131.0000 Family Orchidaceae Juss.; Gen. Pl.: 64 (1789), nom. cons.

over 800 genera, more than 30.000 spp. worldwide, mostly tropical

Species listed in database: 30483; hybrids: 1409

131.4327 Genus Dactylorhiza Neck. ex Nevski; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 4: 332 (1937), nom. cons.

Distribution: Atlantic Islands, Europe, European Russia, few to Russian Far East and Alaska. The species concept in this genus is highly controversial and fluctuating, depending on author, leading to countless synonyms. Recent molecular studies point to relatively few highly variable species. Coeloglossum is phylogenetically embedded in Dactylorhiza.

Species listed in database: 54; hybrids: 78

species :

Dactylorhiza aristata (Fisch. ex Lindl.) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

Name: Fischer´s Orchid

Distribution: Russian Far East (Kamchatka, Khabarovsk, Kuril Isl., Sakhalin); China (Hebei, Henan, Shandong, Shanxi); North Korea; Japan; Aleutes; SW-Alaska

31 RFE 36 CHI 38 JAP NKO 70 ASK

Remarks: sambucina agg.

Synonyms: hidden

= Dactylorchis aristata (Fisch.) Verm.; Stud. Dactylorch. 64 (1947)

= Dactylorhiza aristata f. alba P.M.Br.; North Amer. Native Orchid J. 1(1): 10 (1995)

= Dactylorhiza aristata f. albomaculata P.M.Br.; N. Amer. Native Orchid J. 10: 36 (2004 publ. 2005)

= Dactylorhiza aristata f. perbracteata (Lepage) Catling; Naturaliste Canad. 109: 278 (1982)

= Dactylorhiza aristata f. perbracteata (Lepage) P.M.Br.; North Amer. Native Orchid J. 2(1): 68 (1996), isonym

= Dactylorhiza aristata f. rosea P.M.Br.; North Amer. Native Orchid J. 1(3): 199 (1995)

= Dactylorhiza aristata var. kodiakensis Luer & G.M.Luer; Amer. Orchid Soc. Bull. 41: 207 (1972)

= Dactylorhiza aristata var. perbracteata (Lepage) Catling; Nat. Canad. 109(2): 278 (1982)

= Orchis aristata f. perbracteata Lepage; Amer. Midl. Naturalist 46: 757 (1952)

= Orchis aristata f. punctata Tatew.; Trans. Sapporo Nat. Hist. Soc. 9: 10 (1927)

= Orchis aristata Fisch. ex Lindl.; Gen. Sp. Orchid. Pl.: 262 (1835)

= Orchis aristata var. albiflora Koidz.; Bot. Mag. (Tokyo) 31: 34 (1917)

= Orchis aristata var. immaculata Makino; J. Jap. Bot. 8: 17 (1932)

= Orchis aristata var. maculata Makino; J. Jap. Bot. 8: 6 (1932)

= Orchis beeringiana (Cham.) Kudô; J. Coll. Agric. Hokkaido Imp. Univ. 11: 94 (1922)

= Orchis latifolia var. beeringiana Cham.; Linnaea 3: 26 (1828)

species :

Dactylorhiza armeniaca Hedrén; Pl. Syst. Evol. 229(1-2): 42 (2001)

Distribution: Northern Caucasus; Georgia [Caucasus]; Azerbaijan; NE-Turkey

33 AZE GEO NCS 34 TUR

Synonyms: hidden

= Dactylorhiza euxina subsp. armeniaca (Hedrén) Kreutz; Kompend. Eur. Orchid. 37 (2004)

species :

Dactylorhiza baumanniana J.Hölz. & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(4): 514 (1983)

subspecies :

subsp. baumanniana

Name: Baumanns Knabenkraut

Distribution: Albania; Bulgaria; NW-Greece

13 ALB BUL GRC

Remarks: traunsteineri agg. Included by Pedersen et al. 2019 in D. majalis subsp. pindica, which seems wrong

subspecies :

subsp. smolikana (B.Willing & E.Willing) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 607 (2005)

Distribution: N-Greece

13 GRC

Remarks: traunsteineri agg. Included by Pedersen et al. 2019 in D. majalis subsp. pindica, which seems wrong

Synonyms: hidden

= Dactylorhiza smolikana B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 6(1): 33 (1989)

species :

Dactylorhiza bucovina Kreutz, Bobocea & Bradeanu; Ber. Arbeitskreis. Heimische Orchid. 38(2): 101 (2021)

Distribution: Romania (Bucovina, Maramures)

13 ROM

Remarks: provisionally accepted name, status not confirmed genetically

species :

Dactylorhiza cantabrica H.A.Pedersen; Bot. J. Linn. Soc. 152(4): 428 (2006)

Distribution: Spain (Galicia)

12 SPA

Remarks: rank disputed

Synonyms: hidden

= Dactylorhiza sambucina subsp. cantabrica (H.A.Pedersen) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 155 (2007)

species :

Dactylorhiza cordigera (Fr.) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

subspecies :

subsp. cordigera

Name: Herz-Knabenkraut

Distribution: Croatia; Bosnia & Hercegovina; Serbia; Kosovo; ?North Macedonia; Albania; Romania; Bulgaria; Greece; European Russia

13 ALB BIH BUL CRO GRC KOS ROM SRB 14 RUS

Remarks: majalis agg.

Synonyms: hidden

= Dactylorchis cordigera (Fr.) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza cordigera f. albiflora Landwehr; Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza cordigera var. grisebachii (Pant.) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza cordigera var. rhodopeia Presser; J. Eur. Orch. 36(4): 982 (2004)

= Dactylorhiza cordigera var. rochelii (Griseb. & Schenk) Soó; Nom. Nov. Gen. Dactylorhiza: 5 (1962)

= Dactylorhiza cordigera var. vermionica B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 4(1): 36 (1987)

= Dactylorhiza majalis subsp. cordigera (Fr.) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 82 (1976) [without basionym page]

= Dactylorhiza majalis subsp. cordigera (Fr.) H.Sund.; Europ. Medit. Orchideen 45 (1975)

= Dactylorhiza richteri B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 263 (2012)

= Orchis cordigera lusus immaculata Klinge; Rev. Orch. cordigera, Orchis angustifolia: 19 (1893)

= Orchis cordigera proles rochelii (Griseb. & Schenk) Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 742 (1907)

= Orchis cordigera f. banatica Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 741 (1907)

= Orchis cordigera f. grisebachii (Pant.) Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 910 (1926)

= Orchis cordigera f. macrobracteata (Schur) Soó; Bot. Arch. 23: 72 (1928)

= Orchis cordigera f. rivularis (Heuff.) Pauca; Fl. Republ. Social. Roman. 12: 702 (1972)

= Orchis cordigera f. rochelii (Griseb. & Schenk) Klinge; Rev. Orch. cordigera, Orchis angustifolia: 23 (1893)

= Orchis cordigera f. simonkaiana Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis cordigera f. subtriloba Soó; Bot. Arch. 23: 71 (1928)

= Orchis cordigera Fr.; Novit. Fl. Succ. Mant. 3: 130 (1845)

= Orchis cordigera var. foliosa Klinge; Rev. Orch. cordigera, Orchis angustifolia 37 (1893)

= Orchis cordigera var. grisebachii (Pant.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 33 (1899)

= Orchis cordigera var. klingei E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 174 (1908)

= Orchis cordigera var. macrobracteata Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis cordigera var. rocheliana Klinge; Rev. Orch. cordigera, Orchis angustifolia 20 (1893)

= Orchis cruenta Rochel; Pl. Banat. Rar. 34, t. 1, f. 1 (1828) non O. F. Müll.

= Orchis foliosa Schur; Enum. Pl. Transs. 642 (1866)

= Orchis grisebachii Pant.; Verh. Ver. Natur- u. Heilk. Presb. 2: 27 (1871-72)

= Orchis hungarica Pant. ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 227 (1933)

= Orchis latifolia f. gracilis Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis latifolia var. conica Lindl.; Gen. Sp. Orchid. Pl.: 260 (1835)

= Orchis latifolia var. cordigera (Fr.) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 62 (1851)

= Orchis latifolia var. orbiculatus Zapal.; Consp. Fl. Gallic. Crit.: 209 (1906)

= Orchis latifolia var. rochelii Griseb. & Schenk; Arch. Naturgesch. 18: 355 (1852)

= Orchis monticola Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 178 (1898), nom. superfl.

= Orchis monticola subsp. bosniaca (Beck) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 180 (1898)

= Orchis monticola subsp. cordigera (Fr.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 179 (1898), nom. superfl.

= Orchis rivularis Heuff. ex Schur; Enum. Pl. Transs. 642 (1866)

= Orchis rocheliana (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 179 (1898), nom. inval.

= Orchis traunsteineri var. grisebachii (Pant.) E.G.Camus; Monogr. Orchid. 183 (1908)

subspecies :

subsp. bosniaca (Beck) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

Distribution: Bosnia & Hercegovina

13 BIH

Synonyms: hidden

= Dactylorhiza bosniaca (Beck) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza lagotis (Rchb.f.) H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 50 (1983)

= Orchis bosniaca Beck; Ann. K. K. Naturhist. Hofmus. 2: 53 (1887)

= Orchis cordigera subsp. bosniaca (Beck) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 34 (1898)

= Orchis cordigera var. bosniaca (Beck) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 28 (1899)

= Orchis lagotis Rchb.f.; Icon. Fl. Germ. 13: t. 403 (1851)

= Orchis latifolia var. lagotis Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 58 (1851)

subspecies :

subsp. graeca (H.Baumann) Kreutz; Kompend. Eur. Orchid. 34 (2004)

Name: Griechisches Knabenkraut

Distribution: NW-Greece (Varvousia Mts.)

13 GRC

Synonyms: hidden

= Dactylorhiza graeca H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 59 (1983)

= Dactylorhiza cordigera var. graeca (H.Baumann) Presser; J. Eur. Orch. 36(4): 981 (2004)

subspecies :

subsp. pindica (B.Willing & E.Willing) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 706 (2005)

Distribution: Bulgaria; NE-Greece

13 BUL GRC

Synonyms: hidden

= Dactylorhiza pindica B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 3(2): 215 (1986 publ. 1987)

= Dactylorhiza majalis subsp. pindica (B.Willing & E.Willing) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 161 (2019)

subspecies :

subsp. siculorum (Soó) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

Distribution: Romania; SW-Ukraine

13 ROM 14 UKR

Synonyms: hidden

= Dactylorhiza majalis subsp. siculorum (Soó) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 81 (1976)

= Dactylorhiza siculorum (Soó) Kreutz & Bobocea; Ber. Arbeitskreis. Heimische Orchid. 38(2): 109 (2021)

= Orchis cordigera subsp. siculorum Sóo; Notizbl. Bot. Gart. Berlin-Dahlem 9(89): 910 (1926), et Magyar Bot. Lapok 25: 274 (1927)

= Orchis cordigera var. siculorum (Soó) Pauca; Fl. Republ. Social. Roman. 12: 701 (1972)

= Orchis siculorum (Soó) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

species :

Dactylorhiza cyrnea Foelsche & Cord-Landwehr; J. Eur. Orch. 44(1): 179 (2012)

Distribution: Corsica

12 COR

Remarks: near D. saccifera

species :

Dactylorhiza elata (Poir.) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

subspecies :

subsp. elata

Name: Hohes Knabenkraut

Distribution: Portugal; Spain; France; Corsica; Sardinia; Morocco; Algeria; Tunisia

12 COR FRA POR SAR SPA 20 ALG MOR TUN

Synonyms: hidden

= Dactylorchis elata (Poir.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorchis munbyana (Boiss. & Reut.) Verm.; Stud. Dactylorch. 69, t. 3, fig. 3 (1947)

= Dactylorhiza elata subvar. maroccanica (Soó) Raynaud; Orchid. Maroc: 104 (1985)

= Dactylorhiza elata subvar. munbyana (Boiss. & Reut.) Raynaud; Orchid. Maroc: 103 (1985)

= Dactylorhiza elata subvar. speciosissima (Soó) Raynaud; Orchid. Maroc: 103 (1985)

= Dactylorhiza elata subvar. stephensonii (Maire & Weiller) Raynaud; Orchid. Maroc: 103 (1985)

= Dactylorhiza elata f. alba (Soó) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza elata f. cordata Landwehr; Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza elata f. leucantha (Maire & Weiller) Raynaud; Orchid. Maroc 104 (1985)

= Dactylorhiza elata f. pallida (Maire & Weiller) Raynaud; Orchid. Maroc 104 (1985)

= Dactylorhiza elata f. peltieri (Soó) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza elata f. speciosissima (Soó) Raynaud; Orchid. Maroc: 103 (1985)

= Dactylorhiza elata subsp. africana (Klinge) H.Sund.; Europ. Medit. Orch. ed. 3: 40 (1980)

= Dactylorhiza elata subsp. munbyana (Boiss. & Reut.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 31(2): 161 (2015)

= Dactylorhiza elata var. algerica (Rchb.f.) P.Delforge; Naturalistes Belges 96(Orchid. 28): 14 (2015)

= Dactylorhiza elata var. corsica (E.Rev. ex E.G.Camus) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Dactylorhiza elata var. munbyana (Boiss. & Reut.) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Dactylorhiza elata var. stephensonii (Maire & Weiller) Raynaud; Orchid. Maroc 103 (1985)

= Dactylorhiza incarnata subsp. africana (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980), isonym

= Dactylorhiza incarnata subsp. africana (Klinge) P.D.Sell; Watsonia 6: 317 (1967)

= Dactylorhiza incarnata subsp. elata (Poir.) H.Sund.; Trop. & Mediterr. Orchideen 45 (1975)

= Dactylorhiza munbyana (Boiss. & Reut.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza munbyana (Boiss. & Reut.) Holub; Folia Geobot. Phytotax. 19(2): 214 (1984)

= Gymnadenia elata (Poir.) Lindl.; Gen. Sp. Orchid. Pl.: 277 (1835)

= Orchis africana (Klinge) A.W.Hill; Index Kew., Suppl. 9: 193 (1938)

= Orchis elata subvar. stephensonii Maire & Weiller; Fl. Afrique N. 6: 343 (1959 publ. 1960)

= Orchis elata f. alba Soó; Bull. Soc. Hist. Nat. Afrique Nord 24: 173 (1933)

= Orchis elata f. albiflora Maire; Bull. Soc. Hist. Nat. Afrique N. 31: 42 (1940)

= Orchis elata f. klingeana Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata f. leucantha Maire & Weiller; Fl. Afrique N. 6: 314 (1959 publ. 1960)

= Orchis elata f. mairei Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis elata f. maroccanica Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis elata f. munbyana (Boiss. & Reut.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis elata f. pallida Maire & Weiller; Fl. Afrique N. 6: 313 (1959 publ. 1960)

= Orchis elata f. peltieri Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis elata f. speciosissima Soó; Bull. Soc. Hist. Nat. Afrique Nord 24: 173 (1933)

= Orchis elata f. willdenowiana Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata Poir.; Voy. Barbarie 2: 248 (1789)

= Orchis elata subsp. munbyana (Boiss. & Reut.) E.G.Camus; Iconogr. Orchid. Europe 221 (1928)

= Orchis elata var. algerica (Rchb.f.) Maire & Weiller; Fl. Afrique N. 6: 312 (1960)

= Orchis elata var. corsica (E.Rev. ex E.G.Camus) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata var. munbyana (Boiss. & Reut.) Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis elata var. poiretiana Maire & Weiller; Fl. Afrique N. 6: 312 (1959 publ. 1960)

= Orchis incarnata subsp. munbyana (Boiss. & Reut.) E.G.Camus; Monogr. Orchid.: 178 (1908)

= Orchis incarnata var. algerica Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 53, t. CCCXCVI (1851)

= Orchis incarnata var. elatior E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 178 (1908)

= Orchis kabyliensis G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A 2: 260 (1933)

= Orchis kabyliensis var. rassautae Alleiz.; Bull. Soc. Bot. France 101: 115 (1954)

= Orchis latifolia subsp. elata (Poir.) Maire; Cat. Pl. Maroc 1: 151 (1931)

= Orchis latifolia var. corsica E.Rev. ex E.G.Camus; J. Bot. (Morot) 6: 158 (1892)

= Orchis latifolia var. elata (Poir.) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 59 (1851)

= Orchis latifolia var. elatior Batt.; Batt. & Trab., Fl. Alger: 196 (1884), nom. illeg.

= Orchis latifolia var. labrovaria Brot.; Phytogr. Lusitan. Select. 2: 25 (1827 publ. 1835)

= Orchis latifolia var. munbyana (Boiss. & Reut.) Batt.; Batt. & Trab., Fl. Alger: 196 (1884)

= Orchis munbyana Boiss. & Reut.; Pugill. Pl. Nov. 112 (1852)

= Orchis orientalis subsp. africana Klinge; Acta Hort. Petrop. 17: 1. 156, in clavi, et 186 (1898)

= Orchis rassautae (Alleiz.) Alleiz.; Bull. Soc. Bot. France 109: 94 (1962), no exact basionym page.

= Orchis sesquipedalis var. algerica (Rchb.f.) Briq.; Prodr. Fl. Corse 1: 369 (1910)

= Orchis sesquipedalis var. corsica (E.Rev. ex E.G.Camus) Briq.; Prodr. Fl. Corse 1: 369 (1910)

= Orchis vestita Cav.; Anal. Cienc. Nat. 6: 142 (1803)

= xOrchidactyla ×kabyliensis (G.Keller) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71: 93 (1986)

subspecies :

subsp. ambigua (J.A.Guim.) Kreutz; Kompend. Eur. Orchid. 35 (2004)

Distribution: France

12 FRA

Synonyms: hidden

= Dactylorhiza elata var. ambigua (J.A.Guim.) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Orchis ambigua Martrin-Donos; Fl. Tarn: 705 (1864), nom. illeg.

= Orchis elata subsp. ambigua (J.A.Guim.) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1927)

= Orchis elata var. ambigua (J.A.Guim.) Schltr.; Keller, Monogr. Iconogr. Orchid. Eur. 1: 181 (1927)

= Orchis incarnata var. ambigua J.A.Guim.; Bol. Soc. Brot. 5: 77 (1887)

= Orchis latifolia var. ambigua (J.A.Guim.) Rouy; Rouy & Foucaud, Fl. France 13: 152 (1912)

variant :

var. occitanica (Geniez, Melki, Pain & Soca) P.Delforge; Naturalistes Belges 92(Orchid. 24): 21 (2011)

Distribution: SW-France

12 FRA

Synonyms: hidden

= Dactylorhiza occitanica Geniez, Melki, Pain & Soca; L´Orchidophile 115: 25 (1995)

subspecies :

subsp. atlantica (Kreutz & Vlaciha) Eccarius; Orchideengattung Dactylorhiza 446 (2016)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Dactylorhiza atlantica Kreutz & Vlaciha; Orchidee (Hamburg) 57(5): 584 (2006)

subspecies :

subsp. elongata (Maire) Kreutz, Rebbas & Véla; Ber. Arbeitskreis. Heimische Orchid. 31(2): 165 (2015)

Distribution: Morocco

20 MOR

Synonyms: hidden

= Dactylorhiza elata var. elongata (Maire) Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 216 (1932)

= Dactylorhiza elata var. elongata (Maire) Raynaud; Orchid. Maroc 103 (1985), isonym

= Orchis elata f. elongata (Maire) Soó; Bull. Soc. Hist. Nat. Afrique N. 24: 173 (1933)

= Orchis latifolia var. elongata Maire; Jahand. et al., Cat. Pl. Maroc 1: 151 (1931)

subspecies :

subsp. mauritanica B.Baumann & H.Baumann; Jour. Eur. Orch. 37(4): 919 (2005)

Distribution: Morocco (Middle Atlas)

20 MOR

subspecies :

subsp. sesquipedalis (Willd.) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

Distribution: Spain; +Corsica; Morocco; Algeria

12 COR SPA 20 ALG MOR

Synonyms: hidden

= Dactylorchis durandii (Boiss. & Reut.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorchis sesquipedalis (Willd.) Verm.; Stud. Dactylorch. 68 (1947)

= Dactylorhiza durandii (Boiss. & Reut.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza durandii (Boiss. & Reut.) M.Laínz; Aport. Conocim. Fl. Gallega. 7: 31 (1971)

= Dactylorhiza elata subsp. durandii (Boiss. & Reut.) Molero Mesa & Pérez Raya; Fl. Sierra Nevada: 365 (1987), isonym

= Dactylorhiza elata subsp. durandii (Boiss. & Reut.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza elata subsp. iberica (T.Stephenson) Kreutz; Eurorchis 17: 104 (2005) [basion. illeg.]

= Dactylorhiza elata var. durandii (Boiss. & Reut.) Landwehr; Wilde Orchid. Eur. 1: 210 (1977)

= Dactylorhiza elata var. iberica (T.Stephenson) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

= Dactylorhiza elata var. sesquipedalis (Willd.) Landwehr; Wilde, Orchid. Eur. 1: 204 (1977)

= Dactylorhiza majalis subsp. sesquipedalis (Willd.) H.A.Pedersen & Hedrén; Bot. J. Linn. Soc. 168(2): 191 (2012)

= Dactylorhiza sesquipedalis (Willd.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 895 (1983), isonym

= Dactylorhiza sesquipedalis (Willd.) M.Laínz; Aport. Conocim. Fl. Gallega. 7: 31 (1971), isonym

= Dactylorhiza sesquipedalis (Willd.) Verm.; Bol. Soc. Brot., ser. 2, 44: 89 (1970)

= Dactylorhiza vestita (Lag. & Rodr.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71: 92 (1986)

= Orchis durandii Boiss. & Reut.; Pugill. Pl. Afr. Bor. Hispan.: 111 (1852)

= Orchis elata subvar. durandiana Maire & Weiller; Fl. Afrique N. 6: 313 (1959 publ. 1960)

= Orchis elata subsp. durandii (Boiss. & Reut.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis elata subsp. sesquipedalis (Willd.) P.Fourn.; Quatre Fl. France 208 (1935), isonym

= Orchis elata subsp. sesquipedalis (Willd.) Soó; Repert. Spec. Nov. Regni Veg. 24: 31 (1928)

= Orchis elata var. sesquipedalis (Willd.) Schltr.; Monogr. Iconogr. Orchid. Eur. 1: 180 (1927)

= Orchis elata var. vestita (Lag. & Rodr.) Losa & Rivas Goday; Arch. Inst. Aclim. Almería 13(1): 79 (1968)

= Orchis incarnata subsp. durandii (Boiss. & Reut.) E.G.Camus; Monogr. Orchid. 180 (1908)

= Orchis incarnata subsp. durandii (Boiss. & Reut.) Gand.; Nov. Consp. Fl. Eur. 461 (1910), isonym

= Orchis incarnata subsp. sesquipedalis (Willd.) E.G.Camus; J. Bot. (Morot) 6: 156 (1892)

= Orchis incarnata subsp. sesquipedalis (Willd.) Gand.; Nov. Consp. Fl. Eur. 461 (1910), isonym

= Orchis incarnata var. durandii (Boiss. & Reut.) Willk.; Prodr. Fl. Hispan. 1(1): 170 (1861)

= Orchis incarnata var. sesquipedalis (Willd.) Rchb.f.; Icon. Fl. Germ. Helv. 13/14: 53, tab. 48 (1851)

= Orchis latifolia proles sesquipedala (Willd.) Rouy; Rouy & Foucaud, Fl. France 13: 151 (1912)

= Orchis latifolia subsp. durandii (Boiss. & Reut.) Batt.; Fl. Algérie Monocot.: 30 (1895)

= Orchis latifolia subsp. durandii (Boiss. & Reut.) Malag.; Acta Phytotax. Barcinon. 1: 64 (1968), isonym

= Orchis latifolia subsp. lusitanica (Steud.) Gand.; Nov. Consp. Fl. Eur.: 461 (1910)

= Orchis latifolia subsp. sesquipedalis (Willd.) Maire; Jahand. et al., Cat. Pl. Maroc 1: 151 (1931)

= Orchis latifolia var. durandii (Boiss. & Reut.) Ball; J. Linn. Soc., Bot. Bot. 16: 672 (1878)

= Orchis lusitanica Steud.; Nom. ed. II, 2: 224 (1841)

= Orchis sesquipedalis var. durandii (Boiss. & Reut.) Briq.; Prodr. Fl. Corse 1: 369 (1910)

= Orchis sesquipedalis var. iberica T.Stephenson; Bull. Soc. Bot. France 492 (1928)

= Orchis sesquipedalis Willd.; Sp. Pl. ed. 4. 4: 30 (1805)

= Orchis vestita Lag. & Rodr.; Anales Ci. Nat. 6: 142 (1803)

species :

Dactylorhiza euxina (Nevski) Czerep.; Sosud. Rast. S.S.S.R.: 308 (1981)

subspecies :

subsp. euxina

Name: Schwarzmeer-Knabenkraut

Distribution: Northern Caucasus; Georgia [Caucasus] (Svaneti, Racha-Lechkhumi, Guria, Adzharia, Shiga Kartli, Kartli, Mtiuleti, Kvemo Kartli, Javakheti); Armenia; Azerbaijan; NE-Turkey

33 ARM AZE GEO NCS 34 TUR

Remarks: incarnata agg.

Synonyms: hidden

= Dactylorhiza cataonica subsp. caucasica (Klinge) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962) [nom. illeg.], et Ann. Univ. Sci. Budapest. Rolando Eotvos, Sect. Biol. 11: 69 (1969)

= Dactylorhiza caucasica (Klinge) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962) [nom. inval. illeg.]

= Dactylorhiza euxina (Nevski) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(2): 235 (1981), isonym

= Dactylorhiza euxina (Nevski) Soó; [nom. inval.]

= Dactylorhiza majalis subsp. caucasica (Klinge) H.Sund.; Europ. Medit. Orchideen 45 (1975)

= Orchis angustifolia var. caucasica (Klinge) E.G.Camus; Monogr. Orchid. 183 (1908)

= Orchis caucasica (Klinge) Klinge; Acta Hort. Petrop. 17 1: 155, in clavi, 181 (1898)

= Orchis caucasica (Klinge) Medw.; Vestn. Tiflissk. Bot. Sada 18: 476 (1919), nom. illeg.

= Orchis cordigera Kraenzl.; Repert. Spec. Nov. Regni Veg., Beih. 65: 39 (1931)

= Orchis cordigera var. caucasica (Klinge) Klinge; V. I. Lipsky, Fl. Cauc. Impr. Colch.: 306 (1899)

= Orchis euxina Nevski; Komarov, Fl. URSS 4: 709 (1935)

= Orchis monticola subsp. caucasica (Lipsky) Klinge; Trudy Imp. S.-Petersburgsk Bot. Sada 17: 35 (1898)

variant :

var. markowitschii (Soó) Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 270 (1983)

Distribution: Northern Caucasus; Transcaucasus

33 NCS

Synonyms: hidden

= Dactylorhiza caucasica var. markowitschii Soó; Nom. Nova Gen. Dactylorhiza 5 (1962)

= Dactylorhiza markowitschii (Soó) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 893 (1983)

= Orchis caucasica var. markowitschii Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 909 (1926)

species :

Dactylorhiza foliosa (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

Name: Madeira-Knabenkraut

Distribution: Madeira

21 MDR

Remarks: maculata agg.

Synonyms: hidden

= Dactylorchis foliosa (Rchb.f.) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorchis orientalis subsp. foliosa (Rchb.f.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 157, 189 (1898)

= Dactylorhiza incarnata subsp. foliosa (Rchb.f.) H.Sund.; Europ. Medit. Orchideen 45 (1975)

= Orchis foliosa Sol. ex Lowe; Trans. Cambridge Philos. Soc. 4: 13 (1831), nom. illeg.

= Orchis incarnata var. foliosa Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 52 (1851)

= Orchis latifolia var. foliosa (Rchb.f.) Rouy; Rouy & Foucaud, Fl. France 13: 151 (1912), nom. illeg.

= Orchis maderensis Summerh.; Kew Bull. 3: 123 (1947 publ. 1948)

= Orchis orientalis subsp. foliosa (Rchb.f.) Klinge; Dactylorch. Monogr. Prodr. 11, 43 (1898)

species :

Dactylorhiza fuchsii (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962)

subspecies :

subsp. fuchsii

Name: Fuchs-Fingerwurz, Fuchs-Fingerknabenkraut (DE)

Distribution: Iceland; England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Rheinland-Pfalz, Schleswig-Holstein, Sachsen, Sachsen-Anhalt); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; France; Andorra; ?Spain; Channel Isl.; Italy; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Estonia; Latvia; Lithuania; Belarus; Ukraine; European Russia (N-European Russia, C-European Russia, W-European Russia, E-European Russia); Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Kazakhstan; China (N-Xinjiang); Mongolia

10 DEN FIN GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS FRA 13 BIH CRO ITA KOS MNT NMA ROM SLV SMA SRB 14 BLR EST LAT LIT RUS UKR 30 SIB 32 KAZ 36 CHI 37 MON

Remarks: maculata agg. Separate from D. maculata based on cytological and genetic grounds. In Central Europa, D. fuchsii is much more abundant than D. maculata, and most records for ´D. maculata´ refer to D. fuchsii

Synonyms: hidden

= Dactylorchis fuchsii (Druce) Verm.; Stud. Dactylorch. 69, 147 (1947)

= Dactylorhiza fuchsii f. alpina Landwehr; Orchideeen 37(3): 78 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza fuchsii f. meyeri (Rchb.f.) P.Delforge; Naturalistes Belges 92: 202 (2011)

= Dactylorhiza fuchsii f. rhodochila (Turner Ettl.) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza fuchsii subsp. meyeri (Rchb.f.) Kulikov & E.G.Philippov; Orchid Memories 107 (2004)

= Dactylorhiza fuchsii var. alpina (Landwehr) R.M.Bateman & Denholm; Watsonia 17(3): 341 (1989)

= Dactylorhiza fuchsii var. longibracteata (Schur) Vöth; Linz. Biol. Beitr. 10(1): 189 (1978)

= Dactylorhiza fuchsii var. magnocalcarata Landwehr; Orchideeen 37(3): 79 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza fuchsii var. meyeri (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962)

= Dactylorhiza fuchsii var. rhodochila Turner Ettl.; Watsonia 18: 308 (1991)

= Dactylorhiza fuchsii var. tatrensis Borsos ex Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Bi 3: 351 (1960)

= Dactylorhiza fuchsii var. trilobata (Bréb.) P.D.Sell; Fl. Great Britain & Ireland 5: 364 (1996)

= Dactylorhiza longibracteata (F.W.Schmidt) Holub; Folia Geobot. Phytotax. 18(2): 204 (1983) [p. p.]

= Dactylorhiza maculata subsp. austriaca Vöth; Linz. Biol. Beitr. 10(1): 190 (1978)

= Dactylorhiza maculata subsp. fuchsii (Druce) Hyl.; Nord. Karlvaxtfl. 2: 238 (1966) [nom. inval.]

= Dactylorhiza maculata subsp. meyeri (Rchb.f.) Tournay; Fl. Belg. Nord France Rég. Voise 691 (1967)

= Dactylorhiza meyeri (Rchb.f.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 307 (1982)

= Orchis comosa F.W.Schmidt; Mayer, Samml. Phys. Aufs. 1: 233 (1791), nom. illeg.

= Orchis fuchsii Druce; Bot. Soc. Exch. Club Brit. Isles 4: 105 (1914 publ. 1915)

= Orchis fuchsii f. lancefolia Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis fuchsii f. maxima Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis longibracteata F.W.Schmidt; Mayer, Samml. Phys. Aufs. 1: 233. t. 2 (1791) [would have priority, but conservation of D. fuchsii proposed]

= Orchis maculata f. alba Zapal.; Consp. Fl. Gallic. Crit.: 216 (1906)

= Orchis maculata f. minor Zapal.; Consp. Fl. Gallic. Crit.: 211 (1906)

= Orchis maculata f. paluster E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 50 (1908)

= Orchis maculata f. recurvifolia Brenner; Acta Soc. Fauna Fl. Fenn. 34(4): 10 (1911)

= Orchis maculata f. trilobata (Bréb.) Soó; Bot. Arch. 23: 78 (1928)

= Orchis maculata subsp. meyeri (Rchb.f.) E.G.Camus & A.Camus; isonym

= Orchis maculata subsp. meyeri (Rchb.f.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata subsp. meyeri (Rchb.) O.Bolòs & Vigo; Fl. Països Catalans 4: 629 (2001), isonym

= Orchis maculata var. curvifolia E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 194 (1908)

= Orchis maculata var. elongata Gadeceau; Mem. Soc. Sci. Nat. Ouest 2: 10 (1892)

= Orchis maculata var. media Bréb.; 313 (1836)

= Orchis maculata var. meyeri Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 67 (1851)

= Orchis maculata var. slandzinskii Zapal.; Consp. Fl. Gallic. Crit.: 211 (1906)

= Orchis maculata var. submontana Zapal.; Consp. Fl. Gallic. Crit. 211 (1906)

= Orchis maculata var. triloba E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 50 (1908)

= Orchis maculata var. trilobata Bréb.; Fl. Normandie ed. 5: 387 (1879)

= Orchis mascula var. triloba Bréb.; Fl. Normandie, ed. 5: 387 (1879)

variant :

var. sacciferoides (Klinge) Landwehr; Orchideeen 31: 78 (1975), et Wilde Orchid. Eur. 1: 114 (1977)

Distribution: Netherlands; Germany; etc.

11 GER NET

Synonyms: hidden

= Dactylorhiza maculata var. sacciferoides (Klinge ex Schulze) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Orchis maculata f. sacciferoides (Klinge ex Schulze) Soó; Bot. Arch. 23: 76 (1928)

= Orchis maculata var. sacciferoides Klinge ex Schultze; Mitt. Thür. Bot. Vereins. 4 (1902)

variant :

var. sooana (Borsos) Kreutz; Kompend. Eur. Orchid. 40 (2004)

Distribution: Poland; Czech Republic; Slovakia; Hungary

11 CZE HUN POL SLO

Conservation Status: IUCN V

Synonyms: hidden

= Dactylorhiza fuchsii subsp. sooana (Borsos) Borsos; Ann. Univ. Sci. Budapest. R. Eötvös 4: 76 (1961)

= Dactylorhiza fuchsii subsp. sooana (Borsos) Soó; Nom. Nova Gen. Dactylorhizae 8 (1962), isonym

= Dactylorhiza maculata subsp. sooana Borsos ex Batoušek; Pl. Syst. Evol. 307(4)-51: 16 (2021)

subspecies :

subsp. carpatica (Batousek & Kreutz) Kreutz; Kompend. Eur. Orchid. 38 (2004)

Distribution: Slovakia (Bile Karpaty)

11 SLO

Synonyms: hidden

= Dactylorhiza carpatica (Batousek & Kreutz) P.Delforge; Naturalistes Belges 81(4): 396 (2000)

= Dactylorhiza traunsteineri subsp. carpatica Batousek & Kreutz; J. Eur. Orchid. 31(3): 684, 683 (1999)

subspecies :

subsp. hebridensis (Wilmott) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962)

Distribution: Scotland (incl. Hebrides); Ireland; European Russia

10 GRB IRE 14 RUS

Remarks: frequently treated as bona sp.

Synonyms: hidden

= Dactylorchis fuchsii subsp. hebridensis (Wilmott) Hesl.-Harr.; Ber. Geobot. Forsch. Inst. Rübel Zürich 1953: 55 (1954)

= Dactylorchis maculata subsp. hebridensis (Wilmott) Verm.

= Dactylorhiza fuchsii var. cornubiensis (Pugsley) Soó; Nom. Nov. Gen. Dactylorhiza: 8 (1962)

= Dactylorhiza fuchsii var. hebridensis (Wilmott) R.M.Bateman & Denholm; Watsonia 17(3): 340 (1989)

= Dactylorhiza hebridensis (Wilmott) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 92 (1986), et Spis. Rast. Gerb. Fl. S.S.S.R. 25 (127130): 26 (1986), et Novosti Sist. Vyssh. Rast. 26: 50 (1989)

= Dactylorhiza maculata subsp. hebridensis (Wilmott) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 102 (1976) [without basionym page]

= Dactylorhiza maculata subsp. hebridensis (Wilmott) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20(3): 631 (1988)

= Orchis fuchsii subsp. hebridensis (Wilmott) Clapham; Fl. Brit. Isles 1317 (1952)

= Orchis hebridensis Wilmott; J. Bot. Lond. 77: 192 (1939)

= Orchis maculata var. cornubiensis Pugsley; J. Bot. 78: 178 (1940)

subspecies :

subsp. okellyi (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962)

Distribution: Scotland; Ireland

10 GRB IRE

Synonyms: hidden

= Dactylorchis fuchsii subsp. okellyi (Druce) Verm.; Stud. Dactylorchis 69 (1947)

= Dactylorhiza fuchsii var. okellyi (Druce) R.M.Bateman & Denholm; Watsonia 17(3): 342 (1989)

= Dactylorhiza maculata subsp. okellyi (Druce) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20(3): 632 (1988)

= Dactylorhiza okellyi (Druce) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Orchis maculata subsp. okellyi (Druce) Druce; Bot. Pocket-book ed. 15: 277 (1917)

= Orchis maculata var. okellyi Druce; Irish Naturalist 1909: 211 (1909)

= Orchis okellyi (Druce) Druce; Rep. Bot. Exch. Cl. Brit. Isles 4: 108 (1914 publ. 1915)

subspecies :

subsp. psychrophila (Schltr.) Holub; Preslia 36: 252 (1964)

Distribution: Norway; Sweden; Finland; European Russia

10 FIN NOR SWE 14 RUS

Synonyms: hidden

= Dactylorchis fuchsii subsp. psychrophila (Schltr.) Verm.; Stud. Dactylorchis 69 (1947)

= Dactylorhiza fuchsii var. psychrophila (Schltr.) B.Bock; Bull. Soc. Bot. Centre-Ouest 43: 214 (2012)

= Dactylorhiza fuchsii var. psychrophila (Schltr.) Soó; Nom. nov. Gen. Dactylorhizae 8 (1962)

= Dactylorhiza maculata subsp. psychrophila (Schltr.) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

= Dactylorhiza maculata var. psychrophila (Schltr.) Vöth; Linz. Biol. Beitr. 12(2): 429 (1980)

= Dactylorhiza psychrophila (Schltr.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 308 (1982)

= Orchis maculata var. psychrophila Schltr.; Keller, Monogr. Iconogr. Orchid. Eur. 1: 183 (1927)

subspecies :

subsp. punicea J.Essink, L.Essink & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 79 (2007)

Distribution: Norway

10 NOR

Synonyms: hidden

= Dactylorhiza fuchsii f. andoeyana (Perko) P.Delforge; Naturalistes Belges 89(Orchid. 21): 250 (2008)

= Dactylorhiza andoeyana Perko; Ber. Arbeitskreis. Heimische Orchid. 24(1): 40 (2007)

subspecies :

subsp. sudetica (Poech ex Rchb.f.) Verm.; Orchideeen 37(3): 78 (1975), et Wilde Orchid. Eur. 1: 106 (1977)

Distribution: Germany (Sachsen); ?Austria; Poland; Czech Republic; European Russia

11 CZE GER POL 14 RUS

Remarks: possibly a synonym of ssp. psychrophila or a nothosubspecies fuchsii - psychrophila

Synonyms: hidden

= Dactylorchis fuchsii subsp. sudetica (Poech ex Rchb.f.) Verm.; Wilde Orch. Eur. 1: 106 (1977)

= Dactylorchis maculata var. sudetica (Poech ex Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 236 (1949)

= Dactylorhiza fuchsii var. sudetica (Poech ex Rchb.f.) H.Baumann, Künkele & R.Lorenz; J. Eur. Orch. 36: 771 (2004)

= Dactylorhiza longibracteata subsp. sudetica (Poech ex Rchb.f.) Holub; Folia Geobot. Phytotax. 18: 205 (1983)

= Dactylorhiza maculata subsp. sudetica (Poech ex Rchb.f.) Vöth; Linz. Biol. Beitr. 12(2): 430 (1980)

= Dactylorhiza maculata var. sudetica (Poech ex Rchb.f.) Baumann, Künkele & R.Lorenz; Jour. Eur. Orch. 34(1): 144 (2002), isonym

= Dactylorhiza sudetica (Poech ex Rchb.f.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 310 (1982)

= Orchis maculata f. sudetica (Poech ex Rchb.f.) Neuman; 631 (1901)

= Orchis maculata subsp. sudetica (Poech ex Rchb.f.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata var. sudetica Poech ex Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 66 (1850)

= Orchis maculata var. sudetica Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 66 (1851)

species :

Dactylorhiza gervasiana (Tod.) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(3): 346 (1981)

Distribution: Italy; ?Sicily

13 ITA

Remarks: rank disputed

Synonyms: hidden

= Dactylorhiza saccifera subsp. gervasiana (Tod.) Kreutz; Kompend. Eur. Orchid. 54 (2004)

= Orchis gervasiana Tod.; Orch. Sicul. 57 (1842)

= Orchis maculata var. gervasiana (Tod.) Nyman; Consp. Fl. Eur. 693 (1882)

species :

Dactylorhiza hatagirea (D.Don) Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol. 3: 341 (1960), et Nom. Nov. Gen. Dactylorhiza: 4 (1962)

Distribution: China (Gansu, Heilongjiang, Jilin, Nei Mongol, Ningxia, Qinghai, W-Sichuan, Xinjiang); E-Tibet; Mongolia; Pakistan (Gilgit, Karakorum, Kagan, Astor, Hazara, Swat); Nepal; Bhutan; India (Himachal Pradesh, Jammu & Kashmir, Sikkim, Uttarakhand)

36 CHI TIB 37 MON 40 BHU IND NEP PAK

Synonyms: hidden

= Dactylorhiza graggeriana (Soó) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza hatagirea var. doniana (Soó) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Dactylorhiza hatagirea var. schlaginweitii (Soó) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Dactylorhiza umbrosa var. afghanica (Soó) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Orchis graggeriana Soó; J. Bot. 66: 15 (1928)

= Orchis hatagirea D.Don; Prod. Fl. Nep. 23 (1825)

= Orchis hatagirea var. afghanica Soó; J. Bot. 66: 19 (1928)

= Orchis hatagirea var. doniana Soó; J. Bot. 66: 19 (1928)

= Orchis hatagirea var. schlaginweitii Soó; J. Bot. 66: 19 (1928)

= Orchis latifolia var. indica Lindl.; Gen. Sp. Orchid. Pl. 260 (1835)

species :

Dactylorhiza iberica (M.Bieb. ex Willd.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

Distribution: Greece; European Turkey; Crimea; Northern Caucasus; Georgia [Caucasus] (Shiga Kartli, Kartli); Armenia; Azerbaijan; Turkey; Iraq; Iran; Cyprus; Lebanon; Syria

13 GRC TUE 14 KRY 33 ARM AZE GEO NCS 34 CYP IRN IRQ LEB SYR TUR

Remarks: maculata agg.

Synonyms: hidden

= Dactylorchis iberica (M.Bieb. ex Willd.) Verm.; Stud. Dactylorch. 65 (1947)

= Gymnadenia angustifolia (M.Bieb.) Spreng.; Syst. Veg. ed. 16. 3: 693 (1826)

= Orchis angustifolia M.Bieb.; Fl. Taur. Caucasus 2: 368 (1808)

= Orchis angustifolia var. longifolia (K.Koch) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 34 (1851)

= Orchis angustifolia var. stevenii Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 34 (1851)

= Orchis iberica f. leptophylla E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 165 (1908)

= Orchis iberica M.Bieb. ex Willd.; Sp. Pl. 4: 25 (1805)

= Orchis iberica var. longifolia (K.Koch) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 34 (1851)

= Orchis iberica var. stevenii Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 34 (1851)

= Orchis leptophylla K.Koch; Linnaea 22: 282 (1850)

= Orchis leptophylla var. laxa K.Koch; Linnaea 22: 282 (1850)

= Orchis leptophylla var. longifolia K.Koch; Linnaea 22: 282 (1850)

= Orchis natolica Fisch. & C.A.Mey.; Ann. Sci. Nat., Bot., sér. 4, 1: 30 (1854)

species :

Dactylorhiza incarnata (L.) Soó; Nom. Nov. Gen. Dactylorhizae 3 (1962)

subspecies :

subsp. incarnata

Name: Steifblättrige Fingerwurz, Fleischrote Fingerwurz (DE)

Distribution: ?Faeroe Isl.; England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; +Portugal; Spain; Andorra; France; Channel Isl.; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; ?Greece; Estonia; Latvia; Lithuania; Belarus; Moldova; Ukraine; Crimea; European Russia (N-European Russia, C-European Russia, W-European Russia, E-European Russia); Siberia (Altai, Buryatia, Irkutsk, Krasnoyarsk, W-Siberia, Yakutia); Russian Far East (Khabarovsk); Kazakhstan; Kyrgyzstan; ?Northern Caucasus; N-Turkey; Mongolia

10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS FRA POR SPA 13 ALB BIH BUL CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR EST KRY LAT LIT MLD RUS UKR 30 SIB 31 RFE 32 KAZ KGZ 34 TUR 37 MON

Remarks: incarnata agg.

Synonyms: hidden

= Dactylorchis incarnata (L.) Verm.; Stud. Dactylorch. 65, 101 (1947)

= Dactylorchis incarnata f. multiangulata Verm.; Ned. Kruidk. Arch. 56: 210 (1949)

= Dactylorchis incarnata f. punctata Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. angustispicata Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. beckii Verm.; Ned. Kruidk. Arch. 56: 211 (1949)

= Dactylorchis incarnata var. brachyphylla Verm.; Ned. Kruidk. Arch. 56: 207 (1949)

= Dactylorchis incarnata var. brevicalcarata (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 210 (1949)

= Dactylorchis incarnata var. divaricata (Rich. ex Loisel.) Verm.; Ned. Kruidk. Arch. 56: 208 (1949)

= Dactylorchis incarnata var. latissima (Zapal.) Verm.; Ned. Kruidk. Arch. 56: 205 (1949)

= Dactylorchis incarnata var. leucantha Verm.; Ned. Kruidk. Arch. 56: 212 (1949)

= Dactylorchis incarnata var. longibracteata Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. noncircumscripta Verm.; Ned. Kruidk. Arch. 56: 211 (1949)

= Dactylorchis incarnata var. remota Verm.; Ned. Kruidk. Arch. 56: 207 (1949)

= Dactylorchis incarnata var. rhombeilabia Verm.; Ned. Kruidk. Arch. 56: 209 (1949)

= Dactylorchis incarnata var. sublatifolia (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 205 (1949)

= Dactylorchis incarnata var. triangularis Verm.; Ned. Kruidk. Arch. 56: 210 (1949)

= Dactylorchis incarnata var. vulgaris Verm.; Ned. Kruidk. Arch. 56: 206 (1949)

= Dactylorhiza comosa (Scop.) P.D.Sell; Sell & Murrell, Fl. Great Britain Ireland 5: 364 (1996)

= Dactylorhiza gemmana (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza incarnata lusus albida (Rchb.f.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971), nom. inval.

= Dactylorhiza incarnata f. ochrantha Landwehr; Orchideeën 37: 80 (1975)

= Dactylorhiza incarnata f. punctata (Verm.) R.M.Bateman & Denholm; Watsonia 15: 346 (1985)

= Dactylorhiza incarnata subsp. gemmana (Pugsley) Hesl.-Harr.; Ber. Geobot. Forsch. Inst. Rübel Zürich 1953: 55 (1954)

= Dactylorhiza incarnata subsp. gemmana (Pugsley) P.D.Sell; Watsonia 6: 317 (1967), isonym

= Dactylorhiza incarnata subsp. punctata R.Doll; Feddes Repert. 89(56): 346 (1978)

= Dactylorhiza incarnata subsp. pyrenaica Balayer; Bull. Soc. Bot. France, Lett. Bot. 133(3): 280 (1986)

= Dactylorhiza incarnata var. brachyphylla (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza incarnata var. latissima (Zapal.) Hyl.; Nord. Kärlväxtfl. 2: 387 (1966)

= Dactylorhiza incarnata var. macrophylla (Schur) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza incarnata var. seemenii (Asch. & Graebn.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza incarnata var. sublatifolia (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza incarnata var. trifurca (Rchb.f.) Aver.; Konsp. Sosud. Rast. Zakaz. Mshinskoe Boloto Leningrad. Oblasti: 38 (1988)

= Dactylorhiza intermedia (Serg.) Kulikov & E.G.Philippov; Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol., n.s., 104(2): 31 (1999)

= Dactylorhiza strictifolia (Opiz) Rauschert ex Hudziok; Wiss. Zeitschr. Mart.-Luth.-Univ. Halle, Math.-Nat. 14: 492 (1965)

= Orchis altaica (Rchb.f.) Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 214 (1933)

= Orchis angustifolia Loisel. ex Rchb.; Iconogr. Bot. Pl. Crit. 9: 17 (1831), nom. inval.

= Orchis angustifolia var. divaricata Nyman; Consp. Fl. Eur. 692 (1882)

= Orchis angustifolia var. fraasii Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 34 (1851)

= Orchis angustifolia Wimm. & Grab.; Fl. Siles. 2(1): 252 (1829), nom. illeg.

= Orchis comosa Scop.; Fl. Carniol., ed. 2, 2: 198 (1771)

= Orchis comosa var. angustifolia Ambrosi; Fl. Tirolo Mer. 1: 794 (1854)

= Orchis condensa Schur; Enum. Pl. Transsilv.: 642 (1866)

= Orchis iberica var. fraasii Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 34 (1851)

= Orchis incarnata f. ampla Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 268 (1924 publ. 1925)

= Orchis incarnata f. brevicalcarata (Rchb.f.) Seeland; 51 (1940)

= Orchis incarnata f. elatior Serg.; Sist. Zametki Mater. Gerb. Krylova Tomsk. Gosud. Univ. Kuybysheva 1: 4 (1929), nom. illeg.

= Orchis incarnata f. lanceata (A.Dietr.) Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis incarnata f. macrophylla (Asch. & Graebn.) Seeland; 87 (1929)

= Orchis incarnata f. macrophylla Schur; Enum. Pl. Transsilv.: 641 (1866)

= Orchis incarnata f. praecox Engenst.; Allg. Bot. Z. Syst. 18: 110 (1912)

= Orchis incarnata f. retusa Soó; Bot. Arch. 23: 66 (1928)

= Orchis incarnata f. rostriformis (Zapal.) Soó; Bot. Arch. 23: 65 (1928)

= Orchis incarnata f. stenophylla (Asch. & Graebn.) Seeland; 87 (1929)

= Orchis incarnata f. stenophylla Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 717 (1907), nom. illeg.

= Orchis incarnata f. subfoliosa M.Schulze ex Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 718 (1907)

= Orchis incarnata L.; Fl. Suec. 312 (1755)

= Orchis incarnata subsp. fraasii (Rchb.f.) K.Richt.; Pl. Eur. 1: 270 (1890)

= Orchis incarnata subsp. fraasii K.Richt.; Pl. Eur. 1: 270 (1890)

= Orchis incarnata subsp. lanceata (A.Dietr.) E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 177 (1908)

= Orchis incarnata subsp. lanceata (A.Dietr.) Gsell; 8 (1936)

= Orchis incarnata var. albida Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 183 (1851), nom. inval.

= Orchis incarnata var. altaica Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 53 (1851)

= Orchis incarnata var. angustifolia Blytt; Enum. Pl. Vasc.: 54 (1844)

= Orchis incarnata var. brevicalcarata Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 53 (1851)

= Orchis incarnata var. brevifolia A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 69 (1919)

= Orchis incarnata var. divaricata (Rich. ex Loisel.) St.-Lag.; Cariot, Étude Fl., éd. 8, 2: 806 (1889)

= Orchis incarnata var. fraasii (K.Richt.) Soó; Bot. Arch. 23: 66 (1928)

= Orchis incarnata var. incisiloba Zapal.; Consp. Fl. Gallic. Crit.: 207 (1906)

= Orchis incarnata var. intermedia Serg.; Sist. Zametki Mater. Gerb. Krylova Tomsk. Gosud. Univ. Kuybysheva 3: 685 (1929)

= Orchis incarnata var. lanceata (A.Dietr.) Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 51 (1851)

= Orchis incarnata var. lanceata-fraasii Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 52 (1851)

= Orchis incarnata var. lanceolata Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 176 (1851)

= Orchis incarnata var. latifolia (L.) Blytt; Enum. Pl. Vasc.: 54 (1844)

= Orchis incarnata var. latissima Zapal.; Consp. Fl. Gallic. Crit. 207 (1906)

= Orchis incarnata var. maxima Cuatrec.; Treb. Inst. Catalana Hist. Nat. 12: 253 (1929)

= Orchis incarnata var. pallida Brenner; Florist. Handb.: 185 (1886)

= Orchis incarnata var. platyphylla Snarskis; Lietuvos T.S.R. Fl. 2: 679 (1963)

= Orchis incarnata var. rhombeilabia-acroglossa Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 53 (1851)

= Orchis incarnata var. rostriformis Zapal.; Consp. Fl. Gallic. Crit.: 207 (1906)

= Orchis incarnata var. seemenii Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 721 (1907)

= Orchis incarnata var. subextensa Neuman; Bot. Not. 1909: 159 (1909)

= Orchis incarnata var. sublatifolia Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 51 (1851)

= Orchis incarnata var. trifurca Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 53 (1851)

= Orchis incarnata var. triloba-retusa Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 53 (1851)

= Orchis lanceata A.Dietr.; Fl. Boruss. t. 5 (1833)

= Orchis latifolia L.; Sp. Pl.: 941 (1753), nom. rej.

= Orchis latifolia subsp. impudica Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927), nom. inval.

= Orchis latifolia subsp. incarnata (L.) Corb.; Nouv. Fl. Normandie 557 (1894), isonym

= Orchis latifolia subsp. incarnata (L.) Hook.f.; Student Fl. Brit. Isl. 353 (1870)

= Orchis latifolia var. angustifolia Bab.; Man. Brit. Bot.: 291 (1843), nom. illeg.

= Orchis latifolia var. angustifolia Wimm. & Grab. ex Döll; 225 (1843)

= Orchis latifolia var. cambrica Pugsley; Bot. J. Linn. Soc. 49: 579 (1935)

= Orchis latifolia var. comosa Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. divaricata (Rich. ex Loisel.) Nyman; Consp. Fl. Eur.: 692 (1882)

= Orchis latifolia var. ecalcarata Lej.; Lej. & Court., Comp. Fl. Belg. 3: 185 (1836), nom. superfl.

= Orchis latifolia var. gemmana Pugsley; J. Linn. Soc., Bot. 49: 578 (1935)

= Orchis latifolia var. hocquartiana Dumort.; Fl. Belg.: 132 (1827)

= Orchis latifolia var. incarnata (L.) Bab.; 310 (1847)

= Orchis latifolia var. incarnata (L.) Coss. & Germ.; Fl. Descr. Anal. Paris, ed. 2: 684 (1861)

= Orchis latifolia var. lanceata Brügger; Jahresber Naturf. Ges. Graubündens [Fl. Curiensis] 17: 58 (1874), nom. inval.

= Orchis latifolia var. lineare Tinant; Fl. Luxemb.: 441 (1836)

= Orchis latifolia var. longibracteata Neilr.; Fl. Wien: 129 (1846)

= Orchis latifolia var. maculata Tinant; Fl. Luxemb.: 441 (1836)

= Orchis latifolia var. nemorosa Westerl.; Bot. Not. 1863: 147 (1863)

= Orchis latifolia var. pallida Tinant; Fl. Luxemb.: 441 (1836)

= Orchis latifolia var. rubra Gray; Nat. Arr. Brit. Pl. 2: 202 (1821 publ. 1822)

= Orchis latifolia var. tharandina Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 58 (1851)

= Orchis macra Schur; Enum. Pl. Transsilv.: 641 (1866), nom. illeg.

= Orchis maculata subsp. incarnata (L.) Douin; Fl. Ill. France 11: 30 (1931)

= Orchis matodes Rchb.f.; Icon. Fl. Germ. 13: t. 398 (1851)

= Orchis militaris Puccin. ex Parl.; Fl. Ital. 3: 521 (1860)

= Orchis mixta Retz.; Fl. Scand. Prodr.: 167 (1779), nom. superfl.

= Orchis mixta var. incantata (L.) Retz.; Fl. Scand. Prodr.: 167 (1779)

= Orchis palmata Gilib.; Exerc. Phyt. 2: 479 (1792), opus utique oppr.

= Orchis palmata subsp. incarnata (L.) Syme; 100 (1869), nom. illeg.

= Orchis richardii Tratt.; Freye Auswahl Pflanzenabbild.: no. 517 (1816)

= Orchis sambucina var. incarnata (L.) Hampe; 55 (1836)

= Orchis strictifolia Opiz; Naturalientausch 10: 217 (1825)

= Orchis tharandina Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: t. 403 (1851)

variant :

var. baumgartneriana (B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter) P.Delforge; Naturalistes Belges 85 (Orchid. 17): 250 (2004)

Distribution: Azerbaijan

33 AZE

Synonyms: hidden

= Dactylorhiza incarnata subsp. baumgartneriana B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter; Jour. Eur. Orch. 35(1): 165 (2003)

= Dactylorhiza kafiriana subsp. baumgartneriana (B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter) Sczep. & Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(2): 103 (2007)

variant :

var. hyphaematodes (Neuman) Løjtnant; Feddes Repert. 89(1): 13 (1978)

Distribution: Denmark; Sweden; N-Germany; etc.

10 DEN SWE 11 GER

Synonyms: hidden

= Dactylorhiza incarnata subsp. hyphaematodes (Neuman) Landwehr; Orchideeen 37: 80 (1975)

= Orchis incarnata var. hyphaematodes Neuman; Bot. Not. 1909: 244 (1909)

variant :

var. immaculata Romolini & Sodi; GIROS Notiz. 25: 21 (2004)

Distribution: Italy

13 ITA

variant :

var. lobelii (Verm.) Soó; Nom. nov. Gen. Dactylorhizae 3 (1962)

Distribution: Denmark; Norway; Netherlands

10 DEN NOR 11 NET

Synonyms: hidden

= Dactylorchis incarnata f. dunensis (Druce) Verm.; Ned. Kruid. Arch. 56: 207 (1949)

= Dactylorchis incarnata var. dunensis (Druce) Verm.; Ned. Kruidk. Arch. 56: 207 (1949)

= Dactylorchis incarnata var. lobelii Verm.; Ned. Kruidk. Arch. 56: 206 (1949)

= Dactylorhiza incarnata subsp. lobelii (Verm.) H.A.Pedersen; Nordic J. Bot. 21(2): 184 (2001)

= Dactylorhiza incarnata var. dunensis (Druce) Hyl.; Nord. Karlvaxtfl. 2: 387 (1966)

= Orchis incarnata var. dunensis Druce; Bot. Soc. Exch. Club Brit. Isl. 4: 212 (1916)

variant :

var. reichenbachii Gathoye & D.Tyteca; Lejeunia 143: 51 (1994)

Distribution: Netherlands; Germany; Hungary; etc.

11 GER HUN NET

Synonyms: hidden

= Dactylorchis incarnata var. extensa (Hartm.) Verm.; Ned. Kruidk. Arch. 56: 208 (1949)

= Dactylorchis incarnata var. haematodes (Rchb.) Verm.; Stud. Dactylorch. 117 (1947)

= Dactylorhiza haematodes (Rchb.) G.H.Loos; Jahrb. Bochum. Bot. Vereins 1: 120 (2010)

= Dactylorhiza incarnata f. haematodes (Rchb.) P.Delforge; Naturalistes Belges 87(Orchids. 19): 259 (2006)

= Dactylorhiza incarnata subsp. haematodes (Rchb.) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza incarnata var. extensa (Hartm.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza incarnata var. haematodes (Rchb.) Soó; Magyar Fl. Veg., Rnd. Kézik.: 171 (1973)

= Dactylorhiza incarnata var. haematodes (Rchb.) Verm.; Ned. Kruid. Arch. 56: 208 (1949)

= Orchis angustifolia var. extensa Hartm.; Handb. Skand. Fl., ed. 4: 281 (1843)

= Orchis cruenta var. haematodes (Rchb.) Neuman; Bot. Not. 1909: 157 (1909)

= Orchis extensa (Hartm.) Pritz.; Icon. Bot. Index 2: 768 (1855)

= Orchis haematodes Rchb.f.; Fl. Germ. Excurs. 126 (1830), nom. illeg.

= Orchis incarnata lusus haematodes (Rchb.) Soó; Bot. Arch. 23: 66 (1928)

= Orchis incarnata f. haematodes (Rchb.) M.Schulze; Asch. & Graebn., Syn. Mitteleur. Fl. 3: 717 (1907)

= Orchis incarnata var. extensa (Hartm.) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 51 (1851)

= Orchis incarnata var. haematodes (Rchb.f.) M.Schultze; Asch. & Graebn., Syn. Mitteleur. Fl. 3(5): 717 (1907)

= Orchis incarnata var. haematodes (Rchb.f.) Pauca; Fl. Republ. Social. Roman. 12: 692 (1972), isonym

= Orchis incarnata var. haematodes (Rchb.) Nyman; Consp. Fl. Eur.: 692 (1882)

= Orchis strictifolia var. extensa (Hartm.) Verm.; Hyl., Uppsala Univ. Årsskr. 7: 117 (1945)

subspecies :

subsp. coccinea (Pugsley) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

Distribution: England; Ireland (coastal)

10 GRB IRE

Synonyms: hidden

= Dactylorchis incarnata subsp. coccinea (Pugsley) Hesl.-Harr.; Ber. Geobot. Forsch. Inst. Rübel Zürich 1953: 54 (1954)

= Dactylorhiza coccinea (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Orchis incarnata f. atrirubra Godfery; Monogr. Ic. Br. Nat. Orchid.: 187 (1933)

= Orchis latifolia var. coccinea Pugsley; Bot. J. Linn. Soc. 49: 579 (1935)

= Orchis strictifolia subsp. coccinea (Pugsley) Clapham; Fl. Brit. Isles 1319 (1952) [nom. inval.]

subspecies :

subsp. cruenta (O.F.Müll.) P.D.Sell; Watsonia 6: 317 (1967)

Name: Blutrote Fingerwurz, Blutrotes Fingerknabenkraut (DE)

Distribution: +Denmark; Norway; Sweden; Finland; Slovakia; France; Italy; Croatia; Bosnia & Hercegovina; Estonia; Latvia; Lithuania; +Belarus; Siberia (Altai, Buryatia, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); China (NW-Xinjiang)

10 DEN FIN NOR SWE 11 SLO 12 FRA 13 BIH CRO ITA 14 BLR EST LAT LIT 30 SIB 36 CHI

Synonyms: hidden

= Dactylorchis cruenta (O.F.Müll.) Verm.; Stud. Dactylorch. 66, 121 (1947)

= Dactylorchis incarnata subsp. cruenta (O.F.Müll.) Verm.; Stud. Dactylorh. 66, 124 (1947)

= Dactylorhiza cruenta (O.F.Müll.) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza cruenta f. ochrantha Wief.; J. Landwehr, Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza cruenta var. lanceolata (Neuman) Landwehr; 80 (1975)

= Dactylorhiza incarnata var. cruenta (O.F.Müll.) Hyl.; Nord. Kärlväxtfl. 2: 387 (1966) [nom. inval.]

= Orchis angustifolia subsp. cruenta (O.F.Müll.) C.Hartm.; Handb. Skand. Fl., ed. 5: 225 (1849)

= Orchis cruenta O.F.Müll.; Oeder et al., Fl. Dan.: t. 876 (1782)

= Orchis cruenta var. lanceolata Neuman; Bot. Not. 1909: 157 (1909)

= Orchis cruenta var. subelliptica Neuman; Bot. Not. 1909: 151 (1909)

= Orchis cruenta var. subtriangularis Neuman; Bot. Not. 1909: 156 (1909)

= Orchis cruentiformis Neuman; Bot. Notiser 243 (1909)

= Orchis incarnata f. cruenta (O.F.Müll.) Cajander; 171 (1906)

= Orchis incarnata subsp. cruenta (O.F.Müll.) Asch. & Graebn.; Syn. Mitteleur. Fl. 3(5): 561 (1907), isonym

= Orchis incarnata subsp. cruenta (O.F.Müll.) Hegi; 352 (1909), isonym

= Orchis incarnata subsp. cruenta (O.F.Müll.) Nyman; Consp. Fl. Eur. 692 (1882)

= Orchis incarnata var. rhombeilabia-cruenta Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 53 (1851)

= Orchis latifolia var. cruenta (O.F.Müll.) Lindl.; Gen. Sp. Orchid. Pl. 260 (1835)

= Orchis latifolia var. subincarnata Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 59 (1851)

subspecies :

subsp. cungsii Kreutz; Ber. Arbeitskreis. Heimische Orchid. 32(2): 158 (2016)

Distribution: Luxembourg

11 LUX

subspecies :

subsp. jugicrucis Akhalk., R.Lorenz & Mosul.; Jahresber. Naturwiss. Vereins Wuppertal 12:36 (2007)

Distribution: Georgia [Caucasus] (Guria, Adzharia, Kartli, Meskheti)

33 GEO

Synonyms: hidden

= Dactylorhiza incarnata var. jugicrucis (Akhalk., R.Lorenz & Mosul.) P.Delforge; Naturalistes Belges 96(Orchid. 28): 15 (2015)

subspecies :

subsp. ochroleuca (Wüstnei ex Boll) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965)

Name: Hellgelbe Fingerwurz, Blassgelbes Fingerknabenkraut (DE)

Distribution: England; Denmark; Sweden; Germany (Brandenburg, Baden-Württemberg, Bayern, Mecklenburg-Vorpommern, +Niedersachsen, +Schleswig-Holstein, +Sachsen-Anhalt); Switzerland; Austria; Poland; Slovakia; Hungary; Croatia; Romania; Estonia; Latvia; Lithuania; Belarus; European Russia

10 DEN GRB SWE 11 AUT GER HUN POL SLO SWI 13 CRO ROM 14 BLR EST LAT LIT RUS

Synonyms: hidden

= Dactylorhiza incarnata subsp. ochroleuca (Wüstnei ex Boll) Hesl.-Harr.f.; Ber. Geobot. Inst. Rübel 1953: 55 (1954), nom. inval. (basionymo non cit.)

= Dactylorhiza incarnata var. ochroleuca (Wüstnei ex Boll) Hyl.; Nord. Kärlvöxtfl. 2: 387 (1966)

= Dactylorhiza incarnata var. straminea (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza ochroleuca (Wüstnei ex Boll) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 893 (1983), isonym

= Dactylorhiza ochroleuca (Wüstnei ex Boll) Holub; Folia Geobot. Phytotax. 9(3): 272 (1974)

= Orchis incarnata subsp. ochroleuca (Wüstnei ex Boll) Hegi; Ill. Fl. Mitt.-Eur. ed. 1, 2: 352 (1909)

= Orchis incarnata subsp. ochroleuca (Wüstnei ex Boll) O.Schwarz; Mitt. Thüring. Bot. Ges. 1: 94 (1949), isonym

= Orchis incarnata var. ochroleuca Wüstnei ex Boll; Arch. Vereins Freunde Naturgesch. Mecklenburg 14: 307 (1860)

= Orchis incarnata var. straminea Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 183 (1851)

= Orchis latifolia var. ochroleuca (Wüstnei ex Boll) Pugsley; J. Linn. Soc., Bot. Bot. 49: 578 (1935)

= Orchis strictifolia var. ochroleuca (Wüstnei ex Boll) Nannf.; Festschrift The Svedberg: 601-622 (1944)

= Orchis traunsteineri subsp. ochroleuca (Wüstnei ex Boll) O.Schwarz; Mitt. Thüring. Bot. Ges. n. s. 1: 94 (1949)

subspecies :

subsp. serotina (Hausskn. ex M.Schulze) Soó & D.M.Moore; Bot. J. Linn. Soc. 76: 367 (1978)

Name: Späte Steifblättrige Fingerwurz, Spätes Steifblättriges Knabenkraut (DE)

Distribution: Ireland; Denmark; Norway; Sweden; Germany; Switzerland; Austria; Czech Republic; Hungary; ?Spain; France; European Russia

10 DEN IRE NOR SWE 11 AUT CZE GER HUN SWI 12 FRA 14 RUS

Synonyms: hidden

= Dactylorchis incarnata var. serotina (Hausskn. ex M.Schulze) Verm.; Stud. Dactylorch.: 162 (1947)

= Dactylorhiza incarnata f. brevibracteata (Landwehr) P.Delforge; Naturalistes Belges 87(Orchid. 19): 258 (2006)

= Dactylorhiza incarnata var. borealis (Neuman) Hyl.; Nord. Kärlväxtfl. 2: 387 (1966)

= Dactylorhiza incarnata var. brevibracteata Landwehr; Orchideeen 37(3): 78 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza incarnata var. haussknechtii (Klinge) Buttler; Bot. Naturschutz Hessen 17: 108 (2004)

= Dactylorhiza incarnata var. serotina (Hausskn. ex M.Schulze) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza serotina (Hausskn. ex M.Schulze) G.H.Loos; Jahrb. Bochum. Bot. Vereins 1: 120 (2010), with incorrect basionym ref.

= Orchis angustifolia var. haussknechtii Klinge; Rev. Orch. cordigera, Orchis angustifolia 70 (1893)

= Orchis incarnata f. serotina (A.F.Schwarz) M.Goldschm.; 156 (1912)

= Orchis incarnata subsp. serotina (A.F.Schwarz) Hegi; 352 (1909)

= Orchis incarnata var. borealis Neuman; Bot. Not. 1909: 229 (1909)

= Orchis incarnata var. haussknechtii (Klinge) O.Schwarz; Mitt. Thüring. Bot. Ges. 1: 94 (1949)

= Orchis incarnata var. serotina Hausskn. & M.Schultze; Orch. Deutschl. Nr. 19, 4 (1894), nom. illeg.

= Orchis serotina (Hausskn. ex M.Schulze) A.F.Schwarz; Fl. Nürnb.: 765 (1901)

= Orchis traunsteineri var. serotina Hausskn. ex M.Schulze; Orch. Deutschl.: 4 (1894)

subspecies :

subsp. versicolor (J.C.Schmidt ex Lüscher) Potucek; Klic. Urc. Ceskosl. Vstavac. Pardubice: 29 (1969) [nothossp.]

Distribution: Germany; Austria

11 AUT GER

Remarks: Dactylorhiza incarnata ssp. incarnata x D. incarnata ssp. ochroleuca

Synonyms: hidden

= Dactylorhiza incarnata nothovar. versicolor (J.C.Schmidt ex Lüscher) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Dactylorhiza ×versicolor (Schmidt & Lüscher) Gathoye & D.Tyteca; Lejeunia 143: 60 (1994)

= Orchis incarnata lusus pulchella Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 270 (1924 publ. 1925)

= Orchis incarnata nothovar. versicolor J.C.Schmidt ex Lüscher; Fl. Aargau: 150 (1918)

= Orchis ×pedemontana Gsell; Jahresb. Naturf. Ges. Graub. (n.f.) 74: 16 (1936) (1936)

species :

Dactylorhiza insularis (Sommier) Landwehr; Orchidee (Hamburg) 20(3): 128 (1969)

Name: Insel-Knabenkraut

Distribution: Portugal; Spain; France; Corsica; Sardinia; Italy; Morocco

12 COR FRA POR SAR SPA 13 ITA 20 MOR

Synonyms: hidden

= Dactylorhiza bartonii (Huxley & P.F.Hunt) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 876 (1984)

= Dactylorhiza iberica subsp. insularis (Sommier) Soó; Nom. Nov. Gen. Dactylorhizae 3 (1962)

= Dactylorhiza insularis (Sommier ex Martelli) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 104 (1976), isonym

= Dactylorhiza insularis (Sommier) Ó.Sánchez & Herrero; Fl. Iber. 21: 98 (2005), isonym

= Dactylorhiza insularis f. bartonii (Huxley & P.F.Hunt) Gathoye & D.Tyteca; Lejeunia 143: 52 (1994)

= Dactylorhiza insularis f. castellana (Rivas Goday) Bernardos; Belgian J. Bot. 135(1-2): 81 (2002)

= Dactylorhiza insularis subsp. castellana (Rivas Goday) D.Rivera & López Vélez; Orq. Prov. Albacete 82 (1987)

= Dactylorhiza insularis var. bartonii (Huxley & P.F.Hunt) D.Rivera & Lopez Velez; Orquíd. Prov. Albacete: 82 (1987), isonym

= Dactylorhiza insularis var. bartonii (Huxley & P.F.Hunt) Landwehr; Orchidee (Hamburg) 20(3): 128 (1969)

= Dactylorhiza romana subsp. bartonii Huxley & P.F.Hunt; J. Roy. Hort. Soc. 92: 309 (1967), et Jahresber. Naturwiss. Vereins Wuppertal 21/22: 97 (1968)

= Dactylorhiza sambucina subsp. insularis (Sommier ex Martelli) Soó; Nom. Nov. Gen. Dactylorhizae 3 (1962)

= Orchis insularis Sommier; Bull. Soc. Bot. Ital. 1895: 248 (1895)

= Orchis insularis Sommier ex Martelli; Monocot. Sardoae, Fasc. 1, 58 (1896)

= Orchis pseudosambucina subsp. castellana Rivas Goday; Farmacognosia 6: 193 (1945)

= Orchis romana var. castellana Rivas Goday; Farmacognosia 6: 193, 197 (1945) [nom. altern.]

= Orchis romana var. insularis (Sommier) E.G.Camus; Monogr. Orchid. 172 (1908)

= Orchis sambucina proles insularis (Sommier) Rouy; Rouy & Foucaud, Fl. France 13: 155 (1912)

= Orchis sambucina subsp. insularis (Sommier) Briq.; Prodr. Fl. Corse 1: 371 (1910)

= Orchis sambucina var. castellana Rivas Goday; Farmacognosia 6: 193, 197 (1945) [nom. altern.]

= Orchis sambucina var. insularis (Sommier) Fiori; Fiori et al., Fl. Anal. Italia 1: 245 (1896)

= Orchis sulphurea subsp. castellana (Rivas Goday) Rivas Goday; Anales Jard. Bot. Madrid 21(2): 306 (1963)

species :

Dactylorhiza isculana Seiser; Ber. Arbeitskreis. Heimische Orchid. 19(1): 23 (2002)

Distribution: Austria (Salzkammergut)

11 AUT

Remarks: provisionally accepted name, status unverified, probably of hybridogenous origin

species :

Dactylorhiza kafiriana Renz; Fl. Iranica 126: 125 (1978)

Distribution: Afghanistan (Badakshan, Wakhan); N-Pakistan (Chitral, Hunza, Gilgit, Karakorum); India (Jammu & Kashmir)

34 AFG 40 IND PAK

species :

Dactylorhiza kalopissii E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 43 (1976)

subspecies :

subsp. kalopissii

Name: Kalopissis´ Knabenkraut

Distribution: Bulgaria; N-Greece

13 BUL GRC

Remarks: praetermissa agg.

Conservation Status: IUCN R

Synonyms: hidden

= Dactylorhiza majalis subsp. kalopissii (E.Nelson) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 154 (2019)

subspecies :

subsp. macedonica (J.Hölz. & Künkele) Kreutz; Kompend. Eur. Orchid. 43 (2004)

Distribution: North Macedonia

13 NMA

Synonyms: hidden

= Dactylorhiza kalopissii f. macedonica (J.Hölz. & Künkele) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza macedonica J.Hölz. & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20(1): 191 (1988)

= Dactylorhiza majalis subsp. macedonica (J.Hölz. & Künkele) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 158 (2019)

subspecies :

subsp. pythagorae (Gölz & H.R.Reinhard) Kreutz; Kompend. Eur. Orchid. 43 (-44) (2004)

Distribution: East Aegaean Isl. (Samos)

34 EAI

Synonyms: hidden

= Dactylorhiza majalis subsp. pythagorae (Gölz & H.R.Reinhard) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 165 (2019)

= Dactylorhiza pythagorae Gölz & H.R.Reinhard; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 24(1): 58, 52 (1992)

species :

Dactylorhiza kerryensis (Wilmott) P.F.Hunt & Summerh.; Watsonia 6: 131 (1965), nom. cons.

Distribution: Ireland

10 IRE

Remarks: for the status as full species see Bateman & Denholm 2011.

Synonyms: hidden

= Dactylorchis kerryensis (Wilmott) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis majalis subsp. occidentalis (Pugsley) Hesl.-Harr.f.; Ber. Geobot. Forsch. Inst. Rübel Zürich 1953: 55 (1954)

= Dactylorchis occidentalis (Pugsley) Verm.; Stud. Dactylorch.: 67 (1947)

= Dactylorhiza comosa f. kerryensis (Roberts) P.D.Sell; Fl. Great Britain & Ireland 5: 365 (1996)

= Dactylorhiza comosa f. occidentalis (Pugsley) P.D.Sell; Sell & Murrell, Fl. Great Britain Ireland 5: 365 (1996)

= Dactylorhiza comosa subsp. occidentalis (Pugsley) P.D.Sell; Fl. Great Britain Ireland 5: 364 (1996)

= Dactylorhiza kerryensis subsp. occidentalis (Pugsley) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20(3): 629 (1988)

= Dactylorhiza kerryensis var. occidentalis (Pugsley) Jebb; Watsonia 27(4): 371 (2009)

= Dactylorhiza latifolia subsp. occidentalis (Pugsley) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Dactylorhiza latifolia var. kerryensis (Wilmott) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza majalis subsp. kerryensis (Wilmott) Senghas; Jahresber. Naturwiss. Ver. Wuppertal 21/22: 53 (1968)

= Dactylorhiza majalis subsp. occidentalis (Pugsley) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 90 (1976), isonym

= Dactylorhiza majalis subsp. occidentalis (Pugsley) P.D.Sell; Acta Fac. Rer. Nat. Univ. Comen. Bot. 14: 19 (1968)

= Dactylorhiza majalis var. kerryensis (Wilmott) R.M.Bateman & Denholm; Watsonia 14(4): 368 (1983)

= Dactylorhiza majalis var. occidentalis (Pugsley) R.M.Bateman & Denholm; Watsonia 14(4): 368 (1983)

= Dactylorhiza occidentalis (Pugsley) P.Delforge; Naturalistes Belges 72(3): 100 (1991)

= Dactylorhiza occidentalis var. kerryensis (Wilmott) R.M.Bateman & Denholm; Watsonia 27(3): 247 (2009)

= Orchis kerryensis Wilmott; Proc. Linn. Soc. Lond. Sess. 148, 126 (1936)

= Orchis majalis var. occidentalis Pugsley; Bot. J. Linn. Soc. 49: 586 (1935)

= Orchis occidentalis (Pugsley) Wilmott; Rep. Bot. Exch. Cl. Brit. Isles 11: 458, 551 (1937 publ. 1938), nom. rej.

= Orchis occidentalis subsp. kerryensis (Wilmott) Clapham; Fl. Brit. Isles 1321 (1952), nom. illeg.

species :

Dactylorhiza lapponica (Laest. ex Hartm.) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

Name: Lappland-Fingerwurz, Lappland-Fingerknabenkraut (DE)

Distribution: Norway; Sweden; Finland; Estonia; N-European Russia; ?Siberia (?W-Siberia)

10 FIN NOR SWE 14 EST RUS

Remarks: majalis agg. Distribution of synonyms between this species and D. traunsteineri is chaotic

Synonyms: hidden

= Dactylorchis lapponica (Laest. ex Hartm.) Verm.; Stud. Dactylorch.: 67 (1947)

= Dactylorhiza angustifolia subsp. pycnantha (Neuman) Aver.; Bot. Not. 1909: 232 (1909)

= Dactylorhiza cruenta subsp. lapponica (Laest. ex Rchb.f.) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 72 (1976) [comb. inval.]

= Dactylorhiza majalis subsp. lapponica (Laest. ex Hartm.) H.Sund.; Europ. Medit. Orchid.: 45 (1975)

= Dactylorhiza pycnantha (Neuman) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(9): 1163 (1983)

= Dactylorhiza traunsteineri subsp. lapponica (Laest. ex Hartm.) Soó; Bot. J. Linn. Soc. 76(4): 367 (1978)

= Dactylorhiza traunsteineri subsp. pycnantha (Neuman) Soó; Nom. Nova Gen. Dactylorhizae 6 (1962), isonym

= Dactylorhiza traunsteineri subsp. pycnantha (Neuman) Verm.; Stud. Dactylorchis 66 (1947)

= Dactylorhiza traunsteineri var. blyttii (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

= Dactylorhiza traunsteineri var. nylandri (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza: 5 (1962)

= Dactylorhiza traunsteineri var. recurva (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza: 6 (1962)

= Dactylorhiza traunsteineri var. tarbatonica (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza: 6 (1962)

= Orchis angustifolia subsp. pycnantha Neuman; Bot. Not. 1909: 232 (1909)

= Orchis angustifolia var. lapponica Laest. ex Hartm.; Handb. Skand. Fl. ed. 4: 281 (1843)

= Orchis angustifolia var. nylandri Klinge; Rev. Orch. cordigera, Orchis angustifolia 67 (1893)

= Orchis angustifolia var. sauteri (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 75 (1899)

= Orchis blyttii Soó; Bot. Ark. 23: 160 (1928), nomen

= Orchis cordigera subsp. blyttii (Rchb.f.) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis cordigera var. blyttii Rchb.f.; Icon. Fl. Germ. Helv. 13/14: 60 (1851)

= Orchis cruenta subsp. blyttii (Rchb.f.) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis lapponica (Laest. ex Hartm.) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis latifolia subsp. lapponica (Laest. ex Rchb.f.) Soó; Repert. Spec. Nov. Regni Veg. 24: 25 (1927)

= Orchis latifolia var. blyttii Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 60 (1851)

= Orchis latifolia var. lapponica (Laest. ex Hartm.) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 58 (1851)

= Orchis maculata subsp. lapponica (Laest. ex Hartm.) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis maculata var. recurva (F.Nyl. ex Fr.) Nyman; Consp. Fl. Eur.: 693 (1882)

= Orchis sambucina var. subalpina Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 52 (1851)

= Orchis traunsteineri var. lapponica Hartm. ex Rchb.f.; Icon. Fl. Germ. Helv. 13/14: 77 (1851)

species :

Dactylorhiza maculata (L.) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

subspecies :

subsp. maculata

Name: Gefleckte Fingerwurz, Geflecktes Fingerknabenkraut (DE)

Distribution: Iceland; Faeroe Isl.; England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Niedersachsen, Nordrhein-Westfalen, ?Rheinland-Pfalz); Switzerland; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; ?Baleares; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Estonia; Latvia; Lithuania; Belarus; ?Moldova; Ukraine; European Russia (N-European Russia, W-European Russia, C-European Russia); Siberia (Krasnoyarsk, W-Siberia)

10 DEN FIN FOR GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LUX NET POL SLO SWI 12 FRA POR SPA 13 BIH CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR EST LAT LIT RUS UKR 30 SIB

Remarks: maculata agg.

Synonyms: hidden

= Dactylorchis maculata (L.) Verm.; Stud. Dactylorch. 68, 130 (1947)

= Dactylorchis maculata var. angustiflora Verm.; Ned. Kruidk. Arch. 56: 235 (1949)

= Dactylorchis maculata var. candidissima (Krock.) Verm.; Ned. Kruidk. Arch. 56: 235 (1949)

= Dactylorchis maculata var. compacta Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. concolor Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. densiflora Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. grandiflora Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. hunyadensis Borsos & Soó; Acta Bot. Acad. Sci. Hung. 5: 324 (1959)

= Dactylorchis maculata var. ibericoides Verm.; Ned. Kruidk. Arch. 56: 235 (1949)

= Dactylorchis maculata var. immaculata (Schur) Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. laxiflora Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. lineata Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. ovalifolia (Beck) Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. spathulata Verm.; Ned. Kruidk. Arch. 56: 232 (1949)

= Dactylorchis maculata var. terschellingensis Verm.; Ned. Kruidk. Arch. 56: 233 (1949)

= Dactylorchis maculata var. traunsteineriifolius Verm.; Ned. Kruidk. Arch. 56: 231 (1949)

= Dactylorchis maculata var. vespertilio (Lindm.) Verm.; Ned. Kruidk. Arch. 56: 234 (1949)

= Dactylorchis maculata var. vexillum (Lindm.) Verm.; Ned. Kruidk. Arch. 56: 234 (1949)

= Dactylorchis maculata var. vulgaris Verm.; Ned. Kruidk. Arch. 56: 236 (1949)

= Dactylorhiza maculata lusus albicans (Marchand ex Tinant) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata lusus carnea (Tinant) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata f. angustiflora (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata f. brevicalcarata (Rohlena) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza maculata subsp. averyanovii (Jagiello) Jagiello; Acta Univ. Wratisl. 1055, 45 (1990)

= Dactylorhiza maculata var. averyanovii Jagiello; Fragm. Flor. Geobot. 3132 (34): 369 (198687 publ. 1988)

= Dactylorhiza maculata var. hunyadensis (Borsos & Soó) Soó; Nom. Nov. Gen. Dactylorhiza: 8 (1962)

= Dactylorhiza maculata var. olgae Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Bi 3: 349 (1960)

= Dactylorhiza maculata var. pseudoelata Balayer; Bull. Soc. Bot. France, Lett. Bot. 133(3): 280 (1986)

= Dactylorhiza maculata var. submacrostachys Borsos ex Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Bi 3: 348 (1960)

= Dactylorhiza maculata var. traunsteineriifolius (Harz) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Orchis basilica L. ex Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 190 (1898), nom. superfl.

= Orchis basilica subsp. maculata (L.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 192 (1898), nom. superfl.

= Orchis biermannii Ortmann; Bot. Zeitung (Berlin) 8: 738 (1850)

= Orchis calvellii A.Terracc.; Annuario Reale Ist. Bot. Roma 4: 47 (1889-1890 publ. 1891)

= Orchis candidissima Krock.; Fl. Siles. 3: 16, t. 2 (1814)

= Orchis danguyana P.Fourn.; Bull. Soc. Bot. France 78: 434 (1931)

= Orchis latifolia var. immaculata Regel; Bull. Soc. Imp. Naturalistes Moscou 41(II): 429 (1868)

= Orchis longibracteata Schur; Enum. Pl. Transs. 643 (1866), non F. W. Schmidt

= Orchis macedonica Griseb.; Spicil. Fl. Rumel. 2: 361 (1844)

= Orchis maculata lusus calvellii (A.Terracc.) Soó; Bot. Arch. 23: 76 (1928)

= Orchis maculata f. bracteosa Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 261 (1895 publ. 1896)

= Orchis maculata f. brevicalcarata Rohlena; Preslia 10: 153 (1931)

= Orchis maculata f. carpatica Zapal.; Consp. Fl. Gallic. Crit.: 211 (1906)

= Orchis maculata f. depressa Engenst.; Allg. Bot. Z. Syst. 18: 110 (1912)

= Orchis maculata f. falcata A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 154 (1919)

= Orchis maculata f. lancifolia Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 258 (1895 publ. 1896)

= Orchis maculata f. leucantha Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 4: 213 (1915 publ. 1916)

= Orchis maculata f. macroglossa Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 5: 579 (1919)

= Orchis maculata f. media E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 50 (1908)

= Orchis maculata f. oblongifolia Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 258 (1895 publ. 1896)

= Orchis maculata f. obscura Neuman; Bot. Not. 1909: 153 (1909)

= Orchis maculata f. obtusifolia Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis maculata f. ovalifolia Beck; Fl. Nieder-Österreich: 204 (1890)

= Orchis maculata f. subintegriflora Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 1921: 396 (1921)

= Orchis maculata f. traunsteineriifolius Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 260 (1895 publ. 1896)

= Orchis maculata unranked immaculata Schur; Oesterr. Bot. Z. 20: 366 (1870)

= Orchis maculata L.; Sp. Pl. 942 (1753)

= Orchis maculata var. albicans Marchand ex Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. alpina Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis maculata var. angustata Harz; Fl. Deutschl. ed. 5 Neuer Abdruck 4: 261 (1896)

= Orchis maculata var. angustifolia Hartm.; Handb. Skand. Fl. 328 (1820)

= Orchis maculata var. angustifolia Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. biermannii (Ortm.) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis maculata var. candidissima (Krock.) Nyman; Consp. Fl. Eur. Suppl. 2: 291 (1890)

= Orchis maculata var. carnea Tinant; Fl. Luxemb.: 441 (1836)

= Orchis maculata var. comosa Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis maculata var. concolor Lange; Haandb. Danske Fl.: 502 (1851)

= Orchis maculata var. elabiata R.Keller; Ber. Schweiz. Bot. Ges. 14: 116 (1904)

= Orchis maculata var. foliis-immaculatis Tinant; Fl. Luxemb.: 441 (1836), nom. inval.

= Orchis maculata var. gruenii Van Heurck & Wesm.; Prodr. Fl. Brabant 79 (1861)

= Orchis maculata var. hunyadensis (Borsos & Soó) Pauca; Fl. Republ. Social. Roman. 12: 698 (1972)

= Orchis maculata var. latifolia Hartm.; Handb. Skand. Fl. 328 (1820)

= Orchis maculata var. longibracteata Schur; Oesterr. Bot. Z. 20: 296 (1870)

= Orchis maculata var. lusitanica J.A.Guim.; Bol. Soc. Brot. 5: 79 (1887)

= Orchis maculata var. macrobracteata Borbás; Deutsche Bot. Monatsschr. 1883: 296 (1883)

= Orchis maculata var. minor Bréb.; Fl. Normandie ed. 5: 389 (1879)

= Orchis maculata var. montana Marchand ex Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. olgae (Soó) Pauca; Fl. Republ. Social. Roman. 12: 698 (1972)

= Orchis maculata var. paludosa Marchand ex Tinant; Fl. Luxemb. 441 (1836)

= Orchis maculata var. palustris Cortesi; 193 (1903)

= Orchis maculata var. pumila Neuman; Bot. Not. 1909: 245 (1909)

= Orchis maculata var. sandomiriensis Zapal.; Consp. Fl. Gallic. Crit. 211 (1906)

= Orchis maculata var. vespertilio Lindm.; Kongl. Svenska Vetensk. Acad. Handl., n.s., 23(3, 1): 6 (1897)

= Orchis maculata var. vexillum Lindm.; Kongl. Svenska Vetensk. Acad. Handl. n. s., 23(3, 1): 6 (1897)

= Orchis nemorosa Montandon; Syn. Fl. Jura, ed. 2: 294 (1868)

= Orchis obtusifolia Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis solida Moench; Meth. 713 (1794), nom. illeg.

= Orchis tetragona Heuff.; Flora 16: 363 (1833)

variant :

var. arduennensis (Zadoks) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 153 (2007)

Distribution: Belgium; ?Germany

11 BGM

Synonyms: hidden

= Dactylorchis arduennensis Zadoks; Bull. Soc. Naturalistes Luxemb. 59: 37 (1954)

= Dactylorchis maculata subsp. arduennensis Zadoks; Bull. Soc. Naturalistes Luxemb. 59: 125 (1954)

= Dactylorhiza maculata f. arduennensis (Zadoks) P.Delforge; Naturalistes Belges 92(Orchid. 24): 202 (2011)

= Dactylorhiza maculata subsp. arduennensis (Zadoks) Tournay; Fl. Belg. Nord France Rég. Voise 691 (1967)

subspecies :

subsp. caramulensis Verm.; Bol. Soc. Brot. II 44: 94 (1970)

Distribution: Portugal

12 POR

Synonyms: hidden

= Dactylorhiza caramulensis (Verm.) D.Tyteca; L´Orchidophile 88: 155 (1989)

subspecies :

subsp. elodes (Griseb.) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

Name: Heidemoor-Fingerwurz, Heidemoor-Fingerknabenkraut (DE)

Distribution: Norway; Sweden; Finland; Netherlands; NW-Germany; ?Slovakia; E-France; Estonia

10 FIN NOR SWE 11 GER NET 12 FRA 14 EST

Synonyms: hidden

= Dactylorchis elodes (Griseb.) Verm.; Stud. Dactylorch. 137 (1947)

= Dactylorchis maculata subsp. elodes (Griseb.) Verm.; Stud. Dactylorchis 68 (1947)

= Dactylorhiza elodes (Griseb.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 309 (1982)

= Dactylorhiza elodes (Griseb.) Englmaier; Abh. zool.-bot. Ges. Österr. 22: 107 (1984), isonym

= Dactylorhiza maculata var. dalarnensis Landwehr; Orchideeen 37(3): 77 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza maculata var. elodes (Griseb.) Aver.; Turczaninowia 3(1): 49 (2000)

= Dactylorhiza maculata var. nummularia Pikner; Ber. Arbeitskreis. Heimische Orchid. 30: 295 (2013)

= Orchis elodes Griseb.; Bild. Torfs. (in Goett. Studien) (1845), Gesam. Abh. 67

= Orchis maculata proles elodes (Griseb.) Rouy; Rouy & Foucaud, Fl. France 13: 154 (1912)

= Orchis maculata subsp. elodes (Griseb.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata subsp. elodes (Griseb.) O.Bolòs & Vigo; Fl. Països Catalans 4: 629 (2001), isonym

= Orchis maculata subsp. elodes (Griseb.) P.Fourn.; Quatre Fl. France 206 (1935)

= Orchis maculata var. elodes (Griseb.) Jess.; 482 (1879), isonym

= Orchis maculata var. elodes (Griseb.) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 67 (1851)

subspecies :

subsp. ericetorum (E.F.Linton) P.F.Hunt & Summerh.; Watsonia 6: 132 (1965)

Distribution: Faeroe Isl.; England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; ?Luxembourg; Germany; Slovakia; Portugal; Spain; France; Channel Isl.; ?Romania

10 DEN FIN FOR GRB IRE NOR SWE 11 BGM GER NET SLO 12 CIS FRA POR SPA

Synonyms: hidden

= Dactylorchis maculata subsp. ericetorum (E.F.Linton) Verm.; Stud. Dactylorchis 69 (1947)

= Dactylorchis maculata subsp. rhoumensis (Hesl.-Harr.f.) Hesl.-Harr.f.; Watsonia 4: 48 (1957)

= Dactylorchis maculata var. ericetorum (E.F.Linton) Verm.; Fl. Neerlandica 1(5): 85 (1958)

= Dactylorhiza ericetorum (E.F.Linton) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 309 (1981)

= Dactylorhiza fuchsii subsp. rhoumensis (Hesl.-Harr.f.) A.R.Clapham, Tutin & E.F.Warb.; Fl. Brit. Isles ed. 2: 1046 (1962)

= Dactylorhiza maculata subsp. rhoumensis (Hesl.-Harr.f.) Soó; Nom. Nov. Gen. Dactylorhiza 7 (1962)

= Dactylorhiza maculata var. ericetorum (E.F.Linton) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 3: 349 (1960), nom. inval.

= Dactylorhiza maculata var. praecox (Webster) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Dactylorhiza maculata var. rhoumensis (Hesl.-Harr.f.) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 15 (2015)

= Orchis ericetorum (E.F.Linton) A.Benn.; Ann. Scott. Nat. Hist. 1905: 39 (1905)

= Orchis fuchsii subsp. rhoumensis Hesl.-Harr.f.; Trans. Bot. Soc. Edinburgh 35: 53 (1948)

= Orchis maculata f. ericetorum (E.F.Linton) Hagerup; Dansk Bot. Ark. 11: 3 (1944)

= Orchis maculata subsp. ericetorum E.F.Linton; Fl. Bournemouth 208 (1900)

= Orchis maculata subsp. rhoumensis Hesl.-Harr.f.; Trans. Bot. Soc. Edinburgh 35: 64 (1948)

= Orchis maculata var. praecox Webster; Brit. Orchids: 54 (1886)

subspecies :

subsp. islandica (Á.Löve & D.Löve) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

Distribution: Iceland

10 ICE

Synonyms: hidden

= Dactylorchis maculata subsp. islandica Á.Löve & D.Löve; Chrom. Numb. Nort. Pl. 106 (1948)

= Dactylorhiza islandica (Á.Löve & D.Löve) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza maculata var. islandica (Á.Löve & D.Löve) Hyl.; Nord. Kärlväxtfl. 2: 387 (1966)

subspecies :

subsp. kolaensis (Montell) Kreutz; Kompend. Eur. Orchid. 46 (2004)

Distribution: Norway; Sweden; Finland; European Russia (NW-European Russia)

10 FIN NOR SWE 14 RUS

Synonyms: hidden

= Dactylorchis maculata subsp. montellii Verm.; Stud. Dactylorchis 69, 141 (1947)

= Dactylorhiza kolaensis (Montell) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza maculata subsp. deflexa Landwehr; Orchideeen 37(3): 77 (1975), et Wilde Orchid. Eur. 2: 556 (1977)

= Dactylorhiza maculata subsp. montellii (Verm.) Landwehr; Orchideeen 37(3): 79 (1975)

= Dactylorhiza maculata var. kolaensis (Montell) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

= Dactylorhiza montellii (Verm.) P.Delforge; Naturalistes Belges 71(3): 101 (1990)

= Orchis maculata var. kolaensis Montell; Mem. Soc. Fauna Flora Fenn. 23: 166 (1947)

subspecies :

subsp. podesta (Landwehr) Kreutz; Kreutz & H. Dekker, Orchid. van Nederland 179 (2000)

Distribution: Scotland; Netherlands (Terschelling Isl.)

10 GRB 11 NET

Synonyms: hidden

= Dactylorhiza maculata var. podesta Landwehr; Wilde Orchid. Eur. 2: 556 (1977)

subspecies :

subsp. pyrenaica Kreutz; Ber. Arbeitskreis. Heimische Orchid. 30(2): 279 (2013)

Distribution: Andorra; SW-France (Pyrenees)

12 ADO FRA

Synonyms: hidden

= Dactylorhiza savogiensis f. pyrenaica (Kreutz) P.Delforge; Naturalistes Belges 96 (Orchid. 28): 15 (2015)

subspecies :

subsp. savogiensis (D.Tyteca & Gathoye) Kreutz; Kompend. Eur. Orchid. 46 (2004)

Distribution: ?Germany; Switzerland; France; Italy

11 SWI 12 FRA 13 ITA

Synonyms: hidden

= Dactylorhiza fuchsii var. brachystachys (E.G.Camus) Soó; Nom. Nov. Gen. Dactylorhiza: 8 (1962)

= Dactylorhiza savogiensis D.Tyteca & Gathoye; L´Orchidophile 92: 108 (1990)

= Orchis maculata var. brachystachys E.G.Camus; Camus, Iconogr. Orchid. Europe, Atlas 1: 4 (1921)

subspecies :

subsp. schurii (Klinge) Soó; Rev. Roum. Biol. (Bot.) 12: 229 (1967)

Distribution: ?Poland; ?Slovakia; ?Romania; Estonia; Latvia; Lithuania; European Russia (W-European Russia)

14 EST LAT LIT RUS

Synonyms: hidden

= Dactylorhiza maculata var. schurii (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Dactylorhiza schurii (Klinge) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza traunsteineri subsp. schurii (Klinge) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 31: 106 (2014 publ. 2015)

= Orchis angustifolia f. schurii Klinge; Rev. Orch. cordigera, Orchis angustifolia: 63, 83 (1893)

= Orchis maculata f. schurii (Klinge) Soó; Keller, Monogr. Iconogr. Orchid. Eur. 2: 240, 243 (1933)

= Orchis maculata var. schurii (Klinge) Pauca; Fl. Republ. Social. Roman. 12: 697 (1972)

= Orchis traunsteineri var. schurii (Klinge) Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 729 (1907)

subspecies :

subsp. sooana Borsos ex Batousek; Pl. Syst. Evol. 307(4)-51: 16 (2021)

Distribution: Slovakia

11 SLO

subspecies :

subsp. transsilvanica (Schur) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962)

Name: Siebenbürger Fingerwurz, Siebenbürger Fingerknabenkraut (DE)

Distribution: Czech Republic; Slovakia; Slovenia; Croatia; Bosnia & Hercegovina; Serbia; Kosovo; Romania; Estonia; European Russia

11 CZE SLO 13 BIH CRO KOS ROM SLV SRB 14 EST RUS

Synonyms: hidden

= Dactylorchis maculata subsp. transsilvanica (Schur) Verm.; Stud. Dactylorchis 69 (1947)

= Dactylorhiza fuchsii subsp. transsilvanica (Schur) S.E.Fröhner; Ber. Arbeitsgem. Sächs. Bot. N. F.8: 143 (1968)

= Dactylorhiza maculata var. transsilvanica (Schur) P.Delforge; Naturalistes Belges 81(4): 397 (2000)

= Dactylorhiza maculata var. wettsteinii (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhiza: 8 (1962)

= Dactylorhiza transsilvanica (Schur) Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 309 (1982)

= Orchis maculata f. wettsteinii (H.Fleischm.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis maculata subsp. transsilvanica (Schur) Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 911 (1926)

= Orchis maculata var. transsilvanica (Schur) Nyman; Consp. Fl. Eur., Suppl. 2: 291 (1890)

= Orchis ochrantha (Pancic) H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 176 (1908 publ. 1909)

= Orchis maculata var. ochrantha Pancic; Verh. Zool.-Bot. Ges. Wien 6: 575 (1856)

= Orchis ochrantha var. wettsteinii H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 178 (1909)

= Orchis transsilvanica Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

species :

Dactylorhiza magna (Czerniak.) Ikonn.; Novosti Sist. Vyssh. Rast. 9: 303 (1972)

subspecies :

subsp. magna

Distribution: Uzbekistan; Tajikistan

32 TZK UZB

Synonyms: hidden

= Dactylorhiza baldshuanica Chernyak.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 534 (1983) ex Aver.

= Dactylorhiza umbrosa subsp. magna (Czerniak.) Soó; Ann. Univ. Sci. Budapest. Rolando Eotvos, Sect. Biol. 11: 68 (1969)

= Orchis magna Czerniak.; Fl. Uzbekistan. 1: 546 (1941)

subspecies :

subsp. czerniakowskae (Aver.) Eccarius; Orchideengattung Dactylorhiza 102 (2016)

Distribution: Kazakhstan; Uzbekistan; Tajikistan

32 KAZ TZK UZB

Synonyms: hidden

= Dactylorhiza czerniakowskae Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 536 (1983)

subspecies :

subsp. kulikalonica (Chernyak.) Eccarius; Orchideengattung Dactylorhiza 106 (2016)

Distribution: Kazakhstan; Uzbekistan; Tajikistan

32 KAZ TZK UZB

Synonyms: hidden

= Dactylorhiza kulikalonica Chernyak.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 535 (1983) ex Aver.

species :

Dactylorhiza majalis (Rchb.) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965), nom. cons.

subspecies :

subsp. majalis

Name: Breitblättrige Fingerwurz, Breitblättriges Fingerknabenkraut (DE)

Distribution: Denmark; Norway; Sweden; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Romania; Lithuania; Belarus; Ukraine; European Russia

10 DEN NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO FRA SPA 13 BIH CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR LIT RUS UKR

Remarks: majalis agg.

Synonyms: hidden

= Dactylorchis latifolia (L.) Rothm.; Feddes Repert. Beih. 67 (1-3): 4 (1963) [nom. rej.]

= Dactylorchis latifolia subsp. brevifolia Bisse; Feddes Repert. Spec. Nov. Regni Veg. 67: 184 (1963)

= Dactylorchis majalis (Rchb.) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis majalis f. albolileacea Verm.; Ned. Kruidk. Arch. 56: 217 (1949)

= Dactylorchis majalis f. ampla (Asch. & Graebn.) Verm.; Ned. Kruidk. Arch. 56: 214 (1949)

= Dactylorchis majalis var. aequiloba (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. albiflora (Tinant) Verm.; Ned. Kruidk. Arch. 56: 218 (1949)

= Dactylorchis majalis var. alborosacea (Erdner) Verm.; Ned. Kruidk. Arch. 56: 217 (1949)

= Dactylorchis majalis var. alpestroides Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. atrorubens Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. brachyphylla Verm.; Ned. Kruidk. Arch. 56: 213 (1949)

= Dactylorchis majalis var. brevicalcarata (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. gracilis Verm.; Ned. Kruidk. Arch. 56: 213 (1949)

= Dactylorchis majalis var. immaculata (Regel) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. integra (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. laxa (Harz) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. linguata (Harz) Verm.; Ned. Kruidk. Arch. 56: 216 (1949)

= Dactylorchis majalis var. macrobracteata (Schur) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorchis majalis var. magnicalcarata (Harz) Verm.; Ned. Kruidk. Arch. 56: 217 (1949)

= Dactylorchis majalis var. submaculata (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 215 (1949)

= Dactylorhiza comosa subsp. majalis (Rchb.) P.D.Sell; Fl. Great Britain & Ireland 5: 365 (1996)

= Dactylorhiza fistulosa (Moench) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(4): 456 (1983)

= Dactylorhiza latifolia (L.) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(4): 444 (1983), isonym

= Dactylorhiza latifolia (L.) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962) [nom. illeg.]

= Dactylorhiza latifolia var. gracilis (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Dactylorhiza latifolia var. limbata Hudziok; 707 (1963)

= Dactylorhiza latifolia var. pinguis (Asch. & Graebn.) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Dactylorhiza majalis (Rchb.) Rauschert ex Hudziok; Wiss. Zeitschr. Mart.-Luth.-Univ. Halle, Math.-Nat. 14: 492 (1965)

= Dactylorhiza majalis lusus albiflora (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus alboliliacea (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus alborosacea (Erdner) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus carnea (Neuman) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis lusus pallida (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. alpestroides (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. ampla (Asch. & Graebn.) Landwehr; 79 (1975)

= Dactylorhiza majalis f. aspersa F.Proch.; Preslia 54: 290 (1982)

= Dactylorhiza majalis f. atrorubens (Verm.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. barlae (E.G.Camus) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. brevicalcarata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. chwaszczyniensis Marg. & Tukallo; Richardiana 12(2): 53 (51-58

= Dactylorhiza majalis f. heikeana Gottschalk; 14 (2014)

= Dactylorhiza majalis f. immaculata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. laxa (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. linearis (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. linguata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. maculata (Thielens) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. magnicalcarata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis f. rubra (Winterl) D.Tyteca & Gathoye; L´Orchidophile 92: 112 (1990)

= Dactylorhiza majalis f. tharandina (Rchb.f.) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza majalis subsp. brevifolia (Bisse) Senghas; Jahresber. naturw. Ver. Wuppertal 21/22: 126 (1963)

= Dactylorhiza majalis var. druceoides Balayer; Bull. Soc. Bot. France, Lett. Bot. 133(3): 280 (1986)

= Dactylorhiza majalis var. pinguis (Asch. & Graebn.) Landwehr; 80 (1975)

= Dactylorhiza majalis var. semimaculata R.Doll; Feddes Repert. 89(56): 347 (1978)

= Orchis divaricata Rich. ex Loisel.; Fl. Gall. ed. I, 606 (1807)

= Orchis elata Hartm.; Handb. Skand. Fl., ed. 10, 202 (1870) non Poir. 1789

= Orchis fistulata Stokes; Bot. Mat. Med. 4: 296 (1812)

= Orchis fistulosa Moench; Meth. 713 (1794), nom. illeg.

= Orchis incarnata subsp. latifolia (L.) Cajander; 171 (1906)

= Orchis latifolia f. aequiloba (Harz) Soó; Bot. Arch. 23: 69 (1928)

= Orchis latifolia f. ampla Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 735 (1907)

= Orchis latifolia f. brevicalcarata (Harz) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. densa (Harz) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. inaequiloba (Harz) Soó; Bot. Arch. 23: 69 (1928)

= Orchis latifolia f. laxa (Harz) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. laxiflora Zapal.; 215 (1906)

= Orchis latifolia f. macrobracteata (Schur) Soó; Bot. Arch. 23: 68 (1928)

= Orchis latifolia f. macrochlamys (Asch. & Graebn.) M.Goldschm.; 285 (1908)

= Orchis latifolia f. macrophylla Zapal.; 215 (1906)

= Orchis latifolia f. pinguis Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 735 (1907)

= Orchis latifolia Scop.; Fl. Carniol., ed. 2, 2: 196 (1771), nom. illeg.

= Orchis latifolia subsp. majalis (Rchb.) Ces.; 315 (1844)

= Orchis latifolia subsp. majalis (Rchb.) Gand.; Nov. Consp. Fl. Eur. 461 (1910), isonym

= Orchis latifolia subsp. majalis (Rchb.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 168 (1898), isonym

= Orchis latifolia var. abbreviata Harz; Fl. Deutschl. ed. 5 Neuer Abdruck 4: 242 (1895 publ. 1896)

= Orchis latifolia var. aequiloba Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. aggregata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. albiflora Tinant; Fl. Luxemb. 441 (1836)

= Orchis latifolia var. albiflora Zapal.; 214 (1906)

= Orchis latifolia var. ampla Asch. & Graebn.; 735 (1907)

= Orchis latifolia var. brevibracteata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. brevicalcarata Harz; Fl. Deutschl. ed. 5 Neuer Abdruck 4: 242 (1896)

= Orchis latifolia var. carnea R.Neumann; 9 (1905)

= Orchis latifolia var. densa Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. densiflora Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. elata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896), nom. illeg.

= Orchis latifolia var. gracilis Warnst.; Deutsche Bot. Monatsschr. 1883: 169 (1883), nom. illeg.

= Orchis latifolia var. humilis Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. inaequiloba Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. integra Harz; Fl. Deutschl. ed. 5 Neuer Abdruck 4: 242 (1896)

= Orchis latifolia var. lanceata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. laxa Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. laxiuscula Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. linguata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. macilenta Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. macrobracteata Schur; Mitt. Geogr. Ges. (Thüringen) Jena 2: 220 (1884 publ. 1885)

= Orchis latifolia var. macrochlamys Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 735 (1907)

= Orchis latifolia var. magnicalcarata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. majalis (Rchb.) Kitt.; Taschenb. Fl. Deutschl.: 137 (1837)

= Orchis latifolia var. majalis (Rchb.) Nyman; Consp. Fl. Eur.: 692 (1882)

= Orchis latifolia var. obtusa Gray; Nat. Arr. Brit. Pl. 2: 202 (1822)

= Orchis latifolia var. obtusa Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896), nom. illeg.

= Orchis latifolia var. palustris Gray; Nat. Arr. Brit. Pl. 2: 202 (1822)

= Orchis latifolia var. parvicalcarata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. parvilabia Zapal.; 215 (1906)

= Orchis latifolia var. pinguis (Asch. & Graebn.) Hegi; 355 (1909)

= Orchis latifolia var. praecox M.Loehr in Wirtg.; 164 (1842)

= Orchis latifolia var. pyramidalis Harz; Fl. Deutschl. ed. 5 Neuer Abdruck 4: 242 (1896)

= Orchis latifolia var. robusta Harz; Fl. Deutschl. ed. 5 Neuer Abdruck 4: 242 (1896)

= Orchis latifolia var. sparsiflora Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. submaculata Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 57 (1851)

= Orchis latifolia var. traunsteineriiformis Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. trifida Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. trilobata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 242 (1895 publ. 1896)

= Orchis latifolia var. uncinata Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 176 (1851)

= Orchis maculata subsp. latifolia Bonnier & Layens; Tabl. Syn. Pl. Vasc. France: 311 (1894)

= Orchis maculata subsp. latifolia Douin; Bonnier, Fl. Ill. France 11: 30 (1931)

= Orchis majalis lusus albiflora Thielens; Fl. Luxemb.: 60 (1875)

= Orchis majalis lusus alborosacea Erdner; Ber. Naturwiss. Vereins Schwaben Neuburg 39: 142 (1911)

= Orchis majalis lusus pallida Thielens; Fl. Luxemb.: 60 (1875)

= Orchis majalis carnea Neuman; Mitt. Bad. Landesvereins Naturk. 5: 201 (1905)

= Orchis majalis f. barlae E.G.Camus; Camus, Iconogr. Orchid. Europe: 4 (1921)

= Orchis majalis f. brevicalcarata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. immaculata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. laxa Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. linearis Thielens; Fl. Luxemb.: 60 (1875)

= Orchis majalis f. linguata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis f. maculata Thielens; Fl. Luxemb.: 60 (1875)

= Orchis majalis f. magnicalcarata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 247 (1895 publ. 1896)

= Orchis majalis Rchb.; Iconogr. Bot. Pl. Crit. 6: 7 (1828), nom. cons.

= Orchis majalis var. pinguis (Asch. & Graebn.) Pugsley; 585 (1935)

= Orchis majalis var. tharandina Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 58 (1851)

= Orchis palmata Syme; 99 (1869), nom. illeg.

= Orchis tingitana Durand ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 230 (1933)

= Orchis ×ulehlae Podp.; Domin & Podpera, Klic K Ulpne Kvetene Rep. Ceskoslov. 796 (1928), et ex Soó, in Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 199 (1932)

variant :

var. alpestris (Pugsley) Wucherpf.; Ber. Arbeitskreis. Heimische Orchid. 19(2): 106 (2002 publ. 2003)

Distribution: Norway; Germany; Switzerland; Austria; Spain; France; Italy

10 NOR 11 AUT GER SWI 12 FRA SPA 13 ITA

Remarks: status disputed, probably only a boreoalpine ecovariety

Synonyms: hidden

= Dactylorchis alpestris (Pugsley) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza alpestris (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(9): 1164 (1983)

= Dactylorhiza fistulosa subsp. alpestris (Pugsley) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 20(3): 628 (1988), nom. illeg.

= Dactylorhiza latifolia subsp. alpestris (Pugsley) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza latifolia var. pumila (Freyn ex M.Schultze) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza majalis subsp. alpestris (Pugsley) Senghas; Jahresber. Naturwiss. Ver. Wuppertal 21-22: 51 (1968)

= Dactylorhiza majalis var. pumila (Freyn ex M.Schultze) Landwehr; Orchideeen 37(3): 80 (1975)

= Dactylorhiza majalis var. pumila (Freyn ex M.Schulze) H.Baumann, Künkele & R.Lorenz; J. Eur. Orch. 34: 151 (2002), isonym

= Orchis alpestris Pugsley; Proc. Linn. Soc. Lond. Sess. 146, 98 (1934), in clavi, et J. Linn. Soc., Lond., Bot. 49: 587 (1935) [descr. ampl.]

= Orchis impudica Crantz; Stirp. Austr. ed. II. fasc. vi. 497 (1769), nom. illeg.

= Orchis latifolia f. pumila Freyn ex M.Schulze; Mitth. Thüring. Bot. Vereins, n.f., 17: 51 (1902)

= Orchis latifolia subsp. alpestris (Pugsley) Janch.; 72 (1950)

= Orchis latifolia subsp. impudica (Crantz) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis latifolia var. macrantha Zapal.; Consp. Fl. Gallic. Crit. 210 (1906)

= Orchis latifolia var. pumila (Freyn ex M.Schultze) W.Zimmerm.; Formen Orch. Deutschl., Deutsch-Österr. & Schweiz 36 (1912)

= Orchis latifolius subsp. impudicus (Crantz) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis latifolius var. impudicus (Crantz) Asch. & Graebn.; Syn. Mitteleur. Fl. 3(6): 736 (1907)

= Orchis majalis subsp. alpestris (Pugsley) Hellm.; Boissiera 7: 382 (1943)

variant :

var. brevifolia (Rchb.f.) Kreutz; Kompend. Eur. Orchid. 50 (2004)

Distribution: Europe

Synonyms: hidden

= Dactylorchis cruenta var. brevifolia (Rchb.f.) Verm.; Stud. Dactylorchis 159 (1947)

= Dactylorhiza cruenta var. brevifolia (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza: 4 (1962)

= Dactylorhiza majalis var. gracilis (Warnst.) Landwehr; 79 (1975)

= Orchis angustifolia var. brevifolia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis cruenta f. brevifolia (Rchb.f.) Neuman; 631 (1901)

= Orchis latifolia var. angustifolia W.Becker; 68 (1898), nom. illeg.

= Orchis latifolia var. brevifolia Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 58 (1851)

subspecies :

subsp. baltica (Klinge) Senghas; Jahresber. Naturwiss. Ver. Wuppertal 21/22: 126 (1968)

Name: Baltisches Knabenkraut

Distribution: Denmark; Finland; Poland; Estonia; Latvia; Lithuania; Belarus; ?N-Ukraine; Siberia (Altai, Irkutsk, Krasnoyarsk, W-Siberia); Kazakhstan; ?China (?Xinjiang); European Russia

10 DEN FIN 11 POL 14 BLR EST LAT LIT RUS 30 SIB 32 KAZ

Synonyms: hidden

= Dactylorchis baltica (Klinge) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis longifolia (Neuman) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis traunsteineri var. dunensis (Rchb.f.) Verm.; Ned. Kruidk. Arch. 56: 226 (1949)

= Dactylorhiza baltica (Klinge) N.I.Orlova; Min., Konsp. Fl. Pskovsk. Obl. 57 (1970), isonym

= Dactylorhiza baltica (Klinge) N.I.Orlova ex Aver.; Bot. Zhurn. (Moscow & Leningrad) 68: 1163 (1983), isonym

= Dactylorhiza baltica (Klinge) Nevski; Komarov, Fl. URSS 4: 713 (1935)

= Dactylorhiza baltica var. kuzkenembe Pikner; Ber. Arbeitskreis. Heimische Orchid. 28(2): 260 (2011), et 29(1): 278 (2012)

= Dactylorhiza incarnata f. dunensis (Rchb.f.) P.Delforge; Naturalistes Belges 92(Orchid. 24): 202 (2011), isonym

= Dactylorhiza latifolia subsp. baltica (Klinge) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962) [nom. inval.]

= Dactylorhiza latifolia var. longifolia (Neuman) Soó; Nom. Nov. Gen. Dactylorhiza 5 (1962)

= Dactylorhiza longifolia (Neuman) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza longifolia (Neuman) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza majalis subsp. baltica (Klinge) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 91 (1976), isonym

= Dactylorhiza majalis subsp. baltica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980), isonym

= Dactylorhiza traunsteineri var. dunensis (Rchb.f.) Soó; Nom. Nov. Gen. Dactylorhiza: 6 (1962)

= Orchis baltica (Klinge) A.Fuchs; Ber. Naturwiss. Ver. Schwaben Neuburg 42: 42 (1919), isonym

= Orchis baltica (Klinge) Nevski; Acta Hort. Petrop. 17 1: 153, in clavi, et 170 (1898)

= Orchis lapponica subsp. dunensis (Rchb.f.) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis latifolia f. baltica (Klinge) M.Schulze; Mitth. Thüring. Bot. Vereins, n.f., 17: 51 (1902)

= Orchis latifolia subsp. baltica Klinge; Dactylorch. Monogr. Prodr. 7, 24 (1898), et ex Lehmann, Fl. Pol. Livl. 188 (1895)

= Orchis latifolia subsp. dunensis (Rchb.f.) Krysht.; 420 (1929)

= Orchis latifolia var. dunensis Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 59 (1851)

= Orchis latifolia var. tenuior Neuman; Bot. Not. 1909: 240 (1909)

= Orchis longifolia Neuman; Bot. Not. 1909: 241 (1909)

= Orchis longifolia var. gracilis Neuman; Bot. Not. 1909: 241 (1909)

subspecies :

subsp. brennensis (E.Nelson) D.Tyteca & M.Hedrèn; J. Eur. Orch. 52(1): 59 (2020)

Distribution: France

12 FRA

Synonyms: hidden

= Dactylorhiza brennensis (E.Nelson) D.Tyteca & Gathoye; Orchidophile (Deuil-la-Barre) 81: 58 (1988)

= Dactylorhiza elata subsp. brennensis E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 50 (1976)

= Dactylorhiza elata var. brennensis (E.Nelson) P.Delforge; Naturalistes Belges 81(4): 397 (2000)

subspecies :

subsp. elatior (Fr.) M.Hedrén & H.A.Pedersen; Nordic J. Bot. 30: 271 (2012)

Distribution: Sweden; Estonia (Saaremaa Isl.)

10 SWE 14 EST

Remarks: status and relationship to subsp. baltica uncertain

Synonyms: hidden

= Dactylorhiza elatior (Fr.) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Dactylorhiza osiliensis Pikner; Naturalistes Belges 86(Orchid. 18): 77 (2005)

= Dactylorhiza praetermissa subsp. osiliensis (Pikner) Kreutz; Orchidee (Hamburg) 57(1): 99 (2006)

= Dactylorhiza ruthei subsp. osiliensis (Pikner) Eccarius; Orchideengattung Dactylorhiza: 490 (2016)

= Orchis angustifolia subsp. elata (Hartm.) C.Hartm.; Handb. Skand. Fl., ed. 5: 225 (1849)

= Orchis angustifolia var. elata Hartm.; Handb. Skand. Fl. ed. 4: 281 (1843)

= Orchis elatior (Fr.) Afzel. ex Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 51 (1851)

= Orchis latifolia subsp. elatior Fr.; Summa Veg. Scand.: 217 (1845)

= Orchis strictifolia var. elata (Hartm.) Hyl.; Uppsala Univ. Årsskr. 7: 117 (1945)

subspecies :

subsp. parvimajalis (D.Tyteca & Gathoye) Kreutz; Kompend. Eur. Orchid. 50 (2004)

Distribution: France

12 FRA

Synonyms: hidden

= Dactylorhiza parvimajalis D.Tyteca & Gathoye; J. Eur. Orchid. 32 (3/4): 471-511 (2000)

species :

Dactylorhiza maurusia (Em. & Maire) Holub; Folia Geobot. Phytotax. 8(2): 176 (1973)

Distribution: N-Morocco (Rif Mts.); Algeria (Kabylie)

20 ALG MOR

Synonyms: hidden

= Dactylorchis maurusia (Emb. & Maire) Verm.; Stud. Dactylorch. 69 (1947)

= Dactylorhiza battandieri Raynaud; Orchid. Maroc 62 (1985)

= Dactylorhiza maculata subsp. baborica (Maire & Weiller) Soó; Acta Bot. Acad. Sci. Hung. 16 (3-4): 368 (1970 publ. 1971)

= Dactylorhiza maculata subsp. battandieri (Raynaud) H.Baumann & Künkele; Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 20: 631 (1988)

= Dactylorhiza maculata subsp. maurusia (Emb. & Maire) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza maurusia (Emb. & Maire) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza maurusia (Emb. & Maire) Holub; Folia Geobot. Phytotax. 8: 176 (1973)

= Dactylorhiza maurusia (Emb. & Maire) Raynaud; Orchid. Maroc 59 (1985), isonym

= Orchis incarnata var. xauensis Pau & Font Quer; Font Quer, Exsicc. (Iter Marocc.) 1928: no. 56 (1929)

= Orchis latifolia subsp. maurusia (Emb. & Maire) Maire; Cat. Pl. Maroc 1: 151 (1931)

= Orchis maculata subsp. baborica Maire & Weiller; Fl. Afrique N. 6: 309 (1959 publ. 1960), no Latin descr.

= Orchis maculata subsp. maurusia (Emb. & Maire) Maire & Weiller; Fl. Afrique N. 6: 309 (1960)

= Orchis maurusia Emb. & Maire; Pl. Marocc. Nov. (Arch. Sci. Maroc), Fasc. III. 7 (1930), et Bull. Soc. Sci. Nat. Maroc 11: Nos. 4-6, 109 (1931)

species :

Dactylorhiza nieschalkiorum H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(2): 259 (1981)

Name: Nieschalks Knabenkraut

Distribution: NE-Turkey

34 TUR

Remarks: praetermissa agg. Status doubtful, possibly in Anatolia the same as D. osmanica

Conservation Status: IUCN R

Synonyms: hidden

= Dactylorhiza majalis subsp. nieschalkiorum (H.Baumann & Künkele) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 159 (2019)

species :

Dactylorhiza osmanica (Klinge) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 3: 341, 355 (1960)

Name: Osmanisches Knabenkraut

Distribution: Siberia (Altai, Buryatia, Chita, Tuva); Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Transcaucasus; Turkey; Iran; Lebanon; Syria

30 SIB 32 KGZ TKM TZK UZB 33 NCS 34 IRN LEB SYR TUR

Remarks: incarnata agg.

Synonyms: hidden

= Dactylorhiza cataonica (H.Fleischm.) Holub; Preslia 36: 252 (1964)

= Dactylorhiza caucasica var. cataonica (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

= Dactylorhiza cilicica (Klinge) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965), isonym

= Dactylorhiza cilicica (Klinge) Soó; Ann. Univ. Sci. Budapest. Rolando Eotvos, Sect. Biol. 3: 340 (1960), et Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza elata subsp. anatolica E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 45 (1976), et Taxon 28 (5-6): 593 (1979)

= Dactylorhiza euxina var. cataonica (H.Fleischm.) Soó; [nom. inval.]

= Dactylorhiza incarnata subsp. cilicica (Klinge) H.Sund.; Europ. Medit. Orchideen 45 (1975)

= Dactylorhiza maculata subsp. osmanica (Klinge) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 98 (1976)

= Dactylorhiza maculata subsp. osmanica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 41 (1980)

= Dactylorhiza olocheilos (Boiss.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza osmanica (Klinge) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965), isonym

= Dactylorhiza osmanica subsp. anatolica (E.Nelson) Eccarius; Orchideengattung Dactylorhiza 232 (2016)

= Dactylorhiza osmanica var. anatolica (E.Nelson) Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41: 271 (1983)

= Orchis cataonica H.Fleischm.; Ann. K. K. Naturhist. Hofmus. 28: 34 (1914)

= Orchis cilicica (Klinge) Schltr.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 1: 178 (1927)

= Orchis cilicica Klinge; Acta Hort. Petrop. 17 1: 156, in clavi, et 187 (1898)

= Orchis holocheilos (Boiss.) G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 228 (1933)

= Orchis incarnata var. latifolia Freyn; Bull. Herb. Boissier, sér. 2, 1: 285 (1901), nom. illeg.

= Orchis incarnata var. olocheilos Boiss.; Fl. Orient. 5: 71 (1882)

= Orchis incarnata var. stenophylla Freyn; Bull. Herb. Boissier, sér. 2, 1: 285 (1901)

= Orchis olocheilos (Boiss.) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis olocheilos var. osmanica (Klinge) Soó; Bot. Arch. 23: 73 (1928)

= Orchis orientalis subsp. cilicica Klinge; Acta Horti Petrop. 17(1): 36, 37 (1898)

= Orchis orientalis subsp. osmanica Klinge; Acta Horti Petrop. 17(1): 157, 188 (1898)

= Orchis osmanica (Klinge) A.W.Hill; Index Kew. Suppl. 9: 194 (1938), isonym

= Orchis osmanica (Klinge) Lipsky; Acta Hort. Petrop. 17 1: 157, in clavi, et 188 (1898)

species :

Dactylorhiza praetermissa (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

subspecies :

subsp. praetermissa

Name: Übersehene Fingerwurz, Übersehenes Fingerknabenkraut (DE)

Distribution: England; Wales; Denmark; Norway; Netherlands; Belgium; Luxembourg; Germany (Hessen [I], Hamburg, Mecklenburg-Vorpommern [I], Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland); France; Channel Isl.; Estonia

10 DEN GRB NOR 11 BGM GER LUX NET 12 CIS FRA 14 EST

Remarks: There is considerable and unresolved debate about whether the name integrata really does apply to the taxon formerly known as praetermissa. See discussion in Bateman & Denholm 2012

Synonyms: hidden

= Dactylorchis praetermissa (Druce) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorchis praetermissa var. cuspidata Deinum ex Verm.; Ned. Kruidk. Arch. 56: 230 (1949)

= Dactylorchis praetermissa var. elata Verm.; Ned. Kruidk. Arch. 56: 229 (1949)

= Dactylorchis praetermissa var. longifolia Verm.; Ned. Kruidk. Arch. 56: 230 (1949)

= Dactylorchis praetermissa var. macrantha (Sipkes) Verm.; Ned. Kruidk. Arch. 56: 229 (1949)

= Dactylorchis praetermissa var. parvifructa Verm.; Ned. Kruidk. Arch. 56: 231 (1949)

= Dactylorchis praetermissa var. tenera Verm.; Ned. Kruidk. Arch. 56: 230 (1949)

= Dactylorchis praetermissa var. triangularis Verm.; Ned. Kruidk. Arch. 56: 231 (1949)

= Dactylorhiza incarnata subsp. praetermissa (Druce) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza integrata (E.G.Camus) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza majalis subsp. integrata (E.G.Camus) H.A.Pedersen; J. Eur. Orch. 41(3/4): 504 (2009)

= Dactylorhiza majalis subsp. praetermissa (Druce) D.M.Moore & Soó; J. Linn. Soc., Bot. 76: 367 (1978)

= Dactylorhiza majalis var. integrata (E.G.Camus) H.A.Pedersen; J. Eur. Orch. 43: 80 (2011)

= Dactylorhiza majalis var. macrantha (Sipkes) R.M.Bateman & Denholm; Watsonia 14(4): 372 (1983)

= Dactylorhiza majalis var. praetermissa (Druce) R.M.Bateman & Denholm; Watsonia 14(4): 371 (1983)

= Dactylorhiza praetermissa subsp. integrata (E.G.Camus ex Fourcy) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

= Dactylorhiza praetermissa var. bracteosa Kreutz; J. Eur. Orch. 48: 321 (2016)

= Dactylorhiza praetermissa var. cuspidata (Deinum ex Verm.) Soó; Nom. Nov. Gen. Dactylorhiza: 5 (1962)

= Dactylorhiza praetermissa var. elata (Verm.) Soó; Nom. Nov. Gen. Dactylorhiza: 5 (1962)

= Dactylorhiza praetermissa var. integrata (E.G.Camus ex Fourcy) D.Tyteca & Gathoye; Belg. J. Bot. 126(1): 93 (1993)

= Dactylorhiza praetermissa var. maculosa D.Tyteca & Gathoye; L´Orchidophile 92: 112 (1990)

= Dactylorhiza praetermissa var. robusta Bussche; Liparis 7: 32 (2001)

= Orchis incarnata subsp. integrata (E.G.Camus) E.G.Camus; J. Bot. (Morot) 4: 157 (1892)

= Orchis incarnata var. albiflora Lecoq & Lamotte ex Thielens; Bull. Soc. Roy. Bot. Belgique 12: 82 (1873)

= Orchis incarnata var. integrata E.G.Camus ex Fourcy; Vadem. Herb. Paris, ed. 6: 325 (1891)

= Orchis integrata (E.G.Camus) E.G.Camus; Monogr. Orchid. France: 48 (1894)

= Orchis praetermissa Druce; Rep. Bot. Exch. Cl. Brit. Isles 3: 341 (1913 publ. 1914)

= Orchis praetermissa f. macrantha (Sipkes) Soó; Bot. Arch. 23: 69 (1938)

= Orchis praetermissa subsp. integrata (E.G.Camus) E.G.Camus & A.Camus; Iconogr. Orchid. Europe 1: 219 (1928)

= Orchis praetermissa var. albiflora Druce; Rep. Bot. Soc. Exch. Club Brit. Isles 5: 53 (1919)

= Orchis praetermissa var. macrantha Sipkes; Levende Natuur 26: 52 (1921)

variant :

var. junialis (Verm.) Senghas; Jahresber. Naturwiss. Ver. Wuppertal 21/22: 126 (1968)

Distribution: England; NW-Germany; NE-Canada [I] (Ontario [I], ?Newfoundland [I])

10 GRB 11 GER 72 cdn

Synonyms: hidden

= Dactylorchis praetermissa var. junialis (Verm.) Verm.; Stud. Dactylorchis 67 (1947), nom. inval., et Ned. Kruidk. Arch. 56: 229 (1949)

= Dactylorhiza majalis subsp. pardalina (Pugsley) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 88 (1976) [without basionym page]

= Dactylorhiza majalis var. junialis (Verm.) H.A.Pedersen; Flor. Med. Fra Dansk Herb. I, URT 23(2): 64-71 (1999), isonym

= Dactylorhiza majalis var. junialis (Verm.) R.M.Bateman & Denholm; Watsonia 14: 371 (1983)

= Dactylorhiza praetermissa f. junialis (Verm.) P.D.Sell; Fl. Great Britain & Ireland 5: 365 (1996)

= Dactylorhiza praetermissa subsp. junialis (Verm.) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

= Dactylorhiza praetermissa var. junialis (Verm.) H.A.Pedersen; J. Eur. Orch. 41(3/4): 504 (2009), nom. illeg.

= Dactylorhiza ×pardalina (Pugsley) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 93 (1986)

= Orchis latifolia var. junialis Verm.; Ned. Kruid. Arch. 43: 404, 409 (1933)

= Orchis pardalina Pugsley; Proc. Linn. Soc. Lond. Sess. 146, 97 (1934), in clavi, et J. Linn. Soc., Lond., Bot. 49: 581 (1935) [descr. ampl.]

= Orchis praetermissa subsp. junialis (Verm.) Verm.; Chron. Bot. 4(2): 108 (1934)

subspecies :

subsp. schoenophila R.M.Bateman & Denholm; New J. Bot. 2(1): 47 (2012), nom. nov.

Distribution: S-England

10 GRB

Synonyms: hidden

= Dactylorhiza majalis var. bowmannii Jenk.; (1995)

species :

Dactylorhiza purpurella (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

Name: Purpurblütige Fingerwurz, Purpurblütiges Fingerknabenkraut (DE)

Distribution: Faeroe Isl.; England; S-Sweden; Ireland; Denmark; Norway; Germany [I]

10 DEN FOR GRB IRE NOR SWE 11 ger

Remarks: for the status as full species see Bateman & Denholm 2011

Synonyms: hidden

= Dactylorchis incarnata subsp. pulchella (Druce) Hesl.-Harr.f.; Ber. Geobot. Forsch. Inst. Rübel Zürich 1953: 55 (1954)

= Dactylorchis purpurella (T.Stephenson & T.A.Stephenson) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza incarnata subsp. pulchella (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza incarnata var. obscura (Höppner) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza majalis subsp. purpurella (T.Stephenson & T.A.Stephenson) D.M.Moore & Soó; Bot. J. Linn. Soc. 76(4): 367 (1978)

= Dactylorhiza majalis var. atrata A.J.Richards; Watsonia 18(3): 308 (1991)

= Dactylorhiza majalis var. crassifolia (T.Stephenson & T.A.Stephenson) Landwehr; Orchideeen 37: 80 (1975)

= Dactylorhiza majalis var. pulchella (Druce) R.M.Bateman & Denholm; Watsonia 14: 370 (1983)

= Dactylorhiza majalis var. purpurella (T.Stephenson & T.A.Stephenson) R.M.Bateman & Denholm; Watsonia 14: 370 (1983)

= Dactylorhiza pulchella (Druce) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza purpurella subsp. majaliformis E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 77 (1976), without type, et Taxon 28(56): 593 (1979)

= Dactylorhiza purpurella subsp. majaliformis E.Nelson ex Løjtnant; Bot. Tidsskr. 74: 176 (1979), nom. illeg.

= Dactylorhiza purpurella var. atrata (A.J.Richards) A.J.Richards & L.Lewis; J. Eur. Orch. 50(1): 85 (2018)

= Dactylorhiza purpurella var. crassifolia (T.Stephenson & T.A.Stephenson) Landwehr; 80 (1975)

= Dactylorhiza purpurella var. majaliformis (E.Nelson) Kreutz; Eurorchis 18: 95 (2006)

= Dactylorhiza purpurella var. pulchella (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

= Dactylorhiza vadorum Kreutz & H.Dekker; Gorteria 38: 176 (2016)

= Dactylorhiza vadorum var. picturata Kreutz & H.Dekker; Gorteria 38: 184 (2016)

= Orchis incarnata var. obscura Höppner; Fl. Niederrheins, ed. 3: 106 (1913)

= Orchis incarnata var. pulchella Druce; Bot. Soc. Exch. Club Brit. Isles 5: 167 (1918)

= Orchis latifolia var. pulchella (Druce) Pugsley; J. Linn. Soc., Bot. Bot. 49: 587 (1935)

= Orchis praetermissa var. pulchella Druce; Bot. Soc. Exch. Club Brit. Isles 5: 577 (1920)

= Orchis purpurella T.Stephenson & T.A.Stephenson; J. Bot. 58: 164 (1920)

= Orchis purpurella var. crassifolia T.Stephenson; Rep. botl. Soc. Exch. Club Br. Isl. 11: 356 (1937)

= Orchis purpurella var. pulchella (Druce) Pugsley; Bot. J. Linn. Soc. 49: 583 (1935)

= Orchis strictifolia var. pulchella (Druce) Clapham; Fl. Brit. Isles 1319 (1952)

variant :

var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm; Watsonia 14(4): 368 (1983), et Orchids Brit. Isles 373 (2005)

Distribution: Scotland; Wales; Northern Ireland; Ireland; Denmark

10 DEN GRB IRE

Synonyms: hidden

= Dactylorchis majalis subsp. cambrensis R.H.Roberts; Watsonia 5(1): 41 (1961)

= Dactylorhiza cambrensis (R.H.Roberts) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza comosa subsp. cambrensis (R.H.Roberts) P.D.Sell; Fl. Great Britain & Ireland 5: 365 (1996)

= Dactylorhiza latifolia subsp. cambrensis (R.H.Roberts) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

= Dactylorhiza majalis subsp. cambrensis (R.H.Roberts) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 89 (1976)

= Dactylorhiza majalis var. cambrensis (R.H.Roberts) H.A.Pedersen; J. Eur. Orch. 39(2): 350 (2007), isonym

= Dactylorhiza majalis var. eboracensis (Godfery) R.M.Bateman & Denholm; Watsonia 14: 373 (1983)

= Dactylorhiza purpurella var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm; M. Foley, Orchids Brit. Isles 373 (2005)

= Orchis latifolia var. eboracensis Godfery; Mitt. Geogr. Ges. (Thüringen) Jena 2: 220 (1885)

variant :

var. maculosa (T.Stephenson & T.A.Stephenson) Kreutz, H.Dekker & L.Lewis; J. Eur. Orch. 48(2-4): 304 (2016)

Distribution: Scotland; Netherlands

10 GRB 11 NET

Synonyms: hidden

= Dactylorhiza majalis var. maculosa (T.Stephenson) R.M.Bateman & Denholm; Watsonia 14(4): 370 (1983)

= Orchis purpurella var. maculosa T.Stephenson; Bot. Soc. Exch. Club Brit. Isles 11: 355 (1937)

species :

Dactylorhiza romana (Sebast.) Soó; Nom. Nov. Gen. Dactylorhizae 3 (1962)

subspecies :

subsp. romana

Name: Römisches Knabenkraut (DE); Roman Spotted-orchid (EN)

Distribution: Italy; Sicily; Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (incl. Kyklades); Crete; European Turkey; Crimea; East Aegaean Isl. (Rhodos); Turkey; Cyprus; N-Israel

13 ALB BUL CRO GRC ITA KOS KRI MNT NMA SIC SRB TUE 14 KRY 34 CYP EAI ISR TUR

Remarks: sambucina agg.

Synonyms: hidden

= Dactylorchis romana (Sebast.) Verm.; Stud. Dactylorch.: 65 (1947)

= Dactylorhiza sambucina f. zimmermannii (A.Camus) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza sambucina subsp. pseudosambucina (Ten.) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza sambucina subsp. romana (Sebast.) Bornm.; Beibl. Bot. Jahrb. Syst. 140: 126 (1928)

= Dactylorhiza sulphurea (Spreng.) Franco; J. Linn. Soc., Bot. 76: 336 (1978)

= Dactylorhiza sulphurea subsp. pseudosambucina (Ten.) Franco; Bot. J. Linn. Soc. 76(4): 366 (1978)

= Orchis bracteata Ten.; Fl. Napol. 1(Prodr.): LII (1812), nom. illeg.

= Orchis fasciculata var. obtusifolia Tineo; Guss., Fl. Sicul. Syn. 2: 875 (1845)

= Orchis lucana Spreng.; Pl. Min. Cogn. Pug. 2: 79 (1815)

= Orchis mediterranea Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 163 (1898), nom. illeg.

= Orchis mediterranea subsp. pseudosambucina (Ten.) KIinge; Acta Horti Petrop. 17(1): 151, 164 (1898)

= Orchis natalis Tineo; Pl. Rar. Sicil. 8 (1846)

= Orchis ochroleuca Schur; Enum. Pl. Transsilv.: 641 (1866), nom. illeg.

= Orchis pseudosambucina subsp. romana (Sebast.) Gand.; Nov. Consp. Fl. Eur. 462 (1910)

= Orchis pseudosambucina Ten.; Fl. Napol. 2: 284 (1820)

= Orchis pseudosambucina var. natalis (Tineo) Nyman; Consp. Fl. Eur. 693 (1882)

= Orchis romana lusus bicolor G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A 2: 206 (1933)

= Orchis romana Sebast.; Roman. Pl. Fasc. Prim. 1: 12 (1813)

= Orchis romana var. incarnata E.G.Camus & A.Camus; Iconogr. Orchid. Europe 214 (1928)

= Orchis romana var. lutea E.G.Camus & A.Camus; Iconogr. Orchid. Europe 214 (1928)

= Orchis sambucina Brot.; Fl. Lusit. 1: 21 (1800 publ. 1804)

= Orchis sambucina var. zimmermannii A.Camus; Camus, Iconogr. Orchid. Europe: 212 (1928)

= Orchis sulphurea Spreng.; Syst. 3: 688 (1826) [in syn.]

subspecies :

subsp. georgica (Klotzsch) Soó ex Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 271 (1983)

Name: Georgisches Knabenkraut

Distribution: Turkmenistan; Northern Caucasus; Georgia [Caucasus] (Racha-Lechkhumi, Adzharia, Kartli, Kakheti, Trialeti); Armenia; Azerbaijan (incl. Talysh); E-Turkey; N-Iran

32 TKM 33 ARM AZE GEO NCS 34 IRN TUR

Synonyms: hidden

= Dactylorchis flavescens (K.Koch) Verm.; Stud. Dactylorch. 65 (1947)

= Dactylorhiza flavescens (K.Koch) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(2): 237 (1981), isonym

= Dactylorhiza flavescens (K.Koch) Holub; Folia Geobot. Phytotax. 11(1): 83 (1976)

= Dactylorhiza flavescens (Koch) É.M.Seifulin; Czerep., Opred. khvoshcheobraznykh. rastenii Turkmenistana 278 (1978) [isonym, isonym]

= Dactylorhiza ruprechtii Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(4): 537 (1983)

= Dactylorhiza sambucina subsp. georgica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza sulphurea subsp. georgica (Klinge) Herrero & Ó.Sánchez; Doc. Jard. Bot. Atlántico 7: 80 (2010)

= Orchis flavescens K.Koch; Linnaea 22: 281 (1850)

= Orchis georgica (Klinge) Lipsky; Acta Hort. Petrop. 17 1: 151, in clavi, et 166 (1898)

= Orchis georgica (Klinge) Medw.; Act. Hort. Tiflis 18: 473 (1919), in clavi

= Orchis mediterranea subsp. georgica Klinge; Trudy Imp. S.-Petersb. Bot. Sada 17(1): 19 (1898)

= Orchis pseudosambucina subsp. georgica Klinge; Trudy Imp. S.-Petersb. Bot. Sada 17(1): 151 (1898)

= Orchis romana lusus flavescens (K.Koch) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis romana lusus tenuifolia Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis romana subsp. georgica (Klinge) E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 172 (1908)

= Orchis romana var. georgica (Klinge) Schltr.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A 1: 187 (1927)

= Orchis tenuifolia K.Koch; Linnaea 22: 281 (1850)

subspecies :

subsp. guimaraesii (E.G.Camus) H.A.Pedersen; Bot. J. Linn. Soc. 152(4): 426 (2006)

Distribution: Portugal; Spain; Morocco; Algeria

12 POR SPA 20 ALG MOR

Synonyms: hidden

= Dactylorhiza guimaraesii (E.G.Camus) P.Delforge; Naturalistes Belges 96(Orchid. 28): 15 (2015)

= Orchis guimaraesii (E.G.Camus) Rivas Goday; Farmacognosia 4(6): 193 (1945)

= Orchis romana lusus sulphurea (Link) Soó; Bot. Arch. 23: 63 (1929)

= Orchis romana var. guimaraesii E.G.Camus; Iconogr. Orchid. Europe 1: 216 (1928)

subspecies :

subsp. libanotica (Mouterde) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 154 (2007)

Distribution: Lebanon

34 LEB

Synonyms: hidden

= Dactylorhiza libanotica (Mouterde) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 876 (1984)

= Orchis romana subsp. libanotica Mouterde; Nouv. Fl. Liban Syrie 1: 342 (1966)

subspecies :

subsp. markusii (Tineo) Holub; Folia Geobot. Phytotax. 19(2): 214 (1984)

Name: Sizilianisches Knabenkraut

Distribution: Baleares; Sardinia; Italy; Sicily

12 BAL SAR 13 ITA SIC

Synonyms: hidden

= Dactylorhiza markusii (Tineo) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(4): 461 (1981)

= Dactylorhiza romana subsp. siciliensis (Klinge) Soó; Nom. Nov. Gen. Dactylorhizae 3 (1962)

= Dactylorhiza romana var. markusii (Tineo) Soó; Nom. Nov. Gen. Dactylorhizae 3 (1962)

= Dactylorhiza romana var. sicula (Tineo) Soó; Nom. Nov. Gen. Dactylorhiza 3 (1962)

= Dactylorhiza sambucina subsp. siciliensis (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza siciliensis (Klinge) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 105 (1976) [without basionym page]

= Dactylorhiza sicula (Tineo ex Rchb.f.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza sulphurea (Link) Franco; Bot. J. Linn. Soc. 76: 366 (1978), nom. inval.

= Dactylorhiza sulphurea subsp. siciliensis (Klinge) Franco; J. Linn. Soc., Bot. 76(4): 367 (1978)

= Orchis markusii Tineo; Pl. Rar. Sicil. 9 (1846)

= Orchis mediterranea subsp. siciliensis Klinge; Acta Horti Petrop. 17(1): 165, 166 (1898)

= Orchis pseudosambucina subsp. markusii (Tineo) Gand.; Nov. Consp. Fl. Eur. 462 (1910), isonym

= Orchis pseudosambucina subsp. markusii (Tineo) Nyman; Consp. Fl. Eur. 4: 693 (1882)

= Orchis pseudosambucina subsp. sicula (Tineo) K.Richt.; Pl. Eur. 1: 271 (1890)

= Orchis pseudosambucina subsp. sulphurea Link; Nov. Consp. Fl. Eur. 462 (1910)

= Orchis romana f. markusii (Tineo) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis romana var. markusii (Tineo) E.G.Camus, Bergon & A.Camus; Monogr. Orchid.: 171 (1908)

= Orchis romana var. markusii (Tineo) Rivas Goday; Farmacognosia 4(6): 194, 195 (1945), isonym

= Orchis siciliensis (Klinge) A.W.Hill; Index Kew. Suppl. 9: 194 (1938)

= Orchis sicula Tineo; Pl. Rar. Sicil. 8 (1846)

= Orchis sulphurea Link; J. Bot. (Schrader) 1806: 1 (1806), et Nomencl. Bot. 1: 225 (1821)

= Orchis sulphurea var. markusii (Tineo) Maire; Jahand. et al., Cat. Pl. Maroc 1: 151 (1931)

species :

Dactylorhiza ruthei (M.Schultze ex R.Ruthe) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

Name: Ruthe-Fingerwurz, Ruthe-Fingerknabenkraut (DE)

Distribution: NE-Germany (Mecklenburg-Vorpommern); +NW-Poland

11 GER POL

Remarks: possibly a subspecies of praetermissa (integrata). Populations in Estonia have been separated as D. vironii

Synonyms: hidden

= Dactylorchis ruthei (M.Schultze ex R.Ruthe) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza majalis subsp. ruthei (M.Schultze ex R.Ruthe) H.Kretzschmar; Orchideen Deutschlands [Kretzschmar] 16 (2008)

= Orchis maculata subsp. ruthei (M.Schulze ex R.Ruthe) Hegi; 354 (1909)

= Orchis ruthei M.Schultze ex R.Ruthe; Deutsch. Bot. Monatsschr. 15: 237 (1897)

species :

Dactylorhiza saccifera (Brongn.) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962)

subspecies :

subsp. saccifera

Name: Langähriges Knabenkraut

Distribution: Corsica; Sardinia; Italy; Sicily; Croatia; Bosnia & Hercegovina; Albania; Romania; Bulgaria; Greece; European Turkey; Turkey; Lebanon

12 COR SAR 13 ALB BIH BUL CRO GRC ITA ROM SIC TUE 34 LEB TUR

Remarks: maculata agg.

Synonyms: hidden

= Dactylorchis maculata subsp. saccifera (Brongn.) Verm.; Stud. Dactylorchis 68 (1947)

= Dactylorchis maculata var. heuffelii Borsos & Soó; Acta Bot. Acad. Sci. Hung. 5: 324 (1959)

= Dactylorchis saccifera (Brongn.) Verm.; Fl. Neerl. 1: Afl. 5, 87 (1958)

= Dactylorhiza fuchsii var. nesogenes (Briq.) Soó; Nom. Nov. Gen. Dactylorhiza: 8 (1962)

= Dactylorhiza maculata subsp. saccifera (Brongn.) Diklic; Fl. SR Srbije 8: 77 (1976)

= Dactylorhiza maculata subsp. saccifera (Brongn.) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza maculata var. orophila (Briq.) Soó; Nom. Nov. Gen. Dactylorhiza: 7 (1962)

= Dactylorhiza saccifera subsp. macrostachys (Tineo) Niketic & Djordjevic; Bull. Nat. Hist. Mus. Belgrade 11: 104 (2018)

= Dactylorhiza saccifera var. heuffelii (Borsos & Soó) Soó; Nom. Nov. Gen. Dactylorhiza: 8 (1962)

= Dactylorhiza saccifera var. lanceolata Landwehr; Wilde Orchid. Eur. 2: 556 (1977)

= Orchis basilica subsp. saccifera (Brongn.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 157 (1898)

= Orchis bonanniana Tod.; Orchid. Sicul.: 57 (1842)

= Orchis comosa Schur; Enum. Pl. Transs. 641 (1866)

= Orchis macrostachys Tineo; Pl. Rar. Sicil. 7 (1846)

= Orchis maculata proles macrostachys (Tineo) Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 748 (1907)

= Orchis maculata proles nesogenes (Briq.) Rouy; Rouy & Foucaud, Fl. France 13: 154 (1912)

= Orchis maculata subsp. macrostachys (Tineo) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis maculata subsp. saccifera (Brongn.) E.G.Camus; Icon. Orchid. Europe, Text(1): 242 (1928)

= Orchis maculata subsp. saccifera (Brongn.) K.Richt.; Pl. Eur. 1: 272 (1890)

= Orchis maculata var. bonanniana (Tod.) Nyman; Consp. Fl. Eur.: 693 (1882)

= Orchis maculata var. cartalinoides Klinge ex H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 176 (1908 publ. 1909)

= Orchis maculata var. heuffelii (Borsos & Soó) Pauca; Fl. Republ. Social. Roman. 12: 698 (1972)

= Orchis maculata var. kortiathica Formánek; Verh. Naturf. Vereins Brünn 32: 26 (1894)

= Orchis maculata var. nesogenes Briq.; Prodr. Fl. Corse 1: 371 (1910)

= Orchis maculata var. orophila Briq.; Prodr. Fl. Corse 1: 370 (1912)

= Orchis maculata var. pyramidata Schur; Enum. Pl. Transsilv.: 643 (1866)

= Orchis maculata var. saccifera (Brongn.) Rchb.f.; Icon. Fl. Germ. Helv. 13/14: 67 (1851)

= Orchis maculata var. turfosa Schur; Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4: 72 (1853)

= Orchis nesogenes (Briq.) Rouy; G. Rouy & J. Foucaud, Fl. France 13: 154 (1912)

= Orchis saccifera Brongn.; Bory, Exp. Sci. Morée, Bot.: 259 (1832)

subspecies :

subsp. taurica Kreutz & Isler; Res. Türkiye Fl. 3a: 135 (2022)

Distribution: Turkey

34 TUR

Remarks: provisionally accepted name

species :

Dactylorhiza sajanensis Stepanov; Red Book Krasnoyarsk. Territ. [ed. 3] 304 (2022)

Distribution: Siberia (Krasnoyarsk)

30 SIB

Remarks: provisionally accepted name

species :

Dactylorhiza salina (Turcz. ex Lindl.) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

Distribution: Siberia (Altai, Buryatia, Chita, Krasnoyarsk); Russian Far East (Amur); Kazakhstan; Kyrgyzstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Mongolia; Pakistan (Karakorum)

30 SIB 31 RFE 32 KAZ KGZ 33 ARM GEO NCS 37 MON 40 PAK

Synonyms: hidden

= Dactylorchis salina (Turcz. ex Lindl.) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorhiza cruenta subsp. salina (Turcz. ex Lindl.) E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 70 (1976)

= Dactylorhiza knorringiana (Kraenzl.) Dickoré; Stapfia 39: 221 (1995) isonym

= Dactylorhiza knorringiana (Kraenzl.) Ikonn.; Novit. Syst. Pl. Vasc, Acad. Sci. URSS 6: 267 (1970)

= Dactylorhiza umbrosa subsp. knorringiana (Kraenzl.) Soó; Ann. Univ. Sci. Budapest. Rolando Eotvos, Sect. Biol. 11: 68 (1969)

= Dactylorhiza umbrosa var. knorringiana (Kraenzl.) Soó; Ann. Univ. Sci. Budapest Rolando Eötvös, Sect. Biol. 5: 4 (1962)

= Orchis incarnata var. knorringiana Kraenzl.; Repert. Spec. Nov. Regni Veg. 65: 34 (1931)

= Orchis knorringiana (Kraenzl.) Czerniak.; Fl. Kirgizsk. SSR 3: 138 (1951)

= Orchis orientalis subsp. salina (Turcz. ex Lindl.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 185 (1898), nom. superfl.

= Orchis salina Turcz. ex Lindl.; Gen. Sp. Orchid. Pl. 259 (1835)

species :

Dactylorhiza sambucina (L.) Soó; Nom. Nov. Gen. Dactylorhizae 3 (1962)

Name: Holunder-Knabenkraut

Distribution: Denmark; Norway; Sweden; Finland; Germany (Brandenburg, Baden-Württemberg, Bayern, +Hessen, Rheinland-Pfalz, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Corsica; Sardinia; Italy; Sicily; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece; Estonia; +Belarus; Ukraine; European Russia (W-European Russia, C-European Russia)

10 DEN FIN NOR SWE 11 AUT CZE GER HUN POL SLO SWI 12 ADO COR FRA POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SIC SLV SRB 14 BLR EST RUS UKR

Remarks: sambucina agg.

Synonyms: hidden

= Dactylorchis sambucina (L.) Verm.; Stud. Dactylorch. 65 (1947)

= Dactylorhiza fasciculata (Tineo) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984), isonym

= Dactylorhiza fasciculata (Tineo) H.Baumann & Künkele; Mitt. Arbeitskreis Heimische Orchid. Baden-Württemberg 13: 470 (1981)

= Dactylorhiza latifolia f. rubra (Winterl) D.Tyteca & Gathoye; Orchidophile (Deuil-la-Barre) 92: 112 (1990)

= Dactylorhiza sambucina lusus incarnata (Wahlenb.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza sambucina f. chusae C.E.Hermos.; Estud. Mus. Cienc. Nat. Álava 15: 190 (2000)

= Dactylorhiza sambucina f. hybrida (W.Zimm.) Kümpel; Ber. Arbeitskreis. Heimische Orchid. 18: 27 (2001 publ. 2002)

= Dactylorhiza sambucina f. robusta (Neuman) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza sambucina f. rubra (Winterl) Hyl.; Nord. Kärlväxtfl. 2: 387 (1966)

= Dactylorhiza sambucina f. rubrobracteata (Harz) Soó; Acta Bot. Acad. Sci. Hung. 16: 368 (1970 publ. 1971)

= Dactylorhiza sambucina subsp. laurentina (R.Bolós ex Vayr.) Malag.; Acta Phytotax. Barcinon. 1: 64 (1968)

= Dactylorhiza sambucina var. hungarica (Soó) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Dactylorhiza sambucina var. hybrida (W.Zimm.) Peitz; Jahresber. Naturwiss. Vereins Wuppertal 25: 178 (1972)

= Dactylorhiza sambucina var. laurentina (R.Bolòs ex Vayr.) Soó; Nom. Nov. Gen. Dactylorhiza: 3 (1962)

= Orchis fasciculata Tineo; Guss., Fl. Sicul. Syn. 2: 875 (1845)

= Orchis guffroyi P.Fourn.; Bull. Soc. Bot. France 78: 434 (1931)

= Orchis incarnata var. sambucina (L.) Lapeyr. ex Bubani; Fl. Pyren. 1: 125 (1897)

= Orchis incarnata Willd.; Sp. Pl. ed. 4. 4: 80 (1805), nom. illeg.

= Orchis laurentina R.Bolòs ex Vayr.; Anales Soc. Esp. Hist. Nat. 9: 96, t. 1 (1880)

= Orchis lutea Dulac; Fl. Hautes-Pyr. 125 (1867), nom. illeg.

= Orchis maculata subsp. sambucina (L.) Bonnier & Layens; Tabl. Syn. Pl. Vasc. France 1: 311 (1894)

= Orchis mixta var. sambucina (L.) Retz.; 167 (1779), nom. illeg.

= Orchis pallens Moritzi; Fl. Schweiz 508 (1839), Puccin. Syn. Fl. Luc. 476

= Orchis pseudosambucina var. fasciculata (Tineo) Rchb.f.; Icon. Fl. Germ. Helv. 13-14: 176 (1851)

= Orchis saccata Rchb.; Fl. Germ. Excurs.: 122 (1830), nom. illeg.

= Orchis salina Fronius; Verh. Siebenb. Ver. Naturw. 8: 102 (1857)

= Orchis sambucina lusus hybrida W.Zimm.; Allg. Bot. Z. Syst. 22: 129 (1916)

= Orchis sambucina lusus incarnata (Wahlenb.) Soó; Bot. Arch. 23: 62 (1928)

= Orchis sambucina lusus lutea W.Zimm.; Form. Orch. Deut.: 38 (1912)

= Orchis sambucina lusus luteopurpurea (Rouy) G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A 2: 203 (1933)

= Orchis sambucina lusus rubra (Winterl) Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A 2: 203 (1933)

= Orchis sambucina f. bracteata M.Schulze; Mitt. Geogr. Ges. (Thüringen) Jena 6(Gesamtthüringen): 26 (1889)

= Orchis sambucina f. hungarica Soó; Repert. Spec. Nov. Regni Veg. 24: 29 (1927)

= Orchis sambucina f. lanceolata W.Zimm.; Allg. Bot. Z. Syst. 22: 129 (1916)

= Orchis sambucina f. laurentina (R.Bolòs ex Vayr.) Soó; Repert. Spec. Nov. Regni Veg. 24: 29 (1927)

= Orchis sambucina f. obovata W.Zimm.; Allg. Bot. Z. Syst. 22: 129 (1916)

= Orchis sambucina f. purpurea (W.D.J.Koch) Neuman; Sver. Fl.: 629 (1901)

= Orchis sambucina f. rubrobracteata Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 271 (1895 publ. 1896)

= Orchis sambucina f. tripartita A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 1921: 177 (1921)

= Orchis sambucina L.; Fl. Suec. 312 (1755)

= Orchis sambucina unranked laurentina (R.Bolòs ex Vayr.) Willk.; Suppl. Prodr. Fl. Hispan.: 42 (1893)

= Orchis sambucina subsp. asturica (Gand.) Gand.; Nov. Consp. Fl. Eur.: 462 (1910), nom. inval.

= Orchis sambucina subsp. incarnata (Willd.) Ces.in Car.Catt.; 315 (1844), nom. inval.

= Orchis sambucina subsp. laurentina (R.Bolòs ex Vayr.) Malag.; Acta Phytotax. Barcinon. 1: 64 (1969), nom. superfl.

= Orchis sambucina subsp. laurentina (R.Bolòs ex Vayr.) Willk.; Suppl. Prodr. Fl. Hispan.: 42 (1893), nom. alt.

= Orchis sambucina subvar. luteopurpurea Rouy; Rouy & Foucaud, Fl. France 13: 155 (1912)

= Orchis sambucina subvar. purpurea (W.D.J.Koch) Rouy; Rouy & Foucaud, Fl. France 13: 155 (1912)

= Orchis sambucina var. barlae Rouy; Fl. France 13: 155 (1912)

= Orchis sambucina var. bracteata (M.Schulze) Harz; Schltdl., Fl. Deutschl. ed. 5., Neuer Abdruck, 4: 271 (1895 publ. 1896)

= Orchis sambucina var. candida St.-Lag.; Cariot, Étude Fl., éd. 8, 2: 806 (1889)

= Orchis sambucina var. hungarica (Soó) A.Camus; Camus, Iconogr. Orchid. Europe: 212 (1928)

= Orchis sambucina var. incarnata (Willd.) Amo; Fl. Faner. Penins. Iberica 1: 455 (1871)

= Orchis sambucina var. incarnata (Willd.) Mayrhofer; 23 (1832), isonym

= Orchis sambucina var. incarnata (Willd.) Rchb.; XIII (1833), isonym

= Orchis sambucina var. incarnata (Willd.) Wahlenb.; Fl. Suec. 2: 554 (1826)

= Orchis sambucina var. laurentina (R.Bolòs ex Vayr.) Willk.; Prodr. Fl. Hispan. 42: 356 (1893)

= Orchis sambucina var. lutea E.G.Camus; Camus, Iconogr. Orchid. Europe: 212 (1928), nom. illeg.

= Orchis sambucina var. pallide-flava Schur; 295 (1870), nom. inval.

= Orchis sambucina var. purpurascens R.Hinterh. & J.Hinterh.; Prodr. Fl. Salzburg 192 (1851)

= Orchis sambucina var. purpurea W.D.J.Koch; Syn. Fl. Germ. Helv. ed. 2: 972 (1845)

= Orchis sambucina var. robusta Neuman; Sver. Fl. 629 (1901), nom. inval.

= Orchis sambucina var. rubra Huter; 48 (1872), nom. inval.

= Orchis sambucina var. rubra Winterl; Index Hort. Bot. Univ. Hung.: s.p. (1788)

= Orchis sambucina var. rubriflora St.-Lag.; Cariot, Étude Fl., éd. 8, 2: 806 (1889)

= Orchis schleicheri Sweet; Brit. Fl. Gard. 2: t. 199 (1827)

species :

Dactylorhiza sibirica Efimov; Phytotaxa 258(2): 111 (2016)

Distribution: Siberia (Krasnoyarsk)

30 SIB

species :

Dactylorhiza sphagnicola (Höppner ex Soó) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

Name: Torf-Fingerwurz, Torf-Fingerknabenkraut (DE)

Distribution: Denmark; Norway; Sweden; +Netherlands; Belgium; Germany (Hamburg, Niedersachsen, Nordrhein-Westfalen, Schleswig-Holstein); France

10 DEN NOR SWE 11 BGM GER NET 12 FRA

Remarks: praetermissa agg.

Synonyms: hidden

= Dactylorchis sphagnicola (Höppner) Verm.; Stud. Dactylorch.: 66 (1947)

= Dactylorchis traunsteineri subsp. sphagnicola (Höppner) Verm.; Flora Neerl. I 5: 70 (1958)

= Dactylorhiza incarnata subsp. sphagnicola (Höppner) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza majalis subsp. sphagnicola (Höppner) H.A.Pedersen & Hedrén; Nord. J. Bot. 22(6): 651 (2004)

= Dactylorhiza sphagnicola (Höppner) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69: 875 (1984)

= Orchis angustifolia var. densiflora Neuman; Bot. Not. 1909: 234 (1909), nom. illeg.

= Orchis sphagnicola Höppner; Sitzungsber. Niederrhein. Ges. Natur- Heilk. Bonn 1926D: 23 (1927)

= Orchis ×sphagnicola Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 252, 259 (1933)

form :

f. calcifugiens (H.A.Pedersen) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

Distribution: Denmark

10 DEN

Synonyms: hidden

= Dactylorhiza majalis subsp. calcifugiens H.A.Pedersen; Nordic J. Bot. 22: 655 (2004)

= Dactylorhiza sphagnicola subsp. calcifugiens (H.A.Pedersen) Kreutz & Sczep.; Ber. Arbeitskreis. Heimische Orchid. 24(2): 97 (2007)

variant :

var. deweveri (Verm.) Kreutz; J. Eur. Orch. 48(2-4): 314 (2016)

Distribution: Netherlands; Belgium; N-Germany

11 BGM GER NET

Synonyms: hidden

= Dactylorchis deweveri Verm.; Ned. Kruidk. Arch., Versl. 56: 227 (1949)

= Dactylorhiza deweveri (Verm.) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza sphagnicola f. deweveri (Verm.) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza sphagnicola subsp. deweveri (Verm.) Wenker; Orchideen Nordrhein-Westfalens 105 (2001)

= Dactylorhiza traunsteineri subsp. deweveri (Verm.) Verm.; Fl. Neerl., I, 5, Orchidaceae: 71 (1958)

variant :

var. hoeppneri (A.Fuchs) Kreutz; J. Eur. Orch. 48(2-4): 317 (2016)

Distribution: Netherlands; NW-Germany

11 GER NET

Synonyms: hidden

= Dactylorchis hoeppneri (A.Fuchs) Verm.; Stud. Dactylorch.: 66 (1947)

= Dactylorhiza hoeppneri (A.Fuchs) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 3: 346 (1960)

= Dactylorhiza sphagnicola f. hoeppneri (A.Fuchs) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

= Dactylorhiza sphagnicola subsp. hoeppneri (A.Fuchs) Dalkowski, Wende & Wenker; Orchideen Nordrhein-Westfalens 105 (2001), nom. inval.

= Orchis hoeppneri (A.Fuchs) Höppner ex Verm.; Stud. Dactylorch. 66 (1947)

= Orchis traunsteineri f. hoeppneri (A.Fuchs) Soó; Keller, Monogr. Iconogr. Orchid. Eur. 2: 251 (1933)

= Orchis ×pseudotraunsteineri subsp. hoeppneri A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 149 (1919)

form :

f. sennia (Vollmar) P.Delforge; Naturalistes Belges 88(Orchid. 20): 246 (2007)

Distribution: NC-Germany (E-Nordrhein-Westfalen)

11 GER

Synonyms: hidden

= Dactylorhiza maculata subsp. sennia (Vollmar) Kreutz; Kompend. Eur. Orchid. 47 (2004)

= Dactylorhiza sennia Vollmar; Ber. Arbeitskreis. Heimische Orchid. 18(2): 98 (2001 publ. 2002)

= Dactylorhiza sphagnicola subsp. sennia (Vollmar) Kreutz; Eurorchis 17: 105 (2005)

variant :

var. wirtgenii (Höppner) comb.ined.

Distribution: NW-Germany; France

11 GER 12 FRA

Synonyms: hidden

= Dactylorhiza traunsteineri subsp. wirtgenii (Höppner) Kreutz; Kompend. Eur. Orchid. 57 (2004)

= Dactylorhiza wirtgenii (Höppner) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Orchis wirtgenii Höppner; Abh. Vereins Naturwiss. Erforsch. Niederrheins 2: 57 (1916)

species :

Dactylorhiza stortonii Benigni, Mandozzi, Monaldi, Barigelli & Petroselli; Micol. Marche, N. S. 5(2): 43 (2021)

Distribution: Italy

13 ITA

Remarks: provisionally accepted name

species :

Dactylorhiza traunsteineri (Saut. ex Rchb.) Soó; Nom. Nov. Gen. Dactylorhizae 5 (1962)

subspecies :

subsp. traunsteineri

Name: Traunsteiners Knabenkraut (DE)

Distribution: +Netherlands; Germany (Baden-Württemberg, Bayern); Switzerland; Austria; France; Italy; Slovenia; Croatia

11 AUT GER NET SWI 12 FRA 13 CRO ITA SLV

Remarks: traunsteineri agg. This Central European, mostly alpine species is not identical with the nordic D. lapponica

Synonyms: hidden

= Dactylorchis gracilis Hesl.-Harr.; Vasculum 35: 19 (1950)

= Dactylorchis traunsteineri (Saut. ex Rchb.) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorchis traunsteineri var. klingei Verm.; Ned. Kruidk. Arch. 56: 225 (1949)

= Dactylorchis traunsteineri var. rhombilabris Verm.; Ned. Kruidk. Arch. 56: 225 (1949)

= Dactylorhiza devillersiorum P.Delforge; Naturalistes Belges 92(Orchid. 24): 79 (2011)

= Dactylorhiza majalis subsp. traunsteineri (Saut. ex Rchb.) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza traunsteineri var. mielichhoferi (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza: 6 (1962)

= Dactylorhiza ×gracilis (Höppner) Hesl.-Harr.; Vasculum (Substitute) 35: 19 (1950)

= Dactylorhiza ×gracilis (Höppner) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×rhenana (Höppner) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×rigida (Höppner) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×steegeri (Höppner) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×suevica (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Orchis angustifolia subf. conferta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia subf. densata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia subf. dissita Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia subf. elegantior Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia subf. filiformis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia subf. intermedia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 146 (1899)

= Orchis angustifolia subf. laxata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia subf. maculata Klinge; Rev. Orch. cordigera, Orchis angustifolia: 68 (1893)

= Orchis angustifolia subf. strictior Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia subf. subcurva (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 79 (1899)

= Orchis angustifolia subf. superans Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia subf. venusta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 79 (1899)

= Orchis angustifolia f. acuminata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 76 (1899)

= Orchis angustifolia f. ammatica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 79 (1899)

= Orchis angustifolia f. amplivaginata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia f. angustivaginata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia f. anziensis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 80 (1899)

= Orchis angustifolia f. brevispicata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia f. comosa Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia f. densiflora Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia f. densissima Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. densiuscula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

= Orchis angustifolia f. fichtenbergii Klinge; Rev. Orch. cordigera, Orchis angustifolia: 68 (1893)

= Orchis angustifolia f. friesii Klinge; Rev. Orch. cordigera, Orchis angustifolia: 67 (1893)

= Orchis angustifolia f. gracilis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia f. immaculata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899), nom. illeg.

= Orchis angustifolia f. laxa Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 85 (1899)

= Orchis angustifolia f. laxiflora Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia f. laxiuscula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

= Orchis angustifolia f. lehnertii Klinge; Rev. Orch. cordigera, Orchis angustifolia: 67 (1893)

= Orchis angustifolia f. levaschevoica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. longispicata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia f. oblonga Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 85 (1899)

= Orchis angustifolia f. obtusifolia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 75 (1899)

= Orchis angustifolia f. paanajaervica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia f. papjerwica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia f. perhoica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899)

= Orchis angustifolia f. pulcherrima Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. reichenbachii Klinge; Rev. Orch. cordigera, Orchis angustifolia: 67 (1893)

= Orchis angustifolia f. remota Klinge; Rev. Orch. cordigera, Orchis angustifolia: 67 (1893)

= Orchis angustifolia f. rosiensis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 80 (1899)

= Orchis angustifolia f. sauteri Klinge; Rev. Orch. cordigera, Orchis angustifolia: 67 (1893)

= Orchis angustifolia f. schmidtii Klinge; Rev. Orch. cordigera, Orchis angustifolia: 68 (1893)

= Orchis angustifolia f. spathulata Klinge; Rev. Orch. cordigera, Orchis angustifolia: 67 (1893), et Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 76 (1899)

= Orchis angustifolia f. subcurva Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia f. suberecta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

= Orchis angustifolia f. tiudiensis Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 88 (1899)

= Orchis angustifolia Rchb. ex Klinge; 291 (1893), nom. illeg.

= Orchis angustifolia subsp. friesii (Klinge) Neuman; Bot. Not. 1909: 231 (1909)

= Orchis angustifolia subsp. subcapitata Neuman; Bot. Not. 1909: 233 (1909)

= Orchis angustifolia subsp. traunsteineri (Saut. ex Rchb.) C.Hartm.; Handb. Skand. Fl., ed. 11: 388 (1879) [Unplaced]

= Orchis angustifolia subsp. traunsteineri (Saut. ex Rchb.) F.Aresch.; 409 (1881)

= Orchis angustifolia subsp. traunsteineri (Saut. ex Rchb.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 175 (1898)

= Orchis angustifolia var. acuminata (Klinge) Neuman; Bot. Not. 1909: 232 (1909)

= Orchis angustifolia var. blyttii (Rchb.f.) Klinge; Rev. Orch. cordigera, Orchis angustifolia 67 (1893)

= Orchis angustifolia var. elongata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis angustifolia var. erecta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia var. gracillima Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia var. immaculata Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899), nom. illeg.

= Orchis angustifolia var. mielichhoferi Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 75 (1899)

= Orchis angustifolia var. patens Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899)

= Orchis angustifolia var. poenalica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia var. pusilla Neuman; Bot. Not. 1909: 233 (1909)

= Orchis angustifolia var. recurva Klinge; Rev. Orch. cordigera, Orchis angustifolia 68 (1893)

= Orchis angustifolia var. reolana Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 86 (1899)

= Orchis angustifolia var. rigidula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899)

= Orchis angustifolia var. sanionis Klinge; Rev. Orch. cordigera, Orchis angustifolia 67 (1893)

= Orchis angustifolia var. stricta Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 82 (1899), nom. illeg.

= Orchis angustifolia var. superba Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 80 (1899)

= Orchis angustifolia var. tarbatonica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 78 (1899)

= Orchis angustifolia var. tenuifolia Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 81 (1899)

= Orchis angustifolia var. traunsteineri (Saut. ex Rchb.) Klinge; 329 (1893)

= Orchis incarnata subsp. traunsteineri (Saut. ex Rchb.) Cajander; 171 (1906)

= Orchis incarnata var. angustifolia (Lindl.) Rchb.f.; Rchb., Icon. Fl. Germ. Helv. 13-14: 52 (1851), nom. illeg.

= Orchis incarnata var. traunsteineri (Saut. ex Rchb.) Parl.; Fl. Ital. 3: 521 (1860)

= Orchis latifolia var. traunsteineri (Saut. ex Rchb.) Godr.; Fl. Lorraine 3: 30 (1844)

= Orchis maculata subsp. traunsteineri (Saut. ex Rchb.) Douin; Fl. Ill. France 11: 30 (1931)

= Orchis sambucina var. subalpina Rchb.; 140[hochgestellt:]18 (1832)

= Orchis traunsteineri Saut. ex Koch; Syn. Fl. Germ., ed. I, 688 (1837)

= Orchis traunsteineri Saut. ex Rchb.; Fl. Germ. Excurs.: 140 (1831)

= Orchis ×gracilis Höppner; Sitz. Niederrh. Ges. Natur.- u. Heilk., D. 14 (1926 publ. 1927)

= Orchis ×gracilis Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 252 (1933)

= Orchis ×pseudotraunsteineri subsp. suevica A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 107 (1919)

= Orchis ×rhenana Höppner; Abk. Ver. Naturw. Erforsch. Niederrh. 2: 69 (1916)

= Orchis ×steegeri Höppner; Sitz. Niederrh. Ges. Natur.- u. Heilk., D. 16 (1926 publ. 1927)

= Orchis ×steegeri Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 252 (1933)

variant :

var. vosagiaca (Kreutz & P.Wolff) Sonnb.; Fl. German. [M. Hassler & T. Muer] 1675 (2022)

Name: Wasgau-Fingerwurz (DE)

Distribution: SW-Germany (Rheinland-Pfalz, +Saarland); NE-France (Vosges)

11 GER 12 FRA

Synonyms: hidden

= Dactylorhiza traunsteineri subsp. vosagiaca Kreutz & P.Wolff; Ber. Arbeitskreis. Heimische Orchid. 27(2): 196 (2010 publ. 2011)

= Dactylorhiza traunsteineri subsp. vosagiaca P.Wolff; 77 (1998), nom. inval.

= Dactylorhiza traunsteineri subsp. vosagiaca P.Wolff ex Kreutz & P.Wolff; 196 (2011)

= Dactylorhiza vosagiaca (P.Wolff ex Kreutz & P.Wolff) Herr-Heidtke & Heidtke ex P.Wolff; Kochia 10: 57 (2017)

= Dactylorhiza vosagiaca (P.Wolff ex Kreutz & P.Wolff) Herr-Heidtke & Heidtke; 11 (2012), nom. inval.

subspecies :

subsp. bohemica (Businský) Kreutz & Sczep.; Ber. Arbeitskreis. Heimische Orchid. 24(2): 94 (2007)

Distribution: Czech Republic; Slovakia

11 CZE SLO

Synonyms: hidden

= Dactylorhiza bohemica Businsky; Preslia 61(4): 296 (1989)

subspecies :

subsp. irenica (F.M.Vázquez) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 28(2): 269 (2011)

Distribution: Spain

12 SPA

Synonyms: hidden

= Dactylorhiza irenica F.M.Vázquez; J. Eur. Orch. 40(1): 32 (2008)

subspecies :

subsp. rhaetica (H.Baumann & R.Lorenz) F.Benoît; Orchidophile (Asnières) 205: 228 (2015)

Distribution: S-Germany (Bayern); Switzerland; Austria; Slovakia (Mala Fatra); France; N-Italy; ?Ukraine (Carpathians)

11 AUT GER SLO SWI 12 FRA 13 ITA

Synonyms: hidden

= Dactylorhiza lapponica subsp. rhaetica H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(4): 941 (2005)

= Dactylorhiza rhaetica (H.Baumann & R.Lorenz) Biagioli, Kreutz & De Simoni; GIROS Orch. Spont. Eur. 65(2): 149 (2022)

subspecies :

subsp. russowii (Klinge) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

Distribution: Sweden; Finland; NE-Germany (Mecklenburg-Vorpommern); Poland; Czech Republic; Estonia; Latvia; Lithuania; N-Belarus; European Russia (N-European Russia, C-European Russia); Siberia (Krasnoyarsk, W-Siberia); Kazakhstan

10 FIN SWE 11 CZE GER POL 14 BLR EST LAT LIT RUS 30 SIB 32 KAZ

Synonyms: hidden

= Dactylorchis curvifolia (F.Nyl.) Verm.; Stud. Dactylorch. 151 (1947)

= Dactylorchis pseudocordigera (Neuman) Lid; Norsk Fl., ed. 2, 210 (1952)

= Dactylorchis russowii (Klinge) Á.Löve & D.Löve; Bot. Notiser 114: 38 (1961)

= Dactylorchis traunsteineri subsp. curvifolia (F.Nyl.) Verm.; Stud. Dactylorchis 66 (1947)

= Dactylorchis traunsteineri subsp. russowii (Klinge) Verm.; Stud. Dactylorchis 158 (1947)

= Dactylorhiza curvifolia (F.Nyl.) Czerep.; Sosud. Rast. S.S.S.R.: 307 (1981)

= Dactylorhiza lapponica subsp. pseudocordigera (Neuman) Landwehr; Orchideeen 37(3): 80 (1975), isonym

= Dactylorhiza lapponica subsp. pseudocordigera (Neuman) Verm.; Stud. Dactylorch. 67 (1947)

= Dactylorhiza lapponica subsp. russowii (Klinge) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 706 (2005)

= Dactylorhiza lapponica var. pseudocordigera (Neuman) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 152 (2007)

= Dactylorhiza majalis subsp. russowii (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza pseudocordigera (Neuman) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza russowii (Klinge) Holub; Preslia 36: 253 (1964)

= Dactylorhiza traunsteineri subsp. curvifolia (F.Nyl.) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza traunsteineri var. curvifolia (F.Nyl.) Aver.; Turczaninowia 3(1): 50 (2000)

= Dactylorhiza traunsteineri var. patula (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza: 6 (1962)

= Dactylorhiza traunsteineri var. russowii (Klinge) Aver.; Turczaninowia 3(1): 50 (2000)

= Orchis angustifolia f. arcuata Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia f. curvata Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia f. elongata Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia f. gruneri Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia f. intermedia Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia f. patens Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia f. stricta Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia subsp. curvifolia (F.Nyl.) E.G.Camus & A.Camus; Icon. Orchid. Eur., Text(1): 229 (1928)

= Orchis angustifolia subsp. russowii (Klinge) Klinge; Dactylorch. Monogr. Prodr. 8, 31 (1898)

= Orchis angustifolia subf. concolor Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893)

= Orchis angustifolia subf. immaculata Klinge; Rev. Orch. cordigera, Orchis angustifolia: 69 (1893), nom. illeg.

= Orchis angustifolia var. arcuata (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 85 (1899)

= Orchis angustifolia var. curvata (Klinge) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 84 (1899)

= Orchis angustifolia var. curvifolia (F.Nyl.) Nyman; Consp. Fl. Eur. Suppl. 2: 291 (1890)

= Orchis angustifolia var. patula Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

= Orchis angustifolia var. russowii Klinge; Arch. Naturk. Livl. Estl. Kurl. Ser. 2., Biol. Naturk. 10: 84 (1893)

= Orchis curvifolia F.Nyl.; Spicil. Pl. Fennic. 2: 12. n. 25 (1844)

= Orchis incarnata f. curvifolia (F.Nyl.) Cajander; 172 (1906)

= Orchis maculata subsp. curvifolia (F.Nyl.) E.G.Camus; Monogr. Orch. 194 (1908)

= Orchis pseudocordigera Neuman; Bot. Notiser 236 (1909)

= Orchis recurva F.Nyl. ex Fr.; Summa Veg. Scand. 1: 61 (1814)

= Orchis russowii (Klinge) Klinge; Acta Hort. Petrop. 17 1: 154, in clavi, 177 (1899)

= Orchis russowii (Klinge) Kuppf.; Exsicc. (Herb. Norm.) 44: 107 (1902), isonym

= Orchis russowii (Klinge) Schltr.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 1 (G. Keller & Schltr., Monogr. & Iconogr. Orch. Eur.) 170 (1927), isonym

= Orchis traunsteineri subsp. russowii (Klinge) Soó; Bot. Arch. 23: 83 (1929)

= Orchis traunsteineri var. russowii (Klinge) M.Schulze; Mitth. Thüring. Bot. Vereins, n.f., 17: 50 (1902)

= Orchis traunsteineri var. russowii (Klinge) Vollm.; 164 (1914), isonym

subspecies :

subsp. turfosa (F.Proch.) Kreutz; Kompend. Eur. Orchid. 56 (-57) (2004)

Distribution: Czech Republic

11 CZE

Synonyms: hidden

= Dactylorhiza comosa subsp. turfosa (F.Proch.) Holub; Preslia 70(2): 103 (1998)

= Dactylorhiza majalis subsp. turfosa F.Proch.; Preslia 54(4): 289 (1982)

species :

Dactylorhiza traunsteinerioides (Pugsley) R.M.Bateman & Denholm; List Vasc. Pl. Brit. Isles (D. H. Kent) Suppl. 3: 20 (2006)

subspecies :

subsp. traunsteinerioides

Distribution: NE-England; N-Wales; Ireland

10 GRB IRE

Remarks: for detailed discussion see Bateman & Denholm 2012

Synonyms: hidden

= Dactylorchis traunsteinerioides (Pugsley) Verm.; Stud. Dactylorch. 66 (1947)

= Dactylorhiza majalis subsp. traunsteinerioides (Pugsley) R.M.Bateman & Denholm; Watsonia 14(4): 372 (1983)

= Dactylorhiza majalis var. traunsteinerioides (Pugsley) R.M.Bateman & Denholm; Watsonia 14(4): 373 (1983)

= Dactylorhiza traunsteineri subsp. hibernica Landwehr; Orchideeen 37(3): 79 (1975), et Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza traunsteineri subsp. traunsteinerioides (Pugsley) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza traunsteinerioides (Pugsley) Landwehr; Orchideeen 37(3): 80 (1975), nom. inval.

= Orchis majalis subsp. traunsteinerioides Pugsley; Proc. Linn. Soc. Lond. 148: 124 (1936)

= Orchis traunsteinerioides (Pugsley) Pugsley; J. Bot. 78: 179 (1940)

subspecies :

subsp. francis-drucei (Wilmott) R.M.Bateman & Denholm; New J. Bot. 2(1): 44 (2012)

Distribution: W-Scotland (Hebrides)

10 GRB

Synonyms: hidden

= Dactylorhiza francis-drucei (Wilmott) Aver.; Bot. Zhurn. (Moscow & Leningrad) 69(6): 875 (1984)

= Dactylorhiza majalis var. francis-drucei (Wilmott) R.M.Bateman & Denholm; Watsonia 14(4): 373 (1983)

= Dactylorhiza traunsteineri subsp. francis-drucei (Wilmott) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza traunsteinerioides var. francis-drucei (Wilmott) F.M.Vázquez; J. Eur. Orch. 40(1): 30 (2008)

= Orchis francis-drucei Wilmott; Proc. Linn. Soc. Lond. Sess. 148, 128 (1936)

variant :

var. ebudensis (Wief. ex R.M.Bateman & Denholm) R.M.Bateman & Denholm; New J. Bot. 2(1): 45 (2012)

Distribution: Scotland (Hebrides: North Uist Isl., S-Berneray Isl.)

10 GRB

Synonyms: hidden

= Dactylorhiza comosa subsp. scotica (E.Nelson) P.D.Sell; Sell & Murrell, Fl. Great Britain Ireland 5: 364 (1996)

= Dactylorhiza ebudensis (Wief. ex R.M.Bateman & Denholm) P.Delforge; Naturalistes Belges 81: 397 (2000)

= Dactylorhiza majalis subsp. ebudensis (Wief. ex R.M.Bateman & Denholm) M.R.Lowe; Eurorchis 15: 81 (2003)

= Dactylorhiza majalis subsp. scotica E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 90 (1976), without type, et Taxon 28 (5-6): 593 (1979), with type

= Dactylorhiza majalis var. bowmanii M.N.Jenk.; Watsonia 20(3): 272 (1995)

= Dactylorhiza majalis var. ebudensis Wief.; Landwehr, Wilde Orchid. Eur. 2: 557 (1977)

= Dactylorhiza majalis var. ebudensis Wief. ex R.M.Bateman & Denholm; Edinburgh J. Bot. 52(1): 57 (1995)

= Dactylorhiza majalis var. scotica (E.Nelson) R.M.Bateman & Denholm; Watsonia 44: 368 (1983)

species :

Dactylorhiza umbrosa (Kar. & Kir.) Nevski; Acta Inst. Bot. Acad. Sci. URSS, Ser. I, Fasc. 4, 332 (1937)

Name: Persisches Knabenkraut

Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Tajikistan; Armenia; Azerbaijan; E-Turkey; N-Iraq; Iran; Afghanistan (Badakshan, Wakhan, Baghlan, Bamyan, Faryab, Herat, Kabul, Kunar / Nuristan, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan, Takhar); China (Xinjiang); N-Pakistan (Kurram); India (Jammu & Kashmir)

32 KAZ KGZ TKM TZK UZB 33 ARM AZE 34 AFG IRN IRQ TUR 36 CHI 40 IND PAK

Remarks: incarnata agg. Often included in D. incarnata ssp. cilicica

Synonyms: hidden

= Dactylorchis umbrosa (Kar. & Kir.) Wendelbo; Nytt Mag. Bot., Oslo 1: 24 (1952)

= Dactylorhiza chuhensis Renz & Taubenheim; Fl. Turkey & E. Aegaean Isl. 8: 564 (1984), nom. nov.

= Dactylorhiza incarnata subsp. turcestanica (Klinge) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza incarnata var. kotschyi (Rchb.f.) H.A.Pedersen, P.J.Cribb & Rolf Kühn; Field Guide Orchids Europe Medit. 139 (2019)

= Dactylorhiza kotschyi (Rchb.f.) P.F.Hunt & Summerh.; Watsonia 6: 130 (1965)

= Dactylorhiza merovensis (Grossh.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 894 (1983)

= Dactylorhiza persica (Schltr.) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza renzii Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 893 (1983) [non H. Baumann & Künkele (1981)]

= Dactylorhiza sanasunitensis (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhizae 4 (1962)

= Dactylorhiza umbrosa subsp. persica (Schltr.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 156 (2007)

= Dactylorhiza umbrosa var. chuhensis (Renz & Taubenheim) Kreutz; Orch. Tur. 181 (1998)

= Dactylorhiza umbrosa var. longibracteata Renz; Fl. Iranica 126: 131 (1978)

= Dactylorhiza umbrosa var. ochroleuca (Bornm.) Renz; Fl. Iranica 126: 131 (1978)

= Dactylorhiza vanensis E.Nelson; Monogr. & Ikonogr. Orchid.-Gatt. Dactylorhiza 59 (1976), without type, et Taxon 28 (5-6): 593 (1979), with type

= Orchis incarnata f. ochroleuca Bornm.; Repert. Spec. Nov. Regni Veg. 47: 72 (1939), nom. illeg.

= Orchis incarnata var. kotschyi Rchb.f.; Icon. Fl. Germ. Helv. (H. G. L. Reichenbach) 13-14: 53 (1851)

= Orchis kotschyi (Rchb.f.) Schltr.; Repert. Spec. Nov. Regni Veg. 19: 48 (1923)

= Orchis merovensis Grossh.; Beih. Bot. Centralbl. 44: II. 207 (1927)

= Orchis olocheilos var. sanasunitensis (H.Fleischm.) Soó; Repert. Spec. Nov. Regni Veg. 24: 30 (1927)

= Orchis orientalis subsp. turcestanica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 183 (1898)

= Orchis persica Schltr.; Repert. Spec. Nov. Regni Veg. 15: 290 (1918)

= Orchis sanasunitensis H.Fleischm.; Ann. K. K. Naturhist. Hofmus. 28: 35 (1914)

= Orchis turcestanica (Klinge) O.Fedtsch.; Acta Hort. Petrop. 17 1: 155, in clavi, 183 (1898), et ex B. Fedtsch., J. Russe de Bot., 191 (1908)

= Orchis umbrosa Kar. & Kir.; Bull. Soc. Nat. Mosc. 15: 504 (1842)

species :

Dactylorhiza urvilleana (Steud.) H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(2): 240 (1981)

subspecies :

subsp. urvilleana

Name: D´Urvilles Knabenkraut

Distribution: Northern Caucasus; Georgia [Caucasus] (Abkhazia, Racha-Lechkhumi, Samegrelo, Imereti, Adzharia, Shiga Kartli, Kartli, Mtiuleti, Trialeti, Javakheti); Armenia; Azerbaijan (incl. Talysh); NE-Turkey; Iran

33 ARM AZE GEO NCS 34 IRN TUR

Remarks: praetermissa agg.

Synonyms: hidden

= Dactylorchis amblyoloba (Nevski) Soó; [nom. inval.]

= Dactylorchis lancibracteata (K.Koch) Soó; [nom. inval.]

= Dactylorhiza affinis (K.Koch) Aver.; Bot. Zhurn. (Moscow & Leningrad) 68(7): 895 (1983)

= Dactylorhiza amblyoloba (Nevski) Aver.; Novosti Sist. Vyssh. Rast. 26: 47 (1989)

= Dactylorhiza lancibracteata (K.Koch) Renz; Fl. Iranica 126: 135 (1978)

= Dactylorhiza maculata subsp. triphylla (K.Koch) H.Sund.; Europ. Medit. Orchid., ed. 3: 40 (1980)

= Dactylorhiza pontica (H.Fleischm. & Hand.-Mazz.) P.Delforge; Naturalistes Belges 81(4): 397 (2000)

= Dactylorhiza saccifera subsp. amblyoloba (Nevski) Eccarius; Orchideengattung Dactylorhiza 166 (2016)

= Dactylorhiza saccifera subsp. cartaliniae (Klinge) Soó; Nom. Nov. Gen. Dactylorhizae 7 (1962) [p. p., nom. inval.]

= Dactylorhiza saccifera subsp. lancibracteata (K.Koch) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza saccifera var. pontica (H.Fleischm. & Hand.-Mazz.) Soó; Nom. Nov. Gen. Dactylorhiza 8 (1962)

= Dactylorhiza triphylla (H.Sund.) Czerep.; Sosud. Rast. S.S.S.R.: 309 (1981)

= Dactylorhiza urvilleana var. pontica (H.Fleischm. & Hand.-Mazz.) P.Delforge; Naturalistes Belges 89(Orchid. 21): 250 (2008)

= Orchis affinis K.Koch; Linnaea 22: 284 (1849)

= Orchis amblyoloba Nevski; Komarov, Fl. URSS 4: 707, 753 (1935)

= Orchis basilica subsp. cartaliniae Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(1): 158, 196 (1898)

= Orchis cartaliniae (Klinge) Lipsky; Acta Hort. Petrop. 17 1: 158, in clavi, et 196 (1898)

= Orchis cartaliniae (Klinge) Medw.; Act. Hort. Tiflis 18: 475 (1919) [in clavi]

= Orchis lancibractea Grossh.; Fl. Kavkaza 1: 269 (1928)

= Orchis lancibracteata K.Koch; Linnaea 22: 282 (1850)

= Orchis maculata subsp. lancibracteata (K.Koch) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927) [nom. inval.]

= Orchis maculata var. broteri Sommier & Levier; Trudy Imp. S.-Peterburgsk. Bot. Sada 16: 419 (1900)

= Orchis maculata var. cartaliniae (Klinge) Schltr.; Monogr. Iconogr. Orchid. Eur. 1: 179 (1927)

= Orchis maculata var. pontica (H.Fleischm. & Hand.-Mazz.) Soó; Repert. Spec. Nov. Regni Veg. 24: 32 (1927)

= Orchis pontica H.Fleischm. & Hand.-Mazz.; Ann. K. K. Naturhist. Hofmus. 23: 208 (1909)

= Orchis saccata d´Urv.; Enum. Pl. Ponti-Eux.: 119 (1822), nom. illeg.

= Orchis triphylla K.Koch; Linnaea 22: 283 (1850) [non Spreng. (1826)]

= Orchis urvilleana Steud.; Nomencl. Bot., ed. 2, 2: 225 (1841)

= xDactyloglossum ×ponticum Kohlmüller; Ber. Arbeitskr. Heim. Orch. 16(1): 74 (1999)

subspecies :

subsp. bithynica (H.Baumann) H.Baumann & R.Lorenz; Jour. Eur. Orch. 37(3): 707 (2005)

Distribution: Turkey

34 TUR

Synonyms: hidden

= Dactylorhiza bithynica H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 68 (1983)

= Dactylorhiza saccifera subsp. bithynica (H.Baumann) Kreutz; Kompend. Eur. Orchid. 54 (2004)

subspecies :

subsp. ilgazica (Kreutz) Kreutz; Kompend. Eur. Orchid. 58 (2004)

Distribution: NE-Turkey

34 TUR

Synonyms: hidden

= Dactylorhiza ilgazica Kreutz; Eurorchis 9: 49 (1997)

subspecies :

subsp. phoenissa B.Baumann & H.Baumann; Jour. Eur. Orch. 37(2): 251, 254 (2005)

Distribution: Lebanon

34 LEB

Synonyms: hidden

= Dactylorhiza phoenissa (B.Baumann & H.Baumann) P.Delforge; Naturalistes Belges 96(Orchid. 28): 15 (2015)

species :

Dactylorhiza viridis (L.) R.M.Bateman, Pridgeon & M.W.Chase; Lindleyana 12(3): 113-141 (1997)

Name: Grüne Hohlzunge (DE), Frog Orchid (EN)

Distribution: Iceland; Faeroe Isl.; England; Ireland; Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Baden-Württemberg, Bayern, Hessen, +Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Spain; Andorra; France; Italy; ?Sicily; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Estonia; Latvia; Lithuania; Belarus; Ukraine; Crimea; European Russia (N-European Russia, W-European Russia, C-European Russia); Siberia (Altai, Buryatia, Chita, Irkutsk, Krasnoyarsk, Tuva, W-Siberia, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Magadan, Primorye, Commander Isl.); Kazakhstan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus] (Abkhazia, Samegrelo, Imereti, Adzharia, Shiga Kartli, Kartli, Mtiuleti); Armenia; Azerbaijan (incl. Talysh); N-Turkey; Iran; China (Gansu, Hebei, Heilongjiang, Henan, Hubei, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shanxi, Sichuan, Xinjiang, NW-Yunnan); NE-Tibet; Mongolia; North Korea; South Korea; Japan (Hokkaido, Honshu); Taiwan; Pakistan (Baltistan, Karakorum); Nepal; Bhutan; India (Himachal Pradesh, Jammu & Kashmir); Aleutes; C-Alaska; Canada (Alberta, British Columbia, Manitoba, New Brunswick, Newfoundland, N. W. T., Nova Scotia, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon, Anticosti Isl.); USA (Arizona, Colorado, Connecticut, Illinois, Indiana, Iowa, Maine, Maryland, Massachusetts, Michigan, Minnesota, Missouri, Montana, Nebraska, New Hampshire, New Jersey, New Mexico, New York, North Carolina, North Dakota, Ohio, Pennsylvania, Rhode Island, South Dakota, Tennessee, Utah, Vermont, Virginia, Washington, West Virginia, Wisconsin, Wyoming)

10 DEN FIN FOR GRB ICE IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO FRA SPA 13 ALB BIH BUL CRO ITA KOS MNT NMA ROM SLV SRB 14 BLR EST KRY LAT LIT RUS UKR 30 SIB 31 RFE 32 KAZ KGZ TZK 33 ARM AZE GEO NCS 34 IRN TUR 36 CHI TIB 37 MON 38 JAP KOR NKO TAI 40 BHU IND NEP PAK 70 ASK CDN 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA

Synonyms: hidden

= Chamorchis viridis (L.) Dumort.; Fl. Belg.: 133 (1827)

= Coeloglossum bracteatum (Muhl. ex Willd.) Parl.; Fl. Ital. 3: 409 (1860)

= Coeloglossum bracteatum (Muhl. ex Willd.) Schltr.; Repert. Spec. Nov. Regni Veg. 16 (463-467): 374 (1920), isonym

= Coeloglossum bracteatum var. kashmiricum Soó; Ann. Hist.-Nat. Mus. Natl. Hung. 26: 356 (1929)

= Coeloglossum bracteatum var. majus (Maxim.) Nakai; J. Jap. Bot. 18: 282 (1942)

= Coeloglossum coreanum (Nakai) Schltr.; Repert. Spec. Nov. Regni Veg. 16: 374 (1920)

= Coeloglossum islandicum (Lindl.) Kom.; Fl. Penins. Kamtsch. 1: 313 (1927), isonym

= Coeloglossum islandicum (Lindl.) Nyman; Syll. Fl. Eur. 359 (1855)

= Coeloglossum kaschmirianum Schltr.; Repert. Spec. Nov. Regni Veg. 16: 374 (1920)

= Coeloglossum nankotaizanense (Masam.) Masam.; J. Geobot., Hokuriku 16: 55 (1968)

= Coeloglossum nankotaizanensis (Masam.) S.S.Ying; Quart. J. Chin. Forest. 8(4): 149 (1975), isonym

= Coeloglossum taiwanianum S.S.Ying; Alp. Pl. Taiwan Colour 1: 74 (1975)

= Coeloglossum vaillantii (Ten.) Guss.; Icon. Fl. Germ. 13-14: 130 (1851)

= Coeloglossum viride (L.) Hartm.; Handb. Skand. Fl.: 329 (1820)

= Coeloglossum viride f. bracteatum (Muhl. ex Willd.) Bolzon; Bull. Soc. Bot. Ital. 1909: 70 (1909)

= Coeloglossum viride f. brevibracteatum Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 807 (1907)

= Coeloglossum viride f. collinum (Rupp ex W.Zimm.) Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1924 publ. 1925)

= Coeloglossum viride f. dentatum Zapal.; Consp. Fl. Gallic. Crit. 1: 214 (1906)

= Coeloglossum viride f. gracillimum Schur; Enum. Pl. Transsilv.: 645 (1866)

= Coeloglossum viride f. grandiflorum Zapal.; Consp. Fl. Gallic. Crit. 1: 214 (1906)

= Coeloglossum viride f. integrum Soó; Acta Bot. Acad. Sci. Hung. 16: 367 (1970 publ. 1971), no type indicated.

= Coeloglossum viride f. islandicum (Lindl.) O.Gruss & M.Wolff; Orchid. Atlas: 77 (2007)

= Coeloglossum viride f. lancifolium Rohlena; Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 1912(Beitr.): 118 (1913)

= Coeloglossum viride f. longibracteatum Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 807 (1907)

= Coeloglossum viride f. microbracteatum Schur; Oesterr. Bot. Z. 20: 294 (1870)

= Coeloglossum viride f. parviflorum Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1924 publ. 1925)

= Coeloglossum viride f. purpureum Schur; Enum. Pl. Transsilv.: 646 (1866)

= Coeloglossum viride f. roseum Bolzon; Bull. Soc. Bot. Ital. 1910: 71 (1910)

= Coeloglossum viride f. rubens Norm.; Christ. Vidensk.-Selsk. Forh. 1893 (16): 39 (1893)

= Coeloglossum viride f. subalpinum Neuman; Sver. Fl.: 633 (1901)

= Coeloglossum viride f. thuringiacum (Rupp ex W.Zimm.) Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1924 publ. 1925)

= Coeloglossum viride f. virescens (Muhl. ex Willd.) O.Gruss & M.Wolff; Orchid. Atlas: 77 (2007)

= Coeloglossum viride subsp. bracteatum (Muhl. ex Willd.) Hultén; Acta Univ. Lund. 2, 39, no. 1: 472 (1943)

= Coeloglossum viride subsp. coreanum (Nakai) Satomi; J. Jap. Bot. 44: 127 (1969)

= Coeloglossum viride subsp. eurasiaticum Selander; Acta Phytogeogr. Suec. 28: 10 (1950)

= Coeloglossum viride subsp. islandicum (Lindl.) Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 150 (2007)

= Coeloglossum viride subsp. islandicum (Lindl.) Selander; Acta Phytogeogr. Suec. 28: 10 (1950)

= Coeloglossum viride var. akaishimontanum Satomi; J. Jap. Bot. 44: 127 (1969)

= Coeloglossum viride var. bracteatum (Muhl. ex Willd.) A.Gray; Manual, ed. 5: 500 (1867)

= Coeloglossum viride var. bracteatum (Muhl. ex Willd.) K.Richt.; Pl. Eur. 1: 278 (1890)

= Coeloglossum viride var. brevibracteatum Asch. & Graebn.; 807 (1907)

= Coeloglossum viride var. collinum Rupp ex W.Zimm.; Form. Orch. Deut. 61 (1912)

= Coeloglossum viride var. dentatum Zapal.; 219 (1906)

= Coeloglossum viride var. grandiflorum Zapal.; 219 (1906)

= Coeloglossum viride var. interjecta (Fernald) Miyabe & Kudô; J. Fac. Agric. Hokkaido Imp. Univ. 26: 360 (1932)

= Coeloglossum viride var. islandicum (Lindl.) M.Schultze; Oesterr. Bot. Z. 48: 113 (1898)

= Coeloglossum viride var. labellifidum (Costa ex Vayr.) E.G.Camus; Monogr. Orchid.: 320 (1908)

= Coeloglossum viride var. longibracteatum Asch. & Graebn.; 807 (1907), nom. illeg.

= Coeloglossum viride var. macrobracteatum Schur; 294 (1870)

= Coeloglossum viride var. microbracteatum Schur; 294 (1870)

= Coeloglossum viride var. rhenanum Höppner; Verh. Naturhist. Vereines Preuss. Rheinl. Westphalens 81: 266 (1925)

= Coeloglossum viride var. subalpinum Rupp ex W.Zimm.; Form. Orch. Deut. 61 (1912)

= Coeloglossum viride var. thuringiacum Rupp ex W.Zimm.; Form. Orch. Deut. 61 (1912)

= Coeloglossum viride var. vaillantii (Ten.) Thell.; Schinz & Keller, Fl. Schweiz, ed. 3: 73 (1914)

= Coeloglossum viride var. virescens (Muhl. ex Willd.) Luer; Native Orchids U. S. & Canada excluding Florida 172 (1975)

= Dactylorhiza viridis var. coreana (Nakai) N.S.Lee; Ill. Fl. Korean Orchids 100 (2011)

= Dactylorhiza viridis var. virescens (Muhl. ex Willd.) Baumbach; Orchidee (Hamburg) 64: 137 (2013)

= Entaticus viridis (L.) Gray; Nat. Arr. Brit. Pl. 2: 206 (1821 publ. 1822)

= Gymnadenia bracteata (Muhl. ex Willd.) C.Presl; Reliq. Haenk. 1: 92 (1827)

= Gymnadenia viridis (L.) Rich.; Orch. Eur. Annot. (Mem. Mus. Hist. Nat. Paris 4: 57: 1818) 35 (1817)

= Gymnadenia viridis var. brevibracteata (Bréb.) Bréb.; 259 (1849)

= Habenaria bracteata (Muhl. ex Willd.) R.Br.; W. T. Aiton, Hortus Kew. 5: 192 (1813)

= Habenaria flava var. virescens (Muhl. ex Willd.) Fernald; Rhodora 23: 148 (1921)

= Habenaria virescens (Willd.) Spreng.; Syst. 3: 688 (1826)

= Habenaria viridis (L.) R.Br.; W. T. Aiton, Hortus Kew. 5: 192 (1813)

= Habenaria viridis subsp. bracteata (Muhl. ex Willd.) R.T.Clausen; Cornell Univ. Agric. Exp. Sta. Mem. 291: 11 (1949)

= Habenaria viridis var. bracteata (Muhl. ex Willd.) A.Gray; Manual, ed. 5: 500 (1867)

= Habenaria viridis var. interjecta Fernald; Rhodora 28: 173 (1926)

= Herminium nankotaizanense Masam.; J. Soc. Trop. Agric., Taiwan 4: 194 (1932)

= Himantoglossum viride (L.) Rchb.; Fl. Germ. Excurs. 120 (1830)

= Orchis batrachites Schrank; Baier. Reise 86 (1786)

= Orchis bractealis Salisb.; Parad. Lond. 2: t. 110 (1808), nom. superfl.

= Orchis bracteata Muhl. ex Willd.; Sp. Pl. 4(1): 34 (1805)

= Orchis coreana Nakai; Bot. Mag. (Tokyo) 28: 302 (1914)

= Orchis ferruginea (F.W.Schmidt) F.W.Schmidt; Samml. Phys.-Oekon. Aufsätze 1: 238 (1795)

= Orchis flava var. virescens (Muhl. ex Willd.) Green; Trans. Soc. Promot. Useful Arts 3: 120 (1814)

= Orchis obsoleta Muhl. ex Willd.; Sp. Pl. ed. 4. 4: 34 (1805)

= Orchis virens Scop.; Fl. Carniol., ed. 2, 2: 199 (1771)

= Orchis virescens (L.) Muhl. ex Willd.; Sp. Pl. 4: 37 (1805)

= Orchis viridis (L.) Crantz; Stirp. Austr. ed. II, fasc. 6: 491 (1769)

= Orchis viridis var. major Tinant; Fl. Luxemb. 442 (1836)

= Orchis viridis var. vaillantii Ten.; Syll. Pl. Neapol. 629 (1831)

= Peristylus bracteatus (Muhl. ex Willd.) Lindl.; Gen. Sp. Orchid. Pl. 298 (1835)

= Peristylus bracteatus f. major Franch. & Sav.; Enum. Pl. Jap. 2: 31 (1879)

= Peristylus islandicus Lindl.; Gen. Sp. Orchid. Pl. 297 (1835)

= Peristylus montanus Schur; Enum. Pl. Transs. 645 (1866)

= Peristylus virescens (Muhl. ex Willd.) Lindl.; Gen. Sp. Orchid. Pl.: 298 (1835)

= Peristylus viridis (L.) Lindl.; Syn. Brit. Fl. ed. II, 261 (1835)

= Peristylus viridis var. bracteata (Muhl. ex Willd.) Rchb.f. ex Kurtz; Bot. Jahrb. Syst. 19: 408 (1894)

= Peristylus viridis var. macrobracteatus Schur; Enum. Pl. Transsilv. 645 (1866)

= Perularia virescens (Muhl. ex Willd.) A.Gray; Bot. Gaz. 5: 63 (1880)

= Platanthera bracteata (Muhl. ex Willd.) Torr.; Fl. N. York 2: 279 (1843)

= Platanthera coreana (Nakai) Nakai; Fl. M´t. Paik-Tu-San 57 (1918)

= Platanthera islandica (Lindl.) Kraenzl.; Repert. Spec. Nov. Regni Veg., Beih. 65: 53 (1931)

= Platanthera nankotaizanensis (Masam.) Masam.; Bot. Mag. (Tokyo) 46: 773 (1932)

= Platanthera viridis (L.) Lindl.; Syn. Brit. Fl., ed. I, 261 (1829)

= Platanthera viridis var. bracteata (Muhl. ex Willd.) Rchb.f.; Icon. Fl. Germ. Helv. (H. G. L. Reichenbach) 13/14(2): 130, t. 83 (- t. 435) (1851)

= Platanthera viridis var. densiflora Regel; Bull. Soc. Imp. Naturalistes Moscou 41(I): 430 (1868)

= Satyrium alpinum F.W.Schmidt; Fl. Boem. 1: 63 (1793)

= Satyrium bracteale Salisb.; Trans. Hort. Soc. 1: 290 (1812)

= Satyrium bracteatum (Muhl. ex Willd.) Pers.; Syn. 2: 507 (1807) [non Lindl.]

= Satyrium ferrugineum F.W.Schmidt; Fl. Boem. 1: 62 (1793)

= Satyrium fuscum Huds.; Fl. Angl. ed. I, 337 (1762)

= Satyrium lingulatum Vill.; Hist. Pl. Dauph. 2: 42 (err. typ. 24) (1787)

= Satyrium virescens (Muhl. ex Willd.) Pers.; Syn. 2: 507 (1807)

= Satyrium viride L.; Sp. Pl. 944 (1753)

= Sieberia viridis (L.) Spreng.; Anleit. Kenntn. Gew., ed. 2, 2(1): 282 (1817)

species :

Dactylorhiza vironii Kreutz; Ber. Arbeitskreis. Heimische Orchid. 24(1): 90 (2007)

Distribution: Estonia

14 EST

Remarks: contains the Baltic populations of "Dactylorhiza ruthei". Status unconfirmed

species :

Dactylorhiza ×abantiana H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(2): 261 (1981)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x nieschalkiorum

species :

Dactylorhiza ×abeliana (Klotzsch) Peitz; Jahresber. Naturwiss. Ver. Wuppertal 25: 173 (1972)

Distribution: Europe

Synonyms: hidden

= Orchis traunsteineri f. abeliana Klotzsch

= Orchis angustifolia var. abeliana Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 87 (1899)

species :

Dactylorhiza ×aldenii H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 74 (1983)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza iberica x (majalis ssp.) kalopissii

species :

Dactylorhiza ×altobracensis (Coste & Soulié) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

Distribution: C-Europe; Austria

11 AUT

Remarks: Dactylorhiza fuchsii x sambucina

Synonyms: hidden

= Dactylorhiza ×influenza (Sennholz) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis ×altobracensis Coste & Soulié; Bull. Soc. Bot. France 44(Sess. Extraord.): cxvii (1898)

= Orchis ×influenza Sennholz; Verh. zool.-bot. Ges. Wien 12, Sitzb. 40 (1891), et Oesterr. Bot. Z. 41: 381 (1891)

species :

Dactylorhiza ×alutiiqorum (A.Baum & H.Baum) J.M.H.Shaw; Orchid Rev. Suppl., 122(1306): 37 (2014)

Distribution: Aleutes

70 ASK

Remarks: Dactylorhiza viridis x D. aristata

Synonyms: hidden

= Dactylorhiza ×alutiiqorum (A.Baum & H.Baum) Eccarius; Orchideengattung Dactylorhiza 580 (2016), isonym

= xDactyloglossum ×alutiiqorum A.Baum & H.Baum; Native Orchid Conf. J. 6(2): 3 (2009)

species :

Dactylorhiza ×ambigua (A.Kern.) H.Sund.

Distribution: Europe; Germany (?Baden-Württemberg, ?Bayern, Rheinland-Pfalz, Schleswig-Holstein)

11 GER

Remarks: Dactylorhiza incarnata x Dactylorhiza maculata

Synonyms: hidden

= Dactylorhiza ×carnea nothosubsp. maculatiformis (Rouy) Gathoye & D.Tyteca; Lejeunia 143: 63 (1994)

= Dactylorhiza ×claudiopolitana nothosubsp. maculatiformis (Rouy) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Dactylorhiza ×maculatiformis (Rouy) Borsos & Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orch. Pannon. & Karpat. Fl. 5) 81 (1961)

= Orchis ×maculatiformis Rouy; Rouy & Foucaud, Fl. France 13: 174 (1912)

= Orchis ×ambigua A.Kern.; Verh. K. K. Zool.-Bot. Ges. Wien 15: 205 (1865), nom. illeg.

species :

Dactylorhiza ×aschersoniana (Hausskn.) Borsos & Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol., 3 (Geobot. Monogr. Orch. Pannon. & Karpat. Fl. IV.) 113 (1960) [nothossp.]

subspecies :

subsp. aschersoniana

Distribution: Europe (widespread); European Russia

14 RUS

Remarks: Dactylorhiza incarnata x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorchis ×pseudotraunsteineri (A.Fuchs) Verm.; Stud. Dactylorch.: 66 (1947)

= Dactylorhiza ×bavarica (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×eifliaca (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×gennachensis (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×koningweeniana (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

= Dactylorhiza ×pseudojunialis R.Doll; Feddes Repert. 89: 346 (1978)

= Orchis ×aschersoniana Hausskn.; Mitt. Geogr. Ges. (Thüringen) Jena 2: 223 (1884 publ. 1885)

= Orchis ×pseudotraunsteineri A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 24 (1919)

= Orchis ×pseudotraunsteineri subsp. bavarica A.Fuchs; Ber. Naturwiss. Ver. Schwaben 42: 100 (1919)

= Orchis ×pseudotraunsteineri subsp. eifliaca A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 154 (1919)

= Orchis ×pseudotraunsteineri var. gennachensis A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 141 (1919)

= Orchis ×pseudotraunsteineri var. koningweeniana A.Fuchs; Ber. Naturwiss. Vereins Schwaben Neuburg 42: 158 (1919)

variant :

var. mulignensis (Gsell) Kümpel; Ber. Arbeitskr. Heim. Orch. 18(2): 23 (2001 publ. 2002) [nothovar.]

Distribution: Switzerland

11 SWI

Remarks: Dactylorhiza incarnata var. serotina x Dactylorhiza majalis

Synonyms: hidden

= Dactylorhiza ×mulignensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis ×mulignensis Gsell; Jahresber. Naturf. Ges. Graubündens, n.s., 74: 17 (1936)

variant :

var. uliginosa Kümpel; Ber. Arbeitskr. Heim. Orch. 18(2): 24 (2001 publ. 2002) [nothovar.]

Distribution: Germany

11 GER

Remarks: Dactylorhiza incarnata var. serotina x Dactylorhiza majalis ssp. brevifolia

variant :

var. wisniewskii (Hemke) Gathoye & D.Tyteca; Lejeunia 143: 61 (1994) [nothovar.]

Distribution: Czech Republic; Slovakia

11 CZE SLO

Remarks: Dactylorhiza incarnata ssp. haematodes x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×aschersoniana nothosubsp. wiesniewskii (Hemke) Potucek; Kl´úc na Urcovanie Vstavacovit ch Ceskoslovenska (Rosalia, Mimor. Vyd.): 135 (1990)

= Dactylorhiza ×wisniewskii Hemke; Mitt. Arbeitskr. Heim. Orch. Zentr. Fachausschusses Bot. Kulturb. DDR 8: 51 (1978), contrary to Art. 37.3 ICBN (1994)

= Dactylorhiza ×aschersoniana nothovar. wisniewskii Kümpel; Ber. Arbeitskr. Heim. Orch. 18(2): 24 (2001 publ. 2002)

subspecies :

subsp. hochreutineriana (K.Hellm.) Gathoye & D.Tyteca; Lejeunia 143: 61 (1994) [nothossp.]

Distribution: Switzerland

11 SWI

Synonyms: hidden

= Dactylorhiza ×hochreutinerana (K.Hellm.) Soó; Feddes Repert. 83: 159 (1972)

= Dactylorhiza ×hochreutineriana (K.Hellm.) Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 92 (1986), isonym

= Orchis ×hochreutineriana K.Hellm.; Boissiera 7: 387 (1943)

subspecies :

subsp. ishorica (Aver.) Eccarius; Orchideengattung Dactylorhiza 585 (2016) [nothossp.]

Distribution: European Russia

14 RUS

Remarks: Dactylorhiza incarnata x baltica ssp. longifolia

Synonyms: hidden

= Dactylorhiza ×ishorica Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 93 (1986)

subspecies :

subsp. templinensis (Potucek) H.Kretzschmar; H. Baumann, Orchideen Deutschlands 696 (2005) [nothossp.]

Distribution: Germany

11 GER

Remarks: Dactylorhiza incarnata ssp. ochroleuca x majalis

Synonyms: hidden

= Dactylorhiza ×aschersoniana nothovar. templinensis (Potucek) Kümpel; Ber. Arbeitskr. Heim. Orch. 18(2): 24 (2001 publ. 2002)

= Dactylorhiza ×templinensis Potucek; Gleditschia 3: 31 (1975)

species :

Dactylorhiza ×baicalica Aver.; Bot. Zhurn. (Moscow & Leningrad) 71(1): 92 (1986)

Distribution: Siberia (Buryatia)

30 SIB

Remarks: Dactylorhiza (incarnata ssp.) cruenta x Dactylorhiza salina

species :

Dactylorhiza ×balabaniana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 75 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x urvilleana

Synonyms: hidden

= Dactylorhiza ×balabaniana nothosubsp. mizabensis B.Baumann & H.Baumann; J. Eur. Orch. 37: 254 (2005)

species :

Dactylorhiza ×bayburtiana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 76 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza euxina x umbrosa (incarnata ssp. cilicica)

species :

Dactylorhiza ×beckeriana (Höppner) Soó; Nom. Nov. Gen. Dactylorhizae 6 (1962)

Distribution: Germany; Switzerland

11 GER SWI

Remarks: Dactylorhiza incarnata x D. maculata x D. majalis

Synonyms: hidden

= Dactylorchis ×beckeriana (Höppner) Verm.; Stud. Dactylorch.: 68 (1947)

= Dactylorhiza ×genevensis (Klinge ex M.Schultze) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

= Orchis ×beckeriana Höppner; Sitz. Niederrh. Ges. Natur.-u. Heilk., D. 3 (1926 publ. 1927)

= Orchis incarnata var. genevensis Klinge ex Schulze; Mitth. Thüring. Bot. Vereins n.f. 17: 49 (1902)

= Orchis ×beckeriana Höppner ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 251, 258 (1933)

species :

Dactylorhiza ×boluiana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 76 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza maculata ssp. saccifera x D. majalis ssp. nieschalkiorum

species :

Dactylorhiza ×braunii (Halácsy) Borsos & Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orch. Pannon. & Karpat. Fl. 5): 82 (1961) [nothossp.]

subspecies :

subsp. braunii

Distribution: Europe; European Russia; Germany (?Bayern, ?Mecklenburg-Vorpommern, Rheinland-Pfalz, ?Schleswig-Holstein, ?Sachsen); Czech Republic

11 CZE GER 14 RUS

Remarks: Dactylorhiza fuchsii ssp. fuchsii x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×braunii nothovar. alpina (E.G.Camus) Soó; Nom. Nov. Gen. Dactylorhiza: 9 (1962)

= Dactylorhiza ×dinglensis nothosubsp. braunii (Halácsy) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 1 (2019)

= Orchis ×braunii nothovar. alpina E.G.Camus; Camus, Iconogr. Orchid. Europe 1: 279 (1927)

= Orchis ×braunii Halácsy; Oesterr. Bot. Z. 31: 137 (1881)

subspecies :

subsp. lilacina F.Proch.; Preslia 54(4): 291 (1982) [nothossp.]

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza fuchsii ssp. fuchsii x Dactylorhiza majalis ssp. turfosa

Synonyms: hidden

= Dactylorhiza ×braunii nothosubsp. lilacina (F.Proch.) Holub; Preslia 70(2): 103 (1998)

= Dactylorhiza ×braunii nothovar. lilacina F.Proch.; Preslia 54(4): 291 (1982)

subspecies :

subsp. monticola Potucek; Preslia 48(2): 134 (1976) [nothossp.]

Distribution: Slovakia

11 SLO

Remarks: Dactylorhiza fuchsii ssp. psychrophila x Dactylorhiza majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×braunii nothosubsp. monticola (Potucek) Oddone; GIROS Orch. Spont. Eur. 59: 355 (2016)

= Dactylorhiza ×braunii nothovar. monticola Potucek; Preslia 48: 134 (1976)

= Dactylorhiza ×monticola (Potucek) Gathoye & D.Tyteca; Lejeunia 143: 65 (1994)

= Dactylorhiza ×townsendiana nothosubsp. monticola (Potucek) Eccarius; Orchideengattung Dactylorhiza: 587 (2016)

subspecies :

subsp. smitakii Batousek; J. Eur. Orchid. 29(4): 643 (1997) [nothossp.]

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza fuchsii ssp. sooana x majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×dinglensis nothosubsp. smitakii (Batousek) Batousek, Taraska & Trávn.; Pl. Syst. Evol. 307(4)-51: 18 (2021)

species :

Dactylorhiza ×breviceras Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 269 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza osmanica x urvilleana

species :

Dactylorhiza ×carnea Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: Germany (Niedersachsen); Czech Republic; France; European Russia

11 CZE GER 12 FRA 14 RUS

Remarks: Dactylorhiza elodes x Dactylorhiza incarnata

Synonyms: hidden

= Dactylorhiza ×claudiopolitana nothosubsp. carnea (Soó) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Orchis ×carnea E.G.Camus; J. Bot. (Morot) 6: 419 (1892), nom. illeg.

species :

Dactylorhiza ×conigera (Norman) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 267 (2011 publ. 2012)

Distribution: Europe; Austria

11 AUT

Remarks: Dactylorhiza viridis x Dactylorhiza maculata

Synonyms: hidden

= Coeloglossum ×conigerum Norman; Norges Arktiske Fl. 1: Sp. Pl. 1050 (1900)

= Dactylorhiza ×conigera nothosubsp. evae (R.Breiner) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

= Orchis ×dominiana E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 322 (1908)

= Orchis ×ullmanii P.M.Hall; Bot. Soc. Exch. Club Brit. Isles 8: 790 (1928 publ. 1929), nom. provis.

= xCoeloglossorchis ×dominiana (E.G.Camus) P.Fourn.; Quatre Fl. France 201 (1935)

= xDactyloglossum ×dominianum (E.G.Camus) Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol. 8: 317 (1966)

= xDactyloglossum ×conigerum (Norman) Rauschert; Feddes Repert. 83 (9-10): 658 (1972 publ. 1973)

= xDactyloglossum ×evae R.Breiner; Ber. Arbeitskr. Heim. Orch. 18(2): 90, 89 (2001 publ. 2002)

= xDactyloglossum ×ullmanii (Hall) Peitz; Jahresber. Naturwiss. Ver. Wuppertal 25: 189 (1972)

= xOrchicoeloglossum ×conigerum (Norman) Nordh.; Norsk Fl. (Oslo): 65 (1940)

= xOrchicoeloglossum ×dominianum E.G.Camus; Monogr. Orch. 322 (1908)

species :

Dactylorhiza ×csatoi (Soó) Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orch. Pannon. & Karpat. Fl. 5) 81 (1961)

Distribution: Romania

13 ROM

Remarks: Dactylorhiza (majalis ssp.) cordigera x maculata

Synonyms: hidden

= Orchis ×csatoi Soó; Repert. Spec. Nov. Regni Veg. 24: 36 (1927)

species :

Dactylorhiza ×daunia W.Rossi, Ard., Cianchi, Piemont. & Bullini; Webbia 50(1): 11 (1995)

Distribution: Italy

13 ITA

Remarks: Dactylorhiza maculata ssp. saccifera x D. majalis ssp. pindica

Synonyms: hidden

= Dactylorhiza ×kerasovinensis B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 3(2): 217 (1986 publ. 1987)

species :

Dactylorhiza ×delamainii (G.Keller ex T.Stephenson) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: France

12 FRA

Remarks: Dactylorhiza elata x D. maculata

Synonyms: hidden

= Orchis ×delamainii G.Keller ex T.Stephenson; Bot. Soc. Exch. Club Brit. Isles 8: 494 (1927 publ. 1928)

species :

Dactylorhiza ×dinglensis (Wilmott) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962) [nothossp.]

subspecies :

subsp. dinglensis

Distribution: Ireland

10 IRE

Remarks: Dactylorhiza maculata ssp. ericetorum x majalis ssp. occidentalis

Synonyms: hidden

= Orchis ×dinglensis Wilmott; Proc. Linn. Soc. Lond. Sess. 148. 128 (1936)

subspecies :

subsp. robertsii F.Horsman; Watsonia 18(4): 397 (1991) [nothossp.]

Distribution: England

10 GRB

Remarks: Dactylorhiza maculata ssp. ericetorum x majalis ssp. cambrensis

Synonyms: hidden

= Dactylorhiza ×formosa nothovar. robertsii (F.Horsman) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

species :

Dactylorhiza ×drucei (A.Camus) Eccarius; Orchideengattung Dactylorhiza 589 (2016)

Distribution: England; Czech Republic; etc.

10 GRB 11 CZE

Remarks: Dactylorhiza viridis x Dactylorhiza majalis

Synonyms: hidden

= Dactylorhiza drucei (A.Camus) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

= Orchis tschiebozi G.Keller; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 196 (1932)

= xDactyloglossum ×drucei (A.Camus) Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol. 8: 317 (1966)

= xOrchicoeloglossum ×drucei A.Camus; E. G. Camus & A. Camus, Icon. Orchid. Eur. 377 (1928)

species :

Dactylorhiza ×dubreuilhii (G.Keller & Jeanj.) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: France

12 FRA

Remarks: Dactylorhiza elata (sesquipedalis) x incarnata

Synonyms: hidden

= Dactylorhiza ×bourdonii D.Tyteca & Gathoye; L´Orchidophile 81: 59 (1988)

= Orchis ×dubreuilhii G.Keller & Jeanj.; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 253 (1933)

species :

Dactylorhiza ×dufftii (Hausskn.) Peitz; Jahresber. Naturwiss. Ver. Wuppertal 25: 176 (1972)

subspecies :

subsp. dufftii

Distribution: Austria; Czech Republic; France; European Russia

11 AUT CZE 12 FRA 14 RUS

Remarks: Dactylorhiza incarnata x D. lapponica

Synonyms: hidden

= Dactylorhiza angustata (Arv.-Touv.) D.Tyteca & Gathoye; L´Orchidophile 97: 115 (1991)

= Dactylorhiza delphinensis D.Tyteca & Gathoye; Naturalistes Belges 69(2): 52 (1988)

= Dactylorhiza incarnata nothosubsp. dufftii (Hausskn.) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 269 (2011 publ. 2012) [nothosubsp.]

= Dactylorhiza lapponica subsp. angustata (Arv.-Touv.) Kreutz; Kompend. Eur. Orchid. 44 (2004)

= Dactylorhiza ×approximata (Rouy) P.Delforge; Naturalistes Belges 89(Orchid. 21): 250 (2008)

= Dactylorhiza ×dufftiana (M.Schultze) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

= Orchis angustata Arv.-Touv.; Ess. Pl. Dauph. 62 (1872)

= Orchis divaricata Boreau; Fl. Centr. Fr. ed. II, 522 (1849), ed. III 2: 646 (1857)

= Orchis strictiflora Opiz; Natural. 10: 217 (1825)

= Orchis ×approximata Rouy; Fl. France [Rouy & Foucaud] 13: 172, hybr. (1912)

= Orchis ×dufftiana M.Schultze; Orch. Deutschl. sub No. 21 [p. 9] (1894)

= Orchis ×dufftii Hausskn.; Mitt. Geogr. Ges. (Thüringen) Jena 2: 225 (1884 publ. 1885)

subspecies :

subsp. gotlandica (Kreutz) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016) [nothossp.]

Distribution: Sweden

10 SWE

Remarks: Dactylorhiza incarnata ssp. ochroleuca x D. lapponica

Synonyms: hidden

= Dactylorhiza ×gotlandica Kreutz; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 25(4): 436 (1993)

species :

Dactylorhiza ×elcitoi Benigni, Barigelli & Petroselli; Micol. Marche, N. S. 5(2): 41 (2021)

Distribution: Italy

13 ITA

species :

Dactylorhiza ×erdingeri (A.Kern.) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 267 (2011 publ. 2012)

Distribution: Austria; Czech Republic; France; etc.

11 AUT CZE 12 FRA

Remarks: Dactylorhiza viridis x Dactylorhiza sambucina

Synonyms: hidden

= Coeloglossum ×erdingeri A.Kern.; Oesterr. Bot. Z. 14: 140 (1864)

= Dactylorhiza ×erdingeri f. elongata (E.G.Camus) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 267 (2011 publ. 2012)

= Orchis ×erdingeri (A.Kern.) Sennholz; Verh. K. K. Zool.-Bot. Ges. Wien 41(Sitzb.): 41 (1891)

= Platanthera ×erdingeri (A.Kern.) A.Kern.; Verh. K. K. Zool.-Bot. Ges. Wien 15: 229 (1865)

= xCoeloglossorchis ×erdingeri (A.Kern.) Guétrot; Pl. Hybr. France 2: 57 (1926 publ. 1927)

= xDactyloglossum ×sennholzii Peitz; Jahresber. Naturwiss. Ver. Wuppertal 25: 189 (1972), nom. nov.

= xDactyloglossum ×erdingeri (A.Kern.) Janch. ex Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol. 8: 317 (1966)

= xDactyloglossum ×erdingeri nothof. elongatum (E.G.Camus) Gathoye & D.Tyteca; Lejeunia 143: 72 (1994)

= xOrchicoeloglossum ×erdingeri (A.Kern.) Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 849 (1907)

= xOrchicoeloglossum ×erdingeri var. elongatum E.G.Camus; Iconogr. Orchid. Europe 2: 375 (1929)

species :

Dactylorhiza ×estonica (Klinge) Jagiello & Kuusk; Fragm. Flor. Geobot. 34 (34): 328 (1989)

Distribution: Czech Republic; France; Estonia; European Russia; etc.

11 CZE 12 FRA 14 EST RUS

Remarks: Dactylorhiza fuchsii x D. (majalis ssp.) lapponica

Synonyms: hidden

= Dactylorhiza traunsteineri var. estonica (Klinge) Soó; Nom. Nov. Gen. Dactylorhiza: 6 (1962)

= Dactylorhiza ×silvae-gabretae F.Proch.; Preslia 71: 236 (1999)

= Orchis angustifolia var. estonica Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 77 (1899)

species :

Dactylorhiza ×flixensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: Switzerland

11 SWI

Remarks: Dactylorhiza incarnata ssp. pulchella x traunsteineri

Synonyms: hidden

= Orchis ×flixensis Gsell; Jahresb. Naturf. Ges. Graub. (n.f.) 74: 17 (1936)

species :

Dactylorhiza ×formosa (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

Distribution: England

10 GRB

Remarks: Dactylorhiza ericetorum x purpurella

Synonyms: hidden

= Orchis ×formosa T.Stephenson & T.A.Stephenson; J. Bot. 60: 33 (1922), nom. illeg.

species :

Dactylorhiza ×fourkensis B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 4(1): 61 (1987)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza majalis ssp. pindica x D. sambucina

species :

Dactylorhiza ×gabretana (A.Fuchs) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

Distribution: Germany; etc.

11 GER

Remarks: Dactylorhiza incarnata x maculata x sambucina

Synonyms: hidden

= Orchis ×gabretana A.Fuchs; Mitt. Bayer. Bot. Ges. 3: 381 (1917)

species :

Dactylorhiza ×godferyana (Soó) Peitz; Jahresber. Naturwiss. Ver. Wuppertal 25: 177 (1972)

Distribution: England

10 GRB

Remarks: Dactylorhiza majalis x praetermissa

Synonyms: hidden

= Orchis ×godferyana Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 257 (1933)

= Dactylorhiza majalis nothosubsp. godferyana (Soó) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 3 (2019)

species :

Dactylorhiza ×grandis (Druce) P.F.Hunt; Orchid Rev. 79 (935): 139 (1971) [nothossp.]

subspecies :

subsp. grandis

Distribution: England; France; Channel Isl.

10 GRB 12 CIS FRA

Remarks: Dactylorhiza fuchsii x D. praetermissa

Synonyms: hidden

= Dactylorhiza ×dinglensis nothosubsp. grandis (Druce) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 1 (2019)

subspecies :

subsp. robustior (Hartm.) J.M.H.Shaw; Orchid Rev. 125(1319, Suppl.): 57 (2017) [nothossp.]

Distribution: England; etc.

10 GRB

Remarks: Dactylorhiza fuchsii x D. majalis ssp. elatior

Synonyms: hidden

= Dactylorhiza ×mortonii (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis angustifolia f. robustior (Hartm.) Klinge; Trudy Imp. S.-Peterburgsk. Bot. Sada 17(2): 83 (1899)

= Orchis latifolia var. robustior Hartm.; Handb. Skand. Fl., ed. 3: 204 (1838)

= Orchis ×grandis Druce; Fl. Buckinghamshire 328 (1926)

= Orchis ×mortonensis E.G.Camus & A.Camus; Iconogr. Orchid. Europe: 280 (1928)

= Orchis ×mortonii Druce; Rep. Bot. Exch. Cl. Brit. Isles 8: 419 (1927 publ. 1928)

species :

Dactylorhiza ×guilhotii (E.G.Camus) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 268 (2011 publ. 2012)

Distribution: France

12 FRA

Remarks: Dactylorhiza viridis x Dactylorhiza incarnata

Synonyms: hidden

= xCoeloglossorchis ×guilhotii (E.G.Camus) P.Fourn.; Quatre Fl. France 201 (1935)

= xDactyloglossum ×guilhotii (E.G.Camus) Soó; Ann. Univ. Sci. Budapest. Rolando Eötvös, Sect. Biol. 8: 317 (1966)

= xOrchicoeloglossum ×guilhotii E.G.Camus; E. G. Camus et al., Monogr. Orchid.: 321 (1908)

species :

Dactylorhiza ×guillaumeae C.Bernard; Bull. Soc. Bot. France, Lett. Bot. 130(2): 154 (1983)

Distribution: France

12 FRA

Remarks: Dactylorhiza incarnata x sambucina

species :

Dactylorhiza ×gustavssonii H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 77 (1983)

Distribution: Greece; Turkey

13 GRC 34 TUR

Remarks: Dactylorhiza iberica x saccifera

Synonyms: hidden

= Dactylorhiza ×sultandagi Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 270 (1983)

species :

Dactylorhiza ×hallii (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962) [nothossp.]

subspecies :

subsp. hallii

Distribution: England; Channel Isl.

10 GRB 12 CIS

Remarks: Dactylorhiza maculata ssp. ericetorum x praetermissa

Synonyms: hidden

= Dactylorhiza ×dinglensis nothosubsp. hallii (Druce) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 2 (2019)

= Orchis ×hallii Druce; Rep. Bot. Exch. Cl. Brit. Isles 4: 24 (1914 publ. 1915)

subspecies :

subsp. nummiana (P.Fourn.) Gathoye & D.Tyteca; Lejeunia 143: 67 (1994) [nothossp.]

Distribution: France

12 FRA

Remarks: Dactylorhiza maculata ssp. elodes x praetermissa

Synonyms: hidden

= Dactylorhiza ×nummiana (P.Fourn.) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

= Orchis ×nummiana P.Fourn.; Bull. Soc. Bot. France 78: 433 (1931)

species :

Dactylorhiza ×hepburnii (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: Scotland

10 GRB

Remarks: Dactylorhiza foliosa x D. incarnata x D. fuchsii

Synonyms: hidden

= Orchis ×hepburnii Druce; Rep. Bot. Exch. Cl. Brit. Isles 4: 211 (1915 publ. 1916)

species :

Dactylorhiza ×insignis (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

Distribution: England

10 GRB

Remarks: Dactylorhiza praetermissa x purpurella

Synonyms: hidden

= Dactylorhiza ×salteri (T.Stephenson) Soó; Jahresber. Naturwiss. Ver. Wuppertal 21-22: 18 (1968)

= Orchis ×insignis T.Stephenson & T.A.Stephenson; J. Bot. 60: 33. hybr. (1922)

= Orchis ×salteri T.Stephenson; J. Bot. Lond. 80: 104 (1942)

species :

Dactylorhiza ×jenensis (Brand) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

Distribution: Europe; Germany; France

11 GER 12 FRA

Remarks: Dactylorhiza maculata ssp. ericetorum x traunsteineri (not lapponica)

Synonyms: hidden

= Orchis ×jenensis Brand; W. D. J. Koch, Syn. Deut. Schweiz. Fl., ed. 3, 3: 2430 (1904)

= Orchis ×schulzei C.Richt.; Pl. Eur. 1: 274 (1890), et J. Bot. (Morot) 6: 418 (1892)

species :

Dactylorhiza ×jestrebiensis Businsky; Preslia 61(4): 306 (1989)

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza traunsteineri ssp. bohemica x Dactylorhiza majalis ssp. majalis

species :

Dactylorhiza ×juennensis Perko; Carinthia II, 187/107(1): 91 (1997)

Distribution: Austria

11 AUT

Remarks: Dactylorhiza traunsteineri ssp. rhaetica (not lapponica) x fuchsii

species :

Dactylorhiza ×katarana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 78 (1983)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza maculata ssp. saccifera x D. majalis ssp. kalopissii

species :

Dactylorhiza ×kelleriana (Cif. & Giacom.) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: England; Ireland; etc.

10 GRB IRE

Remarks: Dactylorhiza fuchsii x Dactylorhiza traunsteineri

Synonyms: hidden

= xOrchicoeloglossum ×kellerianum Duffort ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 299 (1935), et adnot.

species :

Dactylorhiza ×kerneriorum (Soó) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962) [nothossp.]

subspecies :

subsp. kerneriorum

Distribution: Europe; Germany; Czech Republic; Slovakia; European Russia; etc.

11 CZE GER SLO 14 RUS

Remarks: Dactylorhiza fuchsii x incarnata

Synonyms: hidden

= Dactylorhiza ×claudiopolitana (Simonk.) Borsos & Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orch. Pannon. & Karpat. Fl. 5) 81 (1961)

= Dactylorhiza ×pomeranica Wucherpf.; J. Eur. Orch. 44(4): 700 (2012)

= Orchis ×ambigua var. claudiopolitana Simonk.; Enum. Fl. Transsilv. 500 (1887)

= Orchis ×curtisii Druce; Bot. Soc. Exch. Club Brit. Isles 9: 669 (1931 publ. 1932)

= Orchis ×kerneriorum Soó; Notizbl. Bot. Gart. Berlin-Dahlem 9: 904 (1926)

= Orchis ×maculatoformis Rouy; Fl. France 13: 174 (1912)

= Orchis ×maculotoformis nothomorph. claudiopolitana (Simonk.) Pauca; Fl. Republ. Social. Roman. 12: 707 (1972)

= Orchis ×rigida Höppner; Verh. Nat. Ver. Preuss. Rheinl. u. Westf. 1924, 270-273

subspecies :

subsp. ampolai (Hautz.) Eccarius; Orchideengattung Dactylorhiza 581 (2016) [nothossp.]

Distribution: Italy

13 ITA

Remarks: Dactylorhiza cruenta x maculata

Synonyms: hidden

= Dactylorhiza ×ampolai Hautz.; Verh. zool.-bot. Ges. Wien 112: 115 (1972)

= Dactylorhiza ×kerneri nothosubsp. ampolai (Hautz.) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

= Dactylorhiza ×kerneriorum nothosubsp. ampolai (Hautz.) Eccarius; Orchideengattung Dactylorhiza: 581 (2016)

subspecies :

subsp. lillsundica (Kreutz) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016) [nothossp.]

Distribution: Sweden

10 SWE

Remarks: Dactylorhiza fuchsii x incarnata ssp. ochroleuca

Synonyms: hidden

= Dactylorhiza ×lillsundica Kreutz; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 25(4): 435 (1993)

subspecies :

subsp. variabilis (Hesl.-Harr.) P.D.Sell; Fl. Great Britain & Ireland 5: 364 (1996) [nothossp.]

Distribution: Scotland

10 GRB

Synonyms: hidden

= Dactylorhiza ×variabilis (Hesl.-Harr.) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis ×variabilis Hesl.-Harr.; J. Bot. Lond. 79: 168 (1941), et Proc. Univ. Durham Phil. Soc. 10: 308 (1941), descr.

species :

Dactylorhiza ×komiensis Aver.; Bot. Zhurn. (Moscow & Leningrad) 67(3): 310 (1982)

Distribution: European Russia

14 RUS

Remarks: Dactylorhiza hebridensis x maculata

species :

Dactylorhiza ×kopdagiana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 78 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x umbrosa

species :

Dactylorhiza ×koutsourana B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 6(1): 36 (1989)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza baumanniana x smolikana

species :

Dactylorhiza ×krylovii (Soó) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962)

Distribution: E-European Russia; Siberia (W-Siberia)

14 RUS 30 SIB

Remarks: Dactylorhiza incarnata ssp. cruenta x D. incarnata ssp. incarnata

Synonyms: hidden

= Orchis ×krylovii Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 211 (1933)

species :

Dactylorhiza ×kuehnensis Presser & Riech.; J. Eur. Orchid. 32(1): 109 (2000)

Distribution: Germany

11 GER

Remarks: Dactylorhiza majalis x ruthei

species :

Dactylorhiza ×kuuskiae E.Breiner & R.Breiner; Ber. Arbeitskr. Heim. Orch. 15(2): 34 (1998)

Distribution: Estonia

14 EST

Remarks: Dactylorhiza (majalis ssp.) baltica x vironii

Synonyms: hidden

= Dactylorhiza ×kuehnensis nothosubsp. kuuskiae (E.Breiner & R.Breiner) Eccarius; Orchideengattung Dactylorhiza 588 (2016)

species :

Dactylorhiza ×latirella (P.M.Hall) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: England

10 GRB

Remarks: Dactylorhiza incarnata x (majalis ssp.) purpurella

Synonyms: hidden

= Orchis ×latirella P.M.Hall; J. Bot. 74: 329 (1936)

species :

Dactylorhiza ×megapolitana (Bisse) Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol. 11: 71 (1969)

Distribution: Germany (Mecklenburg-Vorpommern); European Russia

11 GER 14 RUS

Remarks: Dactylorhiza fuchsii x Dactylorhiza russowii

Synonyms: hidden

= Dactylorchis ×megapolitana Bisse; Feddes Repert. 67: 186 (1963)

= Dactylorhiza ×silvae-gabretae nothosubsp. megapolitana (Bisse) Eccarius; Orchideengattung Dactylorhiza 582 (2016)

species :

Dactylorhiza ×metsowonensis B.Baumann & H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 16(1): 128 (1984)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza latifolia x kalopissii

species :

Dactylorhiza ×mixta (Asch. & Graebn.) B.Bock; Bull. Soc. Bot. Centre-Ouest 42: 268 (2011 publ. 2012)

Distribution: England; Slovakia; France; etc.

10 GRB 11 SLO 12 FRA

Remarks: Dactylorhiza viridis x Dactylorhiza fuchsii

Synonyms: hidden

= Habenaria ×mixta (Asch. & Graebn.) Druce; Bot. Soc. Exch. Club Brit. Isles 8: 876 (1928 publ. 1929)

= Orchis ×kaspari Domin & Podp.; Klic K Uplne Kvetene Rep. Ceskoslov. 796 (1928)

= Orchis ×mixta Domin; Sitzungsber. Königl. Böhm. Ges. Wiss., Math.-Naturwiss. Cl. 22: 7 (1902 publ. 1903), nom. illeg.

= xDactyloglossum ×mixtum (Asch. & Graebn.) P.F.Hunt; Orchid Rev. 79 (935): 140 (1971), isonym

= xDactyloglossum ×mixtum (Asch. & Graebn.) Rauschert; Feddes Repert. 79: 413 (1969)

= xOrchicoeloglossum ×mixtum Asch. & Graebn.; Syn. Mitteleur. Fl. 3: 847 (1907)

species :

Dactylorhiza ×nevskii H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(2): 262 (1981)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza osmanica x umbrosa

species :

Dactylorhiza ×ornonensis (G.Keller & Jeanj.) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: France

12 FRA

Remarks: Dactylorhiza elata ssp. sesquipedalis x incarnata x maculata

Synonyms: hidden

= Orchis ×ornonensis G.Keller & Jeanj.; Act. Soc. Linn. Bordeaux 89: Proc.-Verb. 34 (1937)

species :

Dactylorhiza ×panoramica Willing; Mitt. Arbeitskreis Heim. Orch. Baden-Württ. 17(4): 516 (1985)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza baumanniana x saccifera

species :

Dactylorhiza ×paridaeniana Kreutz; Eurorchis 2: 65 (1990)

Distribution: Netherlands

11 NET

species :

Dactylorhiza ×perez-chiscanoi F.M.Vázquez; J. Eur. Orch. 40(1): 36 (2008)

Distribution: Spain

12 SPA

Remarks: Dactylorhiza elata x D. lapponica

species :

Dactylorhiza ×predaensis (Gsell) Soó ex Rauschert; Feddes Repert. 85 (9-10): 656 (1974)

Distribution: Switzerland; European Russia

11 SWI 14 RUS

Remarks: Dactylorhiza (incarnata ssp.) cruenta x Dactylorhiza majalis

Synonyms: hidden

= Dactylorchis ×predaensis Gsell ex Soó in Rothm.; 88 (1963), nom. inval.

= Dactylorhiza ×aschersoniana nothosubsp. predaensis (Gsell) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Dactylorhiza ×predaensis Gsell ex Soó ex Soó; 656 (1974), nom. inval.

= Dactylorhiza ×scampstonii (Druce) Soó; Nom. Nov. Gen. Dactylorhiza 9 (1962)

= Orchis ×predaensis Gsell; Jahresb. Naturf. Ges. Graub. (n.f.) 74: 17 (1936), nom. inval.

= Orchis ×scampstonii Druce; Rep. Bot. Exch. Cl. Brit. Isles 5: 53 (1917 publ. 1918)

species :

Dactylorhiza ×prochazkana Businsky; Preslia 61(4): 305 (1989)

Distribution: Czech Republic

11 CZE

Remarks: Dactylorhiza bohemica x Dactylorhiza maculata ssp. maculata

species :

Dactylorhiza ×reitaluae Hennecke, E.Breiner & R.Breiner; Ber. Arbeitskreis. Heimische Orchid. 15: 36 (1998)

Distribution: Estonia

14 EST

Remarks: Dactylorhiza incarnata x vironii [described as incarnata x ruthei]

species :

Dactylorhiza ×renzii H.Baumann & Künkele; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 13(2): 261 (1981)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza incarnata x nieschalkiorum

species :

Dactylorhiza ×rizeana Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 269 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza euxina x urvilleana

species :

Dactylorhiza ×rombucina (Cif. & Giacom.) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

Distribution: C-Europe

Remarks: Dactylorhiza romana x sambucina

Synonyms: hidden

= Orchis ×rombucina Cif. & Giacom.; Nomencl. Fl. Ital. 1: 165 (1950)

species :

Dactylorhiza ×ruppertii (M.Schultze) Borsos & Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol., 3 (Geobot. Monogr. Orch. Pannon. & Karpat. Fl. 4) 122 (1960)

Distribution: Germany; Czech Republic; Slovakia; etc.

11 CZE GER SLO

Remarks: Dactylorhiza majalis ssp. majalis x Dactylorhiza sambucina

Synonyms: hidden

= Orchis ×ruppertii M.Schultze; Oesterr. Bot. Z. 49: 264 (1899)

species :

Dactylorhiza ×salictina B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 6(1): 35 (1989)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza pindica x smolikana

Synonyms: hidden

= Dactylorhiza ×souflikensis nothosubsp. salictina (B.Willing & E.Willing) Eccarius; Orchideengattung Dactylorhiza 580 (2016)

species :

Dactylorhiza ×samnaunensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962)

Distribution: Switzerland; European Russia

11 SWI 14 RUS

Remarks: Dactylorhiza cruenta x Dactylorhiza maculata

Synonyms: hidden

= Orchis ×samnaunensis Gsell; Jahresb. Naturf. Ges. Graub. (n.f.) 74: 17 (1936)

species :

Dactylorhiza ×serbica (H.Fleischm.) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: Serbia

13 SRB

Remarks: Dactylorhiza incarnata x saccifera

Synonyms: hidden

= Dactylorhiza ×naris W.Rossi & Mazzola; Atti Soc. Tosc. Sci. Nat. Pisa, Mem., Ser. B, 89: 271 (1982 publ. 1983)

= Orchis ×serbica H.Fleischm.; Mitt. Naturwiss. Vereines Steiermark 45: 179 (1908 publ. 1909)

species :

Dactylorhiza ×serreana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 79 (1983)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza majalis ssp. cordigera x D. cordigera ssp. bosniaca

species :

Dactylorhiza ×sivasiana Baumann & Künkele ex Renz & Taubenheim; Notes Roy. Bot. Gard. Edinburgh 41(2): 270 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza umbrosa x urvilleana

species :

Dactylorhiza ×sooi (Ruppert ex Soó) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

Distribution: C-Europe

Remarks: Dactylorhiza majalis var. alpestris x fuchsii

Synonyms: hidden

= Orchis ×sooi Ruppert ex Soó; G. Keller & Schltr., Monogr. & Iconogr. Orch. Eur. 2 (Repert. Spec. Nov. Regni Veg.,Sonderbeih. A.) 398 (1936)

species :

Dactylorhiza ×souflikensis B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 4(1): 60 (1987)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza baumanniana x pindica

species :

Dactylorhiza ×stagni-novi D.Tyteca & Gathoye; L´Orchidophile 81: 58 (1988)

Distribution: Spain; France

12 FRA SPA

Remarks: Dactylorhiza brennensis x fuchsii

Synonyms: hidden

= Dactylorhiza ×hjertsonii P.P.Ferrer, J.L.Lozano, R.Roselló, Feliu & Peña-Riv.; Flora Montiber. 71: 100 (2018)

species :

Dactylorhiza ×stenostachys (Murr) Rauschert; Feddes Repert. 85 (9- 101: 656 (1974)

subspecies :

subsp. stenostachys

Distribution: Switzerland; European Russia

11 SWI 14 RUS

Remarks: D. incarnata ssp. incarnata x D. (majalis ssp.) lapponica

Synonyms: hidden

= Dactylorhiza ×thellungiana (Braun-Blanq.) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis instabilis G.Keller ex G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 252 (1933)

= Orchis ×stenostachys Murr; Neue Übersicht Farn- & Blütenpfl. Vorarlb. 281 (1923)

= Orchis ×thellungiana Braun-Blanq.; Veröff. Geobot. Inst. Rübel 7: 349 (1932)

subspecies :

subsp. lehmannii (Klinge) Eccarius; Orchideengattung Dactylorhiza 586 (2016) [nothossp.]

Distribution: Germany (Mecklenburg-Vorpommern); European Russia

11 GER 14 RUS

Remarks: Dactylorhiza incarnata var. longebracteata x Dactylorhiza traunsteineri ssp. russowii

Synonyms: hidden

= Dactylorhiza ×lehmannii (Klinge) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis ×lehmannii Klinge; Rev. Orch. Cordig. Angust. 102 (1893)

subspecies :

subsp. stenkyrkae (Hautz.) Eccarius; Orchideengattung Dactylorhiza 586 (2016) [nothossp.]

Distribution: Sweden; Switzerland; etc.

10 SWE 11 SWI

Remarks: Dactylorhiza incarnata subsp. cruenta x traunsteineri

Synonyms: hidden

= Dactylorhiza ×dufftii nothosubsp. stenkyrkae (Hautz.) Oddone; GIROS Orch. Spont. Eur. 59(2): 355 (2016)

= Dactylorhiza ×engadinensis (Cif. & Giacom.) Soó; Nom. Nov. Gen. Dactylorhizae 8 (1962), nom. inval.

= Dactylorhiza ×stenkyrkae Hautz.; Verh. zool.-bot. Ges. Wien 115: 48 (1976)

= Orchis ×engadinensis Cif. & Giacom.; Nomenclator Fl. Ital. 1: 162 (1950)

species :

Dactylorhiza ×surensis (Gsell) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

Distribution: Switzerland

11 SWI

Remarks: provisionally accepted name

Synonyms: hidden

= Orchis ×surensis Gsell; Jahresb. Naturf. Ges. Graub. (n.f.) 74: 17 (1936)

species :

Dactylorhiza ×szaboiana (Soó) Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol., 4 (Geobot. Monogr. Orch. Pannon. & Karpat. Fl. 5) 81 (1961)

Distribution: Hungary; Romania

11 HUN 13 ROM

Remarks: Dactylorhiza maculata ssp. maculata x D. majalis ssp. cordigera

Synonyms: hidden

= Dactylorhiza ×csatoi nothosubsp. szaboiana (Soó) Eccarius; Orchideengattung Dactylorhiza 580 (2016)

= Orchis ×csatoi var. szaboiana Soó; Repert. Spec. Nov. Regni Veg. 24: 36 (1927)

= Orchis ×szaboiana (Soó) Soó; Arch. Bot. (Leipzig) 23: 125 (1928)

species :

Dactylorhiza ×transiens (Druce) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962) [nothossp.]

subspecies :

subsp. transiens

Distribution: Europe; European Russia; Slovakia; Channel Isl.

11 SLO 12 CIS 14 RUS

Remarks: Dactylorhiza fuchsii x Dactylorhiza maculata

Synonyms: hidden

= Dactylorhiza maculata nothosubsp. transiens (Druce) M.H.J.van der Meer; Cact. Phantast. 2019(5)-1: 2 (2019)

= Orchis ×transiens Druce; Bot. Soc. Exch. Club Brit. Isles 4: 213 (1915 publ. 1916)

subspecies :

subsp. corylensis (Hesl.-Harr.) P.D.Sell; Fl. Great Britain & Ireland 5: 364 (1996) [nothossp.]

Distribution: Scotland

10 GRB

Synonyms: hidden

= Dactylorhiza ×corylensis (Hesl.-Harr.) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis ×corylensis Hesl.-Harr.; J. Bot. Lond. 79: 168 (1941), et Proc. Univ. Durham Phil. Soc. 10: 308 (1941), descr.

species :

Dactylorhiza ×turcestanica (G.Keller & Soó) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

Distribution: Tajikistan

32 TZK

Remarks: Dactylorhiza viridis x D. umbrosa

Synonyms: hidden

= xOrchicoeloglossum ×turcestanicum G.Keller & Soó; Repert. Spec. Nov. Regni Veg. Sonderbeih. A. 2: 299 (1935)

= xDactyloglossum ×turcestanicum (G.Keller & Soó) Soó; Ann. Univ. Sci. Budapest. R. Eötvös, Sect. Biol. 8: 317 (1966)

species :

Dactylorhiza ×vallis-peenae Kümpel; Ber. Arbeitskr. Heim. Orch. 18(2): 28 (2001 publ. 2002)

Distribution: Germany

11 GER

Remarks: Dactylorhiza majalis x Dactylorhiza traunsteineri ssp. russowii

species :

Dactylorhiza ×venusta (T.Stephenson & T.A.Stephenson) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962) [nothossp.]

subspecies :

subsp. venusta

Distribution: England

10 GRB

Remarks: Dactylorhiza fuchsii x purpurella

Synonyms: hidden

= Orchis ×venusta T.Stephenson & T.A.Stephenson; J. Bot. 60: 34 (1922)

subspecies :

subsp. hebridella (Wilmott) P.D.Sell; Fl. Great Britain & Ireland 5: 364 (1996) [nothossp.]

Distribution: W-Scotland (Hebrides)

10 GRB

Remarks: Dactylorhiza fuchsii ssp. hebridensis x purpurella

Synonyms: hidden

= Dactylorhiza ×hebridella (Wilmott) Soó; Nom. Nov. Gen. Dactylorhizae 9 (1962)

= Orchis ×hebridella Wilmott; J. Bot. Lond. 77: 193 (1939)

species :

Dactylorhiza ×vermeuleniana Soó; Jahresber. Naturwiss. Ver. Wuppertal 21-22: 17 (1968), et Acta Bot. Acad. Sci. Hung. 18 (1- 2): 174 (1973)

Distribution: Czech Republic; etc.

11 CZE

Remarks: Dactylorhiza incarnata x maculata x majalis

Synonyms: hidden

= Dactylorhiza ×senayi nothosubsp. vermeuleniana (Soó) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

variant :

var. senayi (Alleiz.) Kümpel; Ber. Arbeitskr. Heim. Orch. 18(2): 28 (2001 publ. 2002) [nothovar.]

Distribution: France

12 FRA

Remarks: Dactylorhiza maculata ssp. elodes x majalis

Synonyms: hidden

= Dactylorhiza ×senayi (Alleiz.) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

= Orchis ×senayi Alleiz.; Bull. Soc. Bot. France 94: 338 (1948)

variant :

var. townsendiana (Rouy) Kümpel; Ber. Arbeitskr. Heim. Orch. 18(2): 29 (2001 publ. 2002) [nothovar.]

Distribution: Germany

11 GER

Remarks: Dactylorhiza maculata ssp. maculata x D. majalis ssp. majalis

Synonyms: hidden

= Dactylorhiza ×senayi nothosubsp. townsendiana (Rouy) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

= Dactylorhiza ×townsendiana (Rouy) Soó; Nom. Nov. Gen. Dactylorhizae 10 (1962)

= Orchis ×braunii var. townsendiana Rouy; Fl. France 13: 173 (1912)

species :

Dactylorhiza ×viridella (Hesl.-Harr.) Oddone; GIROS Orch. Spont. Eur. 59(2): 354 (2016)

Distribution: England

10 GRB

Remarks: Dactylorhiza viridis x D. purpurella

Synonyms: hidden

= Orchis ×viridella Hesl.-Harr.; Vasculum (Substitute) 34: 22 (1949)

= xDactyloglossum ×viridellum (Hesl.-Harr.) Soó; Jahresber. Naturwiss. Ver. Wuppertal 21-22: 18 (1968)

species :

Dactylorhiza ×vitosana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 79 (1983)

Distribution: Bulgaria

13 BUL

Remarks: Dactylorhiza lagotis x saccifera

species :

Dactylorhiza ×vogtiana H.Baumann; Mitt. Arbeitskr. Heim. Orch. Baden-Württemberg 15(1): 80 (1983)

Distribution: Turkey

34 TUR

Remarks: Dactylorhiza iberica x incarnata

species :

Dactylorhiza ×vorasica B.Willing & E.Willing; Ber. Arbeitskr. Heim. Orch. 4(1): 63 (1987)

Distribution: Greece

13 GRC

Remarks: Dactylorhiza cordigera x sambucina

species :

Dactylorhiza ×weissenbachiana Perko; Carinthia II, 184/104(1): 211 (1994)

Distribution: Austria

11 AUT

Remarks: Dactylorhiza incarnata x lapponica

species :

Dactylorhiza ×wiefelspuetziana D.Tyteca; Bull. Soc. Roy. Bot. Belg. 114(1): 24 (1981)

Distribution: Belgium

11 BGM

Remarks: Dactylorhiza maculata ssp. arduennensis x sphagnicola

Synonyms: hidden

= Dactylorhiza ×senayi nothosubsp. wiefelspuetziana (D.Tyteca) Oddone; GIROS Orch. Spont. Eur. 59(2): 356 (2016)

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A complete, synonymic checklist of the Higher Plants of the World

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Genus (740)

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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Contact

How to Cite:

Search

+ Family

Filter

Order

Family (14)

Genus (740)

Display options

A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species, 8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms, in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]