512.0000
Family
Asteraceae Dumort.; Comment. Bot. 55 (1822)
1620 genera, ca. 29.000 spp. (35.000 spp. incl. microspecies). Systematics taken mostly from Funk et al. (2009, 2020) and other recent publications
Species listed in database: 35235; hybrids: 679
= Acarnaceae Link; (1829)
= Ambrosiaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820), nom. cons.
= Anthemidaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Aposeridaceae Raf.; New Fl. N. Amer. 4: 106 (1838)
= Arctotaceae Bessey; Nebraska Univ. Stud. 14: 107 (1914)
= Arctotidaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Artemisiaceae Martinov; Tekhno-Bot. Slovar: 48 (1820)
= Athanasiaceae Martinov; Tekhno-Bot. Slovar: 56 (1820)
= Calendulaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Carduaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Carlinaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Centaureaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Cichoriaceae Juss.; Gen. Pl.: 168 (1789), nom. cons.
= Cnicaceae Vest; Anleit. Stud. Bot.: 273, 297 (1818)
= Compositae Giseke; Prael. Ord. Nat. Pl.: 538 (1792), nom. cons.
= Coreopsidaceae Link; Handbuch 1: 768 (1829)
= Cynaraceae Burnett; Outlines Bot.: 926, 931, 1094, 1111 (1835)
= Echinopaceae Bercht. & J.Presl; Prir. Rostlin: 255 (1820)
= Eupatoriaceae Bercht. & J.Presl; Prir. Rostlin: 253 (1820)
= Gnaphaliaceae Link ex F.Rudolphi; Syst. Orb. Veg.: 46 (1830)
= Grindeliaceae Rchb. ex A.Heller; Muhlenbergia 2: 330 (1907)
= Heleniaceae Raf.; Cincinnati Lit. Gaz. 2: 28 (1824)
= Helianthaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Helichrysaceae Link; Handbuch 1: 712 (1829)
= Inulaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Ivaceae Raf.; Fl. Tellur. 1: 31 (1837)
= Lactucaceae Drude; J. A. Schenk, Handb. Bot. 3(2): 369 (1886)
= Lapsanaceae Martinov; (1820)
= Madiaceae A.Heller; Muhlenbergia 2: 332 (1907)
= Matricariaceae Voigt; Hort. Suburb. Calcutt.: 400 (1845)
= Mutisiaceae Burnett; Outlines Bot.: 934, 935, 1094, 1111 (1835)
= Nassauviaceae Burmeist.; Handb. Naturgesch.: 290 (1836)
= Partheniaceae Link; Handbuch 1: 816 (1829)
= Perdiciaceae Link; Handbuch 1: 728 (1829)
= Picridaceae Martinov; Tekhno-Bot. Slovar: 482 (1820)
= Santolinaceae Martinov; Tekhno-Bot. Slovar: 560 (1820)
= Senecionaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Serratulaceae Martinov; Tekhno-Bot. Slovar: 577 (1820)
= Tanacetaceae Vest; Anleit. Stud. Bot.: 273, 298 (1818)
= Tussilagaceae Bercht. & J.Presl; Prir. Rostlin: 254 (1820)
= Vernoniaceae Burmeist.; Handb. Naturgesch.: 296 (1836)
= Xanthiaceae Vest; Anleit. Stud. Bot.: 273, 298 (1818)
= Xeranthemaceae Döll; Rhein. Fl.: 498 (1843)
512.3201
Genus
Cousinia Cass.; Diot. Sci. Nat. 47: 503 (1827)
Distribution: mostly Iran (200), Afghanistan (154), C-Asia, the rest from the E-Mediterranean to Mongolia, China (only 11) and W-Himalayas, north to Caucasus and Middle Asia. The borders between Cousinia and Arctium have been redefined by Lopez-Vinyallonga et al. (2011).
Species listed in database: 674; hybrids: 12
= Auchera DC.; Coll. Mem. 9: t. 11, Prod. 6: 557 (1837)
= Lipskyella Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 323 (1937)
= Tiarocarpus Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 108: 5 (1971 publ. (1972)
species :
Cousinia abbreviata
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 87 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia acanthoblephara
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 70 (1955)
Distribution: Afghanistan (Badakshan, Kabul, Wardak, Parwan)
34 AFG
species :
Cousinia acanthodendron
Rech.f. & Podlech; Pl. Syst. Evol., 142(1-2): 2 (1983)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia acanthostephana
Rech.f.; Fl. Iranica 90: 186 (1972)
Distribution: Afghanistan (Ghazni)
34 AFG
species :
Cousinia acrodroma
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 89 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia actinacantha
Rech.f.; Fl. Iranica 90: 302 (1972)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia actinia
Boiss.; Diagn. Ser. II. 3: 55 (1856)
subspecies :
subsp. actinia
Distribution: Afghanistan (Baghlan, Kabul)
34 AFG
Synonyms: hidden
= Arctium actinia (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
subspecies :
subsp. brachyphylla
Rech.f.; Fl. Iranica 90: 40 (1972)
Distribution: Afghanistan (Kabul)
34 AFG
species :
Cousinia adenophora
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 320 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia adenosticta
Bornm.; Bull. Herb. Boiss., ser. 2, 7: 212, t. 3, f. 2 (1907)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia adnata
Bunge; Mem. Acad. Sci. Pétersb. 7 sir., 9 2: 50 (1865)
Distribution: Iran (Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium adnatum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia affinis
Schrenk; Enum. Pl. Nov. 1: 41 (1841)
Distribution: Kazakhstan; Uzbekistan; China (Xinjiang); Mongolia
32 KAZ UZB 36 CHI 37 MON
Synonyms: hidden
= Arctium affine (Schrenk) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia wolgensis var. affinis (Schrenk) Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 317 (1879)
species :
Cousinia afghanica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 273 (1893)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts.); Afghanistan (Farah, Herat, Kandahar)
34 AFG IRN
Synonyms: hidden
= Cousinia cynaroides C.Winkl. ex Aitch. & Hemsl.; Trans. Linn. Soc. Ser. II. 3: 79 (1886)
species :
Cousinia agelocephala
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 89 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia aggregata
DC.; Prodr. [A. DC.] 6: 553 (1838)
Distribution: Iran (EC-Iran, N-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Cousinia aggregata var. glabra Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 135 (2017)
= Cousinia pauciflora Bunge; Mém. Acad. Imp. Sci. Saint Pétersbourg, Sér. 7, 9(2) [Gatt. Cousinia]: 12 (1865)
species :
Cousinia agridaghensis
Tugay, Ertugrul & Ulukus; Phytotaxa 427(4): 260 (2019)
Distribution: Turkey
34 TUR
species :
Cousinia aintabensis
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 503 (1875)
Distribution: Turkey (S-Anatolia, SE-Anatolia); Iraq (NE-Iraq, NW-Iraq); Syria (C-Syrian Desert)
34 IRQ SYR TUR
Synonyms: hidden
= Arctium aintabense (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia arbelensis C.Winkl. & Bornm.; Bull. Herb. Boiss. 3 11: 567, t. 14 (1895)
= Cousinia arbelensis var. pinnata C.Winkl. & Bornm.; Bull. Herb. Boissier 3: 568 (1895)
= Cousinia birecikensis Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 53 (1972)
= Cousinia handelii Bornm.; Oesterr. Bot. Z. 62: 187 (1912)
= Cousinia sinjarensis Rech.f.; Publ. Cairo Univ. Herb.: 289 (1977)
species :
Cousinia aitchisonii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 275 (1893)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.); Afghanistan (Herat)
32 TKM 34 AFG IRN
Synonyms: hidden
= Cousinia lasiosiphon Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 311 (1937)
species :
Cousinia alaica
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 90 (1962)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia alata
Schrenk ex Fisch. & C.A.Mey.; Enum. Pl. Nov. 1: 40 (1841)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; China (Xinjiang)
32 KAZ TKM UZB 36 CHI
Synonyms: hidden
= Arctium alatum (Schrenk) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
species :
Cousinia albertoregelia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 91 (1887)
Distribution: Tajikistan
32 TZK
species :
Cousinia albescens
C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 223 (1896)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia albescens var. subsphaerica Bornm.; (1906)
species :
Cousinia albida
DC.; Prodr. [A. DC.] 6: 552 (1838)
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium albidum (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia albiflora
(Bornm. & Sint.) Bornm.; Beih. Bot. Centralbl. 34 2: 285 (1916)
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Cousinia hypopolia var. albiflora Bornm. & Sint.; (1907)
species :
Cousinia alepidea
Boiss.; Diagn. Ser. II. 3: 53 (1856)
Distribution: Afghanistan (Ghazni, Kabul, Wardak, Paktia / Khost, Zabul); Pakistan (Baluchistan, Waziristan)
34 AFG 40 PAK
Synonyms: hidden
= Arctium alepidea (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia aleppica
Boiss.; Diagn. Pl. Orient., ser. 1, 10: 101 (1849)
Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon); Syria (C-Syrian Desert, NW-Syria)
34 LEB SYR TUR
species :
Cousinia alexeenkoana
Bornm.; Oesterr. Bot. Zeitschr. 62: 107, t. 2, f. 2 (1912)
Distribution: Iran (EC-Iran, W-Iran)
34 IRN
variant :
var. nekarmanica
(Rech.f.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 101 (1980)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia nekarmanica Rech.f.; Ann. Naturhist. Mus. Wien 57: 78 (1950)
species :
Cousinia alfredii
Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 49: 264 (1940)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Cousinia kelardashtiana Parsa; (1948)
species :
Cousinia algurdina
Rech.f.; Anz. Österr. Akad. Wiss. Math.-Naturwiss. Kl. 101: 344 (1964)
Distribution: Iraq (NE-Iraq)
34 IRQ
Synonyms: hidden
= Cousinia cynaroides var. arlgirdensis Blakelock; (1949)
species :
Cousinia aligudarzensis
Attar & Ghahr.; Novon 17(2): 145 (2007)
Distribution: Iran
34 IRN
species :
Cousinia allolepis
Tscherneva & Vved.; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 91 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia alpina
Bunge; Del. Sem. Hort. Dorp. (1845) 3
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium aciculare (Franch.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium alpicola (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium alpinum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia acicularis Franch.; Ann. Sci. Nat., Bot., sér. 6, 16: 319 (1883)
= Cousinia alpicola I.G.Borshch. ex C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 282 (1892)
= Cousinia alpicola I.G.Borshch. ex Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 8: 484 (1883)
species :
Cousinia ambigens
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 17 (1932), et Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 306, 307 (1937)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia amicorum
Tscherneva, Joharchi & F.Ghahrem.; Bot. Zhurn. (Moscow & Leningrad) 90(3): 411 (2005)
Distribution: Iran
34 IRN
species :
Cousinia ammophila
Rech.f.; Fl. Iranica 90: 67 (1972)
Distribution: Afghanistan (Ghazni)
34 AFG
species :
Cousinia amoena
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 237 (1894)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia andarabensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 284 (1966)
Distribution: Afghanistan (Baghlan)
34 AFG
species :
Cousinia angrenii
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 17 (1932)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia angusticeps
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 323 (1937)
Distribution: Turkmenistan
32 TKM
species :
Cousinia annua
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 88 (1887)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
Synonyms: hidden
= Cousinia dichacantha Lipsky; Exc. bot. au de la Mer Casp. 12 (1889)
= Lipskyella annua (C.Winkl.) Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 324 (1937)
species :
Cousinia anoplophylla
Rech.f.; Fl. Iranica 139a: 140, t. 134 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia antonowi
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 135, t. 2, f. 2 (1889)
Distribution: Turkmenistan; Iran (E-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium antonowi (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia apiculata
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 91 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia aptera
Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 170 (1882)
Distribution: Afghanistan (Bamyan, Ghazni, Ghorat, Kabul, Kandahar, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan); Pakistan (Kurram)
34 AFG 40 PAK
Synonyms: hidden
= Cousinia modesta Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 120 (1955)
species :
Cousinia arachnoidea
Fisch. & C.A.Mey. ex DC.; Prodr. [A. DC.] 6: 553 (1838)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
Synonyms: hidden
= Carduus pterocaulos Stephan ex DC.; Prodr. [A. DC.] 6: 553 (1838)
= Carduus pycnocephalus-alatus Steven ex DC.; Prodr. [A. DC.] 6: 553 (1838)
species :
Cousinia arakensis
Attar & Djavadi; Iranian J. Bot. 16(1): 166 (2010)
Distribution: Iran
34 IRN
species :
Cousinia araneosa
DC.; Prodr. [A. DC.] 6: 556 (1838)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium araneosum (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia araneotexta
Rech.f.; Fl. Iranica 90: 105 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia araxena
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 184 (1937)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
species :
Cousinia archibaldii
Rech.f.; Fl. Iranica 90: 147, t. 73 (1972)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia albescens var. adpressa Bornm.; (1914)
species :
Cousinia arctotidifolia
Bunge; Mém. Acad. Sci. Petersb. 7, ser., 9 2: 42 (1865)
Distribution: Turkmenistan; Iran (NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium arctotidifolium (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia arctotidifolia var. gracilescens Bornm. & Rchb.f.; (1944)
= Cousinia arctotidifolia var. taraxacifolia Bornm. & Gauba; (1944)
= Cousinia praetermissa Bornm.; (1941)
species :
Cousinia ardalensis
Attar & Djavadi; Iranian J. Bot. 16(1): 147 (2010)
Distribution: Iran
34 IRN
species :
Cousinia arenaria
Bunge; Mém. Acad. Sci. Pétersb. 7, sér., 9 2: 29 (1865)
Distribution: Iran (E-Iran)
34 IRN
Synonyms: hidden
= Arctium arenarium (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia argentea
Mehregan & Assadi; Willdenowia 39(2): 268 (2010), nom. nov.
Distribution: Iran
34 IRN
Synonyms: hidden
= Jurinea pungens Boiss.; Fl. Orient. 3: 569 (1875) emend. Rech.f., Repert. Spec. Nov. Regni Veg. 48: 148, t. 306, f. 1 & 2 (1940)
species :
Cousinia ariana
Bornm.; Beih. Bot. Centralbl. 34: 11. 278 (1916)
Distribution: Afghanistan (Baghlan, Bamyan, Wardak, Parwan); Pakistan (Swat, Quetta)
34 AFG 40 PAK
species :
Cousinia arida
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 190 (1893)
Distribution: Iran (E-Iran); Afghanistan (Herat)
34 AFG IRN
Synonyms: hidden
= Cousinia deserti Aitch. & Hemsl.; Trans. Linn. Soc. Ser. II. 3: (1886) 77 (non Bunge)
species :
Cousinia armena
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 182 (1937)
Distribution: Armenia
33 ARM
species :
Cousinia aspera
(Kult.) Karmysch.; Fl. Kazakhst. 9: 243 (1966)
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Cousinia chrysantha var. aspera Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., Ser. 8b, Bot. 6: 25 (1929)
species :
Cousinia assadii
Attar; Iranian J. Bot. 17(2): 153 (2011)
Distribution: Iran
34 IRN
species :
Cousinia assyriaca
Jaub. & Spach; Ill. Pl. Or. 2: 87, t. 169 (1846)
Distribution: Iran (EC-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium assyriacum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia leucochlora Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 24 (1865)
= Cousinia turkmenorum var. flaviflora Parsa; (1948)
species :
Cousinia astracanica
(Spreng.) Tamamsch.; Not. Syst. (Leningrad) 16: 468 (1954)
Distribution: S-European Russia; Kazakhstan; Turkmenistan
14 RUS 32 KAZ TKM
Synonyms: hidden
= Arctium wolgense (M.Bieb. ex Willd.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 198 (1887)
= Carduus astracanicus Biehler; Pl. Nov. Herb. Spreng.: 34 (1807)
= Carduus wolgensis M.Bieb. ex Willd.; Enum. Pl. 2: 839 (1809)
= Cousinia wolgensis (M.Bieb. ex Willd.) C.A.Mey. ex DC.; Prodr [A. DC.] 6: 555 (1838)
species :
Cousinia atrobracteata
Attar; Iranian J. Bot. 17(2): 152 (2011)
Distribution: Iran
34 IRN
species :
Cousinia atropatana
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 17 (1865)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Arctium atropatanum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia attariae
Assadi & Joharchi; Iranian J. Bot. 16(2): 192 (194-195, fig. 2) (2010)
Distribution: Iran
34 IRN
species :
Cousinia aucheri
DC.; Prodr. [A. DC.] 6: 555 (1838)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
Synonyms: hidden
= Arctium aucheri (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia montbretiana Bornm.; Repert. Spec. Nov. Regni Veg. 46: 275 (1939)
species :
Cousinia auriculata
Boiss.; Diagn. Ser. II. 3: 58 (1856)
Distribution: Tajikistan (Pamir-Alai); Afghanistan (Wakhan, Bamyan, Ghorat, Kabul, Wardak); Pakistan (Waziristan, Kurram); NW-India (Jammu & Kashmir)
32 TZK 34 AFG 40 IND PAK
Synonyms: hidden
= Arctium auriculatum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia kotandarica Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 112 (1955)
= Cousinia pamirica Bornm.; Beih. Bot. Centralbl. 34: II. 182 (1916) ex O. Fedtsch., Act. Hort. Petrop. 31: 463 (1915)
= Cousinia semidecurrens C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 574 (1837)
species :
Cousinia austrojordanica
F.G.Davies & Boulos; Publ. Cairo Univ. Herb. 7 & 8: 287 (1977)
Distribution: Jordania (S-Jordania)
34 JOR
species :
Cousinia autranii
C, Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 193 (1892)
Distribution: Afghanistan (Ghazni, Ghorat, Kabul, Wardak, Zabul)
34 AFG
Synonyms: hidden
= Cousinia ghazniensis Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 97 (1955)
species :
Cousinia azerbaidjanica
Djavadi, Attar & Najafi; Iranian J. Bot. 13(1): 44 (2007)
Distribution: Iran
34 IRN
species :
Cousinia azmarensis
S.A.Ahmad, Rastegar & Attar; Harvard Pap. Bot. 22(1): 71 (2017)
Distribution: Iraq
34 IRQ
species :
Cousinia baberi
Boiss.; Diagn. Ser. II. 3: 53 (1856)
Distribution: Afghanistan (Kabul, Logar, Wardak)
34 AFG
Synonyms: hidden
= Arctium baberi (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bachtiarica
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 475 (1875)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium bachtiaricum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia badghysi
Kult.; Korovin, Kult. & Popov, Descr. Pl. Nov. Turkmenistan. (Bodenk. u. Pflanzengeogr. Beob. u. Unters. 1: Flussgeb. d. Amu Darya u. Syr Darya, II.), 76 (1916)
Distribution: Turkmenistan
32 TKM
species :
Cousinia balchanica
Tscherneva; Novosti Sist. Vyssh. Rast. 30: 153 (1996)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia baluchistanica
Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 125: 53 (1988 publ. 1989)
Distribution: Pakistan (Baluchistan)
40 PAK
species :
Cousinia bamianica
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 77 (1955)
Distribution: Afghanistan (Bamyan, Wardak)
34 AFG
Synonyms: hidden
= Cousinia actinostegia Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 282 (1966)
species :
Cousinia baranovii
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR. 17: 92 (1962)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia barbeyi
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 274 (1893)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia barezica
Assadi; Iranian J. Bot. 15(1): 37 (2009)
Distribution: Iran
34 IRN
species :
Cousinia batalinii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 248 (1892)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia bazoftensis
Attar; Iranian J. Bot. 17(2): 154 (2011)
Distribution: Iran
34 IRN
species :
Cousinia beauverdiana
Bornm.; Oesterr. Bot. Zeitschr. 63: 55 (1913)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia arctotidifolia var. laeviseta Winkl. & Bornm.; (1897)
species :
Cousinia beckeri
Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 457 (1886)
Distribution: Turkmenistan
32 TKM
species :
Cousinia belangeri
DC.; Prodr. [A. DC.] 6: 553 (1838)
Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium belangeri (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium verutum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia carlinoides DC.; Prodr. [A. DC.] 6: 554 (1838)
= Cousinia pugionifera var. decurrens Parsa; (1948)
= Cousinia teheranica Bornm.; Beih. Bot. Centralbl. 34(2): 274 (1916)
= Cousinia verutum Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 31 (1865)
= Cousinia verutum subsp. teheranica Bornm.; (1917)
species :
Cousinia bicolor
Freyn & Sint.; Oesterr. Bot. Zeitschr. 42 6: 204 (1892)
Distribution: Turkey (E-Anatolia, NE-Anatolia)
34 TUR
species :
Cousinia bienerti
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 27 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium bienerti (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bijarensis
Rech.f.; Fl. Iranica 90: 152 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia bipinnata
Boiss.; Diagn. Ser. II. 3: 52 (1856)
Distribution: Afghanistan (Kandahar); Pakistan (Quetta, Baluchistan)
34 AFG 40 PAK
Synonyms: hidden
= Arctium bipinnatum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia birandiana
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 49 (1972)
Distribution: Turkey (Inner Anatolia, C-Anatolia)
34 TUR
species :
Cousinia blepharobasis
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 78 (1955)
Distribution: Afghanistan (Bamyan, Kabul, Wardak)
34 AFG
species :
Cousinia bobekii
Rech.f.; Fl. Iranica 90: 238, t. 120 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia bobrovii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 308 (1937)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia boissieri
Buhse; Boiss. & Buhse, Nouv. Mém. Soc. Nat. Moscou 12 18: 127 (1860)
Distribution: Turkey (E-Anatolia, SE-Anatolia); Iraq (SE-Iraq: Mesopotamia); Iran (Iranian Aserbaijan, W-Iran)
34 IRN IRQ TUR
Synonyms: hidden
= Arctium boissieri (Buhse) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia layardii Ball & Barbey; Cousinia layardi 2 (1890)
species :
Cousinia bonvalotii
Franch.; Ann. Sci. Nat. Ser. VI. 16: 320 (1883)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan
32 KAZ KGZ UZB
Synonyms: hidden
= Arctium bonvalotii (Franch.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bornmulleriana
Attar & Maroofi; Nova Biol. Reperta 9(2): 145 (2022), non C. bornmuelleri C. Winkl.
Distribution: Iran
34 IRN
species :
Cousinia botschantzevii
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 93 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia boyerahmadica
Rastegar, Attar & Mirtadz.; Phytotaxa 343(3): 228 (2018)
Distribution: Iran
34 IRN
species :
Cousinia brachyptera
DC.; Prodr. [A. DC.] 6: 554 (1838)
Distribution: Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, NE-Anatolia)
33 ARM GEO 34 TUR
Synonyms: hidden
= Arctium brachypterum (DC.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia gundelia K.Koch; Linnaea 24: 387 (1851)
= Cousinia heterolepis K.Koch; Linnaea 24: 386 (1851)
= Cousinia squarrosa K.Koch; Linnaea 24: 387 (1851)
species :
Cousinia brevicaulis
Attar, Mozaff. & Mirtadz.; Nordic J. Bot. 34(2): 176 (2016)
Distribution: Iran
34 IRN
species :
Cousinia bucharica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: (1887) 473 et 477
Distribution: Uzbekistan (Buchara)
32 UZB
Synonyms: hidden
= Arctium bucharicum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia bungeana
Regel & Schmalh.; Trudy Imp. S.-Peterburgsk. Bot. Sada 6: 318 (1879)
Distribution: Kazakhstan; Uzbekistan; Tajikistan
32 KAZ TZK UZB
Synonyms: hidden
= Arctium bungeanum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia buphthalmoides
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 155
Distribution: Tajikistan; Afghanistan (Badakshan, Wakhan, Baghlan, Bamyan, Ghorat, Kabul, Laghman, Logar, Wardak, Paktia / Khost, Parwan, Samangan); Pakistan (Kurram, Chitral, Drosh, Swat, Gilgit)
32 TZK 34 AFG 40 PAK
Synonyms: hidden
= Arctium buphthalmoides (Regel) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia auriculata Hook.f.; Fl. Brit. India 3: 360 (1881), nom. illeg.
species :
Cousinia butkovii
Tschern. & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 54 (1961)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia caesarea
Boiss. & Balansa; Boiss., Diagn. Pl. Orient., ser. 2, 6: 116 (1859)
Distribution: Turkey (Central Anatolia, Inner Anatolia)
34 TUR
Synonyms: hidden
= Arctium caesareum (Boiss. & Balansa) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia caespitosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 93 (1887)
Distribution: Kazakhstan; Kyrgyzstan; China (W-Xinjiang)
32 KAZ KGZ 36 CHI
species :
Cousinia calcitrapa
Boiss.; Fl. Orient. 3: 490 (1875)
Distribution: Iran (Iranian Aserbaijan, S-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium calcitrapa (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia larvea C.Winkl. & Bornm.; Bull. Herb. Boissier 5: 167 (1897)
variant :
var. interrupta
Heimerl; Denkschr. Kaiserl. Akad. Wiss., Wien. Math.-Naturwiss. Kl. 50: 62 (1885)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia xiphacantha C.Winkl. & Strauss ex C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 225 (1897)
= Cousinia xiphacantha var. lachnopoda Bornm.; (1912)
species :
Cousinia calcitrapella
Bornm.; Notizbl. Bot. Gart. Berlin-Dahlem 15: 118 (1940)
Distribution: India (Punjab)
40 IND
species :
Cousinia calocephala
Jaub. & Spach; Ill. Pl. Or. 2: 96 (1846)
subspecies :
subsp. calocephala
Distribution: Turkey (E-Anatolia, SE-Anatolia); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, W-Iran)
34 IRN TUR
Synonyms: hidden
= Arctium calocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium squarrosum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia squarrosa Boiss.; Diagn. Pl. Orient., ser. 1, 10: 102 (1849)
= Cousinia vanensis Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 55 (1972).
subspecies :
subsp. asterocephala
(Hausskn. & Bornm.) Mehregan; Willdenowia 38(2): 348 (2008)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia asterocephala Hausskn. & Bornm.; Beih. Bot. Centralbl. 20(2): 160 (1906)
subspecies :
subsp. behboudiana
(Rech.f. & Esfand.) Mehregan; Willdenowia 38(2): 347 (2008)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia behboudiana Rech.f. & Esfand.; Ann. Naturh. Mus. Wien 57: 81 (1950)
= Cousinia squarrosa var. behboudiana (Rech.f. & Esfand.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 128 (1980)
species :
Cousinia calolepis
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 116 (1846)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium anisopterum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia anisoptera Jaub. & Spach; Illustr. 2: 88. t. 170 (1846)
species :
Cousinia calva
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 317 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia campyloraphis
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 37 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia candicans
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 300 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia candolleana
Jaub. & Spach; Ill. Pl. Or. 2: 74, t. 156 (1845)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium orientale (DC.) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Auchera orientalis DC.; Coll. Mem. 9: t. 11 (1838)
species :
Cousinia canescens
DC.; Prodr. [A. DC.] 6: 556 (1838)
Distribution: Armenia; Turkey (E-Anatolia, Inner Anatolia, NE-Anatolia, S-Anatolia, N-Anatolia, SE-Anatolia); Iran (Iranian Aserbaijan, W-Iran)
33 ARM 34 IRN TUR
Synonyms: hidden
= Arctium eriocephalum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia eriocephala Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 504 (1875)
= Cousinia satdagensis Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 55 (1972)
= Cousinia wettsteiniana Bornm.; Verh. K. K. Zool.-Bot. Ges. Wien 60: 137 (1910)
species :
Cousinia carduncelloidea
Regel & Schmalh.; Pl. Nov. Fedsch. 50 (1882)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Arctium carduncelloideum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia carduncelloidea var. sonchifolia Bornm.
= Cousinia carduncellus Regel & Schmalh. ex O.Fedtsch. & B.Fedtsch.; Consp. Fl. Turkest. 4: 252 (1902)
= Cousinia sonchifolia (Bornm.) Bornm.; Repert. Spec. Nov. Regni Veg. Beih. 108: 42 (1938)
species :
Cousinia caroli-henrici
Attar & Ghahr.; Rostaniha 7(Suppl. 2): 317 (2007)
Distribution: Iran
34 IRN
species :
Cousinia carthamoides
Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 171 (1882)
Distribution: Afghanistan (Kunar / Nuristan, Paktia / Khost); Pakistan (Kurram)
34 AFG 40 PAK
Synonyms: hidden
= Cousinia carthamoides var. stenorhachis Bornm.; (1938)
species :
Cousinia cataonica
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 483 (1875)
Distribution: Turkey (Inner Anatolia, SE-Anatolia)
34 TUR
Synonyms: hidden
= Arctium cataonicum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia catenata
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 82 (1955)
Distribution: Afghanistan (Balkh, Jawzjan / Sar-e-Pol)
34 AFG
species :
Cousinia cavarae
Bornm.; Mitt. Thür. Bot. Ver. 38: 2, 49 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia centauroides
Fisch. & C.A.Mey.; Bull. Soc. Imp. Naturalistes Moscou (1839) 159 ex Bunge
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Arctium centauroides (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia ceratophora
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 24 (1929)
Distribution: Tajikistan
32 TZK
species :
Cousinia chaetocephala
Kult.; Acta Univ. As. Med., ser. 8b 7: 3 (1929)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia chaetolepis
Rech.f.; Fl. Iranica 90: 207 (1972)
Distribution: C-Afghanistan
34 AFG
species :
Cousinia chamaepeuce
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 120 (1846)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium chamaepeuce (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia chamaepeucides Bornm.; Bull. Herb. Boiss. Ser. 2(7): 212 (1907)
species :
Cousinia chejrabadensis
Kult.; Fl. Turkm. 7: 380 (1960)
Distribution: Turkmenistan
32 TKM
species :
Cousinia chionophila
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 84 (1955)
Distribution: Afghanistan (Badakshan, Wakhan, Baghlan, Bamyan Ghazni, Kunar / Nuristan, Parwan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia chitralensis
Rech.f.; Fl. Iranica 90: 306 (1972)
Distribution: Pakistan (Chitral)
40 PAK
species :
Cousinia chlorocephala
C.A.Mey.; Prodr. [A. DC.] 6: 554 (1838)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Iran (N-Iran)
33 ARM AZE 34 IRN
Synonyms: hidden
= Arctium chlorocephalum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia intermedia C.A.Mey.; Prodr. [A. DC.] 6: 553 (1838)
species :
Cousinia chlorosphaera
Bornm.; Beih. Bot. Centralbl. 28 2: 253 (1911)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia chlorosphaera var. straminea Bornm.; (1914)
= Cousinia chlorosphaera var. producta Bornm.; (1914)
species :
Cousinia chlorothyrsa
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 86 (1955)
Distribution: Afghanistan (Bamyan, Ghorat, Orozgan / Daykundi)
34 AFG
species :
Cousinia chrysacantha
Jaub. & Spach; Ill. Pl. Or. 2: 78, t. 160 (1845)
Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium chrysacanthum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia chrysandra
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 115, t. 11, f. 1 (1941)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia chrysantha
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 25 (1929)
Distribution: Kazakhstan; Uzbekistan
32 KAZ UZB
species :
Cousinia chrysochlora
Rech.f. & Köie; Dansk Biol. Skr. 8, 2: 88 (1955)
Distribution: Iran (E-Iran); Afghanistan (Farah, Herat)
34 AFG IRN
species :
Cousinia cirsioides
Boiss. & Balansa; Boiss., Diagn. Pl. Orient., ser. 2, 6: 117 (1859)
Distribution: Turkey (Inner Anatolia)
34 TUR
Synonyms: hidden
= Arctium cirsioides (Boiss. & Balansa) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia cisdarvasica
V.V.Nikitin; Not. Syst. Herb. Inst. Bot. Acad. Sci. U.R.S.S. 19: 385 (1959)
Distribution: Tajikistan
32 TZK
species :
Cousinia coerulea
Kult. ex Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 51 (1959)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Cousinia subinermis Tschern.; Bot. Mater. Gerb. Inst. Bot. Zool. Akad. Nauk Uzbeksk. S.S.R. 15: 40 (1959)
species :
Cousinia commutata
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 24 (1865)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium commutatum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia campylacme Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 40: 340 (1936)
= Cousinia dichroantha Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 40: 344 (1936)
= Cousinia larica Parsa; Kew Bull. 3: 205 (1948)
species :
Cousinia concinna
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 513 (1875)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium concinnum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia concolor
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 33 (1865)
Distribution: Iran (EC-Iran, E-Iran)
34 IRN
Synonyms: hidden
= Arctium concolor (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia congesta
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 12 (1865)
Distribution: Turkmenistan; Tajikistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran, W-Iran); Afghanistan (Balkh, Faryab, Ghorat, Herat, Kabul, Qunduz, Samangan); Pakistan (Baluchistan)
32 TKM TZK 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium congestum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia consanguinea
(Bornm.) Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 89 (1955)
Distribution: Afghanistan (locality unknown)
34 AFG
Synonyms: hidden
= Cousinia scariosa subsp. consanguinea Bornm.; Beih. Bot. Centralbl. 34: 11. 282 (1916)
species :
Cousinia contumax
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 168, t. 4 (1897)
Distribution: Iran (EC-Iran, S-Iran)
34 IRN
species :
Cousinia cordifolia
Djavadi & Attar; Feddes Repert. 117(7-8): 453 (2006)
Distribution: Iran
34 IRN
species :
Cousinia coronata
Franch.; Ann. Sci. Nat. Ser. VI. 16: 321 (1883)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium coronatum (Franch.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia campylacantha Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 312 (1937)
species :
Cousinia corymbosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 89 (1887)
Distribution: Tajikistan
32 TZK
species :
Cousinia crassipes
Kult.; Acta Univ. As. Med., ser. 8b 7: 2 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia crispa
Jaub. & Spach; Ill. Pl. Or. 2: 81 (1846)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium crispum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia barbarae Gauba; Mitth. Thüring. Bot. Vereins n.f. 47: 116 (1941)
= Cousinia kandewanensis Parsa; Kew Bull. 3(2): 205 (1948)
species :
Cousinia cryptadena
Juz.; Not. Syst. Herb. Hort. Petrop. 5: 111 (1924)
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Cousinia leucocitrina Kult.; Bot. Mater. Gerb. Glavn. Bot. Sada S. S. S. R. 6: 30 (1926)
species :
Cousinia curvibracteata
Mehregan; Iranian J. Bot. 17(2): 144 (2011)
Distribution: Iran
34 IRN
species :
Cousinia cylindracea
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 115 (1846)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium cylindraceum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia cylindracea var. patula Heimerl; (1885)
species :
Cousinia cylindrocephala
Jaub. & Spach; Ill. Pl. Or. 2: 84, t. 166 (1846)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium cylindrocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia cynaroides
(M.Bieb.) C.A.Mey.; Verz. Pfl. Caucasus 230 (1831)
Distribution: Turkmenistan; Northern Caucasus; Armenia; Azerbaijan; Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
32 TKM 33 ARM AZE NCS 34 IRN
Synonyms: hidden
= Arctium cynaroides (M.Bieb.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Carthamus cynaroides M.Bieb.; Beschr. Länder Terek Kasp. 195 (1800)
= Onobroma cynaroides (M.Bieb.) Spreng.; Syst. Veg. ed. 16, 3: 392 (1826)
= Onopordon cynaroides (M.Bieb.) Steven ex Link; Enum. Hort. Berol. 2: 303 (1822)
species :
Cousinia czatkalica
Tscherneva; Novosti Sist. Vyssh. Rast. 29: 122 (1993)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia czerniakowskae
Kult.; Acta Univ. As. Med., ser. 8b 7: 5 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia tscherniakowskiana Kult.; (1930)
variant :
var. adenostegia
(Rech.f.) Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 64 (2017)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia adenostegia Rech.f.; Fl. Iranica 139a: 151, t. 147 (1979)
species :
Cousinia dalahuensis
Attar & Ghahr.; Iranian J. Bot. 8(2): 264 (2000)
Distribution: Iran
34 IRN
species :
Cousinia daralaghezica
Takht.; Trans. Armen. Branck Acad. Sci. USSR, Biol. Ser. 2: 181 (1937)
Distribution: Armenia; Azerbaijan (incl. Nachichevan)
33 ARM AZE
species :
Cousinia darwasica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 249 (1892)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia dasylepis
Kult.; Acta Univ. As. Med., ser. 8b 7: 4 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia davisiana
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 56 (1972)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Cousinia decipiens
Boiss. & Buhse; Nouv. Mém. Soc. Nat. Moscou 12 18: 125 (1860)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium decipiens (Boiss. & Buhse) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia daryoushiana Parsa; Kew Bull. 3(2): 203 (1948)
= Cousinia hyrcanica Bornm.; Oesterr. Bot. Z. 75: 233 (1926)
species :
Cousinia decolorans
Freyn & Sint.; Oesterr. Bot. Zeitschr. 42 6: 206 (1892)
Distribution: Turkey (Inner Anatolia, E-Anatolia)
34 TUR
species :
Cousinia decumbens
Rech.f.; Ann. Naturh. Mus. Wien 57: 77 (1950)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia meyamensis var. decumbens (Rech.f.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 99 (1980)
species :
Cousinia decurrens
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 154
Distribution: Uzbekistan; Tajikistan; Afghanistan (Badakshan, Badghys, Baghlan, Balkh, Bamyan, Faryab, Ghazni, Kabul, Wardak, Qunduz, Takhar); Pakistan (Baluchistan, Quetta)
32 TZK UZB 34 AFG 40 PAK
Synonyms: hidden
= Arctium decurrens (Regel) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia decurrentifolia
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 93 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia deluensis
Attar, Mozaff. & Mirtadz.; Phytotaxa 343(3): 231 (2018)
Distribution: Iran
34 IRN
species :
Cousinia denaensis
Attar & Djavadi; Iranian J. Bot. 16(1): 140 (2010)
Distribution: Iran
34 IRN
species :
Cousinia deserti
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 29 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.); Afghanistan (Farah, Herat, Zabul)
34 AFG IRN
Synonyms: hidden
= Arctium deserti (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia leuconeura Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 324 (1934)
variant :
var. longispinosa
Djavadi & Attar; Feddes Repert. 120(1-2): 33 (2009)
Distribution: Iran
34 IRN
species :
Cousinia dichotoma
Bunge; Del. Sem. Hort. Dorpat. (1845) 3
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
Synonyms: hidden
= Arctium dichotomum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia dichromata
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 14 (1929)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia diezii
Rech.f.; Fl. Iranica 90: 275, t. 149 (1972)
Distribution: Iran (E-Iran, NE-Iran: Mts.)
34 IRN
species :
Cousinia dimoana
Kult.; Korovin, Kult. & Popov, Descr. Pl. Nov. Turkmenistan. (Bodenk. u. Pflanzengeogr. Beob. u. Unters. 1: Flussgeb. d. Amu Darya u. Syr Darya, II.), 79 (1916)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia dipterocarpa
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 147, t. 313, f. 2 (1940)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia discolor
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 34 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium discolor (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia discolor var. glabra Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 66 (2017)
species :
Cousinia dissecta
Kar. & Kir.; Bull. Soc. Imp. Naturalistes Moscou 15: 391 (1842)
Distribution: Kazakhstan; China (NW-Xinjiang)
32 KAZ 36 CHI
Synonyms: hidden
= Arctium dissectum (Kar. & Kir.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia dissectifolia
Kult.; Trans. Sci. Soc. Turk. 1: 111 (1923)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia divaricata
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 334 (1890)
Distribution: Tajikistan
32 TZK
species :
Cousinia dolichoclada
Juz.; Not. Syst. Herb. Hort. Petrop. 2: 57 (1921)
Distribution: Uzbekistan
32 UZB
species :
Cousinia dolicholepis
Schrenk; Bull. Phys.-Math. Acad. Petersb. 3: 107 (1845)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
Synonyms: hidden
= Arctium dolicholepis (Schrenk) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia dshisakensis
Kult.; Bull. Inst. Pedol. & Geobot. Univ. As. Centr. 1: 115 (1925)
Distribution: Uzbekistan
32 UZB
species :
Cousinia dubia
Popov; Popov & Androsov, Rast. Zapovedn. Guralash i Zaaminsk. Lesnich. 37 (1936)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia eburnea
Bornm.; Oesterr. Bot. Zeitschr. 62: 105, t. 2, f. 1 (1912)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia ebrahim Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 121 (1980)
species :
Cousinia ecbatanensis
Bornm.; Oesterr. Bot. Zeitschr. 62: 184, t. 3, f. 3-3b (1912)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia echinocephala
Sennikov; Phytotaxa 5: 24 (2010)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia edelbergii
Rech.f.; Fl. Iranica 90: 182 (1955)
Distribution: C-Afghanistan
34 AFG
Remarks: provisionally accepted name
species :
Cousinia edmondsonii
Rech.f.; Fl. Iranica 90: 321 (1972)
Distribution: Iran (NE-Iran: Mts., N-Iran)
34 IRN
species :
Cousinia egens
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 304 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia elburzensis
Attar, Mahdigholi & Ghahr.; Novon 17(1): 3 (2007)
Distribution: Iran (Mazandaran)
34 IRN
species :
Cousinia eleonorae
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 50 (1972)
Distribution: Turkey (Inner Anatolia)
34 TUR
species :
Cousinia elephantina
Rech.f.; Fl. Iranica 90: 182 (1972)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia elwendensis
Bornm.; Beih. Bot. Centralbl. 28 2: 255 (1911)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia elymaitica
Attar, Mozaff. & Mirtadz.; Nova Biol. Reperta 5(3): 306, fig. 1 (2018)
Distribution: Iran
34 IRN
species :
Cousinia erectispina
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 94 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia erinacea
Jaub. & Spach; Ill. Pl. Or. 2: 83, t. 165 (1846)
Distribution: Iran (N-Iran)
34 IRN
Synonyms: hidden
= Arctium erinaceum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia lindsayana Rech.f.; Oesterr. Bot. Z. 100: 465 (1953)
species :
Cousinia eriobasis
Bunge; Mem. Acad. Sci. Pétersb. 7 sér., 9 2: 37 (1865)
Distribution: Iran (EC-Iran, S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium eriobasis (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia deickii Bornm.; Notizbl. Bot. Gart. Berlin 14: 278 (1938)
species :
Cousinia eriophylla
(Kult.) Bornm.; Rech.f., Repert. Spec. Nov. Regni Veg. 48: 147 (1940)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia freynii var. eriophylla Kult.; Acta Univ. As. Med. Ser. 8b 7: 7 (1929)
species :
Cousinia eriorrhiza
Bornm.; Beih. Bot. Centralbl. 28 2: 251 (1911)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia eriotricha
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 321 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia erivanensis
Bornm.; Vestn. Tiflissk. Bot. Sada 30: 15 (1913)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
species :
Cousinia ermenekensis
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 50 (1972)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Cousinia eryngioides
Boiss.; Diagn. Pl. Orient., ser. 1, 10: 101 (1849)
Distribution: Turkmenistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium eryngioides (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia esfandiarii
Rech.f. & Aellen; Ann. Naturh. Mus. Wien 57: 75 (1950)
Distribution: Iran (EC-Iran, NE-Iran: Mts., N-Iran)
34 IRN
species :
Cousinia euchlora
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 13 1, t. 309, f. 2 (1940)
Distribution: Iran (E-Iran)
34 IRN
Synonyms: hidden
= Cousinia euchlora var. arachnoidea Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 69 (2017)
species :
Cousinia eugenii
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 141 (1924), et Not. Syst. Herb. Hort. Bot. Russia 6: 28 (1926)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia euphratica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 51 (1972)
Distribution: Turkey (SE-Anatolia)
34 TUR
species :
Cousinia eurylepis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 95 (1955)
Distribution: Afghanistan (Bamyan, etc.)
34 AFG
species :
Cousinia fabrorum
Rech.f.; Fl. Iranica 139a: 113 (1979)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia falcinella
Bornm.; Repert. Spec. Nov. Regni Veg. 26: 179 (1929)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia falconeri
Hook.f.; Fl. Brit. India 3(8): 360 (1881)
Distribution: W-Tibet; NW-India (Zanskar, Jammu & Kashmir)
36 TIB 40 IND
Synonyms: hidden
= Arctium falconeri (Hook.f.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia fallax
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 471 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia farimanensis
Assadi; Iranian J. Bot. 17(1): 6 (2011), nom. inval., et Phytotaxa 227(1): 99 (2015)
Distribution: Iran
34 IRN
species :
Cousinia farsistanica
Bornm.; Oesterr. Bot. Zeitschr. 62: 185, t. 3, f. 2 (1912)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia fascicularis
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 299 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia fedorovii
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 183 (1937)
Distribution: Armenia
33 ARM
species :
Cousinia ferghanensis
Bornm.; Beih. Bot. Centralbl. 34: 11. 195 (1916)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Cousinia popovii Kult.; Trudy Turkestansk. Gosud. Univ. 4: 66 (1922)
species :
Cousinia ferruginea
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 142 (1924)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Cousinia truncata Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 318 (1937)
species :
Cousinia fetissowii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 247 (1892)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia finitima
Juz.; Trans. Rubb. & Guttop. Inst., Fasc. 4, 24 (1932)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia firuzkuhensis
Rech.f.; Fl. Iranica 139a: 120, t. 111 (1979)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia foliosa
Boiss. & Balansa; Boiss., Diagn. Pl. Orient., ser. 2, 6: 116 (1859)
Distribution: Turkey (Inner Anatolia, S-Anatolia, SE-Anatolia)
34 TUR
Synonyms: hidden
= Arctium foliosum (Boiss. & Balansa) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia fragilis
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 169, t. 6 (1897)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia fragillima
Rech.f.; Oesterr. Bot. Zeitschr. 100: 458, f. 2 (1953)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia franchetii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 578 (1887), nom. nov.
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium franchetianum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia canescens Franch.; Ann. Sci. Nat. Ser. VI. 16: 323 (1883)
species :
Cousinia fraternella
Bornm.; Beih. Bot. Centralbl. 34: 11. 273 (1916)
Distribution: Afghanistan (Bamyan, Ghorat, Wardak, Parwan)
34 AFG
species :
Cousinia freitagii
Rech.f.; Fl. Iranica 90: 86 (1972)
Distribution: Afghanistan (Baghlan)
34 AFG
species :
Cousinia freynii
Bornm. & Sint.; Russk. Bot. Zhurn. 1911: 4 (1911)
Distribution: Turkmenistan; Iran (NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
species :
Cousinia fursei
Rech.f.; Fl. Iranica 90: 247, t. 128 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia gabrielae
Bornm.; Beih. Bot. Centralbl. 59 2: 301 (1939)
Distribution: Iran (S-Iran); Afghanistan (Farah)
34 AFG IRN
species :
Cousinia gaharensis
Attar & Djavadi; Iranian J. Bot. 16(1): 149 (2010)
Distribution: Iran
34 IRN
species :
Cousinia gatchsaranica
Mehregan, Assadi & Attar; Willdenowia 33(1): 108 (2003)
Distribution: Iran
34 IRN
species :
Cousinia gaubae
Bornm.; Repert. Spec. Nov. Regni Veg. 36: 326, t. 181B (1934)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia gedrosiaca
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 121, t. 15 (1941)
Distribution: Iran (S-Iran, SW-Iran); Pakistan (Baluchistan)
34 IRN 40 PAK
species :
Cousinia ghahremanii
Mirtadz. & Attar; Iranian J. Bot. 15(2): 147 (2009)
Distribution: Iran
34 IRN
species :
Cousinia ghorana
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ci. 346 (1964)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia gigantolepis
Rech.f.; Fl. Iranica 90: 257, t. 136 (1972)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
species :
Cousinia gigantoptera
Rech.f. & Podlech; Pl. Syst. Evol., 142(1-2): 2 (1983)
Distribution: Afghanistan (Takhar)
34 AFG
species :
Cousinia gilanica
Bornm.; Oesterr. Bot. Zeitschr. 62: 108, t. 2, f. 3 (1912)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia gilliatii
Rech.f.; Fl. Iranica 90: 244, t. 124 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia ginuae
Attar; Taxon. Biosyst. 3(6): 60 (2011), et Ann. Bot. Fenn. 57(4-6): 238 (2020)
Distribution: Iran
34 IRN
species :
Cousinia glabriseta
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 7 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia glandulosa
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 13 (1929)
Distribution: Turkmenistan
32 TKM
species :
Cousinia glaphyra
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 99 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia glaphyrocephala
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 94 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia glaucopsis
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 134, t. 310, f. 2 (1940)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
variant :
var. spinellosissima
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 134, t. 310, f. 2 (1940)
Distribution: Iran
34 IRN
species :
Cousinia glochidiata
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 3 (1929)
Distribution: Turkmenistan
32 TKM
species :
Cousinia gmelinii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 241 (1893)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia gnezdilloi
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 47 (1959)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
species :
Cousinia golestanica
Attar; Taxon. Biosyst. 3(6): 63 (2011)
Distribution: Iran
34 IRN
species :
Cousinia gomolitzkii
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 95 (1962)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia gontscharowii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 311 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia gracilis
Boiss.; Diap., ser. 1, 6: 117 (1846)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium gracile (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia elymaea Hausskn. & Winkl. ex C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 225 (1897)
species :
Cousinia graminifolia
Rech.f.; Fl. Iranica 90: 178 (1972)
Distribution: Afghanistan (Baghlan)
34 AFG
species :
Cousinia grandis
C.A.Mey.; Prodr. [A. DC.] 6: 557 (1838)
Distribution: Armenia; Turkey (SE-Anatolia); Iran (Iranian Aserbaijan)
33 ARM 34 IRN TUR
Synonyms: hidden
= Arctium grande (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia grantii
Rech.f.; Fl. Iranica 90, 245, t. 125 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia greuteri
Rech.f.; Fl. Iranica 90: 191 (1972)
Distribution: Afghanistan (Baghlan, Samangan)
34 AFG
species :
Cousinia griffithiana
Boiss.; Diagn. Ser. II. 3: 55 (1856)
Distribution: Afghanistan (Ghazni, Zabul)
34 AFG
Synonyms: hidden
= Arctium griffithianum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia grigoriewii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 313 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia grisea
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 5 (1929)
Distribution: Uzbekistan
32 UZB
species :
Cousinia gulczensis
Kult.; Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 6, 16 (1929)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia gymnoclada
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 102 (1955)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia hablitzii
C.A.Mey.; DC., Prodr 6: 554 (1838)
Distribution: Iran (N-Iran)
34 IRN
Synonyms: hidden
= Arctium hablitzii (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Carduus macranthus (Schrank) Hablitz; Boiss., Fl. Orient. 3: 496 (1875)
= Chamaepeuce macrantha Schrank; Bull. Sci. Acad. Petersb. 10: 354 (1842)
= Cirsium macranthum (Schrenk) B.Fedtsch.; Rastitel´n. Turkestana: 755 (1915)
species :
Cousinia haeckeliae
Bornm.; Repert. Spec. Nov. Regni Veg. Beih. 108: 40 (1938)
Distribution: Afghanistan (Bamyan, Kunar / Nuristan, Parwan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia hakkarica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 54 (1972)
Distribution: Turkey (SE-Anatolia)
34 TUR
species :
Cousinia halysensis
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 51 (1972)
Distribution: Turkey (Inner Anatolia)
34 TUR
species :
Cousinia hamadae
Juz.; Trudy Sk. Bazy 8: 523 (1940)
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TKM TZK UZB
Synonyms: hidden
= Cousinia pseudaffinis var. bucharica Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., Ser. 8b, Bot 6: 11 (1929)
species :
Cousinia hamadanensis
Rech.f.; Fl. Iranica 139a: 142, t. 136 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia hamosa
C.A.Mey.; Prodr. [A. DC.] 6: 555 (1838)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
Synonyms: hidden
= Arctium hamosum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Cousinia buhsei Bunge; Mém. Acad. Imp. Sci. Saint Pétersbourg, Sér. 7, 9(2): 22 (1865)
= Cousinia hamulosa C.A.Mey.; Boiss., Fl. Orient. [Boiss. ] 3: 483 (1875)
species :
Cousinia harazensis
Rech.f.; Fl. Iranica 90: 259, t. 137 (1972)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia hastifolia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 571 (1887)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Cousinia taldykensis Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 299 (1937)
species :
Cousinia haussknechtii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 238 (1893)
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
Synonyms: hidden
= Cousinia serotina Rech.f.; Anz. Österr. Akad. Wiss. Math.-Naturwiss. Kl. 101: 349 (1964)
species :
Cousinia hazarensis
Mirtadz. & Attar; Willdenowia 34(1): 191 (2004)
Distribution: Iran
34 IRN
species :
Cousinia hedgei
Rech.f.; Fl. Iranica 90: 43 (1972)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia heliantha
Bunge; Mém. Acad. Sci. Petersb. 7 ser., 9 2: 39 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium helianthum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hemsleyana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 228 (1892)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia hergtiana
Bornm.; Beih. Bot. Centralbl. 32 2: 400 (1914)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia heteroloba
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ci. 347 (1964)
Distribution: Afghanistan (Baghlan, Bamyan)
34 AFG
species :
Cousinia heterophylla
Boiss.; Diagn. Ser. II. 3: 54 (1856)
Distribution: Afghanistan (Ghazni, Ghorat, Herat, Orozgan / Daykundi, Paktia / Khost, Zabul); Pakistan (Quetta, Baluchistan)
34 AFG 40 PAK
Synonyms: hidden
= Arctium heterophyllum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hilariae
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 22 (1929)
Distribution: Tajikistan
32 TZK
species :
Cousinia hohenackeri
Fisch. & C.A.Mey.; Ind. Sem. Hort. Petrop. 2: 34 (1835)
Distribution: Azerbaijan (Talysh); Iran
33 AZE 34 IRN
Synonyms: hidden
= Arctium hohenackeri (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hololeuca
Bunge; Mém. Acad. Sci. Petersb. 7 sér., 9 2: 27 (1865)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium hololeucum (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hoplites
Bornm.; Beih. Bot. Centralbl. 34: 11. 277 (1916)
Distribution: Afghanistan (Ghazni, Wardak)
34 AFG
species :
Cousinia hoplophylla
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 49 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia horrida
Kult.; Acta Univ. As. Med. Ser. 8b 7: 1, f. 1 (1929)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia humilis
Boiss.; Ann. Sci. Nat., ser. 4, 2: 248 (1854)
Distribution: Turkey (Inner Anatolia)
34 TUR
Synonyms: hidden
= Arctium humile (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia hymenostephanus
(Rech.f.) Podlech; Vasc. Pl. Afghanistan 162 (2013)
Distribution: Afghanistan (Ghorat)
34 AFG
Synonyms: hidden
= Tiarocarpus hymenostephanus Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 108: 7 (1971 publ. 1972)
species :
Cousinia hypochionea
Bornm.; Bull. Herb. Boiss., ser. 2, 7: 217, t. 7, f. 1 (1907)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia hypoleuca
Boiss.; Fl. Orient. 3: 464 (1875)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium hypoleucum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia assasinorum Bornm.; Bull. Herb. Boiss. Ser. 2(7): 210 (1907)
= Cousinia hypoleuca var. congesta Bornm.; (1917)
= Cousinia hypoleuca var. teheranica Parsa; (1948)
= Cousinia tafreshensis Rech.f.; Ann. Naturhist. Mus. Wien lvii. 73 (1950)
species :
Cousinia hypopolia
Bornm. & Sint.; Bull. Herb. Boiss., ser. 2, 7: 214, t. 7, f. 4 (1907)
Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
species :
Cousinia hystricocephala
Sennikov; Phytotaxa 25: 25 (2011)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia iconica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 52 (1972)
Distribution: Turkey (Central Anatolia, Inner Anatolia)
34 TUR
species :
Cousinia ilicifolia
Jaub. & Spach; Ill. Pl. Or. 2: 82 (1846)
Distribution: Iran (S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium ilicifolium (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia iljinii
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 187 (1937)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
species :
Cousinia immitans
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat. 103: 285 (1966)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia immitantiformis
Rech.f.; Fl. Iranica 90: 125 (1972)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia incompta
DC.; Prodr. [A. DC.] 6: 553 (1838)
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium incomptum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia incompta var. continua Assadi; Fl. Iran [ed. M. Assadi et al.] 125: 331 (2017)
species :
Cousinia inflata
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 512 (1875)
Distribution: Iraq (NE-Iraq, W-Iraq); Iran (W-Iran)
34 IRN IRQ
Synonyms: hidden
= Arctium inflatum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia straussii Hausskn. & C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 235 (1897)
species :
Cousinia infundibularis
Rech.f.; Fl. Iranica 90: 60 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia insignis
Rech.f.; Fl. Iranica 90: 106 (1972)
Distribution: Afghanistan (Orozgan / Daykundi)
34 AFG
species :
Cousinia integrifolia
Franch.; Ann. Sci. Nat. Ser. VI. 16: 322 (1883)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Arctium integrifolium (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia intermedia
DC.; Prodr. [A. DC.] 6: 553 (1838)
Distribution: Armenia; Azerbaijan (Nachichevan)
33 ARM AZE
Synonyms: hidden
= Arctium intermedium (C.A.Mey. ex DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891), nom. illeg.
species :
Cousinia intertexta
Freyn & Sint.; Oesterr. Bot. Zeitschr. 42 5: 169 (1892)
Distribution: Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, SE-Anatolia)
34 TUR
species :
Cousinia iranshahriana
Attar & Maroofi; Iranian J. Bot. 16(2): 197 (-199, fig. 1) (2010)
Distribution: Iran
34 IRN
species :
Cousinia iranshahrii
Rech.f.; Fl. Iranica 139a: 152, t. 148 (1979)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia irritans
Rech.f.; Anz. Österr. Akad. Wiss. Math.-Naturwiss. Kl. 57: 72 (1950)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
species :
Cousinia isfahanica
Assadi; Iranian J. Bot. 15(1): 37 (2009)
Distribution: Iran
34 IRN
species :
Cousinia jacobsii
Rech.f.; Fl. Iranica 90: 237, t. 119 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia jassyensis
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 472 (1887)
Distribution: Kyrgyzstan
32 KGZ
Synonyms: hidden
= Arctium jassyense (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia joharchii
Assadi & Mehregan; Phytotaxa 257(3): 276 (2016)
Distribution: Iran
34 IRN
species :
Cousinia juzepczukii
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 95 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia kadereitii
Mehregan & Assadi; Willdenowia 39(2): 268 (2010)
Distribution: Iran
34 IRN
species :
Cousinia kaluensis
Rech.f.; Fl. Iranica 90: 84 (1972)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia kandavanensis
Attar; Taxon. Biosyst. 3(6): 64 (2011)
Distribution: Iran
34 IRN
variant :
var. golydaghi
Parsa; Kew Bull. 3(2): 205 (1948)
Distribution: Iran
34 IRN
species :
Cousinia karkasensis
Mehregan & Djavadi; Iranian J. Bot. 16(2): 200 (-203, fig. 1, map) (2010)
Distribution: Iran
34 IRN
species :
Cousinia kasachstanica
Sennikov; Phytotaxa 5: 21 (2010)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia kataghanica
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 105: 6 (1969)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia kazachorum
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 96 (1962)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia keredjensis
Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 328, t. 227 (1934)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia kermanensis
Mirtadz. & Attar; Nordic J. Bot. 32(3): 279 (2014)
Distribution: Iran
34 IRN
species :
Cousinia kermanshahensis
Attar, Ghahr. & Assadi; Iranian J. Bot. 9(1): 56 (2001)
Distribution: Iran
34 IRN
species :
Cousinia kerstanii
Bornm.; Repert. Spec. Nov. Regni Veg. Beih. 108: 49 (1938)
Distribution: Afghanistan (Kunar / Nuristan); Pakistan (Chitral); NW-India (Jammu & Kashmir)
34 AFG 40 IND PAK
species :
Cousinia khashensis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 105 (1955)
Distribution: Afghanistan (Badakshan, Wakhan)
34 AFG
species :
Cousinia khorasanica
Djavadi & Attar; Rostaniha 7(Suppl. 2): 170 (2007)
Distribution: Iran
34 IRN
species :
Cousinia khorramabadensis
Bornm.; Köie, Danish Scient. Invest. Iran 4, Beitr. Sw. Iran 1: 25, f. 4 (1945)
Distribution: Iran (SW-Iran)
34 IRN
variant :
var. purpurea
Attar & Ghahr.; Iranian J. Bot. 9(2): 166 (2002)
Distribution: Iran
34 IRN
species :
Cousinia kilouyensis
Djavadi & Attar; Willdenowia 39(1): 89 (2009)
Distribution: Iran
34 IRN
species :
Cousinia kirrindica
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 141 (1940)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia lurorum var. recurvata Bornm.; (1936)
= Cousinia lurorum var. lancigera Bornm.; (1936)
species :
Cousinia knorringiae
Bornm.; Beih. Bot. Centralbl. 34: 11. 201 (1916)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia koelzii
Rech.f.; Biol. Skr. 8(2): 109 (1955)
Distribution: Afghanistan (Badakshan)
34 AFG
species :
Cousinia kokanica
Regel & Schmalh. ex Regel; Pl. Nov. Fedsch. 49 (1882)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Arctium kokanicum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia komaroffii
(Kuntze) C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 470 (1889)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
Synonyms: hidden
= Arctium komaroffii Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
species :
Cousinia komidjanensis
Mehregan; Iranian J. Bot. 17(2): 140 (2011)
Distribution: Iran
34 IRN
species :
Cousinia kopi-karadaghensis
Rech.f.; Fl. Iranica 90: 249, t. 130 (1972)
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
species :
Cousinia kornhuberi
Heimerl; Denkschr. Akad. Wiss. Math.-Naturwiss. Kl. 50: 62 (1885)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia medorum Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 40: 345 (1936)
species :
Cousinia korowiakowii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 226 (1897)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia kotschyi
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 120 (1846)
subspecies :
subsp. kotschyi
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Lappa kotschyi (Boiss.) Boiss.; Fl. Orient. 3: 458 (1875)
subspecies :
subsp. khansarica
(Attar & Ghahr.) Mehregan; Willdenowia 38(2): 340 (2008)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia khansarica Attar & Ghahr.; Nordic J. Bot. 23(5): 592 (2005)
species :
Cousinia kovalevskiana
Tscherneva; Novosti Sist. Vyssh. Rast. 26: 162 (1989)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia krauseana
Regel & Schmalh. ex Regel; Pl. Nov. Fedsch. 48 (1882)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
Synonyms: hidden
= Arctium krauseanum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia kuekenthalii
Bornm.; Beih. Bot. Centralbl. 34: 11. 146 (1916)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia kuramensis
Bornm.; Beih. Bot. Centralbl. 34: 11. 150 (1916)
Distribution: Afghanistan (Paktia / Khost); Pakistan (Kurram)
34 AFG 40 PAK
species :
Cousinia kurdistanica
Attar; Iranian J. Bot. 16(1): 169 (2010)
Distribution: Iran
34 IRN
species :
Cousinia lachnobasis
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 113 (1955)
Distribution: Afghanistan (Farah)
34 AFG
species :
Cousinia lachnopoda
Juz.; Trudy sk. Bazy 8: 525 (1940)
Distribution: Tajikistan
32 TZK
species :
Cousinia lachnosphaera
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 36 (1865)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts.); Afghanistan (Bamyan, Ghorat, Herat, Wardak, Orozgan / Daykundi)
34 AFG IRN
Synonyms: hidden
= Arctium lachnosphaerum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia sven-hediniana Bornm.; Beih. Bot. Centralbl. 34: II. 268 (1916)
variant :
var. kashanensis
(Rech.f. & Esfand.) Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 108 (1980)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Cousinia kashanensis Rech.f. & Esfand.; Ann. Naturhist. Mus. Wien, Ivii. 76 (1950)
species :
Cousinia lactiflora
Rech.f.; Fl. Iranica 139a: 143, t. 138 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia laetevirens
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 576 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
species :
Cousinia lanata
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 238 (1894)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
species :
Cousinia lancifolia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 572 (1887)
Distribution: Tajikistan
32 TZK
species :
Cousinia laniceps
(Bornm.) Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 316 (1937)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
Synonyms: hidden
= Cousinia schtschurowskiana var. laniceps Bornm.; Beih. Bot. Centralbl. 34(2): 187 (1916)
species :
Cousinia lasiandra
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 40 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium lasiandrum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia lasiolepis
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 118 (1846)
Distribution: Iran (EC-Iran, E-Iran, S-Iran, SW-Iran); Afghanistan (Bamyan, Ghorat)
34 AFG IRN
Synonyms: hidden
= Arctium lasiolepis (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia machaerophora C.Winkl. & Bornm.; (1897)
species :
Cousinia lasiophylla
C.Shih; Bull. Bot. Res., Harbin 4(2): 59-60 (1984)
Distribution: China (W-Xinjiang)
36 CHI
species :
Cousinia leiocephala
(Regel) Juz.; Not. Inst. Bot. Acad. Sci. U.R.S.S., ser. 1, 3: 314 (1936)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; China (Xinjiang)
32 KAZ KGZ UZB 36 CHI
Synonyms: hidden
= Cousinia sewertzowii var. leiocephala Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 6(2): 314 (1880)
species :
Cousinia lepida
Bunge; Boiss., Fl. Orient. 3: 476 (1875)
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium lepidum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia leptacma
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 35 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia leptocampyla
Bornm.; Beih. Bot. Centralbl. 34: 11. 179 (1916)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia leptocephala
Fisch. & C.A.Mey.; Bull. Soc. Imp. Naturalistes Moscou (1839) 159 ex Bunge
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Arctium leptocephalum (Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia leptoclada
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 15 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia leptocladoides
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 45 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia leptolepis
(Bornm. & Gauba) Rech.f.; Fl. Iranica 90: 1 1 1, t. 49 (1972)
Distribution: Iraq (NE-Iraq); Iran (EC-Iran)
34 IRN IRQ
Synonyms: hidden
= Cousinia adenosticta var. leptolepis Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 33 (1934)
species :
Cousinia leptomera
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ci. 348 (1964)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia leucantha
Bornm. & Sint; J. Russe de Bot., 1 (1911)
Distribution: Turkmenistan
32 TKM
Synonyms: hidden
= Cousinia simplex Kult.; (1960)
species :
Cousinia libanotica
DC.; Prodr. [A. DC.] 6: 556 (1838)
Distribution: Lebanon; Syria; Israel (coastal W-Israel); Jordania (S-Jordania)
34 ISR JOR LEB SYR
Synonyms: hidden
= Arctium hermonis (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium libanoticum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia dayi Post; Pl. Postianae 2: 13 (1891)
= Cousinia hermonis Boiss.; Diagn. Pl. Orient., ser. 1, 10: 102 (1849)
species :
Cousinia lignosissima
Rech.f.; Fl. Iranica 139a: 113, t. 106 (1979)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia linczewskii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., ser. 1, 3 (1936)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia litwinowiana
Bornm.; Bull. Herb. Boiss. Ser. II. 7: 215 (1907)
Distribution: Uzbekistan
32 UZB
species :
Cousinia lomakinii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 220 (1897)
Distribution: Armenia
33 ARM
species :
Cousinia longibracteata
Attar & Mirtadz.; Iranian J. Bot. 15(2): 149 (2009)
Distribution: Iran
34 IRN
species :
Cousinia longifolia
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 168, t. 8 (1897)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
species :
Cousinia lordeganensis
Mehregan; Willdenowia 38(2): 341 (2008)
Distribution: Iran
34 IRN
species :
Cousinia lucida
DC.; Prodr. [A. DC.] 6: 553 (1838)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium lucidum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia lucida var. arachnoidea Assadi & Mehregan; Fl. Iran [ed. M. Assadi et al.] 125: 132 (2017)
species :
Cousinia lurestanica
Attar & Djavadi; Iranian J. Bot. 16(1): 153 (2010)
Distribution: Iran
34 IRN
species :
Cousinia lurorum
Bornm.; Repert. Spec. Nov. Regni Veg. 40: 346 (1936)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia cymbolepis var. subintegriloba Bornm.; (1914)
species :
Cousinia lyrata
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 53 (1865)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.); NW-Afghanistan
32 TKM 34 AFG IRN
Synonyms: hidden
= Arctium lyratum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia maassoumii
Assadi; Iranian J. Bot. 15(1): 40 (2009)
Distribution: Iran
34 IRN
species :
Cousinia macrocephala
C.A.Mey.; Verz. Pfl. Caucasus 231 (1831)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Iran
33 ARM AZE 34 IRN
Synonyms: hidden
= Arctium macrocephalum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia gabrieljaniae Takht. & Tamanian; Bot. Zhurn. (Moscow & Leningrad) 73(11): 1609 (1988)
= Cousinia takhtajanii Tamanian; Fl. Rast. Rast. Res. Armenii (Sborn. Nauch. Trud. 13): 17 (1991)
species :
Cousinia macrolepis
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 505 (1875)
Distribution: Iraq (NE-Iraq)
34 IRQ
Synonyms: hidden
= Arctium macrolepis (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia gigantosphaera Rech.f.; Anz. österr. Akad. Wiss. Math.-Naturwiss. Kl. 101: 345 (1964)
species :
Cousinia macroptera
C.A.Mey.; Prodr. [A. DC.] 6: 555 (1838)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Turkey (E-Anatolia); Iran (Iranian Aserbaijan, W-Iran)
33 ARM AZE 34 IRN TUR
Synonyms: hidden
= Arctium macropterum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium obovatum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia carlinoides Boiss. & Buhse; Nouv. Mém. Soc. Imp. Naturalistes Moscou 12: 126 (1860)
= Cousinia compacta Turrill; Bull. Misc. Inform. Kew 1929(7): 229 (1929)
= Cousinia macroptera var. obovata (Boiss.) Winkl.; (1893)
= Cousinia obovata Boiss.; Fl. Orient. 3: 492 (1875)
species :
Cousinia magnifica
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 319 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia malacophylla
Rech.f.; Fl. Iranica 90: 176 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia malurensis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 119 (1955)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia manouchehrii
Rech.f. & Esfand.; Ann. Naturh. Mus. Wien 57: 79 (1950)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia maracandica
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 30 (1932)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia margaritae
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 5 (1929)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia margiana
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 303 (1937)
Distribution: Turkmenistan
32 TKM
species :
Cousinia masulehensis
Attar & Rastegar; Phytotaxa 343(3): 231 (2018)
Distribution: Iran
34 IRN
species :
Cousinia mattfeldii
Bornm.; Repert. Spec. Nov. Regni Veg. Beih. 108: 38 (1938)
Distribution: Pakistan (Chitral); NW-India (Jammu & Kashmir)
40 IND PAK
species :
Cousinia megalomastix
Rech.f.; Fl. Iranica 90: 316 (1972)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia meghrica
Takht.; Trans. Armen. Branch Acad. Sci. USSR, Biol. Ser. 2: 186 (1937)
Distribution: Armenia; Azerbaijan (Nachichevna)
33 ARM AZE
species :
Cousinia mehreganii
Assadi; Iranian J. Bot. 17(1): 8 (2011)
Distribution: Iran
34 IRN
species :
Cousinia meshhedensis
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 144, t. 312, f. 1 (1940)
Distribution: Iran (EC-Iran, NE-Iran: Mts., N-Iran)
34 IRN
Synonyms: hidden
= Cousinia strictissima Rech.f.; Ann. Naturhist. Mus. Wien 57: 80 (1950)
species :
Cousinia microcarpa
Boiss.; Diagn. Pl. Orient., ser. 2, 3: 59 (1856)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Iran (E-Iran, NE-Iran: Mts.); Afghanistan (Badakshan, Badghys, Baghlan, Balkh, Bamyan, Farah, Faryab, Ghorat, Herat, Jawzjan / Sar-e-Pol, Kabul, Orozgan / Daykundi, Paktia / Khost, Parwan, Qunduz, Samangan, Takhar); Pakistan (Waziristan, Kurram, Pakistani Punjab); NW-India (Jammu & Kashmir)
32 KAZ KGZ TKM TZK UZB 34 AFG IRN 40 IND PAK
Synonyms: hidden
= Arctium microcarpum (Boiss.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Cousinia acanthoptera Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 72 (1955)
= Cousinia hystrix C.B.Clarke; Compos. Ind. 214 (1876)
= Cousinia xeranthemoides Bunge; Boiss., Fl. Orient. 3: 497 (1865)
species :
Cousinia microcephala
C.A.Mey.; Prodr. [A. DC.] 6: 552 (1838)
Distribution: Armenia; Azerbaijan (incl. Nachichevan); Iran (Iranian Aserbaijan)
33 ARM AZE 34 IRN
Synonyms: hidden
= Arctium microcephalum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia grossheimii Bornm.; Oesterr. Bot. Z. 75: 235 (1926)
= Cousinia microcephala var. glabrescens Turrill; (1930)
species :
Cousinia millefontana
Rech.f.; Fl. Iranica 90: 248 (1972)
Distribution: Iran (Iranian Aserbaijan, W-Iran)
34 IRN
species :
Cousinia mindshelkensis
B.Fedtsch.; Repert. Spec. Nov. Regni Veg. 10: 164 (1911)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia minkwitziae
Bornm.; Beih. Bot. Centralbl. 34: 11. 158 (1916)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia minuticapitata
Attar; Taxon. Biosyst. 3(6): 66 (2011)
Distribution: Iran
34 IRN
species :
Cousinia miserabilis
Rech.f.; Fl. Iranica 90: 171 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia moabitica
Bornm. & Nábelek; Bornm., Österr. Bot. Zeitschr. 63: 60 (1913)
Distribution: Jordania (S-Jordania)
34 JOR
species :
Cousinia mobayenii
Ghahr. & Attar; Iranian J. Bot. 8(2): 260 (2000)
Distribution: Iran
34 IRN
species :
Cousinia mogensii
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 107 (1955)
Distribution: Afghanistan (Bamyan, Ghazni)
34 AFG
species :
Cousinia mogoltavica
Tscherneva & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 55 (1961)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia mollis
Schrenk; Bull. Phys-Math. Acad. Petersb. 3: 108 (1845)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Afghanistan (Samangan)
32 KAZ KGZ TKM TZK UZB 34 AFG
Synonyms: hidden
= Arctium molle (Schrenk) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia monocephala
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 52 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium monocephalum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia mozaffarianii
Attar, Assadi & Ghahr.; Pakistan J. Bot. 32(2): 293 (2000)
Distribution: Iran
34 IRN
species :
Cousinia mozdouranensis
Djavadi & Attar; Feddes Repert. 116(5-6): 287 (2005), nom. inval., et Phytotaxa 227(1): 99 (2015)
Distribution: Iran
34 IRN
species :
Cousinia mucida
Kult. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 97 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia mulgediifolia
Bornm.; Beih. Bot. Centralbl. 34: 11. 139 (1916)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia multiloba
DC.; Prodr. [A. DC.] 6: 554 (1838)
Distribution: Turkmenistan; Tajikistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran, S-Iran, W-Iran); Afghanistan (Badghys, Bamyan, Ghazni, Ghorat, Jawzjan / Sar-e-Pol, Kabul, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan); Pakistan (Kurram, Baluchistan, Chitral)
32 TKM TZK 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium multilobum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia elegans Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 172 (1882)
= Cousinia gnaphalodes Bornm.; Repert. Spec. Nov. Regni Veg. 46: 281 (1939)
= Cousinia multiloba var. afghanorum Bornm.
= Cousinia palmatiloba Jaub. & Spach; Illustr. 2: 80. t. 162 (1846)
species :
Cousinia murgabica
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 97 (1962)
Distribution: Turkmenistan
32 TKM
species :
Cousinia mutehensis
Rech.f.; Fl. Iranica 139a: 129, t. 122 (1979)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia myrioglochis
Rech.f.; Fl. Iranica 90: 57 (1972)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia myriolepis
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 124 (1955)
Distribution: Afghanistan (Logar, Paktia / Khost, Parwan)
34 AFG
species :
Cousinia myriotoma
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 126 (1955)
Distribution: Iran (NE-Iran: Mts.); Afghanistan (Farah, Herat)
34 AFG IRN
species :
Cousinia nabelekii
Bornm.; Oesterr. Bot. Zeitschr. 63: 59, t. 1, f. 5, 6 (1913)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
species :
Cousinia nana
Attar, Ghahr. & Assadi; Nordic J. Bot. 20(6): 698 (2000 publ. 2001)
Distribution: Iran
34 IRN
species :
Cousinia neglecta
Juz.; Not. Syst. Herb. Hort. Petrop. 4: 78 (1923)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia neubaueri
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 128 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
Synonyms: hidden
= Tiarocarpus neubaueri (Rech.f.) Rech.f.; Anz. Österr. Akad. Wiss., Mat.-Naturwiss., 108: 6 (1971 publ. 1972)
species :
Cousinia neurocentra
Bunge; Mém. Acad. Sci. Pétersb. 7 ser., 9 2: 29 (1865)
Distribution: Iran (EC-Iran, E-Iran)
34 IRN
Synonyms: hidden
= Arctium neurocentrum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia newesskyana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 333 (1896)
Distribution: Uzbekistan
32 UZB
species :
Cousinia ninae
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 315 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia niveocoronata
C.Jeffrey ex Grey-Wilson; Kew Bull. 29(1): 50 (1974)
Distribution: Afghanistan (Balkh)
34 AFG
species :
Cousinia noeana
Boiss.; Diagn. Pl. Orient., ser. 2, 3: 57 (1856)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium noeanum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia novissima
Rech.f.; Fl. Iranica 90: 310 (1972)
Distribution: Afghanistan (Ghazni, Ghorat)
34 AFG
species :
Cousinia nujianensis
Attar, Ghahr., Saber & Zarre; Iranian J. Bot. 11(1): 66 (2005)
Distribution: Iran
34 IRN
species :
Cousinia odontolepis
DC.; Prodr. [A. DC.] 6: 556 (1838)
subspecies :
subsp. odontolepis
Distribution: Iraq (NE-Iraq, NW-Iraq); Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN IRQ
Synonyms: hidden
= Arctium cymbolepis (Boiss.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium odontolepis (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia acanthophysa Rech.f.; Fl. Iranica 90: 227, t. 115 (1972)
= Cousinia baueri Bornm. & Nábelek ex Bornm.; Oesterr. Bot. Z. 63: 61 (1913)
= Cousinia cymbolepis Boiss.; Diagn. Pl. Orient., ser. 1, 6: 119 (1846)
= Cousinia leatherdalei Rech.f.; Oesterr. Bot. Zeitschr. 100: 462, f. 3-4 (1953)
= Cousinia mazu-shirinensis Rech.f.; Fl. Iranica 90: 226, t. 114 (1972)
= Cousinia odontolepis var. leiolepis Bornm.; Repert. Spec. Nov. Regni Veg. 46: 282 (1939)
= Cousinia schultziana Jaub. & Spach; Illustr. 2: 95. t. 177 (1846)
subspecies :
subsp. kurdica
(C.Winkl. & Bornm.) Mehregan; Willdenowia 38(2): 315 (2008)
Distribution: Iraq (NE-Iraq)
34 IRQ
Synonyms: hidden
= Cousinia carduchorum C.Winkl. & Bornm.; Bull. Herb. Boiss., ser. 1, 3: 568, t. 15 (1895)
= Cousinia carduchorum var. major C.Winkl. & Bornm.; Bull. Herb. Boissier 3: 569 (1895)
= Cousinia carduchorum var. minor C.Winkl. & Bornm.; Bull. Herb. Boissier 3: 568 (1895), nom. inval.
= Cousinia kurdica C.Winkl. & Bornm.; Bull. Herb. Boiss. 3: 567, t. 13 (1895)
= Cousinia qandilica Rech.f.; Fl. Iranica 90: 223 (1972)
species :
Cousinia olgae
Regel & Schmalh.; Regel, Descr. Pl. Nov. Fedtsch., Izv. Obsh. Ljub. Estv. Antrop. Etnogr. 34 2: 48 (1882)
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Iran (E-Iran); Afghanistan (Badghys, Baghlan, Balkh, Faryab, Herat, Jawzjan / Sar-e-Pol)
32 KGZ TKM TZK UZB 34 AFG IRN
species :
Cousinia oligocephala
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 116 (1846)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium oligocephalum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia olivieri
DC.; Prodr [A. DC.] 6: 552 (1838)
Distribution: ?Iran
Synonyms: hidden
= Arctium olivieri (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia pyrrhocephala Jaub. & Spach; Ill. Pl. Orient. 2(17-18): 85 (1846)
species :
Cousinia omissa
Tscherneva; Novosti Sist. Vyssh. Rast. 30: 155 (1996)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia omphalodes
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 98 (1962)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia onopordioides
Ledeb.; Eichw., Pl. Nov. It. Caspio-Caucasus 40 (1833)
Distribution: Kazakhstan; Turkmenistan; Northern Caucasus; Azerbaijan (incl. Talysh); Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran); Afghanistan (Ghazni, Logar, Paktia / Khost, Parwan); Pakistan (Baluchistan)
32 KAZ TKM 33 AZE NCS 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium albicaule (Boiss. & Buhse) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium grandiceps (Bunge) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium karelinii (Less. ex Kar.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium polyacanthum (Eichw.) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Cousinia albicaulis Boiss. & Buhse; Nouv. Mém. Soc. Nat. Moscou 12 18: 128, t. 37 (1860)
= Cousinia bornmuelleri C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 235 (1897)
= Cousinia grandiceps Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 54 (1865)
= Cousinia karelinii Less. ex Kar.; Bull. Soc. Imp. Naturalistes Moscou 13: 159 (1839)
= Cousinia onopordioides var. glabra Assadi & Mehregan; Fl. Iran [ed. M. Assadi et al.] 125: 224 (2017)
= Cousinia subzevarensis Rech.f.; Fl. Iranica 139a: 134 (1979)
= Onopordum polyacanthum Eichw.; Pl. Casp.-Caucas.: 3 (1831), nom. nud.
species :
Cousinia onopordon
Freyn & Sint.; Oesterr. Bot. Zeitschr. 42: 207 (1892)
Distribution: ?Transcaucasus
species :
Cousinia oophora
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 133 (1955)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia oopoda
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 310 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia oreodoxa
Bornm. & Sint.; Mitt. Thür. Bot. Ver. 23: 99 (1908)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia oreoxerophila
Kult.; Fl. Turkm. 7: 380 (1960)
Distribution: Turkmenistan
32 TKM
species :
Cousinia orientalis
(Adams) K.Koch; Linnaea 24: 386 (1851)
Distribution: Northern Caucasus; Georgia [Caucasus]; Azerbaijan
33 AZE GEO NCS
Synonyms: hidden
= Arctium carduiforme (Cass.) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium orientale (Adams) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Carduus orientalis Adams; Weber & Mohr, Beitr. 1: 63 (1805)
= Cousinia carduiformis Cass.; Dict. Sci. Nat. 47: 503 (1827)
= Cousinia orientalis (Adams) Grossh.; Grossh. & Schischk., Sched. Pl. Or. Exsicc. Fasc. X. 14 (1928)
= Cousinia orientalis (Adams) O.Hoffm.; Engl. et Prantl, Natürl. Pflanzenfam. 4: 5 319 (1890)
species :
Cousinia orthacantha
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 98 (1962)
Distribution: Uzbekistan; Tajikistan; Afghanistan (Balkh, Qunduz, Samangan)
32 TZK UZB 34 AFG
species :
Cousinia orthoclada
Hausskn. & Bornm.; Beih. Bot. Centralbl. 20 2: 159 (1906)
Distribution: Iran (S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia gracilis var. erecto-patula Parsa; (1948)
= Cousinia orthoclada var. subappendiculata Bornm.; (1914)
= Cousinia orthoclada var. subrecurvata Bornm.; (1917)
species :
Cousinia ortholepis
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 99 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia orthoneura
Rech.f.; Fl. Iranica 90: 120 (1972)
Distribution: Afghanistan (Laghman, Parwan)
34 AFG
species :
Cousinia oshtorankuhensis
Attar; Iranian J. Bot. 17(2): 155 (2011)
Distribution: Iran
34 IRN
species :
Cousinia ottonis
Bornm.; Oesterr. Bot. Zeitschr. 62: 106, t. 3, f. 484a (1912)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia outichaschensis
Franch.; Ann. Sci. Nat. Ser. VI. 16: 322 (1883)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Arctium outichaschense (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia commixta Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., Ser. 8b, Bot. 6: 18 (1929)
= Cousinia outichaschensis f. commixta (Kult.) Kamelin; Fl. Ushchel. Reki Varzob 343 (2021)
= Cousinia outichaschensis f. glauciifolia (Kult.) Kamelin; Fl. Ushchel. Reki Varzob 343 (2021)
species :
Cousinia ovczinnikovii
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 99 (1962)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia oxiana
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 100 (1962)
Distribution: Turkmenistan; Uzbekistan; Tajikistan; Iran (NE-Iran: Mts.); Afghanistan (Badghys, Balkh, Farah, Faryab)
32 TKM TZK UZB 34 AFG IRN
species :
Cousinia oxytoma
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 135 (1955)
Distribution: Tajikistan (Petr I Mts.); Afghanistan (Badakshan, Baghlan, Kunar / Nuristan); Pakistan (Chitral)
32 TZK 34 AFG 40 PAK
Synonyms: hidden
= Cousinia batalinii var. virescens Bornm.; Beih. Bot. Centralbl. 34(2): 168 (1916)
species :
Cousinia panjshirensis
Rech.f.; Fl. Iranica 90: 128 (1972)
Distribution: Afghanistan (Parwan)
34 AFG
species :
Cousinia pannosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 578 (1887)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Cousinia resupinata Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 318 (1937)
species :
Cousinia pannosiformis
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 101 (1962)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
species :
Cousinia papillosa
Djavadi & Attar; Rostaniha 8(2): 64 (2007)
Distribution: Iran
34 IRN
species :
Cousinia parjumanensis
Rech.f.; Fl. Iranica 90: 101 (1972)
Distribution: Afghanistan (Farah, Ghorat)
34 AFG
species :
Cousinia parsana
Ghahr., Iranshahr & Attar; Iranian J. Bot. 8(1): 16 (1999)
Distribution: Iran
34 IRN
species :
Cousinia parviceps
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 136 (1955)
Distribution: Afghanistan (Badghys, Bamyan, Faryab, Ghazni, Ghorat, Herat, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan)
34 AFG
species :
Cousinia parviziana
Parsa; Flore de l´Iran (Publ. Univ. Teheran 1777) 10: 110 (1980)
Distribution: Iran
34 IRN
species :
Cousinia pasargardensis
Attar; Iranian J. Bot. 16(1): 139 (2010)
Distribution: Iran
34 IRN
species :
Cousinia patentispina
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 101 (1962)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan
32 KAZ TKM UZB
species :
Cousinia pauciramosa
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 139 (1924)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia pectinata
Rech.f.; Ann. Naturhist. Mus. Wien 57: 76 (1950)
Distribution: Afghanistan (Kabul)
34 AFG
Synonyms: hidden
= Cousinia gillii Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 99 (1955)
species :
Cousinia peduncularis
Juz. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 101 (1962)
Distribution: Tajikistan
32 TZK
species :
Cousinia pergamacea
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 513 (1875)
subspecies :
subsp. pergamacea
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
Synonyms: hidden
= Arctium pergamaceum (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia wheeler-hainesii Rech.f.; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 101: 353 (1964)
subspecies :
subsp. sardashtensis
(Rech.f.) Mehregan; Willdenowia 38(2): 331 (2008)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Cousinia sardashtensis Rech.f.; Fl. Iranica 139a: 139, t. 132 (1979)
species :
Cousinia perovskiensis
(Bornm.) Juz. ex Tscherneva; Kom., Fl. U.R.S.S. 27: 161 (1962)
Distribution: Kazakhstan
32 KAZ
Synonyms: hidden
= Cousinia dissecta var. perovskiensis Bornm.
species :
Cousinia persarum
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 227 (1893)
Distribution: Iran (S-Iran)
34 IRN
species :
Cousinia persica
Djavadi & Attar; Feddes Repert. 117(7-8): 455 (2006)
Distribution: Iran
34 IRN
species :
Cousinia perspolitana
Attar & Ghahr.; Nordic J. Bot. 23(5): 589 (2005)
Distribution: Iran
34 IRN
species :
Cousinia pestalozzae
Boiss.; Fl. Orient. 3: 471 (1875)
Distribution: Lebanon (Antilebanon); Syria (C-Syrian Desert)
34 LEB SYR
Synonyms: hidden
= Arctium pestalozzae (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia phyllocephala
Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 36: 330, t. 181 (1934)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia koieana Bornm.; Danish Sci. Invest. Iran 4: 26 (1945)
= Cousinia koeieana var. adenoloba Bornm.; (1945)
species :
Cousinia pichleriana
Bornm.; Repert. Spec. Nov. Regni Veg. 46: 282 (1939), in adnot., et ex Rech.f., Fl. Iranica 139a: 141, t. 135 (1979)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia ecbatanensis f. inermis Bornm.
species :
Cousinia pinarocephala
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 121 (1846)
Distribution: Iran (EC-Iran, N-Iran, Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Arctium pinarocephalum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia declinata Parsa; Kew Bull. 3(2): 204 (1948)
= Cousinia pinarocephala var. totschalensis Bornm.; (1907)
species :
Cousinia pineticola
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 138 (1955)
Distribution: Afghanistan (Baghlan, Ghazni, Laghman, Orozgan / Daykundi, Paktia / Khost, Parwan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia piptocephala
Bunge; Mém. Acad. Sci. Petersb. 7 sér., 9 2: 26 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts., S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium piptocephalum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia platyacantha
Bunge; Mém. Acad. Sci. Petersb. 7 sér., 9 2: 37 (1865)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia centauroides Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 152
species :
Cousinia platylepis
Schrenk ex Fisch. & C.A.Mey.; Index Sem. (St. Petersburg) 9: 10 (1843)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; China (NW-Xinjiang)
32 KAZ KGZ TZK UZB 36 CHI
Synonyms: hidden
= Arctium platylepis (Schrenk ex Fisch. & C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia platyptera
Bornm. & C.Winkl.; Oesterr. Bot. Zeitschr. 62: 181, t. 3, f. 1 (1912)
Distribution: Iran (S-Iran)
34 IRN
species :
Cousinia platyraphis
Kult.; Acta Univ. As. Med., ser. 8b 7: 1 (1929)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia platystegia
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 103 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia podlechii
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 286 (1966)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia podophylla
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 43 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia polycephala
Rupr.; Mem. Acad. Imp. Sci. Saint Petersbourg (Ser. 7) 14(4): 54 (1869)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; China (Xinjiang)
32 KAZ KGZ TKM TZK UZB 36 CHI
Synonyms: hidden
= Arctium polycephalum (Rupr.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia polyneura
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 140 (1955)
Distribution: Afghanistan (Kabul)
34 AFG
species :
Cousinia polytimetica
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 103 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia porphyrochrysea
Rech.f.; Fl. Iranica 90: 278 (1972)
Distribution: Afghanistan (Badakshan, Takhar)
34 AFG
species :
Cousinia porphyrostephana
Rech.f.; Fl. Iranica 90: 266 (1972)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia postiana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 223 (1892)
Distribution: Lebanon (Antilebanon); Syria (C-Syrian Desert); Jordania (S-Jordania)
34 JOR LEB SYR
species :
Cousinia praestans
Tscherneva & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 57 (1961)
Distribution: Uzbekistan
32 UZB
species :
Cousinia prasina
Jaub. & Spach; Ill. Pl. Or. 2: 86, t. 168 (1846)
Distribution: Iran (EC-Iran, W-Iran)
34 IRN
Synonyms: hidden
= Arctium prasinum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia princeps
Franch.; Ann. Sci. Nat. Ser. VI. 16: 324 (1888)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Arctium princeps (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia prolifera
Jaub. & Spach; Ill. Pl. Or. 2: 75, t. 157 (1845)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; Iraq (NW-Iraq); Iran (EC-Iran, E-Iran, NE-Iran: Mts., Iranian Aserbaijan, S-Iran); Afghanistan (Balkh, Farah, Helmand, Herat, Kabul, Kandahar, Kunar / Nuristan, Nangarhar, Paktia / Khost, Parwan); Pakistan (Baluchistan, Waziristan, Kurram, Pakistani Punjab, Peshawar, Jhelum, Rawalpindi, Swat, Hazara); NW-India (Jammu & Kashmir, Rajasthan); Myanmar
32 KAZ TKM TZK UZB 34 AFG IRN IRQ 40 IND PAK 41 MYA
Synonyms: hidden
= Arctium minutum (Boiss.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Arctium proliferum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Cousinia aralensis Bunge; Mem. Sav. Etr. Petersb. 7: 354 (1851), et Beitr. Fl. Russl.: 178 (1852)
= Cousinia calcitrapiformis Jaub. & Spach; Illustr. 2: 76. t. 158 (1845)
= Cousinia minuta Boiss.; Diagn. Pl. Orient., ser. 1, 6: 117 (1846)
species :
Cousinia proxima
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 301 (1937)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia psammophila
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 15 (1929)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia pseudactinia
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 142 (1955)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia pseudaffinis
Kult.; Trans. Sci. Soc. Turk. 1: 27 (1923), et in Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 3, 105 (1928)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan
32 KAZ KGZ UZB
species :
Cousinia pseudobonvalotii
Juz.; Trudy sk. Bazy 8: 525 (1940)
Distribution: ?Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia pseudocandolleana
Assadi; Iranian J. Bot. 15(1): 42 (2009)
Distribution: Iran
34 IRN
species :
Cousinia pseudocirsium
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 144 (1955)
Distribution: Afghanistan (Badakshan)
34 AFG
species :
Cousinia pseudodshizakensis
Tschern. & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 59 (1961)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia pseudolanata
Popov ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 104 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia pseudomollis
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 470 (1887)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan (Pamir-Alai, Tianshan); Tajikistan; Afghanistan (Badakshan, Baghlan, Balkh, Samangan, Takhar)
32 KAZ KGZ TZK UZB 34 AFG
Synonyms: hidden
= Arctium pseudomolle (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia pseudomollis var. platyacantha Bornm.; (1916)
species :
Cousinia pseudostenolepis
Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 147 (1955)
Distribution: Afghanistan (Badakshan, Baghlan, Bamyan, Orozgan / Daykundi, Samangan, Takhar)
34 AFG
species :
Cousinia pterocarpa
Boiss.; Fl. Orient. 3: 501 (1875)
Distribution: Afghanistan (Ghazni, Wardak)
34 AFG
Synonyms: hidden
= Arctium pterocarpum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pterocaulos
(C.A.Mey.) Rech.f.; Fl. Iranica 90: 95 (1972)
Distribution: Azerbaijan (Talysh); Iran (N-Iran, Iranian Aserbaijan, W-Iran)
33 AZE 34 IRN
Synonyms: hidden
= Arctium hystrix (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Carthamus pterocaulos C.A.Mey.; Verz. Pfl. Caucasus 67 (1831)
= Cousinia hystrix C.A.Mey.; Ind. Sem. Horti Petrop. 2: 34 (1835)
species :
Cousinia pugionifera
Jaub. & Spach; Ill. Pl. Or. 2: 92, t. 174 (1846)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium pugioniferum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pulchella
Bunge; Del. Sem. Hort. Dorpat. (1845) 3
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan (Pamir-Alai); Afghanistan (Badakshan, Baghlan, Orozgan / Daykundi)
32 KGZ TZK UZB 34 AFG
Synonyms: hidden
= Arctium pulchellum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pulcherantha
Attar & Mirtadz.; Nordic J. Bot. 32(3): 281 (2014)
Distribution: Iran
34 IRN
species :
Cousinia pulchra
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 478 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
Synonyms: hidden
= Arctium pulchrum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pulvinaris
Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 147 (1955)
Distribution: Afghanistan (Farah)
34 AFG
species :
Cousinia pungens
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 19 (1932), et Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 306, 308 (1937)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
species :
Cousinia purpurea
C.A.Mey.; Prodr. [A. DC.] 6: 555 (1838)
Distribution: Northern Caucasus; Armenia; Azerbaijan (incl. Nachichevan); Iran (Iranian Aserbaijan)
33 ARM AZE NCS 34 IRN
Synonyms: hidden
= Arctium purpureum (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pusilla
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 468 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Arctium pusillum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia pycnocephala
Rech.f.; Fl. Iranica 90: 195 (1972)
Distribution: Tajikistan; Afghanistan (Faryab)
32 TZK 34 AFG
species :
Cousinia pycnoloba
Boiss.; Diagn. Ser. II. 3: 57 (1856)
Distribution: Afghanistan (Ghazni, Wardak, Orozgan / Daykundi, Paktia / Khost); Pakistan (Baluchistan, Kurram)
34 AFG 40 PAK
Synonyms: hidden
= Cousinia scala Aitch. & Hemsl.; J. Linn. Soc., Bot. 19: 172 (1882)
species :
Cousinia pygmaea
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 467 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan; Afghanistan (Baghlan, Ghazni)
32 TKM TZK UZB 34 AFG
Synonyms: hidden
= Arctium pygmaeum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia qaisarensis
Rech.f.; Fl. Iranica 90: 261 (1972)
Distribution: Afghanistan (Faryab)
34 AFG
species :
Cousinia qaradaghensis
Rech.f.; Fl. Iranica 90: 258 (1972)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
species :
Cousinia qarehbilensis
Rech.f.; Fl. Iranica 139a: 131, t. 123 (1979)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia quettensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 287 (1966)
Distribution: S-Afghanistan; Pakistan
34 AFG 40 PAK
species :
Cousinia racemosa
Boiss.; Diagn. Ser. II. 3: 56 (1856)
Distribution: Afghanistan (Bamyan, Ghorat, Kabul, Wardak, Parwan); Pakistan (Chitral, Drosh)
34 AFG 40 PAK
Synonyms: hidden
= Arctium racemosum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia raddeana
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: (1887) 473 in obs., et 11: (1889) 132 t. 1
Distribution: Turkmenistan; Tajikistan (S-Pamir-Alai); Afghanistan (Badghys, Balkh, Faryab, Samangan)
32 TKM TZK 34 AFG
Synonyms: hidden
= Arctium raddeanum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia nobilis Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 132 (1955)
species :
Cousinia radians
Bunge; Del. Sem. Hort. Dorpat. (1845) 3
Distribution: Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TKM TZK UZB
Synonyms: hidden
= Arctium radians (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium stenolepis (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia interrupta C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 229 (1892)
= Cousinia stenolepis Regel & Schmalh.; Izv. Imp. Obshch. Lyubit. Estestv. Moskovsk. Univ. 34(2): 49 (1882)
species :
Cousinia ramosissima
DC.; Predr. 6: 552 (1838)
Distribution: Turkey (Inner Anatolia, S-Anatolia); Lebanon (Antilebanon, C-Lebanon); Syria (C-Syrian Desert, NW-Syria, Jbel Druze)
34 LEB SYR TUR
Synonyms: hidden
= Arctium ramosissimum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia drusorum Bornm. & Sam.; Repert. Spec. Nov. Regni Veg. 40: 342 (1936)
species :
Cousinia ramulosa
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 149 (1955)
Distribution: Tajikistan (Pamir); Afghanistan (Badakshan, Kunar / Nuristan, Parwan)
32 TZK 34 AFG
species :
Cousinia raphiostegia
Rech.f.; Fl. Iranica 139a: 128, t. 120 (1979)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia rava
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 227 (1895)
Distribution: Tajikistan
32 TZK
species :
Cousinia rawanduzensis
Mehregan; Iranian J. Bot. 17(2): 144 (2011)
Distribution: Iraq
34 IRQ
species :
Cousinia rechingerae
Bornm.; Repert. Spec. Nov. Regni Veg. 48: 133, t. 309, f. 1 (1940)
Distribution: Iran (NE-Iran: Mts., N-Iran)
34 IRN
Synonyms: hidden
= Cousinia ahmed-adlii Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 49: 263 (1940)
= Cousinia kandewanensis var. golydaghi Parsa; (1948)
species :
Cousinia rechingerorum
Bornm.; Repert. Spec. Nov. Regni Veg. 48: 145, t. 313, f. 1 (1940)
Distribution: Iran (E-Iran)
34 IRN
Synonyms: hidden
= Cousinia malekiana Parsa; Fl. Iran 3: 539 (1949)
species :
Cousinia recurvata
DC.; Prodr. [A. DC.] 6: 555 (1838)
Distribution: Iran (NE-Iran: Mts., SW-Iran)
34 IRN
Synonyms: hidden
= Arctium recurvatum (DC.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia regelii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 229 (1895)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KAZ KGZ TZK UZB
species :
Cousinia renominata
Rech.f.; Fl. Iranica 90: 163 (1972)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Cousinia turkmenorum Bornm. & Gauba; (1939) (nec Bornm. 1916
species :
Cousinia resinosa
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 24 (1932), et Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KAZ KGZ TKM TZK UZB
species :
Cousinia rhabdodes
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 137, t. 310, f. 1 (1940)
Distribution: Iran (EC-Iran)
34 IRN
species :
Cousinia rhaphiocephala
Rech.f.; Fl. Iranica 90: 319 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia rhodantha
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 22 (1929)
Distribution: Uzbekistan
32 UZB
species :
Cousinia rhombiformis
C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 232 (1897)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia rigida
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 26 (1929)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia rigidissima
Rech.f.; Fl. Iranica 90: 200 (1972)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia robusta
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 151 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia rosea
Kult.; Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 3, 105 (1928), et Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 17 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia rotundifolia
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 579 (1887)
Distribution: Turkmenistan; Uzbekistan; Tajikistan
32 TKM TZK UZB
Synonyms: hidden
= Cousinia kultiasowii Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 317 (1937)
= Cousinia rubrarenosa Kult.; Trudy Sredne-Aziatsk. Gosud. Univ., Ser. 8b, Bot. 6: 18 (1929)
species :
Cousinia rubiginosa
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 23 (1929)
Distribution: Tajikistan
32 TZK
species :
Cousinia rudis
Rech.f.; Fl. Iranica 90: 42 (1972)
Distribution: Afghanistan (Balkh)
34 AFG
species :
Cousinia rufidula
Bornm.; Beih. Bot. Centralbl. 34: II. 270 (1916)
Distribution: Afghanistan (Ghorat, Herat)
34 AFG
species :
Cousinia sabalanica
Attar, Ghahr. & Assadi; Nordic J. Bot. 20(6): 699 (2000 publ. 2001)
Distribution: Iran
34 IRN
species :
Cousinia sabzevarensis
Rech.f.; Fl. Iranica 139a: 134, t. 126 (1979)
Distribution: Iran (NE-Iran: Mts., N-Iran)
34 IRN
species :
Cousinia sagittata
C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 233 (1897)
subspecies :
subsp. sagittata
Distribution: Iran (SW-Iran)
34 IRN
subspecies :
subsp. iranica
(C.Winkl. & Strauss) Mehregan; Willdenowia 38(2): 328 (2008)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia iranica C.Winkl. & Strauss; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 233 (1897)
species :
Cousinia sahandica
Attar & Djavadi; Iranian J. Bot. 16(1): 161 (2010)
Distribution: Iran
34 IRN
species :
Cousinia sakawensis
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 489 (1875)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Arctium sakawense (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia salangensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ciii. 288 (1966)
Distribution: Afghanistan (Baghlan, Parwan)
34 AFG
species :
Cousinia saloukensis
Mehregan; Willdenowia 41(2): 263 (2011)
Distribution: Iran
34 IRN
species :
Cousinia sanandajensis
Rech.f.; Fl. Iranica 90.235, t. 118 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia sarawschanica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 424 (1886)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia sarzehensis
Attar, Ghahr. & Assadi; Nordic J. Bot. 20(6): 697 (2000 publ. 2001)
Distribution: Iran
34 IRN
species :
Cousinia scabrida
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia scariosa
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 154
Distribution: Afghanistan (Ghorat, Jawzjan / Sar-e-Pol, Wardak); Pakistan (Chitral)
34 AFG 40 PAK
Synonyms: hidden
= Arctium scariosum (Regel) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia lamprophyllaria Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 114 (1955)
species :
Cousinia scheibeana
Bornm.; Repert. Spec. Nov. Regni Veg. Beih. 108: 43 (1938)
Distribution: Afghanistan (Badakshan, Bamyan, Kunar / Nuristan, Parwan); Pakistan (Chitral); NW-India (Jammu & Kashmir)
34 AFG 40 IND PAK
species :
Cousinia schepsaica
Karmysch.; Fl. Kazakhst. 9: 244, 569 (1966)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia schindleriana
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 124, t. 11, f. 2 (1941)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia schiraziana
Attar; Iranian J. Bot. 16(1): 139 (2010)
Distribution: Iran
34 IRN
species :
Cousinia schischkinii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 316 (1937)
Distribution: Kyrgyzstan (Tianshan)
32 KGZ
species :
Cousinia schistoptera
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 301 (1937)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan
32 KAZ TKM UZB
species :
Cousinia schistosa
Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 154 (1955)
Distribution: Afghanistan (Kunar / Nuristan)
34 AFG
species :
Cousinia schtschurowskiana
Regel & Schmalh. ex Regel; Pl. Nov. Fedsch. 50 (1882)
Distribution: Uzbekistan
32 UZB
Synonyms: hidden
= Arctium schtschurowskianum (Regel & Schmalh.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia scleracantha
Kult. ex Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 104 (1962)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
species :
Cousinia sclerolepis
C.Shih; Bull. Bot. Res., Harbin 4(2): 60-61, photo. 1 (1984)
Distribution: China (W-Xinjiang)
36 CHI
species :
Cousinia sclerophylla
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 308 (1937)
Distribution: Tajikistan
32 TZK
species :
Cousinia sefidiana
(Pau) Rech.f.; An. Jard. Bot. Madrid, 47(2): 365 (1989 publ. (1990)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia cymbolepis var. sefidiana Pau; Trab. Mus. Nac. Ci. Nat., Ser. Bot. 14: 40 (1918)
species :
Cousinia seidlitzii
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 20 (1865)
Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN
Synonyms: hidden
= Arctium seidlitzii (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia semilacera
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 322 (1937)
Distribution: Tajikistan (Pamir-Alai); Afghanistan (Badakshan)
32 TZK 34 AFG
Synonyms: hidden
= Cousinia jacea Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 104 (1955)
species :
Cousinia semnanensis
Attar, Mirtadz. & Sotoodeh; Nova Biol. Reperta 9(20: 147 (2022)
Distribution: Iran
34 IRN
species :
Cousinia serratuloides
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 118 (1846)
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium serratulodes (Boiss.) Kuntze; Revis. Gen. Pl. 1: 367 (1891)
= Cousinia olivieri Jaub. & Spach; Illustr. t. 172 (1846) (non DC. 1838)
species :
Cousinia sewerzovii
Regel; Bull. Soc. Imp. Naturalistes Moscou 40: (1867) II. 151 (sewerzowii)
Distribution: Kazakhstan; Uzbekistan; Tajikistan; Afghanistan (Parwan)
32 KAZ TZK UZB 34 AFG
Synonyms: hidden
= Arctium sewerzovii (Regel) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia shahrestanica
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., cv. 7 (1969)
Distribution: Afghanistan (Orozgan / Daykundi)
34 AFG
species :
Cousinia shahuensis
Attar; Taxon. Biosyst. 3(6): 61 (2011)
Distribution: Iran
34 IRN
species :
Cousinia shahvarica
Rech.f.; Ann. Naturh. Mus. Wien 57: 71 (1950)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
species :
Cousinia sharifii
Attar & Amini Rad; Phytotaxa 395(4): 287 (2019)
Distribution: Iran
34 IRN
species :
Cousinia shebliensis
Ghahr., Iranshahr & Attar; Iranian J. Bot. 8(1): 19 (1999)
Distribution: Iran
34 IRN
species :
Cousinia sheidaii
Attar, Ghahr. & Mahdigholi; Novon 17(1): 5 (2007)
Distribution: Iran
34 IRN
species :
Cousinia shibarensis
Rech.f.; Fl. Iranica 90: 271 (1972)
Distribution: Afghanistan (Parwan)
34 AFG
species :
Cousinia shorlughensis
Rech.f.; Fl. Iranica 139a: 116 (1979)
Distribution: E-Afghanistan
34 AFG
species :
Cousinia shugnanica
Juz.; Trudy Tadz. Bazy Akad. Nauk 8: 525, 564 (1940)
Distribution: Tajikistan; Afghanistan (Badakshan, Wakhan, Kunar / Nuristan, Takhar)
32 TZK 34 AFG
Synonyms: hidden
= Cousinia lindbergii Rech.f.; Ann. Naturhist. Mus. Wien 65: 36 (1962)
species :
Cousinia shulabadensis
Attar & Ghahr.; Iranian J. Bot. 9(2): 162 (2002)
Distribution: Iran
34 IRN
species :
Cousinia sicigera
C.Winkl. & Bornm.; Bull. Herb. Boiss. 5: 166, t. 7 (1897)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
species :
Cousinia silvanica
Attar; Iranian J. Bot. 16(1): 153 (2010)
Distribution: Iran
34 IRN
species :
Cousinia silyboides
Jaub. & Spach; Illustr. 2: 94. t. 176 (1846)
subspecies :
subsp. silyboides
Distribution: Iran
34 IRN
Synonyms: hidden
= Arctium silybodes (Jaub. & Spach) Kuntze; Revis. Gen. (1891) 307 et 308
subspecies :
subsp. disfulensis
(Bornm.) Mehregan; Willdenowia 38(2): 311 (2008)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Cousinia disfulensis Bornm.; Köie, Danish Scient. Invest. Iran 4, Beitr. Sw. Iran 1: 23, f. 3 (1945)
subspecies :
subsp. zardkuhensis
(Attar & Ghahr.) Mehregan; Willdenowia 38(2): 310 (2008)
Distribution: Iran
34 IRN
Synonyms: hidden
= Cousinia zardkuhensis Attar & Ghahr.; Iranian J. Bot. 9(2): 164 (2002)
species :
Cousinia simulatrix
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 573 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia singularis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ci. 350 (1964)
Distribution: Afghanistan (Ghazni)
34 AFG
species :
Cousinia sintenisii
Freyn; Oesterr. Bot. Zeitschr. 42 6: 205 (1892)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
species :
Cousinia sivasica
Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 32: 53 (1972)
Distribution: Turkey (Inner Anatolia)
34 TUR
species :
Cousinia smirnowii
Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 8: 488 (1883)
Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium smirnowii (Trautv.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia fulgens Bornm. & Rchb.f.; Repert. Spec. Nov. Regni Veg. 48: 139 (1940)
= Cousinia globosa (Kult.) Bornm.; (1940)
= Cousinia russeola Juz.; Trudy Bot. Inst. Akad. Nauk S. S. S. R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 319 (1937)
= Cousinia smirnowii subsp. sintenisiana Bornm.; (1940)
= Cousinia smirnowii subsp. trautvetteriana Bornm.; (1940)
= Cousinia smirnowii var. globosa Kult.; (1929)
species :
Cousinia sogdiana
Bornm.; Beih. Bot. Centralbl. 34: II. 154 (1916)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia sororia
Juz.; Trudy sk. Bazy 8: 525 (1940)
Distribution: Tajikistan
32 TZK
species :
Cousinia spathulata
Kult.; Acta Univ. As. Med., ser. 8b 7: 3, f. 2 (1929)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia speciosa
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 579 (1887)
Distribution: Kyrgyzstan; Tajikistan
32 KGZ TZK
species :
Cousinia sphaerocephala
Jaub. & Spach; Ill. Pl. Or. 2: 79 (1845)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Arctium sphaerocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
species :
Cousinia spiridonovii
Juz.; Trans. Rubb. & Guttap. Inst., Fasc. 4, 12 (1932)
Distribution: Kazakhstan; Uzbekistan; Tajikistan
32 KAZ TZK UZB
species :
Cousinia splendida
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 13: 240 (1894)
Distribution: Tajikistan
32 TZK
Synonyms: hidden
= Cousinia lipskyi C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 14: 237 (1895)
species :
Cousinia sporadocephala
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 307 (1937)
Distribution: Uzbekistan; Kyrgyzstan
32 KGZ UZB
species :
Cousinia spryginii
Kult.; Korovin, Kult. & Popov, Descr. Pl. Nov. Turkmenistan. (Bodenk. u. Pflanzengeogr. Beob. u. Unters. 1: Flussgeb. d. Amu Darya u. Syr Darya, II.), 78 (1916)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia stahliana
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 117, t. 13 (1941)
Distribution: Turkmenistan; Iran (NE-Iran: Mts.)
32 TKM 34 IRN
species :
Cousinia stapfiana
Freyn & Sint.; Oesterr. Bot. Zeitschr. 44: 218 (1894)
Distribution: Turkey (Inner Anatolia, N-Anatolia)
34 TUR
Synonyms: hidden
= Cousinia stapfiana var. ankarensis Wagenitz; Willdenowia 3 2: 276 (1962)
species :
Cousinia stechmanniopsis
Rech.f.; Fl. Iranica 90: 52 (1972)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia stellaris
Bornm.; Beih. Bot. Centralbl. 34: II. 161 (1916)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
species :
Cousinia stenocalathia
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 156 (1955)
Distribution: Afghanistan (Badakshan, Baghlan, Takhar)
34 AFG
species :
Cousinia stenocephala
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 115 (1846)
Distribution: Turkey (SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); Iraq (NE-Iraq, NW-Iraq); Iran (W-Iran); Syria (Jazira)
34 IRN IRQ SYR TUR
Synonyms: hidden
= Arctium involucratum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Arctium stenocephalum (Boiss.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia involucrata Boiss.; Diagn. 1 6: 114 (1846)
species :
Cousinia stenophylla
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 7 (1929)
Distribution: Turkmenistan
32 TKM
species :
Cousinia stephanophora
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 575 (1887)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia stereolepis
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 158 (1955)
Distribution: Afghanistan (Ghorat, Jawzjan / Sar-e-Pol)
34 AFG
species :
Cousinia stereoneura
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 159 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia stocksii
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 12: 216 (1893)
Distribution: Iran (S-Iran, SW-Iran); Afghanistan (Farah, Helmand, Kandahar, Orozgan / Daykundi); Pakistan (Baluchistan)
34 AFG IRN 40 PAK
species :
Cousinia stricta
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 60 (1961)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia strobilocephala
Tscherneva & Vved.; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 16: 62 (1961)
Distribution: Uzbekistan
32 UZB
species :
Cousinia stroterolepis
Rech.f.; Fl. Iranica 90: 250 (1972)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia subappendiculata
Kult.; Not. Syst. Herb. Hort. Petrop. 5: 140 (1924), et Not. Syst. Herb. Hort. Bot. Russia 6: 27 (1926)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia subcandicans
Tscherneva; Not. Syst. Herb. Inst. Bot. Acad. Sci. . 15: 41 (1959)
Distribution: Uzbekistan; Tajikistan
32 TZK UZB
species :
Cousinia subinflata
Bornm.; Oesterr. Bot. Zeitschr. 63: 291 (1913)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia submutica
Franch.; Ann. Sci. Nat. Ser. VI. 16: 314 (1883)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Arctium submuticum (Franch.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia subpectinata
Mirtadz., Attar & Assadi; Iranian J. Bot. 10(2): 144 (2004)
Distribution: Iran
34 IRN
species :
Cousinia subscaposa
Rech.f.; Fl. Iranica 90: 270 (1972)
Distribution: Afghanistan (Badakshan, Baghlan, Takhar); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia sylvicola
Bunge; Del. Sem. Hort. Dorp. (1845) 3
Distribution: Uzbekistan
32 UZB
species :
Cousinia tabrisiana
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 16 (1865)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
Synonyms: hidden
= Arctium tabrisianum (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia takharensis
Rech.f.; Fl. Iranica 90: 116 (1972)
Distribution: Afghanistan (Wakhan Kunar / Nuristan, Takhar)
34 AFG
species :
Cousinia talassica
(Kult.) Juz. ex Tscherneva; Komolrov, Fl. U.R.S.S. 27: 154 (1962)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
Synonyms: hidden
= Cousinia affinis var. talassica Kult.
species :
Cousinia tamarae
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 297 (1937)
Distribution: Kyrgyzstan
32 KGZ
Synonyms: hidden
= Cousinia krauseana var. macrocephala C.Winkl.
species :
Cousinia tashkurghanica
Rech.f.; Fl. Iranica 90: 42 (1972)
Distribution: Afghanistan (Samangan)
34 AFG
species :
Cousinia taybadensis
Djavadi & Attar; Rostaniha 7(Suppl. 2): 168 (2007)
Distribution: Iran
34 IRN
species :
Cousinia tedshenica
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 105 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia tenella
Fisch. & C.A.Mey.; Ind. Sem. Horti Petrop. 1: 25 (1834), et Linnaea 10: Litt.-Ber. 86 (1836)
Distribution: Germany [I]; Latvia [I]; Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Armenia; Azerbaijan (incl. Nachichevan); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan); Afghanistan (Badghys, Baghlan, Faryab, Qunduz, Samangan); Pakistan (Baluchistan)
11 ger 14 lat 32 KAZ KGZ TKM TZK UZB 33 ARM AZE 34 AFG IRN 40 PAK
Synonyms: hidden
= Arctium tenellum (Fisch. & C.A.Mey.) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Lappa tenella (Fisch. & C.A.Mey.) DC.; Prodr [A. DC.] 6: 662 (1838)
species :
Cousinia tenuifolia
C.A.Mey.; Prodr. [A. DC.] 6: 554 (1838)
Distribution: Armenia; Iran (Iranian Aserbaijan)
33 ARM 34 IRN
Synonyms: hidden
= Arctium tenuifolium (C.A.Mey.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia tenuiramula
Rech.f.; Fl. Iranica 139a: 124, t. 113 (1979)
Distribution: Iran (SW-Iran)
34 IRN
species :
Cousinia tenuisecta
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia tenuispina
Rech.f.; Fl. Iranica 90: 83 (1972), nom. nov.
Distribution: Afghanistan (Bamyan, Ghazni, Ghorat, Herat, Wardak)
34 AFG
Synonyms: hidden
= Cousinia leptacantha Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 116 (1955)
species :
Cousinia termei
Rech.f.; Fl. Iranica 139a: 150, t. 146 (1979)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia tetanocephala
Bornm. & Gauba; Mitt. Thür. Bot. Ver. 47: 119, t. 14 (1941)
Distribution: Iran (E-Iran)
34 IRN
species :
Cousinia thamnodes
Boiss. & Hausskn.; Boiss., Fl. Orient. 3: 471 (1875)
Distribution: Iran (S-Iran, SW-Iran)
34 IRN
Synonyms: hidden
= Arctium thamnodes (Boiss. & Hausskn.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia thomsonii
C.B.Clarke; Compos. Ind. 213 (1876)
Distribution: Afghanistan (Kunar / Nuristan, Laghman); S-Tibet; Pakistan (Swat, Dir, Chitral, Gilgit, Astor); Nepal; N-India (Dras, Ladakh, Jammu & Kashmir, Zanskar, Himachal Pradesh, Uttarakhand)
34 AFG 36 TIB 40 IND NEP PAK
Synonyms: hidden
= Arctium thomsonii (C.B.Clarke) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia brandisii C.B.Clarke; Compos. Ind. 214 (1876)
= Cousinia elasoonensis Rech.f. & Edelb.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 91 (1955)
= Cousinia xanthophoenicea Bornm.; Repert. Spec. Nov. Regni Veg. Beih. 108: 46 (1938)
species :
Cousinia tianschanica
Kult.; Bull. Inst. Pedol. & Geobot. Univ. As. Centr. 1: 113 (1925)
Distribution: Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan
32 KAZ KGZ TZK UZB
species :
Cousinia tirinensis
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., cv. 8 (1969)
Distribution: Afghanistan (Kandahar, Orozgan / Daykundi)
34 AFG
species :
Cousinia touchalensis
Attar; Taxon. Biosyst. 3(6): 65 (2011)
Distribution: Iran
34 IRN
species :
Cousinia trachylepis
Bunge; Mém. Acad. Sci. Pétersb. 7 sér., 9 2: 22 (1865)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium trachylepis (Bunge) Kuntze; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 197 (1887)
= Cousinia masandarana Bornm.; Oesterr. Bot. Z. 75: 232 (1926)
= Cousinia meyameyensis Parsa; Kew Bull. 3(2): 206 (1948)
= Cousinia masanderana var. mallophora Bornm. & Gauba; (1944)
species :
Cousinia trachyphylla
Juz.; Trudy sk. Bazy 8: 524 (1940)
Distribution: Tajikistan
32 TZK
species :
Cousinia trachyphyllaria
Bornm. & Rech.f.; Repert. Spec. Nov. Regni Veg. 48: 143, t. 312, f. 2 (1940)
Distribution: Iran (NE-Iran: Mts.)
34 IRN
species :
Cousinia tragacanthoides
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 161 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
Synonyms: hidden
= Tiarocarpus tragacanthoides (Rech.f. & Gilli) Rech.f.; Anz. Österr. Akad. Wiss., Mat. Naturwiss., 108: 6 (1971 publ. 1972)
species :
Cousinia transiliensis
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 298 (1937)
Distribution: Kazakhstan; Kyrgyzstan
32 KAZ KGZ
species :
Cousinia transoxana
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 105 (1962)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia triceps
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 8 (1929)
Distribution: Turkmenistan; Uzbekistan
32 TKM UZB
species :
Cousinia trichophora
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 26 (1929)
Distribution: Uzbekistan
32 UZB
species :
Cousinia tricolor
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ci. 351 (1964)
Distribution: Afghanistan (Ghorat)
34 AFG
species :
Cousinia trollii
Bornm.; Repert. Spec. Nov. Regni Veg. Beih. 108: 45 (1938)
Distribution: Afghanistan (Kunar / Nuristan); Pakistan (Chitral)
34 AFG 40 PAK
species :
Cousinia tscherneviae
Berdyev; Novit. Syst. Pl. Vasc., Akad. Sci. U.R.S.S. 6: 200 (1970)
Distribution: Turkmenistan
32 TKM
species :
Cousinia turcomanica
C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 11: 133 (1889)
Distribution: Turkmenistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Synonyms: hidden
= Arctium turcomanicum (C.Winkl.) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia turcomanica var. leiophylla Bornm. & Sint.; (1912)
= Serratula microcephala Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 458 (1886)
species :
Cousinia turkestanica
(Regel) Juz.
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan
32 KAZ KGZ TKM UZB
Synonyms: hidden
= Cousinia decurrens var. turkestanica Regel
= Cousinia syrdarjensis Kult.; Acta Univ. As. Med. Ser. VIII b, Bot. Fasc. 3, 106 (1928)
species :
Cousinia turkmenorum
Bornm.; Beih. Bot. Centralbl. 34 2: 153 (1917)
Distribution: Turkmenistan; Iran (EC-Iran, E-Iran, NE-Iran: Mts., N-Iran); Afghanistan (Farah, Herat)
32 TKM 34 AFG IRN
Synonyms: hidden
= Cousinia dissecta var. microcephala Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 456 (1886)
= Cousinia dissecta var. simplicior Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 456 (1886)
= Cousinia simplicior Juz.; Not. Syst. Herb. Hort. Petrop. 4: 77 (1923), in adnot.
= Cousinia stenoptera Juz.; Not. Syst. Herb. Hort. Petrop. 2: 59 (1921)
species :
Cousinia ulotoma
Bornm.; Beih. Bot. Centralbl. 34: II. 148 (1916)
Distribution: Tajikistan
32 TZK
species :
Cousinia umbilicata
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 306 (1937)
Distribution: Kazakhstan; Uzbekistan
32 KAZ UZB
species :
Cousinia unaiensis
Rech.f.; Fl. Iranica 90: 136 (1972)
Distribution: Afghanistan (Ghazni, Wardak)
34 AFG
species :
Cousinia urumiensis
Bornm.; Verh. K. K. Zool.-Bot. Ges. Wien 60: 135 (1910)
Distribution: Turkey (E-Anatolia); Iran (Iranian Aserbaijan)
34 IRN TUR
Synonyms: hidden
= Cousinia azerbaijanensis Parsa; Kew Bull. 3(2): 202 (1948)
= Cousinia azerbaijanensis var. elongata Parsa; (1948)
species :
Cousinia verbascifolia
Bunge; Mém. Acad. Sci. Pétersb. 7 s&., 9 2: 52 (1865)
Distribution: Iran (EC-Iran, NE-Iran: Mts.)
34 IRN
Synonyms: hidden
= Arctium verbascifolium (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
species :
Cousinia verticillaris
Bunge; Mem. Sav. Etr. Petersb. 6: 357 (1851), et Beitr. Fl. Russl.: 181 (1852)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Arctium verticillare (Bunge) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia hissarica C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 93 (1887)
= Cousinia poliothrix C.Winkl.; Trudy Imp. S.-Peterburgsk. Bot. Sada 10: 577 (1887)
species :
Cousinia vicaria
Kult.; Trans. Sci. Soc. Turkmenistan. 1: 113 (1923), et in Bull. Univ. As. Centr. 7: Suppl. 32 (1924)
Distribution: Kazakhstan; Uzbekistan
32 KAZ UZB
species :
Cousinia volkii
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Nat., ci. 352 (1964)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia vvedenskyi
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 106 (1962)
Distribution: Uzbekistan
32 UZB
species :
Cousinia waldheimiana
Bornm.; Beih. Bot. Centralbl. 34: II. 140 (1916)
Distribution: Kyrgyzstan
32 KGZ
species :
Cousinia wendelboi
Rech.f.; Fl. Iranica 90: 167, t. 75 (1972)
Distribution: Iran (N-Iran)
34 IRN
species :
Cousinia wesheni
Post; Pl. Postianae 2: 13 (1891)
Distribution: Turkey (SE-Anatolia: Mesopotamian Anatolia); Iraq (NE-Iraq, NW-Iraq); Syria (C-Syrian Desert, Jazira)
34 IRQ SYR TUR
Synonyms: hidden
= Cousinia chaborasica Bornm. & Hand.-Mazz.; Oesterr. Bot. Z. 62: 183 (1912)
species :
Cousinia wilhelminae
Rech.f.; Fl. Iranica 90: 142 (1972)
Distribution: Iran (Iranian Aserbaijan)
34 IRN
species :
Cousinia winkleriana
Aitch. & Hemsl.; Trans. Linn. Soc. Ser. II. 3: 79 (1886)
Distribution: Afghanistan (Herat)
34 AFG
species :
Cousinia woronowii
Bornm.; Oesterr. Bot. Zeitschr. 63: 290 (1913)
Distribution: Turkey (E-Anatolia, NE-Anatolia)
34 TUR
species :
Cousinia xanthacantha
Regel; Bull. Soc. Imp. Naturalistes Moscou 11 2: 156 (1867)
Distribution: Afghanistan (Baghlan, Bamyan, Kabul, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan)
34 AFG
Synonyms: hidden
= Arctium xanthacanthum (Regel) Kuntze; Revis. Gen. Pl. 1: 308 (1891)
= Cousinia neurophylla Rech.f. & Köie; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 130 (1955)
species :
Cousinia xanthina
Bornm.; Beih. Bot. Centralbl. 34: II. 190 (1916)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Cousinia curviloba Juz.; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 3: 321 (1937)
species :
Cousinia xanthiocephala
Tscherneva; Bot. Mater. Gerb. Inst. Bot. Akad. Nauk Uzb. SSR 17: 107 (1962)
Distribution: Kazakhstan
32 KAZ
species :
Cousinia xanthothyrsa
Rech.f. & Gilli; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 165 (1955)
Distribution: Afghanistan (Ghazni, Kabul, Logar, Wardak)
34 AFG
species :
Cousinia xanthula
Rech.f.; K. Danske Vid. Selsk., Biol. Skrift. 8: No. 2 (Symb. Afgh. II.) 167 (1955)
Distribution: Afghanistan (Bamyan)
34 AFG
species :
Cousinia xiphiolepis
Boiss.; Diagn. Pl. Orient., ser. 1, 6: 121 (1846)
Distribution: Iran (EC-Iran)
34 IRN
Synonyms: hidden
= Arctium actinocephalum (Jaub. & Spach) Kuntze; Revis. Gen. Pl. 1: 307 (1891)
= Arctium xiphiolepis (Boiss.) Kuntze; Revis. Gen. (1891)
= Cousinia actinocephala Jaub. & Spach; Illustr. 2: 93. t. 175 (1846)
species :
Cousinia yasujensis
Attar; Iranian J. Bot. 17(2): 153 (2011)
Distribution: Iran
34 IRN
species :
Cousinia zagrica
Attar, Ghahr. & Assadi; Sendtnera 8: 5 (2002)
Distribution: Iran
34 IRN
species :
Cousinia ×androssovii
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 305 (1937)
Distribution: Turkmenistan; Tajikistan; Afghanistan (Badakshan, Baghlan, Takhar)
32 TKM TZK 34 AFG
species :
Cousinia ×atripurpurea
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 313 (1937)
Distribution: Tajikistan
32 TZK
Remarks: Cousinia coronata Franch. x Cousinia microcarpa Boiss.
species :
Cousinia ×cana
Juz.; Acta Inst. Bot. Acad. Sci. U.R.S.S., Ser. I. Fasc. 3, 316 (1937)
Distribution: Kazakhstan
32 KAZ
Remarks: Cousinia ninae Juz. x Cousinia sewerzowii Regel
species :
Cousinia ×heterogenetos
Bornm.; Beih. Bot. Centralbl. 34: 11. 163 (1916)
Distribution: Tajikistan
32 TZK
Remarks: Cousinia outichaschensis Franch. x Cousinia pulchella Bunge
species :
Cousinia ×heteromorpha
Bornm.
Distribution: C-Asia
Remarks: Cousinia alpestris (Bornm.) Juz. x Cousinia submutica Franch.
species :
Cousinia ×hybrida
Rech.f.; Ann. Naturh. Mus. Wien 57: 83 (1950)
Distribution: Iran (N-Iran)
34 IRN
Remarks: Cousinia crispa x sphaerocephala
species :
Cousinia ×iskanderi
Bornm.; Beih. Bot. Centralbl. 34: 11. 168 (1916)
Distribution: Tajikistan
32 TZK
Remarks: Cousinia outichaschensis Franch. x Cousinia submutica Franch.
species :
Cousinia ×kamarbandensis
Rech.f.; Ann. Naturh. Mus. Wien 57: 82 (1950)
Distribution: Iran (N-Iran)
34 IRN
Remarks: Cousinia crispa x gmelini
species :
Cousinia ×kurubasgecidiensis
Ilçim & Özçelik; Biol. Diversity Conservation 6(1): 71 (2013)
Distribution: Turkey
34 TUR
species :
Cousinia ×mesomorpha
Rech.f.; Candollea 31: 87, f. 1 (1976)
Distribution: Iran (EC-Iran)
34 IRN
Remarks: Cousinia behboudiana x belangeri
species :
Cousinia ×paraatripurpurea
Kamelin; Bot. Zhurn. (Moscow & Leningrad) 58(5): 632 (1973), without latin descr.
Distribution: C-Asia
Remarks: Cousinia microcarpa Boiss. x Cousinia radians Bunge
species :
Cousinia ×triacantha
Kult.; Acta Univ. As. Med. Ser. VIII. Bot. Fasc. VI. 2 (1929)
Distribution: Turkmenistan
32 TKM
Remarks: Cousinia alberti x umbrosa
