothoperanema (Tagawa) Ching; Acta Phytotax. Sin. 11: 25 (1966) (fide Zhang (2012))
= Oligocampia Trevis.; Atti Ist. Veneto II, 2: 165 (1851)
= Pachyderris J.Sm.ms. ex T.Moore; Index Fil. (T. Moore) 146 (1857)
= Peranema D.Don; Prodr. Fl. Nepal. 12 (1825) (fide Zhang (2012))
= Podeilema R.Br.ms. ex Wall.; Ic. pl. as. rar. 1. 42 (1830)
= Psidopodium Neck.; Element. Bot. 3: 315 (1790)
= Pteris Gled.; Syst. Pl. 289 (1764)
= Pycnopteris T.Moore; Gard. Chr. 468 (1855)
= Revwattsia D.L.Jones; Fl. Australia 48: 711 (1998)
= Rumohra sect.Acrorumohra H.Itô; J. Jap. Bot. 11: 583 (1935)
= Sphaeropteris R.Br. ex Wall.; Pl. asiat. rar. 1: 41 (1830) (nom. illeg.)
= Syneuron J.Sm.ms. ex Hook.; Ic. Pl. t.950 (1854)
= Stenolepia Alderw.; Bull. Dépt. agric. Ind. néerl. 27: 45 (1909)
= Dryopteris sect.Stenolepia (Alderw.) L.Y.Kuo & Y.H.Chang; Syst. Bot. 41(3): 603 (2016)
species :
Dryopteris acutodentata Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 432 (1938)
Distribution: China (W-Sichuan, NW-Yunnan); Tibet; Taiwan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal); Bhutan; Nepal; Myanmar (Kachin, etc.)
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA
Synonyms: hidden
= Dryopteris filix-mas subsp. kingii (Hope) C.Chr.; Ind. Fil. 5: 265 (1905) (non Dryopteris kingii (Bedd.) C. Chr.)
= Dryopteris silaensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 65 (1941)
= Nephrodium kingii Hope; J. Bombay Nat. Hist. Soc. 12: 621, t. 9 (1899) (non Dryopteris kingii (Bedd.) C. Chr.)
species :
Dryopteris adnata (Blume) Alderw.; Mal. Ferns 191 (1909) & Suppl. 501 (1917)
Distribution: Java; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Sumatra; Lesser Sunda Isl. (Timor)
42 BOR JAW LSI SUL SUM
Remarks: Subgen. Dryopsis
Synonyms: hidden
= Aspidium adnatum Blume; Enum. Fil. 162 (1828)
= Ctenitis adnata (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 289 (1938)
= Dryopsis adnata (Blume) Holttum & P.J.Edwards; Kew Bull. 41(1): 190 (1986)
= Polystichum truncatulum Alderw.; Bull. Jard. Bot. Buitenz. II, 16: 60, t. 8 (1914)
species :
Dryopteris aemula (Aiton) Kuntze; Rev. Gen. Pl. 1: 812 (1891)
subspecies :
subsp. aemula
Name: Heuduftender Wurmfarn (DE); Ninfa de los bosques (ES)
Distribution: England; Wales; Scotland; France; Ireland; Spain; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl.); Canary Isl. (La Gomera)
10 GRB IRE 12 FRA SPA 21 AZO CNY MDR
Remarks: ser. Aemulae
Synonyms: hidden
= Allantodia aemula (Aiton) Desv.; Mem. Soc. Linn. Paris 6, 2: 265 (1827)
= Aspidium aemulum (Aiton) Sw.; Schrad., J. Bot. 1800(2): 42 (1801)
= Aspidium recurvum (Newman) Bree; Phytol. 1: 773 (1843)
= Lastrea aemula (Aiton) Brack.; U. S. Expl. Exp. 16: 200 (1854)
= Lastrea foenisecii Lowe ex Watson; Phytol. 2: 568 (1846)
= Lastrea recurva Newman; Natur. Alm. 23 (1844)
= Lophodium foenisecii Newman; Phyt. 4: app. 16 (1851)
= Lophodium recurvum (Newman) Newman; Phyt. 4: 371 (1851)
= Nephrodium aemulum (Aiton) Baker; Syn. 279 (1867)
= Nephrodium foenisecii Lowe; Trand. Cambridge Philos. Soc. 4: 7 (1830)
= Polypodium aemulum Aiton; Hort. Kew 3: 466 (1789) (basion.)
= Polystichum aemulum (Aiton) Corb.; Nouv. Fl. Normandie: 676 (1894)
= Polystichum aemulum (Aiton) C.Presl; Tent. Pterid. 83 (1836)
= Polystichum foenisecii (Lowe) P.Fourn.; Quatre Fl. France 17 (1934)
= Polystichum recurvum (Newman) Gand.; Fl. Eur. 27: 174 (1891)
subspecies :
subsp. liliana (Golitsin) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)
Distribution: Georgia [Caucasus] (Adzharia); Turkey (Anatolia)
33 GEO 34 TUR
Synonyms: hidden
= Dryopteris liliana Golitsin; Fedde, Repert. 31: 388 (1933)
species :
Dryopteris affinis (Lowe) Fraser-Jenk.; Fern Gaz. 12: 56 (1979)
subspecies :
subsp. affinis
Name: Spreuschuppiger Wurmfarn (DE); Dentabron (ES)
Distribution: Madeira (Madeira Isl.); Azores (Sao Miguel Isl., Terceira, Sao Jorge, Pico, Faial, Flores Isl.); Canary Isl. (La Gomera, Tenerife); Portugal; Spain; Andorra; France; Corsica; Belgium; Luxembourg; Netherlands; England; Channel Isl.; Scotland; Wales; Norway; Germany (Baden-Württemberg, Bayern, Niedersachsen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt); Switzerland; Liechtenstein; Italy (incl. Elba Isl.); Hungary; Sicily; Sardinia; Slovakia; Slovenia; Austria; Romania; Turkey (Anatolia); European Turkey; Greece; Northern Caucasus; ?Armenia; Georgia [Caucasus]; Azerbaijan; New Zealand [I]
10 GRB NOR 11 AUT BGM GER HUN LIE LUX NET SLO SWI 12 ADO CIS COR FRA POR SAR SPA 13 GRC ITA ROM SIC SLV TUE 21 AZO CNY MDR 33 AZE GEO NCS 34 TUR 51 nzl
Remarks: affinis agg.
Synonyms: hidden
= Aspidium affine Lowe; J. Bot. 1: 25 (1834) (nomen)
= Aspidium filix-mas var. subintegra Döll; (1857) non Boreau (1840)
= Dryopteris affinis f. valongensis Res.-Pinto ex Fraser-Jenk.; Anal. Jard. Bot. Madrid 41(1): 209 (1984), nom. nov. for D. resendeana M. Pinto 1969
= Dryopteris affinis subsp. cluthensis A.R.Church; Fern Gaz. 21(2): 89 (2019) [fide Fraser-Jenkins et al. 2020]
= Dryopteris affinis var. azorica Fraser-Jenk.; Willdenowia 10(1): 110 (1980)
= Dryopteris affinis var. subintegra (Döll) Askerov; Zametki Sist. Geogr. Rast. 39: 7 (1983)
= Dryopteris borreri subsp. resendeana (M.Pinto) Malag.
= Dryopteris borreri var. atlantica Tavel; 153 (1937), nom. inval.
= Dryopteris borreri var. subintegra (Döll ex Asch. & Graebn.) Tavel; (1937)
= Dryopteris pseudomas (Woll.) Holub & Pouzar; Holub, Folia Geobot. Phytotax. 2: 332 (1967)
= Dryopteris resendeana M.Pinto; Portug. Acta Biol. 9: 320, f. 1-2 (1969)
= Lastrea filix-mas var. paleacea (T.Moore) Syme & N.E.Br.; 59 (1886)
= Lastrea pseudomas Woll.; Phytol. n. s. 1: 172 (1855)
= Nephrodium affine Lowe; Tr. Cambridge Philos. Soc. 6. 525 (1838)
= Nephrodium filix-mas var. affine (Lowe) Hook.f.; 465 (1870)
variant :
var. disjuncta (Fomin) Fraser-Jenk.; 110 (1980)
Distribution: Germany; etc.
11 GER
Remarks: the predominant form in Central Europe
Synonyms: hidden
= Dryopteris borreri var. disjuncta (Fomin) Tavel; 52 (1937)
= Dryopteris mediterranea f. disjuncta Fomin
subspecies :
subsp. atropes Fraser-Jenk.; Indian Fern J. 39(1): 127 (2022)
Distribution: Ireland
10 IRE
subspecies :
subsp. jessenii (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)
Distribution: SC-Romania; Serbia
13 ROM SRB
Synonyms: hidden
= Dryopteris affinis var. jessenii Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)
subspecies :
subsp. kerryensis (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)
Distribution: Ireland
10 IRE
Synonyms: hidden
= Dryopteris affinis var. kerryensis Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)
= Dryopteris kerryensis (Fraser-Jenk.) P.D.Sell; Fl. Great Britain and Ireland 1: 671 (2018)
subspecies :
subsp. paleaceolobata (T.Moore) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)
Distribution: England; Scotland; Wales; Ireland; ?Germany
10 GRB IRE
Synonyms: hidden
= Dryopteris affinis var. paleaceolobata (T.Moore) Fraser-Jenk.; Indian J. Forestry 43(1): 83 (2020), et Indian J. Forest. 43(1): 83 (2021)
= Dryopteris paleaceolobata (T.Moore) P.D.Sell; Fl. Great Britain and Ireland 1: 671 (2018)
= Lastrea filix-mas var. paleaceolobata T.Moore; Handb. Brit. Ferns, ed. 3: 112 (1857)
subspecies :
subsp. punctata (Oberh. & Tavel) Fraser-Jenk.; Fern Gaz. 18(1): 5 (2007)
Name: Punktierter Spreuschuppiger Wurmfarn (DE)
Distribution: N-Italy; Switzerland; S-Germany (Bayern); France; Austria; Slovenia; Croatia; Turkey (Anatolia)
11 AUT GER SWI 12 FRA 13 CRO ITA SLV 34 TUR
Synonyms: hidden
= Dryopteris affinis var. punctata Oberh. & Tavel ex Fraser-Jenk.; Willdenowia 10(1): 110 (1980)
= Dryopteris affinis var. splendens (Ehrler ex Bech.) Fraser-Jenk.; Willdenowia 10(1): 111 (1980)
= Dryopteris borreri subvar. splendens Ehrler ex Bech.
species :
Dryopteris aitoniana Pic.Serm.; Webbia 8: 154, f. 3 (1951), nom. nov.
Distribution: Madeira (Madeira Isl.)
21 MDR
Synonyms: hidden
= Aspidium elongatum Sw.; Schrad., J. Bot. 1800(2): 38 (1801) (nec Willd.)
= Dryopteris elongata (Aiton) A.Chev.; Rev. Bot. Appl. Agric. Trop. 15: 1060 (1935) (non (Wall. ex Hook.) Kuntze)
= Dryopteris macaronesica Romariz; Rev. Fac. Cienc. Lisboa ser. 2, C. 3: 92, t. 5(1) (1953) (non Dryopteris elongata (Wall. ex Hook.) Kuntze)
= Lastrea elongata (Aiton) C.Presl; Tent. Pterid. 76 (1836)
= Lastrea grandis T.Moore; Ind. Fil. 93 (1858)
= Nephrodium boryanum Hook.; Sp. 4: 126 (1862) (non Aspidium boryanum Willd. (1810))
= Nephrodium elongatum (Aiton) Hook. & Grev.; Icon. Fil. t. 234 (1831)
= Polypodium elongatum Aiton; Hort. Kew 3: 465 (1789) (non Dryopteris elongata (Wall. ex Hook.) Kuntze)
= Tectaria elongata (Aiton) Cav.; Anal. Ci. Nat. 4: 101 (1801)
species :
Dryopteris alpestris Tagawa; Acta Phytotax. Geobot. 3: 88 (1934)
Distribution: Taiwan; Tibet; China (Yunnan); NE-India (Arunachal Pradesh, Sikkim); Nepal; Bhutan; Myanmar (Kachin, etc.)
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris alpestris Tagawa ex Ching & S.K.Wu; Fl. Xizangica 1: 259 (1983)
= Dryopteris doshunglaensis Ching & S.K.Wu; Enum. vasc. Pl. Tibet (Tibet) 26 (1981), nom. nud.
species :
Dryopteris alpina Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 173 (1913)
Distribution: Sulawesi; Borneo (Sabah: Mt. Kinabalu); Java; New Guinea (Papua New Guinea, Irian Jaya); Philippines (Mindanao); Bismarck Arch. (New Ireland)
42 BOR JAW NWG PHI SUL 43 BIS
Remarks: Subgen. Nothoperanema, sect. Stenolepia
Synonyms: hidden
= Allantodia scabra Kunze; Bot. Zeit. (Berlin) 192: 192 (1848)
= Alsophila tristis (Blume) Blume; T. Moore, Ind. 58 (1857)
= Aspidium triste Blume; Enum. Fil. 169 (1828)
= Athyrium atropurpureum Copel.; Phil. J. Sci. 12C: 59 (1917)
= Athyrium scabrum C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 67 (1851)
= Ctenitis alpina (Rosenst.) Copel.; Gen. Fil. 124 (1947)
= Cyathea tristis (Blume) Domin; Pterid. 263 (1929)
= Cystopteris tristis (Blume) Mett.; Ann. Mus. Bot. Lugd.-Bat. 1: 241 (1864)
= Davallia tristis (Blume) Racib.; Pter. Bull. 131 (1898)
= Dryopteris robur Christenh.; The Global Flora, Special edition, part 1(4): 40 (2018), nom. nov. for Aspidium triste Blume
= Lastrea tristis (Blume) T.Moore; Ind. 107 (1858)
= Stenolepia tristis (Blume) Alderw.; Bull. Dépt. agric. Ind. néerl. 27: 46 (1909)
species :
Dryopteris ambigens (Nakai) Koidz.; Fl. Symb. Orient. As. 39 (1930)
Distribution: Japan
38 JAP
Synonyms: hidden
= Dryopteris erythrosora var. ambigens Nakai; Bot. Mag. (Tokyo) 39: 119 (1925)
species :
Dryopteris amblyodonta J.P.Roux; Kew Bull. 59(2): 207 (2004)
Distribution: Sierra Leone
22 SIE
species :
Dryopteris amurensis (Milde) Christ; Bull. Ac. Géogr. Bot. Mans 35 (1909), et Mém. 20: 164 (1909)
Distribution: Russian Far East (Amur, Khabarovsk, Primorye, Sakhalin); Japan; China (Heilongjiang, Jilin, Liaoning, Nei Mongol); North Korea; ?South Korea
31 RFE 36 CHI 38 JAP NKO
Synonyms: hidden
= Aspidium spinulosum var. amurense Milde; Fil. Eur. 133 (1867)
= Dryopteris amurensis (Milde) Takeda; Bot. Mag. (Tokyo) 24: 113 (1910)
= Leptorumohra amurensis (Milde) Tzvelev; Novosti Sist. Vyssh. Rast. 26: 6 (1989)
= Nephrodium amurense (Milde) B.Fedtsch.; Acta Hort. Petrop. 81: 99 (1912)
species :
Dryopteris anadroma Mitsuta; nom. nud.
Distribution: Japan (Yakushima Isl.)
38 JAP NNS
Conservation Status: VU in Japan
species :
Dryopteris aneitensis (Hook.) C.Chr.; Ind. 252 (1905)
Distribution: Vanuatu
60 VAN
Synonyms: hidden
= Phegopteris aneitensis (Hook.) Kuhn; Verh. Zool.-Bot. Ges. Wien 19: 577 (1869)
= Polypodium aneitense Hook.; Sp. 4: 268 (1862)
= Dryopteris layardii (Baker) C.Chr.; Ind. 274 (1905)
= Nephrodium layardii Baker; Ann. Bot. 6: 323 (1891)
species :
Dryopteris angustifrons (Hook.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: China (Yunnan); Myanmar
36 CHI 41 MYA
Remarks: not in India
Synonyms: hidden
= Lastrea angustifrons (Hook.) Bedd.; Handb. Ferns Brit. India 2: t. 226 (1867), non (Mett. ex Kunze) T. Moore (1858)
= Nephrodium angustifrons (Hook.) Baker; Syn. 283 (1867)
= Nephrodium splendens var. angustifrons Hook.; Sp. Fil. 4: 126 (1862)
species :
Dryopteris angustipalea Darnaedi; M. Kato & K. Iwats., J. Jap. Bot. 64(10): 308 (1989)
Distribution: Moluccas
42 MOL
species :
Dryopteris annamensis (Tagawa) Li Bing Zhang; Taxon 61(6): 1210 (2012), nom. nov.
Distribution: China (Hainan, Yunnan); Tibet; Vietnam
36 CHI TIB 41 VIE
Remarks: subgen. Nothoperanema. Probably to be included in D. aspidioides fide Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Diacalpe annamensis Tagawa; Acta Phytotax. Geobot. 14: 46 (1950)
= Peranema annamensis (Tagawa) Fraser-Jenk.; Panjab Univ. Res. J., Sci. 59: 18 (2010)
species :
Dryopteris antarctica (Baker) C.Chr.; Ind. Fil., Suppl. 29 (1913), et Suppl. III: 81 (1934)
Distribution: D.R.Congo [Zaire]; E-Uganda; Rwanda; Kenya; Tanzania; S-Malawi; E-Zimbabwe; South Africa (W-Cape Prov., E-Cape Prov.); Madagascar; Ile St. Paul; La Réunion
23 RWA ZAI 25 KEN TAN UGA 26 MLW ZIM 27 SAF 29 MDG REU 91 ASP
Synonyms: hidden
= Aspidium antarcticum (Baker) E.Fourn.; Compt. rend. 81: 1140 (1875)
= Dryopteris callolepis C.Chr.; Notizbl. bot. Gart. Berlin-Dahlem 9: 177 (1924)
= Nephrodium antarcticum Baker; J. Linn. Soc. 14: 479 (1875)
species :
Dryopteris anthracinisquama Miyam.; J. Jap. Bot. 59(8): 238 (1984)
Distribution: Japan (Kyushu)
38 JAP
Conservation Status: CR in Japan
species :
Dryopteris apiciflora (Wall. ex Mett.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: Nepal; Bhutan; India (Arunachal Pradesh, Manipur, Nagaland, Sikkim, Uttarakhand, West Bengal); Myanmar (Kachin, etc.); China (Yunnan); Tibet; Taiwan
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA
Remarks: Subgen. Dryopsis
Synonyms: hidden
= Aspidium apiciflorum Wall.; Num. List no. 345 (1828) (nomen), ex Mett., Pheg. u. Asp. 54, n. 128 (1858)
= Ctenitis apiciflora (Wall. ex Mett.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 284 (1938)
= Dryopsis apiciflora (Wall. ex Mett.) Holttum & P.J.Edwards; Kew Bull. 41(1): 189 (1986)
= Lastrea apiciflora (Wall. ex Mett.) Bedd.; Ferns Brit. India t. 40 (1865), isonym
= Lastrea apiciflora C.Presl; Tent. Pterid. 76 (1836), nom. nud.
= Lastrea filix-mas subvar. apiciflora (Wall. ex Mett.) Bedd.; Ferns brit. India, 249 (1883)
= Nephrodium apiciflorum (Wall. ex Mett.) Hook.; Sp. Fil. 4: 112-113, t. 248 (1862)
species :
Dryopteris approximata Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 5: 11, f. 1 (1973)
Distribution: S-India (Kerala, Tamil Nadu); Sri Lanka
40 IND SRL
species :
Dryopteris aquilinoides (Desv.) C.Chr.; Ind. 252 (1905)
Distribution: La Réunion; Mauritius
29 MAU REU
Synonyms: hidden
= Aspidium aquilinoides (Desv.) Mett. ex Kuhn; Fil. Afr. 126 (1868)
= Aspidium boryanum Bojer; Hort. Maur. 391 (1837), Kuhn, Fil. Afr. 127 (pt.) (1868)
= Nephrodium aquilinoides Desv.; Mem. Soc. Linn. Paris 6, 2: 261 (1827)
= Aspidium grande Fée; Gen. 295 (1850-52)
= Nephrodium aquilinoides var. angustatum Desv.
species :
Dryopteris ardechensis Fraser-Jenk.; Fern Gaz. 12: 184 (1981)
Distribution: France; Corsica; Sardinia; Italy; ?Spain
12 COR FRA SAR 13 ITA
Remarks: tyrrhena agg.
Conservation Status: IUCN R
species :
Dryopteris arguta (Kaulf.) Watt; Canad. Naturalist & Quart. J. Sci., ser. 2, 13: 159 (1867)
Name: Coastal woodfern
Distribution: USA (Arizona, California, Oregon, Washington State); Canada (Vancouver Isl.); Mexico (Baja California Norte)
73 USA 76 USA 79 MEX
Synonyms: hidden
= Aspidium argutum Kaulf.; Enum. Fil. 242 (1824)
= Dryopteris arguta (Kaulf.) Maxon; Amer. Fern J. 11: 3 (1921)
= Dryopteris arguta f. bella Ewan; Amer. Fern J. 34: 112 (1944)
= Dryopteris arguta f. cristata Ewan; Amer. Fern J. 34: 112 (1944)
= Dryopteris arguta f. flocculosa Ewan; Amer. Fern J. 34: 112 (1944)
= Dryopteris arguta f. nudata Ewan; Amer. Fern J. 34: 111 (1944)
= Dryopteris rigida var. arguta (Kaulf.) Underw.; Native Ferns, ed. 4 116 (1893)
= Filix rigida var. arguta (Kaulf.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1916)
= Lastrea arguta (Kaulf.) Brack.; U. S. Expl. Exp. 16: 196 (1854)
= Nephrodium argutum (Kaulf.) Diels; Nat. Pfl. 1(4): 173 (1899)
= Nephrodium rigidum var. americanum Hook.; Brit. Ferns [W. J. Hooker] sub tab. 16 (1861)
= Thelypteris arguta (Kaulf.) Moxley; Bull. South. Calif. Acad. Sci. 19: 57 (1920)
species :
Dryopteris arunachalensis (Fraser-Jenk. & Benniamin) comb.ined.
Distribution: India (Arunachal Pradesh, ?Meghalaya)
40 IND
Remarks: previously reported as D. paucisora
Synonyms: hidden
= Dryopsis arunachalensis Fraser-Jenk. & Benniamin; Annot. Checkl. Ind. Pterid. 2: 219 (2018)
species :
Dryopteris ascensionis (Hook.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: Ascension Isl.
28 ASC
Conservation Status: IUCN Ex/E (a single plant seen 1976)
Synonyms: hidden
= Aspidium ascensionis (Hook.) Kuhn; Fil. Afr. 126 (1868)
= Lastrea ascensionis (Hook.) J.Sm.; Hist. Fil. 215 (1875)
= Nephrodium ascensionis Hook.; Sp. 4: 124, t. 257 (1862)
species :
Dryopteris assamensis (Hope) C.Chr. & Ching; Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. 6: 4 (1933)
Distribution: China (Guangdong, Hongkong, Guizhou, Guangxi); NE-India (Assam, Arunachal Pradesh, Meghalaya, +West Bengal); Vietnam; Thailand; Myanmar
36 CHI 40 IND 41 MYA THA VIE
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris filix-mas subsp. assamense (C.Hope) C.Chr.; Ind. Fil. 5: 266 (1905)
= Lastrea filix-mas var. assamensis (C.Hope) Bedd.; Suppl. Ferns Brit. Ind. 57 (1892)
= Nephrodium assamense C.Hope; J. Bot. 326 (1890)
species :
Dryopteris athamantica (Kunze) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Ghana; Nigeria; Cameroon; Congo [Brazzaville]; Central African Republic; D.R.Congo [Zaire]; Sudan; Ethiopia; Uganda; Rwanda; Burundi; Kenya; Tanzania; Angola; Zambia; Malawi; Mozambique; Zimbabwe; South Africa (Limpopo, North West, Gauteng, Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho; Eswatini [Swaziland]
22 GHA GUI IVO LBR NGA SIE 23 BUR CAF CMN CON RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 LES SAF SWZ
Synonyms: hidden
= Aspidium athamanticum Kunze; Linn. 18: 133 (1844)
= Lastrea athamantica (Kunze) T.Moore; J. Bot. 5: 311 (1853)
= Lastrea plantii T.Moore; J. Bot. 5: 227 (1853)
= Nephrodium athamanticum (Kunze) Hook.; Sp. 4: 125, t. 258 (1862)
= Nephrodium eurylepium Peter; Repert. Spec. Nov. Regni Veg. Beih. 40(1): 57 (1929)
species :
Dryopteris atrata (Wall) Ching; Sinensia 3: 326 (1933)
Distribution: China (Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan, Zhejiang); Taiwan; Tibet; Bhutan; India (Arunachal Pradesh, Assam State, Kerala, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, West Bengal); Myanmar (Kachin, Chin, etc.); Nepal; Thailand; Vietnam
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA THA VIE
Remarks: many Himalayan records of Dryopteris atrata refer to D. stenolepis (gamblei) fide Fraser-Jenkins but true D. atrata occurs at least in China, Nepal and N-India
Synonyms: hidden
= Aspidium atratum Wall. ex Kunze; Linn. 24: 279 (1851)
= Dryopteris hirtipes subsp. atrata (Wall. ex Kunze) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 333 (1989)
= Dryopteris hirtipes var. atrata (Wall. ex Kunze) C.Chr.; Contr. U. S. Natl. Herb. 26(6): 278 (1931)
= Nephrodium atratum (Wall. ex Kunze) Hand.-Mazz.; Symb. Sin. 6: 22 (1929)
species :
Dryopteris aurantiaca J.P.Roux; Syst. Biodivers. 2(2): 164 (2004 publ. 2005)
Distribution: Annobon Isl.
23 ANN
species :
Dryopteris austroindica Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 370 (1989), nom. nov.
Distribution: S-India (Tamil Nadu)
40 IND
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Lastrea intermedia Bedd.; Ferns South. India, t. 113. 1863 (non C. Presl (1836))
species :
Dryopteris bamleriana Rosenst.; Repert. Spec. Nov. Regni Veg. 10: 334 (1912)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
species :
Dryopteris barbigera (Hook.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: Pakistan (Kurram, Swat, Hazara, Gilgit); Tajikistan; Kazakhstan; Uzbekistan; China (Yunnan, Sichuan, Qinghai); Nepal; Bhutan; Myanmar; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Arunachal Pradesh, West Bengal)
32 KAZ TZK UZB 36 CHI 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Aspidium barbigerum (T.Moore ex Hook.) Christ; Farnkr. 260 (1897)
= Dryopteris barbigera var. falconeri (T.Moore ex Hook.) R.R.Stewart; Bull. Torrey Bot. Club 72(4): 407 (1945)
= Dryopteris falconeri (T.Moore ex Hook.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
= Lastrea barbigera (T.Moore ex Hook.) Bedd.; Handb. Ferns Brit. India, t. 227 (1867)
= Lastrea falconeri (T.Moore ex Hook.) Bedd.; Handb. Ferns Brit. India, t. 41 (1865)
= Nephrodium barbigerum T.Moore ex Hook.; Sp. Fil. 4: 113 (1862)
= Nephrodium falconeri Hook.; Sp. 4: 123, t. 254 (1862)
species :
Dryopteris basisora Christ; Not. syst. 1: 44 (1909)
Distribution: Bhutan; Nepal; N-India (Uttarakhand, Arunachal Pradesh); China (Yunnan, Sichuan)
36 CHI 40 BHU IND NEP
Remarks: D. lachoongensis is included here by Fraser-Jenkins et al. (2018) but kept separate by Knapp & Hsu (2017)
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium pseudovarium Christ; Bull. Soc. Bot. France 52, Mem. 1. 42 (1905)
= Aspidium varium var. obtusum Christ; Mem. Soc. Bot. France 52(Mem. 1): 43 (1905)
= Dryopteris pseudovaria (Christ) C.Chr.; Ind. 287 (1905)
= Dryopteris fructuosa var. integriloba Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 446 (1938) (non Dryopteris integriloba Ching 1933)
= Dryopteris tieluensis Ching & Y.P.Hsu; Fl. Tsinling. 2: 228 (1974) [fide Fraser-Jenkins (1986)]
species :
Dryopteris bernieri Tardieu; Notul. Syst. (Paris) 16: 161 (1956)
Distribution: La Réunion
29 REU
species :
Dryopteris blanfordii (Hope) C.Chr.; Ind. Fil. 254 (1905)
subspecies :
subsp. blanfordii
Distribution: China (W-Sichuan); Tibet; E-Afghanistan; Pakistan (Kurram, Swat, Hazara); India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand); Nepal
34 AFG 36 CHI TIB 40 IND NEP PAK
Synonyms: hidden
= Lastrea spinulosa Bedd.; Handb. Ferns Brit. India 336 (1870) (non C. Presl (1836))
= Lastrea spinulosa var. remota (C.B.Clarke) Bedd.; Handb. Ferns Brit. India 252 (1883)
= Nephrodium blanfordii Hope; J. Bombay Nat. Hist. Soc. 12: 624, t. 11 (1899)
= Nephrodium remotum C.B.Clarke; Tr. Linn. Soc. II, 1: 523 (1880)
= Dryopteris mehrae Khullar; (nom. nud., fide Fraser-Jenkins 1997)
subspecies :
subsp. nigrosquamosa (Ching) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 388 (1989)
Distribution: China (Gansu, Sichuan, Yunnan); Tibet
36 CHI TIB
Synonyms: hidden
= Dryopteris blanfordii subsp. gongboensis (Ching) Fraser-Jenk.; Pakistan Syst. 5(1-2): 86 (1991) (fide Fraser-Jenkins 1997)
= Dryopteris gongboensis Ching; Fl. Xizangica 1: 269 (1983)
= Dryopteris gushaingensis Ching; Fl. Xizangica 1: 269 (1983)
= Dryopteris nigrosquamosa Ching; Bull. Fan Mem. Inst. Biol. Bot. 2: 194 (1931)
= Dryopteris sublaeta Ching & Y.P.Hsu; Fl. Tsinling. 2: 227 (1974)
species :
Dryopteris bodinieri (Christ) C.Chr.; Ind. 254 (1905)
Distribution: China (Guizhou, Yunnan, Sichuan, Hunan)
36 CHI
Synonyms: hidden
= Aspidium bodinieri Christ; Bull. Ac. Géogr. Bot. Mans 248 (1902)
= Nephrodium cyclodioides Baker; Bull. Misc. Inform. Kew 1900: 10 (1900)
species :
Dryopteris bojeri (Baker) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: +Mauritius
29 MAU
Synonyms: hidden
= Nephrodium bojeri Baker; Syn. 280 (1867)
= Nephrodium amplum Bory; Bél., Voy. Bot. 2: 62 (1833)
species :
Dryopteris borreri (Newman) Oberh. & Tavel; Verh. Schweiz. Naturf. Ges. 118: 153 (1937)
Name: Borrers Wurmfarn
Distribution: Portugal; Spain; Andorra; France; Corsica; Belgium; Luxembourg; Netherlands; Scotland; England; Channel Isl.; Wales; Scotland; Ireland; Northern Ireland; Denmark; SW-Norway; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Italy (incl. Elba Isl.); Sicily; Austria; Czech Republic; Bosnia & Hercegovina; Montenegro; Croatia; Macedonia; Serbia & Kosovo; Slovenia; Romania; Bulgaria; European Turkey; Turkey (Anatolia); European Russia (Kaliningrad district); Ukraine; ?Crimea; Northern Caucasus (Krasnodar, Daghestan, Severo-Ossetia); Georgia [Caucasus] (incl. Adzharia); Azerbaijan; NW-Iran; Morocco
10 DEN GRB IRE NOR 11 AUT BGM CZE GER LUX NET SWI 12 ADO CIS COR FRA POR SPA 13 BIH BUL CRO ITA MNT NMA ROM SIC SLV SRB TUE 14 RUS UKR 20 MOR 33 AZE GEO NCS 34 IRN TUR
Remarks: affinis agg.
Synonyms: hidden
= Aspidium blackwellianum Ten.; Atti Ist. Incor. Napoli 6: 133 (1830), nom. prov. inval.
= Aspidium filix-mas subvar. ursinum Walther Zimm.; 53 (1916)
= Aspidium filix-mas var. stilluppense Sabr.; Österr. Bot. Zeit. 52: 144 (1902) (basion.)
= Dryopteris abbreviata (DC.) Newman; Phytol. app. 21 (1851) ex Manton (1950) (p. p. incl. typo, non D. abbreviata (Schrad.) Kuntze (1891))
= Dryopteris affinis subsp. borreri (Newman) Fraser-Jenk.; Willdenowia 10(1): 110 (1980)
= Dryopteris affinis subsp. stilluppensis (Sabr.) Fraser-Jenk.; Willdenowia 10(1): 112 (1980)
= Dryopteris affinis var. rubiginosa (Fomin) Askerov; Zametki Sist. Geogr. Rast. 39: 7 (1983)
= Dryopteris borreri f. aurea Tavel; 154 (1937), nom. inval.
= Dryopteris borreri var. melanothrix Tavel; 153 (1937), nom. inval.
= Dryopteris borreri var. robusta Tavel; 154 (1937), nom. inval.
= Dryopteris borreri var. tenuis Tavel; 154 (1937), nom. inval.
= Dryopteris borreri var. ursina (Walther Zimm.) Tavel; 153 (1937)
= Dryopteris filix-mas f. subintegra (Döll) Stefánsson; 5 (1924)
= Dryopteris filix-mas nothosubsp. tavelii (Rothm.) Schidlay
= Dryopteris filix-mas subsp. abbreviata (DC.) Schidlay; (p. p. incl. typo)
= Dryopteris filix-mas subsp. borreri (Newman) Bech. & Tavel
= Dryopteris filix-mas var. abbreviata (DC.) Newman; (p. p. incl. typo)
= Dryopteris filix-mas var. borreri Newman; Hist. br. Ferns ed. III: 189 (1854) (basion.)
= Dryopteris paleacea (T.Moore) Fomin; Kuzn. & al., Fl. Caucas. Crit. 1(1): 52 (1911) (non (Sw.) C. Chr., nec (D. Don) Hand.-Mazz.)
= Dryopteris paleacea var. borreri (Newman) H.Wolff
= Dryopteris paleacea var. rubiginosa Fomin
= Dryopteris stilluppensis (Sabr.) Holub; Preslia 58(4): 302 (1986)
= Dryopteris tavelii f. concinna (Jones) Rothm.; 93 (1946)
= Dryopteris tavelii f. crispa (Sim) Rothm.; 93 (1946)
= Dryopteris tavelii f. cristata (T.Moore & Houlston) Rothm.; 93 (1946)
= Dryopteris tavelii f. pinderi (T.Moore) Rothm.; 93 (1946)
= Dryopteris ×tavelii Rothm.; Candollea 10: 92 (1946) (fide Fraser-Jenkins (2007))
= Filix filix-mas var. paleacea (Sw.) Farw.; Amer. Midl. Naturalist 12: 254 (1931)
= Lastrea abbreviata (DC.) Woll.; Phytol. n. s. 1: 172 (1855)
= Lastrea filix-mas var. abbreviata (DC.) T.Moore; 138 (1848)
= Lophodium abbreviatum (DC.) Newman; Phyt. 4: app. 21 (1851)
= Nephrodium abbreviatum (DC.) Lowe; Fern Grow. 6: 161 (1895)
= Nephrodium borreri Rouy; Fl. France 14: 408 (1913)
= Nephrodium filix-mas f. paleaceum (Lag. ex Sw.) Fiori; Fl. It. 1: 8 (1896)
= Nephrodium filix-mas var. abbreviatum (DC.) Hook.f.; 465 (1870)
= Nephrodium filix-mas var. borreri (Newman) Hook.f.; 465 (1870)
= Polystichum abbreviatum DC.; Fl. franc. 2: 560 (1805)
= Polystichum filix-mas var. abbreviatum (DC.) Gren.; 631 (1856)
= Polystichum induratum Schur; Enum. Pl. Transsilv. 834 (1866)
variant :
var. robusta Oberh. & Tavel; Verh. Schweiz. Naturf. Ges. 118: 153-154 (1937) nom. nud.
Distribution: England; Wales; Scotland; Germany; C-Europe etc.
10 GRB 11 GER
Remarks: as an apomictic taxon it possibly deserves recognition as species, see Flora of Great Britain.
Synonyms: hidden
= Dryopteris pseudomas subsp. robusta (Oberh. & Tavel ex Fraser-Jenk.) Holub; Folia Geobot. Phytotax. 19(2): 214 (1984)
= Dryopteris mediterranea var. robusta (Fraser-Jenk.) Holub; Preslia 58(4): 301 (1986)
= Dryopteris affinis var. robusta (Oberh. & Tavel ex Fraser-Jenk.) Fraser-Jenk. & Salvo; Anal. Jard. Bot. Madrid 41(1): 195 (1984)
= Dryopteris affinis subsp. robusta Oberh. & Tavel ex Fraser-Jenk.; Willdenowia 10(1): 111 (1980)
= Dryopteris robusta (Oberh. & Tavel ex Fraser-Jenk.) P.D.Sell; Fl. Great Britain and Ireland 1: 671 (2018)
species :
Dryopteris cacaina Tagawa; Acta Phytotax. Geobot. 6: 155 (1937)
Distribution: Taiwan; Philippines; Japan (Yakushima Isl.)
38 JAP NNS TAI 42 PHI
Synonyms: hidden
= Dryopteris yakusilvicola Kurata; J. Geobot. 15: 85 (1967)
species :
Dryopteris cambrensis (Fraser-Jenk.) Beitel & W.R.Buck; Fiddlehead Forum 15(2): 15 (1988)
subspecies :
subsp. cambrensis
Name: Walisischer Wurmfarn (DE)
Distribution: England; Scotland; Wales; France; Ireland; Northern Ireland; Norway; Croatia; Turkey (Anatolia)
10 GRB IRE NOR 12 FRA 13 CRO 34 TUR
Remarks: affinis agg.
Synonyms: hidden
= Dryopteris affinis subsp. cambrensis Fraser-Jenk.; Sommerfeltia 6: xi (1987)
= Dryopteris affinis var. paleaceocrispa (T.Moore) Fraser-Jenk.; Indian J. Forest. 43(1): 83 (2021)
= Lastrea filix-mas subvar. paleaceocrispa T.Moore; Brit. Ferns [T. Moore] 1: 196 (1859)
subspecies :
subsp. distans (Viv.) Fraser-Jenk.; Fern Gaz. 18(1): 6 (2007)
Distribution: Corsica; Italy; France
12 COR FRA 13 ITA
Remarks: affinis agg.
Synonyms: hidden
= Polystichum distans (Viv.) Trevis.; Syll. Sporoph. Ital. 19 (1874)
= Nephrodium distans (Viv.) Ces.; Pass. & Gib. Comp. Fl. ltal. (?) (1870)
= Aspidium distans Viv.; App. ad Fl. Cors. Prod. 8 (1825)
subspecies :
subsp. insubrica (Oberh. & Tavel ex Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 6 (2007)
Name: Insubrischer Wurmfarn (DE)
Distribution: France; Corsica; Belgium; Luxembourg; Netherlands; Denmark; Norway; S-Switzerland (Tessin); Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Italy (incl. Elba Isl.); Sicily; Sardinia; Austria; Czech Republic; Slovenia; Croatia; Bulgaria; S-Poland
10 DEN NOR 11 AUT BGM CZE GER LUX NET POL SWI 12 COR FRA SAR 13 BUL CRO ITA SIC SLV
Remarks: affinis agg.
Synonyms: hidden
= Dryopteris affinis var. insubrica Oberh. & Tavel ex Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)
species :
Dryopteris campyloptera (Kunze) Clarkson; Amer. Fern J. 20: 118 (1930)
Name: Mountain woodfern
Distribution: USA (Alabama, Connecticut, Massachusetts, Maryland, Maine, Michigan, North Carolina, New Hampshire, New York, Pennsylvania, Tennessee, Virginia, Vermont, West Virginia); Canada (Labrador, New Brunswick, Newfoundland, Nova Scotia, Prince Edward Isl., Québec)
75 USA 78 USA
Synonyms: hidden
= Aspidium campylopterum Kunze; Amer. J. Sci. Arts, ser. 2, 6: 84 (1848)
= Aspidium spinulosum var. americanum Fisch. ex Kunze; Amer. J. Sci. Arts, Ser. 2, 6: 84 (1848)
= Aspidium spinulosum var. concordianum (Davenp.) Eastman; Ferns & Fern Allies New England 20 (1904)
= Dryopteris austriaca (Jacq.) Woyn.; Schinz & Thell., Vierteljahrsschr. Naturf. Ges. Zürich 60: 339 (1915)
= Dryopteris austriaca var. concordiana (Davenp.) C.V.Morton; Amer. Fern J. 40: 222 (1950)
= Dryopteris dilatata subsp. americana (Fisch. ex Kunze) Hultén
= Dryopteris intermedia f. concordiana (Davenp.) Clute; Ferns in Their Haunts 184 (1938)
= Dryopteris intermedia var. concordiana (Davenp.) M.Broun; Index N. Amer. Ferns 71 (1938)
= Dryopteris spinulosa var. americana (Fisch. ex Kunze) Fernald; Rhodora 17: 48 (1915)
= Dryopteris spinulosa var. concordiana (Davenp.) Eastman; Ferns & Fern Allies New England 20 (1904)
= Filix spinulosa var. americana (Fisch. ex Kunze) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1916)
= Filix spinulosa var. concordiana (Davenp.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 83 (1916)
= Filix-mas rigida var. americana (Fisch. ex Kunze) Farw.; Amer. Midl. Naturalist 12: 256 (1931)
= Filix-mas spinulosa var. americana (Fisch. ex Kunze) Farw.; Amer. Midl. Naturalist 12: 257 (1931)
= Nephrodium spinulosum f. concordianum (Davenp.) Clute
= Nephrodium spinulosum var. concordianum Davenp.; Rhodora 6: 33 (1904)
= Thelypteris dilatata var. americana (Fisch. ex Kunze) House; New York State Mus. Bull. 233-234: 69 (1922)
= Thelypteris spinulosa var. americana (Fisch. ex Kunze) Weath.; Rhodora 21: 178 (1919)
= Thelypteris spinulosa var. concordiana (Davenp.) Weath.; Rhodora 21: 178 (1919)
species :
Dryopteris camusiae Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 131 (1997)
Distribution: NE-India (Nagaland); China (Yunnan); ?Myanmar
36 CHI 40 IND
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
species :
Dryopteris caperata J.P.Roux; Syst. Biodivers. 2(2): 164 (2004 publ. 2005)
Distribution: São Tomé
23 SAO
species :
Dryopteris caroli-hopei Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 422 (1989), nom. nov.
Distribution: India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Assam State, Arunachal Pradesh, Nagaland, Meghalaya, Manipur, Mizoram, Sikkim, West Bengal); Bhutan; Nepal; Tibet; China (C-Yunnan); Myanmar (Chin, etc.)
36 CHI TIB 40 BHU IND NEP 41 MYA
Synonyms: hidden
= Aspidium dilatatum var. patuloides Christ; Bull. Soc. Bot. France 52(Mem. 1): 41 (1905)
= Dryopteris pseudomarginata Ching; Sporae Pterid. Sin. 327 (1976), nom. nud.
species :
Dryopteris carpatica S.Jess. & Bujnoch; Fern Gaz. 21(8): 462 (2022)
Name: Carpathian Scaly Male Fern
Distribution: Romania
13 ROM
Remarks: affinis agg.
species :
Dryopteris carthusiana (Vill.) H.P.Fuchs; Bull. Soc. Bot. France 105: 339 (1959)
Name: Dorniger Wurmfarn, Gewöhnlicher Dornfarn (DE), Toothed woodfern (EN)
Distribution: Albania; Austria; Belgium; Luxembourg; England; Wales; Scotland; Bulgaria; Corsica; Czech Republic; Slovakia; Denmark; Finland; France; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Greece; Ireland; Northern Ireland; Switzerland; Liechtenstein; Netherlands; Spain; Hungary; Italy; Bosnia & Hercegovina; Montenegro; Croatia; Macedonia; Serbia & Kosovo; Slovenia; Norway; Poland; Romania; Sweden; ?European Turkey; Turkey (Anatolia); Estonia; Latvia; Lithuania; N-European Russia; C-European Russia; W-European Russia; E-European Russia; Belarus; Ukraine; Kazakhstan; Crimea; Northern Caucasus; Georgia [Caucasus]; Azerbaijan; Armenia; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, W-Siberia); China (Xinjiang); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Idaho, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Montana, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Oregon, Pennsylvania, Rhode Island, South Carolina, South Dakota, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia); Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); St. Pierre et Miquelon
10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 COR FRA SPA 13 ALB BIH BUL CRO GRC ITA MNT NMA ROM SLV SRB 14 BLR EST KRY LAT LIT RUS UKR 30 SIB 32 KAZ 33 ARM AZE GEO NCS 34 TUR 36 CHI 72 SPM 73 USA 74 USA 75 USA 78 USA
Synonyms: hidden
= Aspidium callipteris F.H.Wilms; Verh. Rhein. Ges. 9: 577 (1852), nom. illeg., non Fée
= Aspidium euspinulosum J.T.Jansen; 289 (1932), nom. inval.
= Aspidium groenlandicum Gand.; Bull. Soc. Bot. France 66: 305 (1920)
= Aspidium pringleanum Gand.; Bull. Soc. Bot. France 66: 305 (1920)
= Aspidium spinulosum (O.F.Müll.) Sw.; 38 (1801), nom. illeg.
= Aspidium spinulosum f. anadenium B.L.Rob.; Rhodora 9: 84 (1907)
= Aspidium spinulosum var. elevatum A.Braun ex Döll; 17 (1843)
= Dryopteris austriaca subsp. spinulosa (O.F.Müll.) Schinz & Thell.; 340 (1915)
= Dryopteris austriaca var. spinulosa (O.F.Müll.) Fiori; Filic. Equiset. Lycopod. 115 (1943)
= Dryopteris eu-spinulosa (Asch.) Fomin; Mon. Jard. Bot. Tiflis 20: 53 (1911)
= Dryopteris lanceolatocristata (Hoffm.) Alston; Watsonia 4: 41 (1957)
= Dryopteris spinulosa (O.F.Müll.) Kuntze; Rev. Gen. Pl. 2: 810 (1891) (comb. superfl.)
= Dryopteris spinulosa var. elevata (A.Braun ex Döll) Vollm.; (1914)
= Dryopteris spinulosa var. exaltata (Lasch) Vollm.; (1914)
= Dryopteris spinulosa var. glandulosa Gilbert; Fern Bull. 8: 11 (1900)
= Dryopteris spinulosa var. patuloides Christ; Bull. Soc. Bot. France 62, Mem. 1: 41 (1905)
= Filix spinulosa (O.F.Müll.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix-Mas spinulosa (O.F.Müll.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Lastrea dilatata var. linearis Bab.; 386 (1843)
= Lastrea spinosa (Schrank) Newman; Natur. Alm. 21 (1844)
= Lastrea spinulosa (O.F.Müll.) C.Presl; Tent. Pterid. 76 (1836)
= Lastrea spinulosa var. elevata (A.Braun ex Döll) Syme & N.E.Br.; 78 (1886)
= Lastrea spinulosa var. exaltata (Lasch) Syme & N.E.Br.; 78 (1886)
= Lophodium spinosum (Weiss) Newman; Phyt. 4: 371 (1851)
= Nephrodium jordanii Rouy; Fl. France 14: 411 (1913)
= Nephrodium spinulosum (Retz.) Strempel; Fil. Berol. Syn.: 30 (1822) (nom. illeg.)
= Nephrodium spinulosum f. anadenium (B.L.Rob.) Clute; Fern Bull. 17: 20 (1909)
= Nephrodium spinulosum var. elevatum (A.Braun ex Döll) Murr; 9 (1923)
= Nephrodium spinulosum var. swarzii F.W.Schultz; 566 (1845), nom. inval.
= Polypodium carthusianum Vill.; Hist. pl. Dauph. 1: 292 (1786) (basion.)
= Polypodium filix-femina var. spinosa F.G.Weiss; (1770)
= Polypodium lanceolatocristatum Hoffm.; 9 (1790)
= Polypodium spinosum Schrank; Baier. Fl. 2: 424 (1789)
= Polypodium spinulosum O.F.Müll.; Oeder, Fl. Dan. 12: 7 (1777) (non Burman fil., Fl. Indica: 233 (1768))
= Polypodium tanacetifolium Hoffm.; Deutschl. Fl. 2: 8 (1795)
= Polystichum angustatum Schum.; Enum. Pl. Saell. 2: 22 (1803)
= Polystichum carthusianum (Vill.) Sanio; Verh. Bot. Vereins Prov. Brandenburg 25. 84 (1884)
= Polystichum conifolium Schum.; Enum. Pl. Saell. 2: 22 (1803)
= Polystichum muelleri Schum.; Enum. Pl. Saell. 2: 20 (1803)
= Polystichum pseudocristatum Schur; Sert. Pl. Transs. 94 (1852)
= Polystichum spinosum (F.G.Weiss) Roth; Röm., Arch. 2(1): 106 (1799)
= Polystichum spinulosum (O.F.Müll.) Lam. & DC.; Fl. franc. 2: 561 (1805)
= Thelypteris spinulosa (O.F.Müll.) Nieuwl.; Midl. Naturalist 1: 226 (1910)
variant :
var. enigmatica Harmaja; Memoranda Soc. Fauna Fl. Fenn. 88: 23 (2012)
Distribution: Finland
10 FIN
species :
Dryopteris caucasica (A.Braun) Fraser-Jenk. & Corley; Brit. Fern Gaz. 10: 222 (1972)
Distribution: European Turkey; Turkey (Anatolia); NW-Iran; Northern Caucasus; Georgia [Caucasus]; Armenia; ?Azerbaijan; Crimea
13 TUE 14 KRY 33 ARM GEO NCS 34 IRN TUR
Remarks: filix-mas agg.
Synonyms: hidden
= Aspidium affine Fisch. & C.A.Mey.; Bull. Soc. Nat. Moscou 240 (1888) (non Blume (1828))
= Aspidium caucasicum A.Braun; Flora (Regensburg) 24: 707 (1841) (basion.)
= Aspidium filix-mas var. incisum (T.Moore) Döll; 27 (1855)
= Dryopteris affinis Newman; Hist. br. Ferns ed. III: 187 (1854)
= Dryopteris filix-mas subsp. caucasica (A.Braun) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)
= Dryopteris filix-mas var. affinis (Fisch. & C.A.Mey.) B.Fedtsch.; 9 (1927), nom. illeg.
= Dryopteris filix-mas var. affinis (Fisch. & Mey.) Askerov; Zametki Sist. Geogr. Rast. 39: 7 (1983)
= Lastrea filix-mas var. incisa T.Moore; 138 (1848)
species :
Dryopteris caudipinna Nakai; Bot. Mag. (Tokyo) 45: 96 (1931)
Distribution: Japan
38 JAP
Synonyms: hidden
= Dryopteris erythrosora var. caudipinna (Nakai) H.Itô; (1939)
species :
Dryopteris celsa (W.Palmer) Knowlt., W.Palmer & Pollard; Proc. Biol. Soc. Wash. 13: 202 (1900)
Name: Log fern
Distribution: USA (Alabama, Arkansas, District of Columbia, Delaware, Georgia, Illinois, Indiana, Kentucky, Louisiana, Maryland, Michigan, Missouri, North Carolina, New Jersey, New York, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)
74 USA 75 USA 78 USA
Synonyms: hidden
= Aspidium goldieanum f. celsum (W.Palmer) Clute; Ferns in Their Haunts 313 (1901)
= Aspidium goldieanum var. celsum (W.Palmer) B.L.Rob.; Rhodora 10: 30 (1908)
= Dryopteris clintoniana f. atropalustris (Small) Wherry; Amer. Fern J. 40: 120 (1950)
= Dryopteris clintoniana f. celsa (W.Palmer) Wherry; Amer. Fern J. 40: 119 (1950)
= Dryopteris clintoniana var. atropalustris (Small) C.F.Reed; Bull. Nat. Hist. Soc. Maryland 13: 51 (1943)
= Dryopteris goldieana subsp. celsa W.Palmer; Proc. Biol. Soc. Wash. 13: 65 (1899)
= Dryopteris goldieana f. celsa (W.Palmer) Clute; Ferns in Their Haunts 172, 385 (1938)
= Dryopteris wherryi Crane; Amer. Fern J. 50: 271, t. 23(6) (1960)
= Dryopteris ×atropalustris Small; Ferns S. E. States 274 cum tab. (1938)
= Filix goldieana var. celsa (W.Palmer) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1917)
species :
Dryopteris chaerophyllifolia (Zippel) C.Chr.; Ind. 257 (1905)
Distribution: Borneo (Mt. Kinabalu, Sabah); Java; Vanuatu
42 BOR JAW 60 VAN
Synonyms: hidden
= Aspidium chaerophyllifolium Zippel; Mett., Ann. Mus. Bot. Lugd.-Bat. 1: 228 (1864)
species :
Dryopteris championii (Benth.) C.Chr.apud Ching; Sinensia 8: 327 (1933)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shandong, Sichuan, Yunnan, Zhejiang); Taiwan; Japan; South Korea (incl. Cheju Isl.)
36 CHI 38 JAP KOR TAI
Synonyms: hidden
= Aspidium championii Benth.; Fl. Hongk. 456 (1861)
= Aspidium erythrosorum var. amoyense Christ; Warb., Monsunia 1: 80 (1900)
= Dryopteris bullatipaleacea Ching; Bull. Bot. Res. Harbin 2(2): 65-66, pl. 2, f. 1 (1982)
= Dryopteris changii Ching; Wuyi Sci. J. 1: 7 (1981)
= Dryopteris conferta Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 17 (1987) (nom. illeg.)
= Dryopteris erythrosora subsp. chiliolepis Hand.-Mazz.; Akad. Anz. Akad. Wiss. Wien no. 7 (1922)
= Dryopteris erythrosora var. cavaleriei Rosenst.; Repert. Spec. Nov. Regni Veg. 13(355-358): 131-132 (1914)
= Dryopteris grandiosa Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 8 (1987)
= Dryopteris huanglungensis Ching; Bot. Res. Acad. Sin. 1987(2): 10 (1987)
= Dryopteris infrahirtella Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 3(4): 22 (1983)
= Dryopteris jinfoshanensis Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 6(1): 179 (1986), nom. nov.
= Dryopteris laoshanensis J.X.Li & S.T.Ma; Bull. Bot. Res. Harbin 3(4): 139 (1983)
= Dryopteris lepigera C.Chr.; Ind. 274 (1905) (non (Baker) Kuntze)
= Dryopteris linganensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 23 (1983)
= Dryopteris nanchuanensis Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(3): 19 (1984) (non 3(4) (1983))
= Dryopteris neofuscipes Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(3): 22 (1984)
= Dryopteris paleifera Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 2 (1984)
= Dryopteris pseudo-erythrosora Kodama; Matsum., Ic. pl. Koisik. 1. 165, t. 83 (1913)
= Dryopteris qinyuanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 7 (1987)
= Dryopteris wangii Ching; Bot. Res. Acad. Sin. 1987(2): 13 (1987)
= Dryopteris wuyishanensis Ching; Fl. Fujianica 1: 600 (1982)
= Dryopteris yandongensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 13 (1983)
= Dryopteris zunyiensis Ching; Bot. Res. Acad. Sin. 1987(2): 18 (1987)
species :
Dryopteris chinensis (Baker) Koidz.; Fl. Symb. Orient. As. 39 (1930)
Distribution: China (Anhui, Henan, Jiangsu, Jiangxi, Liaoning, Shandong, Zhejiang); Japan; North Korea; South Korea (incl. Cheju Isl.); Russian Far East (Primorye)
31 RFE 36 CHI 38 JAP KOR NKO
Synonyms: hidden
= Aspidium forbesii Hance; J. Bot. 198 (1876)
= Nephrodium chinense Baker; Hook. & Baker, Syn. Fil. 278 (1867)
species :
Dryopteris christensenae (Ching) Li Bing Zhang; Taxon 61(6): 1209 (2012), nom. nov.
Distribution: China (Yunnan)
36 CHI
Remarks: subgen. Nothoperanema
Synonyms: hidden
= Diacalpe christensenae Ching; Bull. Fan Mem. Inst. Biol. Bot. 2, 1: 291 (1949)
species :
Dryopteris chrysocoma (Christ) C.Chr.; Ind. 257 (1905)
Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; India (Jammu & Kashmir, Himachal Pradesh, Arunachal Pradesh, Meghalaya, Mizoram, Sikkim, Uttarakhand, West Bengal); Nepal; Bhutan; Pakistan (Hazara); Philippines; Myanmar; Vietnam
36 CHI TIB 40 BHU IND NEP PAK 41 MYA VIE 42 PHI
Synonyms: hidden
= Aspidium chrysocoma (Christ) Christ; Bull. Acad. Int. Geogr. Bot. Il 253 (1902)
= Aspidium filix-mas var. chrysocoma Christ; Bull. Herb. Boiss. 6: 966 (1898)
= Dryopteris alpicola Ching & Z.R.Wang; Acta Phytotax. Sin. 23(5): 349 (1985), nom. nov. for D. chrysocoma var. alpina Ching 1937
= Dryopteris chrysocoma var. alpina Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 438 (1937), not D. alpina Rosenst. (1913) (fide Fraser-Jenkins et al. 2015)
= Dryopteris chrysocoma var. majus Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 438 (1938)
= Dryopteris chrysocoma var. squamosa (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 438 (1938)
= Dryopteris cochleata var. squamosa C.Chr.; Acta Hort. Gothob. 1: 59 (1924)
= Dryopteris daliensis (Z.R.Wang) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 119 (1997) (pro hybr.)
= Dryopteris gracilofrons C.Chr.; Ind. 268 (1905) (nom. nud., fide Fraser-Jenkins (1997))
= Dryopteris macrocarpa Stewart; Bull. Torrey Bot. Cl. 72: 406 (1945) (fide Fraser-Jenkins 1997)
= Dryopteris parachrysocoma Ching & Z.R.Wang; Acta Phytotax. Sin. 23(5): 344 (1985) (fide Fraser-Jenkins 1997)
= Dryopteris pseudochrysocoma Ching; Acta Phytotax. Sin. 23(5): 349 (1985), nom. nud.
= Dryopteris ×daliensis Z.R.Wang; Acta Phytotax. Sin. 23(5): 346 (1985)
= Lastrea filix-mas var. elongata Bedd.; Handb. Ferns Brit. India 250 (1883) (non Dryopteris elongata (Wall. ex Hook.) Kuntze, fide Fraser-Jenkins (1997))
= Lastrea filix-mas var. schimperiana Bedd.; Handb. Ferns Brit. India, Suppl. 58 (1892)
= Nephrodium chrysocoma (Christ) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)
= Nephrodium filix-mas var. schimperiana C.B.Clarke; Tr. Linn. Soc. II, 1: 520, t. 69, f. 2 (1880)
= Nephrodium schimperianum Hope; J. Bombay Nat. Hist. Soc. 14: 733 (1903) (non Aspidium schimperianum Hochst. (1841))
species :
Dryopteris cicatricata J.P.Roux; Syst. Biodivers. 2(2): 167 (2004 publ. 2005)
Distribution: São Tomé
23 SAO
species :
Dryopteris cinnamomea (Cav.) C.Chr.; Amer. Fern J. 1: 95 (1911)
Name: Cinnamon woodfern
Distribution: USA (Arizona, SW-Texas); Mexico (Aguascalientes, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas)
76 USA 79 MEX
Synonyms: hidden
= Aspidium agatholepis Fée; Mém. Foug. 8: 106 (1857)
= Aspidium athyrioides M.Martens & Galeotti; Mém. Acad. Roy. Sci. Bruxelles 15: 67, t. 18 (1842)
= Aspidium flaccidum E.Fourn.; Bull. Soc. Bot. France 27: 328 (1880) (non Blume (1828))
= Aspidium indecorum (Liebm.) E.Fourn.; Mex. Pl. 1: 97 (1872)
= Aspidium inquinans (T.Moore) Fée; Mém. Foug. 8: 106 (1867)
= Aspidium mexicanum (C.Presl) Kunze; Linn. 13: 147 (1839)
= Aspidium mexicanum var. acutilobum Kunze; Linnaea 13: 147 (1839)
= Aspidium mexicanum var. obtusilobum Kunze; Linnaea 13: 147 (1839)
= Aspidium roseum E.Fourn.; Mex. Pl. 1: 97 (1872)
= Aspidium sphaerocarpum (Fée) Christ; Bull. Herb. Boiss. II, 6: 726 (1905)
= Athyrium sphaerocarpon Fée; Gen. 186 (1850-52)
= Dryopteris athyrioides (M.Martens & Galeotti) Kuntze; Rev. Gen. Pl. 2: 811 (1891)
= Dryopteris fournieri (Baker) C.Chr.; Ind. 266 (1905)
= Dryopteris glandulifera (Liebm.) C.Chr.; Ind. 267 (1905)
= Dryopteris indecora (Liebm.) C.Chr.; Ind. 272 (1905)
= Dryopteris mexicana (C.Presl) C.Chr.; Vid. Selsk. Skr. VII, 10: 68, f. 1 (1913), isonym
= Dryopteris mexicana (C.Presl) Kuntze; Rev. Gen. Pl. 2: 811 (1891)
= Dryopteris patula var. mexicana (C.Presl) Bonap.; Notes Pteridol. 2: 130 (1915)
= Dryopteris rosea (E.Fourn.) Mickel & Beitel; Pterid. Fl. Oaxaca (Mem. New York Bot. Gard. 46): 165 (1988)
= Lastrea agatholepis (Fée) T.Moore; Ind. 85 (1858)
= Lastrea athyrioides (M.Martens & Galeotti) Liebm.; Vid. Selsk. Skr. V, 1: 274 (1849)
= Lastrea indecora Liebm.; Vid. Selsk. Skr. V, 1: 272 (seors. 120) (1849)
= Lastrea inquinans T.Moore; Ind. 94 (1858)
= Lastrea mexicana (C.Presl) C.Presl; Tent. Pterid. 76 (1836)
= Lophodium glanduliferum (Liebm.) Newman; Phytol. 4: 371 (1851)
= Nephrodium agatholepis (Fée) Kiaersk.; Cat. Fil. hb. ht. Haun. 62 (1874)
= Nephrodium fournieri Baker; Ann. Bot. 6: 317 (1891) (non p. 328)
= Nephrodium glanduliferum (Liebm.) Kiaersk.; Cat. Fil. hb. ht. Haun. 63 (1874)
= Nephrodium indecorum (Liebm.) Kiaersk.; Cat. Fil. hb. ht. Haun. 63 (1874)
= Nephrodium mexicanum C.Presl; Reliq. Haenk. 1: 38 (1825), nom. illeg., non auct.
= Nephrodium sphaerocarpum Hook.; Sp. 4: 139 (1862)
= Phegopteris glanduligera (Liebm.) E.Fourn.; Mex. Pl. 1: 90 (1872) (err.)
= Polypodium glanduliferum Liebm.; Vid. Selsk. Skr. V, 1: 206 (1849)
= Polystichum cystopteroides Nees; Linn. 19: 685 (1847)
= Tectaria cinnamomea Cav.; Descr. Pl. 252 (1802)
species :
Dryopteris clarkei (Baker) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: China (Sichuan, Guizhou, Yunnan, Guangxi); Tibet; India (Arunachal Pradesh, West Bengal, Sikkim); Myanmar (Kachin, etc.); Nepal; Bhutan
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: Subgen. Dryopsis
Synonyms: hidden
= Ctenitis clarkei (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 287 (1938)
= Dryopsis clarkei (Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 181 (1986)
= Dryopteris fibrillosa Hand.-Mazz.; Akad. Anz. Akad. Wiss. Wien no. 7: 2 (1922) (non Dryopteris fibrillosa (Baker) C. Chr.)
= Lastrea filix-mas subvar. clarkei (Baker) Bedd.; Handb. Ferns Brit. India 250 (1883)
= Lastrea filix-mas var. clarkei (Baker) Bedd.; Suppl. Ferns Brit. Ind. 17 (1876)
= Nephrodium clarkei Baker; Hook. & Baker, Syn. Fil. 497 (1874)
species :
Dryopteris clintoniana (D.C.Eaton) Dowell; Proc. Staten Island Assoc. Arts 1: 64 (1906)
Name: Clinton´s woodfern
Distribution: USA (Connecticut, District of Columbia, Delaware, Illinois, Indiana, Massachusetts, Maine, Michigan, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Vermont, Wisconsin); Canada (New Brunswick, Ontario, Québec)
74 USA 75 USA
Synonyms: hidden
= Aspidium cristatum var. clintonianum D.C.Eaton; Gray´s Manual, ed. 5: 665 (1867)
= Dryopteris cristata var. clintoniana (D.C.Eaton) Underw.; Native Ferns, ed. 4 115 (1893)
= Dryopteris ×poyseri Wherry; Amer. Fern J. 50: 90 (1960)
= Filix cristata var. clintoniana (D.C.Eaton) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81 (1916)
= Filix-mas cristata var. clintoniana (D.C.Eaton) Farw.; Amer. Midl. Naturalist 12: 253 (1931)
= Nephrodium cristatum var. clintonianum (D.C.Eaton) Gilbert; Working List N. Amer. Pterid. 17 (1901)
= Thelypteris clintoniana (D.C.Eaton) House; New York State Mus. Bull. 233-234. 69 (1920)
= Thelypteris cristata var. clintoniana (D.C.Eaton) Weath.; Rhodora 21: 177 (1919)
species :
Dryopteris cochleata (D.Don) C.Chr.; Ind. Fil. 258 (1905)
Distribution: China (Yunnan, Guizhou, Sichuan); Bangladesh; Bhutan; Philippines; Thailand; Vietnam; India (Jammu & Kashmir, Andhra Pradesh, Arunachal Pradesh, Assam State, Bihar, Chhattisgarh, Himachal Pradesh, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, Tripura, Uttarakhand, Uttar Pradesh, West Bengal); Java; Lesser Sunda Isl. (Bali); Myanmar (Chin, etc.); peninsular Malaysia; Nepal; Pakistan (Swat)
36 CHI 40 BAN BHU IND NEP PAK 41 MYA THA VIE 42 JAW LSI MLY PHI
Synonyms: hidden
= Aspidium erythrosorum var. souliei Christ; Bull. Soc. Bot. France 52(Mém. 1): 40 (1905)
= Arthrobotrys avara Wall.; Num. List no. 1034 (1828)
= Arthrobotrys macrocarpa Wall.; Num. List no. 395 (1828)
= Aspidium cochleatum (D.Don) Christ; Bull. Herb. Boiss. 6: 967 (1898), isonym
= Aspidium cochleatum (D.Don) Spreng.; Syst. 4: 106 (1827)
= Aspidium filix-mas var. cochleatum (D.Don) Christ; Farnkr. Erde 257 (1897)
= Dryopteris filix-mas var. cochleata (D.Don) Alderw.; Mal. Ferns 193 (1909)
= Dryopteris heleopteroides Christ; Philipp. J. Sci., Bot.2: 212 (1907)
= Lastrea cochleata (D.Don) T.Moore; Index Fil. (T. Moore) 88 (1857)
= Lastrea filix-mas var. cochleata (D.Don) Bedd.; Handb. Ferns Brit. India 250 (1883)
= Lastrea macrocarpa (Wall.) C.Presl; Tent. Pterid. 77 (1836)
= Nephrodium cochleatum D.Don; Prodr. Fl. Nepal. 6 (1825)
= Nephrodium filix-mas var. cochleatum (D.Don) Hook.; Sp. Fil. 4: 117 (1862)
species :
Dryopteris cognata (C.Presl) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Name: Large Kidney Fern (EN)
Distribution: St. Helena
28 STH
Conservation Status: IUCN E
Synonyms: hidden
= Aspidium cognatum (C.Presl) Fée; Gen. 292 (1850-52), nomen, ex Mett., Pheg. u. Asp. 61, n. 145 (1858)
= Lastrea cognata C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 40 (1851)
= Nephrodium cognatum (C.Presl) Hook.; Sp. 4: 123, t. 256 (1862)
= Dryopteris fuliginosa C.Chr.; Vid. Selsk. Skr. 8(6): 120, f. 29 (1920)
species :
Dryopteris commixta Tagawa; Acta Phytotax. 2: 190 (1933)
Distribution: China (Fujian, Guangxi, Jiangxi, Sichuan, Zhejiang); Japan
36 CHI 38 JAP
Synonyms: hidden
= Dryopteris shangqianensis Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(3): 20 (1984)
= Dryopteris sinodickinsii Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 46 (1990)
species :
Dryopteris comorensis (Tardieu) Fraser-Jenk.; Thaiszia 4(1): 34 (1994)
Distribution: Madagascar; Comores (Grande Comore)
29 COM MDG
Synonyms: hidden
= Dryopteris inaequalis var. comorensis Tardieu; Notul. Syst. (Paris) 15(2): 162 (1956)
species :
Dryopteris conjugata Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 63 (1941)
Distribution: China (Yunnan); India (Himachal Pradesh, Uttarakhand, Arunachal Pradesh, Meghalaya); Nepal; Myanmar
36 CHI 40 IND NEP 41 MYA
Synonyms: hidden
= Dryopteris subconjugata S.G.Lu; Guihaia 11(3): 224 (1991)
species :
Dryopteris conversa Alderw.; Bull. Jard. Bot. Buitenz. II, 20: 13 (1915)
Distribution: Sumatra
42 SUM
Remarks: provisionally accepted name
species :
Dryopteris corleyi Fraser-Jenk.; Bol. Soc. Broter. Ser. 2, 55: 248 (1982)
Distribution: N-Spain (Asturias)
12 SPA
Conservation Status: IUCN I
species :
Dryopteris costalisora Tagawa; Acta Phytotax. Geobot. 3: 88 (1934)
Distribution: Taiwan; E-Nepal; India (West Bengal, ?Arunachal Pradesh, Sikkim)
38 TAI 40 IND NEP
Remarks: bona sp. fide Fraser-Jenkins (1997) and Knapp (Ferns of Taiwan)
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris chrysocoma var. costalisora (Tagawa) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 2: 306 (1975)
species :
Dryopteris crassirhizoma Nakai; Cat. Sem. Hort. Bot. Univ. Imp. Tokyo 32 (1920)
Distribution: China (Hebei, Heilongjiang, Jilin, Liaoning); Japan; North Korea; South Korea (incl. Cheju Isl.); Russian Far East (Amur, Khabarovsk, Kuril Isl., Primorye, Sakhalin)
31 RFE 36 CHI 38 JAP KOR NKO
Synonyms: hidden
= Aspidium filix-mas var. maackii Milde; Fil. Eur. 122 (1867)
= Dryopteris buschiana Fomin; Fl. Sib. & Or. extr. 6: 52, fig. (1930)
= Dryopteris buschiana var. maackii (Milde) Fomin; Fl. Sib. & Or. extr. 5: 54 (1930)
= Dryopteris crassirhizoma var. maackii (Milde) Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 426 (1938)
= Dryopteris setosa Kudô; Medical Fl. Hokkaido no. 1
species :
Dryopteris crinalis (Hook. & Arn.) C.Chr.; Ind. 269 (1905)
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
63 HAW
Conservation Status: IUCN R
Synonyms: hidden
= Polypodium crinale Hook. & Arn.; Bot. Beech. Voy. 105 (1832)
= Phegopteris crinalis (Hook. & Arn.) Mann; Proc. Amer. Acad. 7: 218 (1868)
variant :
var. podosora (W.H.Wagner) D.D.Palmer; Amer. Fern J. 92(2): 100 (2002)
Distribution: Hawaii (Kauai)
63 HAW
Synonyms: hidden
= Dryopteris podosora W.H.Wagner & Flynn; Contr. Univ. Michigan Herb. 20: 252 (1995)
species :
Dryopteris crispifolia Rasbach, Reichst. & Vida; J. Linn. Soc., Bot. 74(3): 270 (1977)
Distribution: Azores (Sao Miguel Isl., Pico, Faial)
21 AZO
species :
Dryopteris cristata (L.) A.Gray; Man. 631 (1848)
Name: Kamm-Wurmfarn, Kammfarn (DE), Crested woodfern (EN)
Distribution: Austria; Belgium; England; Czech Republic; Slovakia; Denmark; Finland; France; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Netherlands; Hungary; Italy; Serbia & Kosovo; Slovenia; Norway; Poland; Romania; Sweden; Croatia; Estonia; Latvia; Lithuania; Belarus; Ukraine; N-European Russia; C-European Russia; W-European Russia; E-European Russia; Siberia (Altai, Krasnoyarsk, W-Siberia); Kazakhstan; USA (Alabama, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Idaho, Illinois, Indiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Montana, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Tennessee, Virginia, Vermont, Washington State, Wisconsin, West Virginia); Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon
10 DEN FIN GRB NOR SWE 11 AUT BGM CZE GER HUN LIE NET POL SLO SWI 12 FRA 13 CRO ITA ROM SLV SRB 14 BLR EST LAT LIT RUS UKR 30 SIB 32 KAZ 72 SPM 73 USA 74 USA 75 USA 78 USA
Synonyms: hidden
= Acrostichum callipteris (Ehrh.) Ehrh.; Beitr. 8: 77 (1788)
= Aspidium cristatum (L.) Sw.; Schrad., J. Bot. 1800(2): 37 (1801)
= Aspidium cristatum var. crenatum Christ; 145 (1900)
= Aspidium lancastriense Spreng.; Anleit. Kenntn. Gew. 3: 134 (1804)
= Dryopteris affinis var. cristata (T.Moore) Fraser-Jenk.; Indian J. Forestry 43(1): 83 (2020)
= Dryopteris cristata var. lancastriensis (Spreng.) Tidestr.; Elys. Marian., Ferns 1: 44, pl. 6 (1906)
= Dryopteris cristata var. mariana Tidestr.; Elys. Marian., Ferns 1: 44, pl. 6 (1906)
= Filix cristata (L.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix-Mas cristata (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Lastrea callipteris (Ehrh.) Newman; Hist. br. Ferns ed. II: 12 (1844)
= Lastrea cristata (L.) C.Presl; Tent. Pterid. 77 (1836)
= Lastrea lancastriensis (Spreng.) J.Sm.; Cat. Kew Ferns 6 (1856)
= Lophodium callipteris (Ehrh.) Newman; Phyt. 4: 421 (1851)
= Nephrodium cristatum (L.) Michx.; Fl. bor. Amer. 2: 269 (1803)
= Nephrodium cristatum f. lancastriense (Spreng.) Gilbert; List N. Amer. Pterid. 17 (1901)
= Nephrodium lancastriense (Spreng.) Desv.; Mém. Soc. Linn. Paris 6(3): 260 (1827)
= Polypodium callipteris Ehrh.; Hannov. Mag. 8-9: 127, 138 (1784)
= Polypodium cristatum L.; Sp. Pl. 2: 1090 (1753) (basion.)
= Polystichum callipteris (Ehrh.) Bernh.; Schrad., J. Bot. 1799(1): 307 (1799)
= Polystichum cristatum (L.) Roth; Tent. Fl. Germ. 3: 84 (1799), Röm., Arch. 2(1): 106 (1799)
= Tectaria cristata (L.) Lag., Garcia & Clem.; Anal. Ci. Nat. 5: 147 (1802)
= Thelypteris cristata (L.) Nieuwl.; Midl. Naturalist 1: 226 (1910)
species :
Dryopteris cycadina (Franch. & Sav.) C.Chr.; Ind. 260 (1905)
Distribution: China (Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan; Japan; South Korea
36 CHI 38 JAP KOR TAI
Synonyms: hidden
= Aspidium cycadinum Franch. & Sav.; Enum. Pl. Jap. 2: 236 (1879)
= Dryopteris fengyangshanensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 33 (1983)
= Dryopteris longirostrata Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 47 (1990)
= Dryopteris pseudatrata Ching; Fl. Fujianica 1: 601 (1982)
= Dryopteris rigidiuscula Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 47 (1990)
= Dryopteris cycadina var. melanolepis Nakai; Bot. Mag. (Tokyo) 45: 99 (-100) (1931)
species :
Dryopteris cyclopeltidiformis C.Chr.; Dansk Bot. Ark. 9(3). 60, t. 6(3-4) (1937)
Distribution: China (Hainan); Vietnam
36 CHI 41 VIE
species :
Dryopteris damingshanensis Li Bing Zhang & H.M.Liu; Novon 23(1): 119 (2014)
Distribution: China (Guangxi)
36 CHI
species :
Dryopteris daozhenensis P.S.Wang & X.Y.Wang; Pterid. Fl. Guizhou 310 (2001)
Distribution: China (Guizhou)
36 CHI
Remarks: provisionally accepted name, status uncertain
species :
Dryopteris decipiens (Hook.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Jiangxi, Hunan, Sichuan, Zhejiang); Taiwan; Japan; Ryukyu Isl.
36 CHI 38 JAP NNS TAI
Synonyms: hidden
= Aspidium decipiens (Hook.) Luerss.; Engl. Jahrb. 4: 360 (1883)
= Nephrodium decipiens Hook.; Sp. 4: 86, t. 243 (1862)
variant :
var. diplazioides (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 476 (1938)
Distribution: China (Anhui, Fujian, Guizhou, Jiangsu, Jiangxi, Sichuan, Zhejiang); Taiwan; Japan; South Korea
36 CHI 38 JAP KOR TAI
Remarks: included in type variety by Knapp (Ferns of Taiwan)
Synonyms: hidden
= Dryopteris cordipinna Ching & K.H.Shing; Fl. Fujianica 1: 599 (1982)
= Dryopteris decipiens var. subbipinnata Ching; Wuyi Sci. J. 1: 8 (1981)
= Dryopteris fuscipes var. diplazioides (Christ) Ching; Fl. Jiangsuensis 1: 66 (1977), nom. nud. ex L. K. Ling, Q. Q. Zhang & Y. R. Hwang (1991)
= Dryopteris integripinnula Ching; Wuyi Sci. J. 1: 7 (1981)
= Dryopteris metafuscipes Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 19 (1983)
= Dryopteris mimetica Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 21 (1983)
= Dryopteris retrorsopaleacea Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 20 (1983)
= Polystichum diplazioides Christ; Bull. Ac. Géogr. Bot. Mans 260 (1902)
species :
Dryopteris dehuaensis Ching & K.H.Shing; Fl. Fujianica 1: 601 (1982)
Distribution: China (Fujian, Jiangxi, Zhejiang)
36 CHI
Synonyms: hidden
= Dryopteris gushanica Ching & K.H.Shing; Fl. Fujianica 1: 601 (1982)
= Dryopteris neosordidipes Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990)
species :
Dryopteris deparioides (T.Moore) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
subspecies :
subsp. deparioides
Distribution: +S-India (+Tamil Nadu); Sri Lanka
40 IND SRL
Conservation Status: IUCN CR or EX for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium deparioides (T.Moore) Hook.; Fil. Exot. t. 3 (1858)
= Diclisodon deparioides T.Moore; Index Fil. (T. Moore) 96 (1857)
= Lastrea deparioides (T.Moore) Bedd.; Ferns South. India 104 (1865)
= Nephrodium deparioides (T.Moore) Hook.; Fil. Exot. t. 53 &, t. 60 (1859) in obs.
subspecies :
subsp. ambigua (Sledge) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 447 (1989)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Dryopteris ambigua Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 5: 23 (1973)
subspecies :
subsp. concinna [Bedd.] C.Chr.; Ind. Fil., Suppl. 3: 84 (1934)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Aspidium concinnum Thwaites; Enum. pl. Zeyl. 392 (1864) (non Link (1833))
= Dryopteris emigrans Copel.; Univ. Calif. Publ. Bot. 12: 392 (1931)
= Dryopteris thwaitesii (Baker) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
= Lastrea concinna Bedd.; Handb. Ferns Brit. India, t. 207 (1864) (non L. concinna (Willd.) T. Moore)
= Lastrea thwaitesii (Baker) Bedd.; Ferns br. Ind. Suppl. 18. (1876)
= Nephrodium thwaitesii Baker; Syn. 277 (1867)
species :
Dryopteris devriesei Rosenst.; Meded. Rijks Herb. no. 31: 1 (1917)
Distribution: Java
42 JAW
Remarks: provisionally accepted name
species :
Dryopteris diacalpioides (Ching) Li Bing Zhang; Taxon 61(6): 1206 (2012)
Distribution: China (S-Yunnan)
36 CHI
Synonyms: hidden
= Nothoperanema diacalpioides Ching; Acta Phytotax. Sin. 11: 28 (1966)
species :
Dryopteris dickinsii (Franch. & Sav.) C.Chr.; Ind. 262 (1905)
Distribution: China (Anhui, Fujian, Guangxi, Guizhou, Hubei, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan; South Korea (Cheju Isl.); NW-India (Jammu & Kashmir); Pakistan (Azad Kashmir); Japan
36 CHI 38 JAP KOR TAI 40 IND PAK
Conservation Status: IUCN CR or EX for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium dickinsii Franch. & Sav.; Enum. Pl. Jap. 2: 236, 621 (1879)
= Aspidium thibeticum Franch.; Nouv. Arch. Mus. II, 10: 118 (1887)
= Dryopteris basiaurita Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 405 (1938)
= Dryopteris hirtipes var. japonica Nakai; Bot. Mag. (Tokyo) 45: 100 (1931)
= Dryopteris okushirensis Miyabe & Kudô; Tr. Sapporo Nat. Hist. Soc. 7. 23 (1918)
= Dryopteris thibetica (Franch.) C.Chr.; Ind. 298 (1905) (fide Fraser-Jenkins et al. 2018)
= Dryopteris yungtzeensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 60 (1941) (fide Fraser-Jenkins et al. 2018)
= Lastrea thibetica (Franch.) Bedd.; Handb. Suppl. 52 (1892)
= Nephrodium dickinsii (Franch. & Sav.) Baker; Hook., Ic. Pl. t. 1651 (1886)
= Nephrodium thibeticum (Franch.) Baker; Ann. Bot. 6: 318 (1891)
= Thelypteris thibetica (Franch.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 57 (1974)
species :
Dryopteris dicksonioides (Mett. ex Kuhn) Copel.; Bull. Bernice P. Bishop Mus. 93: 35 (1932)
Distribution: Society Isl. (Tahiti, Raiatea, Moorea, Huahine)
61 SCI
Synonyms: hidden
= Phegopteris dicksonioides Mett. ex Kuhn; Linn. 36: 118 (1869)
= Thelypteris dicksonioides (Mett. ex Kuhn) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)
species :
Dryopteris diffracta (Baker) C.Chr.; Ind. 262 (1905)
Distribution: China (Guangxi, Guizhou, Hainan, Yunnan); Tibet; Taiwan; NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.); Vietnam; N-Thailand
36 CHI TIB 38 TAI 40 IND 41 MYA THA VIE
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Arachniodes diffracta (Baker) Ching; Acta Bot. Sin. 10: 257 (1962)
= Aspidium diffractum (Baker) Christ; Bull. Herb. Boiss. 7: 17 (1899)
= Acrorumohra diffracta (Baker) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 101 (1939)
= Dryopteris reflexipinna Hayata; Ic. Pl. Formos. 4: 174, f. 113 (1914)
= Nephrodium diffractum Baker; Bull. Misc. Inform. Kew 1898: 230 (1898)
= Polystichum diffractum (Baker) Masam.; Short Fl. Formos. 26 (1936)
= Rumohra diffracta (Baker) Ching; Sinensia 6: 69, t. 18 (1934)
species :
Dryopteris dilatata (Hoffm.) A.Gray; Man. 631 (1848)
subspecies :
subsp. dilatata
Name: Breitblättriger Wurmfarn, Breitblättriger Dornfarn (DE)
Distribution: Austria; Belgium; Luxembourg; England; Channel Isl.; Wales; Scotland; Bulgaria; Corsica; Czech Republic; Slovakia; Denmark; Faeroe Isl.; Finland; France; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Greece; Ireland; Northern Ireland; Switzerland; Liechtenstein; Netherlands; Spain; Hungary; Italy; Bosnia & Hercegovina; Croatia; Montenegro; Macedonia; Serbia & Kosovo; Slovenia; Portugal; Norway; Poland; Romania; Sweden; Turkey (Anatolia); N-European Russia; C-European Russia; W-European Russia; Estonia; Latvia; Lithuania; Belarus; Ukraine; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Siberia (Altai); Japan; Mongolia; Falkland Isl. [I]; Argentina [I] (Tierra del Fuego [I]); New Zealand [I]; Azores (Pico)
10 DEN FIN FOR GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 CIS COR FRA POR SPA 13 BIH BUL CRO GRC ITA MNT NMA ROM SLV SRB 14 BLR EST LAT LIT RUS UKR 21 AZO 30 SIB 33 ARM AZE GEO NCS 34 TUR 37 MON 38 JAP 51 nzl 85 arg 91 fal
Remarks: carthusiana agg.
Synonyms: hidden
= Aspidium dilatatum (Hoffm.) Sm.; Fl. Brit. 8: 1125 (1804)
= Aspidium dilatatum var. deltoideum (Milde) Borbás; 449 (1877)
= Aspidium dilatatum var. oblongum (Milde) Borbás; 449 (1877)
= Aspidium drepanum Schkuhr; Kr. Gew. 1: t. 47 (1806)
= Aspidium dumetorum Sm.; Engl. Fl. 4: 281 (1828)
= Aspidium spinulosum f. deltoideum (Milde) Zapal.; 79 (1905)
= Aspidium spinulosum f. oblongum (Milde) Zapal.; 79 (1905)
= Aspidium spinulosum f. recurvatum (Lasch) Asch.; 926 (1864)
= Aspidium spinulosum subsp. recurvatum Lasch; 80 (1861)
= Aspidium spinulosum var. deltoideum Milde; 57 (1865)
= Aspidium spinulosum var. dilatatum (Hoffm.) Hampe; 63 (1836)
= Aspidium spinulosum var. muticum A.Braun ex Döll; 18 (1843)
= Aspidium spinulosum var. oblongum Milde; 57 (1865)
= Aspidium tanacetifolium (Lam. & DC.) Opiz; Kratos 2: 10 (1820)
= Dryopteris alexeenkoana Fomin; Kuzn. & al., Fl. Caucas. Crit. l(1): 67 (1911), et Moniteur Jard. Bot. Tiflis, livr. 20: 46 (1911)
= Dryopteris aristata (Vill.) Druce; List Brit. Pl. 87 (1908)
= Dryopteris austriaca (Jacq.) Woyn.; 339 (1915), nom. illeg.
= Dryopteris austriaca f. deltoidea (Milde) Tavel; 73 (1918)
= Dryopteris austriaca f. oblonga (Milde) Tavel; 74 (1918)
= Dryopteris austriaca subsp. dilatata (Hoffm.) Schinz & Thell.; 340 (1915)
= Dryopteris austriaca var. dilatata Hoffm.; (1943)
= Dryopteris carthusiana subsp. dilatata (Hoffm.) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans: 1213 (1990)
= Dryopteris dilatata f. deltoideum (Milde) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)
= Dryopteris dilatata f. oblongum (Milde) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)
= Dryopteris dilatata subsp. alexeenkoana (Fomin) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)
= Dryopteris dilatata subsp. remotiloba (Fomin) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)
= Dryopteris dilatata var. remotiloba Fomin
= Dryopteris spinulosa subsp. austriaca Bergdolt; 29 (1936)
= Dryopteris spinulosa subsp. dilatata (Hoffm.) C.Chr.; Ind. Fil. 293 (1905)
= Dryopteris spinulosa var. dilatata (Hoffm.) Underw.; Native Ferns, ed. 4: 116 (1893)
= Filix spinulosa var. dilatata (Hoffm.) Farw.; Ann. Rep. Mich. Acad. Sci 6: 209 (1904)
= Lastrea aristata (Vill.) Britton & Rendle; List Brit. Pl. 39 (1907)
= Lastrea collina Newman; Hist. br. Ferns ed. II: 224 (1844)
= Lastrea dilatata (Hoffm.) C.Presl; Tent. Pterid. 77 (1836)
= Lastrea glandulosa (Newman) Newman; Phytol. 4: 258 (1851)
= Lastrea lepidota T.Moore; Handb. Brit. Ferns ed. III: 130 (1857)
= Lastrea maculata Deakin; Fl. Brit. 4: 110 (1848)
= Lastrea multiflora (Roth) Newman; Hist. br. Ferns ed. II: 215 (1844)
= Lophodium collinum (Newman) Newman; Phyt. 4: app. 18 (1851)
= Lophodium glandulosum Newman; Phyt. 4: app. 18 (1851)
= Lophodium multiflorum (Roth) Newman; Phyt. 4: 371 (1851)
= Nephrodium austriacum (Jacq.) Fritsch; Mitt. Naturw. Ver. Steiermark 46 (1908)
= Nephrodium dilatatum (Hoffm.) Desv.; Mem. Soc. Linn. Paris 6, 2: 261 (1827)
= Nephrodium spinulosum subsp. dilatatum (Hoffm.) Arcang.; Comp. Fl. It. 811 (1882)
= Nephrodium spinulosum subsp. dilatatum (Hoffm.) Hook.fil.; (1870)
= Nephrodium spinulosum var. dilatatum (Hoffm.) F.W.Schultz; 566 (1845)
= Nephrodium spinulosum var. dumetorum (Sm.) Hook.f.; 466 (1870)
= Nephrodium tanacetifolium (Lam. & DC.) Rouy; Fl. France 14: 413 (1913)
= Polypodium aristatum Vill.; Hist. pl. Dauph. 3: 844 (1789) (non G. Forst. (1786))
= Polypodium dilatatum Hoffm.; Deutschl. Fl. 2: 7 (1796) (basion.)
= Polypodium filix-femina var. cristata Weiss; 317 (1770)
= Polypodium lanceolata-cristatum Hoffm.; Bot. Mag. (Römer & Usteri) 9: 9 (1790)
= Polypodium multiflorum Roth; Catal. Bot. 1: 135 (1797)
= Polystichum dilatatum (Hoffm.) Schumacher; Enum. Pl. Saell. 2: 24 (1803)
= Polystichum multiflorum (Roth) Roth; Tent. Fl. Germ. 3: 87 (1799), Röm., Arch. 2(1): 106 (1799)
= Polystichum spinulosum subsp. dilatatum (Hoffm.) F.Aresch.; 552 (1881)
= Polystichum spinulosum subsp. dilatatum (Hoffm.) P.Fourn.; Quatre Fl. France 17 (1934), isonym
= Polystichum spinulosum var. dilatatum (Hoffm.) W.D.J.Koch; 979 (1845)
= Polystichum spinulosum var. muticum (A.Braun ex Döll) Schur; 168 (1851)
= Polystichum tanacetifolium Lam. & DC.; Fl. franc. 2: 562 (1805)
= Thelypteris dilatata (Hoffm.) House; New York State Mus. Bull. 233-234. 69 (1920)
= Thelypteris spinulosa var. dilatata (Hoffm.) H.St.John; Prelim. List Pl. Kaniksu Natl. Forest 1: 1 (1925)
subspecies :
subsp. lusitanica Fraser-Jenk., Andreis & Vasco Silva; Indian J. Forestry 43(1): 82 (2020), et Indian J. Forest. 43(1): 82 (2021)
Distribution: Portugal; France; Spain
12 FRA POR SPA
species :
Dryopteris dracomontana Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 147 (1982)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov., northeastern W-Cape Prov., Free State); Lesotho
27 LES SAF
Synonyms: hidden
= Aspidium inaequale var. montanum Kunze; Linnaea 10: 549 (1836)
= Dryopteris pentheri var. montana (Kunze) Alston; Bol. Soc. Broter., ser. 2, 30: 14 (1956)
species :
Dryopteris drakei C.Chr.; Ind. 262 (1906)
Distribution: Society Isl. (Tahiti)
61 SCI
Synonyms: hidden
= Polypodium vescoi Drake; Fl. Polyn. franc. 307 (1893) (non Nephrodium vescoi Drake (1893))
species :
Dryopteris dulongensis (S.K.Wu & X.Cheng) Zhang; Phytotaxa 71: 17-27 (2012)
Distribution: China (Yunnan)
36 CHI
Remarks: Possibly a hybrid D. apiciflora x heterolaena. Subgen. Dryopsis
Synonyms: hidden
= Dryopsis ×dulongensis (S.K.Wu & X.Cheng) S.Y.Dong; Acta Bot. Yunnan. 23: 187 (2001)
= Ctenitis dulongensis S.K.Wu & X.Cheng; Acta Phytotax. Sin. 23: 402 (1985)
species :
Dryopteris edwardsii Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 139 (1997)
Distribution: India (Uttarakhand, Himachal Pradesh, Jammu & Kashmir); Bhutan; Pakistan; Nepal; Taiwan; Tibet
36 TIB 38 TAI 40 BHU IND NEP PAK
Remarks: not mentioned in Flora of China but recorded for Taiwan by Knapp (Ferns of Taiwan, 2011 ff.)
species :
Dryopteris enneaphylla (Baker) C.Chr.; Ind. 263 (1905)
Distribution: Taiwan; China (Zhejiang, Hubei)
36 CHI 38 TAI
Synonyms: hidden
= Nephrodium enneaphyllum Baker; J. Bot. 170 (1887)
species :
Dryopteris erythrosora (D.C.Eaton) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Name: Autumn fern
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; South Korea (incl. Cheju Isl.); Ryukyu Isl.
36 CHI 38 JAP KOR NNS
Synonyms: hidden
= Aspidium cystolepidotum Miq.; Ann. Mus. Bot. Lugd.-Bat. 3: 177 (1867)
= Aspidium erythrosorum D.C.Eaton; Perry, Narr. Exp. to China 2: 330 (1856)
= Aspidium filix-mas var. erythrosorum (D.C.Eaton) Christ; Farnkr. Erde 257 (1897)
= Aspidium prolificum Maxim.; Franch. & Sav., Enum. Pl. Jap. 2: 239, 632 (1879)
= Dryopteris bulligera Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 483 (1938)
= Dryopteris cystolepidota (Miq.) C.Chr.; Ind. 260 (1905)
= Dryopteris distantipinna Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 4(3): 27, f. 27 (1984)
= Dryopteris erythrosora (D.C.Eaton) C.Chr.; Ind. 260 (1905)
= Dryopteris erythrosora f. viridisora (Nakai & H.Itô) H.Itô; Nov. Fl. Jap. 4: 41 (1939)
= Dryopteris erythrosora var. dilatata (Koidz.) Sugim.; (1962)
= Dryopteris erythrosora var. viridisora Nakai ex H.Itô; Bot. Mag. (Tokyo) 50: 69 (1936)
= Dryopteris linyingensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 30 (1983)
= Dryopteris nipponensis Koidz.; Mayebara, Fl. Austro-Higoensis 5 (1931), Acta Phytotax. 1: 28 (1932)
= Dryopteris oblongipinnula Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 5 (1987)
= Dryopteris oppositipinna Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(3): 25 (1984), nom. illeg., non (Hayata) Hayata (1911)
= Dryopteris paraerythrosora Ching & C.F.Zhang; Bull. Bot. Res., Harbin 3: 27 (1983)
= Dryopteris pseudoerythrosora Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 27 (1983), nom. illeg., non Kodama 1913
= Dryopteris remotipinnula Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 15 (1983), non Bonap. 1925
= Dryopteris sinoerythrosora Ching & K.H.Shing; Fl. Fujianica 1: 600 (1982)
= Dryopteris squamistipes Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 7 (1984)
= Dryopteris taquetii Christ; Fedde, Repert. Spec. Nov. 5: 284 (1908)
= Lastrea erythrosora (D.C.Eaton) T.Moore; Ind. 91 (1858)
= Lastrea prolifica (Maxim.) T.Moore; Gard. Chr. n. s. 18: 744 (1882)
= Nephrodium erythrosorum (D.C.Eaton) Hook.; Sp. 4: 120, t. 253 (1862)
= Nephrodium filix-mas var. erythrosorum (D.C.Eaton) Christ ex Matsum.; Ind. Pl. Jap. 1: 319 (1904)
= Nephrodium prolificum (Maxim.) Diels; Nat. Pfl. 1(4): 173 (1899)
= Nephrodium prolificum (Maxim.) G.Nicholson; Dict. Gard. 4: 573 (1888)
species :
Dryopteris erythrovaria K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(2): 86 (2018)
Distribution: China (Jiangsu, Zhejiang, Anhui); Japan (Honshu, Kyushu); South Korea
36 CHI 38 JAP KOR
species :
Dryopteris esterhuyseniae Schelpe & N.C.Anthony; Contr. Bolus Herb. 10: 148 (1982)
Distribution: South Africa (W-KwaZulu-Natal: Drakensberg)
27 SAF
Remarks: included by Roux (2009) in D. dracomontana but kept separate by Ferns of South Africa
species :
Dryopteris expansa (C.Presl) Fraser-Jenk. & Jermy; Fern Gaz. 11(5): 338 (1977)
Name: Feingliedriger Wurmfarn (DE), Spreading woodfern (EN)
Distribution: Iceland; England; Wales; Scotland; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, SLf Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Hungary; Czech Republic; Slovakia; Norway; Sweden; Andorra; Portugal; Spain; France; Corsica; Italy; Serbia & Kosovo; Slovenia; Croatia; Montenegro; Bulgaria; Greece (mostly in mountains); N-European Russia; C-European Russia; Estonia; Latvia; Lithuania; Belarus; Ukraine; Northern Caucasus; Georgia [Caucasus]; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); Japan; China (Hebei, Heilongjiang, Jilin, Liaoning); North Korea; South Korea; Taiwan; Mongolia; Turkey (Anatolia); Alaska; USA (California, Colorado, Idaho, Michigan, Minnesota, Montana, Oregon, Washington State, Wisconsin, Wyoming); Canada (Alberta, British Columbia, Labrador, Newfoundland, Northern Territories, Ontario, Québec, Yukon); Greenland
10 GRB ICE NOR SWE 11 CZE GER HUN LIE SLO SWI 12 ADO COR FRA POR SPA 13 BUL CRO GRC ITA MNT SLV SRB 14 BLR EST LAT LIT RUS UKR 30 SIB 31 RFE 33 GEO NCS 34 TUR 36 CHI 37 MON 38 JAP KOR NKO TAI 70 ASK GNL 73 USA 74 USA 75 USA
Remarks: carthusiana agg.
Synonyms: hidden
= Aspidium spinulosum var. oblongum Milde; Fil. Eur. & Atl. 137 (1867)
= Dryopteris assimilis S.Walker; Amer. J. Bot. 48: 607 (1961)
= Dryopteris austriaca subsp. assimilis (S.Walker) O.Bolòs & Vigo; Fl. Paisos Catalans: 170 (1984)
= Dryopteris carthusiana subsp. assimilis (S.Walker) O.Bolòs, Vigo, Masalles & Ninot; Fl. Man. Paisos Catalans: 1213 (1990)
= Dryopteris expansa subsp. assimilis (S.Walker) Tzvelev; Sosud. Rast. Sovet. Dal´nego Vostoka 5: 54 (1991)
= Dryopteris expansa var. alpina (T.Moore) Viane; 90 (1986)
= Dryopteris extremiorientalis V.N.Vassil.; nom. inval.
= Dryopteris manshurica Ching ex Wang & al.; Liou, Herb. Fl. N.-E. China 1: 69 (1958)
= Dryopteris minimisora Nakai; Bull. Natl. Sci. Mus. (Tokyo) 27: 19 (1949)
= Dryopteris morrisonensis (Hayata) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30. 450 (1911), Ic. Fl. Taiwan 5. 281 f. 108 (1915)
= Dryopteris siranensis Nakai; Bull. Natl. Sci. Mus. (Tokyo) 33: 3 (1953)
= Dryopteris spinulosa subsp. assimilis (S.Walker) Schidlay; Fl. Slovenska 2: 217 (1966)
= Dryopteris spinulosa var. morrisonensis Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 450 (1911) [fide Fraser-Jenkins]
= Lastrea expansa (C.Presl) C.Presl; Tent. Pterid. 76 (1836)
= Nephrodium expansum C.Presl; Reliq. Haenk. 1: 38 (1825) (basion.)
variant :
var. alpina (T.Moore) Viane; Pl. Syst. Evol. 153(1-2): 90 (1986)
Distribution: England
10 GRB
Synonyms: hidden
= Lastrea dilatata var. alpina T.Moore; Ferns Brit. & Ireland Nature-Printed, sub t. 22 (1856) (non Dryopteris alpina Rosenst.)
variant :
var. willeana (Lid) Elven; Nordic J. Bot. 16(1): 4 (1996)
Distribution: Scandinavia
Synonyms: hidden
= Dryopteris austriaca var. willeana Lid
species :
Dryopteris fadenii Pic.Serm.; Webbia 37(2): 333 (1984)
Distribution: Ethiopia; Uganda; Kenya; Tanzania
24 ETH 25 KEN TAN UGA
species :
Dryopteris fangii Ching ex Fraser-Jenk. & Z.R.Wang; Acta Phytotax. Sin. 23(5): 349 (1985), nom. nov.
Distribution: NE-India (Arunachal Pradesh); China (Sichuan, etc.); Tibet
36 CHI TIB 40 IND
Remarks: included in D. chrysocoma by Flora of China but Fraser-Jenkins et al. (2018) keep it separate
Synonyms: hidden
= Dryopteris chrysocoma var. gracilis Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 437 (1938) (non (T. Moore ex Bedd.) Ching)
species :
Dryopteris fatuhivensis E.D.Br.; Bull. Bernice P. Bishop Mus. 89 [Fl. S. E. Polynesia II]: 27, t. 8 (1931) (fide Fraser-Jenkins et al. 2018)
Distribution: Marquesas Isl. (Nuku Hiva, Fatu Hiva, Ua Huka, Ua Pou, Hiva Oa); Cook Isl. (Rarotonga)
61 COO MRQ
Remarks: belongs possibly to D. hirtipes
Synonyms: hidden
= Thelypteris fatuhivensis (E.D.Br.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)
species :
Dryopteris filipaleata J.P.Roux; Bothalia 34(1): 28 (2004)
Distribution: Kenya; Tanzania
25 KEN TAN
species :
Dryopteris filix-mas (L.) Schott; Gen. Fil. tab. 9 (1834)
subspecies :
subsp. filix-mas
Name: Gewöhnlicher Wurmfarn, Männlicher Wurmfarn (DE), Male Fern (EN)
Distribution: Albania; Austria; Belgium; Luxembourg; England; Channel Isl.; Wales; Scotland; Bulgaria; Corsica; Czech Republic; Slovakia; Denmark; Finland; France; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Greece; Ireland; Northern Ireland; Switzerland; Liechtenstein; Netherlands; Spain; ?Andorra; Hungary; Italy; San Marino; Iceland; Bosnia & Hercegovina; Montenegro; Croatia; Macedonia; Serbia & Kosovo; Slovenia; Portugal; Malta; Norway; Poland; Romania; ?Sardinia; Sicily; Sweden; European Turkey; Estonia; Latvia; Lithuania; Crimea; N-European Russia; C-European Russia; W-European Russia; E-European Russia; Belarus; Ukraine; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Siberia (Altai, Buryatia, Chita, Irkutsk, Tuva, W-Siberia, Yakutia); Tajikistan; China (Xinjiang); Kazakhstan; Kyrgyzstan; Uzbekistan; Pakistan (Dir, Chitral, Swat, Hazara, Baltistan); NW-India (Jammu & Kashmir: Ladakh); E-Afghanistan; Turkey (Anatolia); Iran; Tunisia; Algeria; Morocco; North America; New Zealand [I]; Chatham Isl. [I]; Chile [I] (Los Lagos [I]); Argentina [I] (Neuquen [I])
10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 CIS COR FRA POR SPA 13 ALB BIH BUL CRO GRC ITA MLT MNT NMA ROM SIC SLV SMA SRB TUE 14 BLR EST KRY LAT LIT RUS UKR 20 ALG MOR TUN 30 SIB 32 KAZ KGZ TZK UZB 33 ARM AZE GEO NCS 34 AFG IRN TUR 36 CHI 40 IND PAK 51 ctm nzl 85 arg cle
Remarks: filix-mas agg.
Synonyms: hidden
= Aspidium abbreviatum Poir.; Encycl. suppl. 4: 516 (1816)
= Aspidium depastum Schkuhr; Kr. Gew. 1: 50, t. 51 (1806)
= Aspidium erosum Schkuhr; Kr. Gew. 1: 46, t. 45 (1806)
= Aspidium expansum Dietr.; Fl. Jenensis (1826)
= Aspidium filix-mas (L.) Sw.; Schrad., J. Bot. 1800(2) 38 (1801)
= Aspidium filix-mas f. crenatum Milde; 51 (1865)
= Aspidium filix-mas f. erosum (Döll) Zapal.; 78 (1905)
= Aspidium filix-mas f. heleopteris (Borkh.) Milde; 52 (1865)
= Aspidium filix-mas var. crenatum Milde
= Aspidium filix-mas var. erosum Döll; 16 (1843)
= Aspidium filix-mas var. heleopteris (Borkh.) Milde; 121 (1867)
= Aspidium filix-mas var. subintegrum Döll; 27 (1855)
= Aspidium mildeanum Göpp.; Denkschr. 50jähr. Best. vaterl. Ges. Schles. 193 (1853)
= Aspidium nemorale (Salisb.) Gray; Nat. Arr. br. Fl. 2 (1821)
= Aspidium opizii Wierzb.; Flora 379 (1840)
= Aspidium umbilicatum (Poir.) Desv.; Berl. Mag. 5: 320 (1811)
= Aspidium veselskii Hassl. ex Domin; Vestn. Kral. Ceske Spolecn. Nauk, Tr. Mat.-Prir. 1941(21): 4 (1942)
= Dryopteris filix-mas f. crenatum (Milde) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)
= Dryopteris filix-mas var. recurva (Francis) Rothm.; 97 (1946)
= Dryopteris patagonica Diem; Darwiniana 12: 70, f. 2 (1960)
= Dryopteris ×bohemica Domin; Vestn. Kral. Ceske Spolecn. Nauk, Tr. Mat.-Prir. 1941(21): 2-3 (1942)
= Dryopteris ×digenea Domin; Preslia 13-15: 8 (1936)
= Filix filix-mas (L.) Farw.; Rep. (Annual) Michigan Acad. Sci. 6: 201 (1904)
= Filix pinnata Gilib.; Exerc. Phyt. 2: 557 (1792)
= Filix-Mas filix-mas (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Filix-Mas vulgaris Hill; Br. Herbal 527, t. 74 (1756)
= Lastrea erosa (Schkuhr) Deakin; Fl. Brit. 4. 101 (1848)
= Lastrea filix-mas (L.) C.Presl; Tent. Pterid. 76 (1836)
= Lastrea officinalis Bubani; Fl. Pyr. 4: 433 (1901)
= Lastrea propinqua Woll. ex Lowe; Our nat. Ferns 1: 234, 280 (1865)
= Lophodium erosum (Schkuhr) Newman; Phyt. 4: app. 21 (1851)
= Lophodium filix-mas (L.) Newman; Phyt. 4: app. 20 (1851)
= Nephrodium crenatum Stokes; Bot. Mat. Med. 4: 606 (1812), based on Polystichum filix-mas (L.) Roth, nom. superfl.
= Nephrodium filix-mas (L.) Rich.; Marthe, Cat. Pl. Jard. Med. Paris 129 (1801)
= Nephrodium filix-mas (L.) Strempel; Fil. Berol. Syn.: 30 (1822)
= Nephrodium filix-mas var. crenatum (Milde) Murr; 8 (1923)
= Nephrodium filix-mas var. deorsilobatum (T.Moore) Murr; 8 (1923)
= Nephrodium filix-mas var. erosum (Schkuhr) Murr; 8 (1923)
= Nephrodium filix-mas var. subintegrum (Döll) Murr; 8 (1923)
= Nephrodium propinquum (Woll. ex Lowe) Lowe; Brit. Ferns 133 (1891)
= Nephrodium rigidoforme Rouy; Fl. France 14: 408 (1913)
= Nephrodium umbilicatum (Poir.) Desv.; Mem. Soc. Linn. Paris 6, 2: 260 (1827)
= Polypodium filix-mas L.; Sp. Pl. 2: 1090 (1753)
= Polypodium heleopteris Borkh.; Röm., Arch. 1(3): 19 (1798)
= Polypodium mas L.; Mant. Pl. (2): 506 (1771)
= Polypodium nemorale Salisb.; Prod. 403 (1796), nom. nov. for P. filix-mas L., nom. superfl.
= Polypodium umbilicatum Poir.; Encycl. 5: 528 (1804)
= Polystichum callipteris Bernh.; Schrad., J. Bot. 1799(1): 307 (1799) (p. p.)
= Polystichum filix-mas (L.) Roth; Tent. Fl. Germ. 3: 82 (1799)
= Polystichum filix-mas var. erosum (Döll) Sendtn.; 907 (1854)
= Polystichum obtusum Dulac; Fl. Dépt. Hautes-Pyrenees 33 (1867)
= Polystichum patentissimum Trevis.; Syll. Sporoph. Ital. 18 (1874)
= Polystichum polysorum Tod.; Giorn. sci. nat. Palermo 1: 240 (1866)
= Tectaria filix-mas (L.) Cav.; Descr. Pl. 251 (1802)
= Thelypteris filix-mas (L.) Nieuwl.; Midl. Naturalist 1: 226 (1910)
subspecies :
subsp. brittonii Fraser-Jenk. & Widén; Advances Forest. Res. India 29: 139-160 (2006)
Distribution: USA (Arizona, California, Colorado, Idaho, Illinois, Maine, Michigan, Montana, New Hampshire, New Mexico, Nevada, New York, Oklahoma, Oregon, Pennsylvania, Rhode Island, South Dakota, Texas, Utah, Vermont, Washington State, Wisconsin, Wyoming); Canada (Alberta, British Columbia, New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec, Saskatchewan); S-Greenland; St. Pierre et Miquelon; Mexico (Coahuila)
70 GNL 72 SPM 73 USA 74 USA 75 USA 76 USA 77 USA 79 MEX
subspecies :
subsp. pseudorigida (Christ) Askerov; Zametki Sist. Geogr. Rast. 39: 6 (1983)
Distribution: European Russia
14 RUS
Synonyms: hidden
= Dryopteris filix-mas var. pseudorigida Christ
species :
Dryopteris flaccisquama A.Rojas; Revista Biol. Trop. 49(2): 436 (2001)
Distribution: Costa Rica; Panama; Bolivia (Cochabamba, La Paz); Colombia (Cundinamarca); Haiti
80 COS PAN 81 HIS 82 CLM 83 BOL
species :
Dryopteris flemingii (Fraser-Jenk.) Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 125 (1997), pro hybr.
Distribution: India (Himachal Pradesh, Uttarakhand); C-Nepal; Bhutan; China (Yunnan); Tibet
36 CHI TIB 40 BHU IND NEP
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris ×flemingii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 458 (1989)
species :
Dryopteris formosana (Christ) C.Chr.; Ind. 266 (1905)
Distribution: China (Sichuan, Zhejiang); Taiwan; Japan; South Korea (Cheju Isl.); Ryukyu Isl. (Tokunoshima); Philippines
36 CHI 38 JAP KOR NNS TAI 42 PHI
Synonyms: hidden
= Aspidium formosanum Christ; Bull. Herb. Boiss. II, 4: 615 (1904)
= Dryopteris constantissima (Hayata) Hayata; Icon. Pl. Formosan. 4: 191 (1914)
= Dryopteris kodamai Hayata; Icon. Pl. Formosan. 4: 156, f. 97 (1914)
= Dryopteris liui Ching; Bull. Bot. Res. Harbin 4(4): 12 (1984)
= Dryopteris phaeolepis Hayata; Ic. Pl. Formos. 4: 169, t. 109 (1914)
= Polystichum constantissimum Hayata; Ic. Pl. Formos. 4: 191, f. 129 (1914)
species :
Dryopteris fragrans (L.) Schott; Gen. Fil. ad t. 9 (1834)
Name: Duftender Wurmfarn, Fragrant woodfern
Distribution: N-Finland; N-European Russia; Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye, Sakhalin); Kazakhstan; Japan; China (Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Xinjiang); Mongolia; North Korea; South Korea (incl. Cheju Isl.); Alaska; USA (Minnesota, New York)
10 FIN 14 RUS 30 SIB 31 RFE 32 KAZ 36 CHI 37 MON 38 JAP KOR NKO 70 ASK 75 USA
Synonyms: hidden
= Aspidium fragrans (L.) Sw.; Schrad., J. Bot. 1800(2): 35 (1801)
= Dryopteris aquilonaris Maxon; Bull. Torrey Bot. Cl. 27. 638 (1900)
= Dryopteris fragrans f. aquilonaris (Maxon) M.Broun; Index N. Amer. Ferns 68 (1938)
= Dryopteris fragrans var. hookeriana (Fernald) A.Prince ex Weath.; Amer. Fern J. 26: 62 (1936)
= Dryopteris fragrans var. remotiuscula (Kom.) Kom.; Fl. USSR 1: 38 (1934)
= Filix aquilonaris (L.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix fragrans (L.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix-Mas fragrans (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Filix-Mas fragrans var. hookeriana (Fernald) Farw.; Amer. Midl. Naturalist 12: 254 (1931)
= Lastrea fragrans (L.) C.Presl; Tent. Pterid. 76 (1836)
= Nephrodium fragrans (L.) Desv.; Mem. Soc. Linn. Paris 6: 260 (1827), isonym
= Nephrodium fragrans (L.) Rich.; Frankl., Narr. Journey Polar Sea 754 (1823)
= Nephrodium fragrans f. aquilonare (Maxon) Clute; Fern Bull. 15: 21 (1907)
= Nephrodium fragrans var. aquilonare (Maxon) Gilbert; List N. Amer. Pterid. 17 (1901)
= Nephrodium fragrans var. remotiusculum Kom.; Repert. Spec. Nov. Regni Veg. 9: 94 (1911)
= Polypodium fragrans L.; Sp. Pl. 2: 1089 (1753)
= Polystichum fragrans (L.) Ledeb.; Fl. ross. 4: 514 (1853)
= Polystichum fragrans (L.) Roth; Catal. Bot. 2: 147 (1800)
= Thelypteris fragrans (L.) Nieuwl.; Midl. Naturalist 1: 226 (1910)
= Thelypteris fragrans var. hookeriana Fernald; Rhodora 25: 3 (1923)
= Woodsia xanthosporangia Ching; Bull. Fan Mem. Inst. Biol. Bot. 1: 101, c. t (1929)
species :
Dryopteris fragrantiformis Tzvelev; Novosti Sist. Vyssh. Rast. 26: 5 (1989)
Distribution: Russian Far East (Kuril Isl.)
31 RFE
species :
Dryopteris fructuosa (Christ) C.Chr.; Ind. 267 (1905)
Distribution: China (Hubei, Sichuan, Yunnan); Tibet; Taiwan; Bhutan; India (Arunachal Pradesh, Nagaland, Sikkim); Nepal; Myanmar
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA
Remarks: D. lachoongensis is kept separate by Flora of China and probably a valid species fide Knapp (in litt. 2016)
Synonyms: hidden
= Aspidium fructuosum (Christ) Christ; Bull. Soc. Bot. France 52, Mem. 1. 38 (1905)
= Aspidium varium var. fructuosum Christ; Bull. Soc. Bot. France 52, Mem. 1: 38 (1905)
= Dryopteris adenorachis C.Chr.; Ind. Fil., Suppl. 2, 13 (1917), based on D. cavaleriei H. Lévl.
= Dryopteris apicifixa Ching; Boufford & K. H. Shing, J. Arnold Arbor. 64(1): 27 (1983)
= Dryopteris cavaleriei Lév.; Flore du Kouy-tcheou 490 (1915), nom. illeg., non C. Chr. (1905)
= Dryopteris hypophlebia Hayata; Ic. Pl. Formos. 4: 154, f. 96 (1914)
= Dryopteris pseudosabaei Hayata; Ic. Pl. Formos. 5: 283, f. 110 (1915)
= Nephrodium fructuosum (Christ) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)
species :
Dryopteris fuscipes C.Chr.; Ind. Fil., Suppl. 2: 14 (1917)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan (MaZu Isl.); Japan; South Korea; Vietnam; New Guinea (Papua New Guinea)
36 CHI 38 JAP KOR TAI 41 VIE 42 NWG
Synonyms: hidden
= Aspidium erythrosorum var. obtusum Christ; Bull. Soc. Bot. France 52(Mem. 1): 40 (1905)
= Dryopteris bipinnata C.Chr.; Lév., Cat. Pl. Yun-Nan 102 (1916) (non Copel. (1914))
= Dryopteris confertipinna Ching & K.H.Shing; Fl. Fujianica 1: 600 (1982)
= Dryopteris erythrosora var. obtusa (Makino) Makino; Bot. Mag. (Tokyo) 23(270): 144 (1909)
= Dryopteris fuscipes f. major Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990)
= Dryopteris makinoi f. latipinnula H.Itô; Bot. Mag. (Tokyo) 50(590): 71 (1936)
= Dryopteris makinoi Koidz.; Acta Phytotax. 1: 26 (1932), nom. nov.
= Dryopteris medialisora Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 6 (1987)
= Dryopteris medioxima Koidz.; Acta Phytotax. 1: 32 (1932)
= Dryopteris multijugata Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 12 (1983)
= Dryopteris neofuscipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 4 (1987), non Ching & Z. Y. Liu (1984)
= Dryopteris obtusissima Makino; J. Jap. Bot. 2: 47 (1918), nom. nov.
= Dryopteris parafuscipes Ching & Z.Y.Liu; (1984), not Ching & P. S. Chiu (1987)
= Dryopteris persimilis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 16 (1983), nom. illeg., non (Baker) C. Chr.
= Dryopteris stenochlamys Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990)
= Nephrodium erythrosorum var. obtusum Makino; Bot. Mag. (Tokyo) 13: 79 (1899)
= Nephrodium filix-mas f. obtusum (Makino) Makino ex Matsum.; Ind. Pl. Jap. 1: 319 (1904)
= Nephrodium fuscipes (C.Chr.) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)
species :
Dryopteris fuscoatra (Hillebr.) W.J.Rob.; Bull. Torrey Bot. Cl. 39: 592, t. 42 (1912)
Distribution: Hawaii (Kauai, Oahu, Lanai, Molokai, Maui, Hawaii Isl.)
63 HAW
variant :
var. lamoureuxii Fraser-Jenk.; Thaiszia 4(1): 22 (1994)
Distribution: Hawaii (Kauai, Maui, Hawaii Isl.)
63 HAW
species :
Dryopteris futura A.R.Sm.; Proc. Calif. Acad. Sci. ser, 4, 40: 216 (1975)
Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Mexico State, Michoacan, Morelos, Oaxaca); Guatemala; Honduras
79 MEX 80 GUA HON
species :
Dryopteris gamblei (Hope) C.Chr.; Ind. Fil. 267 (1906)
Distribution: India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Uttarakhand, Sikkim, West Bengal); Bhutan; Myanmar; Nepal; Taiwan; Tibet; China; Japan
36 CHI TIB 38 JAP TAI 40 BHU IND NEP 41 MYA
Remarks: not a synonym of D. stenolepis fide Fraser-Jenkins et al. (2015, 2018)
Synonyms: hidden
= Nephrodium gamblei Hope; J. Bombay Nat. Hist. Soc. 12: 533, t. 7 (1899)
= Dryopteris darjeelingensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 335 (1989)
species :
Dryopteris gaoligongensis Z.Y.Zuo, J.Mei Lu & D.Z.Li; Pl. Diversity 44(2): 185 (2021)
Distribution: China (Yunnan)
36 CHI
species :
Dryopteris gemmifera S.Y.Dong; Bot. Stud. (Taipei) 51(4): 537 (2010)
Distribution: China (Hainan)
36 CHI
species :
Dryopteris glabra (Brack.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Remarks: ser. Aemulae
Synonyms: hidden
= Aspidium glabrum (Brack.) Mett.; Pheg. u. Asp. 59, t. 140 (1858)
= Lastrea glabra Brack.; U. S. Expl. Exp. 16: 200 (1854)
= Nephrodium glabrum (Brack.) Baker; Syn. 278 (1867)
= Thelypteris glabra (Brack.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 251 (1941)
variant :
var. alboviridis (W.H.Wagner) D.D.Palmer; Amer. Fern J. 92(2): 100 (2002)
Distribution: Hawaii (Kauai)
63 HAW
Synonyms: hidden
= Dryopteris alboviridis W.H.Wagner; Contr. Univ. Michigan Herb. 22: 177 (1999)
variant :
var. flynnii D.D.Palmer; Amer. Fern J. 92(2): 101 (2002)
Distribution: Hawaii (Kauai)
63 HAW
variant :
var. hobdyana (W.H.Wagner) D.D.Palmer; Amer. Fern J. 92(2): 101 (2002)
Distribution: Hawaii (Maui)
63 HAW
Synonyms: hidden
= Dryopteris hobdyana W.H.Wagner; Contr. Univ. Michigan Herb. 22: 177 (1999)
variant :
var. nuda (Underw.) Fraser-Jenk.; Thaiszia 4(1): 38 (1994)
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Dryopteris nuda Underw.ap.A.Heller; Minn. bot. St. 1: 780 (1897)
variant :
var. pusilla (Hillebr.) Fraser-Jenk.; Thaiszia 4(1): 35 (1994)
Name: Na Pali-Kona Woodfern
Distribution: Hawaii (Kauai)
63 HAW
Synonyms: hidden
= Dryopteris parvula W.J.Rob.; Bull. Torrey Bot. Cl. 39: 593, t. 44 (1912)
= Aspidium glabrum var. pusillum Hillebr.; Fl. Haw. 577 (1888)
= Dryopteris tenebrosa W.H.Wagner; Contr. Univ. Michigan Herb. 19: 68 (1993)
variant :
var. soripes (Hillebr.) Fraser-Jenk.; Thaiszia 4(1): 36 (1994)
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Aspidium glabrum var. soripes Hillebr.
species :
Dryopteris glandulosopaleata J.P.Roux; Kew Bull. 59(2): 211 (2004)
Distribution: Cameroon; Nigeria
22 NGA 23 CMN
species :
Dryopteris goeringiana (Kunze) Koidz.; Bot. Mag. (Tokyo) 48: 382 (1929)
Distribution: China (Gansu, Hebei, Henan, Liaoning, Nei Mongol, Shaanxi, Shanxi, Sichuan, Xinjiang); Japan; North Korea; South Korea; Russian Far East (Khabarovsk, Primorye)
31 RFE 36 CHI 38 JAP KOR NKO
Synonyms: hidden
= Aspidium filix-mas var. giraldii Christ; Baroni & Christ, Nuov. Giorn. Bot. Ital. n. s. 4(1): 94-95 (1897)
= Aspidium goeringianum Kunze; Bot. Zeit. (Berlin) 557 (1848)
= Aspidium oxyodon Franch.; Nouv. Arch. Mus. II, 7: 163 (1883) (non Dryopteris oxyodon (Baker) C. Chr.)
= Asplenium goeringianum (Kunze) Mett.; Aspl. 198, n. 243, t. 6, f. 11-12 (1859)
= Athyrium goeringianum (Kunze) T.Moore; Ind. 185 (1860)
= Athyrium nakaii Tagawa; Acta Phytotax. 2: 195 (1933), nom. nov.
= Athyrium triangulare Nakai; Bot. Mag. (Tokyo) 41: 91 (1931) (non Alderw. (1915))
= Dryopteris filix-mas subsp. oxyodon (Franch.) C.Chr.; Ind. Fil. 5: 266 (1905)
= Dryopteris giraldii (Christ) C.Chr.; J. Wash. Acad. Sci. 17: 500 (1927)
= Dryopteris laeta (Kom.) C.Chr.; Ind. 267 (1905)
= Dryopteris laeta f. triangulare (Nakai) Kitag.; Neo-Lineam. Fl. Manshur.: 35 (1979)
= Dryopteris laeta var. alpina Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 466 (1938)
= Dryopteris laeta var. giraldii (Christ) Ching; Fl. Tsinlingensis 2: 170 (1974)
= Dryopteris laeta var. oblongifrons Kitag.; Bot. Mag. (Tokyo) 48(566): 91, f. 8 (1934)
= Dryopteris oxyodon (Franch.) Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 58 (1935)
= Dryopteris oxyodon var. oblongifrons (Kitag.) Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 59 (1935)
= Dryopteris subramosa Christ; Not. syst. 1: 42 (1909)
= Dryopteris wladiwostokensis B.Fedtsch.; Acta Hort. Petrop. 31: 4 (1912)
= Lastrea goeringiana (Kunze) T.Moore; Ind. 93 (1858)
= Nephrodium laetum Kom.; Fl. Manch. 1: 124 (1901)
= Nephrodium wladiwostokense (B.Fedtsch.) B.Fedtsch.; Acta Hort. Petrop. 31: 99 (1912)
species :
Dryopteris goldieana (Hook.) A.Gray; Man. ed. 1: 631 (1848)
Name: Goldie´s woodfern
Distribution: USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia); Canada (New Brunswick, Ontario, Québec, Nova Scotia)
74 USA 75 USA 78 USA
Synonyms: hidden
= Aspidium americanum Torrey; Cat. Pl. N. York 80 (1818) (nomen)
= Aspidium filix-mas var. goldieanum (Hook.) C.Chr.; Farnkr. Erde 258 (1897)
= Aspidium goldieanum Hook. ex Goldie; Edinburgh New Philos. J. 6: 333 (1822)
= Filix goldieana (Hook.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix-Mas goldieana (Hook.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Lastrea goldieana (Hook.) C.Presl; Tent. Pterid. 76 (1836)
= Nephrodium goldieanum Hook. & Grev.; Icon. Fil. t. 102 (1829)
= Polystichum goldieanum (Hook.) Keyserl.; Pol. Cyath. Hb. Bung. 44 (1873)
= Thelypteris goldieana (Hook.) Nieuwl.; Midl. Naturalist 1: 226 (1910)
species :
Dryopteris gonggaensis H.S.Kung, Li Bing Zhang & X.S.Guo; Acta Bot. Yunnan. 17(2): 159-160, f. 1 (1995)
Distribution: China (Sichuan)
36 CHI
Remarks: included by Fraser-Jenkins (1997) in D. blanfordii ssp. nigrosquamosa, but kept separate in Flora of China
species :
Dryopteris gorgonea J.P.Roux; Bothalia 34(1): 33 (2004)
Distribution: Cape Verde Isl. (Fogo)
21 CVI
species :
Dryopteris grandifrons Li Bing Zhang; Taxon 61(6): 1207 (2012), nom. nov.
Distribution: China (NW-Yunnan)
36 CHI
Synonyms: hidden
= Nothoperanema giganteum Ching; Acta Phytotax. Sin. 11: 28 (1966)
species :
Dryopteris guanchica Gibby & Jermy; J. Linn. Soc., Bot. 74: 256 (1977)
Name: Guanchen-Wurmfarn (DE); Helecho penco dentado (ES)
Distribution: Spain; Portugal; Canary Isl. (Tenerife, La Gomera, Hierro)
12 POR SPA 21 CNY
Remarks: carthusiana agg.
species :
Dryopteris guangxiensis S.G.Lu; Guihaia 19(1): 27 (1999)
Distribution: China (Guangxi)
36 CHI
Synonyms: hidden
= Dryopteris parafuscipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 4 (1987), nom. illeg., non Ching & Z. Y. Liu (1984)
species :
Dryopteris gymnophylla (Baker) C.Chr.; Ind. 269 (1905)
Distribution: China (Anhui, Guizhou, Hebei, Henan, Hubei, Jiangsu, Jiangxi, Shandong, Zhejiang); Japan; North Korea; South Korea; N-Thailand
36 CHI 38 JAP KOR NKO 41 THA
Synonyms: hidden
= Dryopteris sakuraii (Rosenst.) Tagawa; J. Jap. Bot. 13: 185 (1937)
= Dryopteris subtripinnata var. sakuraii Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 132 (1914)
= Nephrodium gymnophyllum Baker; J. Bot. 170 (1887)
= Dryopteris koraiensis Tagawa; Acta Phytotax. Geobot. 6: 254 (1936)
species :
Dryopteris gymnosora (Makino) C.Chr.; Ind. 269 (1905)
Distribution: China (Anhui, Fujian, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan
36 CHI 38 JAP
Synonyms: hidden
= Nephrodium gymnosorum Makino; Bot. Mag. (Tokyo) 13: 64 (1899)
variant :
var. angustata H.Itô; (1938)
Distribution: Japan (Yakushima Isl.)
38 JAP NNS
Conservation Status: CR in Japan
species :
Dryopteris habaensis Ching; Sunyatsenia 6: 2 (1941)
Distribution: China (NW-Yunnan)
36 CHI
species :
Dryopteris hadanoi Kurata; J. Geobot. 14: 83 (1966)
Distribution: Japan
38 JAP
Conservation Status: IUCN R. NT in Japan
species :
Dryopteris handeliana C.Chr.; Dansk Bot. Ark. 9(3): 62, t. 6(5-6) (1937)
Distribution: China (Guizhou, Hubei, Hunan, Sichuan, Yunnan, Zhejiang); Japan
36 CHI 38 JAP
Synonyms: hidden
= Dryopteris tasiroi Tagawa; Acta Phytotax. Geobot. 9: 236 (1940)
species :
Dryopteris hangchowensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 414 (1938)
Distribution: China (NE-Zhejiang); ?Japan; South Korea
36 CHI 38 KOR
species :
Dryopteris hasseltii (Blume) C.Chr.; Ind. 269 (1905)
Distribution: Borneo (Mt. Kinabalu, Sabah); India (Assam, Arunachal Pradesh, Meghalaya, Mizoram); Vietnam; Philippines; Thailand; Sumatra; Sulawesi; ?Lesser Sunda Isl.; Moluccas; New Guinea (Irian Jaya, Papua New Guinea); Java; peninsular Malaysia (Malacca, Perak, Pahang, Johor); Solomon Isl. (Guadalcanal); China (Yunnan, Hainan); Taiwan; S-Japan (Yakushima Isl.); Ryukyu Isl. (Tokara Isl.); Australia (NE-Queensland); Fed. States of Micronesia (Pohnpei); Fiji (Viti Levu, Vanua Levu, Ovalau, Taveuni)
36 CHI 38 JAP NNS TAI 40 IND 41 THA VIE 42 BOR JAW MLY MOL NWG PHI SUL SUM 43 SOL 50 AUS 60 FIJ 62 MIC
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Arachniodes hasseltii (Blume) Ching; Acta Bot. Sin. 10: 258 (1962)
= Athyrium hasseltii (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 6 (1940)
= Byrsopteris hasseltii (Blume) Morton; Amer. Fern J. 50: 154 (1960)
= Dryopteris adiantoides Suzuki; J. Jap. Bot. 11: 644, f. 1 (1935)
= Dryopteris gillespiei Copel.; Bull. Bernice P. Bishop Mus. 69: 10 (1929)
= Acrorumohra hasseltii (Blume) Ching; Acta Phytotax. Sin. 9: 385 (1964)
= Dryopteris hatusimae H.Itô; Bot. Mag. (Tokyo) 67: 216, f. 2 (1954)
= Dryopteris laserpitiifolia (Scort.) C.Chr.; Ind. 273 (1905), non (Mett.) Kuntze (1891)
= Dryopteris obovata (Baker) C.Chr.; Ind. 280 (1905)
= Dryopteris odontophora Copel.; Phil. J. Sci. 60: 106 (1936)
= Dryopteris psilosora Tagawa; Acta Phytotax. 2: 191 (1932)
= Dryopteris sparsa var. raapii Alderw.; Mal. Ferns 197 (1908)
= Lastrea obovata (Baker) Bedd.; Handb. Suppl. 60 (1892)
= Nephrodium obovatum Baker; J. Bot. 265 (1890)
= Phegopteris hasseltii (Blume) Mett.; Pheg. u. Asp. 13, n. 14 (1858)
= Phegopteris laserpitiifolia Bedd.; Handb. Suppl. 84 (1892)
= Polypodium anisopterum Kunze; Bot. Zeit. (Berlin) 118 (1848)
= Polypodium hasseltii Blume; Fl. Jav. Fil. 195, t. 92 (1829)
= Polypodium laserpitiifolium Scort. ex Bedd.; J. 324 (1887)
= Polystichopsis hasseltii (Blume) Holttum; Rev. Fl. Mal. 2: 487, f. 288 (1955)
= Rumohra hasseltii (Blume) Ching; Sinensia 6: 61 (1934)
species :
Dryopteris hawaiiensis (Hillebr.) W.J.Rob.; Bull. Torrey Bot. Cl. 39: 594 (1912)
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
63 HAW
Remarks: ser. Aemulae
Synonyms: hidden
= Aspidium hawaiiense Hillebr.; Fl. Haw. 575 (1888)
species :
Dryopteris hendersonii (Bedd.) C.Chr.; Ind. 270 (1905)
Distribution: India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, West Bengal); Myanmar; Bhutan; Thailand; Vietnam; Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon); Nepal; China (Yunnan, Guizhou); Taiwan; Japan
36 CHI 38 JAP TAI 40 BHU IND NEP 41 MYA THA VIE 42 BOR PHI
Remarks: subgen. Nothoperanema
Synonyms: hidden
= Aspidium spectabile Wall.; Num. List no. 372 (1828) (nomen, non Blume (1828))
= Ctenitis hendersonii (Bedd.) H.Itô ex Nakai & Honda; Nov. Fl. Jap. 489, t. 5 (106) (1939)
= Ctenitis leptorhachis (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 280 (1938)
= Ctenitis mearnsii Copel.; Phil. J. Sci. 81: 24 (1952)
= Dryopteris leptorhachis Hayata; Ic. Pl. Formos. 4: 162, f. 102 (1914)
= Lastrea hendersonii Bedd.; Handb. Ferns Brit. India, Suppl., t. 377 (1876)
= Lastrea spectabilis (C.B.Clarke) Bedd.; Handb. Ferns Brit. India 257 (1883)
= Lastrea wallichii T.Moore; Ind. 104 (1858)
= Nephrodium spectabile C.B.Clarke; Tr. Linn. Soc. II, 1: 526 (1880)
= Nothoperanema hendersonii (Bedd.) Nakaike; Enum. Pterid. Jap., Fil. 239 (1975), isonym
= Nothoperanema hendersonii (Bedd.) Ching; Acta Phytotax. Sin. 11: 28 (1966)
species :
Dryopteris herbacea Alderw.; Bull. Jard. Bot. Buitenz. II, 20: 13 (1915)
Distribution: Java
42 JAW
Remarks: provisionally accepted name
species :
Dryopteris heterolaena C.Chr.; Acta Hort. Gothob. 1: 62, t. 17 (1924)
Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Sichuan, Yunnan, Zhejiang); Tibet; NE-India (Arunachal Pradesh)
36 CHI TIB 40 IND
Remarks: Subgen. Dryopsis
Conservation Status: IUCN CR for India (as C. fengiana, Fraser-Jenkins 2012)
Synonyms: hidden
= Ctenitis chingii Z.Y.Liu & J.I.Chang; Bull. Bot. Res. Harbin 4(4): 14, f. 42 (1984)
= Ctenitis heterolaena (C.Chr.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 293 (1938)
= Ctenitis kwangsiensis Ching & P.S.Chiu; Acta Phytotax. Sin. 19(1): 120 (1981)
= Ctenitis omeiensis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 121 (1981)
= Ctenitis tibetica Ching & S.K.Wu; Fl. Xizangica 1: 274 (1983)
= Ctenitis zhejiangensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 34-35, f. 25 (1983)
= Dryopsis fengiana (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 183 (1986)
= Ctenitis fengiana Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 69 (1941)
= Dryopsis kwangsiensis (Ching & Chu H.Wang) Holttum & P.J.Edwards; Kew Bull. 41(1): 186 (1986)
= Dryopsis heterolaena (C.Chr.) Holttum & P.J.Edwards; Kew Bull. 41(1): 185 (1986)
= Dryopteris nobilis var. fengiana Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 67 (1941)
species :
Dryopteris himachalensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 367 (1989)
Distribution: India (Himachal Pradesh); China (NW-Yunnan, SW-Sichuan)
36 CHI 40 IND
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris pectinatopinnata Ching; Acta Phytotax. Sin. 23(5): 349-350 (1985), nom. nud.
species :
Dryopteris hirtipes (Blume) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: peninsular Malaysia (Cameron Highlands, Pahang); S-India (Kerala, Tamil Nadu); N-Thailand; Sumatra; Java; Lesser Sunda Isl. (Bali); Sulawesi; Borneo (Sabah); Philippines; Myanmar; Vietnam; Sri Lanka; Fiji (Viti Levu); [Western] Samoa (Savaii, Upolu); New Guinea (Irian Jaya, Papua New Guinea)
40 IND SRL 41 MYA THA VIE 42 BOR JAW LSI MLY NWG PHI SUL SUM 60 FIJ SAM
Synonyms: hidden
= Aspidium aduncum Wall.; Num. List no. 384 (1828)
= Aspidium hirtipes Blume; Enum. Fil. 148 (1828)
= Lastrea hirtipes (Blume) T.Moore; Ind. 85 (1858)
species :
Dryopteris hondoensis Koidz.; Acta Phytotax. 1: 31 (1932)
Distribution: China (Sichuan, Zhejiang); Japan
36 CHI 38 JAP
Synonyms: hidden
= Dryopteris scabripes Ching; Bot. Res. Acad. Sin. 1987(2): 9 (1987)
species :
Dryopteris huberi (Christ) C.Chr.; Ind. Fil., Suppl. 34 (1913)
Distribution: Brazil (Goiás, Mato Grosso, Pará)
84 BRA
Remarks: a species close to D. futura, recognized by Sessa et al. 2012
Synonyms: hidden
= Aspidium huberi Christ; Hedwigia 46: 192 (1906)
species :
Dryopteris hypolepioides Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 175 (1913)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Phegopteris hypolepioides (Rosenst.) Alderw.; Mal. Ferns Suppl. 309 (1917)
= Ctenitis hypolepioides (Rosenst.) Copel.; Gen. Fil. 124 (1947)
species :
Dryopteris immixta Ching; Fl. Tsinlingensis 2: 225 (1974)
Distribution: China (Fujian, Gansu, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang)
36 CHI
species :
Dryopteris inaequalis (Schltdl.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: South Africa (KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Eswatini [Swaziland]; Lesotho
27 LES SAF SWZ
Synonyms: hidden
= Aspidium amplum Kuhn; Fil. Afr. 125 (1868)
= Aspidium inaequale Schltdl.; Adumbr. Fl. Aethiop. 23, t. 12 (1825)
= Lastrea inaequalis (Schltdl.) C.Presl; Tent. Pterid. 77 (1836)
= Nephrodium inaequale (Schltdl.) Hook.; Sp. 4: 125 (1862) (non Schrad. (1824))
= Polystichum inaequale (Schltdl.) Keyserl.; Pol. Cyath. Hb. Bung. 44 (1873)
species :
Dryopteris indonesiana Darnaedi; M. Kato & K. Iwats., J. Jap. Bot. 64(10): 304 (1989)
Distribution: Moluccas; Java; Sumatra
42 JAW MOL SUM
species :
Dryopteris indusiata (Makino) Makino & Yamam.; Suppl. Ic. Pl. Formos. V. 2 (1932)
Distribution: China (Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan
36 CHI 38 JAP
Synonyms: hidden
= Dryopteris gymnosora var. indusiata (Makino) Makino ex Bonap.
= Dryopteris labordei var. indusiata (Makino) Seriz.; J. Jap. Bot. 52(1): 16 (1977)
= Nephrodium gymnosorum var. indusiatum Makino; Bot. Mag. (Tokyo) 13: 65 (1899)
species :
Dryopteris insularis Kodama; Matsum., Ic. pl. Koisik. 2: 49, t. 109 (1913)
Distribution: Bonin Isl.
38 OGA
Conservation Status: VU in Japan. IUCN R
variant :
var. chichishimensis (Nakai ex Itô) Itô; (1939)
Distribution: Bonin Isl. (Chichijima); Japan (Izu Isl.)
38 JAP OGA
Synonyms: hidden
= Dryopteris chichishimensis Nakai ex H.Itô; Bot. Mag. (Tokyo) 40: 435, f. 6) 7(14) (1935)
species :
Dryopteris integriloba C.Chr.; Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. no. 6: 5 (1933)
Distribution: Japan; Taiwan; China (Guangdong, Hainan, Guangxi, Yunnan); N-Vietnam; peninsular Malaysia (Negeri Sembilan); Thailand
36 CHI 38 JAP TAI 41 THA VIE 42 MLY
Synonyms: hidden
= Dryopteris subintegriloba Seriz.; J. Jap. Bot. 46: 17, f.1a (1971)
species :
Dryopteris intermedia (Muhl. ex Willd.) A.Gray; Man. 630 (1848)
subspecies :
subsp. intermedia
Name: Fancy fern, Intermediate woodfern
Distribution: USA (Alabama, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia); Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Prince Edward Isl., Québec); St. Pierre et Miquelon
72 SPM 74 USA 75 USA 78 USA
Synonyms: hidden
= Aspidium americanum Davenp.; Amer. Naturalist 12: 714 (1878)
= Aspidium grayanum Regel; Ind. Sem. ht. Petr. (1855)
= Aspidium intermedium Muhl. ex Willd.; Sp. Pl. 5(1): 262 (1810)
= Dryopteris austriaca var. intermedia (Muhl. ex Willd.) C.V.Morton; Amer. Fern J. 40: 222 (1950)
= Dryopteris intermedia f. furcata C.F.Reed; Castanea 16: 70, fig. 5 (1951)
= Dryopteris intermedia f. sparsifrondosa C.F.Reed; Castanea 16: 71, fig. 6 (1951)
= Dryopteris spinulosa var. intermedia (Muhl. ex Willd.) Underw.; Native Ferns, ed. 4 116 (1893)
= Filix spinulosa var. intermedia (Muhl. ex Willd.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 83 (1916)
= Filix-mas spinulosa var. intermedia (Muhl. ex Willd.) Farw.; Amer. Midl. Naturalist 12: 257 (1931)
= Lastrea intermedia (Muhl. ex Willd.) C.Presl; Tent. Pterid. 77 (1836)
= Nephrodium intermedium (Muhl. ex Willd.) C.Presl; Reliq. Haenk. 1: 38 (1826)
= Nephrodium spinulosum f. intermedium (Muhl. ex Willd.) Gilbert
= Nephrodium spinulosum var. intermedium (Muhl. ex Willd.) Davenp.; Gilbert, List N. Amer. Pterid. 17 (1901)
= Thelypteris intermedia (Muhl. ex Willd.) House; New York State Mus. Bull. 233-234 (1920)
= Thelypteris spinulosa var. intermedia (Muhl. ex Willd.) Nieuwl.; Amer. Midl. Naturalist 2: 278 (1912)
subspecies :
subsp. azorica (Christ) Jermy; Webbia 38: 408 (1984)
Distribution: Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.)
21 AZO
Synonyms: hidden
= Aspidium dilatatum var. azoricum Christ; Bull. Ac. Géogr. Bot. Mans 17 (212): 157 (1907)
= Dryopteris azorica (Christ) Alston; Bol. Soc. Broter. Ser.2. 30: 14 (1956)
subspecies :
subsp. maderensis (Milde ex Alston) Fraser-Jenk.; Arquipelago, Ser. G. Nat. 2: 85 (1981)
Distribution: Madeira (Madeira Isl.)
21 MDR
Synonyms: hidden
= Aspidium spinulosum f. maderensis Milde; Fil. Eur. & Atl. 140 (1867)
= Dryopteris maderensis Milde ex Alston; Bol. Soc. Broter. ser. 2, 30: 14 (1956)
species :
Dryopteris inuyamensis H.Itô; J. Jap. Bot. 9: 54 (1933)
Distribution: Japan
38 JAP
species :
Dryopteris iranica Fraser-Jenk.; Fern Gaz. 18(1): 9 (2007)
Distribution: Iran; Turkey (Anatolia)
34 IRN TUR
Remarks: affinis agg.
Synonyms: hidden
= Dryopteris wallichiana subsp. coriacea (Fraser-Jenk.) Fraser-Jenk.; Thaiszia 4(1): 23 (1994)
= Dryopteris affinis subsp. coriacea Fraser-Jenk.; Willdenowia 10(1): 112 (1980)
= Dryopteris coriacea (Fraser-Jenk.) Tzvelev; Novosti Sist. Vyssh. Rast. 35: 17 (2003), nom. illeg., non (Brause) Ching
= Dryopteris affinis var. gigantea Fraser-Jenk.; (1980)
species :
Dryopteris jishouensis G.X.Chen & D.G.Zhang; Life Sci. Res. 17(5): 412, figs. 1-2 (2013)
Distribution: China (Hunan)
36 CHI
species :
Dryopteris jiucaipingensis P.S.Wang, Q.Luo & Li Bing Zhang; Novon 22(2): (2012)
Distribution: China (NW-Guizhou)
36 CHI
species :
Dryopteris juxtaposita Christ; Bull. Ac. Géogr. Bot. Mans 138 (1907)
Distribution: China (Yunnan, Sichuan, Guizhou, Gansu); Tibet; Myanmar; India (Himachal Pradesh, Arunachal Pradesh, Karnataka, Kerala, Manipur, Meghalaya, Nagaland, Sikkim, Tamil Nadu, Uttarakhand, West Bengal, Jammu & Kashmir); Nepal; Bhutan; Vietnam
36 CHI TIB 40 BHU IND NEP 41 MYA VIE
Synonyms: hidden
= Nephrodium filix-mas var. normale C.B.Clarke; Trans. Linn. Soc. London, 2 Bot., 1: 519, t. 68, f. 2 (1880)
= Aspidium filix-mas var. normale (C.B.Clarke) Christ; Bull. Soc. Bot. France 52, Mem. 1: 38 (1905)
species :
Dryopteris karwinskyana (Mett.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: Mexico (Chiapas, Ciudad de Mexico, Durango, Guerrero, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Sinaloa); +Easter Isl.; Guatemala; Honduras; El Salvador; Nicaragua; Costa Rica
61 EAS 79 MEX 80 COS ELS GUA HON NIC
Synonyms: hidden
= Aspidium karwinskyanum Mett.; Abh. Senckenb. Naturf. Ges. 2: 343 (1858)
= Dryopteris espinosae Hicken; Rev. Chil. Hist. Nat. 17: 93, t. 9 (1913)
= Lastrea karwinskiana (Mett.) T.Moore; Ind. 94 (1858)
= Nephrodium karwinskyanum (Mett.) Baker; Syn. 279 (1867)
= Nephrodium karwinskyanum f. grandis Donn.Sm.; Enum. Pl. Guatem. 4: 172 (1895)
= Thelypteris espinosae (Hicken) R.R.Rodriguez; Bol. Soc. Biol. Concepcion 45: 153 (1972)
species :
Dryopteris kashmiriana Fraser-Jenk. & Widén; Ferns and Fern-allies of Nepal 1: 26 (2015)
Distribution: India (Jammu & Kashmir, Uttarakhand)
40 IND
species :
Dryopteris katangaensis J.P.Roux; Bot. J. Linn. Soc. 142(2): 237 (2003)
Distribution: Zambia; D.R.Congo [Zaire]
23 ZAI 26 ZAM
species :
Dryopteris kawakamii Hayata; J. Coll. Sci. Imp. Univ. Tokyo 80: 416 (1911)
Distribution: China (Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Tibet; Taiwan; Myanmar
36 CHI TIB 38 TAI 41 MYA
Remarks: Subgen. Dryopsis. D. mariformis is possibly a separate species
Synonyms: hidden
= Aspidium polylepis Christ; (1905) (non Franch. & Sav. 1879)
= Ctenitis jiulungshanensis P.C.Chiu & G.H.Yao; Bull. Bot. Res. Harbin 2(2): 66-67, pl. 2, f. 2 (1982)
= Ctenitis kawakamii (Hayata) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 286 (1938)
= Ctenitis mariformis (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 286 (1938)
= Ctenitis sacholepis (Hayata) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 80 (1939)
= Ctenitis truncata Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 123 (1981)
= Dryopsis kawakamii (Hayata) Holttum & P.J.Edwards; Kew Bull. 41(1): 186 (1986)
= Dryopsis mariformis (Rosenst.) Holttum & P.J.Edwards; Kew Bull. 41(1): 188 (1986)
= Dryopsis truncata (Ching & H.S.Kung) Holttum & P.J.Edwards; Kew Bull. 41(1): 188 (1986)
= Dryopteris filix-mas var. rosthornii C.Chr.
= Dryopteris mariformis Rosenst.; Repert. Spec. Nov. Regni Veg. 18: 131 (1914)
= Dryopteris sacholepis Hayata; Ic. Pl. Formos. 6: 285, f. 111 (1916)
species :
Dryopteris khullarii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 362 (1989)
Distribution: NW-India (Himachal Pradesh, Uttarakhand); Nepal
40 IND NEP
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
species :
Dryopteris kilemensis (Kuhn) Kuntze; Rev. Gen. Pl. 2: 813 (1891) (kilmensis)
Distribution: Cameroon; D.R.Congo [Zaire]; Sudan; Ethiopia; Uganda; Rwanda; Burundi; Kenya; Tanzania; Zambia; Malawi; Mozambique; Zimbabwe; Madagascar
23 BUR CMN RWA ZAI 24 ETH SUD 25 KEN TAN UGA 26 MLW MOZ ZAM ZIM 29 MDG
Synonyms: hidden
= Aspidium kilemense Kuhn; v. Decken, Reis. 83: Bot. 46 (1879)
= Aspidium lastii (Baker) Hieron.; Engl., Pflanzenw. Ostafr. C: 85 (1895)
= Dryopteris cordipinnula C.Chr.; Cat. Pl. Madag. Pterid. 27 (1932) (nomen), Dansk Bot. Ark. 7: 55, t. 11, f. 10-12 (1932)
= Dryopteris lastii (Baker) C.Chr.; Ind. 274 (1905)
= Dryopteris platylepis Rosenst.; Feddes Repert. 4: 4, 5 (1907)
= Dryopteris remotipinnula Bonap.; Notes pterid. 16: 178 (1925)
= Nephrodium kilemense (Kuhn) Baker; Syn. Fil. (Hook. & Baker) ed. 2: 498 (1874)
= Nephrodium lastii Baker; Ann. Bot. 6: 324 (1891)
species :
Dryopteris kinkiensis Koidz.; Tagawa, Acta Phytotax. 2: 200 (1933)
Distribution: China (Fujian, Guangdong, Zhejiang, Jiangxi, Sichuan); Japan; South Korea (incl. Cheju Isl.); Myanmar (Kachin, etc.)
36 CHI 38 JAP KOR 41 MYA
Synonyms: hidden
= Dryopteris championii var. tenuifrons (H.Itô) H.Itô; Nov. Fl. Jap. 4: 48 (1939)
= Dryopteris neoassamensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 480 (1938)
= Dryopteris pseudoerythrosora var. tenuifrons H.Itô
= Dryopteris zhenangensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 11 (1987)
= Dryopteris hunanica Ching; Bull. Fan Mem. Inst. Biol., Bot. 10: 179 (1940) (fide Fraser-Jenkins 1986)
species :
Dryopteris kinokuniensis Kurata; J. Geobot. 18: 6 (1970)
Distribution: Japan
38 JAP
species :
Dryopteris knoblochii A.R.Sm.; Mem. New York Bot. Gard. 88 (Pterid. Mexico): 276 (2004)
Distribution: USA (Arizona); Mexico (Chihuahua, Durango, Sinaloa, Sonora)
76 USA 79 MEX
species :
Dryopteris koidzumiana Tagawa; Acta Phytotax. 2: 190 (1933)
Distribution: Japan
38 JAP
Synonyms: hidden
= Dryopteris erythrosora var. koidzumiana (Tagawa) Hara; (1939)
species :
Dryopteris komarovii Kossinsky; Not. syst. Hort. Petr. 2: 1 (1921)
Distribution: China (Gansu, Qinghai, Shaanxi, Sichuan, Yunnan); Taiwan; Tibet; Bhutan; India (Arunachal Pradesh, Jammu & Kashmir, Himachal Pradesh, Sikkim, Uttarakhand); Myanmar; Nepal; Pakistan (Kurram, Hazara); Tajikistan; Kyrgyzstan; Turkmenistan; Uzbekistan
32 KGZ TKM TZK UZB 36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Dryopteris barbigera subsp. komarovii (Kossinsky) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 384 (1989)
= Dryopteris subbarbigera Ching; Fl. Xizangica 1: 260 (1983)
species :
Dryopteris kominatoensis Tagawa; Acta Phytotax. Geobot. 6: 154 (1937)
Distribution: Japan
38 JAP
species :
Dryopteris kwanzanensis Tagawa; Acta Phytotax. Geobot. 7: 186 (1938)
Distribution: Taiwan
38 TAI
Synonyms: hidden
= Dryopteris amoena Ching; Bull. Fan Mem. Inst. Biol., Bot. 8: 454 (1938) (fide Fraser-Jenkins 1986)
species :
Dryopteris labordei (Christ) C.Chr.; Ind. 273 (1905)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan
36 CHI 38 JAP
Remarks: not in Taiwan
Synonyms: hidden
= Aspidium labordei Christ; Bull. Soc. Bot. France 62, Mem. 1. 40 (1905)
species :
Dryopteris lacera (Thunb.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: China (Heilongjiang, Hubei, Hunan, Jiangxi, Sichuan, Zhejiang); Japan; North Korea; South Korea; Taiwan
36 CHI 38 JAP KOR NKO TAI
Synonyms: hidden
= Aspidium filix-mas var. lacerum (Thunb.) Christ; Farnkr. Erde 257 (1897)
= Aspidium lacerum (Thunb.) Sw.; Schrad., J. Bot. 1800(2): 39 (1801)
= Aspidium lacerum var. bipinnatum Franch. & Sav.; Enum. Pl. Jap. 2: 238 (1876)
= Aspidium lacerum var. obtusum Christ; Bull. Soc. Bot. France 52(Mem. 1): 39 (1905)
= Dryopteris filix-mas var. lacerum (Thunb.) Christ; (1897)
= Dryopteris lacera var. chinensis Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 439 (1938)
= Lastrea lacera (Thunb.) D.C.Eaton; Proc. Amer. Acad. 4: 110 (1858)
= Nephrodium lacerum (Thunb.) Baker; Syn. 273 (1867)
= Polypodium lacerum Thunb.; Fl. Jap. 337 (1784) et Linn. Syst. Veg. ed. 14: 938 (1784)
= Polystichum lacerum C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 56 (1851)
species :
Dryopteris lachoongensis (Bedd.) B.K.Nayar & Kaur; Comp. to Bedd. Handb. Ferns Brit. India 61 (1974)
Distribution: China (Yunnan); Tibet; Taiwan; India; Nepal; ?Bhutan
36 CHI TIB 38 TAI 40 IND NEP
Remarks: valid species separate from D. fructuosa - where it was included by Fraser-Jenkins - fide Knapp & Hsu (2017)
Synonyms: hidden
= Lastrea filix-mas var. lachoongensis Bedd.; Handb. Ferns Brit. India, Suppl. 58 (1892) (fide Fraser-Jenkins 1997)
= Dryopteris pseudodontoloma Ching; Fl. Xizangica 1: 265 (1983)
= Dryopteris venosa Ching & S.K.Wu; Fl. Xizangica 1: 264 (1983)
species :
Dryopteris lacunosa S.Jess., Zenner, Chr.Stark & Bujnoch; Kochia 5: 21 (2011)
Name: Lückiger Wurmfarn, Lückiger Schuppen-Wurmfarn (DE)
Distribution: Switzerland; England; Ireland; France; Germany (Baden-Württemberg, Bayern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland); Italy; Austria; England; Scotland; Ireland
10 GRB IRE 11 AUT GER SWI 12 FRA 13 ITA
species :
Dryopteris lepidopoda Hayata; Ic. Pl. Formos. 4: 161 (1914)
Distribution: China (N-Yunnan, W-Sichuan); Taiwan; Tibet; Bhutan; Nepal; India (Jammu & Kashmir, Himachal Pradesh, Arunachal Pradesh, Assam State, ?Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Pakistan (Hazara); Myanmar (Chin)
36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Christella khasiana (C.B.Clarke) Lév.; Flore du Kouy-tcheou 474-476 (1915)
= Dryopteris filix-mas subsp. khasiana (C.B.Clarke) C.Chr.; Ind. Fil. 5: 265 (1905)
= Dryopteris junlianensis H.S.Kung; Acta Bot. Yunnan. 4(4): 340 (1982)
= Dryopteris latibasis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 61 (1941) (fide Fraser-Jenkins et al. 2018)
= Dryopteris lepidopoda var. phaeocoma Ching & S.K.Wu; Fl. Xizangica 1: 258 (1983)
= Dryopteris longistipes Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 59 (1941)
= Dryopteris neolepidopoda Ching & S.K.Wu; Fl. Xizangica 1: 258 (1983)
= Dryopteris nigra Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 430 (1938)
= Dryopteris paleacea var. khasiana (C.B.Clarke) C.Chr.; (1931) (non D. khasiana C. Chr. 1905)
= Dryopteris taiwanicola Tagawa; Acta Phytotax. Geobot. 8: 230 (1939)
= Dryopteris yigongensis Ching; Fl. Xizangica 1: 253 (1983) (fide Fraser-Jenkins 2008)
= Nephrodium filix-mas var. khasiana C.B.Clarke; Tr. Linn. Soc. II, 1: 519, t. 69, f. 1 (1880)
= Nephrodium parallelogrammum subsp. khasiana (C.B.Clarke) Hope; J. Bombay Nat. Hist. Soc. 14: 729 (1903)
species :
Dryopteris lepidorachis C.Chr.; Ind. Fil. 5: 274 (1905), nom. nov.
Distribution: China (Anhui, Fujian, Jiangsu, Jiangxi, Zhejiang)
36 CHI
Synonyms: hidden
= Dryopteris basitripinnatifida Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 6 (1984)
= Dryopteris championii var. rheosora (Baker) K.H.Shing; Fl. Jiangxi 1: 282 (1993)
= Dryopteris lepidocaulon Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 19 (1987), non Kuntze (1891)
= Nephrodium rheosorium (Baker) Hand.-Mazz.; Symb. Sin. 6: 24 (1929), based on Polypodium rheosorum Baker (1891), nom. nud.
= Polypodium rheosorum Baker; Ann. Bot. 5: 457 (1891) (non Baker (1884))
species :
Dryopteris lepinei (Kuhn) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: Society Isl. (Tahiti)
61 SCI
Synonyms: hidden
= Aspidium lepinei Kuhn; Linn. 36: 118 (1869)
= Nephrodium lepinei (Kuhn) Baker; Syn. Fil. (Hook. & Baker) ed. 2: 501 (1874)
species :
Dryopteris leucorhachis (Cheeseman) C.Chr.; Ind. 274 (1905)
Distribution: Cook Isl. (Rarotonga)
61 COO
Synonyms: hidden
= Acrophorus leucorhachis (Cheeseman) Brownlie; Brownlie & Philipson, Pac. Sci. 25: 507 (1971)
= Nephrodium leucorhachis Cheeseman; Tr. Linn. Soc. II, Bot. 6. 309 (1903)
species :
Dryopteris lewalleana Pic.Serm.; Bull. Jard. Bot. Belg. 55(1-2): 158 (1985)
Distribution: D.R.Congo [Zaire]; Ethiopia; Uganda; Burundi; Kenya; Tanzania; Zambia; Malawi; Mozambique; Zimbabwe; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov., W-Cape Prov.); Lesotho; Eswatini [Swaziland]
23 BUR ZAI 24 ETH 25 KEN TAN UGA 26 MLW MOZ ZAM ZIM 27 LES SAF SWZ
Synonyms: hidden
= Dryopteris inaequalis var. atropaleacea Schelpe; Bol. Soc. Broter., ser. 2, 41: 213 (1967)
= Lastrea pentagona T.Moore; J. Bot. 5: 227 (1853)
= Aspidium pentagonum (T.Moore) Kuhn; Fil. Afr. 139 (1868)
species :
Dryopteris liangkwangensis Ching; Icon. Fil. Sin. 4: t. 180 (1937) (liankwangensis)
Distribution: China (Guangxi, Guangdong, SE-Yunnan); Vietnam
36 CHI 41 VIE
species :
Dryopteris liboensis P.S.Wang, X.Y.Wang & Li Bing Zhang; Novon 22(2): 256 (2012)
Distribution: China (S-Guizhou)
36 CHI
species :
Dryopteris lijianxiui Xiao J.Li; Bangladesh J. Bot. 50(3): 747 (2021)
Distribution: China (Shandong)
36 CHI
species :
Dryopteris loxoscaphoides (Baker) C.Chr.; Ind. 276 (1905)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Acrophorus loxoscaphoides (Baker) Alston; J. Bot. 77: 288 (1939)
= Davallia forbesii A.Gepp; J. Bot. Suppl. 59 (1923)
= Leucostegia loxoscaphoides (Baker) C.Chr.; Ind. Fil., Suppl. III: 120 (1934)
= Phegopteris loxoscaphoides (Baker) Alderw.; Mal. Ferns 493 (1909)
= Polypodium loxoscaphoides Baker; J. Bot. 107 (1890)
species :
Dryopteris ludens (Baker) C.Chr.; Ind. 276 (1906)
Distribution: New Caledonia; [Western] Samoa (Savaii); American Samoa (Ta´u); Solomon Isl. (Guadalcanal, Kolombangara, Makira, Vella Lavella)
43 SOL 60 ASA NWC SAM
Synonyms: hidden
= Aspidium arborescens (Baker) Luerss.; Fil. Graeff. 176 (1871)
= Aspidium ludens (Baker) Christ; Engl. Jahrb. 28: 352 (1896)
= Aspidium powellii (Baker) Christ; Engl. Jahrb. 28: 354 (1896)
= Dryopteris arborescens (Baker) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
= Dryopteris powellii (Baker) C.Chr.; Ind. 285 (1905)
= Nephrodium arborescens Baker; Syn. 286 (1867)
= Nephrodium ludens Baker; Ann. Bot. 6: 325 (1891)
= Nephrodium powellii Baker; Ann. Bot. 5: 325 (1891)
species :
Dryopteris ludoviciana (Kunze) Small; Ferns S. E. States 281 (1938)
Name: Southern woodfern
Distribution: USA (Alabama, Arkansas, Florida, Georgia, Kentucky, Louisiana, Mississippi, North Carolina, South Carolina, Texas)
77 USA 78 USA
Synonyms: hidden
= Aspidium cristatum var. floridanum (Hook.) Christ; 258 (1897)
= Aspidium floridanum (Hook.) D.C.Eaton; Chapm. Fl. South. U. St. 595 (1860)
= Aspidium ludovicianum Kunze; Amer. J. Sci. Arts, ser. 2, 6: 84 (1848)
= Dryopteris floridana (Hook.) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
= Filix floridana (Hook.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix-mas cristata var. floridana (Hook.) Farw.; Amer. Midl. Naturalist 12: 254 (1931)
= Lastrea floridanum (Hook.) J.Sm.; Ferns Brit. For., ed. II: 306 (1877)
= Lastrea petiolata Riddell; New Orleans Med. Surg. J. 9: 617 (1853)
= Nephrodium floridanum Hook.; Fil. Exot. t. 99 (1859)
species :
Dryopteris lunanensis (Christ) C.Chr.; Ind. 276 (1905)
Distribution: China (Gansu, Guizhou, Sichuan, Yunnan); Bhutan; India (Arunachal Pradesh)
36 CHI 40 BHU IND
Synonyms: hidden
= Aspidium lunanense Christ; Bull. Herb. Boiss. 6: 966 (1898)
= Dryopteris semipinnata Ching; Fl. Tsinlingensis 2: 226 (1974)
= Dryopteris lungnanensis Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990), orth. var.
species :
Dryopteris lustrata (Hieron.) C.Chr.; Ind. Fil. 276 (1905)
Distribution: Colombia (Nariño)
82 CLM
Synonyms: hidden
= Nephrodium lustratum Hieron.; Bot. Jahrb. Syst. 34(4): 443 (1904)
species :
Dryopteris macrochlamys (Fée) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 453 (1989)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Acrorumohra obtusissima (Mett. ex Kuhn) Ching; Acta Phytotax. Sin. 9: 385 (1964) (fide Fraser-Jenkins 1997)
= Acrorumohra undulata (Bedd.) Ching; Acta Phytotax. Sin. 9: 385 (1964)
= Arachniodes obtusissima (Mett. ex Kuhn) Ching; Acta Bot. Sin. 10: 259 (1962)
= Arachniodes zeylanica (Ching) Ching; Acta Bot. Sin. 10: 260 (1962) (fide Fraser-Jenkins 1997)
= Aspidium macrochlamys Fée; Gen. 296 (1850-52)
= Aspidium obtusissimum Mett. ex Kuhn; Linn. 36: 119 (1869)
= Aspidium undulatum (Bedd.) Thwaites; Enum. pl. Zeyl. 444 (1864) (non Afzel. ex Sw. (1801))
= Dryopteris deparioides subsp. gracillima (Ching) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 449 (1989) (fide Fraser-Jenkins 1997)
= Dryopteris gracillima Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 452 (1938) (fide Fraser-Jenkins 1997)
= Dryopteris gracillima var. prolongata Sledge; Bull. Brit. Mus. (Nat. Hist.) Bot. 5(1): 19 (1973)
= Dryopteris gracillima var. triangularis Sledge
= Dryopteris obtusissima (Mett. ex Kuhn) Christ; J. de Bot. 21: 231 (1908)
= Dryopteris simulans Kuntze; Rev. Gen. Pl. 2: 813 (1891)
= Dryopteris sri-lankensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 451 (1989), nom. nov. superfl.
= Dryopteris undulata (Bedd.) Kuntze; Rev. Gen. Pl. 2: 814 (1891)
= Lastrea deltoidea Bedd.; Ferns South. India, t. 248 (1863) (non (Sw.) T. Moore)
= Lastrea filix-mas var. minor Bedd.; Handb. Ferns Brit. India 254 (1883)
= Lastrea undulata Bedd.; Ferns South. India, t. 271 (1863)
= Nephrodium sparsum var. minus Thwaites ex Baker; Hook. & Baker, Syn. Fil. ed.2. 498 (1874)
= Nephrodium undulatum (Bedd.) Baker; Syn. 276 (1867)
= Rumohra obtusissima (Mett. ex Kuhn) Ching; Sinensia 6: 62 (1934)
= Rumohra zeylanica Ching; Sinensia 5: 70 (1934) (fide Fraser-Jenkins 1997)
species :
Dryopteris macrolepidota Copel.; Bull. Bernice P. Bishop Mus. 98: 8, t. 7 B (1932)
Distribution: Society Isl. (Tahiti)
61 SCI
species :
Dryopteris macropholis Lorence & W.L.Wagner; PhytoKeys 4: 15 (2011)
Distribution: Marquesas Isl. (Ua Huka, Nuku Hiva, Ua Pou, Hiva Oa, Tahuata)
61 MRQ
species :
Dryopteris mangindranensis Tardieu; Notul. Syst. (Paris) 15: 161 (1956)
Distribution: Madagascar
29 MDG
species :
Dryopteris manipurensis (Bedd.) C.Chr.; Ind. 276 (1905)
Distribution: NE-India (Manipur, Arunachal Pradesh, Meghalaya); Nepal; Philippines; New Guinea (Papua New Guinea)
40 IND NEP 42 NWG PHI
Remarks: subgen. Nothoperanema, sect. Shiehia
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Ctenitis copelandii (Christ) Copel.; Gen. Fil. 124 (1947)
= Ctenitis manipurensis (Bedd.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 297 (1938)
= Dryopteris copelandii Christ; Philipp. J. Sci., Bot.2: 216 (1907)
= Dryopsis manipurensis (Bedd.) Holttum & P.J.Edwards; Kew Bull. 41(1): 200 (1986)
= Phegopteris copelandii (Christ) Alderw.; Mal. Ferns 492 (1909)
= Phegopteris manipurensis (Bedd.) Bedd.; Handb. Suppl. 83 (1892)
= Polypodium manipurense Bedd.; J. Bot. 235 (1888)
species :
Dryopteris manniana (Hook.) C.Chr.; Ind. 276 (1905)
Distribution: Guinea; Sierra Leone; Liberia; Ivory Coast; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; Equatorial Guinea (Rio Muni); Uganda; Rwanda; Burundi; Kenya; Tanzania; Malawi; Mozambique; Zimbabwe; Madagascar
22 GUI IVO LBR NGA SIE 23 BIO BUR CMN EQG RWA SAO 25 KEN TAN UGA 26 MLW MOZ ZIM 29 MDG
Synonyms: hidden
= Dryopteris raynalii Tardieu; Adansonia 5: 200, t. 2, f. 1-4 (1965)
= Phegopteris manniana (Hook.) Kuhn; Fil. Afr. 123 (1868)
= Polypodium mannianum Hook.; Sp. 4: 253 (1862)
species :
Dryopteris marginalis (L.) A.Gray; Man. ed. 1: 632 (1848)
Name: Marginal woodfern
Distribution: USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, Wisconsin, West Virginia); Canada (British Columbia, New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec); Greenland
70 GNL 74 USA 75 USA 78 USA
Synonyms: hidden
= Aspidium filix-mas var. marginale (L.) Christ; Farnkr. Erde 257 (1897)
= Aspidium marginale (L.) Sw.; Schrad., J. Bot. 1800(2): 35 (1801)
= Aspidium marginale f. davenportii Eastman; Ferns & Fern Allies New England 27 (1904)
= Aspidium marginale var. elegans J.Rob.; Bull. Essex Inst. 7(3): 51 (1875)
= Aspidium marginatum (L.) Schkuhr; Kr. Gew. 1: 195, t. 45b (1809), emend.
= Dryopteris marginalis f. churchiae (Gilbert) Leeds; Amer. Fern J. 23: 12 (1933)
= Dryopteris marginalis f. davenportii (Eastman) A.Prince ex Weath.; Amer. Fern J. 26: 62 (1936)
= Dryopteris marginalis f. elegans (J.Rob.) F.W.Gray; Amer. Fern J. 14: 9 (1924)
= Dryopteris marginalis f. traillae (G.Lawson) Gruber; Amer. Fern J. 30: 89 (1940)
= Dryopteris marginalis f. tripinnatifida (Clute) Weath.; Amer. Fern J. 26: 61 (1936)
= Dryopteris marginalis var. elegans (J.Rob.) Carhart; Amer. Fern J. 6: 52 (1916)
= Filix marginalis (L.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix marginalis var. bipinnatifida Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 82 (1916)
= Filix-Mas marginalis (L.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Lastrea marginalis (L.) C.Presl; Tent. Pterid. 77 (1836)
= Lastrea marginalis var. traillae G.Lawson
= Nephrodium marginale (L.) Michx.; Fl. bor. Amer. 2: 267 (1803)
= Nephrodium marginale f. bipinnatifidum Clute
= Nephrodium marginale f. davenportii (Eastman) Floyd; Rhodora 4: 245 (1902)
= Nephrodium marginale f. tripinnatifidum Clute; Fern Bull. 15: 21 (1907)
= Nephrodium marginale var. distans Waters; Ferns (Waters) 238 (1903)
= Nephrodium marginale var. elegans (J.Rob.) Gilbert; Working List N. Amer. Pterid. 17 (1901)
= Nephrodium marginale var. traillae (G.Lawson) Gilbert; List N. Amer. Pterid. 17 (1901)
= Nephrodium marginale var. umbilicatum Desv.; Mém. Soc. Linn. Paris 6(3): 260 (1827)
= Polypodium marginale L.; Sp. 2: 1091 (1753)
= Polystichum marginale (L.) Keyserl.; Pol. Cyath. Hb. Bung. 44 (1873)
= Thelypteris marginalis (L.) Nieuwl.; Midl. Naturalist 1: 226 (1910)
= Thelypteris marginalis f. churchiae (Gilbert) Dole; Fl. Vermont, ed. 3: 11 (1937)
= Thelypteris marginalis f. davenportii (Eastman) L.B.Sm.; Rhodora 30: 17 (1928)
= Thelypteris marginalis f. elegans (J.Rob.) Weath.; Proc. Boston Soc. Nat. Hist. 36: 197 (1922)
species :
Dryopteris marginata (Wall. ex Christ) Christ; Phil. J. Sci. 2: 212 (1907)
Distribution: China (Guangxi, Guizhou, Jiangxi, Sichuan, Yunnan); Taiwan; India (Arunachal Pradesh, Assam State, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Pakistan; peninsular Malaysia; Nepal; Bhutan; Myanmar; Vietnam
36 CHI 38 TAI 40 BHU IND NEP PAK 41 MYA VIE 42 MLY
Synonyms: hidden
= Aspidium filix-mas var. marginatum (Wall. ex C.B.Clarke) Christ; Bull. Herb. Boiss. 6: 967 (1898)
= Aspidium marginatum Wall.; Num. List no. 391 (1828) (p. p. ex Christ 1905)
= Dryopteris blinii Lév.; Flore du Kouy-tcheou 490 (1915)
= Dryopteris chiui Ching; Bot. Res. Acad. Sin. 1987(2): 32 (1987)
= Dryopteris grandissima Tagawa; Acta Phytotax. Geobot. 3: 89 (1934)
= Dryopteris leveillei Nakai; Bull. Natl. Sci. Mus. (Tokyo) 31: 17 (1952), non Christ (1909)
= Dryopteris marginata Christ; Philipp. J. Sci., Bot.2: 212 (1907)
= Dryopteris metcalfei Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 463 (1938)
= Lastrea marginata Wall. ex T.Moore; Ind. 96 (1858)
= Nephrodium filix-mas var. marginata Wall. ex C.B.Clarke; Tr. Linn. Soc. II, Bot. I: 521, t. 71 (1880)
= Nephrodium marginatum (C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14. 740, t. 33 (1903)
= Polystichum blinii (H.Lév.) H.Lév. & C.Chr.; Lév., Cat. Pl. Yun-Nan 110 (1916)
species :
Dryopteris mauiensis C.Chr.; Ind. 277 (1905)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Goniopteris clarkei (Baker) Ching; Sunyatsenia 5: 239 (1940)
= Polypodium clarkei Baker; Ann. Bot. 5: 457 (1891) (non Thelypteris clarkei (Bedd.) C. F. Reed)
= Thelypteris mauiensis (C.Chr.) C.F.Reed; Phytologia 17: 291 (1968)
= Thelypteris pacifica C.F.Reed; Phytologia 17: 300 (1968), nom. nov.
species :
Dryopteris maxima (Baker) C.Chr.; Ind. 277 (1905)
Distribution: Fiji (Viti Levu, Ovalau, Taveuni)
60 FIJ
Synonyms: hidden
= Nephrodium maximum Baker; J. Bot. 182 (1886)
= Arachniodes maxima (Baker) Brownlie; Nova Hedwigia Beih. 55 (Pterid. Fl. Fiji): 280 (1977)
species :
Dryopteris maximowicziana (Miq.) C.Chr.; Acta Hort. Gothob. 1(2): 63 (1924)
Distribution: China (Chongqing, Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Zhejiang); Japan; South Korea; Taiwan
36 CHI 38 JAP KOR TAI
Remarks: subgen. Nothoperanema, sect. Shiehia
Synonyms: hidden
= Aspidium matsumurae Makino; Bot. Mag. (Tokyo) (1892)
= Aspidium maximowiczianum Miq.; Ann. Mus. Bot. Lugd.-Bat. 8: 178 (1867)
= Aspidium spectabile Christ; (1901) (non Wall. 1828)
= Ctenitis aureovestita (Rosenst.) Ching; Bull. Fan Mem. Inst. Biol. Bot.8, t. 295 (1938)
= Ctenitis matsumurae (Makino) Koidz.; Acta Phytotax. Geobot. 8: 189 (1939)
= Ctenitis maximowicziana (Miq.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 294 (1938)
= Ctenitis whankanshanensis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 123 (1981)
= Dryopteris aureo-vestita Rosenst.; Hedwigia 66: 343 (1915)
= Dryopteris matsumurae (Makino) C.Chr.; Ind. 277 (1905)
= Dryopsis maximowicziana (Miq.) Holttum & P.J.Edwards; Kew Bull. 41(1): 197 (1986)
= Dryopteris maximowicziana (Miq.) Koidz.; Fl. Symb. Orient. As. 21 (1931) (non Dryopteris maximowiczii)
= Nephrodium matsumurae Makino; Bot. Mag. (Tokyo) 13: 48 (1899)
species :
Dryopteris maximowiczii (Baker) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: Japan; South Korea (incl. Cheju Isl.)
38 JAP KOR
Synonyms: hidden
= Aspidium callopsis Franch. & Sav.; Enum. Pl. Jap. 2: 240, 634 (1879)
= Aspidium commutatum Franch. & Sav.; Enum. Pl. Jap. 2: 240 (1876)
= Byrsopteris maximowiczii (Baker) Morton; Amer. Fern J. 50: 154 (1960)
= Dryopteris callipteris C.Chr.; Bull. Ac. Géogr. Bot. Mans 1909, Mem. 20: 153 (1909)
= Dryopteris callopsis Franch. & Sav.; C. Chr., Ind. 256 (1905) (p. p.)
= Arachniodes maximowiczii (Baker) Ohwi; J. J. Jap. Bot. 37: 76 (1962)
= Lastrea maximowiczii (Baker) T.Moore; Gard. Chr. n. s. 15: 626 (1881)
= Nephrodium callopsis (Franch. & Sav.) Palib.; Acta Hort. Petrop. 19: 140 (1901)
= Nephrodium maximowiczii Baker; Syn. Fil. (Hook. & Baker) ed. 2: 499 (1874)
= Polystichopsis maximowiczii (Baker) Tagawa; J. Jap. Bot. 33: 94 (1958)
= Rumohra callipteris (C.Chr.) C.Chr. ex Tardieu & C.Chr.; Notul. Syst. (Paris) 7: 84 (1938)
= Rumohra maximowiczii (Baker) Ching; Sinensia 5: 62, t. 13 (1934)
species :
Dryopteris maxonii Underw. & C.Chr.; Amer. Fern J. 1: 96 (1911)
Distribution: Guatemala; El Salvador; Costa Rica; Mexico (Colima, Guanajuato, Guerrero, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Sinaloa)
79 MEX 80 COS ELS GUA
Synonyms: hidden
= Dryopteris patula var. moreliae Christ; Notul. Syst. (Paris) 1: 234 (1910)
species :
Dryopteris mayebarae Tagawa; Acta Phytotax. Geobot. 7: 185 (1938)
Distribution: Japan
38 JAP
species :
Dryopteris meghalaica Fraser-Jenk. & Gibby; Acta Bot. Fenn. 164: 6 (1999)
Distribution: NE-India (Manipur, Meghalaya, Nagaland); Myanmar (Kachin)
40 IND 41 MYA
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
species :
Dryopteris melanocarpa Hayata; Ic. Pl. Formos. 4: 163, f. 104 (1914)
Distribution: Taiwan
38 TAI
Remarks: kept separate from D. sparsa by Knapp (Ferns of Taiwan)
species :
Dryopteris microlepis (Bak) C.Chr.; Ind. Fil., Suppl. 36 (1913)
Distribution: China (Guizhou, Yunnan); India (Arunachal Pradesh); Myanmar
36 CHI 40 IND 41 MYA
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris nakanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 413 (1938)
= Nephrodium microlepis Baker; Bull. Misc. Inform. Kew 1908: 10 (1908)
species :
Dryopteris mindshelkensis Pavlov; Vestn. Akad. Nauk Kaz. SSR. 8: 129, f. 31 (1954)
Distribution: Georgia [Caucasus]; ?Armenia; ?Azerbaijan; ?Northern Caucasus; Kazakhstan; Spain; ?Andorra; France; Italy; Serbia & Kosovo; Montenegro; Albania; Macedonia; Greece; Croatia; NW-England; Wales; Scotland; Turkey (Anatolia); N-Iran; E-Afghanistan; Algeria
10 GRB 12 FRA SPA 13 ALB CRO GRC ITA MNT NMA SRB 20 ALG 32 KAZ 33 GEO 34 AFG IRN TUR
Synonyms: hidden
= Dryopteris submontana (Fraser-Jenk. & Jermy) Fraser-Jenk.; Candollea 32(2): 311 (1977)
= Dryopteris villarii subsp. submontana Fraser-Jenk. & Jermy; Fern Gaz. 11: 339 (1977) (basion.)
= Dryopteris villarii subsp. mindshelkensis (Pavlov) Fraser-Jenk.; Candollea 32(2): 311 (1977)
species :
Dryopteris monticola (Makino) C.Chr.; Ind. Fil. 278 (1905)
Distribution: Russian Far East (Kuril Isl., Sakhalin); Japan; North Korea; South Korea (incl. Cheju Isl.); China (Liaoning)
31 RFE 36 CHI 38 JAP KOR NKO
Synonyms: hidden
= Aspidium goldieanum Christ; Bull. Herb. Boiss. II, 2: 828 (1902) (non Hook. (1822))
= Aspidium monticola (Makino) Christ; Bull. Soc. Bot. France 62 Mem. 1. 39 (1905)
= Dryopteris goldieana subsp. monticola (Makino) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 192 (1986)
= Dryopteris submonticola Nakai; Bot. Mag. (Tokyo) 46: 98 (1931)
= Nephrodium erythrosorum var. manshuricum Kom.; Trudy Imp. S.-Peterburgsk. Bot. Sada 20: 120 (1901)
= Nephrodium monticola Makino; Bot. Mag. (Tokyo) 13: 80 (1899)
species :
Dryopteris montigena Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 64 (1941)
Distribution: China (Yunnan)
36 CHI
Synonyms: hidden
= Dryopteris likiangensis Ching; Sunyatsenia 6: 1 (1941) (non Dryopteris lichiangensis (Wright) C. Chr. 1913)
species :
Dryopteris munchii A.R.Sm.; Proc. Calif. Acad. Sci. ser. 4, 40: 218 (1975)
Distribution: Mexico (Chiapas, Hidalgo, Puebla)
79 MEX
species :
Dryopteris namegatae (Kurata) Kurata; J. Geobot. 17: 89 (1969)
Distribution: China (Gansu, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Taiwan; N-India (Arunachal Pradesh)
36 CHI 38 JAP TAI 40 IND
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris dickinsii var. namegatae Kurata; J. Geobot. 7: 115 (1958)
= Dryopteris infrapuberula Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64(1): 28 (1983)
species :
Dryopteris napoleonis (Bory) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: St. Helena
28 STH
Conservation Status: IUCN R
Synonyms: hidden
= Aspidium napoleonis (Bory) Fée; Gen. 292 (1850-52)
= Aspidium petersenii Kunze ex Kuhn; (1868)
= Lastrea napoleonis (Bory) C.Presl; Tent. Pterid. 77 (1836)
= Nephrodium napoleonis Bory; Bél., Voy. Bot. 2: 61, t. 6 (1833)
= Polystichum napoleonis (Bory) Keyserl.; Pol. Cyath. Hb. Bung. 44 (1873)
= Polystichum riparium C.Presl; Tent. Pterid. 8 (1836)
species :
Dryopteris nidus (Baker) Zhang; Phytotaxa 71: 17-27 (2012)
Distribution: China (Guizhou, Sichuan, Yunnan); Tibet; India (Arunachal Pradesh, Sikkim, West Bengal); E-Nepal; Bhutan; Myanmar
36 CHI TIB 40 BHU IND NEP 41 MYA
Remarks: Subgen. Dryopsis. The lumping in Flora of China might be too much in this case, and some species might be valid (Fraser-Jenkins et al. 2018)
Synonyms: hidden
= Aspidium filix-mas var. nidus (Baker) Christ; Bull. Soc. Bot. France 52(Mem. 1): 38 (1905)
= Ctenitis contigua Ching; Acta Phytotax. Sin. 19(1): 119 (1981)
= Ctenitis crassirachis Ching; Acta Phytotax. Sin. 19(1): 119 (1981)
= Ctenitis crenata Ching; Acta Phytotax. Sin. 19(1): 120 (1981)
= Ctenitis dentisora Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 67 (1941)
= Ctenitis nidus (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 287 (1938)
= Ctenitis silaensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 68 (1941)
= Ctenitis submariformis Ching & Chu H.Wang; Acta Phytotax. Sin. 19(1): 122 (1981)
= Ctenitis wantsingshanica Ching & K.H.Shing; Ching & Chu H. Wang, Acta Phytotax. Sin. 123 (1981)
= Ctenitis zayuensis Ching & S.K.Wu; Fl. Xizangica 1: 273 (1983)
= Dryopsis contigua (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 193 (1986)
= Dryopsis crassirachis (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 195 (1986)
= Dryopsis crenata (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 193 (1986)
= Dryopsis dentisora (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 195 (1986)
= Dryopsis nidus (Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 192 (1986)
= Dryopsis silaensis (Ching) Holttum & P.J.Edwards; Kew Bull. 41(1): 197 (1986)
= Dryopsis submariformis (Ching & Chu H.Wang) Holttum & P.J.Edwards; Kew Bull. 41(1): 194 (1986)
= Dryopsis wantsingshanica (Ching & K.H.Shing) Holttum & P.J.Edwards; Kew Bull. 41(1): 194 (1986)
= Dryopteris crassirachis (Ching) Li Bing Zhang; Phytotaxa 71: 21 (2012)
= Dryopteris filix-mas subsp. nidus (Baker) C.Chr.; Ind. Fil. 5: 265 (1905)
= Dryopteris leiboensis Li Bing Zhang; Phytotaxa 71: 22 (2012), nom. nov.
= Dryopteris nokoensis Tagawa; Acta Phytotax. Geobot. 3: 91 (1934)
= Dryopteris nushanensis Li Bing Zhang; Phytotaxa 71: 24 (2012)
= Dryopteris submariformis (Ching & Chu H.Wang) Li Bing Zhang; Phytotaxa 71: 25 (2012)
= Dryopteris wantsingshanica (Ching & K.H.Shing) Li Bing Zhang; Phytotaxa 71: 26 (2012)
= Lastrea filix-mas subvar. nidus (Baker) Bedd.; Handb. Ferns Brit. India 250 (1883)
= Lastrea filix-mas var. nidus (Baker) Bedd.; Handb. Ferns Brit. India, Suppl. 171, t. 372 (1876)
= Lastrea filix-mas var. parallelogramma subvar.nidus Bedd.; Handb. Ferns Brit. India t. 249, t. 129 (1883)
= Nephrodium apiciflorum var. nidus (Baker) C.B.Clarke; Tr. Linn. Soc. II, 1: 518 (1880)
= Nephrodium filix-mas var. nidus Baker; Syn. Fil. (Hook. & Baker) ed. 2: 498 (1874)
= Nephrodium nidus (Baker) C.B.Clarke; J. Linn. Soc. 161: 156 (1876)
species :
Dryopteris nigropaleacea (Fraser-Jenk.) Fraser-Jenk.; Bol. Soc. Broter. 55: 238 (1982)
Distribution: ?Tibet; Pakistan (Kurram, Peshawar, Waziristan, Swat, Chitral, Pakistani Punjab, Hazara, Chitral, Dir, Swat, Murree, Azad Kashmir, Gilgit); India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand); W-Nepal; Bhutan
40 BHU IND NEP PAK
Remarks: reports of D. odontoloma for the S-Himalaya belong here fide Fraser-Jenkins 2008
Synonyms: hidden
= Dryopteris pallida subsp. nigropaleacea Fraser-Jenk.; Candollea 32(2): 316 (1977)
= Dryopteris juxtaposita subsp. nigropaleacea (Fraser-Jenk.) Khullar; J. Indian Bot. Soc. 88(3-4): 207 (2009), et in Indian Fern J. 26(1-2): 91 (2010), without basionym reference
species :
Dryopteris nobilis Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 65 (1941)
Distribution: China (NW-Yunnan); ?Tibet; N-India (West Bengal, Sikkim, Arunachal Pradesh, Assam State, Meghalaya)
36 CHI 40 IND
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
species :
Dryopteris nodosa (C.Presl) Li Bing Zhang; Taxon 61(6): 1208 (2012)
Distribution: Japan (Yakushima Isl.); Borneo (Mt. Kinabalu, Sabah, Sarawak); N-Thailand; Philippines; Sulawesi; Vietnam; Sumatra; Lesser Sunda Isl. (Bali, Flores); Myanmar; Fiji (Viti Levu); Solomon Isl. (Kolombangara, Rendova, Guadalcanal); Bismarck Arch. (New Ireland); New Guinea (Irian Jaya, Papua New Guinea)
38 JAP NNS 41 MYA THA VIE 42 BOR LSI NWG PHI SUL SUM 43 BIS SOL 60 FIJ
Remarks: subgen. Nothoperanema. Separated from D. paleolata by molecular data (Zhang et al. 2012). Cited for Taiwan by Flora of China but excluded by Knapp (Ferns of Taiwan, 2011 ff.)
Synonyms: hidden
= Acrophorus blumei Ching ex C.Chr. & Holttum; Gard. Bull. Str. Settl. 7: 226 (1934), nom. nov.
= Acrophorus nodosus (Blume) C.Presl; Tent. Pterid. 94, t. 3, f. 2 (1836)
= Acrophorus stipellatus (Wall.) T.Moore; Gard. Chr. 135 (1864)
= Aspidium nodosum Blume; Enum. pl. Javae 171 (1828), non Willd.
= Cystopteris gigantea C.Presl; Tent. Pterid. 93 (1836)
= Cystopteris nodosa (C.Presl) Mett.; Ann. Mus. Bot. Lugd.-Bat. 1: 241 (1864)
= Davallia ligulata Wall.; Num. List no. 254 (1828)
= Davallia nodosa (C.Presl) Hook.; Sp. 1: 157 (1846)
= Leucostegia ligulata J.Sm.; Lond. J. Bot. 1: 426 (1842)
= Leucostegia nodosa (C.Presl) Bedd.; Handb. Ferns Brit. India, Suppl. 4 (1870)
= Peranema nodosum (C.Presl) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 317 (2008)
species :
Dryopteris nubigena Maxon & C.V.Morton; Proc. Biol. Soc. Wash. 50: 179 (1937)
Distribution: Mexico (Chiapas, Guerrero, Mexico State, Oaxaca); Guatemala; Honduras; Costa Rica; Panama; El Salvador
79 MEX 80 COS ELS GUA HON PAN
species :
Dryopteris nyingchiensis Ching; Fl. Xizang. 1: 262 (1983)
Distribution: Tibet; Taiwan; etc.
36 TIB 38 TAI
Remarks: kept separate from D. sublacera by Taiwan checklist (2019)
species :
Dryopteris obtusiloba (Baker ex Hook. & Baker) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: Sri Lanka
40 SRL
Remarks: subgen. Nothoperanema, sect. Shiehia
Synonyms: hidden
= Aspidium obtusilobum (Baker ex Hook. & Baker) Prantl; Verh. zool. Bot. Ges. 81: 119 (1882)
= Ctenitis ferruginea var. obtusiloba (Baker) Sledge; Kew Bull. 27: 409 (1972)
= Ctenitis obtusiloba (Baker ex Hook. & Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 296 (1938)
= Dryopsis obtusiloba (Baker ex Hook. & Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 202 (1986)
= Dryopteris peranemiformis subsp. obtusiloba (Baker ex Hook. & Baker) C.Chr.; Ind. 284 (1905)
= Dryopteris zeylanica Alderw.; Mal. Ferns 203 (1909) (nom. nov. superfl.)
= Lastrea obtusiloba (Baker ex Hook. & Baker) Bedd.; Handb. Ferns Brit. India, t. 296 (1868)
= Nephrodium obtusilobum Baker; Syn. 284 (1867)
species :
Dryopteris occidentalis J.P.Roux; Kew Bull. 59(2): 214 (2004)
Distribution: Guinea; Ivory Coast; Nigeria
22 GUI IVO NGA
species :
Dryopteris odontoloma (T.Moore) C.Chr.; Acta Hort. Gothob. 1: 59 (1924)
Distribution: S-India (Tamil Nadu)
40 IND
Conservation Status: IUCN NT for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Lastrea filix-mas var. odontoloma (T.Moore) Bedd.; Handb. Ferns Brit. India, Suppl. 55 (1892)
= Lastrea odontoloma (T.Moore) Bedd.; Ferns South. India 39, pl. 114 (1864)
= Lastrea odontoloma T.Moore; Ind. Fil. 90 (1858)
= Nephrodium odontoloma (T.Moore) Hope; J. Bombay Nat. Hist. Soc. 14: 736 (1903)
species :
Dryopteris oligodonta (Desv.) Pic.Serm.; Webbia 8: 150, f. 2 (1951)
Name: Kanaren-Wurmfarn (DE); Helecho macho (ES)
Distribution: Canary Isl. (Gran Canaria, Tenerife, La Gomera, Hierro, La Palma Isl.); Cape Verde Isl.
21 CNY CVI
Conservation Status: IUCN R (Capverdes)
Synonyms: hidden
= Allantodia oligantha (Desv.) Desv.; Mem. Soc. Linn. Paris 6, 2: 265 (1827)
= Aspidium aquilinoides Bolle; Zeitschr. fil. Erdk. n. Folge 1: 221 (1866)
= Aspidium canariense A.Braun; Flora 708 (1841)
= Aspidium elongatum Willd.; Sp. 5: 269 (1810) (non Sw. (1801) nec All.)
= Aspidium oliganthum Desv.; Berl. Mag. 5: 321 (1811)
= Aspidium oligodonton Desv.; Berl. Mag. 5: 322 (1811)
= Dryopteris elongata var. simplex A.Chev.; Revue de botanique appliquée et d’agriculture tropicale 15,170/171: 328 (seors 1060) (1935)
= Dryopteris oligantha (Desv.) C.Chr.; Ind. Fil., Suppl. III: 93 (1934)
= Lastrea canariensis (A.Braun) T.Moore; Ind. 87 (1858)
= Nephrodium canariense (A.Braun) Kiaersk.; Cat. Fil. hb. ht. Haun. 62 (1874)
= Nephrodium oligodontum (Desv.) Desv.; Mem. Soc. Linn. Paris 6, 2: 261 (1827)
= Polystichum canariense (A.Braun) Keyserl.; Pol. Cyath. Hb. Bung. 45 (1873)
species :
Dryopteris oreades Fomin; Vestn. Tiflissk. Bot. Sada 18: 20 (1911)
Name: Geröll-Dornfarn (DE)
Distribution: N-England; Wales; Scotland; Ireland; Faeroe Isl.; France; Spain; ?Andorra; Italy; Portugal; Norway; Turkey (Anatolia); Northern Caucasus; Azerbaijan; Georgia [Caucasus]; Armenia; Corsica; Sardinia; Germany (Nordrhein-Westfalen)
10 FOR GRB IRE NOR 11 GER 12 COR FRA POR SAR SPA 13 ITA 33 ARM AZE GEO NCS 34 TUR
Remarks: filix-mas agg.
Synonyms: hidden
= Dryopteris abbreviata (DC.) Newman ex Manton; (1950) (p. p. excl. typo, non (Schrad.) Kuntze (1891), nec Polystichum abbreviatum DC. (1805))
= Dryopteris filix-mas subsp. oreades (Fomin) O.Bolòs & Vigo; Fl. Paisos Catalans: 171 (1984)
= Lastrea filix-mas var. pumila T.Moore
= Aspidium pumilum (T.Moore) Lowe; Ferns 6, t. 15 (1857)
= Lastreapumila (T. Moore) J.Sm.
species :
Dryopteris otomasui Kurata; J. Geobot. 15: 83 (1967)
Distribution: Japan
38 JAP
species :
Dryopteris paleolata (Pic.Serm.) Li Bing Zhang; Taxon 61(6): 1208 (2012)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Jiangxi, Sichuan, Yunnan); Taiwan; Tibet; Bhutan; Philippines; Vietnam; India (Arunachal Pradesh, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Uttarakhand, West Bengal); Nepal; Myanmar (Kachin, etc.)
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA VIE 42 PHI
Remarks: subgen. Nothoperanema. The name Acrophorus stipellatus (Wall.) T. Moore, based on Davallia stipellata Wall., a nomen nudum, is not validly published. Synonymy fide Fraser-Jenkins et al. 2018
Synonyms: hidden
= Acrophorus diacalpioides Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 381 (1983)
= Acrophorus dissectus Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 382 (1983) (fide Fraser-Jenkins et al. 2018)
= Acrophorus emeiensis Ching; S. H. Wu, Acta Phytotax. Sin. 21(4): 380 (1983) (fide Fraser-Jenkins et al. 2018)
= Acrophorus exstipellatus Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 383 (1983) (fide Fraser-Jenkins et al. 2018)
= Acrophorus paleolatus Pic.Serm.; Webbia 31 (1): 252 (1977) (paleolulatus auct., errore)
= Davallia stipellata Wall.; Num. List no. 260 (1828), nom. nud.
= Dryopteris acrophorus Li Bing Zhang; Taxon 61(6): 1208 (2012), nom. nov. for Acrophorus diacalpoides Ching & S. H. Wu (fide Fraser-Jenkins et al. 2018)
= Dryopteris emeiensis (Ching) Li Bing Zhang; Taxon 61(6): 1208 (2012) (fide Fraser-Jenkins et al. 2018), non Dryopteris omeiensis (Baker) C. Chr. 1905
= Dryopteris exstipellata (Ching & S.H.Wu) Li Bing Zhang; Taxon 61(6): 1209 (2012) (fide Fraser-Jenkins et al. 2018)
= Dryopteris wusugongii Li Bing Zhang; Taxon 61(6): 1208 (2012), nom. nov. for Acrophorus dissectus Ching & S. H. Wu (fide Fraser-Jenkins et al. 2018)
= Peranema paleolatum (Pic.Serm.) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 317 (2008)
species :
Dryopteris pallida (Bory) Maire & Petitm.; Bull. Séances Soc. Sci. Nancy ser. 3, 9: 480 (1908)
subspecies :
subsp. pallida
Name: Fahler Wurmfarn (DE); Pale Male-fern (EN)
Distribution: Baleares; Sardinia; Sicily; Italy; Bosnia & Hercegovina; Montenegro; Macedonia; Croatia; Albania; Greece (incl. Kiklades); Crete; East Aegaean Isl. (incl. Rhodos); Turkey (Anatolia); Cyprus; Lebanon; Syria; Israel; Palestine; Iran; Tunisia; Algeria
12 BAL SAR 13 ALB BIH CRO GRC ITA KRI MNT NMA SIC 20 ALG TUN 34 CYP EAI IRN ISR LEB PAL SYR TUR
Synonyms: hidden
= Aspidium affine Reichb.; Vis., Fl. Dalm. 1: 39 (1842)
= Aspidium pallidum (Bory) Link; Fil. Sp. 107 (1841) (non Blume (1828))
= Aspidium rigidum var. australe Ten.; Atti Ist. lncor. Napoli 6: 144, t. 2, f. 4 B (1830)
= Dryopteris pallida (Bory) Fomin; Moniteur Jard. Bot. Tiflis, livr. 20, 3: 2 (1910)
= Dryopteris villarii subsp. pallida (Bory) Heywood; Fedde, Repert. 69: 44 (1964)
= Lastrea nivalis T.Moore; Ind. 98 (1858)
= Nephrodium aristatum C.Presl; Fl. Sic. 46 (1826)
= Nephrodium pallidum Bory; Exp. Sci. Morée Bot. 287 (1832) (basion.)
= Nephrodium villarii subsp. pallidum (Bory) Hayek; Fedde, Repert. Beih. 30(1): 27 (1924)
= Polystichum pallidum (Bory) Tod.; Syn. Pl. acot. vasc. Sic. 35 (1866) non Gardner 1842
subspecies :
subsp. balearica (Litard.) Fraser-Jenk.; Candollea 32: 314 (1977)
Distribution: Baleares
12 BAL
Conservation Status: IUCN R
Synonyms: hidden
= Dryopteris balearica (Litard.) E.Nardi; Webbia 30(1): 28 (1976)
= Dryopteris rigida var. balearica Litard.; Bull. Ac. Géogr. Bot. Mans 21: 23 (1911) (basion.)
= Dryopteris villarii subsp. balearica (Litard.) O.Bolòs & Vigo; Fl. Paisos Catalans: 172 (1984)
subspecies :
subsp. libanotica (Rosenst.) E.Nardi; Webbia 32: 97 (1900)
Distribution: Turkey (Anatolia); Cyprus; Lebanon; Syria; Israel; ?Jordania
34 CYP ISR LEB SYR TUR
Synonyms: hidden
= Aspidium libanoticum Rosenst.; Mém. Herb. Boissier 9: 1 (1900) (basion.)
= Dryopteris libanotica (Rosenst.) C.Chr.; Ind. Fil. 275 (1905)
= Dryopteris pallida subsp. libanotica (Rosenst.) Fraser-Jenk.; Candollea 32(2): 315 (1977)
= Nephrodium libanoticum (Rosenst.) Bornm.; Beih. Bot. Centralbl. 86. 11. 279 (1914)
subspecies :
subsp. raddeana (Fomin) Fraser-Jenk.; Candollea 32(2): 315 (1977)
Distribution: Azerbaijan (Talysh)
33 AZE
Synonyms: hidden
= Dryopteris pallida subsp. raddeana (Fomin) E.Nardi; Webbia 32(1): 97 (1977) (comb. superfl.)
= Dryopteris raddeana (Fomin) Fomin; Kuzn. & al., Fl. Caucas. Crit. l(1): 57 (1911), et Mon. Jardin bot. Tiflis, livr. 20. 33 (1911)
= Nephrodium raddeanum Fomin; Moniteur Jard. Bot. Tiflis, livr. 12: 8, t. 2 (1908)
= Dryopteris talyschensis (Askerov & A.E.Bobrov) Askerov & Aktcay; Turczaninowia 19(1): 81 (2017)
= Dryopteris raddeana var. talyschensis Askerov & A.E.Bobrov; Bot. Zhurn. (Moscow & Leningrad) 57(10): 1297 (1972)
species :
Dryopteris panda (C.B.Clarke) C.Chr.; Ind. Fil., Suppl. III: 94 (1934)
Distribution: China (Gansu, Guizhou, Sichuan, Yunnan); Tibet; India (Himachal Pradesh, Uttarakhand, Sikkim, West Bengal, Jammu & Kashmir); Nepal; Pakistan (Azad Kashmir)
36 CHI TIB 40 IND NEP PAK
Synonyms: hidden
= Dryopteris bonatiana (Brause) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 363 (1989) (fide Fraser-Jenkins 1997)
= Dryopteris filix-mas subsp. panda (C.B.Clarke) C.Chr.; Ind. Fil. 5: 265 (1905) ex Ching (1938)
= Dryopteris yui Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 417 (1938) (fide Fraser-Jenkins 1997)
= Lastrea filix-mas var. panda (C.B.Clarke) Bedd.; Handb. Ferns Brit. India 251 (1883)
= Nephrodium filix-mas var. panda C.B.Clarke; Tr. Linn. Soc. II, 1: 519, t. 68, f. 1 (1880)
= Nephrodium pandum (C.B.Clarke) Hope; J. Bombay Nat. Hist. Soc. 12: 623 (1899)
= Polystichum bonatianum Brause; Hedwigia 54. 200, t. 4 B (1914) (fide Fraser-Jenkins 1997)
species :
Dryopteris papuae-novae-guineae Li Bing Zhang; Taxon 61(6): 1209 (2012), nom. nov.
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Acrophorus stipellatus var. montanus Rosenst.; Repert. Spec. Nov. Regni Veg. 12: 166 (1913)
species :
Dryopteris papuana C.Chr.; Ind. 282 (1905)
Distribution: Solomon Isl. (Kolombangara, Rendova); Bismarck Arch. (New Ireland); New Guinea (Papua New Guinea)
42 NWG 43 BIS SOL
species :
Dryopteris paralunanensis S.G.Lu ex W.M.Chu; Guihaia 11(3): 225 (1991)
Distribution: China (Yunnan)
36 CHI
species :
Dryopteris patula (Sw.) Underw.; Native Ferns ed. 4: 177 (1893)
Distribution: Mexico (Chiapas, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, Sinaloa, Veracruz); Guatemala; Honduras; El Salvador; Nicaragua; Costa Rica; Panama; Colombia (Antioquia, Boyacá, Cundinamarca, Guainía, Magdalena, Nariño); Venezuela (Aragua, Lara, Merida, Miranda, Portuguesa, Sucre, Tachira); Ecuador; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Brazil (Acre, Goiás, Maranhão, Minas Gerais, Mato Grosso, Pará, Rio de Janeiro, Rondônia, Rio Grande do Sul, São Paulo); Jamaica; Hispaniola; Galapagos
79 MEX 80 COS ELS GUA HON NIC PAN 81 HIS JAM 82 CLM VEN 83 BOL ECU GAL PER 84 BRA
Synonyms: hidden
= Aspidium apertum Fée; Mém. Foug. 8: 106 (1857)
= Aspidium chaerophylloides Moritz; Mett., Pheg. u. Asp. 66 (1858) (syn.)
= Aspidium diastematocarpon (Fée) Fée; Mém. Foug. 8: 107 (1857)
= Aspidium imbecile Kunze ex Ettingsh.; Farnkr. 176, t. 107, f. 10, t. 109, f. 1 (1865)
= Aspidium leptorachis Kunze; Linn. 18: 346 (1844)
= Aspidium mexicanum var. serratum Mett.; Abh. Senckenberg. Naturf. Ges. 2: 349 (1858)
= Aspidium patulum Sw.; Kongl. Vetensk. Acad. Handl. 1817: 64 (1817)
= Aspidium patulum var. chaerophylloides (Baker ex Mart.) J.Bommer & Christ; Bull. Soc. Roy. Bot. Belgique 35, pt. 1: 214 (1896)
= Aspidium paupertinum Roem.; Kunze, Linn. 18. 345 (1844)
= Aspidium polystichoides (C.Presl) Mett.; Salomon, Nom. 67 (1883)
= Dryopteris aperta (Fée) C.Chr.; Ind. 252 (1905)
= Dryopteris patula var. chaerophylloides (Baker ex Mart.) Hieron.; Hedwigia 46: 345 (1907)
= Dryopteris patula var. serrata (Mett.) C.V.Morton; Contr. U. S. Natl. Herb. 38(6): 219 (1973)
= Dryopteris simplicior Mickel & Beitel; Pterid. Fl. Oaxaca (Mem. New York Bot. Gard. 46): 166 (1988)
= Filix patula (Sw.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix-mas patula (Sw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Filix-mas patula var. chaerophylloides (Baker ex Mart.) Farw.; Amer. Midl. Naturalist 12: 256 (1931)
= Lastrea aperta (Fée) T.Moore; Ind. 85 (1858)
= Lastrea diastematocarpa (Fée) T.Moore; Ind. 90 (1858)
= Lastrea leptorachis (Kunze) Liebm.; Vid. Selsk. Skr. V, 1: 272 (seors. 120) (1849)
= Lastrea patula (Sw.) T.Moore; Ind. 99 (1858)
= Lastrea paupertina (Roem.) Liebm.; Vid. Selsk. Skr. V, 1: 272 (seors. 120) (1849)
= Lastrea polystichoides C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 38 (1851)
= Nephrodium patulum (Sw.) Baker; Mart., Fl. Bras. 1(2): 480 (1870)
= Nephrodium patulum var. chaerophylloides Baker; Mart., Fl. Bras. 1(2): 481 (1870)
= Nephrodium patulum var. robustior Donn.Sm.; Enum. Pl. Guatem. 3: 101 (1893)
species :
Dryopteris paucisora Copel.; Philipp. J. Sci., Bot.6: 136 (1911)
Distribution: Borneo (Mt. Kinabalu, Sabah, Sarawak)
42 BOR
Remarks: Subgen. Dryopsis
Synonyms: hidden
= Ctenitis paucisora (Copel.) Copel.; Gen. Fil. 124 (1947)
= Dryopsis paucisora (Copel.) Holttum & P.J.Edwards; Kew Bull. 41(1): 183 (1986)
species :
Dryopteris pauliae Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 140 (1997)
Distribution: India (Himachal Pradesh, Uttarakhand); Nepal
40 IND NEP
species :
Dryopteris peninsulae Kitag.; Rep. First Sci. Exped. Manchoukuo 4(2): 54 (1935)
Distribution: China (Gansu, Guizhou, Henan, Hubei, Jiangxi, Liaoning, Shaanxi, Shandong, Sichuan, NE-Yunnan)
36 CHI
Synonyms: hidden
= Dryopteris apicisora Ching & Y.T.Hsieh; Fl. Tsinlingensis 2: 228 (1974)
= Dryopteris cathayana Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 1 (1984)
= Dryopteris lacera var. peninsulae Tagawa
= Dryopteris neolacera Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 440 (1938)
= Dryopteris shensicola Ching & Y.T.Hsieh; Fl. Tsinlingensis 2: 229 (1974)
species :
Dryopteris pentheri (Krasser) C.Chr.; Ind. 284 (1905)
Distribution: Cape Verde Isl. (Ilha de Sao Nicolau); Guinea; Guinea-Bissau; Sierra Leone; Liberia; Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; São Tomé; D.R.Congo [Zaire]; Sudan; Ethiopia; Uganda; Rwanda; Burundi; Kenya; Tanzania; Angola; Zambia; Malawi; Mozambique; Zimbabwe; South Africa (Gauteng, Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov., W-Cape Prov., Free State); Lesotho; Eswatini [Swaziland]; Madagascar; La Réunion
21 CVI 22 GNB GUI LBR NGA SIE 23 BIO BUR CMN RWA SAO ZAI 24 ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 LES SAF SWZ 29 MDG REU
Synonyms: hidden
= Nephrodium pentheri Krasser; Ann. Hofmus. Wien 16. 5, t. 2 t 1-5 (1900)
= Dryopteris elongata Sim; Ferns S. Afr. ed. 2: 104 (1916) (non Dryopteris elongata (Wall. ex Hook.) Kuntze)
species :
Dryopteris peranema Li Bing Zhang; Taxon 61(6): 1211 (2012), nom. nov.
Distribution: China (Guizhou, Guangxi, Hubei, Hunan, Sichuan, Yunnan); Philippines (Benguet: Mt. Pulag); Tibet; Taiwan; Myanmar (Chin, etc.); Vietnam; India (Arunachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Bhutan; Nepal
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA VIE 42 PHI
Remarks: subgen. Nothoperanema. Inclusion of D. zhuweimingii fide Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Dryopteris zhuweimingii Li Bing Zhang; Taxon 61(6): 1211 (2012), nom. nov. for Peranema luzonicum Copel.
= Peranema cyatheoides D.Don; Prodr. Fl. Nepal. 12 (1825)
= Peranema cyatheoides var. luzonicum (Copel.) Ching & S.H.Wu; Acta Phytotax. Sin. 21(4): 373-374 (1983)
= Peranema formosanum Hayata; Bot. Mag., Tokyo 26: 110 (1912), non Dryopteris formosana (Christ) C. Chr. 1905
= Peranema luzonicum Copel.; Phil. J. Sci. 4(2): 111 (1909)
= Sphaeropteris barbata Wall.; Num. List no. 183 (1828)
species :
Dryopteris peranemiformis C.Chr.; Ind. 284 (1905)
Distribution: S-India (Kerala, Tamil Nadu: Nilgiri Hills, Annamalai Hills)
40 IND
Remarks: Subgen. Nothoperanema, sect. Shiehia
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Ctenitis ferruginea (T.Moore ex Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 296 (1938)
= Dryopsis ferruginea (T.Moore ex Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 201 (1986)
= Dryopteris ferruginea (T.Moore ex Baker) Kuntze; Rev. Gen. Pl. 2: 812 (1891), nom. inval.
= Lastrea ferruginea (T.Moore ex Baker) Bedd.; Ferns South. India, t. 100 (1863) (non Aspidium ferrugineum Fée (1850))
= Nephrodium ferrugineum T.Moore ex Baker; Syn. 283 (1867)
species :
Dryopteris permagna M.G.Price; Gard. Bull. Singapore 30: 249 (1977)
Distribution: Philippines (Kalinga, Benguet, Mountain Province, South Cotabato)
42 PHI
Conservation Status: EN in the Philippines
species :
Dryopteris podophylla (Hook.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: China (Fujian, Guangdong, Guangxi, Hainan, Yunnan)
36 CHI
Synonyms: hidden
= Aspidium podophyllum Hook.; J. Bot. 6: 236, t. 1 (1863)
= Dryopteris podophylla f. auriculata Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 400 (1938)
= Lastrea podophylla (Hook.) Hook.; J. Bot. (Hook.) Kew Misc. 9: 339 (1857), non J. Sm. (1857)
= Nephrodium podophyllum (Hook.) Hook.; Sp. 4: 87 (1862)
species :
Dryopteris poilanei Tardieu; Bull. Soc. Bot. France 87: 368 (1941)
Distribution: Vietnam
41 VIE
Remarks: provisionally accepted name
species :
Dryopteris polita Rosenst.; Repert. Spec. Nov. Regni Veg. 18: 218 (1914)
Distribution: China (Hainan, Yunnan); Taiwan; S-Japan (Yakushima Isl.); Sumatra; New Guinea (Papua New Guinea); peninsular Malaysia (Cameron Highlands, Pahang); Borneo (Mt. Kinabalu, Sabah); Thailand; Cambodia; Vietnam; Philippines; Sulawesi
36 CHI 38 JAP NNS TAI 41 CBD THA VIE 42 BOR MLY NWG PHI SUL SUM
Remarks: subgen. Acrorumohra
Synonyms: hidden
= Dryopteris chapaensis C.Chr. & Ching ex Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 474 (1938)
= Dryopteris exindusiata Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 32 (1987)
= Dryopteris laurisilvicola T.Suzuki; J. Jap. Bot. 11: 646, f. 2 (1935)
= Dryopteris lividis Ching & Chu H.Wang; Acta Phytotax. Sin. 8: 168. t.22(26)
= Dryopteris taitoensis Tagawa; Acta Phytotax. Geobot. 4: 134 (1935)
= Dryopteris viridis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 474 (1938)
= Phegopteris polita (Rosenst.) Alderw.; Mal. Ferns Suppl. 309 (1917)
species :
Dryopteris polylepis (Franch. & Sav.) C.Chr.; Ind. 285 (1905)
Distribution: Japan; South Korea
38 JAP KOR
Remarks: not in China
Synonyms: hidden
= Aspidium polylepis Franch. & Sav.; Enum. Pl. Jap. 2: 236, 631 (1879)
= Nephrodium filix-mas var. polylepis (Franch. & Sav.) Makino; Bot. Mag. (Tokyo) 15: 66 (1901)
= Nephrodium polylepis (Franch. & Sav.) Baker; Ann. Bot. 5: 320 (1891)
species :
Dryopteris pontica (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 7 (2007)
Distribution: NW-Iran; N-Turkey (Anatolia); ?Armenia
34 IRN TUR
Remarks: affinis agg.
Synonyms: hidden
= Dryopteris affinis subsp. pontica Fraser-Jenk.; Ann. Bot. Fenn. 33: 73 (1996)
species :
Dryopteris porosa Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 460 (1938)
Distribution: China (Guizhou, Sichuan, Yunnan); N-Thailand; ?Bhutan; ?
36 CHI 41 THA
species :
Dryopteris protobissetiana K.Hori & N.Murak.; Acta Phytotax. Geobot. 66(1): 54 (2015)
Distribution: Japan (Kyushu)
38 JAP
species :
Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang; Taxon 61(6): 1209 (2012)
Distribution: China (Sichuan, Hainan, Yunnan); Tibet; Taiwan; India (Arunachal Pradesh, ?Assam, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, Tripura, West Bengal); Nepal; Bhutan; Myanmar (Kachin, Chin, etc.); Sri Lanka; Vietnam; Sumatra; Java; Borneo (Sabah); New Guinea (Irian Jaya, Papua New Guinea); Thailand (widespread); Laos; peninsular Malaysia; Philippines; Sulawesi; Lesser Sunda Isl. (Sumbawa)
36 CHI TIB 38 TAI 40 BHU IND NEP SRL 41 LAO MYA THA VIE 42 BOR JAW LSI MLY NWG PHI SUL SUM
Remarks: subgen. Nothoperanema. Taxonomy fide Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Cyathea aspidioides (Blume) Moritz; Syst. Verz. 1842-1844: 108 (1854)
= Diacalpe adscendens Ching ex S.H.Wu; Acta Phytotax. Sin. 21(4): 378 (1983), nom. illeg.
= Diacalpe aspidioides Blume; Enum. Pl. Jav. 241 (1828)
= Diacalpe aspidioides var. himalayensis Wall. ex Ching & S.K.Wu; S. H. Wu, Acta Phytotax. Sin. 21: 376 (1983), nom. nud.
= Diacalpe aspidioides var. hookeriana Wall. ex Ching & S.K.Wu; Fl. Xizangica 1: 194, t. 43, f. 1-3, t. 5, f. 7-8 (1983), nom. nud.
= Diacalpe aspidioides var. minor Ching ex S.K.Wu; Acta Phytotax. Sin. 21(4): 375, t. 1, f. 1 (1983)
= Diacalpe caudifolia Ching & S.K.Wu; Fl. Xizangica 1: 194 (1983)
= Diacalpe chinensis Ching & S.H.Wu; Acta Phytotax. Sin. 21(4): 376 (1983)
= Diacalpe hookeriana Wall. ex T.Moore; Gard. Chr. 1854: 135 (1854), nom. nud.
= Diacalpe laevigata Ching & S.H.Wu; Acta Phytotax. Sin. 21(4): 377 (1983)
= Diacalpe medogensis Ching & S.K.Wu; Fl. Xizangica 1: 196 (1983)
= Diacalpe omeiensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 2, 1: 293 (1949) (fide Fraser-Jenkins et al. 2018)
= Diacalpe pseudocaenopteris Kunze; Bot. Zeit. (Berlin) 4: 457 (1846)
= Dryopteris adscendens (Ching ex S.H.Wu) Li Bing Zhang; Taxon 61(6): 1210 (2012)
= Dryopteris diacalpe Li Bing Zhang; Taxon 61(6): 1210 (2012), nom. nov. for Diacalpe chinensis Ching & S. H. Wu, non Dryopteris chinensis (Baker) Koidz.
= Dryopteris hookeriana (Wall. ex T.Moore) Li Bing Zhang; Taxon 61(6): 1210 (2012)
= Dryopteris kungiana Li Bing Zhang; Taxon 61(6): 1210 (2012), nom. nov. for Diacalpe omeiensis Ching
= Dryopteris medogensis (Ching & S.K.Wu) Li Bing Zhang; Taxon 61(6): 1210 (2012)
= Peranema aspidioides (Blume) Mett.; Fil. Lechl. 2: 33 (1850)
= Peranema pseudocaenopteris (Kunze) Panigrahi; Pl. Sci. Res., Orissa 20(1&2): 43 (1998), as ´pseudocystopteris´
= Physematium aspidioides (Blume) Kunze; Analecta Pteridogr. 43-44 (1837)
= Sphaeropteris hookeriana Wall.; List 248 (1828) (incorrect number, nom. nud.)
species :
Dryopteris pseudocomplexa (Fraser-Jenk.) P.D.Sell; Fl. Great Britain and Ireland 1: 671 (2018)
Distribution: Scotland (Hebrides); Germany (Sachsen-Anhalt); Ireland; France (Vosges); Norway; ?Austria
10 GRB IRE NOR 11 GER 12 FRA
Remarks: affinis agg., a fertile tetraploid probably deserving species rank
Synonyms: hidden
= Dryopteris affinis subsp. pseudocomplexa (Fraser-Jenk.) Fraser-Jenk.; Indian Fern J. 39(1): 120 (2022)
= Dryopteris cambrensis subsp. pseudocomplexa Fraser-Jenk.; Fern Gaz. 18(1): 6 (2007)
= Dryopteris pseudocomplexa (Fraser-Jenk.) Fraser-Jenk.; Indian J. Forestry 43(1): 83 (2020), isonym
species :
Dryopteris pseudodisjuncta (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 7 (2007)
Distribution: Switzerland; Austria; S-Germany (Baden-Württemberg, Bayern, Hessen, Nordrhein-Westfalen); Belgium; Luxembourg; Norway; SW-Scotland
10 GRB NOR 11 AUT BGM GER LUX SWI
Remarks: affinis agg.
Synonyms: hidden
= Dryopteris affinis subsp. pseudodisjuncta (Fraser-Jenk.) Fraser-Jenk.; Nordic J. Bot. 16(1): 4 (1996)
= Dryopteris affinis var. pseudodisjuncta Fraser-Jenk.; Hegi, Ill. Fl. Mitteleur. ed. 3 l(1): 147 (1984 publ. 1983)
= Dryopteris affinis var. setosa (Christ) Fraser-Jenk.; Sommerfeltia 6: xi (1987)
= Aspidium filix-mas var. setosum Christ
species :
Dryopteris pseudofilix-mas (Fée) Rothm.; Candollea 10: 96 (1945)
Distribution: Mexico (Chiapas, Ciudad de Mexico, Hidalgo, Mexico State, Michoacan, Morelos, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sonora, Tamaulipas, Veracruz); Guatemala
79 MEX 80 GUA
Synonyms: hidden
= Aspidium chrysocarpon Fée; Mém. Foug. 8: 103 (1857)
= Aspidium paleaceum var. majus E.Fourn.; Mexic. Pl. 1: 93 (1872)
= Aspidium pseudofilix-mas Fée; Mém. Foug. 8: 103 (1857)
= Dryopteris ×chrysocarpa (Fée) Rothm.; Candollea 10: 95 (1945)
= Lastrea chrysocarpa (Fée) T.Moore; Ind. 88 (1858)
species :
Dryopteris pseudolunanensis Tagawa; Acta Phytotax. Geobot. 9: 145 (1940)
Distribution: Taiwan
38 TAI
Conservation Status: very rare
Synonyms: hidden
= Dryopteris kuratae Nakaike ex Hoshiz. & K.A.Wilson; Amer. Fern J. 89(1): 10 (1999), nom. superfl.
species :
Dryopteris pseudoparasitica Alderw.; Nova Guinea 14: 19 (1924)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Synonyms: hidden
= Cyclosorus pseudoparasiticus (Alderw.) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 248 (1941)
= Thelypteris pseudoparasitica (Alderw.) C.F.Reed; Phytologia 17: 306 (1968)
species :
Dryopteris pseudosieboldii Hayata; Ic. Pl. Formos. 4: 171-172, f. 111 (1914)
Distribution: Taiwan
38 TAI
Remarks: treated as full species by Knapp (2013), included as variety in D. enneaphylla by Flora of China
Synonyms: hidden
= Dryopteris enneaphylla var. pseudosieboldii (Hayata) Tagawa & K.Iwats.; Amer. Fern J. 50: 102 (1960)
= Dryopteris heteroneura (Tagawa) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 398 (1938)
= Dryopteris sieboldii var. heteroneura Tagawa; J. Jap. Bot. 12(7): 487 (1936)
species :
Dryopteris pseudosparsa Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 489 (1938)
Distribution: China (Yunnan, Guizhou, Guangxi, Sichuan); N-Thailand; India (Meghalaya); Myanmar (Kachin, etc.)
36 CHI 40 IND 41 MYA THA
Synonyms: hidden
= Dryopteris cuneatiloba Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 11 (1987)
species :
Dryopteris pudouensis Ching; Bull. Bot. Res. Harbin 3(3): 11 (1983)
Distribution: China (Anhui, Fujian, Hunan, Jiangsu, Jiangxi, Zhejiang); Japan; South Korea (incl. Cheju Isl.)
36 CHI 38 JAP KOR
Synonyms: hidden
= Dryopteris hikonensis (H.Itô) Nakaike; New Fl. Japan, Pterid. 841 (1992)
= Dryopteris hikonensis monstr.cristata (T.Ueno) Nakaike; New Fl. Japan, Pterid. 841 (1992)
= Dryopteris lungjingensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 27 (1987)
= Dryopteris pacifica (Nakai) Tagawa; Colenso Ill. Jap. Pterid. 211 (1959), nom. illeg., non Christ (1912)
= Dryopteris quadrifida Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3), 49 (1990)
= Dryopteris varia var. hikonensis (H.Itô) Kurata; Enum. Jap. Pterid. 304 (1961)
= Dryopteris yushanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 28 (1987)
= Polystichum hololepis var. hikonensis H.Itô; J. Jap. Bot. 10: 451 (1961)
= Polystichum pacificum Nakai; Bot. Mag. (Tokyo) 39: 119 (1925)
species :
Dryopteris pulvinulifera (Bedd.) Kuntze; Rev. Gen. Pl. 2: 213 (1891)
Distribution: Sri Lanka; China (NW-Yunnan); Philippines; India (Uttarakhand, Sikkim, Darjeeling, Meghalaya, Nagaland, Arunachal Pradesh, Assam State, Mizoram, West Bengal); Nepal; Bhutan
36 CHI 40 BHU IND NEP SRL 42 PHI
Synonyms: hidden
= Dryopteris harae Itô; Hara, Fl. E. Himal. 476, t. 26 (1966)
= Dryopteris reholttumii M.G.Price; Fiddlehead Forum 8(3): 19 (1981), nom. nud.
= Lastrea pulvinulifera Bedd.; Ferns Brit. India t. 333 (1870)
= Lastrea pulvinulifera var. zeylanica Bedd.; Handb. Ferns Brit. India, Suppl. 17 (1876)
= Lastrea sparsa var. zeylanica (Bedd.) Bedd.; Handb. Ferns Brit. India 254 (1883)
= Nephrodium pulvinuliferum (Bedd.) Baker; Syn. Fil. 500 (1874) (p. p.)
= Nephrodium sparsum var. squamulosum C.B.Clarke; Tr. Linn. Soc. II, 1: 524 (1880) (p. p.)
= Polypodium marginatum T.Moore; Ind. 297 (1861)
species :
Dryopteris purpurascens (Blume) Christ; Philipp. J. Sci., Bot.2: 213 (1907)
Distribution: Borneo (Mt. Kinabalu, Sabah); Philippines; Sulawesi; New Guinea (Papua New Guinea)
42 BOR NWG PHI SUL
Synonyms: hidden
= Aspidium purpurascens Blume; Enum. Fil. 169 (1828)
= Nephrodium purpurascens (Blume) Hook.; Sp. 4: 132, t. 262
species :
Dryopteris pycnopteroides (Christ) C.Chr.; Ind. Fil., Suppl. 38 (1913)
Distribution: China (Sichuan, Guizhou, Yunnan, Hubei); ?Japan
36 CHI
Synonyms: hidden
= Aspidium pycnopteroides Christ; Bull. Ac. Géogr. Bot. Mans 116 (1906)
= Dryopteris sunii Ching; Bull. Fan Mem. Inst. Biol., n. s., 1: 295 (1949) (fide Fraser-Jenkins (1986))
species :
Dryopteris raiateensis (J.W.Moore) Li Bing Zhang; Taxon 61(6): 1209 (2012)
Distribution: Society Isl. (Raiatea)
61 SCI
Remarks: sometimes included in D. leucorhachis from Rarotonga
Synonyms: hidden
= Acrophorus raiateensis J.W.Moore; Bull. Bernice P. Bishop Mus. 102: 6 (1933)
species :
Dryopteris ramosa (Hope) C.Chr.; Ind. Fil. 266 (1905)
Distribution: India (Himachal Pradesh, Jammu & Kashmir, Uttarakhand); Nepal; Bhutan; E-Afghanistan; Pakistan (Kurram, Chitral, Swat, Hazara, Gilgit)
34 AFG 40 BHU IND NEP PAK
Synonyms: hidden
= Lastrea rigida Bedd.; Handb. Ferns Brit. India 251 (1883)
= Nephrodium ramosum Hope; J. Bot. 1896 126 (1896)
species :
Dryopteris rarissima Kurata; J. Geobot. 13: 43 (1964)
Distribution: Japan
38 JAP
species :
Dryopteris redactopinnata Basu & Panigrahi; Indian J. Forest. 3 (3): 270, t. 2 (1980)
Distribution: Bhutan; India (Jammu & Kashmir, Sikkim, Himachal Pradesh, Uttarakhand, Arunachal Pradesh); Nepal; Pakistan (Hazara); China (Yunnan, Sichuan); Tibet; Taiwan
36 CHI TIB 38 TAI 40 BHU IND NEP PAK
Synonyms: hidden
= Dryopteris pseudofibrillosa Ching; Fl. Xizangica 1: 252 (1983)
species :
Dryopteris reflexosquamata Hayata; Ic. Pl. Formos. 4: 176, f. 114 (1914)
Distribution: Taiwan; China (Sichuan, Guizhou, Hunan, NE-Yunnan); ?India (?Sikkim)
36 CHI 38 TAI
Synonyms: hidden
= Dryopteris mingetsuensis Hayata; Ic. Pl. Formos. 6: 281, f. 109 (1916)
= Dryopteris paludicola Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(3): 21 (1984)
species :
Dryopteris remota (Döll) Druce; List Brit. Pl.: 87 (1908)
Name: Entferntfiedriger Wurmfarn, Entferntfiedriger Dornfarn (DE)
Distribution: +Ireland; +Scotland; Germany (Baden-Württemberg, Bayern); Switzerland; ?Liechtenstein; Czech Republic; ?Austria; Spain; France; Slovenia; Croatia; Turkey (Anatolia); European Russia; Northern Caucasus; Georgia [Caucasus]
10 GRB IRE 11 CZE GER SWI 12 FRA SPA 13 CRO SLV 14 RUS 33 GEO NCS 34 TUR
Remarks: filix-mas agg.
Synonyms: hidden
= Aspidium filix-mas var. remotum (Döll) Jess.; 613 (1879)
= Aspidium filix-mas × spinulosum A.Braun
= Aspidium rigidum var. remotum A.Braun; Döll, Rhein. Fl. 16 (1843)
= Aspidium rigidum var. remotum Döll; Rhein. Fl.: 16 (1843) (basion.)
= Dryopteris kemulariae Mikhel.; Zametki Sist. Geogr. Rast. 23: 56, f. 1 (1963)
= Dryopteris ×remota (Döll) Hayek; Fl. Steiermark 56 (1908)
= Lastrea remota (Döll) T.Moore; Ind. 102 (1858)
= Nephrodium borbasio E.Walter in Guétrot; 104 (1929)
= Nephrodium remoti Guétrot; 37, var. orthogr. (1927)
= Nephrodium remotum (Döll) Murr; (1923), non Heward nec Hook.
= Nephrodium spinulosum (A.Braun) Baker; (1867)
= Nephrodium spinulosum subsp. remotum (Döll) Hook.fil.; (1870)
species :
Dryopteris remotissima (Christ) Koidz.; Bot. Mag. (Tokyo) 43: 386 (1929)
Distribution: Japan
38 JAP
Synonyms: hidden
= Aspidium spinulosum var. remotissimum Christ; Bot. Mag. (Tokyo) 24: 240 (1910)
species :
Dryopteris renchangiana Z.Y.Zuo & D.Z.Li; Pl. Diversity 44(2): 185 (2021)
Distribution: China (Chongqing, Sichuan)
36 CHI
Synonyms: hidden
= Dryopteris glabrior Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 11 (1984), nom. illeg.
species :
Dryopteris rhomboideo-ovata H.Itô; J. Jap. Bot. 9: 55 (1933)
Distribution: Japan
38 JAP
species :
Dryopteris rodolfii J.P.Roux; Webbia 59(1): 143 (2004)
Distribution: Ethiopia; Kenya
24 ETH 25 KEN
species :
Dryopteris rossii C.Chr.; H. Ross, Mem. Acad. Nac. Ci. Antonio Alzate 32: 178 (1912)
Name: Ross´ woodfern
Distribution: USA (Arizona); Nicaragua; Costa Rica; Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chihuahua, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Queretaro, San Luis Potosi, Sinaloa, Zacatecas)
76 USA 79 MEX 80 COS NIC
Conservation Status: IUCN V (Arizona: E)
Synonyms: hidden
= Dryopteris patula var. rossii (C.Chr.) C.Chr.; (1912 publ. 1913)
species :
Dryopteris rosthornii (Diels) C.Chr.; Ind. 289 (1905)
Distribution: China (Gansu, Guizhou, Hubei, Shanxi, Sichuan, Yunnan)
36 CHI
Remarks: not in India
Synonyms: hidden
= Aspidium filix-mas var. omeiense Christ; Bull. Ac. Géogr. Bot. Mans 1906: 117 (1906)
= Dryopteris centrochinensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 423 (1938)
= Dryopteris filix-mas var. rosthornii (Diels) C.Chr.; Acta Hort. Gothob. 1: 58 (1924)
= Dryopteris hupehensis Ching; Fl. Tsinlingensis 2: 226 (1974)
= Dryopteris omeicola Ching; Acta Phytotax. Sin. 10: 303 (1965)
= Nephrodium rosthornii Diels; Engl. Jahrb. 29: 190 (1900)
species :
Dryopteris rubiginosa (Brack.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Remarks: subgen. Nothoperanema
Synonyms: hidden
= Aspidium rubiginosum (Brack.) Mann; Proc. Amer. Acad. 7: 217 (1868)
= Ctenitis rubiginosa (Brack.) Copel.; Gen. Fil. 125 (1947)
= Nothoperanema rubiginosum (Brack.) A.R.Sm. & D.D.Palmer; Amer. Fern J. 85(2): 63 (1995)
= Lastrea rubiginosa Brack.; U. S. Expl. Exp. 16: 201 (1854)
= Nephrodium rubiginosum (Brack.) Hook.; Sp. 4: 143 (1862)
species :
Dryopteris rubrobrunnea W.M.Chu; Acta Bot. Yunnan., Suppl. 5: 46 (1992)
Distribution: China (Yunnan); NE-India (Arunachal Pradesh); Myanmar (Kachin, etc.)
36 CHI 40 IND 41 MYA
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Dryopteris rubripes Ching & Chu ex Fraser-Jenk.; (1986)
species :
Dryopteris ruwenzoriensis C.Chr. ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 204 (1986)
Distribution: D.R.Congo [Zaire]; Uganda; Rwanda
23 RWA ZAI 25 UGA
species :
Dryopteris ryo-itoana Kurata; J. Geobot. 15: 84 (1967)
Distribution: China (Jiangxi, Zhejiang); Japan; Taiwan
36 CHI 38 JAP TAI
Remarks: new record for Taiwan see Knapp (2013)
Synonyms: hidden
= Dryopteris dispar Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 17 (1983)
= Dryopteris gutishanensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 29 (1983)
= Dryopteris kaihuaensis Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 24 (1983)
= Dryopteris lushanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 20 (1987)
= Dryopteris pseudofuscipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 3 (1987)
= Dryopteris triangularifrons Ching; Bull. Bot. Res. Harbin 2(2): 64 (1982)
species :
Dryopteris sabaei (Franch. & Sav.) C.Chr.; Ind. 290 (1905)
Distribution: S-Japan (incl. Yakushima Isl.)
38 JAP NNS
Synonyms: hidden
= Aspidium sabaei Franch. & Sav.; Enum. Pl. Jap. 2: 259, 632 (1879)
= Nephrodium sabaei Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)
species :
Dryopteris sacrosancta Koidz.; Bot. Mag. (Tokyo) 88: 108 (1924)
Distribution: China (Liaoning, Shandong, Zhejiang); Japan; North Korea; South Korea (incl. Cheju Isl.)
36 CHI 38 JAP KOR NKO
Synonyms: hidden
= Dryopteris bissetiana var. sacrosancta (Koidz.) H.Itô; Bot. Mag. (Tokyo) 50: 36 (1936)
= Dryopteris kobayashii Kitag.; Rep. Exped. Manchoukuo Sect. IV, Pt. 2, Contr. Cogn. Fl. Manshuricae, 56 (1935) (fide Fraser-Jenkins (1986))
= Dryopteris tieanzuensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 24 (1987)
= Dryopteris varia subsp. sacrosancta (Koidz.) Sugim.
= Dryopteris varia var. sacrosancta (Koidz.) Ohwi; Fl. Jap. Pterid. 88 (1957)
= Polystichum bissetianum var. sacrosanctum (Koidz.) Nakai
= Polystichum sacrosanctum (Koidz.) Koidz.; Bot. Mag. (Tokyo) 43: 388 (1929)
species :
Dryopteris saffordii C.Chr.; Amer. Fern J. 1: 94 (1911)
Distribution: Peru; Bolivia (La Paz, Tarija)
83 BOL PER
species :
Dryopteris sandwicensis (Hook. & Arn.) C.Chr.; Ind. 290 (1905)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Conservation Status: IUCN E
Synonyms: hidden
= Phegopteris sandwicensis (Hook. & Arn.) Mann; Proc. Amer. Acad. 7: 218 (1868)
= Polypodium sandwicense Hook. & Arn.; Bot. Beech. Voy. 105 (1832)
= Thelypteris sandwicensis (Hook. & Arn.) Fosberg; Occ. Pap. Bishop Mus. 23: 30 (1962)
species :
Dryopteris saxifraga H.Itô; Bot. Mag. (Tokyo) 60: 125 (1936)
Distribution: China (Jilin, Liaoning); Japan; South Korea; North Korea (incl. Cheju Isl.)
36 CHI 38 JAP KOR NKO
Synonyms: hidden
= Dryopteris varia subsp. saxifraga (H.Itô) Sugim.
= Dryopteris varia var. saxifraga (H.Itô) H.Ohba; Sci. Rep. Tohoku Univ., Ser. 4, Biol. 36: 111 (1971)
species :
Dryopteris saxifragivaria Nakai; J. Jap. Bot. 18: 286 (1942), pro hybr.
Distribution: South Korea (incl. Cheju Isl.)
38 KOR
Remarks: stabilized hybrid derived from Dryopteris saxifraga x bissetiana
Synonyms: hidden
= Dryopteris saxifraga var. deltoidea H.Itô; Bot. Mag. (Tokyo) 50: 126 (1936)
= Dryopteris saxifragivaria f. cristata Nakai
= Dryopteris varia f. cristata (Nakai) S.Y.Oh
species :
Dryopteris scabrosa (Kunze) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: S-India (Kerala, Tamil Nadu)
40 IND
Remarks: Subgen. Nothoperanema, sect. Shiehia
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium scabrosum Kunze; Linn. 24: 286 (1856)
= Ctenitis scabrosa (Kunze) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 292 (1938)
= Dryopsis scabrosa (Kunze) Holttum & P.J.Edwards; Kew Bull. 41(1): 199 (1986)
= Gymnothalamium aspidioides Zenker; ms.
= Lastrea scabrosa (Kunze) T.Moore; Ind. 103 (1858)
= Nephrodium scabrosum (Kunze) Baker; Syn. 283 (1867)
= Polypodium marginale Wall.; Num. List no. 322 (1828) (p. p.)
= Polypodium nigrocarpum Bedd.; Ferns South. India, t. 169 (1863) non Blume 1828
species :
Dryopteris schimperiana (Hochst. ex A.Braun) C.Chr.; Ind. 291 (1905)
Distribution: D.R.Congo [Zaire]; Sudan; Eritrea; Ethiopia (Ahmar Mts.); Uganda; Rwanda; Burundi; Kenya; Tanzania; Malawi; N-Yemen
23 BUR RWA ZAI 24 ERI ETH SUD 25 KEN TAN UGA 26 MLW 35 YEM
Synonyms: hidden
= Aspidium schimperianum Hochst. ex A.Braun; Flora 708 (1841)
= Polystichum schimperianum (Hochst. ex A.Braun) Keyserl.; Pol. Cyath. Hb. Bung. 44 (1873)
species :
Dryopteris scottii (Bedd.) Ching; Bull. Dept. Biol. Coll. Sci. Sun Yatsen Univ. 6: 3 (1933)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan; Bhutan; Myanmar (Kachin, Chin, etc.); N-Thailand; Vietnam; Japan; Borneo (Mt. Kinabalu, Sabah); India (Arunachal Pradesh, Assam State, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal, Jammu & Kashmir); Sumatra; Nepal
36 CHI 38 JAP TAI 40 BHU IND NEP 41 MYA THA VIE 42 BOR SUM
Synonyms: hidden
= Dryopteris grossa (Christ) C.Chr.; Ind. 269 (1906)
= Dryopteris hirtipes var. exinvolucrata (C.B.Clarke) Sledge ex Panigrahi & Basu; Proc. Indian Sci. Congr. Assoc. 69(3/6): 71 (1982) (fide Fraser-Jenkins 1997)
= Dryopteris nigrisquama f. subdecipiens (Hayata) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 16. 1938 (1939)
= Dryopteris nigrisquama Hayata; Ic. Pl. Formos. 4: 167, f. 106 (1914)
= Dryopteris subdecipiens Hayata; Ic. Pl. Formos. 4: 181, f. 119 (1914)
= Nephrodium hirtipes var. exinvolucrata C.B.Clarke; J. Linn. Soc., Bot. 25: 93 (1889)
= Phegopteris grossa Christ; Bull. Herb. Boiss. 7: 13 (1899)
= Phegopteris melanolepis Alderw.; Bull. Jard. Bot. Buitenz. II, 16: 25 (1914)
= Phegopteris scottii (Bedd.) Bedd.; Handb. Ferns Brit. India, Suppl. 19 (1876)
= Polypodium scottii Bedd.; Ferns brit. India, t. 345 (1870)
species :
Dryopteris sericea C.Chr.; Bot. Gaz. 64. 336 (1913)
Distribution: China (Gansu, Hubei, Shaanxi, Shanxi)
36 CHI
species :
Dryopteris serratodentata (Bedd.) Hayata; Ic. Pl. Formos. 4: 179, f. 116 (1914)
Distribution: China (W-Sichuan, NW-Yunnan); Tibet; Taiwan; India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal); Bhutan; Myanmar; Pakistan (Hazara); Nepal
36 CHI TIB 38 TAI 40 BHU IND NEP PAK 41 MYA
Synonyms: hidden
= Dryopteris filix-mas subsp. serratodentatum (Bedd.) C.Chr.; Ind. Fil. 5: 265 (1905)
= Dryopteris filix-mas var. serratodentata (Bedd.) Hayata; J. Coll. Sci. Imp. Univ. Tokyo 30: 416 (1911)
= Lastrea filix-mas var. odontoloma Baker; Hook. & Baker, Syn. Fil. 498 (1874) (excl. fig. of Bedd. cited and the var. nidus)
= Lastrea filix-mas var. serratodentata Bedd.; Handb. Suppl. 55 (1892)
= Lastrea odontoloma (Baker) Bedd.; Handb. Ferns Brit. India, t. 115 (1865)
= Nephrodium filix-mas var. odontoloma Baker; Syn. Fil. (Hook. & Baker) ed. 2: 498 (1874)
= Nephrodium odontoloma (Baker) C.B.Clarke; Tr. Linn. Soc. II, 1: 521 (1880)
= Nephrodium serratodentatum (Bedd.) Hope; J. Bombay Nat. Hist. Soc. 12: 622, t. 10 (1899)
= Woodsia veitchii Christ; Bull. Ac. Géogr. Bot. Mans 121 (1906)
species :
Dryopteris setosa (Thunb. ex Murray) Akasawa; Bull. Kochi Wom. Univ. 7: 27 (1959)
Distribution: China (Anhui, Fujian, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shandong, Shaanxi, Shanxi, Sichuan, Yunnan, Zhejiang); Japan; South Korea (incl. Cheju Isl.); North Korea
36 CHI 38 JAP KOR NKO
Remarks: valid name disputed, possibly D. bissetiana (Baker) C. Chr.
Synonyms: hidden
= Aspidium setosum (Thunb. ex Murray) Sw.; J. Bot. (Schrader) 1800 [2]. 39 (1800 publ. 1801)
= Dryopteris bissetiana (Baker) C.Chr.; Ind. 254 (1905)
= Dryopteris paravaria Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 22 (1987)
= Dryopteris pseudobissetiana Ching ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 49 (1990)
= Dryopteris shanghaiensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 24 (1987)
= Dryopteris sinobissetiana Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 8 (1984)
= Dryopteris thunbergii Koidz.; Bot. Mag. (Tokyo) 38: 106 (1924)
= Dryopteris varia subsp. setosa (Thunb. ex Murray) Sugim.
= Dryopteris varia var. setosa (Thunb. ex Murray) Ohwi; Fl. Jap. Pterid. 88 (1957)
= Nephrodium bissetianum Baker; J. Bot. 366 (1877)
= Polypodium setosum Thunb.; Linn. Syst. Veg. ed. 14: 938 (1784)
= Polystichum bissetianum (Baker) Nakai; Bot. Mag. (Tokyo) 45: 102 (1931)
= Polystichum thunbergii Koidz.; Bot. Mag. (Tokyo) 38: 106 (1924)
species :
Dryopteris shiakeana H.Shang & Y.H.Yan; Phytotaxa 218(2): 158 (2015)
Distribution: China (Guangdong)
36 CHI
species :
Dryopteris shibipedis Kurata; J. Geobot. 14: 84 (1966)
Distribution: Japan (Kyushu)
38 JAP
Conservation Status: EW in Japan (listed as EX)
species :
Dryopteris shikokiana (Makino) C.Chr.; Ind. 292 (1905)
Distribution: China (Guangxi, Guizhou, Hunan, Sichuan, Yunnan); Japan
36 CHI 38 JAP
Remarks: not in India. subgen. Nothoperanema
Synonyms: hidden
= Aspidium shikokianum Makino; Bot. Mag. (Tokyo) (1892)
= Ctenitis shikokiana (Makino) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 88 (1939)
= Dryopteris hirtosparsa Christ; Bull. Ac. Géogr. Bot. Mans (1909), Mém. 20: 176 (1909)
= Nephrodium shikokianum Makino; Bot. Mag. (Tokyo) 18: 62 (1899)
= Nothoperanema shikokianum (Makino) Ching; Acta Phytotax. Sin. 11: 28 (1966)
= Nothoperanema shikokianum (Makino) Nakaike; Enum. Pterid. Jap., Fil. 240 (1975), isonym
species :
Dryopteris shiroumensis Kurata & Nakam.; J. Geobot. 9: 54 (1960)
Distribution: Japan (Honshu)
38 JAP
Conservation Status: IUCN R
species :
Dryopteris shorapanensis Askerov; Izv. Akad. Nauk Az. SSR Biol. Nauk. 1978(4): 4 (1978 publ. 1979), ex Fraser-Jenkins, Fern Gaz. 18(1): 7 (2007), pro hybr.
Distribution: Northern Caucasus (Krasnodar, Daghestan); Georgia [Caucasus] (incl. Abkhazia); Azerbaijan; NW-Iran; Turkey (Anatolia)
33 AZE GEO NCS 34 IRN TUR
Remarks: affinis agg.
Synonyms: hidden
= Dryopteris affinis subsp. persica Fraser-Jenk.; Willdenowia 10(1): 113 (1980)
species :
Dryopteris sichotensis Kom.; Bull. Jard. Bot. Petr. 16: 146 (1916)
Distribution: China (Heilongjiang, Jilin); North Korea; Japan; Russian Far East (Sakhalin, Amur, Khabarovsk, Primorye)
31 RFE 36 CHI 38 JAP NKO
Synonyms: hidden
= Dryopteris barbellata Fomin; Fl. Sib. & Or. extr. 6: 59, fig. (1930)
= Dryopteris crassirhizoma var. setosa (Christ) Miyabe & Kudô; Fl. Hokkaido 15 (1930)
= Dryopteris filix-mas var. setosa Christ; Bull. Ac. Géogr. Bot. Mans 1909: 163 (1909)
= Dryopteris coreanomontana Nakai; Bot. Mag. (Tokyo) 36. 132 (1921)
= Dryopteris sichotensis subsp. coreano-montana (Nakai) Vorosch.; A. K. Skvortsov (ed.), Florist. issl. v razn. raionakh SSSR: 144 (1985)
species :
Dryopteris sieboldii (T.Moore) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Zhejiang); Japan
36 CHI 38 JAP
Synonyms: hidden
= Aspidium podophyllum Lowe; Ferns 6, t. 34-35 (1857) (non Hook. (1853))
= Aspidium sieboldii (T.Moore) Van Houtte ex Mett.; Fil. Hort. Bot. Lips. 87, t. 20, f. 1-4 (1856)
= Lastrea podophylla J.Sm.; Cat. cult. ferns 57 (1857)
= Lastrea sieboldii (T.Moore) T.Moore; Index Fil. (T. Moore) 88 (1857)
= Nephrodium sieboldii (T.Moore) Hook.; Fil. Exot. t. 31 (1859)
= Polystichum sieboldii (T.Moore) Keyserl.; Pol. Cyath. Hb. Bung. 41 (1873)
= Pycnopteris sieboldii T.Moore; Gard. Chr. 468 (1855)
species :
Dryopteris sikkimensis (Bedd.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: China (Yunnan); Tibet; India (Sikkim, Arunachal Pradesh); Bhutan; Myanmar (Kachin, etc.)
36 CHI TIB 40 BHU IND 41 MYA
Conservation Status: IUCN VU for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium sikkimense Bedd. ex Baker; Hook. & Baker, Syn. Fil. 256 (1867)
= Dryopteris pseudosikkimensis Ching & S.K.Wu; Fl. Xizangica 1: 267 (1983)
= Lastrea sikkimensis (Bedd.) C.B.Clarke; J. Linn. Soc., Bot. 17: 403 (1879)
= Lastrea sikkimensis (Bedd.) Bedd.; Handb. Ferns Brit. India 259 (1883), isonym
= Nephrodium sikkimense (Bedd.) C.B.Clarke; Tr. Linn. Soc. II, 1: 525 (1880)
= Polystichum sikkimensis Bedd.; Handb. Ferns Brit. India, t. 1270 (1866)
= Thelypteris sikkimensis (Bedd.) C.F.Reed; Phytologia 17: 313 (1968)
species :
Dryopteris simasakii (H.Itô) Kurata; J. Geobot. 18: 5 (1970)
Distribution: China (Guangxi, Guizhou, Sichuan, Yunnan, Zhejiang); Japan
36 CHI 38 JAP
Synonyms: hidden
= Dryopteris erythrochlamys Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 5 (1984)
= Dryopteris excelsior Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 8 (1987)
= Dryopteris indusiata var. simasakii H.Itô; J. Jap. Bot. 9: 57, f. 6 (1933)
= Dryopteris labordei var. simasakii (H.Itô) H.Itô
= Dryopteris laodianensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 16 (1987)
= Dryopteris occidentalizhejiangensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 17 (1987)
variant :
var. paleacea (H.Itô) Sa.Kurata; J. Geobot. 18: 5 (1970)
Distribution: China (Guangxi, Guizhou, Sichuan, Yunnan); Japan
36 CHI 38 JAP
Synonyms: hidden
= Dryopteris indusiata var. paleacea H.Itô; J. Jap. Bot. 9: 57 (1933)
= Dryopteris labordei f. paleacea (H.Itô) H.Itô
= Dryopteris paomowanensis Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 10 (1984)
= Dryopteris rufosquamosa Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 6 (1987)
species :
Dryopteris sinonepalensis Z.Y.Zhuo & Fraser-Jenk.; Pl. Diversity 44(2): 185 (2021)
Distribution: Nepal; Tibet; China (Chongqing, Sichuan, Yunnan, ?Guangxi, ?Guizhou)
36 CHI TIB 40 NEP
Synonyms: hidden
= Aspidium nitidulum Wall. ex Kuhn; [Num. List no. 392 (1828)] ex Kuhn, Linnaea 36: 117 (1869)
= Dryopteris nitidula (Wall. ex Kuhn) Mitsuta & S.K.Wu; Vasc. Pl. Hengduan Mount. 1: 131 (1993), nom. illeg., non (C. Presl) Kuntze 1891
= Dryopteris sparsa subsp. nitidula (Wall. ex Kuhn) C.Chr.; Ind. Fil. 293 (1905)
= Dryopteris sparsa var. nitidula (Wall. ex Kuhn) Ching; Bull. Fan Mem. Inst. Biol., Bot. 8: 472 (1938)
= Lastrea sparsa var. nitidula Wall. ex Kuhn; Suppl. Ferns South. India 17, t. 374 (1876)
= Nephrodium sparsum var. nitidulum (Wall. ex Kuhn) C.B.Clarke; Tr. Linn. Soc. , Bot. 1: 524 (1880)
species :
Dryopteris sledgei Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 361 (1989)
Distribution: +Sri Lanka; S-India (Tamil Nadu)
40 IND SRL
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
species :
Dryopteris sordidipes Tagawa; Acta Phytotax. Geobot. 3: 29 (1934)
Distribution: S-Japan (incl. Yakushima Isl.); Ryukyu Isl.; Taiwan; Vietnam
38 JAP NNS TAI 41 VIE
Synonyms: hidden
= Dryopteris yabei Hayata; J. Coll. Sci. Imp. Univ. Tokyo 80: 424 (1911)
species :
Dryopteris sororia (Maxon) M.A.McHenry, Sundue & Barrington; Taxon 62(6): 1158 (2013)
Distribution: Guatemala; Mexico (Chiapas)
79 MEX 80 GUA
Synonyms: hidden
= Adenoderris sororia Maxon; Bot. Gaz. 39: 368 (1905)
species :
Dryopteris sparsa (D.Don) Kuntze; Rev. Gen. Pl. 2: 613 (1891)
subspecies :
subsp. sparsa
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Jiangxi, Shanxi, Sichuan, Yunnan, Zhejiang); Tibet; Taiwan; Japan; Ryukyu Isl.; India (Arunachal Pradesh, Himachal Pradesh, Meghalaya, Sikkim, Uttarakhand, West Bengal); Bhutan; Sri Lanka; Myanmar (Kachin, etc.)
36 CHI TIB 38 JAP NNS TAI 40 BHU IND SRL 41 MYA
Synonyms: hidden
= Aspidium densum Wall.; Num. List no. 390 (1828) ex Mett., Abh. Senckenb. Naturf. Ges. 2(2): 349 (1858)
= Aspidium oppositum Wall.; Cat. n. 7080 (1832), nom. nud., non (Vahl) Sw. (1829)
= Aspidium sparsum (D.Don) Spreng.; Syst. 4: 106 (1827)
= Dryopteris parasparsa Ching & S.K.Wu; Fl. Xizangica 1: 267 (1983)
= Dryopteris sinosparsa Ching & K.H.Shing; Fl. Fujianica 1: 601 (1982)
= Dryopteris sparsa var. ryukyuensis Seriz.; J. Jap. Bot. 46: 279 (1971)
= Lastrea densa (Wall.) C.Presl; Tent. Pterid. 76 (1836)
= Lastrea sparsa (D.Don) T.Moore; Ind. 87, 104 (1858)
= Lastrea sparsa (D.Don) Bedd.; Ferns South. India, t. 103 (1863) (excl. parte)
= Nephrodium sparsum D.Don; Prodr. Fl. Nepal. 6 (1825)
= Polypodium oppositum Wall.; Num. List no. 7080 (1829)
= Polystichum sparsum (D.Don) Keyserl.; Pol. Cyath. Hb. Bung. 43 (1873)
subspecies :
subsp. lehalii Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 322 (2008) [nothossp.]
Distribution: India (Darjeeling)
40 IND
Remarks: Dryopteris sparsa ssp. sparsa x sparsa ssp. viridescens
subspecies :
subsp. rectipinnula Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 320 (2008)
Distribution: China (Yunnan, Sichuan, Guizhou); Tibet; S-Japan; India (Arunachal Pradesh, Assam State, Chhattisgarh, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Tamil Nadu, Sikkim, Tripura, West Bengal); Nepal; Bhutan; Myanmar; Thailand; ?Cambodia; ?Laos; Vietnam; New Guinea (Irian Jaya, Papua New Guinea); Philippines; Palawan; peninsular Malaysia (Maxwell Hill, Perak, Fraser´s Hill & Cameron Highlands, Pahang); Australia (NE-Queensland); Borneo (Mt. Kinabalu, Sabah); Java; Sumatra; Lesser Sunda Isl. (Bali, Flores); Sulawesi; ?Solomon Isl.; Fed. States of Micronesia (Pohnpei, Kosrae)
36 CHI TIB 38 JAP 40 BHU IND NEP 41 MYA THA VIE 42 BOR JAW LSI MLY NWG PHI PLW SUL SUM 50 AUS 62 MIC
Conservation Status: IUCN R (Queensland)
Synonyms: hidden
= Aspidium catophorum Kunze; Bot. Zeit. (Berlin) 262 (1848)
= Aspidium viridescens (Baker) Miq.; Cat. Mus. Bot. Lugd.-Bat. 127 (1870)
= Nephrodium sparsum var. latisquamum C.B.Clarke; Tr. Linn. Soc. II, 1: 524 (1880)
= Dryopteris rheophila Mitsuta ex Darnaedi; M. Kato & K. Iwats., J. Jap. Bot. 64(10): 302 (1989) (fide Fraser-Jenkins et al. 2018)
= Aspidium weigleanum Kunze; Linn. 24: 284 (1851)
= Dryopteris sparsa subsp. viridescens (Baker) Fraser-Jenk.; Taxon. Revis. Indian Subcontinental Pteridophytes 321 (2008)
= Dryopteris sparsa var. viridescens (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot.8: 473 (1938)
= Dryopteris viridescens (Baker) Kuntze; Rev. Gen. Pl. 2: 814 (1891)
= Nephrodium viridescens Baker; Syn. 276 (1847)
species :
Dryopteris speciosissima Copel.; Univ. Calif. Publ. Bot. 18: 219. (1942)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Remarks: Subgen. Nothoperanema, sect. Stenolepia
Synonyms: hidden
= Stenolepia speciosissima (Copel.) Holttum ex P.J.Edwards; Guide Alpine Subalpine Fl. Mount Jaya [R. J. Johns & al.] 88 (2006)
= Ctenitis speciosissima (Copel.) Copel.; Gen. Fil. 125 (1947)
species :
Dryopteris sphaeropteroides (Baker) C.Chr.; Ind. 293 (1905)
Distribution: China (Sichuan, Yunnan)
36 CHI
Remarks: Subgen. Dryopsis
Synonyms: hidden
= Aspidium sphaeropteroides (Baker) Christ; Bull. Ac. Géogr. Bot. Mans 119 (1900)
= Athyrium sphaeropteroides (Baker) C.Chr.; Acta Hort. Gothob. 1: 77 (1924)
= Ctenitis sphaeropteroides (Baker) Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 295 (1938)
= Dryopsis sphaeropteroides (Baker) Holttum & P.J.Edwards; Kew Bull. 41(1): 199-200 (1986)
= Phegopteris sphaeropteroides (Baker) Christ; Bull. Herb. Boiss. 7: 14 (1899)
= Polypodium sphaeropteroides Baker; Bull. Misc. Inform. Kew 1895: 55 (1895)
species :
Dryopteris splendens (Hook. ex Bedd.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: China (Yunnan); Tibet; N-India (Arunachal Pradesh, Sikkim, West Bengal); peninsular Malaysia (Malacca); Bhutan; Nepal
36 CHI TIB 40 BHU IND NEP 42 MLY
Synonyms: hidden
= Aspidium splendens (Hook. ex Bedd.) Christ; Farnkr. Erde 259 (1897), non Willd. (1810)
= Lastrea splendens Hook. ex Bedd.; Handb. Ferns Brit. India, t. 42 (1865)
= Nephrodium splendens Hook.; Sp. Fil. 4: 126 (1862 (excl. parte)
species :
Dryopteris squamiseta (Hook.) Kuntze; Rev. Gen. Pl. 2: 813 (1891)
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; Equatorial Guinea (Rio Muni); Sudan; Ethiopia; Somalia; Uganda; Rwanda; Kenya; Tanzania; Zambia; Malawi; E-Zimbabwe; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Eswatini [Swaziland]; Madagascar; La Réunion; Rwanda; China (Yunnan); Tibet; Taiwan; Bhutan; N-India (Arunachal Pradesh, Sikkim, West Bengal); Nepal
23 BIO CMN EQG RWA 24 ETH SOM SUD 25 KEN TAN UGA 26 MLW ZAM ZIM 27 SAF SWZ 29 MDG REU 36 CHI TIB 38 TAI 40 BHU IND NEP
Remarks: subgen. Nothoperanema
Conservation Status: IUCN R (Fernando Poo)
Synonyms: hidden
= Aspidium squamisetum (Hook.) Kuhn; Fil. Deck. 24 (1867)
= Ctenitis trichorhachis (Hayata) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 90 (1939)
= Dryopteris atrosetosa Rosenst.; Hedwigia 66: 342 (1915)
= Dryopteris buchananii (Baker) Kuntze; Rev. Gen. Pl. 2: 812 (1891)
= Dryopteris trichorhachis Hayata; Ic. Pl. Formos. 4: 185, f. 123 (1914) (thrichorhachis)
= Lastrea buchananii (Baker) Bedd.; Handb. Ferns Brit. India 255 (1883)
= Nephrodium buchananii Baker; Syn. Fil. (Hook. & Baker) ed. 2: 498 (1874)
= Nephrodium squamisetum Hook.; Sp. 4: 140, t. 268 (1862)
= Nothoperanema squamisetum (Hook.) Ching; Acta Phytotax. Sin. 11: 27 (1966)
species :
Dryopteris stanley-walkeri Fraser-Jenk.; Pl. Divers. Himalaya 140 (2001)
Distribution: India (Himalaya)
40 IND
species :
Dryopteris stenolepis (Baker) C.Chr.; Ind. 294 (1905)
Distribution: China (Gansu, Guangxi, Sichuan, Yunnan); Taiwan; Tibet; Vietnam; ?Laos; Myanmar (Kachin, etc.); India (Arunachal Pradesh, Uttarakhand); Nepal; Bhutan
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA VIE
Synonyms: hidden
= Aspidium stenolepis Christ; Bull. Herb. Boiss. 7: 22 (1899)
= Dryopteris atrata var. stenolepis (Baker) Tagawa; Acta Phytotax. Geobot. 10(4): 281 (1941)
= Dryopteris hirtipes var. stenolepis (Baker) C.Chr.; Contr. U. S. Natl. Herb. 26(6): 279 (1931)
= Lastrea hirtipes (Hook.) Bedd.; Handb. Ferns Brit. India 232 (1883) (excl syn.)
= Nephrodium hirtipes Hook.; Sp. Fil. 4: 115, t. 2,49 (1862) (excl syn. Aspidium hirtipes Blume)
= Polypodium stenolepis Baker; Bull. Misc. Inform. Kew 1898: 231 (1898)
= Thelypteris stenolepis (Baker) C.F.Reed; Phytologia 17: 315 (1968)
species :
Dryopteris stewartii Fraser-Jenk.; Kalikasan 7(3): 272 (1978 publ. 1979)
Distribution: E-Afghanistan; Pakistan (Quetta, Kurram, Chitral, Swat, Pakistani Punjab, Hazara, Gilgit); India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand); W-Nepal
34 AFG 40 IND NEP PAK
Synonyms: hidden
= Dryopteris odontoloma f. brevifolia Mehra & Khullar; (1974 publ. 1980) (nom. nud., fide Fraser-Jenkins (1997))
species :
Dryopteris subarborea (Baker) C.Chr.; Ind. 295 (1905)
Distribution: peninsular Malaysia (Cameron Highlands, Pahang); Sumatra; Borneo (Mt. Kinabalu, Sabah); Philippines; Sulawesi; New Guinea (Irian Jaya, Papua New Guinea); Fiji (Viti Levu)
42 BOR MLY NWG PHI SUL SUM 60 FIJ
Synonyms: hidden
= Alsophila warburgii Christ; Ann. Jard. Bot. Buitenz. 15: 80, t. 13, f. 3 (1897)
= Aspidium filix-mas var. megaphylla (Baker) C.Chr.; Bull. Herb. Boiss. 6: 194 (1898) (non Mett. (1864))
= Dryopteris tabacicoma Alderw.; Bull. Jard. Bot. Buitenz. II, 16: 11 (1914)
= Lastrea megaphylla (Baker) Bedd.; Handb. Suppl. 63 (1892)
= Nephrodium megaphyllum Baker; J. Linn. Soc. 22: 227 (1886)
= Polypodium subarboreum Baker; J. Linn. Soc. 24: 259 (1887)
species :
Dryopteris subatrata Tagawa; Acta Phytotax. Geobot. 9: 207 (1940)
Distribution: Taiwan
38 TAI
species :
Dryopteris subbipinnata W.H.Wagner & Hobdy; Contr. Univ. Michigan Herb. 19: 69 (1993)
Distribution: Hawaii (Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Dryopteris palikuensis Fraser-Jenk.; Thaiszia 4(1): 24 (1994)
species :
Dryopteris subcochleata Fraser-Jenk. & K.C.; Ferns and Fern-allies of Nepal 1: 27 (2015)
Distribution: India (Meghalaya, Manipur, Mizoram, Nagaland); W-Myanmar
40 IND 41 MYA
species :
Dryopteris subexaltata (Christ) C.Chr.; Ind. 295 (1905)
Distribution: Taiwan; Japan; Ryukyu Isl.; South Korea (Cheju Isl.)
38 JAP KOR NNS TAI
Synonyms: hidden
= Aspidium subexaltatum Christ; Bull. Herb. Boiss. II, 4: 616 (1904)
= Dryopteris hayatae Tagawa; Acta Phytotax. Geobot. 1: 156 (1932) (hayatai)
species :
Dryopteris subhikonensis K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(2): 99 (2018)
Distribution: Japan (Honshu)
38 JAP
species :
Dryopteris subimpressa Loyal; Nova Hedwigia 16: 467, t. 177, 178, f. 9-16 (1969)
Distribution: China (W-Yunnan); India (Himachal Pradesh, Arunachal Pradesh, Sikkim, Uttarakhand); Nepal; Myanmar; Bhutan
36 CHI 40 BHU IND NEP 41 MYA
Synonyms: hidden
= Dryopteris lancipinnula Ching; Sporae Pterid. Sin. 326 (1976), nom. nud.
= Dryopteris submarginata Loyal; Nova Hedwigia 16: 465, t. 175, 176, f. 1-8 (1969)
= Dryopteris subodontoloma Ching ex Loyal; (1961) (nom. nud., fide Fraser-Jenkins (1997))
species :
Dryopteris sublacera Christ; Lecomte, Not. syst. 1: 43 (1909)
Distribution: China (Hubei, Shaanxi, Sichuan, Yunnan); Tibet; C-Taiwan; India (Himachal Pradesh, Uttarakhand, ?Arunachal Pradesh); Nepal; Bhutan
36 CHI TIB 38 TAI 40 BHU IND NEP
Synonyms: hidden
= Dryopteris blepharolepis C.Chr.; Lév., Cat. Pl. Yun-Nan 103 (1916)
= Dryopteris juxtaposita f. multica Lév.; Cat. Pl. Yun-Nan 104 (1916)
= Dryopteris minjiangensis H.S.Kung; Acta Bot. Yunnan. 4(4): 341 (1982)
= Dryopteris schneideriana Hand.-Mazz.; Akad. Anz. Akad. Wiss. Wien no. 7.1 (1922)
= Nephrodium sublacerum (Christ) Hand.-Mazz.; Symb. Sin. 6: 21-25 (1929)
species :
Dryopteris submarginata Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 132 (1914)
Distribution: China (Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Zhejiang); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Dryopteris nudistipes Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 30 (1987)
= Dryopteris sessilipinna Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 29 (1987), nom. illeg., non Copel.
species :
Dryopteris subpycnopteroides Ching ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 199 (1986)
Distribution: China (NW-Yunnan)
36 CHI
species :
Dryopteris subreflexipinna Ogata; J. Jap. Bot. 11: 30, f. 2 (1935)
Distribution: N-Thailand; Taiwan
38 TAI 41 THA
Remarks: status uncertain, possible syn. of A. diffracta
Conservation Status: IUCN R
Synonyms: hidden
= Arachniodes subreflexipinna (Ogata) Ching; Acta Bot. Sin. 10: 260 (1962)
= Acrorumohra subreflexipinna (Ogata) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4 102 (1939)
= Rumohra subreflexipinna (Ogata) H.Itô; J. Jap. Bot. 11: 583 (1935)
species :
Dryopteris subtriangularis (Hope) C.Chr.; Ind. Fil. 296 (1905)
Distribution: China (Guangxi, Guizhou, Hainan, Sichuan, Yunnan); Ryukyu Isl.; Taiwan; Tibet; Myanmar; India (Assam, Arunachal Pradesh, Meghalaya); Philippines; Thailand; Vietnam
36 CHI TIB 38 NNS TAI 40 IND 41 MYA THA VIE 42 PHI
Remarks: often confused with D. tenuicula
Synonyms: hidden
= Dryopteris acutidens (C.Chr.) Ching; Ching & Chu H. Wang, Acta Phytotax. Sin. 8: 168, t. 2 (24) (1959), not C. Chr. (1905)
= Dryopteris caudifrons Ching; Ching & Chu H. Wang, Acta Phytotax. Sin. 8: 168, t. 22(25) (1959)
= Dryopteris subassamensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 451 (1938)
= Dryopteris uropinna M.G.Price; Gard. Bull. Singapore 30: 244, f. 1A-C (1977)
= Lastrea filix-mas var. subtriangularis (Hope) Bedd.; Handb. Ferns Brit. India, Suppl. 56 (1892)
= Nephrodium subtriangulare Hope; J. Bot. 1890: 327 (1890)
species :
Dryopteris subtsushimensis K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(3): 154 (2018)
Distribution: Japan (Kyushu); China (Jiangsu)
36 CHI 38 JAP
species :
Dryopteris sukungiana Z.Y.Zuo; Phytotaxa 533(5): 260 (2022)
Distribution: China (Yunnan)
36 CHI
species :
Dryopteris sweetiorum Lorence & W.L.Wagner; PhytoKeys 4: 19 (2011)
Distribution: Marquesas Isl. (Fatu Hiva)
61 MRQ
species :
Dryopteris tahmingensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 480 (1938)
Distribution: China (Fujian, Guangdong, Guangxi, Yunnan)
36 CHI
Synonyms: hidden
= Dryopteris hwangii Ching; Fl. Fujianica 1: 600 (1982)
species :
Dryopteris takeuchiana Koidz.; Acta Phytotax. 1: 30 (1932)
Distribution: Japan
38 JAP
species :
Dryopteris tenompokensis C.Chr.; Gard. Bull. Str. Settl. 7: 248 (1934)
Distribution: Borneo (Sabah)
42 BOR
species :
Dryopteris tenuicula Matthew & Christ; Not. syst. 1: 56 (1909)
Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Sichuan, Zhejiang); Taiwan; Myanmar; Japan; Ryukyu Isl. (Okinawa)
36 CHI 38 JAP NNS TAI 41 MYA
Synonyms: hidden
= Dryopteris erythrosora var. purpurascens H.Itô
= Dryopteris indusiata var. purpurascens (H.Itô) Sa.Kurata; (1962)
= Dryopteris jiulungshanensis P.S.Chiu & G.H.Yao ex Ching; Bull. Bot. Res. Harbin 2(2): 62 (1982)
= Dryopteris labordei var. purpurascens (H.Itô) Seriz.; J. Jap. Bot. 52(1): 17 (1977)
= Dryopteris neglecta Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 4(4): 4 (1984)
= Dryopteris purpurella Tagawa; Acta Phytotax. 1: 307 (1932)
= Dryopteris rubristipes Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 3(4): 19 (1983) (fide Fraser-Jenkins 1997)
= Dryopteris subchampionii Ching; Bot. Res. Acad. Sin. 1987(2): 14 (1987)
= Dryopteris subtenuicula Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 29 (1987)
= Dryopteris yaoi Ching; Bull. Bot. Res. Harbin 2(2): 64 (1982)
= Dryopteris zhangii Ching; Bull. Bot. Res., Harbin 3(3): 26 (1983) (fide Fraser-Jenkins (1986)
= Nephrodium tenuiculum (Matthew & Christ) Tutcher; Fl. Kwantung and Hongkong 348 (1912)
species :
Dryopteris tenuipes (Rosenst.) Seriz.; J. Jap. Bot. 46: 19 (1971)
Distribution: Taiwan
38 TAI
Remarks: Subgen. Dryopsis
Synonyms: hidden
= Dryopteris erythrosora var. tenuipes Rosenst.; Hedwigia 56: 341 (1915)
species :
Dryopteris tetrapinnata W.H.Wagner & Hobdy; Contr. Univ. Michigan Herb. 22: 173 (1999)
Name: Wakamoi Woodfern
Distribution: Hawaii (Maui)
63 HAW
Synonyms: hidden
= Dryopteris unidentata var. ukelelensis Fraser-Jenk.; Thaiszia 4(1): 33 (1994)
species :
Dryopteris tetsu-yamanakae Kurata; J. Geobot. 10: 100 (1962)
Distribution: Japan
38 JAP
species :
Dryopteris tingiensis Ching & S.K.Wu ex Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 202 (1986)
Distribution: Tibet; Nepal
36 TIB 40 NEP
species :
Dryopteris tokyoensis (Matsum.) C.Chr.; Ind. 298 (1905)
Distribution: China (Fujian, Hubei, Hunan, Jiangxi, Zhejiang); Japan; South Korea
36 CHI 38 JAP KOR
Synonyms: hidden
= Aspidium transitorium Christ; Bull. Herb. Boiss. 7: 822 (1899)
= Dryopteris sanmingensis Ching; Fl. Fujianica 1: 600 (1982)
= Nephrodium tokyoense Matsum. ex Makino; Bot. Mag. (Tokyo) 12: 87 (1898)
species :
Dryopteris toyamae Tagawa; Acta Phytotax. Geobot. 8: 167 (1939)
Distribution: Japan; Taiwan
38 JAP TAI
Synonyms: hidden
= Dryopteris enneaphylla var. toyamae (Tagawa) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 12: 305 (1975)
species :
Dryopteris transmorrisonense (Hayata) Hayata; Ic. Pl. Formos. 4: 187 (1914)
Distribution: Taiwan; S-China; Nepal; India (Sikkim)
36 CHI 38 TAI 40 IND NEP
Remarks: subgen. Dryopsis. Synonymized into D. nidus by Flora of China, but Fraser-Jenkins et al. (2018) and Knapp (Ferns of Taiwan) keep the species separate
Synonyms: hidden
= Polystichum transmorrisonense Hayata; J. Coll. Sci. Imp. Univ. Tokyo 80: 427 (1911)
= Dryopsis transmorrisonensis (Hayata) Holttum & P.J.Edwards; Kew Bull. 41(1): 191 (1986)
= Ctenitis transmorrisonensis (Hayata) Tagawa; Acta Phytotax. Geobot. 8: 95 (1939)
species :
Dryopteris tricellularis J.P.Roux; Kew Bull. 57(3): 735 (2002)
Distribution: Ethiopia; Uganda; Kenya; Tanzania
24 ETH 25 KEN TAN UGA
species :
Dryopteris tsoongii Ching; Bot. Res. Acad. Sin. 1987(2): 14 (1987)
Distribution: China (Anhui, Jiangxi, Zhejiang)
36 CHI
Synonyms: hidden
= Dryopteris wuyuanensis Ching; Ching & P. C. Chiu, Bot. Res. Acad. Sin. 2: 15-16, pl. 6, f. 2 (1987)
species :
Dryopteris tsugiwoi Kurata; J. Geobot. 17: 34 (1969)
Distribution: Japan
38 JAP
Conservation Status: IUCN R. CR in Japan
species :
Dryopteris tsushimensis K.Hori & N.Murak.; Acta Phytotax. Geobot. 69(3): 149 (2018)
Distribution: Japan (Kyushu)
38 JAP
species :
Dryopteris tsutsuiana Kurata; J. Geobot. 17: 35 (1969)
Distribution: Japan (Kyushu)
38 JAP
Conservation Status: CR in Japan. IUCN V
species :
Dryopteris tyrrhena Fraser-Jenk. & Reichst.; Fern Gaz. 11: 180 (1975), nom. nov.
Distribution: Spain (Sa. Nevada); France; Corsica; Sardinia; Italy
12 COR FRA SAR SPA 13 ITA
Remarks: tyrrhena agg.
Synonyms: hidden
= Aspidium nevadense Boiss.; Elench. pl. nov. 93 (1838) (non Dryopteris nevadensis (Baker) Underw.)
= Dryopteris ×cebennae Fraser-Jenk.; Candollea 32(2): 318 (1977) [fide Jeßen (in litt.)]
species :
Dryopteris unidentata (Hook. & Arn.) C.Chr.; Ind. 299 (1905)
Distribution: Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Dryopteris acutidens C.Chr.; Ind. 250 (1905)
= Dryopteris unidentata var. acutidens (C.Chr.) Fraser-Jenk.; Thaiszia 4(1): 32 (1994)
= Nephrodium unidentatum (J.Sm.) Diels; Nat. Pfl. 1(4): 174 (1899)
= Phegopteris spinulosa Hillebr.; Fl. Haw. 566 (1888)
= Phegopteris unidentata J.Sm.; Ferns Brit. For. 170 (1866)
= Polypodium spinulosum (Hillebr.) Baker; Ann. Bot. 5: 459 (1891)
= Polypodium unidentatum (J.Sm.) Hook. & Arn.; Bot. Beech. Voy. 105 (1832)
variant :
var. paleacea (Hillebr.) Fraser-Jenk.; Thaiszia 4(1): 31 (1994)
Distribution: Hawaii (Kauai, Oahu, Molokai, Maui, Hawaii Isl.)
63 HAW
Synonyms: hidden
= Phegopteris unidentata var. paleacea Hillebr.
species :
Dryopteris uniformis (Makino) Makino; Bot. Mag. (Tokyo) 23: 145 (1909)
Distribution: China (Anhui, Fujian, Jiangsu, Jiangxi, Zhejiang); Japan; South Korea (incl. Cheju Isl.)
36 CHI 38 JAP KOR
Synonyms: hidden
= Dryopteris decurrentiloba Ching & C.F.Zhang; Bull. Bot. Res. Harbin 3(3): 31 (1983)
= Dryopteris dentipalea Nakai; Bot. Mag. (Tokyo) 34: 36 (1920)
= Dryopteris jiangshanensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 34 (1987)
= Dryopteris pseudouniformis Ching; Fl. Fujianica 1: 601 (1982)
= Dryopteris uniformis f. coreana (Nakai & H.Itô) H.Itô; Nov. Fl. Jap. 4: 14 (1939)
= Dryopteris uniformis var. coreana Nakai ex H.Itô; Nov. Fl. Jap. 4: 14 (1939)
= Dryopteris uniformis var. rufomarginata K.H.Shing; Jiangxi Sci. 8(3): 47 (1990)
= Nephrodium lacerum var. uniforme Makino; Bot. Mag. (Tokyo) 17: 79 (1903)
species :
Dryopteris varia (L.) Kuntze; Rev. Gen. Pl. 2: 814 (1891)
Distribution: China (Anhui, Fujian, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang); Taiwan; South Korea (Cheju Isl.); Japan; Ryukyu Isl.; Bonin Isl. (Chichijima); Volcano Isl. (Kita-Iwojima, Minami-Iwojima); India; Philippines; Vietnam; Myanmar; NE-India (Assam, Meghalaya); Sulawesi
36 CHI 38 JAP KOR KZN NNS OGA TAI 40 IND 41 MYA VIE 42 PHI SUL
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Aspidium lacerum var. subtripinnatum Franch. & Sav.; Enum. Pl. Jap. 2: 238 (1879)
= Aspidium opacum (Hook.) Benth.; Fl. Hongk. 456. 456 (1861)
= Aspidium varium (L.) Sw.; Schrad., J. Bot. 1800(2): 35 (1801)
= Dryopteris caudifolia Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 21 (1987)
= Dryopteris consimilis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 25 (1987)
= Dryopteris fuyangensis Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 26 (1987)
= Dryopteris glabrescens Ching & P.S.Chiu ex K.H.Shing & J.F.Cheng; Jiangxi Sci. 8(3): 48 (1990)
= Dryopteris hololepis (Hayata) Tagawa; Acta Phytotax. Geobot. 4: 133 (1935)
= Dryopteris lingii Ching; Bot. Res. Acad. Sin. 1987(2): 23 (1987)
= Dryopteris matsuzoana Koidz.; Bot. Mag. (Tokyo) 39: 15 (1925)
= Dryopteris nanchuanensis Ching & Z.Y.Liu; Bull. Bot. Res. Harbin 3(4): 20 (1983)
= Dryopteris ogawai H.Itô; Ic. Pl. As. Or. 1: 18, t. 9 (1935)
= Dryopteris sinovaria Ching & Z.Y.Liu; Bull. Bot. Res., Harbin 6(1): 179, nom. nov.,
= Dryopteris yabei f. ogawae (H.Itô) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 59. 1938 (1939)
= Dryopteris yabei var. hololepis (Hayata) H.Itô; Bot. Mag. (Tokyo) 50(591): 128 (1936)
= Dryopteris yabei var. matsuzoana (Koidz.) H.Itô; Bot. Mag. (Tokyo) 50(591): 128 (1936)
= Dryopteris yabei var. ogawae (H.Itô) H.Itô; Bot. Mag. (Tokyo) 50: 128 (1936)
= Lastrea opaca Hook.; J. Bot. 9: 339 (1857)
= Lastrea varia (L.) T.Moore; Ind. Fil. 107 (1858)
= Nephrodium coriaceum Hope; J. Bot. 328 (1890)
= Nephrodium filix-mas f. subtripinnata (Franch. & Sav.) Matsum.; Ind. Pl. Jap. 1: 319 (1904)
= Nephrodium varium (L.) Desv.; Mem. Soc. Linn. Paris 6, 2: 269 (1827)
= Nephrodium varium (L.) C.Presl; Reliq. Haenk. 1: 36 (1826)
= Nephrodium villosum var. opaca (Hook.) Hieron.; Bot. Jahrb. Syst. 34(4): 446 (1904)
= Polypodium varium L.; Sp. Pl. 2: 1090 (1753)
= Polystichum hololepis Hayata; Ic. Pl. Formos. 5: 332 (1915)
= Polystichum varium (L.) C.Presl; Abh. Königl. Böhm. Ges. Wiss. 5(6) (Epim. Bot.): 57 (1851)
= Polystichum varium var. eurylepidota Rosenst.; Hedwigia 56(5): 340 (1915)
= Polystichum varium var. subtripinnatum (Franch. & Sav.) Nakai; Bot. Mag. (Tokyo) 39(461): 119 (1925)
species :
Dryopteris vescoi (Drake) C.Chr.; Ind. 300 (1905)
Distribution: Society Isl. (Tahiti)
61 SCI
Synonyms: hidden
= Nephrodium vescoi Drake; Fl. Polyn. franc. 301 (1893)
species :
Dryopteris vidyae Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 466 (1989), pro hybr.
Distribution: N-India (West Bengal, Sikkim, Arunachal Pradesh); Myanmar (Kachin, etc.)
40 IND 41 MYA
Conservation Status: IUCN EN for India (Fraser-Jenkins 2012)
species :
Dryopteris villarii (Bellardi) Woyn. ex Schinz & Thell.; Vierteljahrsschr. Naturf. Ges. Zürich 60: 339 (1915)
Name: Starrer Wurmfarn (DE)
Distribution: Albania; Austria; Baleares; England; Bulgaria; Corsica; France; Germany (Bayern); Greece; Switzerland; Liechtenstein; Italy; Bosnia & Hercegovina; Montenegro; Croatia; Macedonia; Serbia & Kosovo; Slovenia; Sardinia; Sicily; Northern Caucasus; Georgia [Caucasus]
10 GRB 11 AUT GER LIE SWI 12 BAL COR FRA SAR 13 ALB BIH BUL CRO GRC ITA MNT NMA SIC SLV SRB 33 GEO NCS
Synonyms: hidden
= Aspidium fragrans (Vill.) Gray; Nat. Arr. br. Fl. 2: 9 (1821)
= Aspidium rigidum (Hoffm.) Sw.; Schrad., J. Bot. 1800(2): 37 (1801)
= Aspidium rigidum f. fallax Milde
= Dryopteris rigida (Sw.) Underw.; Native Ferns ed. 4: 116 (1893) (non A. Gray, Man. 631 (1848))
= Dryopteris villarsii f. fallax (Milde) Niketic; Fl. Serbia 1: 155 (1992)
= Dryopteris villarsii var. australe (Ten.) Diklic & V.Nikolic; Fl. SR Srbije 10: 10 (1986)
= Filix rigida (Sw.) Farw.; 18. Ann. Repert. Michigan Acad. Sci. 81-83 (1916)
= Filix-Mas rigida (Sw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Hypodematium nivale Fée; Gen. 298 (1850-52)
= Lastrea rigida (Sw.) C.Presl; Tent. Pterid. 77 (1836)
= Lophodium fragrans Newman; xxi (1871)
= Lophodium rigidum (Sw.) Newman; Phyt. 4: app. 21 (1851)
= Nephrodium meridionale (Lojac.) Rouy; Fl. France 14: 409 (1913)
= Nephrodium rigidum (Hoffm.) Desv.; Mém. Soc. Linn. Paris 6: 261 (1827)
= Nephrodium villarsii (Bellardi) Beck; Glasn. Zemal isk. Muz. Bosni Hercegovini 28: 332 (1917)
= Nephrodium villarsii subsp. rigidum (Hoffm.) Hayek; Fedde, Repert. Beih. 30(1): 26 (1924)
= Polypodium asplenioides Scop.; Fl. Carn. ed. 2, 2: 298, t. 62 (1772)
= Polypodium fragrans Vill.; Hist. pl. Dauph. 3: 843 (1789)
= Polypodium odoratum Poir.; Encycl. 5: 541 (1804)
= Polypodium rigidum (Sw.) Hoffm.; Deutschl. Fl. 2: 6 (1795)
= Polypodium villarsii Bellardi; App. Fl. Pedem.: 49 (1792)
= Polystichum meridionale Lojac.; Fl. Sic. 3. 405 (1909)
= Polystichum rigidum (Sw.) DC.; Lam. & DC., Fl. France, ed. 3, 2: 560 (1805)
= Polystichum strigosum Roth; Röm., Arch. 2(1): 106 (1799), nom. illeg.
species :
Dryopteris wallichiana (Spreng.) Hyl.; Bot. Notis. 1963: 352 (1953)
subspecies :
subsp. wallichiana
Distribution: Mexico (Chiapas, Chihuahua, Colima, Ciudad de Mexico, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tamaulipas, Tlaxcala, Veracruz); Guatemala; El Salvador; Honduras; Costa Rica; Panama; Colombia (Antioquia, Boyacá, Caldas, Cauca, Cesar, Cundinamarca, Magdalena, Meta, Nariño, Norte de Santander, Quindío, Risaralda, Santander, Tolima); Venezuela (Lara, Merida, Tachira, Trujillo); Ecuador; Galapagos; Peru; Bolivia (Chuquisaca, Cochabamba, La Paz, Santa Cruz, Tarija); Brazil (Espírito Santo, Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo); NE-Argentina (Catamarca, Jujuy, Salta, Tucuman); Jamaica; Hispaniola; Tibet; China (Yunnan, Guizhou, Sichuan, Fujian, Jiangxi, Shanxi); Taiwan; Japan; Philippines; Vietnam; Nepal; Bhutan; Borneo (Mt. Kinabalu, Sabah); India (Arunachal Pradesh, Himachal Pradesh, Manipur, Meghalaya, Nagaland, Sikkim, ?Tamil Nadu, Uttarakhand, West Bengal); Sri Lanka; Myanmar (Kachin, Chin, etc.); Java; New Guinea (Irian Jaya, Papua New Guinea); Sulawesi; Turkey (Anatolia); Hawaii (Kauai, Oahu, Molokai, Lanai, Maui, Hawaii Isl.); Gough Isl.
34 TUR 36 CHI TIB 38 JAP TAI 40 BHU IND NEP SRL 41 MYA VIE 42 BOR JAW NWG PHI SUL 63 HAW 79 MEX 80 COS ELS GUA HON PAN 81 HIS JAM 82 CLM VEN 83 BOL ECU GAL PER 84 BRA 85 ARG 91 TDC
Synonyms: hidden
= Aspidium crinitum M.Martens & Galeotti; Mém. Acad. Roy. Sci. Bruxelles 16: 66, t. 17, f. 2 (1842)
= Aspidium donianum Spreng.; Syst. 4: Suppl. 320 (1827), nom. superfl. nov. for Aspidium paleaceum D. Don
= Aspidium filix-mas f. paleaceum (Lag. ex Sw.) Asch.; Syn. Mitteleur. Fl. 1: 28 (1896)
= Aspidium filix-mas var. paleaceum (T.Moore) Mett.; Abh. Senckenb. Naturf. Ges. 2(2): 339 (1858)
= Aspidium filix-mas var. parallelogrammum (Kunze) Hillebr.; Fl. Haw. 574 (1888)
= Aspidium paleaceum (T.Moore) D.Don; Prodr. Fl. Nepal. 4-5 (1825), non Sw. (1806), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806
= Aspidium paleaceum (T.Moore) Dalla Torre & Sarnth.; Fl. Tirol. 6(1): 46 (1906), non Sw. 1806
= Aspidium paleaceum D.Don; Prodr. Fl. Nepal 4 (1824) non Sw. 1806
= Aspidium paleaceum Lag. ex Sw.; Synopsis filicum: 52 (1806)
= Aspidium parallelogrammum Kunze; Linn. 13: 146 (1839)
= Aspidium patentissimum (Wall. ex Kunze) Franch.; Nouv. Arch. Mus. II, 10: 118 (1887) (non Wall. (1829, nom. nud.)) (fide Fraser-Jenkins 2008)
= Aspidium patentissimum Wall.; Num. List no. 340 (1828) ex Kunze, Linn. 13: 146 (1839), nom. superfl.
= Aspidium wallichianum Spreng.; Syst. Veg. 4: 104 (1827), non C. Presl ex Kunze (1851)
= Dichasium parallelogrammum (Kunze) Fée; Gen. 302, t. 23 B, f. 1 (1850-52)
= Dichasium patentissimum (Wall. ex Kunze) Fée; Gen. 302, t. 23 B, f. 2 (1850-52)
= Dryopteris cyrtolepis Hayata; Ic. Pl. Formos. 4: 149 t 89 (1914)
= Dryopteris cyrtolepis var. doiana (Tagawa) H.Itô; Nakai & Honda (1938 publ. 1939)
= Dryopteris doiana Tagawa; Acta Phytotax. Geobot. 5: 253 (1936)
= Dryopteris doniana (Spreng.) Ching; Sunyatsenia 6: 3 (1941)
= Dryopteris filix-mas subsp. paleacea (Lag. ex Sw.) W.Koch ex Braun-Blanq.; Veröff. Geobot. Inst. Rübel Zürich 7: 34 (1932), isonym
= Dryopteris filix-mas subsp. paleacea (Sw.) C.Chr.; Rosenvinge, Biol. Arbejd. Til. Eug. Warming 76 (1911)
= Dryopteris filix-mas subsp. parallelogramma (Kunze) Christ; Ann. Jard. Cons. Bot. Genève 15-16: 187 (1912)
= Dryopteris filix-mas subsp. patentissima (Wall. ex Kunze) C.Chr.; Ind. Fil. 5: 265 (1905) (p. p., excl. Nephrodium filix-mas var. fibrillosa C. B. Clarke (1880)) (fide Fraser-Jenkins 2008)
= Dryopteris filix-mas var. crinita (M.Martens & Galeotti) Rosenst.
= Dryopteris filix-mas var. paleacea (Lag. ex Sw.) Druce; List Brit. Pl. 87 (1908)
= Dryopteris filix-mas var. parallelogramma (Kunze) Conz.; Fl. Taxon. Mexic. 1: 19. (1939), nom. superfl.
= Dryopteris filix-mas var. parallelogramma (Kunze) Christ; Phil. J. Sci. 2: 212 (1907)
= Dryopteris himalaica (Ching & S.K.Wu) S.G.Lu; Acta Bot. Yunnan. 13(1): 40 (1991)
= Dryopteris kweichowicola Ching ex P.S.Wang; J. Arnold Arbor. 71(1): 119 (1990)
= Dryopteris mediterranea Fomin; Komarov, Fl. URSS 1: 35 (1934), nom. nov. for Lastrea filix-mas var. paleacea T. Moore
= Dryopteris paleacea (Lag. ex Sw.) C.Chr.; Amer. Fern J. 1: 94 (1911), non Hand.-Mazz. (1908)
= Dryopteris paleacea (T.Moore) Hand.-Mazz.; Verh. zool. Bot. Ges. Wien 58 (100) (1908), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806
= Dryopteris parallelogramma (Kunze) Alston; Amer. Fern J. 47: 92 (1957)
= Dryopteris patentissima (Wall. ex Kunze) N.C.Nair; Indian Forester 94: 169 (1968), isonym
= Dryopteris patentissima (Wall. ex Kunze) Panigrahi & Basu; Indian J. Forest. 3(3): 266, f. 1, t. 2. 1. & 4 (1980), isonym
= Dryopteris patentissimum (Wall. ex Kunze) Fée; Mém. Foug. 5: 303, pl. 23B, f. 2 (1852)
= Dryopteris quatanensis Ching; Wuyi Sci. J. 1: 6 (1981)
= Dryopteris reichsteinii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 191 (1986) (nom. nov., syn. fide Fraser-Jenkins (1997))
= Dryopteris ursipes Hayata; Ic. Pl. Formos. 5: 291, f. 116 (1915)
= Dryopteris wallichiana (Spreng.) Alston & Bonner; Candollea 15(7): 216 (1956), nom. superfl.
= Dryopteris wallichiana var. himalaica Ching & S.K.Wu; Fl. Xizangica 1: 257 (1983)
= Dryopteris wallichiana var. kweichowicola (Ching ex P.S.Wang) S.K.Wu; Fl. Reipubl. Popularis Sin. 5(1): 149 (2000)
= Lastrea filix-mas var. paleacea T.Moore; Nat. Print. Brit. Ferns 8vo. ed. 1: 178, 193, t. 33(A) (1859), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806
= Lastrea filix-mas var. parallelogramma (Kunze) Bedd.; Handb. Ferns Brit. India 249 (1883)
= Lastrea filix-mas var. parallelogramma Bedd.; Handb. Ferns Brit. India 249 (1883 (pro parte quoad. var. clarkei Bedd.)) (fide Fraser-Jenkins 2008)
= Lastrea filix-mas var. patentissima (Wall. ex Kunze) Bedd.; Suppl. Ferns South. India 29 (1876)
= Lastrea paleacea (Sw.) T.Moore; Ind. 99 (1858)
= Lastrea parallelogramma (Kunze) Liebm.; Vid. Selsk. Skr. V, 1: 271 (seors. 119) (1849)
= Lastrea patentissima (Wall. ex Kunze) J.Sm.; J. Bot. (Hook.) 4: 193 (1842)
= Nephrodium filix-mas f. paleaceum (T.Moore) Fiori; Fl. Anal. Italia 1: 8 (1896)
= Nephrodium filix-mas var. paleaceum (T.Moore) Hook.; Fil. Exot. t. 98 (1859), nom. illeg., non Aspidium paleaceum Lag. ex Sw. 1806
= Nephrodium filix-mas var. parallelogrammum (Kunze) Hook.; Sp. Fil. 4: 116 (1862)
= Nephrodium filix-mas var. patentissimum (Wall. ex Kunze) C.B.Clarke; Tr. Linn. Soc. II, 1: 520 (1880 (pro parte quoad sp.))
= Nephrodium parallelogrammum (Kunze) C.Hope; J. Bombay Nat. Hist. Soc. 14: 728 (1903)
= Nephrodium parallelogrammum f. patentissimum (Wall. ex Kunze) C.Hope; J. Bombay Nat. Hist. Soc. 14: 728 (1903)
= Nephrodium patentissimum (Wall. ex Kunze) C.B.Clarke; J. Linn. Soc. 15: 156 (1876) (incl. descr. in part, excl. type)
= Polystichum filix-mas var. paleaceum (T.Moore) Fiori; Nuov. Fl. Anal. Italia 1: 23 (1923)
subspecies :
subsp. bhutanica Fraser-Jenk.; Ferns and Fern-allies of Nepal 1: 28 (2015)
Distribution: Bhutan; Nepal; India (Arunachal Pradesh, Meghalaya, Uttarakhand); China (Yunnan); Nepal
36 CHI 40 BHU IND NEP
subspecies :
subsp. himalaica Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 134 (1997)
Distribution: India (Arunachal Pradesh, Nagaland, Sikkim, Uttarakhand, West Bengal); Nepal
40 IND NEP
subspecies :
subsp. madrasensis (Fraser-Jenk.) Fraser-Jenk.; Thaiszia 4(1): 22 (1994)
Distribution: S-India (Tamil Nadu); Sri Lanka
40 IND SRL
Synonyms: hidden
= Dryopteris madrasensis Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 359 (1989)
subspecies :
subsp. nepalensis Fraser-Jenk.; New Sp. Syndr. Indian Pteridol. 134 (1997)
Distribution: Bhutan; India (Uttarakhand, Sikkim, Arunachal Pradesh, Mizoram, Nagaland); C-Nepal; Myanmar; China (Yunnan, Sichuan); Tibet
36 CHI TIB 40 BHU IND NEP 41 MYA
subspecies :
subsp. madagascariensis (C.Chr.) J.P.Roux; Strelitzia 23: 124 (2009)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Dryopteris paleacea var. madagascariensis C.Chr.; Dansk Bot. Ark. 7: 53, t. 11, f. 5-9 (1932)
subspecies :
subsp. pachyphylla Fraser-Jenk. & Ralf Knapp; Annot. Checkl. Ind. Pterid. 2: 276 (2018), nom. nov.
Distribution: Taiwan; China; Bhutan; Myanmar; India (Arunachal Pradesh, Meghalaya, Nagaland)
36 CHI 38 TAI 40 BHU IND 41 MYA
Synonyms: hidden
= Dryopteris neorosthornii Ching; Bull. Fan Mem. Inst. Biol. Bot. 11: 62 (1941)
= Dryopteris pachyphylla Hayata; Ic. Pl. Formos. 4: 168, f. 108 (1914), non (Kunze) Kuntze (1891)
subspecies :
subsp. parrisiae (Fraser-Jenk.) Fraser-Jenk.; Indian Fern J. 39(1): 119 (2022)
Distribution: New Guinea (Papua New Guinea); Moluccas (Seram)
42 MOL NWG
Synonyms: hidden
= Dryopteris parrisiae Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 199 (1986)
subspecies :
subsp. reichsteinii Fraser-Jenk.; Ann. Bot. Fenn. 33: 71 (1996), non D. reichsteinii Fraser- Jenk. (1986)
Distribution: Zimbabwe
26 ZIM
species :
Dryopteris wardii (Baker) Kuntze; Rev. Gen. Pl. 2: 814 (1891)
Distribution: Seychelles
29 SEY
Synonyms: hidden
= Araiostegia wardii (Baker) Tardieu; Adansonia 5: 493 (1965)
= Aspidium wardii (Baker) Kuhn; v. Decken, Reis. 88, Bot. 69 (1879)
= Nephrodium wardii Baker; Syn. Fil. (Hook. & Baker) ed. 2: 500 (1874)
= Polystichopsis wardii (Baker) Tardieu; Notul. Syst. (Paris) 15: 176 (1956)
species :
Dryopteris wattsii McKeown, Sundue & Barrington; PhytoKeys 14: 43-56 (2012), nom. nov.
Distribution: Australia (NE-Queensland)
50 AUS
Conservation Status: very rare, known only from 2 localities
Synonyms: hidden
= Revwattsia fragilis (Watts) D.L.Jones; Fl. Australia 48: 711 (1998)
= Polystichum fragile Watts; Tr. Linn. Soc. N. S. Wales 39: 775, t. 88, f. 9 (1915)
species :
Dryopteris whangshanensis Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 421 (1938)
Distribution: China (Anhui, Fujian, Hubei, Jiangxi, Zhejiang)
36 CHI
Synonyms: hidden
= Dryopteris crassirhizoma subsp. whangshangensis (Ching) Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 14(3): 191 (1986) (whangshangensis)
= Dryopteris huangangshanensis Ching; Fl. Fujianica 1: 602 (1982)
= Dryopteris submarginalis Ching, Boufford & K.H.Shing; J. Arnold Arbor. 64: 30. (1983), not (Langsdorf & Fisch.) C. Chr. (1905)
species :
Dryopteris wideniana Fraser-Jenk.; Ann. Bot. Fenn. 52(1-2): 63 (2015)
Distribution: Zimbabwe
26 ZIM
species :
Dryopteris woodsiisora Hayata; Ic. Pl. Formos. 6, 158 (1916)
Distribution: China (Guangdong, Guizhou, Jiangxi, Liaoning, Shandong, Sichuan, Yunnan); Taiwan; Tibet; North Korea; India (Himachal Pradesh, Arunachal Pradesh, ?Manipur, Meghalaya, Nagaland, Sikkim, Uttarakhand, West Bengal); Nepal; Bhutan; Myanmar (Kachin, etc.); N-Thailand; Vietnam
36 CHI TIB 38 NKO TAI 40 BHU IND NEP 41 MYA THA VIE
Synonyms: hidden
= Dryopteris neochrysocoma Ching; Lingnan Sci. J. 21: 31 (1945)
= Dryopteris taishanensis F.Z.Li; Acta Phytotax. Sin. 20(3): 344 (1982)
= Dryopteris tenuissima f. serrata (Tagawa) H.Itô; Nakai & Honda, Nov. Fl. Jap. 4: 71. 1938 (1939)
= Dryopteris tenuissima Tagawa; Acta Phytotax. 1: 308 (1932)
= Dryopteris tenuissima var. serrata Tagawa; J. Jap. Bot. 13(3): 185 (1937)
= Dryopteris woodsiisora var. tenuissima (Tagawa) Li & J.Z.Wang; Bull. Bot. Res. Harbin 9(4): 80 (1989)
= Dryopteris zinongii Z.R.Wang & Fraser-Jenk.; Acta Phytotax. Sin. 23(5). 345 (1985)
species :
Dryopteris wulingshanensis J.P.Shu, Y.H.Yan & R.J.Wang; PhytoKeys 185: 21, figs. 3-4 (2021)
Distribution: China (Hunan)
36 CHI
species :
Dryopteris wuyishanica Ching & P.S.Chiu; Bot. Res. Acad. Sin. 1987(2): 34 (1987)
Distribution: China (Fujian)
36 CHI
species :
Dryopteris wuzhaohongii Li Bing Zhang; Taxon 61(6): 1208 (2012), nom. nov.
Distribution: China (Yunnan); Taiwan; Tibet
36 CHI TIB 38 TAI
Remarks: subgen. Nothoperanema. Probably to be included in Peranema paleolatum fide Fraser-Jenkins et al. (2018)
Synonyms: hidden
= Acrophorus macrocarpus Ching & S.H.Wu; S. H. Wu, Acta Phytotax. Sin. 21(4): 381 (1983)
species :
Dryopteris xanthomelas (Christ) C.Chr.; Ind. Fil., Suppl. 41 (1913)
Distribution: China (Anhui, Gansu, Hebei, Henan, Shanxi, Sichuan); Tibet; Taiwan; Bhutan; Nepal; Pakistan (Swat, Baltistan, Gilgit); India (Jammu & Kashmir, Himachal Pradesh, Uttarakhand, Sikkim, Darjeeling, Arunachal Pradesh, West Bengal)
36 CHI TIB 38 TAI 40 BHU IND NEP PAK
Remarks: D. sinofibrillosa is kept separate by Flora of China but sunk into D. xanthomelas by Fraser-Jenkins (2008) and Knapp (2012)
Synonyms: hidden
= Aspidium xanthomelas Christ; Bull. Acad. Geogr. Bot. 117 (1906)
= Dryopteris canaliculata Ching; Fl. Xizangica 1: 251 (1983)
= Dryopteris chingii N.C.Nair; Indian Forester 94: 169 (1968) for D. fibrillosa (C. B. Clarke) Hand.-Mazz., nom. superfl. (fide Fraser-Jenkins 1997)
= Dryopteris fibrillosa (C.B.Clarke) Hand.-Mazz.; Akad. Anz. Akad. Wiss. Wien no. 72. 1922 (non Christ (1905))
= Dryopteris filix-mas subsp. fibrillosa (C.B.Clarke) C.Chr.; Acta Hort. Gothob. 1: 57 (1924)
= Dryopteris filix-mas var. fibrillosa (C.B.Clarke) C.Chr.; Ind. Fil. 5: 265 (1905) (fide Fraser-Jenkins 1997)
= Dryopteris fibrillosissima Ching; Fl. Xizangica 1: 254 (1983) (fide Fraser-Jenkins et al. 2018)
= Dryopteris pulcherrima Ching; Bull. Fan Mem. Inst. Biol. Bot. 8: 422 (1938) (fide Fraser-Jenkins et al. 2018)
= Dryopteris sinofibrillosa Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 180 (1940) for D. fibrillosa (C. B. Clarke) Hand.-Mazz.
= Lastrea filix-mas var. parallelogramma subvar.fibrillosa (C.B.Clarke) Bedd.; Handb. Ferns Brit. India 250 (1883)
= Nephrodium clarkei var. fibrillosum (C.B.Clarke) Hand.-Mazz.; Symb. Sin. 6: 24 (1929)
= Nephrodium filix-mas var. fibrillosum C.B.Clarke; Tr. Linn. Soc. II, Bot. 1 520, t. 70 (1880)
= Nephrodium parallelogrammum f. fibrillosum (C.B.Clarke) C.Hope; J. Bombay Nat. Hist. Soc. 14: 731 (1903)
species :
Dryopteris xunwuensis Ching & K.H.Shing; Jiangxi Sci. 8(3): 48 (1990)
Distribution: China (Guangxi, Jiangxi)
36 CHI
species :
Dryopteris yongdeensis W.M.Chu ex S.G.Lu; Acta Phytotax. Sin. 28(6): 473 (1990)
Distribution: China (Yunnan)
36 CHI
species :
Dryopteris yoroii Seriz.; J. Jap. Bot. 46: 20 (1971)
Distribution: Taiwan; Tibet; India (Sikkim); China (Sichuan, Yunnan)
36 CHI TIB 38 TAI 40 IND
Remarks: partially confused with D. sinonepalensis, true distribution therefore not completely clear. Some material from India, Bhutan & Myanmar might refer to this species
Synonyms: hidden
= Acrorumohra yoroii (Seriz.) W.C.Shieh; J. Sci. Engin. (Natl. Chung-Hsing Univ.) 12: 301 (1975)
species :
Dryopteris zayuensis Ching & S.K.Wu; Fl. Xizangica 1: 255 (1983)
Distribution: China (Qinghai, Sichuan, Yunnan); Tibet; Taiwan; India (Jammu & Kashmir, Uttarakhand, Arunachal Pradesh, Himachal Pradesh, Sikkim, West Bengal); Nepal; Bhutan; Pakistan (Azad Kashmir)
36 CHI TIB 38 TAI 40 BHU IND NEP PAK
Remarks: D. zayuensis is the correct name fide Fraser-Jenkins et al. (2015)
Synonyms: hidden
= Dryopteris discreta Ching & S.K.Wu; Fl. Xizangica 1: 251 (1983)
= Dryopteris incisolobata Ching; Fl. Xizangica 1: 249 (1983) (fide Fraser-Jenkins 1997)
= Dryopteris nyalamense Ching & S.K.Wu; Fl. Xizangica 1: 256 (1983)
= Dryopteris nyalamense var. angustipinna Ching & S.K.Wu; Fl. Xizangica 1: 256 (1983)
= Dryopteris qamdoensis Ching; Fl. Xizangica 1: 252 (1983)
= Dryopteris squamifera Ching & S.K.Wu; Fl. Xizangica 1: 250 (1983)
= Dryopteris tsangpoensis Ching; Fl. Xizangica 1: 250 (1983)
species :
Dryopteris ×alejandrei Pérez Carro & Fern.Areces; Flora Montiber. 37: 33 (2007)
Distribution: Spain
12 SPA
species :
Dryopteris ×algonquinensis D.M.Britton; Canad. Field-Naturalist 89(2): 165 (1975)
Name: Algonquin woodfern
Distribution: Canada (Ontario)
Remarks: Dryopteris fragrans x marginalis
species :
Dryopteris ×alpirsbachensis Freigang, Zenner, Bujnoch, S.Jess. & Magauer; Kochia 10: 27 (2017)
Distribution: Germany (Baden-Württemberg, Bayern); Austria; France; Italy
11 AUT GER 12 FRA 13 ITA
Remarks: Dryopteris carthusiana x remota
species :
Dryopteris ×ambroseae Fraser-Jenk. & Jermy; Fern Gaz. 11(5): 338 (1977)
Distribution: England; Czech Republic; Germany (Baden-Württemberg, Bayern, Hessen, Nordrhein-Westfalen, Sachsen); Europe; European Russia
10 GRB 11 CZE GER 14 RUS
Remarks: Dryopteris dilatata (Hoffm.) A. Gray x Dryopteris expansa (C. Presl) Fraser-Jenk.
species :
Dryopteris ×apuana Gibby, S.Jess. & Marchetti; Atti Convegn. Internaz. 1994 - Pteridofite Lunigiana storica (Mem. Accad. Lunig. Sci., 66) 92 (1996)
Distribution: Italy
13 ITA
Remarks: Dryopteris mindshelkensis x villarii
species :
Dryopteris ×arecesiae Pérez Carro & T.E.Díaz; Anal. Jard. Bot. Madrid 47(1). 239 (1989 publ. 1990)
Distribution: Spain
12 SPA
Remarks: Dryopteris aemula x corleyi
species :
Dryopteris ×asturiensis Fraser-Jenk. & Gibby; Fern Gaz. 13(2): 113 (1986)
Distribution: Spain
12 SPA
Remarks: Dryopteris affinis ssp. affinis x corleyi
species :
Dryopteris ×australis (Wherry) Small; Ferns S. E. States 279 cum tab., 477 (1938)
Distribution: USA (Alabama, Arkansas, Georgia, Louisiana, Maryland, Mississippi, North Carolina, South Carolina, Tennessee, Virginia)
78 USA
Remarks: Dryopteris celsa x Dryopteris ludoviciana
Synonyms: hidden
= Dryopteris clintoniana var. australis Wherry; Amer. Fern J. 27: 2 (1937)
= Dryopteris clintoniana f. australis (Wherry) Wherry; Amer. Fern J. 40: 120 (1950)
= Dryopteris cristata var. australis (Wherry) Blomq. & Correll
species :
Dryopteris ×benedictii (Farw.) Wherry; Amer. Fern J. 51: 34 (1961)
Distribution: USA (Connecticut, Massachusetts, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Vermont); Canada (Ontario)
75 USA
Remarks: Dryopteris clintoniana x Dryopteris spinulosa
Synonyms: hidden
= Filix benedictii Farw.; Papers Michigan Acad. Sci. 2. 16 (1922)
= Filix-Mas benedictii (Farw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Filix ×benedictii (Farw.) Farw.; Pap. Michigan Acad. Sci. 2: 15 (1923)
species :
Dryopteris ×boottii (Tuck.) Underw.; Our nat. Ferns ed. IV: 117 (1893)
Distribution: USA (Connecticut, Delaware, Iowa, Illinois, Indiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, Tennessee, Virginia, Vermont, Wisconsin, West Virginia); Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec)
74 USA 75 USA 78 USA
Remarks: Dryopteris cristata x Dryopteris intermedia
Synonyms: hidden
= Aspidium boottii Tuck.; Hovey´s Mag. 9. 145 (1843)
= Aspidium spinulosum var. boottii (Tuck.) A.Gray; Manual (Gray), ed. 2. 598 (1856)
= Aspidium uliginosum (Newman) Nym.; Consp. Fl. Eur. 866 (1884)
= Dryopteris boottii f. multiflora (Gilbert) M.Broun
= Dryopteris ×rigida A.Gray; Man. ed. 1: 631 (1848)
= Filix boottii (Tuck.) Farw.; Rep. (Annual) Michigan Acad. Sci. 18: 81 (1916)
= Filix-Mas boottii (Tuck.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
= Lastrea bootii (Tuck.) Nieuwl.; Midl. Naturalist 2: 278 (1912)
= Lastrea uliginosa Newman; Phytol. 3: 679 (1849)
= Lophodium uliginosum (Newman) Newman; Phyt. 4: 371 (1851)
= Nephrodium boottii Baker; Syn. 275 (1867)
= Nephrodium boottii f. multiflorum Gilbert; List N. Amer. Pterid. 17, 36 (1901)
= Nephrodium cristatum var. uliginosum Baker; Syn. 273 (1867)
= Nephrodium uliginosum (Newman) Baker; Syn. 273 (1887)
= Nephrodium uliginosum (Newman) Rouy; Fl. France 14: 414 (1913)
= Polystichum boottii (Tuck.) Gand.; Fl. Eur. 27. 168 (1891)
= Polystichum ×uliginosum (Newman) P.Fourn.; Quatre Fl. France 17 (1934)
= Thelypteris bootii (Tuck.) Nieuwl.; Midl. Naturalist 1: 226 (1910)
= Thelypteris boottii f. multiflora (Gilbert) Dole; Fl. Vermont, ed. 3 10 (1937)
= Thelypteris boottii var. multiflora (Gilbert) Mousley
species :
Dryopteris ×brathaica Fraser-Jenk. & Reichst.; Fern Gaz. 11(5): 337 (1977)
Distribution: England; Germany (Baden-Württemberg, Bayern, Niedersachsen); Austria; Macedonia; +NW-England; European Russia; Canada (Québec)
10 GRB 11 AUT GER 13 NMA 14 RUS
Remarks: Dryopteris carthusiana x filix-mas
species :
Dryopteris ×burgessii Boivin; Nat. Canad. 93: 272 (1966)
Distribution: USA (Connecticut, Massachusetts, Maryland, New Hampshire, New Jersey, New York, Pennsylvania, Vermont); Canada (Ontario, Québec)
75 USA 78 USA
Remarks: ? Dryopteris cristata var. clintoniana x marginalis
species :
Dryopteris ×cantabrica Alejandre, Pérez Carro & Fern.Areces; Flora Montiber. 63: 69 (2016)
Distribution: Spain
12 SPA
Remarks: D. corleyi x D. guanchica
species :
Dryopteris ×cedroensis Gibby & Widén; Fern Gaz. 12(5): 267 (1983)
Distribution: Canary Isl. (La Gomera)
21 CNY
Remarks: Dryopteris guanchica x oligodonta
species :
Dryopteris ×complanata Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007)
Distribution: Germany (Baden-Württemberg, Nordrhein-Westfalen)
11 GER
Remarks: Dryopteris filix-mas x pseudodisjuncta
species :
Dryopteris ×complexa Fraser-Jenk.; Sommerfeltia 6: xi (1987) [nothossp.]
subspecies :
subsp. complexa
Distribution: Germany (Baden-Württemberg, Nordrhein-Westfalen, Saarland, Sachsen-Anhalt); Portugal; Spain; France; Belgium; Luxembourg; England; Scotland; Wales; Switzerland; Italy; Austria; Romania
10 GRB 11 AUT BGM GER LUX SWI 12 FRA POR SPA 13 ITA ROM
Remarks: affinis agg., Dryopteris affinis ssp. affinis x D. filix-mas
subspecies :
subsp. contorta Fraser-Jenk.; Sommerfeltia 6: xii (1987) [nothossp.]
Distribution: Wales; England
10 GRB
Remarks: affinis agg., Dryopteris affinis ssp. paleaceolobata x D. filix-mas
subspecies :
subsp. eschelmuelleri Fraser-Jenk.; Fern Gaz. 18(1): 10 (2007) [nothossp.]
Distribution: Germany (Bayern); Switzerland; Croatia; Turkey (Anatolia)
11 GER SWI 13 CRO 34 TUR
Remarks: affinis agg., Dryopteris affinis ssp. punctata x D. filix-mas
Synonyms: hidden
= Dryopteris ×complexa var. eschelmuelleri (Fraser-Jenk.) Fraser-Jenk. & Parris; Indian J. Forest. 43(3): 279 (2021)
subspecies :
subsp. transsilvanica Tlalka, S.Jess., A.Rostañski & Rojek; Fern Gaz. 21(1): 23-34 (2019) [nothossp.]
Distribution: S-Poland; C-Romania; Macedonia
11 POL 13 NMA ROM
Remarks: affinis agg., Dryopteris affinis ssp. jessenii x D. filix-mas
species :
Dryopteris ×convoluta Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007) [nothossp.]
subspecies :
subsp. convoluta
Distribution: Corsica; Switzerland; France; Germany (Baden-Württemberg, Bayern)
11 GER SWI 12 COR FRA
Remarks: Dryopteris filix-mas x cambrensis ssp. insubrica
subspecies :
subsp. inconspicua Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007) [nothossp.], non D. inconspicua Copel. 1917
Distribution: France; Scotland; Scotland (Arran)
10 GRB 12 FRA
Remarks: Dryopteris filix-mas x cambrensis ssp. pseudocomplexa
Synonyms: hidden
= Dryopteris ×neoinconspicua P.D.Sell; Fl. Great Britain and Ireland 1: 671 (2018), nom. nov.
= Dryopteris ×complexa nothosubsp. inconspicua (Fraser-Jenk.) Fraser-Jenk.; Indian Fern J. 39(1): 121 (2022)
subspecies :
subsp. occidentalis Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007) [nothossp.]
Distribution: Scotland; Wales; England; France; Corsica; Ireland
10 GRB IRE 12 COR FRA
Remarks: Dryopteris filix-mas x cambrensis ssp. cambrensis
species :
Dryopteris ×correllii W.H.Wagner; Amer. Fern J. 91(1): 11 (2001)
Distribution: USA (Vermont)
75 USA
species :
Dryopteris ×critica (Fraser-Jenk.) Fraser-Jenk.; Fern Gaz. 18(1): 11 (2007)
Distribution: France; Belgium; Luxembourg; Scotland; Wales; England; Norway; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Saarland, Sachsen); Switzerland; Italy; Austria; Hungary; Croatia; Czech Republic; Romania; Bulgaria; European Turkey; ?Spain
10 GRB NOR 11 AUT BGM CZE GER HUN LUX SWI 12 FRA 13 BUL CRO ITA ROM TUE
Remarks: Dryopteris borreri x filix-mas
Synonyms: hidden
= Dryopteris ×complexa nothosubsp. critica Fraser-Jenk. [Sommerfeltia 6 (1987)]
species :
Dryopteris ×deweveri (Jansen) Jansen & Wacht.; Heukels & Wachter, Geillustr. Schoolfl. Nederl. ed. II: 93 (1934)
Distribution: Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Mecklenburg-Vorpommern, Rheinland-Pfalz, Sachsen); Czech Republic; Slovakia; England; Scotland; Wales; Ireland; etc.
10 GRB IRE 11 CZE GER SLO
Remarks: Dryopteris carthusiana x dilatata
Synonyms: hidden
= Aspidium ×deweveri Jansen; Nederl. Kruidk. Arch. 1932: 298 (1932)
species :
Dryopteris ×doeppii Rothm.; 93 (1946)
Distribution:
Remarks: Dryopteris affinis x carthusiana
species :
Dryopteris ×doluchanovii Askerov; Izv. Akad. Nauk Az. SSR Biol. Nauk. 1978(4): 4 (1978 publ. 1979)
Distribution: Northern Caucasus
33 NCS
Remarks: Dryopteris dilatata x pseudomas
species :
Dryopteris ×dowellii (Farw.) Wherry; Amer. Fern J. 51: 34 (1961)
Distribution: USA (Connecticut, Massachusetts, Maine, Michigan, New Hampshire, New Jersey, New York, Pennsylvania, Vermont); Canada (Ontario)
75 USA
Remarks: Dryopteris clintoniana x Dryopteris intermedia
Synonyms: hidden
= Filix dowellii Farw.; Papers Michigan Acad. Sci. 2. 16 (1922)
= Filix-Mas dowellii (Farw.) Farw.; Amer. Midl. Naturalist 12: 234-251 (1931)
species :
Dryopteris ×euxinensis Fraser-Jenk. & Corley; Brit. Fern Gaz. 10: 225 (1972)
Distribution: Turkey (Anatolia); Northern Caucasus
33 NCS 34 TUR
Remarks: Dryopteris caucasica x filix-mas
species :
Dryopteris ×fraser-jenkinsii Gibby & Widén; Fern Gaz. 12(5): 268 (1983)
Distribution: Spain
12 SPA
Remarks: Dryopteris affinis ssp. affinis x D. dilatata or D. guanchica
species :
Dryopteris ×fujipedis Kurata; J. Geobot. 14: 35 (1965)
Distribution: Japan
38 JAP
Remarks: Dryopteris crassirhizoma x lacera
species :
Dryopteris ×furadensis Bennert, Rasbach, K.Rasbach & Viane; Fern Gaz. 14(1): 10 (1991)
Distribution: Madeira (Madeira Isl.)
21 MDR
Remarks: Dryopteris aitoniana x intermedia ssp. maderensis
species :
Dryopteris ×ghatakii Fraser-Jenk.; Bull. Brit. Mus. (Nat. Hist.) Bot. 18(5): 460 (1989)
Distribution: S-India (Tamil Nadu)
40 IND
Remarks: Dryopteris austroindica x cochleata
species :
Dryopteris ×gomerica Gibby & Widén; Fern Gaz. 12(5): 267 (1983)
Distribution: Canary Isl. (La Gomera)
21 CNY
Remarks: Dryopteris aemula x guanchica
species :
Dryopteris ×gotenbaensis Nakaike; J. Geobot. 17: 36, fig. (1969)