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Ver. 19.2
April 17th, 2024

453.0000 Family Gentianaceae Juss.; Gen. Pl.: 141 (1789), nom. cons.
95 genera, ca. 1850 spp.
Species listed in database: 1852; hybrids: 23
= Chironiaceae Bercht. & J.Presl; Prir. Rostlin 1(104-114): 1, 36 (1823)
= Coutoubeaceae Martinov; Tekhno-Bot. Slovar: 168 (1820)
= Obolariaceae Martinov; Tekhno-Bot. Slovar: 427 (1820)
= Potaliaceae Mart.; Nov. Gen. Sp. Pl. 2: 89, 91, 133 (1827)
= Saccifoliaceae Maguire & Pires; Mem. New York Bot. Gard. 29: 242 (1978)
453.0203 Genus Exacum L.; Sp. Pl. 1: 112 (1753)
Distribution: Africa, Madagascar (39), Oman, India, Indochina, Malesia to New Guinea, N-Australia
Species listed in database: 74
= Chondropis Raf.; Fl. Tellur. 3: 29 (1837)
= Floyera Neck.; Elem. 1: 386 (1790)
= Paracelsea Zoll. & Moritzi; Nat. en Geneesk. Arch. Neer. Indie. 2: 18 (1845)
= Cotylanthera Blume; Bijdr. 707 (1825)
= Eophylon A.Gray; J. Linn. Soc., Bot. 11: 22 (1869)
species :
Exacum affine Balf.f.; Regel, Gartenfl. (1883) 34. t. 1108
Distribution: Soqotra
24 SOC
Synonyms: hidden
= Exacum gracilipes Balf.f.; Proc. Roy. Soc. Edinb. 12: 80 (1884)
species :
Exacum alberti-grimaldii Wohlh. & Callm.; Candollea 67(2): 374 (2012)
Distribution: Madagascar
29 MDG
species :
Exacum amplexicaule Klack.; Op. Bot. 84: 127 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum anamallayanum Bedd.; Ic. Pl. Ind. Or. 1: 33 (1868-1874)
Distribution: India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum atropurpureum var. anamallayanum (Bedd.) C.B.Clarke; Hook.f., Fl. Brit. India [J. D. Hooker] 4: 97 (1883)
= Exacum saulierei Dunn; Bull. Misc. Inform. Kew 1914: 30 (1914)
species :
Exacum anisopterum Klack.; Op. Bot. 84: 132 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum appendiculatum Klack.; Op. Bot. 84: 79 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum arabicum Thulin; Nordic J. Bot. 21(3): 245 (2001)
Distribution: S-Yemen
35 YEM
species :
Exacum atropurpureum Bedd.; Ic. Pl. Ind. Or. 1: 24 (1868-1874)
Distribution: India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum beddomei C.B.Clarke; J. Linn. Soc., Bot. 14: 427 (1875)
= Exacum atropurpureum var. palghatense Gamble; Fl. Madras 874 (1935)
species :
Exacum axillare Thwaites; Enum. Pl. Zeyl. 203 (1859)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Exacum axillare var. pentamera C.B.Clarke; Hook.f., Fl. Brit. India 4: 96 (1885)
species :
Exacum bulbilliferum Baker; J. Linn. Soc., Bot. 20: 209 (1883)
Distribution: Madagascar
29 MDG
species :
Exacum caeruleum Balf.f.; Proc. Roy. Soc. Edinb. 12: 80 (1884)
Distribution: Soqotra
24 SOC
Synonyms: hidden
= Exacum forbesii Balf.f.; H. O. Forbes, Nat. Hist. Sokotra, 487 (1903)
species :
Exacum conglomeratum Klack.; Op. Bot. 84: 81 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum courtallense Arn.; Ann. Sci. Nat., ser. II. 11: 175 (1839)
Distribution: S-India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum courtallense var. boneccordense M.Mohanan; M. Mohanan & A. N. Henry, Fl. Thiruvananthapuram: 305 (1994)
species :
Exacum darae Hul; Edinburgh J. Bot. 67(1): 155 (2010)
Distribution: Thailand; Cambodia; Vietnam
41 CBD THA VIE
species :
Exacum decapterum Klack.; Fl. Madagascar, fam. 168 & 168 bis: 32 (1990)
Distribution: Madagascar
29 MDG
species :
Exacum dipterum Klack.; Op. Bot. 84: 134 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum divaricatum Schinz; Bull. Herb. Boiss., ser. II. 6: 716 (1906)
subspecies :
subsp. divaricatum
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Belmontia divaricata Baker; Bull. Misc. Inform. Kew 1897: 274 (1897)
subspecies :
subsp. latifolium (Klack.) Klack.; Fl. Madagascar, fam. 168 & 168 bis: 58 (1990)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum latifolium Klack.; Op. Bot. 84: 109 (1985)
species :
Exacum dolichantherum Klack.; Op. Bot. 84: 94 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum emirnense (Baker) Schinz; Bull. Herb. Boiss., ser. II. 6: 716 (1906)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Belmontia emirnensis Baker; J. Linn. Soc., Bot. 22: 507 (1887)
species :
Exacum exiguum Klack.; Op. Bot. 84: 87 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum fruticosum Humbert; Mem. Inst. Sci. Madagascar, ser. B, 6: 130 (1955)
Distribution: Madagascar
29 MDG
species :
Exacum giganteum Klack.; Op. Bot. 84: 137 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum gracile Klack.; Op. Bot. 84: 80 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum hoffmannii Schinz; Viertelj. Naturforsch. Gesells. Zürich 36: 327 (1891)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum quinquenervium subsp. hoffmannii (Vatke ex Schinz) Humbert; Adansonia n.s. 4: 31 (1964)
= Exacum quinquenervium var. hoffmannii (Vatke ex Schinz) Humbert; Adansonia n.s. 4: 36 (1964)
species :
Exacum humbertii Klack.; Op. Bot. 84: 118 (1985)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum tsimihety Humbert; Adansonia n. s. 4: 35 (1964) (p. p.)
species :
Exacum idukkianum Geethakum., Pandur., Ravich. & Deepu; Phytotaxa 282(3): 226 (2016)
Distribution: S-India (Kerala)
40 IND
species :
Exacum intermedium Klack.; Op. Bot. 84: 125 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum keralense Geethakum., K.M.P.Kumar, Pandur. & Deepu; Acta Bot. Gallica Bot. Lett. 163(4): 429 (2016)
Distribution: S-India (Kerala)
40 IND
species :
Exacum klackenbergii Gopalan; J. Bombay Nat. Hist. Soc. 99(2): 271 (2002), nom. nov.
Distribution: S-India (Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum wightianum var. uniflorum A.N.Henry & Swamin.; J. Bombay Nat. Rist. Soc. 80: 459 (1983)
species :
Exacum lawii C.B.Clarke; Hook.f. Fl. Brit. Ind. 4: 98 (1883), nom. nov.
Distribution: India (Maharashtra, Kerala, Rajasthan, Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum sessile C.B.Clarke; J. Linn. Soc., Bot. 14: 427 (1875)
species :
Exacum laxiflorum (Gamble) Geethakum., Deepu, Kissling & Pandur.; Phytotaxa 559(2): 190 (2022)
Distribution: S-India
40 IND
Synonyms: hidden
= Exacum courtallense var. laxiflorum Gamble; Fl. Madras 873 (1935)
species :
Exacum linearifolium (Humbert) Klack.; Op. Bot. 84: 76 (1985)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum quinquenervium subsp. linearifolium Humbert; Adansonia n.s. 4: 32 (1964)
species :
Exacum loheri (H.Hara) Klack.; Bot. Jahrb. Syst. 126(4): 478 (2006)
Distribution: Philippines
42 PHI
Synonyms: hidden
= Cotylanthera loheri H.Hara; J. Jap. Bot., 50(11): 327 (1975)
species :
Exacum lokohense Humbert; Mem. Inst. Sci. Madagascar, ser. B, 6: 128 (1955)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum lichenisylvae Humbert; Adansonia n. s. 4: 31 (1964)
= Exacum lichenisylvae var. macranthum Humbert; Adansonia n. s. 4: 31 (1964)
species :
Exacum macranthum Arn. ex Griseb.; Gen. Sp. Gent.: 111 (1838)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Exacum macranthum Arn.; Ann. Sci. Nat., ser. II. 11: 176 (1889)
= Exacum ovale Griseb.; Gen. et Sp. Gent. 111 (1839)
= Exacum trinervium subsp. macranthum (Arn. ex Griseb.) L.H.Cramer; Dassanayake et al., Revis. Handb. Fl. Ceylon 3: 62 (1981)
= Exacum zeylanicum var. macrantha (Arn. ex Griseb.) C.B.Clarke; Hook.f., Fl. Brit. India 4: 97 (1885)
species :
Exacum marojejyense Humbert; Mem. Inst. Sci. Madagascar, ser. B, 6: 132 (1955)
Distribution: Madagascar
29 MDG
species :
Exacum microcarpum Klack.; Op. Bot. 84: 120 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum millotii Humbert; Mem. Inst. Sci. Madagascar, ser. B, 6: 130 (1955)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum millotii var. protractum Humbert; Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 6: 130 (1955)
species :
Exacum nanum Klack.; Bot. Jahrb. Syst. 126(4): 478 (2006)
Distribution: India (Arunachal Pradesh, Assam); Myanmar; SW-Thailand
40 IND 41 MYA THA
Synonyms: hidden
= Cotylanthera caerulea Lace; Bull. Misc. Inform. Kew 1914: 154 (1914)
species :
Exacum naviculare Klack.; Op. Bot. 84: 77 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum nossibeense Klack.; Op. Bot. 84: 95 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum nummularifolium Humbert; Mem. Inst. Sci. Madagascar, ser. B, 6: 132 (1955)
Distribution: Madagascar
29 MDG
species :
Exacum oldenlandioides (S.Moore) Klack.; Op. Bot. 84: 88 (1985)
Distribution: Senegal; Guinea-Bissau; Guinea; Sierra Leone; Mali; Ivory Coast; Ghana; Togo; Benin; Nigeria; Cameroon; Annobon Isl.; Congo [Brazzaville]; Central African Republic; Burundi; W-Chad; South Sudan; Uganda; Kenya; Tanzania; Angola; Zambia; Zambia; Malawi; Zimbabwe; Mozambique; Namibia; South Africa (Limpopo, Mpulamanga)
22 BEN GHA GNB GUI IVO MLI NGA SEN SIE TOG 23 ANN BUR CAF CMN CON 24 CHA SSU 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 NAM SAF
Synonyms: hidden
= Sebaea oldenlandioides S.Moore; J. Bot. 15: 68 (1877)
species :
Exacum pallidum (Trimen) Klack.; Nordic J. Bot. 3(3): 365 (1983)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Exacum trinervium subsp. pallidum (Trimen) L.H.Cramer; Dassanayake et al., Revis. Handb. Fl. Ceylon 3: 61 (1981)
= Exacum zeylanicum var. pallidum Trimen; Handb. Fl. Ceylon 3: 181 (1895)
species :
Exacum paucisquamum (C.B.Clarke) Klack.; Bot. Jahrb. Syst. 126(4): 478 (2006)
Distribution: China (Sichuan, Yunnan); Tibet; Nepal; Bhutan; N-India (Arunachal Pradesh, Sikkim, Uttarakhand, West Bengal)
36 CHI TIB 40 BHU IND NEP
Synonyms: hidden
= Cotylanthera paucisquama C.B.Clarke; Hook.f. Fl. Brit. Ind. 4: 94 (1883)
= Cotylanthera yunnanensis W.W.Sm.; Notes Roy. Bot. Gard. Edinburgh 13: 158 (1921)
species :
Exacum pedunculatum L.; Sp. Pl. 112 (1753)
Distribution: India (Andhra Pradesh, Bihar, Karnataka, Maharashtra, Odisha, Rajasthan, Tamil Nadu, Telangana); Bangladesh
40 BAN IND
Synonyms: hidden
= Exacum carinatum Roxb.; Hort. Beng. 10 (1814), nomen, Fl. Ind., ed. Carey & Wall. 1: 415 (1820), Fl. Ind., ed. Carey 1: 399 (1832)
= Exacum pedunculare Arn.; Ann. Sci. Nat., ser. II. 11: 176 (1839)
= Exacum punctatum L.f.; Suppl. 124 (1782)
= Exacum rivulare Edgew.; J. Asiat. Soc. Bengal 21: 175 (1853), nom. inval.
= Exacum stellatum Buch.-Ham. ex Wall.; Numer. List: no. 4359 (1831), nom. inval.
= Exacum sulcatum Roxb.; Hort. Beng. 10 (1814), nomen, Fl. Ind., ed. Carey & Wall. 1: 415 (1820), Fl. Ind., ed. Carey 1: 400 (1832)
= Pladera decussata Wall.; Numer. List: no. 4364 (1831), nom. inval.
= Sebaea carinata (Roxb.) Spreng.; Syst. 1: 426 (1824)
= Sebaea sulcata (Roxb.) Spreng.; Syst. 1: 426 (1824)
species :
Exacum penninerve Klack.; Op. Bot. 84: 136 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum petiolare Griseb.; Prodr. [A. DC.] 9: 46 (1845)
Distribution: India (Andhra Pradesh, Bihar, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Tamil Nadu, West Bengal); Sri Lanka
40 IND SRL
Synonyms: hidden
= Exacum pedunculatum var. petiolare (Griseb.) Trimen; Handb. Fl. Ceylon 3: 182 (1895)
species :
Exacum pteranthum Wall.; Numer. List no. 4360 (1832), et ex G. Don, Gen. Syst. 4: 212 (1837), descr.
Distribution: Myanmar (Bago, Taninthayi); N-Thailand; Laos
41 LAO MYA THA
species :
Exacum pumilum Griseb.; Prodr. [A. DC.] 9: 46 (1845)
Distribution: India (Dadra, Goa, Karnataka, Kerala, Madhya Pradesh, Odisha, Tamil Nadu, Uttar Pradesh)
40 IND
species :
Exacum quinquenervium Griseb.; Gen. et Sp. Gent. 112 (1839)
Distribution: Madagascar; ?Mauritius
29 MDG
Remarks: restricted to Madagascar, not in tropical Africa
species :
Exacum radicans Humbert; Adansonia n. s. 4: 32 (1964)
Distribution: Madagascar
29 MDG
species :
Exacum rotundifolium Klack.; Op. Bot. 84: 97 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum sessile L.; Sp. Pl. 112 (1753)
Distribution: S-India (Maharashtra, Karnataka, Kerala, Tamil Nadu, Telangana); Sri Lanka
40 IND SRL
Synonyms: hidden
= Exacum dichotomum Wall.; Numer. List: no. 4359 (1831), nom. inval.
species :
Exacum socotranum Vierh.; Österr. Bot. Zeitschr., 286 (1904)
Distribution: Soqotra
24 SOC
species :
Exacum spathulatum Baker; J. Linn. Soc., Bot. 20: 210 (1883)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum rutenbergiana (Vatke) Schinz; Mitt. Geogr. Ges. Naturhist. Mus. Lübeck 17: 134 (1903)
= Sebaea rutenbergiana Vatke; Abh. Naturwiss. Vereins Bremen 9: 126 (1885)
species :
Exacum speciosum Klack.; Op. Bot. 84: 128 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum stenophyllum Klack.; Op. Bot. 84: 82 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum stenopterum Klack.; Op. Bot. 84: 84 (1985)
Distribution: Madagascar; Comores (Mayotte, etc.)
29 COM MDG
species :
Exacum subacaule Humbert; Adansonia n. s. 4: 33 (1964)
Distribution: Madagascar
29 MDG
species :
Exacum subteres Klack.; Op. Bot. 84: 107 (1985), nom. nov.
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Exacum carinatum Humbert; Mem. Inst. Sci. Madagascar, ser. B, 6: 128 (1955) (non Roxb.)
species :
Exacum subverticillatum Humbert; Adansonia n. s. 4: 33 (1964)
Distribution: Madagascar
29 MDG
species :
Exacum sutaepense Hosseus ex Craib; Bull. Misc. Inform. Kew 1914: 8 (1914)
Distribution: Thailand
41 THA
Synonyms: hidden
= Exacum sutaepense f. gracile (Toyok.) Toyok.; Acta Phytotax. Geobot. 36: 124 (1985)
= Exacum sutaepense Hosseus; Bot. Jahrb. Syst. 40: Beibl. 93, 99 (1808), nomen
= Exacum sutaepense var. gracile Toyok.; Acta Phytotax. Geobot., 32(5-6): 200 (1981)
species :
Exacum tenue (Blume) Klack.; Bot. Jahrb. Syst. 126(4): 478 (2006)
Distribution: India (Arunachal Pradesh, Assam, Odisha); Borneo; Sulawesi; Palawan; Philippines (Luzon, Panay, Negros, Mindanao); Java
40 IND 42 BOR JAW PHI PLW SUL
Synonyms: hidden
= Cotylanthera tenella (A.Gray) Gilg; Engl. & Prantl, Nat. Pflanzenfam. 4(2): 64 (1895)
= Cotylanthera tenuis Blume; Bijdr. 708 (1825)
= Eophylon lobbii A.Gray; J. Linn. Soc., Bot. 11: 23 (1869)
= Eophylon tenellum A.Gray; J. Linn. Soc., Bot. 11: 23 (1869)
species :
Exacum teres Wall.; Roxb. Fl. Ind., ed. Carey & Wall. 1: 414 (1820), et Cat. n. 4354 (1832)
Distribution: China (Yunnan); Nepal; Bhutan; India (Arunachal Pradesh, Assam, Bihar, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Bangladesh; Myanmar (Bago, Rakhine, Shan)
36 CHI 40 BAN BHU IND NEP 41 MYA
species :
Exacum tetragonum Roxb.; Hort. Beng. 10 (1814), nomen, Fl. Ind., ed. Carey & Wall. 1: 413 (1820), Fl. Ind., ed. Carey 1: 398 (1832)
Distribution: China (Guangdong, Guangxi, Guizhou, Jiangxi, Yunnan); Nepal; Bhutan; India (Arunachal Pradesh, Assam, Andhra Pradesh, Bihar, Goa, Karnataka, Kerala, Madhya Pradesh, Mizoram, Odisha, Uttar Pradesh, Tamil Nadu); Bangladesh; Myanmar (widespread); Thailand; Laos; Cambodia; Vietnam; peninsular Malaysia (Perak northward); Sulawesi; Philippines (Luzon, Mindoro, Panay, Mindanao); Java; Lesser Sunda Isl.; New Guinea (Irian Jaya, Papua New Guinea); Australia (Northern Territory)
36 CHI 40 BAN BHU IND NEP 41 CBD LAO MYA THA VIE 42 JAW LSI MLY NWG PHI SUL 50 AUS
Synonyms: hidden
= Canscora justicioides Griff. ex Voigt; Hort. Suburb. Calc. 520 (1845)
= Exacum albens Blanco; Fl. Filip. ed. I. 58 (1837)
= Exacum bellum Hance; Hook. Lond. J. Bot. 7: 472 (1848)
= Exacum bicolor Roxb.; Hort. Beng. 83 (1814), nomen, Fl. Ind., ed. Carey & Wall. 1: 413 (1820), Fl. Ind, ed. Carey 1: 397 (1832)
= Exacum cambodianum Dop; Bull. Soc. Bot. France 59: 145 (1912)
= Exacum chironioides Griseb.; Gen. et Sp. Gent. 109 (1839)
= Exacum chulasiuma Buch.-Ham. ex Wall.; Numer. List: no. 4356 (1831), nom. inval.
= Exacum grandiflorum Wall.; Numer. List no. 4358 (1832) ex G. Don, Gen. Syst. 4: 212 (1837), descr.
= Exacum hamiltonii G.Don; Gen. Hist. 4: 212 (1837)
= Exacum horsfieldianum Miq.; Fl. Ind. Bat. 2: 556 (1856)
= Exacum metzianum Hohen. ex C.B.Clarke; Hook.f., Fl. Brit. India 4: 96 (1883)
= Exacum parviflorum Merr.; Philipp. J. Sci., Bot. 7: 96 (1912)
= Exacum perrottetii Griseb.; Prodr. [A. DC.] 9: 45 (1845)
= Exacum roseum Royle; Illustr. Bot. Himal. 276 (1835)
= Exacum stylosum Wall.; Numer. List no. 4355 (1832) ex G. Don, Gen. Syst. 4: 212 (1837), descr.
= Exacum sulcatum C.B.Clarke; J. Linn. Soc., Bot. 14: 425 (1875)
= Exacum tetragonum var. bicolor (Roxb.) Hook.; Bot. Mag. 73: t. 4340 (1847)
= Exacum tetragonum var. khasianum C.B.Clarke; J. Linn. Soc., Bot. 14: 424 (1875)
= Exacum tetragonum var. roseum Royle ex D.Don; Trans. Linn. Soc. London 17: 532 (1837)
= Exacum tetragonum var. stylosum (Wall. ex G.Don) C.B.Clarke; Hook.f., Fl. Brit. India [J. D. Hooker] 4: 95 (1883)
= Exacum tetrapterum Meyen ex Griseb.; Prodr. [A. DC.] 9: 561 (1845)
= Exacum tricolor Zoll. & Moritzi; Moritzi, Syst. Verz. Java: 56 (1846)
= Paracelsea amoena Zoll. & Moritzi; Natuur- Geneesk. Arch. Ned.-Indië 2: 18 (1845)
variant :
var. grande (Klack.) Shahina & Nampy; Int. J. Advanced Res. 4(3): 1677 (2016)
Distribution: India (Andhra Pradesh, Karnataka, Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum grande Klack.; Op. Bot. 84: 43 (1985)
species :
Exacum travancoricum Bedd.; Ic. Ind. Or. 1: 24 (1868-1874)
Distribution: S-India (Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum courtallense var. travancorica (Bedd.) C.B.Clarke; Hook.f., Fl. Brit. India [J. D. Hooker] 4: 97 (1883)
species :
Exacum trinervium (L.) Druce; Rep. Bot. Exch. Cl. Brit. Isles 3: 418 (1913 publ. 1914)
Distribution: Sri Lanka
40 SRL
Synonyms: hidden
= Chironia trinervia L.; Sp. Pl. 189 (1753)
= Chondropis trinervis (L.) Raf.; Fl. Tellur. 3: 29 (1837)
= Exacum trinervium subsp. ritigalensis (Willis) L.H.Cramer; Dassanayake et al., Revis. Handb. Fl. Ceylon 3: 60 (1981)
= Exacum trinervium var. ritagalense Willis; Ann. Roy. Bot. Gard. (Peradeniya) 3: 282 (1907)
= Exacum zeylanicum Roxb.; Hort. Beng. 83 (1814), nomen, Fl. Ind., ed. Carey & Wall. 1: 414 (1820), Fl. Ind., ed. Carey 1: 398 (1832)
= Exacum zeylanicum var. lewisii Petch; Ann. Roy. Bot. Gard. (Peradeniya) 7: 46 (1919)
= Exacum zeylanicum var. ritigalense Willis; Ann. Roy. Bot. Gard. (Peradeniya) 3: 282 (1907)
= Lisianthus zeylanicus (Roxb.) Spreng.; Syst. 1: 586 (1825)
species :
Exacum umbellatum Klack.; Op. Bot. 84: 116 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum undulatum Klack.; Op. Bot. 84: 124 (1985)
Distribution: Madagascar
29 MDG
species :
Exacum walkeri Arn. ex Griseb.; Gen. et Sp. Gent. 110 (1839)
Distribution: Sri Lanka
40 SRL
species :
Exacum wightianum Arn.; Ann. Sci. Nat., ser. II. 11: 176 (1839)
Distribution: S-India (Karnataka, Kerala, Tamil Nadu)
40 IND
Synonyms: hidden
= Exacum foliosum Griseb.; Prodr. [A. DC.] 9: 45 (1845)
= Lisianthus zeylanicus Spreng.; Syst. Veg. ed. 16. 1: 586 (1824)
species :
Exacum zombense N.E.Br.; Oliv. et al. (eds.), Fl. Trop. Afr. 4: I. 46 (1903)
Distribution: Malawi; N-Mozambique
26 MLW MOZ
species :
Exacum zygomorphum Aver. & K.S.Nguyen; Phytotaxa 183(2): 110, figs. 1-3 (2014)
Distribution: Vietnam
41 VIE
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A complete, synonymic checklist of the Higher Plants of the World


Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life