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009.0000 Family Hymenophyllaceae Mart.; Consp. Regn. Veg. [Martius] 3 (1835)
9 genera, ca. 620 spp. Circumscription of genera see Ebihara et al. (2006)
Species listed in database: 559; hybrids: 4
= Cardiomanioideae K.Iwats.; Acta Phytotax. Geobot. 35(4-6): 177 (1984)
= Trichomanaceae Burmeist.; Handb. Naturgesch. 1, 196: 12-17 (1836)
009.1001 Genus Cephalomanes C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 17 (1843)
Distribution: Tropical Old World
Species listed in database: 8

Search for: Cephalomanes javanicum-var.-sumatranum

Nothing found!
species :
Cephalomanes atrovirens C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 18, t. 5 (1843)
Distribution: Ryukyu Isl. (Iriomote); Borneo (Sabah); Sulawesi; Palawan; Philippines; Moluccas (Seram, Halmaheira, Tanimbar, Aror); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Britain, Admiralty Isl., New Hanover); Solomon Isl. (Bougainville, Choiseul, Gizo, Guadalcanal, Santa Ysabel, Kirakira, Kolombangara, Makira, Malaita, Mono, New Georgia, Nggatokae, Ranongga, Rendova, Russell, Shortland, Tetepare, Ulawa, Vangunu, Vella Lavella, Wagina); Australia (NE-Queensland, NE-New South Wales); Lord Howe Isl.; Santa Cruz Isl.
38 NNS 42 BOR MOL NWG PHI PLW SUL 43 BIS SOL 50 AUS LHI 60 SCZ
Synonyms: hidden
= Cephalomanes acranthum (H.Itô) Tagawa; Acta Phytotax. Geobot. 14: 45 (1950)
= Trichomanes acranthum H.Itô; Iconogr. Pl. Asiae Orient. 2: 110, t. 45 (1937)
= Trichomanes atrovirens (C.Presl) Kunze; Bot. Zeitung (Berlin) 371 (1847)
= Trichomanes javanicum var. atrovirens (C.Presl) C.Chr.; Index Filic. 635 (1906)
form :
f. acrosorum (Copel.) K.Iwats.; J. Jap. Bot. 66(3): 141 (1991)
Distribution: New Guinea; Solomon Isl. (Rendova)
42 NWG 43 SOL
Synonyms: hidden
= Cephalomanes acrosorum (Copel.) Copel.; Philipp. J. Sci. 67: 68 (1938)
= Trichomanes acrosorum Copel.; Philipp. J. Sci. 6: 72 (1911)
= Trichomanes acrosorum var. alatum Alderw.; Bull. Jard. Bot. Buitenzorg, Sér. 2, 11: 23 (1913)
form :
f. kingii (Copel.) K.Iwats.; J. Jap. Bot. 66(3): 141 (1991)
Distribution: New Guinea; Santa Cruz Isl. (Vanikoro)
42 NWG 60 SCZ
Synonyms: hidden
= Cephalomanes kingii (Copel.) Copel.; Philipp. J. Sci. 67: 68 (1938)
= Trichomanes kingii Copel.; Philipp. J. Sci. 6: 72 (1911)
species :
Cephalomanes boryanum (Kunze) Bosch; Ned. Kruidk. Arch. 4: 351 (1859)
Distribution: Bismarck Arch.; Solomon Isl.; New Caledonia; Vanuatu; Wallis & Futuna (Alofi, Futuna); Tonga (Late Isl., Tofua, ´Eua); Fiji (Viti Levu, Vanua Levu, Kadavu, Ovalau, Taveuni, Waya); American Samoa (Tutuila, Ofu, Olosega, Ta´u); [Western] Samoa (Savai´i, ´Upolu); Palau (Babeldaob, Ngarakabesang); Northern Marianas (Rota Isl.); Guam; Fed. States of Micronesia (Yap, Pohnpei, Kosrae)
43 BIS SOL 60 ASA FIJ NWC SAM TON VAN WAL 62 GUM MIC MRN PLU
Remarks: full species fide Zhao et al. 2025
Synonyms: hidden
= Cephalomanes alatum (Bory) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 5: 334 (1848)
= Cephalomanes atrovirens subsp. boryanum (Kunze) K.Iwats.; J. Jap. Bot. 66(3): 142 (1991)
= Cephalomanes australicum Bosch; Ned. Kruidk. Arch. 5(2): 139 (1861)
= Cephalomanes wilkesii Bosch; Ned. Kruidk. Arch. 5(2): 140 (1861)
= Lacostea boryana Prantl; Unters. Morph. Gefäßkrypt. 1: 50 (1875)
= Trichomanes alatum Bory; Voy. Monde, Crypt. 1: 282, t. 38, f. 2 (1828) (non Sw. (1801))
= Trichomanes australicum (Bosch) Copel.; Bull. Bernice P. Bishop Mus. 69: 27 (1929)
= Trichomanes boryanum Kunze; Farrnkräuter 1: 237, t. 97 (1847)
= Trichomanes javanicum var. australicum (Bosch) C.Chr.; Index Filic. 169 (1905)
= Trichomanes javanicum var. boryanum (Kunze) Fosberg; Amer. Fern J. 40(1): 136 (1950)
= Trichomanes javanicum var. wilkesii (Bosch) C.Chr.; Index Filic. 169 (1905)
species :
Cephalomanes crassum (Copel.) M.G.Price; Contr. Univ. Michigan Herb. 17: 268 (1990)
Distribution: Philippines (Samar, Leyte); Sulawesi
42 PHI SUL
Synonyms: hidden
= Trichomanes crassum Copel.; Philipp. J. Sci. 61: 256, t. 54, 55 t 3 (1933)
species :
Cephalomanes densinervium (Copel.) Copel.; Philipp. J. Sci. 67: 67 (1938)
Distribution: New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Santa Ysabel, Malaita, Shortland); Palau (Babeldaob); Fed. States of Micronesia (Yap)
42 NWG 43 SOL 62 MIC PLU
Synonyms: hidden
= Trichomanes densinervium Copel.; Philipp. J. Sci. 6: 71 (1911)
= Trichomanes infundibulare Alderw.; Nova Guinea 14: 55 (1924)
species :
Cephalomanes javanicum (Blume) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 5: 334 (1848)
Distribution: India (Assam State, Manipur, Mizoram, Tripura); Bangladesh; Nicobar Isl.; Myanmar; Thailand (SE-Thailand, peninsular Thailand); Vietnam; peninsular Malaysia; Singapore; Sumatra; Borneo (Sabah, Sarawak, Kalimantan); Brunei; Java
40 BAN IND 41 MYA NCB THA VIE 42 BOR BRN JAW MLY SNG SUM
Remarks: some formerly listed distribution belongs to C. laciniatum
Conservation Status: endangered (Singapore)
Synonyms: hidden
= Lacostea javanica (Blume) Prantl; Unters. Morph. Gefäßkrypt. 1: 50 (1875)
= Trichomanes foersteri Rosenst.; Repert. Spec. Nov. Regni Veg. 13: 213 (1914)
= Trichomanes javanicum Blume; Enum. Pl. Javae 2: 224 (1828)
= Trichomanes rigidum Wall.; Numer. List [Wallich] no. 161 (1829), nomen
= Trichomanes setigerum Wall.; Numer. List [Wallich] no. 158 (1829), nomen
species :
Cephalomanes laciniatum (Roxb.) De Vol; Fl. Taiwan 1: 98 (1975)
Distribution: Ryukyu Isl.; Taiwan; peninsular Malaysia (only collected on Pulau Tioman, Pahang); Borneo (Sabah, Kalimantan); Brunei; Sulawesi; Palawan; Philippines; Java; Lesser Sunda Isl. (Bali, Sumbawa); Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea, Louisiade Arch.: Rossel Isl.); Bismarck Arch. (Umboi Isl.); Solomon Isl. (Kolombangara, Makira, New Georgia, Nggatokae, Ranongga, Rendova, Santa Ysabel, Vangunu)
38 NNS TAI 42 BOR BRN JAW LSI MLY MOL NWG PHI PLW SUL 43 BIS SOL
Remarks: full species fide Zhao et al. 2025
Synonyms: hidden
= Cephalomanes asplenioides C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 5: 334 (1848)
= Cephalomanes curvatum Bosch; Ned. Kruidk. Arch. 4: 351 (1859), nom. nud.
= Cephalomanes javanicum var. asplenioides (C.Presl) K.Iwats.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13(5): 549 & 548 (1985) [errore (C. Chr.)]
= Cephalomanes ledermannii (Brause) Copel.; J. Arnold Arbor. 24: 440 (1943)
= Cephalomanes oblongifolium C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 5: 334 (1848)
= Cephalomanes rhomboideum J.Sm. ex Bosch; Ned. Kruidk. Arch. 4: 350 (1859), nom. nud.
= Cephalomanes zollingeri (Bosch) Bosch; Ned. Kruidk. Arch. 4(4): 351 (1859)
= Lacostea rhomboidea (Bosch) Prantl; Unters. Morph. Gefäßkrypt. 1: 50 (1875), nom. nud.
= Lacostea zollingeri (Bosch) Prantl; Unters. Morph. Gefäßkrypt. 1: 50 (1875)
= Trichomanes asplenioides C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 15, 37 (1843) (non Sw. (1788))
= Trichomanes curvatum J.Sm.; J. Bot. [Hooker] 3: 417 (1841), nom. nud.
= Trichomanes javanicum var. asplenioides (C.Presl) C.Chr.; Index Filic. 635 (1906)
= Trichomanes javanicum var. curvatum (Bosch) C.Chr.; Index Filic. 638 (1906), nom. nud.
= Trichomanes javanicum var. intercalatum Christ; Philipp. J. Sci., C 2: 156 (1907)
= Trichomanes laciniatum Roxb.; Calcutta J. Nat. Hist. 4: 518 (1844)
= Trichomanes ledermannii Brause; Bot. Jahrb. Syst. 56: 35 (1920)
= Trichomanes maluense Brause; Bot. Jahrb. Syst. 56: 36 (1920)
= Trichomanes oblongifolium C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 129 (1843)
= Trichomanes preslii C.V.Morton; Contr. U.S. Natl. Herb. 38: 190 (1968), nom. nov. for T. asplenioides C. Presl
= Trichomanes rhomboideum J.Sm.; J. Bot. [Hooker] 3: 417 (1841), nomen
= Trichomanes suffrutex Alderw.; Nova Guinea 14: 66 (1924)
= Trichomanes zollingeri Bosch; Pl. Jungh. [Miquel] 5: 552 (1857)
species :
Cephalomanes singaporianum Bosch; Ned. Kruidk. Arch. 4: 351 (1859)
Distribution: peninsular Malaysia (Malacca); Singapore; Sumatra; Borneo (Sabah, Sarawak, Kalimantan); Brunei
42 BOR BRN MLY SNG SUM
Conservation Status: vulnerable (Singapore)
Synonyms: hidden
= Lacostea singaporiana (Bosch) Prantl; Unters. Morph. Gefäßkrypt. 50 (1875)
= Trichomanes borneense Alderw.; Bull. Jard. Bot. Buitenzorg, Sér. 2, 20: 25 (1915)
= Trichomanes christii Rosenst.; Bull. Jard. Bot. Buitenzorg, Sér. 2, 11: 27 (1911), nom. illeg., non Copel. (1906)
= Trichomanes javanicum Hook. & Grev.; Icon. Filic. t. 240 (1829) (non Blume)
= Trichomanes rosenstockii Alderw.; Bull. Jard. Bot. Buitenzorg, Sér. 2, 7: 27 (1912)
= Trichomanes singaporianum (Bosch) Alderw.; Bull. Jard. Bot. Buitenzorg, Sér. 2, 20: 25 (1925)
species :
Cephalomanes sumatranum (Alderw.) Copel.; Philipp. J. Sci. 67: 67 (1938)
Distribution: China (Hainan); Thailand; Laos; Cambodia; Vietnam; ?peninsular Malaysia; Sumatra; Borneo (Sabah, Sarawak, Kalimantan); Brunei; Java
36 CHI 41 CBD LAO THA VIE 42 BOR BRN JAW SUM
Remarks: full species fide Zhao et al. 2025
Synonyms: hidden
= Cephalomanes javanicum var. sumatranum (Alderw.) K.Iwats.; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 13(5): 549 (1985)
= Trichomanes sumatranum Alderw.; Bull. Dépt. Agric. Indes Néerl. 18: 4 (1908)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.11 (Nov. 11th, 2025)

Covering 357,753 accepted species,
8,806 hybrids, 48,075 accepted infraspecific taxa and 1,004,944 synonyms,
in total 1,419,578* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

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