002.0000 Family Isoetaceae Dumort.; Anal. Fam. Pl. 67 (1829)

1 genus, ca. 190 spp.

Species listed in database: 212; hybrids: 26

002.0001 Genus Isoetes L.; Sp. Pl. [Linnaeus] 2: 1100 (1753)

Distribution: Worldwide. Many undescribed species remain (for China see Gu et al. 2023). Latest checklist: Troia et al. (2016). Taxonomy especially in North America and in the Mediterranean is quite disputed.

Species listed in database: 212; hybrids: 26

species :

Isoetes abyssinica Chiov.; Atti Soc. Naturalisti Mat. Modena 64: 45-47 (1933)

Distribution: Ghana; Ethiopia; Eritrea; Kenya; Tanzania

22 GHA 24 ERI ETH 25 KEN TAN

species :

Isoetes aemulans J.P.Roux; Bothalia 43(2): 218 (2013)

Distribution: South Africa (Free State, Mpulamanga, KwaZulu-Natal)

27 SAF

species :

Isoetes aequinoctialis Welw. ex A.Braun; Sitzungsber. Ges. Naturf. Freunde Berlin 1867: 7 (1867)

Distribution: Mali; Ghana; Niger; Central African Republic; Tanzania; Angola; Zambia; Zimbabwe; Namibia; South Africa (N-Cape Prov., Mpulamanga); Eswatini [Swaziland]

22 GHA MLI NGR 23 CAF 25 TAN 26 ANG ZAM ZIM 27 NAM SAF SWZ

Synonyms: hidden

= Calamaria aequinoctialis (Welw. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes erongensis Wannt.; Svensk Bot. Tidskr. 64(2): 150-152 (1970)

species :

Isoetes afloramientorum D.F.Brunt. & J.B.S.Pereira; Phytotaxa 451(3): 219 (2020)

Distribution: Bolivia (Santa Cruz)

83 BOL

species :

Isoetes alcalophila Halloy; Lilloa 35(2): 68 (1980)

Distribution: Argentina (Catamarca, Tucuman)

85 ARG

species :

Isoetes alpina Kirk; Trans. & Proc. New Zealand Inst. 7: 377, t. 25 (1875)

Distribution: New Zealand (South Isl., North Isl.)

51 NZL

Synonyms: hidden

= Calamaria alpina (Kirk) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes kirkii var. alpina (Kirk) C.R.Marsden & Chinnock; not validly published (Thesis only)

= Isoetes multiangularis Colenso; Trans. & Proc. New Zealand Inst. 22: 449 (1890)

species :

Isoetes alstonii C.F.Reed & Verdc.; Kirkia 5: 19 (1965)

Distribution: Tanzania; S-Zambia; Zimbabwe; Mozambique; N-Namibia; Madagascar

25 TAN 26 MOZ ZAM ZIM 27 NAM 29 MDG

species :

Isoetes amazonica A.Braun; J. Bot. 18: 109 (1880)

Distribution: Brazil (Pará)

84 BRA

Synonyms: hidden

= Calamaria amazonica (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes anamariae J.B.C.Pereira; Amer. Fern J. 112(3): 194-199 (2022)

Distribution: Brazil (Bahia, Pernambuco)

84 BRA

species :

Isoetes anatolica Prada & Rolleri; Bot. J. Linn. Soc. 147(2): 215 (2005)

Distribution: Turkey (Anatolia)

34 TUR

species :

Isoetes andicola (Amstutz) L.D.Gómez; Brenesia 18: 4 (1980)

Distribution: Peru; Bolivia (La Paz)

83 BOL PER

Conservation Status: IUCN R

Synonyms: hidden

= Isoetes andicola (Amstutz) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 177 (1981), isonym

= Isoetes andicola var. gemmifera (Rauh) L.D.Gómez; Brenesia 18: 4 (1980)

= Stylites andicola Amstutz; Ann. Missouri Bot. Gard. 44: 121 (1957)

= Stylites gemmifera Rauh; Sitzungsber. Heidelberger Akad. Wiss., Math. Naturwiss. Kl. 1959: 11 (1959)

species :

Isoetes andina Spruce ex Hook.; Brit. Ferns [W.J.Hooker] ad t. 55 (1861)

Distribution: Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, Magdalena, Meta, Nariño, Putumayo, Risaralda); Venezuela (Merida, Tachira); Ecuador; Peru

82 CLM VEN 83 ECU PER

Synonyms: hidden

= Calamaria triquetra (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1893)

= Isoetes andina Spruce ex Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 325 (1883), nomen

= Isoetes lechleri var. triquetra (A.Braun) L.D.Gómez; Brenesia 18: 5 (1980)

= Isoetes triquetra A.Braun; Verh. Bot. Vereins Prov. Brandenburg 4: 332 (1862)

species :

Isoetes appalachiana D.F.Brunt. & D.M.Britton; Rhodora 99: 118-133 (1997), nom. nov.

Name: Appalachian quillwort

Distribution: USA (Alabama, Delaware, Florida, Georgia, Kentucky, New Jersey, North Carolina, Pennsylvania, South Carolina, Tennessee, Virginia)

75 USA 78 USA

Synonyms: hidden

= Isoetes engelmannii f. fontana (A.A.Eaton) Clute; Fern Allies 258 (1905)

= Isoetes engelmannii f. georgiana (Engelm.) Clute; Fern Allies 258 (1905)

= Isoetes engelmannii var. fontana A.A.Eaton; Fern Bull. 13: 52 (1905)

= Isoetes engelmannii var. georgiana Engelm.; Trans. Acad. Sci. St. Louis 4: 384 (1882)

species :

Isoetes araucaniana Macluf & Hickey; Amer. Fern J. 97(4): 220 (2008)

Distribution: Chile (Biobío, Araucanía)

85 CLE

Synonyms: hidden

= Isoetes favulata Hickey; nom. nud.

species :

Isoetes attenuata C.R.Marsden & Chinnock; Fl. Australia 48: 704 (1998)

Distribution: Australia (SE-South Australia)

50 AUS

species :

Isoetes australis S.Williams; Proc. Roy. Soc. Edinburgh, B, Biol. 62(1): 1-8 (1943)

Distribution: Australia (SW-Western Australia)

50 AUS

species :

Isoetes azorica Durieu; Fil. Eur. 278 (1867)

Name: Azoren-Brachsenkraut

Distribution: Azores (Terceira, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.)

21 AZO

Conservation Status: IUCN E

Synonyms: hidden

= Calamaria azorica (Durieu) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes baculata Hickey & H.P.Fuchs; Syst. Bot. 11(2): 310 (1986)

Distribution: Colombia (Vaupés); Brazil (Amazonas)

82 CLM 84 BRA

species :

Isoetes baodongii Y.F.Gu, Y.H.Yan & Yi Jun Lu; Novon 29: 207 (2021)

Distribution: China (Zhejiang)

36 CHI

species :

Isoetes bhimii M.Patel & H.K.Goswami; Indian Fern J. 41(2): 26 (2024)

Distribution: India (Madhya Pradesh)

40 IND

species :

Isoetes biafrana Alston; Bol. Soc. Brot., Ser. 2A, 30: 15 (1956)

Distribution: Benin; Cameroon; Bioko Isl. [Fernando Poo]

22 BEN 23 BIO CMN

species :

Isoetes bilaspurensis Panigrahi; Biol. Mem. 6(2): 135, f. 1-10 (1981)

Distribution: India (Chhattisgarh, Karnataka, Madhya Pradesh)

40 IND

Remarks: bona sp. fide Shukla et al. 2024

Synonyms: hidden

= Isoetes mahadevensis G.K.Srivast., D.D.Pant & P.K.Shukla; Amer. Fern J. 83(4): 106 (1993)

species :

Isoetes bischlerae H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(1): 113 (1982)

Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Nariño)

82 CLM

Remarks: provisionally accepted name

species :

Isoetes bolanderi Engelm.; Amer. Naturalist 8: 214 (1874)

Name: Bolander´s Quillwort

Distribution: Canada (Alberta); USA (Arizona, California, Colorado, Idaho, Montana, New Mexico, Nevada, Oregon, Utah, Washington State, Wyoming)

71 CDN 73 USA 76 USA 77 USA

Remarks: not in Florida, obviously a BONAP error

Conservation Status: IUCN R

Synonyms: hidden

= Calamaria bolanderi (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Calamaria pygmaea (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes bolanderi f. pygmaea (Engelm.) Clute; Fern Bull. 16: 54 (1908)

= Isoetes bolanderi var. parryi Engelm.; Amer. Naturalist 8: 214 (1874)

= Isoetes bolanderi var. pygmaea (Engelm.) Clute; Fern Allies 228, 258 (1905)

= Isoetes bolanderi var. sonnei L.F.Hend.; Bull. Torrey Bot. Club 27(6): 358 (1900)

= Isoetes californica Engelm.; Manual [A.Gray], ed. 5: 677 (1867)

= Isoetes parryi Engelm.; Actes Soc. Linn. Bordeaux 36: 338 (1883)

= Isoetes pygmaea Engelm.; Amer. Naturalist 8: 214 (1874)

species :

Isoetes boliviensis U.Weber; Hedwigia 63: 247, fig. 14-15 (1922)

Distribution: Peru; Bolivia (Cochabamba, La Paz)

83 BOL PER

species :

Isoetes boomii Luebke; Amer. Fern J. 82: 23 (1992)

Name: Boom´s quillwort

Distribution: USA (Florida, Alabama, C-Georgia: Laurens Co., Worth Co.)

75 USA 78 USA

Conservation Status: IUCN E

species :

Isoetes boryana Durieu; Bull. Soc. Bot. France 8: 164 (1861)

Distribution: +Spain; SW-France

12 FRA SPA

Conservation Status: IUCN V

Synonyms: hidden

= Calamaria boryana (Durieu) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes boryana subsp. brevis Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes boryana subsp. condensata Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes boryana subsp. frigida Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes boryana subsp. gredensis Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes boryana subsp. legitima Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes boryana subsp. praetervisa Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes variabilis proles boryana (Durieu) Rouy; Fl. France [Rouy & Foucaud] 14: 477 (1913), nom. illeg.

= Isoetes variabilis subsp. boryana (Durieu) P.E.Fourn.; Quatre Fl. France 5 (1934), nom. illeg.

= Isoetes velata var. boryana (Durieu) M.I.Romero & Real; Bot. J. Linn. Soc. 148(4): 464 (2005)

species :

Isoetes boyacensis H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(1): 107 (1982)

Distribution: Colombia (Boyacá, Cundinamarca, Meta, Nariño, Santander)

82 CLM

Remarks: provisionally accepted name

species :

Isoetes bradei Herter; Revista Sudamer. Bot. 8: 19 (1949)

Distribution: Brazil (São Paulo, ?Minas Gerais)

84 BRA

Conservation Status: considered extinct

species :

Isoetes brasiliensis H.P.Fuchs; Fl. Ilustr. Catarin. (Isoetaceas) 27 (1986)

Distribution: Brazil (Santa Catarina, Rio Grande do Sul)

84 BRA

Synonyms: hidden

= Isoetes reitzii H.P.Fuchs; (1979), nom. nud. pro syn.

species :

Isoetes brevicula E.R.L.Johnson; J. Roy. Soc. Western Australia 67(1): 34 (1984)

Distribution: Australia (SW-Western Australia)

50 AUS

species :

Isoetes butleri Engelm.; Bot. Gaz. 3: 1 (1878)

Name: Limestone quillwort

Distribution: USA (Alabama, Arkansas, Georgia, Illinois, Kansas, Kentucky, Missouri, Oklahoma, Tennessee, Texas)

74 USA 75 USA 77 USA 78 USA

Synonyms: hidden

= Calamaria butleri (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes butleri f. immaculata (Engelm.) Clute; Fern Allies 258 (1905)

= Isoetes butleri var. immaculata Engelm.; Trans. Acad. Sci. St. Louis 4: 388 (1882)

species :

Isoetes candelariensis C.C.Macluf & G.J.Marquez; Anais Acad. Brasil. Ci. 96(1)-e20230087: 3 (2024)

Distribution: Argentina (Misiones)

85 ARG

species :

Isoetes cangae J.B.S.Pereira, Salino & Stützel; Phytotaxa 272(2): 145 (2016)

Distribution: Brazil (Pará)

84 BRA

species :

Isoetes caparaoensis J.B.S.Pereira; Bot. J. Linn. Soc. 189(1): 15 (2018)

Distribution: Brazil (Minas Gerais, Espírito Santo)

84 BRA

species :

Isoetes capensis A.V.Duthie; Trans. Roy. Soc. South Africa 17: 330 (1929)

Distribution: South Africa (W-Cape Prov.)

27 SAF

species :

Isoetes caroli E.R.L.Johnson; J. Roy. Soc. Western Australia 67(1): 34 (1984)

Distribution: Australia (SW-Western Australia)

50 AUS

species :

Isoetes caroliniana (A.A.Eaton) Luebke; Amer. Fern J. 82: 26 (1992)

Distribution: SE-USA (North Carolina, Tennessee, Virginia, West Virginia)

75 USA 78 USA

Remarks: bona species fide Troia et al. 2016, not a synonym of I. valida

Synonyms: hidden

= Isoetes engelmannii var. caroliniana A.A.Eaton; Fern Bull. 8: 60 (1900)

species :

Isoetes changleensis Yu C.Chen & Xing Liu; Phytotaxa 642(1): 41 (2024)

Distribution: China (Zhejiang)

36 CHI

species :

Isoetes changxingensis Y.F.Gu & J.H.Shen; Guihaia 44(4): 613 (2024)

Distribution: China (Zhejiang)

36 CHI

species :

Isoetes chubutiana Hickey, Macluf & W.C.Taylor; Amer. Fern J. 93(3): 130 (2003)

Distribution: Argentina (Chubut, Neuquen, Rio Negro); Chile (Biobío, Los Lagos, Aisén, Magallanes)

85 ARG CLE

Synonyms: hidden

= Isoetes savatieri subsp. valdiviensis H.P.Fuchs; (1992), nom. nud.

= Isoetes valdiviensis H.P.Fuchs; (1960), nom. nud.

= Isoetes meyeri H.P.Fuchs; (1982), nom. nud.

species :

Isoetes cipoensis J.B.S.Pereira; Syst. Bot. 47(2): 301 (2022)

Distribution: Brazil (Minas Gerais)

84 BRA

species :

Isoetes coreana Y.H.Chung & H.K.Choi; Korean J. Pl. Taxon. 16(1): 2 (1986)

Distribution: North Korea; South Korea; Japan

38 JAP KOR NKO

Synonyms: hidden

= Isoetes sinensis var. coreana (Y.H.Chung & H.K.Choi) M.Takamiya, M.Watanaba & K.Ono; Acta Phytotax. Geobot. 48(2): 112 (1997 publ. 1998)

species :

Isoetes coromandelina L.f.; Suppl. Pl. [Linnaeus f.] 447 (1782)

subspecies :

subsp. coromandelina

Distribution: Nepal; India (Andhra Pradesh, Bihar, Chhattisgarh, Goa, Gujarat, Haryana, Jharkand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Odisha, Rajasthan, Tamil Nadu, Uttar Pradesh, West Bengal); Sri Lanka; Bangladesh; Myanmar; Thailand; Vietnam

40 BAN IND NEP SRL 41 MYA THA VIE

Synonyms: hidden

= Calamaria coromandelina (L.f.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes brachyglossa A.Braun; Verh. Bot. Vereins Prov. Brandenburg 4: 328 (1862), nom. nov. for I. capsularis Roxb. ex Griff. (fide Fraser-Jenkins 1997, 2017)

= Isoetes capsularis Roxb. ex Griff.; Posth. Papers Cryptog. Pl. 572-575, t. 116-118 (1849), non Roxb. 1844

= Isoetes coromandelina subsp. brachyglossa (A.Braun) Panigrahi; Biol. Mem. 6(2): 131 (1981) (fide Fraser-Jenkins 1997)

= Isoetes coromandelina subsp. thanensis S.K.Shukla, Sarv.K.Singh, P.K.Shukla, N.K.Dubey, H.Khanam & G.K.Srivast.; Taiwania 62(2): 122 (2017)

= Isoetes coromandelina var. raipurensis Unni; J. Bombay Nat. Hist. Soc. 64: 590-592 (1967) (fide Fraser-Jenkins 1997)

= Isoetes debii P.K.Shukla, G.K.Srivast. & S.K.Shukla; Frontiers Bot. 4-6: 69-72 (2002), nom. inval.

= Isoetes debii S.C.Sinha; Frontiers Bot. 4-6: 69 (1992)

= Isoetes fuchsii Bhu, H.K.Goswami, U.S.Sharma & A.K.Bajpai; J. Advancem. Pl. Sci. 2(1)-1: 4 (2019)

= Isoetes fuchsii H.K.Goswami & U.S.Sharma; Bionature 21: 11-18 (2001), nom. inval.

= Isoetes indica D.D.Pant & G.K.Srivast.; Proc. Indian Acad. Sci., B 28(3) 246, t. XIV9, f. 16-259, t. XV, f. 30-35, t. XIV, f. 30-40 text-fig. 4 A-D, 5 A-H, 7 A-M, 12 (1962) (fide Fraser-Jenkins 1997)

= Isoetes indica var. harotiensis Bhardwaja & Gena; Aspects Pl. Sci. 14: 495-499 (1992), nom. nud.

= Isoetes mirzapurensis Panigrahi & R.D.Dixit; Proc. Indian Sci. Congr. Assoc., pt. III, Abstract, Botany 243 (1965) (fide Fraser-Jenkins 1997)

= Isoetes narsinghgarhensis H.K.Goswami & Mazumdar; J. Advancem. Pl. Sci. 2(1)-1: 5 (2019), nom. nov. for I. pantii var. hybrida Goswami

= Isoetes pantii H.K.Goswami & B.S.Arya; J. Indian Bot. Soc. 49: 30 (1970) (fide Fraser-Jenkins 1997)

= Isoetes pantii var. jaideepii M.Patel, M.N.Reddy & H.K.Goswami; Indian Fern J. 41(2): 25 (2024)

= Isoetes pantii var. hybrida H.K.Goswami; Indian Fern J. 21(1-2): 152(-153) (2004), nom. inval. [fide Fraser-Jenkins et al. 2020]

= Isoetes semilocularis Sm.; Cycl. [A.Rees] (London ed.) 19: 150 (1819)

= Isoetes tuberculata Gena & Bhardwaja; J. Bombay Nat. Hist. Soc. 81(1): 167 (1984) (fide Fraser-Jenkins 1997)

= Isoetes unilocularis Roxb. ex Sm.; Cycl. [A.Rees] (London ed.) 19: no. 3 (1811)

subspecies :

subsp. macrotuberculata C.R.Marsden; Contr. Herb. Austral. 24: 2 (1976)

Distribution: Australia (Western Australia, Northern Territory, Queensland)

50 AUS

species :

Isoetes creussensis Lazare & S.Riba; J. Bot. Soc. Bot. France 50: 21 (2010)

Distribution: Andorra; S-France

12 ADO FRA

Remarks: status somewhat uncertain, near I. lacustris but cytologically separate

species :

Isoetes cristata C.R.Marsden & Chinnock; Fl. Australia 48: 704 (1998)

Distribution: Australia (Northern Territory)

50 AUS

Conservation Status: Known from type locality only

species :

Isoetes cubana Engelm. ex Baker; J. Bot. 18: 110 (1880)

Distribution: Mexico (Yucatan); Belize; Cuba

79 MEX 80 BLZ 81 CUB

Synonyms: hidden

= Calamaria cubana (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes delalandei J.Lloyd; Notes Fl. Ouest France 25 (1851)

Distribution: NW-France

12 FRA

Synonyms: hidden

= Isoetes histrix proles delalandei (J.Lloyd) Rouy; Fl. France [Rouy & Foucaud] 14: 481 (1913)

= Isoetes histrix subsp. delalandei (J.Lloyd) P.E.Fourn.; Quatre Fl. France 6 (1934)

= Isoetes histrix var. delalandei (J.Lloyd) Rouy; Fl. France [Rouy & Foucaud] 14: 481 (1913)

species :

Isoetes delilei (Bory) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 72 (1944)

Distribution: Portugal; Spain (W-Spain); SE-France; ?Corsica; ?Sardinia; Morocco; ?Algeria

12 FRA POR SPA 20 MOR

Synonyms: hidden

= Isoetes setacea Bosc ex Delile; Mém. Mus. Hist. Nat. 14(110): 117, t. 6-7 (1827) (non Lam.)

= Isoetes setacea var. delilei Bory; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1165 (1844)

= Isoetes palustris var. setacea F.Muell.; Fil. Eur. 9: 280 (1867)

= Isoetes montana U.Weber; Hedwigia 63: 254 (1922)

species :

Isoetes dispora Hickey; Fieldiana, Bot. 34: 93 (1994)

Distribution: Peru

83 PER

Conservation Status: Known from type only

species :

Isoetes divyadarshanii P.K.Shukla, G.K.Srivast. & S.K.Shukla; Taxon 54(1): 113 (2005)

Distribution: India (Karnataka, +Maharashtra)

40 IND

Remarks: recently rediscovered: Singh et al. 2025

species :

Isoetes dixitii Shende; J. Univ. Bombay 14: 50 (1945)

Distribution: India (Karnataka, Maharashtra)

40 IND

Remarks: bona sp. fide Fraser-Jenkins et al. 2017

species :

Isoetes drummondii A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 593 (1863)

subspecies :

subsp. drummondii

Distribution: Australia (SW-Western Australia, SE-South Australia, SE-Queensland, Victoria, NE-Tasmania)

50 AUS

Synonyms: hidden

= Calamaria drummondii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

subspecies :

subsp. anomala C.R.Marsden & Chinnock; Muelleria 8(1): 58 (1993)

Distribution: Australia (SE-South Australia, SW-Western Australia, Victoria, S-New South Wales)

50 AUS

species :

Isoetes dubsii J.B.S.Pereira; PhytoKeys 131: 59 (2019)

Distribution: Brazil (Mato Grosso do Sul)

84 BRA

species :

Isoetes duriei Bory; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1166 (1844) (durieai auct., durieui auct.)

Name: Durieu-Brachsenkraut (DE)

Distribution: Portugal; Spain; Baleares; France; Corsica; Sardinia; Italy; Sicily; Pantelleria; Malta; Greece (Evvoia, Sporades, Kiklades, Samothraki); Crete (rare); ?Morocco; Algeria; Tunisia; East Aegaean Isl. (Lesvos, Rhodos, Karpathos); Turkey (Anatolia)

12 BAL COR FRA POR SAR SPA 13 GRC ITA KRI MLT PTL SIC 20 ALG TUN 34 EAI TUR

Synonyms: hidden

= Calamaria durieui (Bory) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetella duriei (Bory) Gennari; Comment. Soc. Crittog. Ital. 1(3): 115 (1862)

= Isoetes duriaei subsp. balearica Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. felix Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. insularis Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. kabylica Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. leptophylla Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. ligustica De Not.; Index Filic. Cult. [Kunze] 51 (1853)

= Isoetes duriaei subsp. litorea Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. lusitanica Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. maritima Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes duriaei subsp. sardoa Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes ligustica De Not. ex A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin Sitzung 7 Dec (1863)

= Isoetes tridentata Durieu; Filic. Afr. [Kuhn] 195 (1867)

species :

Isoetes echinospora Durieu; Bull. Soc. Bot. France 8: 164 (1861)

Name: Igelsporiges Brachsenkraut, Stachelsporiges Brachsenkraut (DE), Spiny-spore quillwort (EN)

Distribution: Iceland; England (SW- & NW-England, Wales, Scotland, Shetland Isl.); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Germany (Baden-Württemberg, Schleswig-Holstein+); Switzerland; Austria; Poland; Czech Republic; Slovakia; Spain; Andorra; S-France; Italy; Estonia; Latvia; European Russia (N-European Russia, C-European Russia); Morocco; Siberia (Buryatia, Chita, Irkutsk, W-Siberia); Russian Far East (Kamchatka, Kuril Isl., Primorye, Sakhalin); ?China; Japan; Alaska; Greenland; Canada (Alberta, British Columbia, Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Nunavut, Ontario, Québec, Saskatchewan, Yukon); St. Pierre et Miquelon; USA (California, Colorado, Connecticut, Delaware, Idaho, Massachusetts, Maine, Michigan, Minnesota, Montana, New Hampshire, New Jersey, New York, Ohio, Oregon, Pennsylvania, Rhode Island, Utah, Vermont, Washington State, Wisconsin, Wyoming)

10 DEN FIN GRB ICE IRE NOR SWE 11 AUT CZE GER NET POL SLO SWI 12 ADO FRA SPA 13 ITA 14 EST LAT RUS 20 MOR 30 SIB 31 RFE 38 JAP 70 ASK CDN GNL 71 CDN 72 CDN SPM 73 USA 74 USA 75 USA 76 USA 78 USA

Remarks: not in Bulgaria or Greece. Inclusion of I. asiatica fide Grigoryan et al. (2025)

Synonyms: hidden

= Calamaria echinospora (Durieu) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes ambigua A.Braun; Trans. Acad. Sci. St. Louis 4: 380 (1882)

= Isoetes asiatica (Makino) Makino; Bot. Mag. (Tokyo) 28: 184 (1914)

= Isoetes boottii A.Braun; Manual [A.Gray], ed. 5: 676 (1867)

= Isoetes braunii Durieu; Bull. Soc. Bot. France 11: 101 (1864)

= Isoetes braunii f. astoma (Iversen) M.Broun; Index No. Amer. Ferns: 99 (1938)

= Isoetes braunii f. oligostoma (Iversen) M.Broun; Index No. Amer. Ferns: 99 (1938)

= Isoetes braunii f. polystoma (Iversen) M.Broun; Index No. Amer. Ferns: 99 (1938)

= Isoetes echinospora subf. angustivelata Iversen; Bot. Tidsskr. 40: 128, fig. a-c (1928)

= Isoetes echinospora subf. lativelata Iversen; Bot. Tidsskr. 40: 129 (1928)

= Isoetes echinospora f. astoma Iversen; Bot. Tidsskr. 40: 129 (1928)

= Isoetes echinospora f. boottii (Engelm.) Clute; Fern Allies 258 (1905)

= Isoetes echinospora f. oligostoma Iversen; Bot. Tidsskr. 40: 129 (1928)

= Isoetes echinospora f. polystoma Iversen; Bot. Tidsskr. 40: 128 (1928)

= Isoetes echinospora subsp. admixta Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. asiatica (Makino) Á.Löve; Amer. Fern J. 52: 121 (1962)

= Isoetes echinospora subsp. belgica Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. borealis Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. centralis Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. discedens Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. echinospora Durieu; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. fennica Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. gothica Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. montana Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. muricata (Durieu) Á.Löve & D.Löve; Bot. Not. 114: 49 (1961)

= Isoetes echinospora subsp. oligophylla Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. petropolitana Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. praebens Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. pulchella Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. pygmaea Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. remissa Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora subsp. vogesiaca Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes echinospora var. asiatica Makino; Bot. Mag. (Tokyo) 18: 129 (1904) ex Iversen, Dansk Bot. Ark. 5(23): 2 (1928)

= Isoetes echinospora var. boothii Clute; Ferns Northwest [Frye] 39 (pro syn.) (1934)

= Isoetes echinospora var. boottii Engelm.; Manual [A.Gray], ed. 5: 676 (1867)

= Isoetes echinospora var. braunii (Durieu) Á.Löve & D.Löve; Bot. Not. 114: 49 (1961), isonym

= Isoetes echinospora var. braunii (Durieu) Engelm.; Manual [A.Gray], ed. 5: 676 (1867)

= Isoetes echinospora var. brittoni Cockerell; Muhlenbergia 3: 9 (1907)

= Isoetes echinospora var. elatior Fliche; Mém. Acad. Stanislas, Sér. 4, 11: 182 (1878 publ. 1879)

= Isoetes echinospora var. hesperia (C.F.Reed) Á.Löve; Amer. Fern J. 52: 121 (1962)

= Isoetes echinospora var. muricata (Durieu) Engelm.; Manual [A.Gray], ed. 5: 676 (1867)

= Isoetes lacustris subsp. curvifolia (Casp.) Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. echinospora (Durieu) Hook.f.; Student´s Fl. Brit. Isl., ed. 1: 471 (1870), isonym

= Isoetes lacustris subsp. echinospora (Durieu) Syme & N.E.Br.; Engl. Bot., ed. 3B, 12: 7 (1886), isonym

= Isoetes lacustris subsp. echinospora (Durieu) T.Moore & More; Contr. Cybele Hibern. 381 (1866)

= Isoetes lacustris var. crassa Léman; Dict. Sci. Nat., ed. 2 [F.Cuvier] 24: 24 (1822)

= Isoetes lacustris var. elatior (Fliche) Mer; Bull. Soc. Bot. France 28: 73 (1881)

= Isoetes lacustris var. longifolia Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)

= Isoetes lacustris var. minor (A.Braun ex Milde) Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)

= Isoetes lacustris var. recurvata Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)

= Isoetes muricata Durieu; Bull. Soc. Bot. France 11: 100 (1864)

= Isoetes muricata subsp. asiatica (Makino) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982)

= Isoetes muricata var. braunii (Durieu) C.F.Reed; Amer. Fern J. 35: 83 (1945)

= Isoetes muricata var. hesperia C.F.Reed; Amer. Fern J. 35: 84 (1945)

= Isoetes setacea subsp. asiatica (Makino) Holub; Acta Horti Bot. Prag. 1962: 55 (1962)

= Isoetes setacea subsp. monspeliensis Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes setacea subsp. muricata (Durieu) Holub; Acta Horti Bot. Prag. 1964: 55 (1964)

= Isoetes setacea subsp. stricta Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes setacea subsp. turgida Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes setacea subsp. valida Gand.; Fl. Eur. [Gandoger] 27: 196 (1891)

= Isoetes setacea subsp. vicina Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes tenella f. astoma (Iversen) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 72 (1944)

= Isoetes tenella f. iversenii Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)

= Isoetes tenella f. lativelata (Iversen) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 72 (1944)

= Isoetes tenella f. oligostoma (Iversen) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)

= Isoetes tenella f. polystoma (Iversen) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)

= Isoetes tenella var. braunii (Durieu) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)

= Isoetes tenella var. oligostoma (Iversen) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 72 (1944)

species :

Isoetes ecuadoriensis Aspl.; Bot. Not. 1925: 357, f. (1925)

Distribution: Ecuador

83 ECU

Synonyms: hidden

= Isoetes keeleyorum H.P.Fuchs; (1988), nom. nud.

species :

Isoetes ekmanii U.Weber; Hedwigia 63: 251 (1922)

Distribution: Argentina (Buenos Aires, Corrientes, Misiones)

85 ARG

species :

Isoetes elatior F.Muell. & A.Braun; Linnaea 25: 722 (1852)

Distribution: Australia (Tasmania)

50 AUS

Conservation Status: IUCN R

Synonyms: hidden

= Calamaria elatior (F.Muell.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes tasmanica F.Muell.; Bull. Soc. Bot. France 11: 104 (1864)

species :

Isoetes eludens J.P.Roux, Hopper & Rhian J.Sm.; Kew Bull. 64(1): 124 (2009)

Distribution: South Africa (N-Cape: Namaqualand)

27 SAF

Conservation Status: extremely local in a single 2 meter diameter rock pool

species :

Isoetes engelmannii A.Braun; Flora 29: 178 (1846)

Name: Engelmann´s quillwort

Distribution: Canada (Ontario); USA (Alabama, Arkansas, Connecticut, Delaware, Georgia, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Missouri, Mississippi, North Carolina, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, Rhode Island, South Carolina, Tennessee, Virginia, Vermont, West Virginia)

72 CDN 74 USA 75 USA 78 USA

Synonyms: hidden

= Calamaria engelmannii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes engelmannii var. gracilis Engelm.; Manual [A.Gray], ed. 5: 677 (1867)

= Isoetes lacustris var. microspora Engelm.; Flora 29: 179 (1846)

= Isoetes ×fairbrothersii J.D.Montgom. & W.C.Taylor; Amer. Fern J. 84(4): 120 (1994)

species :

Isoetes escondidensis Halloy; Lilloa 35(2): 67 (1980)

Distribution: Argentina (Tucuman)

85 ARG

Conservation Status: IUCN E

Synonyms: hidden

= Isoetes castillonii H.P.Fuchs; (1965), nom. nud.

= Isoetes halloyi H.P.Fuchs; (1981), nom. nud.

= Isoetes filipovichii H.P.Fuchs; (1965), nom. nud.

species :

Isoetes eshbaughii Hickey & H.P.Fuchs; Syst. Bot. 11: 310 (1986)

Distribution: Bolivia (Cochabamba)

83 BOL

species :

Isoetes fengii Y.F.Gu & Y.H.Yan; Pl. Sci. J. 41(2): 168 (2023)

Distribution: China (Yunnan)

36 CHI

species :

Isoetes flaccida Shuttlew. ex A.Braun; Flora 29: 178 (1846)

Name: Southern quillwort

Distribution: USA (Alabama, Florida, Georgia)

78 USA

Conservation Status: IUCN R

Synonyms: hidden

= Calamaria flaccida (Shuttlew) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes alata (N.Pfeiff.) Small; Ferns Florida 22 (1932)

= Isoetes flaccida f. rigida (Engelm.) Clute; Fern Allies 258 (1905)

= Isoetes flaccida var. alata N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 137 (1922)

= Isoetes flaccida var. rigida Engelm.; Trans. Acad. Sci. St. Louis 4: 386 (1882)

variant :

var. chapmanii Engelm.; Trans. Acad. Sci. St. Louis 4: 386 (1882)

Name: Chapman´s quillwort

Distribution: USA (Florida, Georgia)

78 USA

Synonyms: hidden

= Isoetes chapmanii (Engelm.) Small; Ferns Florida 222 (1931 publ. 1932)

= Isoetes flaccida f. chapmanii (Engelm.) Clute; Fern Allies 258 (1905)

species :

Isoetes fluitans M.I.Romero; Bot. J. Linn. Soc. 146(2): 233 (2004)

Distribution: Spain

12 SPA

species :

Isoetes fuliginosa R.L.Small & Hickey; Amer. Fern J. 91(2): 55 (-57, figs. 5-7, 14-15) (2001)

Distribution: Colombia (Cauca)

82 CLM

species :

Isoetes fuscomarginata H.P.Fuchs; Fl. Ilustr. Catarin. (Isoetaceas) 12 (1986)

Distribution: Brazil (Rio Grande do Sul)

84 BRA

species :

Isoetes gardneriana Kunze ex Mett.; Fil. Lechl. 2: 36 (1859)

Distribution: Brazil (Goiás, Mato Grosso do Sul, Tocantins, Bahia, Rio de Janeiro)

84 BRA

Synonyms: hidden

= Calamaria gardneriana (Kuntze ex Mett.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes georgiana Luebke; Amer. Fern J. 82: 24 (1992)

Name: Georgia quillwort

Distribution: USA (Georgia)

78 USA

species :

Isoetes giessii Launert; Mitt. Bot. Staatssamml. München 2: 309 (1957)

Distribution: Namibia

27 NAM

Conservation Status: known from two locations

species :

Isoetes gigantea U.Weber; Nova Hedwigia 63: 245 (1922)

Distribution: Brazil (Bahia, Minas Gerais, Rio de Janeiro, Mato Grosso)

84 BRA

species :

Isoetes graniticola D.F.Brunt.; Rhodora 118(975): 269 (2016)

Distribution: USA (Alabama)

78 USA

species :

Isoetes gunnii A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 535 (1868)

Distribution: Australia (Tasmania)

50 AUS

Synonyms: hidden

= Calamaria gunnii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes lacustris (L.) Rodway; Tasman. Fl. 279 (1903)

species :

Isoetes gymnocarpa (Gennari) A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 555 (1864)

Name: Nacktsamiges Brachsenkraut; Naked-fruited Quillwort

Distribution: S-Corsica; Italy; Sicily; Montenegro; ?Algeria; East Aegaean Isl. (Karpathos, Lesvos, Rhodos)

12 COR 13 ITA MNT SIC 34 EAI

Remarks: probably to be merged into I. histrix fide Strid (2024)

Synonyms: hidden

= Cephaloceraton gymnocarpum Gennari; Comment. Soc. Crittog. Ital. 1(3): 113 (1862)

= Cephaloceraton histrix var. subinerme Gennari; Comment. Soc. Crittog. Ital. 1: 112 (1862)

= Cephaloceraton pseudohystrix Gennari; Ann. Agric. Siciliana 11: 175 (1865)

= Isoetes histrix proles sicula (Tod.) Rouy; Fl. France [Rouy & Foucaud] 14: 482 (1913)

= Isoetes histrix f. subinermis (Gennari) Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30(1): 12 (1924)

= Isoetes histrix subsp. sicula (Tod.) P.E.Fourn.; Quatre Fl. France 6 (1934)

= Isoetes histrix var. subinermis (Gennari) Durieu; Comp. Fl. Ital. [Cesati] 25 (1867), et Motelay & Vendryès, Actes Soc. Linn. Bordeaux 36: 4009, t. 17), f. 1-5 (1883)

= Isoetes histrix var. subinermis (Gennari) Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 400 (1883)

= Isoetes hystrix var. subinermis Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes pseudohystrix (Gennari) Levier ex Barbey; Fl. Sard. Comp. 195 (1885)

= Isoetes sicula Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 251 (1866)

= Isoetes subinermis (Gennari) Cesca & Peruzzi; Fl. Medit. 11: 308 (2001)

species :

Isoetes habbemensis Alston; J. Arnold Arbor. 26: 180 (1945)

Distribution: New Guinea (Irian Jaya: Mt. Wilhelmina-Lake Habbema)

42 NWG

Conservation Status: IUCN R (twice found)

species :

Isoetes hallasanensis H.K.Choi, Ch.Kim & J.Jung; J. Pl. Biol. 51(5): 357 (2008)

Distribution: South Korea (Cheju Isl.)

38 KOR

species :

Isoetes hallei Eb.Fisch. & Lobin; Willdenowia 52(3): 330 (2022)

Distribution: W-D.R.Congo [Zaire]

23 ZAI

Remarks: known only from type locality

species :

Isoetes harleyi J.B.C.Pereira; Amer. Fern J. 112(3): 199-200 (2022)

Distribution: Brazil (Bahia)

84 BRA

species :

Isoetes haussknechtii Troia & Greuter; Willdenowia 45(3): 395 (2015)

Name: Haussknechts Brachsenkraut (DE)

Distribution: Greece; East Aegaean Isl. (Lesvos)

13 GRC 34 EAI

Remarks: probably a terrestrial form of I. heldreichii fide Strid (2024)

species :

Isoetes hawaiiensis W.C.Taylor & W.H.Wagner; Amer. Fern J. 83(2): 67 (1993)

Distribution: Hawaii (Maui, Hawaii Isl.)

63 HAW

species :

Isoetes heldreichii Wettst.; Verh. K.K. Zool.-Bot. Ges. Wien 36: 239, t. 8 (1886)

Name: Heldreichs Brachsenkraut

Distribution: +C-Greece (+Palaiokastro near Karditsa)

13 GRC

Remarks: extinct fide Troia & Greuter 2015

species :

Isoetes hemivelata R.L.Small & Hickey; Amer. Fern J. 91(2): 57 (60, figs. 9-11, 16-17) (2001)

Distribution: Colombia (Cauca, Meta, Risaralda)

82 CLM

species :

Isoetes herzogii U.Weber; Hedwigia 63: 250 (1922)

Distribution: Peru (Cuzco); Bolivia (Cochabamba, La Paz, Santa Cruz)

83 BOL PER

species :

Isoetes hewitsonii Hickey; Fieldiana, Bot. 34: 94 (1994)

Distribution: Peru (Cajamarca)

83 PER

species :

Isoetes hieronymi U.Weber; Hedwigia 63: 251, fig. 24-25 (1922)

Distribution: Argentina (Cordoba, Rio Negro); Chile (Coquimbo)

85 ARG CLE

Synonyms: hidden

= Isoetes argentina A.A.Eaton; Amer. Fern J. 19: 18 (1929), nom. nud.

= Isoetes lechleri N.Pfeiff.; Ann. Missouri Bot. Gard. 9(2): 138 (1922), non Mett.

species :

Isoetes histrix Bory & Durieu; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1167 (1844)

Name: Stacheliges Brachsenkraut

Distribution: SW-England; Portugal; Spain; France; Channel Isl. (Guernsey, Alderney); Corsica; Sardinia; Italy; Sicily; Malta; Montenegro; North Macedonia; Albania; Greece (scattered, incl. Kiklades); Crete (W-Crete); Morocco; Algeria; Tunisia; East Aegaean Isl. (Lesvos, Ikaria, Rhodos, Karpathos); Turkey (Anatolia); Lebanon

10 GRB 12 CIS COR FRA POR SAR SPA 13 ALB GRC ITA KRI MLT MNT NMA SIC 20 ALG MOR TUN 34 EAI LEB TUR

Synonyms: hidden

= Calamaria histrix (Bory) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Cephaloceraton histrix (Bory) Gennari; Comment. Soc. Crittog. Ital. 1: 111 (1862)

= Isoetes chaeturetii Mendes; Agron. Lusit. 23: 7 (1961) (fide Musselman 2003)

= Isoetes histrix subvar. armata Batt. & Trab.; Fl. Algérie Tunisie 407 (1902)

= Isoetes histrix subvar. inermis Batt. & Trab.; Fl. Algérie Tunisie 407 (1902)

= Isoetes histrix unranked grandis A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)

= Isoetes histrix unranked longispina A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 618 (1864)

= Isoetes histrix unranked nana A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)

= Isoetes histrix unranked solitaria A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 618 (1864)

= Isoetes histrix f. desquamata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)

= Isoetes histrix f. longispina (A.Braun) Fiori; Fl. Ital. Crypt. 5: 425 (1943)

= Isoetes histrix f. loricata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)

= Isoetes histrix var. desquamata (A.Braun) Batt. & Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes histrix var. desquamata f.longispina (A.Braun) Fiori; Fl. Ital. Crypt. 5: 425, f. 154 (1943)

= Isoetes histrix var. grandis (A.Braun) Batt. & Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes histrix var. lacunosa Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 251 (1866)

= Isoetes histrix var. loricata (A.Braun) Batt. & Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes histrix var. scutellata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 617 (1864)

= Isoetes histrix var. subinermis Durieu; Bull. Soc. Bot. France 8: 164 (1861), nom. nud.

= Isoetes histrix var. submersa Trab.; Fl. Algérie Tunisie 407 (1902)

= Isoetes hystrix subsp. algarbiensis Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. arcuata Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. cesatii Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. collina Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. corsica Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. hispanica Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. horridula Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. longifolia Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. longiseta Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. numidica Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. oceanica Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. oranensis Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. phrygia Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. todaroi Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes hystrix subsp. transiens Gand.; Fl. Eur. [Gandoger] 27: 197 (1891)

= Isoetes setacea Moris & De Not.; Fl. Caprar. 161 (1839) non Lam. 1789

species :

Isoetes hopei J.R.Croft; Blumea 26: 177-190 (1980)

Distribution: New Guinea (Irian Jaya)

42 NWG

Conservation Status: EN in Papua New Guinea

species :

Isoetes howellii Engelm.; Trans. Acad. Sci. St. Louis 4: 385 (1882)

Name: Howell´s Quillwort

Distribution: Canada (British Columbia); USA (California, Idaho, Montana, Oregon, Utah, Washington State); Mexico (Baja California Norte, Baja California Sur)

71 CDN 73 USA 76 USA 79 MEX

Synonyms: hidden

= Isoetes melanopoda f. californica (A.A.Eaton) Clute; Fern Allies 260 (1905)

= Isoetes melanopoda var. californica A.A.Eaton; Working List N. Amer. Pterid. 27 (1901)

= Isoetes nuda Engelm.; Trans. Acad. Sci. St. Louis 4: 385 (1882)

= Isoetes underwoodii L.F.Hend.; Bot. Gaz. 23: 124 (1897)

species :

Isoetes humilior F.Muell. ex A.Braun; Linnaea 25: 722 (1852)

Distribution: Australia (Tasmania)

50 AUS

Conservation Status: IUCN R

Synonyms: hidden

= Calamaria humilior (F.Muell. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes hookeri A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 538 (1868)

= Isoetes tasmanica Durieu; Bull. Soc. Bot. France 11: 104 (1864), non F. Muell.

species :

Isoetes hyemalis D.F.Brunt.; Castanea 59(1): 13 (1994)

Name: Winter Quillwort

Distribution: E-USA (North Carolina, South Carolina, Virginia)

75 USA 78 USA

Remarks: sometimes included in I. riparia, but accepted by Troia et al. 2016

Synonyms: hidden

= Isoetes canadensis var. amesii (A.A.Eaton) Clute; Fern Bull. 16: 54 (1908)

= Isoetes riparia var. amesii (A.A.Eaton ex Gilbert) Proctor; Amer. Fern J. 39: 119 (1949)

= Isoetes saccharata var. amesii A.A.Eaton; Rhodora 5(59): 278-279 (1903)

species :

Isoetes hypsophila Hand.-Mazz.; Anz. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 60: 95 (1923), et Symb. Sin. 6: 13 (1929)

Distribution: China (SW-Sichuan, NW-Yunnan)

36 CHI

species :

Isoetes inflata E.R.L.Johnson; J. Roy. Soc. Western Australia 67(1): 37 (1984)

Distribution: Australia (SW-Western Australia)

50 AUS

species :

Isoetes jaegeri Pitot; Bull. Inst. Franç. Afrique Noire, A 30: 58, f. 1-19 (1968)

Distribution: Burkina Faso

22 BKN

species :

Isoetes jamaicensis Hickey; Amer. Fern J. 71(3): 69 (1981)

Distribution: Jamaica

81 JAM

species :

Isoetes japonica A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1: 459 (1861)

Distribution: China; South Korea (Pyeongtaek area); Japan

36 CHI 38 JAP KOR

Synonyms: hidden

= Calamaria japonica (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes edulis Sieber ex Miq.; Prodr. Fl. Jap. 390 (1866-67)

species :

Isoetes jarmaniae D.F.Brunt., Kantvilas & M.Garrett; Phytotaxa 521(1): 29 (2021)

Distribution: Australia (Tasmania)

50 AUS

species :

Isoetes jejuensis H.K.Choi, Ch.Kim & J.Jung; J. Pl. Biol. 51(5): 356 (2008)

Distribution: South Korea (Cheju Isl.)

38 KOR

species :

Isoetes junciformis D.F.Brunt. & D.M.Britton; Amer. Fern J. 89(3): 187-197 (1999)

Name: Rush quillwort

Distribution: USA (S-Georgia)

78 USA

species :

Isoetes karstenii A.Braun; Verh. Bot. Vereins Prov. Brandenburg 4: 332 (1862)

Distribution: Colombia (Arauca, Boyacá, Cauca, Cundinamarca, Huila, Magdalena, Meta, Nariño, Risaralda, Santander, Tolima, Valle); Venezuela (Merida, Trujillo)

82 CLM VEN

Synonyms: hidden

= Isoetes colombiana (T.C.Palmer) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 176 (1981)

= Isoetes lechleri var. colombiana Palmer; Amer. Fern J. 19: 18 (1929)

= Isoetes steyermarkii H.P.Fuchs; (1960), nom. nud.

species :

Isoetes kashyapii M.Patel, M.N.Reddy & H.K.Goswami; Indian Fern J. 39(1): 231 (2022)

Distribution: India (Gujarat)

40 IND

species :

Isoetes killipii C.V.Morton; Amer. Fern J. 35: 49 (1945)

Distribution: Colombia (Antioquia, Boyacá, Cauca, Cundinamarca, Meta, Nariño, Putumayo, Risaralda, Santander, Tolima, Valle); Venezuela (Amazonas, Apure, Bolívar, Tachira); Guyana (Mt. Roraima); Ecuador

82 CLM GUY VEN 83 ECU

Synonyms: hidden

= Isoetes roncadorensis H.P.Fuchs; (1981), nom. nud.

= Isoetes roraimensis H.P.Fuchs; (1981), nom. nud.

species :

Isoetes kirkii A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 649 (1869)

Distribution: New Zealand (North Isl., South Isl.)

51 NZL

Synonyms: hidden

= Calamaria kirkii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes labri-draconis N.R.Crouch; Ferns S. Afr. Compreh. Guide: 746 (2011)

Distribution: South Africa (KwaZulu-Natal: E-Drakensberg)

27 SAF

species :

Isoetes lacustris L.; Sp. Pl. [Linnaeus] 2: 1100 (1753)

Name: Gewöhnliches Brachsenkraut, See-Brachsenkraut (DE), Lake Quillwort (EN)

Distribution: England (SW- & NW-England, Wales, Scotland, Orkney Isl., Shetland Isl.); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Germany (Baden-Württemberg, Bayern, +Mecklenburg-Vorpommern, Niedersachsen, Schleswig-Holstein); Switzerland; Austria; Poland; Czech Republic; Spain; Andorra; France; ?Romania; +Bulgaria; Estonia; Latvia; Lithuania; Belarus; European Russia (N-European Russia, C-European Russia); Siberia (Altai, Irkutsk, W-Siberia); ?Japan; Greenland; Canada (Labrador, Manitoba, New Brunswick, Newfoundland, Nova Scotia, Northern Territories, Ontario, Prince Edward Isl., Québec, Saskatchewan); St. Pierre et Miquelon; USA (Connecticut, Massachusetts, Maine, Michigan, Minnesota, New Hampshire, New Jersey, New York, Rhode Island, Tennessee, Virginia, Vermont, Wisconsin)

10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER NET POL SWI 12 ADO FRA SPA 13 BUL 14 BLR EST LAT LIT RUS 30 SIB 70 CDN GNL 71 CDN 72 CDN SPM 74 USA 75 USA 78 USA

Synonyms: hidden

= Calamaria lacustris (L.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Calamaria setacea (Lam.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes atrovirens Th.Fr.; Bot. Not. (1863): 48 (1865)

= Isoetes hieroglyphica A.A.Eaton; Fernwort Pap. 10 (1900)

= Isoetes lacustris subsp. abbreviata Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. arvernensis Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. cambrica Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. editorum Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. errabunda Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. hieroglyphica (A.A.Eaton) Taylor & Luebke

= Isoetes lacustris subsp. lacustris Hook.f.; Student´s Fl. Brit. Isl., ed. 1: 471 (1870)

= Isoetes lacustris subsp. lamottei Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. langei Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. latifolia Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. noltei Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. obscura Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. pervaga Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. rivalis Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. rossica Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. scanica Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. scotica Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. septentrionalis Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. spreta Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. succedanea Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris subsp. suecica Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes lacustris var. curvifolia Casp.; Süsswasserflora 15: 10 (pro syn.) (1936)

= Isoetes lacustris var. elatior Fliche; Isoetes des Vosges 7 (1879)

= Isoetes lacustris var. falcata Gay; Bull. Soc. Bot. France 10: 386 (1863)

= Isoetes lacustris var. falcatum Fliche; Isoetes des Vosges 7 (1879)

= Isoetes lacustris var. fragilis Gray; Nat. Arr. Brit. Pl. 2: 24 (1821 publ. 1822)

= Isoetes lacustris var. gracilis Gray; Nat. Arr. Brit. Pl. 2: 24 (1821 publ. 1822)

= Isoetes lacustris var. humilior Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)

= Isoetes lacustris var. humilis Mer; Bull. Soc. Bot. France 28: 72 (1881)

= Isoetes lacustris var. intermedia Mer; Bull. Soc. Bot. France 28: 72 (1881)

= Isoetes lacustris var. leiospora Klinggr.; Schriften Naturf. Ges. Danzig 6, 1: 20 (1884)

= Isoetes lacustris var. minor Milde; Sporenpfl. 141 (1865)

= Isoetes lacustris var. moorei (D.Moore) Hook.; Student´s Fl. Brit. Isl., ed. 3, 526 (1884)

= Isoetes lacustris var. moorei (D.Moore) Syme & N.E.Br.; Engl. Bot., ed. 3B, 12: 5 (1886), isonym

= Isoetes lacustris var. rectifolia Casp.; Süsswasserflora 15: 10 (1936)

= Isoetes lacustris var. recurva Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)

= Isoetes lacustris var. setacea Léman; Dict. Sci. Nat., ed. 2 [F.Cuvier] 24: 24 (1822)

= Isoetes lacustris var. stricta (J.Gay) Mer; Bull. Soc. Bot. France 28: 72 (1881)

= Isoetes lacustris var. stricta (J.Gay) Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883), isonym

= Isoetes lacustris var. strictum J.Gay; Bull. Soc. Bot. France 10: 392 (1863)

= Isoetes lacustris var. tenella Léman; Dict. Sci. Nat., ed. 2 [F.Cuvier] 24: 24 (1822)

= Isoetes lacustris var. tenuifolia A.Braun ex Milde; Sporenpfl. 141 (1865)

= Isoetes lacustris var. terrestris Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 327 (1883)

= Isoetes lacustris unranked vergens Gand.; Fl. Eur. [Gandoger] 27: 193 (1891)

= Isoetes leiospora Klinggr.; Schriften Naturf. Ges. Danzig 6(1): 20 (1884)

= Isoetes macrospora Durieu; Bull. Soc. Bot. France 11: 101 (1864)

= Isoetes macrospora f. hieroglyphica (A.A.Eaton) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 194 (1922)

= Isoetes morei D.Moore; J. Bot. 16: 353, t. 199 (1878)

= Isoetes setacea Lam.; Encycl. [J.Lamarck et al.] 3: 314 (1789)

= Isoetes tenella Lem. ex Desv.; Mém. Soc. Linn. Paris 6(2): 179 (1827)

= Isoetes tuckermani var. harveyi (A.A.Eaton) Clute; Fern Allies 226, 227, 260 (1905)

species :

Isoetes laosiensis C.Kim & H.K.Choi; Amer. Fern J. 100(1): 48 (2010)

Distribution: Laos

41 LAO

species :

Isoetes laurentiana D.F.Brunt.; Botany (Ottawa) 97(11): 579 (2019)

Distribution: Canada (Québec)

72 CDN

species :

Isoetes lechleri Mett.; Fil. Lechl. 2: 36 (1859)

Distribution: Colombia (Arauca, Boyacá, Cauca, Cundinamarca, Huila, Meta, Nariño, Santander); Ecuador (Chimborazo, Pinchincha, Napo); Peru (Junín, Puno, Ayacucho, Ancash); Bolivia (Cochabamba, La Paz)

82 CLM 83 BOL ECU PER

Synonyms: hidden

= Calamaria lechleri (Mett.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes arumiana H.P.Fuchs; (1959), nom. nud.

= Isoetes glacialis Aspl.; Ark. Bot. 20A(7): 35, fig. 7 (1926)

= Isoetes karstenii var. anomala (T.C.Palmer) R.L.Small & Hickey; Amer. Fern J. 22: 130 (1933)

= Isoetes laevis U.Weber; Hedwigia 63: 252, fig. 30-31 (1922)

= Isoetes lechleri var. anomala T.C.Palmer; Amer. Fern J. 22: 130 (1933)

= Isoetes peruviana U.Weber; Hedwigia 63: 246, fig. 12-13 (1922)

= Isoetes rimbachiana H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(2): 226 (1982), nom. nov. pro I. lechleri var. anomala

= Isoetes socia A.Braun; Verh. Bot. Vereins Prov. Brandenburg 4: 332 (1862)

species :

Isoetes libanotica Musselman, Bolin & R.D.Bray; Novon 21(3): 295 (2011)

Distribution: Lebanon

34 LEB

species :

Isoetes lithophila N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 135 (1922) (lithophylla)

Name: Rock Quillwort

Distribution: USA (SC-Texas: Burnett Co., Mason Co., Llano Co.)

73 USA 77 USA

Conservation Status: IUCN V

species :

Isoetes longifolia J.B.S.Pereira; Syst. Bot. 47(2): 304 (2022)

Distribution: Brazil (Bahia)

84 BRA

species :

Isoetes longissima Bory & Durieu; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1165 (1844)

subspecies :

subsp. longissima

Distribution: Portugal; Spain; Baleares; France; Corsica; Sardinia; Italy; Sicily; Morocco; Algeria; Tunisia; Libya

12 BAL COR FRA POR SAR SPA 13 ITA SIC 20 ALG LBY MOR TUN

Synonyms: hidden

= Calamaria adspersa (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Calamaria longissima (Bory & Durieu) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Calamaria tenuissima (Boreau) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes adspersa A.Braun; Expl. Sci. Algérie, Atlas: tab. 37, rig. 3 (1849)

= Isoetes adspersa var. perreymondii A.Braun; Bull. Soc. Bot. France 12: 261 (1865)

= Isoetes aspersum Asch. & Graebn.; Syn. Mitteleur. Fl. [Ascherson & Graebner] 1: 169 Hauptreg. 23 (1897)

= Isoetes baetica Willk.; Prodr. Fl. Hispan. [Willkomm & Lange] 1: 14 (1861)

= Isoetes boryana var. lereschii Rchb.f. ex Ler. & H.Lév.; Deux Excurs. Bot. 118 (1880)

= Isoetes capillacea Bory; Compt. Rend. Hebd. Séances Acad. Sci. 23: 620 (1846)

= Isoetes chaboissaei Hy; J. Bot. [Morot] 8: 97 (1894)

= Isoetes chaboissaei Nym.; Consp. Fl. Eur. Suppl. 1: 871 (1884)

= Isoetes decipiens Bory; Flora 29: 719 (1846) (nom. prov)

= Isoetes dubia var. emersa Gennari; Nuovo Giorn. Bot. Ital. 2: 96 (1870)

= Isoetes dubia var. maculosa Gennari; Comment. Soc. Crittog. Ital. 1(2): 105 (1861)

= Isoetes grammontii Bory; Fil. Eur. 280 (1867)

= Isoetes iluliana Gennari; Comment. Soc. Crittog. Ital. 1: 4 (1861) (febr.) (syn. subst.)

= Isoetes lacustris Moris; Stirp. Sard. Elench. 3: 12 (1829) non L. 1753

= Isoetes lineolata Durieu ex Motelay & Vendryès; Actes Soc. Linn. Bordeaux 36: 381 (1883)

= Isoetes longissima subsp. adspersa (A.Braun) Troia & Greuter; Willdenowia 44(2): 288 (2014) (synonymy fide Troia & Rouhan 2018)

= Isoetes longissima subsp. intermedia (Trab.) Troia & Greuter; Willdenowia 44(2): 288 (2014)

= Isoetes longissima subsp. tenuissima (Boreau) Troia & Greuter; Willdenowia 44(2): 288 (2014) (synonymy fide Troia & Rouhan 2018)

= Isoetes perreymondii (Bory) Duval-Jouve; Bull. Soc. Bot. France 16: 213 (1869)

= Isoetes setacea Bory; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1165 (1844)

= Isoetes setacea subsp. longissima (Bory & Durieu) Greuter & Burdet; Willdenowia 14(1): 37 (1984)

= Isoetes setacea var. perreymondii Bory; Compt. Rend. Hebd. Séances Acad. Sci. 18: 1165 (1844)

= Isoetes sicula Tod. ex Ces. & De Not.; Comment. Soc. Crittog. Ital. 1(2): 103 (1861)

= Isoetes tenuissima Boreau; Bull. Soc. Industr. Angers 21: 269 (1850)

= Isoetes tenuissima var. capillifolia Le Grand; Bull. Soc. Bot. France 42: 50 (1895)

= Isoetes tenuissima var. chaboissaei Le Grand; Bull. Soc. Bot. France 42: 50 (1895)

= Isoetes tenuissima var. latifolia Le Grand; Bull. Soc. Bot. France 42: 50 (1895)

= Isoetes tenuissima var. longifolia Le Grand; Bull. Soc. Bot. France 42: 50 (1895)

= Isoetes tenuissima var. tenuissima Le Grand; Bull. Soc. Bot. France 42: 50 (1895)

= Isoetes variabilis proles tenuissima (Boreau) Rouy; Fl. France [Rouy & Foucaud] 14: 477 (1913), nom. illeg.

= Isoetes variabilis proles viollaei (Hy) Rouy; Fl. France [Rouy & Foucaud] 14: 478 (1913)

= Isoetes variabilis Le Grand; Mém. Soc. Imp. Sci. Nat. Cherbourg 1896: 11 (1896) (nom. illeg.)

= Isoetes variabilis subsp. tenuissima (Boreau) R.C.V.Douin; Fl. Ill. France 12: 118 (1934), nom. illeg.

= Isoetes variabilis var. chaboissaei (Hy) P.E.Fourn.; Quatre Fl. France 5 (1934)

= Isoetes variabilis var. tenuissima (Boreau) P.E.Fourn.; Quatre Fl. France 5 (1934), nom. illeg.

= Isoetes variabilis var. viollaei (Hy) P.E.Fourn.; Quatre Fl. France 5 (1934)

= Isoetes velata A.Braun; Expl. Sci. Algérie, Atlas: tab. fig. 1 (1849)

= Isoetes velata f. baetica (Willk.) Prada; Acta Bot. Malac. 8: 88 (1983)

= Isoetes velata f. immaculata Maire; Encycl. Biol. 33: 90 (1952)

= Isoetes velata f. lereschii (Rchb.f. ex Leresche & Levier) Prada; Lazaroa 3: 33 (1982)

= Isoetes velata f. longissima (Bory & Durieu) N.Pfeiff.; Ann. Missouri Bot. Gard. 9(2): 120 (1922)

= Isoetes velata subsp. adspersa (A.Braun) Batt. & Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes velata subsp. algeriensis Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes velata subsp. australis Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes velata subsp. barbara Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes velata subsp. calaostiensis Gennari; Sp. Sard. 31 (1867)

= Isoetes velata subsp. divergens Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes velata subsp. gallica Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes velata subsp. intermedia (Trab.) Maire & Weiller; Fl. Afrique N. 1: 89 (1952)

= Isoetes velata subsp. longissima (Bory & Durieu) Greuter & Burdet; Willdenowia 14(1): 37 (1984)

= Isoetes velata subsp. redux Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes velata subsp. robusta Gand.; Fl. Eur. [Gandoger] 27: 194 (1891)

= Isoetes velata subsp. tenuicula Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes velata subsp. tenuissima (Boreau) O.Bolòs & Vigo; Butl. Inst. Catalana Hist. Nat., Secc. Bot. 1: 64 (1974)

= Isoetes velata subsp. trinacria Gand.; Fl. Eur. [Gandoger] 27: 195 (1891)

= Isoetes velata var. intermedia Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes velata var. brevifolia (A.Braun) Franch.; Bull. Soc. Bot. France 31: 349 (1884)

= Isoetes velata var. brevifolia A.Braun; Akad. Wiss. Wien Sitzungsber., Math.-Naturwiss. Kl., Abt. 1: 605 (1863), et Fl. Algérie Tunisie 407 (1902)

= Isoetes velata var. humilior A.Braun ex Mares & Vigin.; Pl. Vasc. Iles Baleares 322 (1890)

= Isoetes velata var. intermedia A.Braun; Expl. Sci. Algérie, t. 37 (1848)

= Isoetes velata var. lereschii (Rchb.f. ex Leresche & Levier) Molina Abril & Sard.Rosc.; Guineana 10: 346 (2004)

= Isoetes velata var. longissima A.Braun; Expl. Sci. Algérie, t. 37, fig. 2 (1848)

= Isoetes velata var. perreymondii (Bory) Franch.; Süsswasserflora 15: 17 (1936)

= Isoetes velata var. sicula Gennari; Fil. Eur. 282 (1867)

= Isoetes velata var. sicula Tod.; Giorn. Sci. Nat. Econ. Palermo 1: 250 (1866)

= Isoetes vellata var. boryana (Durieu) M.I.Romero & Real; Bot. J. Linn. Soc. 148(4): 464 (2005)

= Isoetes viollaei Hy; Bull. Herb. Boissier App. 1: 23 (1894)

subspecies :

subsp. asturicensis (M.Laínz) L.Síez, Nava & Fern.Prieto; Naturalia Cantabricae 8(Especial): 64 (2020)

Distribution: Spain

12 SPA

Conservation Status: IUCN R

Synonyms: hidden

= Isoetes asturicensis (M.Laínz) M.Laínz; Anuário Soc. Brot. 39: 118 (1973)

= Isoetes velata subsp. asturicensis (M.Laínz) Rivas Mart. & Prada; Lazaroa 2: 327 (1980)

= Isoetes boryana subsp. asturicensis M.Laínz; Bol. Inst. Estud. Asturianos, Supl. Ci. 15: 6 (1970)

= Isoetes velata var. asturicense (M.Laínz) E.Rico & X.Giráldez; Anales Jard. Bot. Madrid 46(2): 585 (1989 publ. 1990)

subspecies :

subsp. perralderiana (Milde) Troia & Greuter; Willdenowia 44(2): 288 (2014)

Distribution: Algeria

20 ALG

Synonyms: hidden

= Calamaria perralderiana (Durieu & Letourn. ex Milde) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes perralderiana Durieu & Letourn.; Pl. Algérie [Exsicc.] 157 (1862)

= Isoetes perralderiana Durieu & Letourn. ex Milde; Fil. Eur. 282 (1867)

= Isoetes velata subsp. perralderiana (Durieu & Letourn.) Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes velata var. perralderiana (Durieu & Letourn.) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 121 (1922)

species :

Isoetes longpingii Y.H.Yan, Y.F.Gu & J.P.Shu; Guihaia 42(10): 1625 (2022)

Distribution: China (Hunan)

36 CHI

species :

Isoetes louisianensis Thieret; Sida, Bot. Misc. 5: 129 (1973)

Name: Louisiana Quillwort

Distribution: USA (S-Louisiana, Mississippi, Alabama)

78 USA

Remarks: may be a hybrid I. engelmannii x melanopoda

Conservation Status: IUCN E

species :

Isoetes luetzelburgii U.Weber ex Luetzelb.; Estud. Bot. Nordeste 3: 257 (1926)

Distribution: Brazil (Paraíba, Rio Grande do Norte)

84 BRA

species :

Isoetes malinverniana Ces. & De Not.; Index Sem. (Geneva) 1858: 36 (1858)

Distribution: NW-Italy (Vercelli, Novara, Torino)

13 ITA

Conservation Status: IUCN E

Synonyms: hidden

= Calamaria malinverniana (Ces. & De Not.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes maritima Underw.; Bot. Gaz. 13: 94 (1888)

Name: Spiny-spored Quillwort

Distribution: Russian Far East (Commander Isl.); Alaska; Aleutes; C-Canada (Vancouver Isl., British Columbia); NW-USA (Washington)

31 RFE 70 ASK 71 CDN 73 USA

Synonyms: hidden

= Isoetes beringensis Kom.; Bull. Jard. Bot. Acad. Sci. USSR 30(1-2): 196 (1932)

= Isoetes braunii var. maritima (Underw.) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 174 (1922)

= Isoetes echinospora f. maritima (Underw.) Clute; Fern Bull. 16: 55 (1908)

= Isoetes echinospora subsp. maritima (Underw.) Á.Löve; Amer. Fern J. 52: 121 (1962)

= Isoetes echinospora var. maritima (Underw.) A.A.Eaton; Fern Bull. 13: 52 (1905)

= Isoetes echinospora var. savilei B.Boivin; Amer. Fern J. 51: 85 (1961)

= Isoetes macounii A.A.Eaton; Fern Bull. 8: 12 (1900)

= Isoetes muricata subsp. maritima (Underw.) Hultén; Ark. Bot., n. s. 7(1): 8 (1968)

= Isoetes setacea subsp. maritima (Underw.) Holub; Acta Horti Bot. Prag. 1964: 55 (1964)

species :

Isoetes martii A.Braun ex Kuhn; Fl. Bras. [Martius] 1(2): 646, t. 78 (1884)

Distribution: Brazil (Espírito Santo, Minas Gerais, Rio de Janeiro, São Paulo)

84 BRA

Synonyms: hidden

= Calamaria martii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes mattaponica Musselman & W.C.Taylor; Novon 11(2): 200 (-204, figs. 1-6) (2001)

Name: Mattaponi quillwort

Distribution: USA (Virginia: Mattaponi River)

78 USA

species :

Isoetes maxima Hickey, Macluf & Link-Pérez; Amer. Fern J. 99(3): 194 (2010)

Distribution: Brazil (Rio Grande do Sul)

84 BRA

species :

Isoetes melanopoda Gay & Durieu; Bull. Soc. Bot. France 11: 102 (1864)

subspecies :

subsp. melanopoda

Name: Black-footed Quillwort

Distribution: USA (Alabama, Arkansas, Georgia, Iowa, Idaho, Illinois, Indiana, Kansas, Kentucky, Louisiana, Minnesota, Missouri, Mississippi, Montana, North Carolina, Nebraska, New Jersey, Oklahoma, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia)

73 USA 74 USA 75 USA 76 USA 77 USA 78 USA

Synonyms: hidden

= Calamaria melanopoda (J.Gay) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes melanopoda f. pallida (Engelm.) Fernald; Rhodora 51: 103 (1949)

= Isoetes melanopoda var. pallida Engelm.; Trans. Acad. Sci. St. Louis 4: 387 (1882)

subspecies :

subsp. silvatica D.F.Brunt. & D.M.Britton; Castanea 71(1): 26 (2006)

Distribution: E-USA (North Carolina)

75 USA 78 USA

species :

Isoetes melanospora Engelm.; Trans. Acad. Sci. St. Louis 3: 395 (1877)

Name: Blackspore Quillwort

Distribution: USA (C-Georgia, South Carolina: Lancaster Co.)

78 USA

Synonyms: hidden

= Calamaria melanospora (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes melanotheca Alston; Bol. Soc. Brot., Ser. 2A, 30: 15 (1956)

Distribution: Guinea-Bissau; Guinea; Togo; Benin

22 BEN GNB GUI TOG

Conservation Status: IUCN R

species :

Isoetes mexicana Underw.; Bot. Gaz. 13: 93 (1888)

Distribution: Mexico (Aguascalientes, Chihuahua, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Queretaro, Zacatecas); N-Guatemala

79 MEX 80 GUA

species :

Isoetes microvela D.F.Brunt.; Rhodora 100(903): 270-274, f. 1-3, 4e-h, 5 (1998)

Name: Thin-wall quillwort

Distribution: USA (North Carolina: coastal)

78 USA

species :

Isoetes minima A.A.Eaton; Fern Bull. 6: 30 (1898)

Name: Midget quillwort

Distribution: SW-Canada (British Columbia); USA (Washington State)

71 CDN 73 USA

Remarks: valid species fide Taylor et al. 2003

Synonyms: hidden

= Isoetes howellii var. minima (A.A.Eaton) N.Pfeiff.; Ann. Missouri Bot. Gard. 9(2): 142 (1922)

species :

Isoetes mississippiensis S.W.Leonard, W.C.Taylor, Musselman & R.D.Bray; PhytoKeys 74: 99 (2016)

Distribution: USA (S-Mississippi)

78 USA

species :

Isoetes mongerensis E.R.L.Johnson; J. Roy. Soc. Western Australia 67(1): 39 (1984)

Distribution: Australia (SW-Western Australia)

50 AUS

species :

Isoetes montezumae A.A.Eaton; Fern Bull. 5: 25 (1897)

Distribution: Mexico (México State)

79 MEX

Remarks: recognized in Troia et al. 2016, included in I. mexicana by other authors

species :

Isoetes mourabaptistae J.B.S.Pereira, P.G.Windisch, Lorscheitt. & Labiak; Amer. Fern J. 102(2): 175 (2012)

Distribution: Brazil (Rio Grande do Sul)

84 BRA

species :

Isoetes muelleri A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 541 (1868)

Distribution: Australia (Australian Capital Territory, Western Australia, Northern Territory, South Australia, Queensland, Victoria, Tasmania)

50 AUS

Synonyms: hidden

= Calamaria muelleri (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes stuartii A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1868: 539 (1868)

= Isoetes tenuissima F.Muell.; (1868), nom. nud.

species :

Isoetes naipiana P.G.Windisch, Lorscheitt. & Nervo; Willdenowia 44: 394 (2014)

Distribution: Brazil (Rio Grande do Sul)

84 BRA

species :

Isoetes nana J.B.S.Pereira; PhytoKeys 89: 93 (2017)

Distribution: Brazil (Rio de Janeiro)

84 BRA

species :

Isoetes neoguineensis Baker; Bull. Misc. Inform. Kew 1899: 122 (1899), et ex Nat. Pflanzenfam. [Engler & Prantl] I, 4: 776 (1902)

Distribution: New Guinea (Papua New Guinea: Owen Stanley Ranges)

42 NWG

Conservation Status: IUCN R (twice found)

Synonyms: hidden

= Isoetes neoguineensis Baker ex F.Muell.; Ann. Rep. Brit. N. Guinea 1897-8: 149 (1898)

= Isoetes neoguineensis var. rheophila J.R.Croft; Blumea 26: 177-190 (1980)

species :

Isoetes nigritiana A.Braun; Filic. Afr. [Kuhn] 196 (1868)

Distribution: Mali; Burkina Faso; Ivory Coast; Nigeria; Central African Republic; Sudan

22 BKN IVO MLI NGA 23 CAF 24 SUD

Synonyms: hidden

= Isoetes garnieri Chev. & Monnier; Bull. Mus. Natl. Hist. Nat. 18(1): 91-979 1 fig. (1946)

= Calamaria nigritiana (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes nigroreticulata Verdc.; Fl. Trop. E. Africa, Isoet. 2 (2005)

Distribution: Kenya

25 KEN

species :

Isoetes novogranadensis H.P.Fuchs; Caldasia 8: 314 (1969)

Distribution: Colombia (Bolívar, Boyacá, Cauca, Cundinamarca, Huila, Nariño, Putumayo, Valle); Ecuador; Peru

82 CLM 83 ECU PER

Synonyms: hidden

= Isoetes dichotoma L.E.Mora & W.Hagemann; Mutisia 43: 5 (1977)

= Isoetes hagemannii H.P.Fuchs; (1981), nom. nud.

= Isoetes leistii H.P.Fuchs; (1981), nom. nud.

= Isoetes narinensis H.P.Fuchs; (1981), nom. nud.

= Isoetes pichichensis H.P.Fuchs; (1981), nom. nud.

= Isoetes quilindanensis H.P.Fuchs; (1981), nom. nud.

species :

Isoetes nuttallii A.Braun ex Engelm.; Amer. Naturalist 8: 215 (1874)

Name: Nuttall´s Quillwort

Distribution: C-Canada (Vancouver Isl.); W-USA (California, Oregon, Washington)

71 CDN 73 USA 76 USA

Synonyms: hidden

= Calamaria nuttallii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Calamaria suksdorfii (Baker) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes opaca Nutt. ex Engelm.; Trans. Acad. Sci. St. Louis 4: 389 (1886), pro syn.

= Isoetes suksdorfii Baker; Handb. Fern-Allies 132 (1887)

species :

Isoetes occidentalis Hend.; Bull. Torrey Bot. Club 27: 358 (1900)

Name: Western Quillwort

Distribution: Alaska; Aleutes; C-Canada (British Columbia); NW-USA (Washington, Oregon, C-California, Idaho, Montana, Wyoming, Colorado)

70 ASK 71 CDN 73 USA 76 USA

Synonyms: hidden

= Isoetes echinospora f. flettii (A.A.Eaton) Clute; Fern Bull. 16: 55 (1908)

= Isoetes echinospora subsp. flettii (A.A.Eaton) Piper & Beattie; Fl. N.W. Coast [Piper & Beattie] 16 (1915)

= Isoetes echinospora var. flettii A.A.Eaton; Fern Bull. 13: 51 (1905)

= Isoetes flettii (A.A.Eaton) N.Pfeiff.; Ann. Missouri Bot. Gard. 9(2): 186 (1922)

= Isoetes howellii f. piperi (A.A.Eaton) Clute; Fern Bull. 16: 56 (1908)

= Isoetes howellii var. piperi (A.A.Eaton) Clute; Fern Allies 246, 258 (1905)

= Isoetes lacustris subsp. paupercula (Engelm.) Feilberg; Nordic J. Bot. 4: 155 (1984)

= Isoetes lacustris var. paupercula Engelm.; Trans. Acad. Sci. St. Louis 4: 377 (1882)

= Isoetes macrospora f. paupercula (Engelm.) Clute; Fern Bull. 16: 56 (1908)

= Isoetes maritima var. flettii (A.A.Eaton) C.F.Reed; Amer. Fern J. 35: 85 (1945)

= Isoetes occidentalis subsp. paupercula (Engelm.) Feilberg; Nordic J. Bot. 4(2): 155 (1984)

= Isoetes occidentalis var. piperi (A.A.Eaton) A.Nelson & J.F.Macbr.; Bot. Gaz. 61(1): 30 (1916)

= Isoetes paupercula (Engelm.) A.A.Eaton; Proc. U.S. Natl. Mus. 23 (1226): 649 (1901)

= Isoetes piperi A.A.Eaton; Fern Bull. 11: 85 (nom. nud.) (1903), 13: 51 (1905)

species :

Isoetes olympica A.Br.; Fil. Eur. 285 (1867)

Distribution: Turkey (Anatolia); Lebanon; S-Syria

34 LEB SYR TUR

Remarks: histrix agg.

Synonyms: hidden

= Calamaria olympica (A.Br.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes orcuttii A.A.Eaton; Fern Bull. 6: 30 (1898) (nom. nud.), et 8: 13 (1900)

Name: Orcutt´s Quillwort

Distribution: USA (S-California); Mexico (Baja California Norte)

76 USA 79 MEX

Synonyms: hidden

= Isoetes nuttallii var. orcuttii (A.A.Eaton) Clute; Fern Allies 253 (1905)

species :

Isoetes organensis U.Weber; Estud. Bot. Nordeste 3: 256 (1922)

Distribution: Brazil (Minas Gerais, Rio de Janeiro)

84 BRA

species :

Isoetes orientalis Hong Liu & Q.F.Wang; Novon 15(1): 164 (2005)

Distribution: China (Zhejiang)

36 CHI

Remarks: status uncertain, probably a hybrid Isoetes sinensis x yunguiensis

species :

Isoetes ovata N.Pfeiff.; Ann. Missouri Bot. Gard. 8(2): 109 (1922)

Distribution: Guyana; French Guiana

82 FRG GUY

Synonyms: hidden

= Isoetes clavata U.Weber; Ber. Deutsch. Bot. Ges. 52: 121 (1934)

species :

Isoetes pallida Hickey; Amer. Fern J. 78(1): 35 (1988)

Distribution: Mexico (Campeche, Guerrero, Oaxaca); Belize

79 MEX 80 BLZ

Conservation Status: IUCN R

species :

Isoetes palmeri H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 168 (1981)

Distribution: Colombia (Arauca, Boyacá, Cauca, Cundinamarca); Venezuela (Merida); Ecuador (Chimborazo)

82 CLM VEN 83 ECU

Synonyms: hidden

= Isoetes cleeffii H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 84(2): 177 (1981)

species :

Isoetes panamensis Maxon & C.V.Morton; Ann. Missouri Bot. Gard. 26(4): 272 (1939)

Distribution: Guatemala; Nicaragua; Costa Rica; Panama; Colombia; Ecuador; E-Peru; Bolivia (Cochabamba, La Paz); Brazil (Bahia, Maranhão, Mato Grosso do Sul, Mato Grosso, Paraná, Tocantins); Paraguay (Alto Paraná, Caaguazu)

80 COS GUA NIC PAN 82 CLM 83 BOL ECU PER 84 BRA 85 PAR

Synonyms: hidden

= Isoetes pacifica Svenson; Amer. Fern J. 34: 123 (1944)

= Isoetes savannarum L.D.Gómez; Phytologia 49(4): 339 (1981)

species :

Isoetes parvula Hickey; Fieldiana, Bot. 34: 92 (1994)

Distribution: Peru (Ayacucho)

83 PER

Conservation Status: Known from type only

species :

Isoetes pedersenii H.P.Fuchs ex E.I.Meza & Macluf; Anais Acad. Brasil. Ci. 82(2): 354 (2010)

Distribution: Brazil (Mato Grosso do Sul); Paraguay; Argentina (Corrientes)

84 BRA 85 ARG PAR

species :

Isoetes perrieriana Iversen; Dansk Bot. Ark. 7: 200-201 (1932)

Distribution: Madagascar

29 MDG

species :

Isoetes philippinensis Merr. & L.M.Perry; Amer. Fern J. 30: 19 (1940)

Distribution: Sulawesi; Philippines (Lanao del Norte)

42 PHI SUL

Conservation Status: CR in the Philippines, recently rediscovered (Amoroso et al. 2022)

species :

Isoetes phrygia Hausskn.; Mitth. Thüring. Bot. Vereins, N. F. 13: 72 (1899)

Distribution: Greece (S-Peloponnisos, Kiklades); Crete (rare); Turkey (Anatolia)

13 GRC KRI 34 TUR

Synonyms: hidden

= Isoetes histrix var. phrygia Boiss.; Fl. Orient. [Boissier] 5: 748 (1884)

species :

Isoetes pichisermollii Eb.Fisch. & Lobin; Willdenowia 52(3): 330-331 (2022)

Distribution: Burundi

23 BUR

Remarks: cited by Pichi-Sermolli 1983/1985 as ´Isoetes abyssinica´

species :

Isoetes pirinica D.F.Brunt. & D.Ivanova; Fern Gaz. 21(6): 247 (2021)

Distribution: Bulgaria

13 BUL

Remarks: close to I. lacustris; to be searched for in N-Greece

species :

Isoetes pitotii Alston; Bol. Soc. Brot., Ser. 2, 30: 15 (1956)

Distribution: Guinea; Mali; Sudan

22 GUI MLI 24 SUD

species :

Isoetes precocia R.L.Small & Hickey; Amer. Fern J. 91(2): 61 (-62, figs. 12-13, 18-19) (2001)

Distribution: Colombia (Boyaca, Cundinamarca); Venezuela (Merida, Trujillo)

82 CLM VEN

species :

Isoetes pringlei Underw.; Zoë 1: 98 (1890)

Distribution: Mexico (Jalisco, Mexico State, Chiapas)

79 MEX

Conservation Status: IUCN R

species :

Isoetes prototypus D.M.Britton & Goltz; Canad. J. Bot. 69: 278 (1991)

Name: Spike quillwort

Distribution: Canada (New Brunswick, Nova Scotia); USA (Maine, New Hampshire)

72 CDN 75 USA

Conservation Status: IUCN V

species :

Isoetes pseudojaponica M.Takamiya, Watan. & Ono; Acta Phytotax. Geobot. 48(2): 107 (1998)

Distribution: Japan

38 JAP

Conservation Status: VU in Japan

species :

Isoetes pusilla C.R.Marsden & Chinnock; Muelleria 8(1): 57 (1993)

Distribution: Australia (Victoria)

50 AUS

species :

Isoetes quiririensis J.B.S.Pereira & Labiak; Syst. Bot. 38(4): 869 (2014)

Distribution: Brazil (Paraná, Santa Catarina)

84 BRA

species :

Isoetes ramboi Herter; Revista Sudamer. Bot. 8 (1-2): 19-25 (1949)

Distribution: Brazil (Mato Grosso, Rio Grande do Sul, Santa Catarina)

84 BRA

species :

Isoetes riparia Engelm.; Flora 29: 178 (1846)

Name: Shore Quillwort

Distribution: USA (Delaware, District of Columbia, Maryland, North Carolina, New Jersey, New York, Pennsylvania, Rhode Island, South Carolina, Vermont, Virginia, West Virginia)

75 USA 78 USA

Remarks: riparia agg.

Synonyms: hidden

= Calamaria riparia (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes amesii A.A.Eaton; Fern Bull. 11: 103 (1903)

= Isoetes foveolata var. plenospora A.A.Eaton; Rhodora 5: 280 (1903)

= Isoetes riparia f. saccharata (Engelm.) Clute; Fern Bull. 16: 57 (1908)

= Isoetes riparia f. saccharata (Engelm.) Proctor; Amer. Fern J. 39: 119 (1949)

= Isoetes riparia var. palmeri (A.A.Eaton) Proctor; Amer. Fern J. 39: 119 (1949)

= Isoetes riparia var. reticulata (A.A.Eaton) Proctor; Amer. Fern J. 39: 119 (1949)

= Isoetes riparia var. riparia Proctor; Amer. Fern J. 39: 119 (1949)

= Isoetes saccharata f. palmeri (A.A.Eaton) A.A.Eaton; Fern Bull. 11: 25 (1903)

= Isoetes saccharata f. reticulata A.A.Eaton; Fern Bull. 11: 25 (1903)

= Isoetes saccharata var. palmeri A.A.Eaton; Proc. Biol. Soc. Washington 14(10): 49 (1901)

= Isoetes saccharata var. reticulata A.A.Eaton; Proc. Biol. Soc. Washington 14(10): 49 (1901)

species :

Isoetes rwandensis Eb.Fisch. & Lobin; Willdenowia 52(3): 332 (2022)

Distribution: Rwanda

23 RWA

species :

Isoetes sabatina Troìa & Azzella; Pl. Biosystems 147(4): 1052 (2013)

Distribution: Italy

13 ITA

species :

Isoetes saccharata Engelm.; Manual [A.Gray], ed. 5: 676 (1867)

Name: Sugar quillwort

Distribution: USA (District of Columbia, Delaware, Maryland, Virginia)

78 USA

Synonyms: hidden

= Calamaria saccharata (Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

species :

Isoetes sahyadrii Mahab.; [Curr. Sci. 7(2): 62, f. 1 (1938), nom. inval.] ex L. N. Rao, Curr. Sci. 13(11): 286-287 (1944) (sahyadriensis auct.), nom. cons. prop.

Distribution: India (Andhra Pradesh, Karnataka, Maharashtra, Madhya Pradesh, Rajasthan)

40 IND

Synonyms: hidden

= Isoetes panchananii D.D.Pant & Srivast.; Proc. Indian Acad. Sci., B 28: 243, t. 13, 14, f. 1, 2 (1962)

= Isoetes panchananii var. panchmarhiensis G.K.Srivast., Mritt.Srivast. & P.K.Shukla; Indian Fern J. 14(1-2): 44 (1998)

= Isoetes panchaganiensis G.K.Srivast., D.D.Pant & P.K.Shukla; Amer. Fern J. 83(4): 107 (1993) (fide Fraser-Jenkins 1997)

= Isoetes panchaganiensis var. kemangundiensis G.K.Srivast., D.D.Pant & P.K.Shukla; Amer. Fern J. 83(4): 114 (1993) (fide Fraser-Jenkins 1997)

= Isoetes rajasthanensis Gena & Bhardwaja; J. Bombay Nat. Hist. Soc. 81(1): 165 (1984)

= Isoetes reticulata Gena & Bhardwaja; J. Bombay Nat. Hist. Soc. 81(1): 166 (1984)

= Isoetes sampathkumaranii L.N.Rao; Curr. Sci. 13: 286 (1944)

species :

Isoetes sajanensis Stepanov; Red Book Krasnoyarsk. Territ. [ed. 3] 2(2): 497 (2022)

Distribution: Siberia (Krasnoyarsk)

30 SIB

Remarks: provisionally accepted name, status unconfirmed

species :

Isoetes salishensis D.F.Brunt.; Amer. Fern J. 114(2): 122 (2024)

Distribution: Canada (British Columbia: +Mitlenatch Isl., Lasqueti Isl.)

71 CDN

Remarks: believed extinct but recently rediscovered: Brunton et al. 2025. Currently known from only one small rock pool

species :

Isoetes santacruzensis J.B.S.Pereira; PhytoKeys 131: 61 (2019)

Distribution: Bolivia (Santa Cruz)

83 BOL

species :

Isoetes saracochensis H.P.Fuchs ex Hickey; Fieldiana, Bot. 34: 94 (1994)

Distribution: Peru

83 PER

Synonyms: hidden

= Isoetes saracochensis H.P.Fuchs; (1960), nom. nud.

species :

Isoetes savatieri Franch.; Bull. Soc. Bot. France 31: 395 (1884)

Distribution: Argentina (Chubut, Neuquen, Rio Negro, Tierra del Fuego); Chile (Valparaíso, Maule, Ñuble, Biobío, Araucanía, Los Lagos, Aisén, Magallanes)

85 ARG CLE

Conservation Status: IUCN R

Synonyms: hidden

= Calamaria savatieri (Franch.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes lechleri var. savatieri (Franch.) L.D.Gómez; Brenesia 18: 5 (1980)

= Isoetes chilensis H.P.Fuchs; (1981), nom. nud.

species :

Isoetes schweinfurthii A.Braun; J. Bot. 18: 108 (1880)

Distribution: Senegal; Burkina Faso; Ivory Coast; Benin; Nigeria; Central African Republic; Chad; Sudan; Ethiopia; Tanzania; S-Angola; Zambia; Malawi; Zimbabwe; Mozambique; N-Namibia; S-Botswana; South Africa (Limpopo); Madagascar

22 BEN BKN IVO NGA SEN 23 CAF 24 CHA ETH SUD 25 TAN 26 ANG MLW MOZ ZAM ZIM 27 BOT NAM SAF 29 MDG

Synonyms: hidden

= Calamaria schweinfurthii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes rhodesiana Alston; Bol. Soc. Brot., Ser. 2, 30: 17 (1956)

= Isoetes kersii Wanntorp; Svensk Bot. Tidskr. 64: 146, t. 1, 3 (1970)

species :

Isoetes sehnemii H.P.Fuchs; Fl. Ilustr. Catarin. (Isoetaceas) 38 (1986)

Distribution: Brazil (Rio Grande do Sul)

84 BRA

species :

Isoetes septentrionalis D.F.Brunt.; Canad. Field-Naturalist 129: 176 (2015)

Name: Northern Quillwort

Distribution: E-Canada (SE-Ontario, SW-Québec); USA (Connecticut, Massachusetts, New Hampshire, ?Maine, Pennsylvania, Vermont)

72 CDN 75 USA

Remarks: riparia agg.

species :

Isoetes serracarajensis J.B.S.Pereira, Salino & Stützel; Phytotaxa 272(2): 142 (2016)

Distribution: Brazil (Pará)

84 BRA

species :

Isoetes shangrilaensis Xiang Li, Yuqian Huang, X.Dai & Xing Liu; Phytotaxa 397(1): 71 (2019)

Distribution: China (Yunnan)

36 CHI

species :

Isoetes sinensis T.C.Palmer; Amer. Fern J. 17: 111 (1927)

Distribution: China (Anhui, Guangxi, Jiangsu, Zhejiang); S-Japan

36 CHI 38 JAP

Remarks: allotetraploid originated from I. yunguiensis x taiwanensis

species :

Isoetes smithii H.P.Fuchs; Fl. Ilustr. Catarin. (Isoetaceas) 17 (1986)

Distribution: Brazil (Rio Grande do Sul, Santa Catarina)

84 BRA

species :

Isoetes spannagelii H.P.Fuchs; Fl. Ilustr. Catarin. (Isoetaceas) 33 (1986)

Distribution: Brazil (Santa Catarina)

84 BRA

species :

Isoetes spinulospora C.Jermy & Schelpe; Contr. Bolus Herb. 10: 151 (1982)

Distribution: Bioko Isl. [Fernando Poo]

23 BIO

Conservation Status: IUCN R

species :

Isoetes stellenbossiensis A.V.Duthie; Trans. Roy. Soc. South Africa 17: 328 (1929)

Distribution: South Africa (W-Cape Prov., N-Cape Prov.)

27 SAF

species :

Isoetes stephanseniae A.V.Duthie; Trans. Roy. Soc. South Africa 17: 330 (1929)

Distribution: South Africa (W-Cape Prov.)

27 SAF

Synonyms: hidden

= Isoetes capensis var. stephanseniae (A.V.Duthie) Schelpe & N.C.Anthony; Bothalia 15(3-4): 555 (1985)

species :

Isoetes stevensii J.R.Croft; Blumea 26: 177-190 (1980)

Distribution: New Guinea (Papua New Guinea)

42 NWG

Conservation Status: IUCN R

species :

Isoetes storkii T.C.Palmer; Amer. Fern J. 22(4): 136 (1932)

Distribution: Costa Rica; Panama

80 COS PAN

Conservation Status: IUCN R

Synonyms: hidden

= Isoetes montana T.C.Palmer; Amer. Fern J. 21(4): 134 (1931) (nom. illeg., non Isoetes montana U. Weber (1922))

= Isoetes tryoniana Gómez Pompa; Revista Biol. Trop. 17: 108 (1970)

species :

Isoetes taiwanensis De Vol; Taiwania 17(1): 1-7, f. 1 (1972)

Distribution: Taiwan

38 TAI

Conservation Status: IUCN E (extremely rare, only in one single pond)

variant :

var. kinmenensis F.Y.Lu, H.H.Chen & Y.L.Hsueh; Fl. Kinmen 26 (2011)

Distribution: Taiwan

38 TAI

Conservation Status: IUCN E (extremely rare)

species :

Isoetes tamaulipana Mora-Olivo, A.Mend. & Mart.-Aval.; Phytotaxa 267(2): 115 (2016)

Distribution: Mexico (Tamaulipas)

79 MEX

species :

Isoetes tegetiformans Rury; Amer. Fern J. 68(4): 100 (1978)

Name: Mat-forming Merlin´s-grass (EN)

Distribution: USA (N-Georgia)

78 USA

Conservation Status: IUCN E. Extremely rare, known from only a few vernal pools on granite outcrops

species :

Isoetes tennesseensis Luebke & Budke; Amer. Fern J. 93(4): 184-190 (2003)

Name: Tennessee Quillwort

Distribution: USA (Tennessee)

78 USA

species :

Isoetes tenuifolia Jermy; Bot. J. Linn. Soc. 64: 118 (1971)

Distribution: Ghana; ?Kenya

22 GHA

Conservation Status: IUCN R (Ghana), E (Kenya)

species :

Isoetes texana Singhurst, Rushing & W.C.Holmes; Phytoneuron 22: 1 (2011)

Distribution: USA (Texas)

77 USA

species :

Isoetes tiguliana Gennari; Comment. Soc. Crittog. Ital. 1(1): 42 (1861)

Distribution: Italy; Morocco; Algeria; Tunisia

13 ITA 20 ALG MOR TUN

Remarks: bona sp. fide Troia et al. 2016

Synonyms: hidden

= Calamaria dubia (Gennari) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Calamaria tegulensis (Genn.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes dubia Gennari; Comment. Soc. Crittog. Ital. 1: 104 (1861)

= Isoetes tegulensis f. submersa Glück; Süsswasserflora 15: 20 (1936)

= Isoetes tegulensis f. terrestris Glück; Süsswasserflora 15: 20 (1936)

= Isoetes tegulensis Gennari; Comment. Soc. Crittog. Ital. 1: 106 (1861) (sept.) (nom. illeg.)

= Isoetes velata subsp. dubia (Gennari) Batt. & Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes velata subsp. tegulensis (Gennari) Batt. & Trab.; Fl. Algérie Tunisie 407 (1905)

= Isoetes velata var. tegulensis (Gennari) Fiori & Paol.; Fl. Anal. Italia 1: 25 (1896)

= Isoetes velata f. maculosa Gennari; Ann. Agric. Siciliana 11: 175 (1865)

= Isoetes velata f. emersa Gennari; Giorn. Bot. Ital. 2: 96 (1870)

= Isoetes velata var. tegulensis Fiori; Fl. Anal. Italia 1: 25 (1896)

species :

Isoetes todaroana Troìa & Raimondo; Amer. Fern J. 99(4): 238 (2009 publ. 2010)

Distribution: S-Italy; Sicily

13 ITA SIC

Remarks: probably to be merged into I. histrix fide Strid (2024)

Synonyms: hidden

= Isoetes iapygia Ernandes, Beccarisi & V.Zuccarello; Pl. Biosystems 144(4): 807 (2010) (fide Troia & Greuter 2015)

species :

Isoetes toximontana Musselman & J.P.Roux; Novon 12: 504-507 (2002)

Distribution: South Africa (N-Cape Prov., W-Cape: Gifberg to Kamiesberg)

27 SAF

species :

Isoetes transvaalensis C.Jermy & Schelpe; Contr. Bolus Herb. 10: 150 (1982)

Distribution: South Africa (Limpopo, Gauteng, Mpulamanga, E-Cape Prov., Free State, KwaZulu-Natal); Lesotho

27 LES SAF

species :

Isoetes triangula U.Weber; Hedwigia 63: 253, f. 32, 33 (1922)

Distribution: Mexico (Oaxaca); Venezuela (Amazonas, Bolívar, Cojedes, Guarico); Guyana; French Guiana; Brazil (Acre, Amazonas, Roraima)

79 MEX 82 FRG GUY VEN 84 BRA

Synonyms: hidden

= Isoetes schinzii H.P.Fuchs; (1960), nom. nud.

species :

Isoetes tripus A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1868: 559 (1868)

Distribution: Australia (SW-Western Australia)

50 AUS

Synonyms: hidden

= Calamaria tripus (A.Braun) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes phaeospora Durieu; Bull. Soc. Bot. France 11: 103 (1864)

species :

Isoetes tuckermanii A.Braun ex Engelm.; Manual [A.Gray], ed. 5: 676 (1867)

subspecies :

subsp. tuckermanii

Name: Tuckermans Quillwort (EN)

Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario, Québec); USA (Connecticut, Delaware, Massachusetts, Maryland, Maine, New Hampshire, New Jersey, New York, Rhode Island, Vermont)

72 CDN 75 USA 78 USA

Remarks: inclusion of I. acadiensis fide Troia et al. 2016

Synonyms: hidden

= Calamaria tuckermanii (A.Braun ex Engelm.) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Isoetes tuckermanii var. borealis A.A.Eaton; Fern Bull. 6: 7 (1898)

subspecies :

subsp. acadiensis (Kott) D.F.Brunt.; Canad. Field-Naturalist 132(4): 366 (2018)

Distribution: E-Canada (Nova Scotia)

72 CDN

Synonyms: hidden

= Isoetes acadiensis Kott; Canad. J. Bot. 59: 2592 (1981)

species :

Isoetes tuerckheimii Brause; Symb. Antill. [Urban] 7: 161 (1912)

Distribution: Hispaniola (Dominican Republic)

81 HIS

species :

Isoetes udupiensis P.K.Shukla, G.K.Srivast., S.K.Shukla & P.K.Rajagopal; Taxon 54(1): 112 (2005)

Distribution: India (Karnataka)

40 IND

Remarks: bona sp. fide Fraser-Jenkins et al. 2017

species :

Isoetes ulei U.Weber; Hedwigia 63: 255, fig. 40-41 (1922)

Distribution: Brazil (Rio de Janeiro)

84 BRA

Synonyms: hidden

= Isoetes goebelii U.Weber; Hedwigia 63: 247, fig. 16-17 (1922)

species :

Isoetes valida (Engelm.) Clute; Fern Allies 236 (1905)

Name: Strong quillwort

Distribution: USA (Alabama, Delaware, Georgia, North Carolina, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)

75 USA 78 USA

Synonyms: hidden

= Isoetes engelmannii var. valida Engelm.; Manual [A.Gray], ed. 5: 677 (1867)

species :

Isoetes vanensis M.Keskin & G.Zare; Phytotaxa 269(4): 295 (2016)

Distribution: Turkey (E-Anatolia)

34 TUR

species :

Isoetes vermiculata Hickey; Ann. Missouri Bot. Gard. 76(4): 1160 (1989)

Distribution: Venezuela (Amazonas)

82 VEN

species :

Isoetes virginica N.Pfeiff.; Claytonia 3: 29 (1937)

Name: Piedmont quillwort (EN)

Distribution: SE-USA (Alabama, Georgia, North Carolina, South Carolina, Virginia)

78 USA

Conservation Status: IUCN R

Synonyms: hidden

= Isoetes piedmontana (N.Pfeiff.) C.F.Reed; Phytologia 12: 392 (1965)

= Isoetes virginica var. piedmontana N.Pfeiff.; Bull. Torrey Bot. Club 66: 411 (1939)

species :

Isoetes viridimontana M.A.Rosenthal & W.C.Taylor; Amer. Fern J. 104(1): 8 (2014)

Distribution: USA (Vermont)

75 USA

species :

Isoetes weberi Herter ex U.Weber; Hedwigia 63: 255-256, fig. 44-45 (1922)

Distribution: Brazil (Rio Grande do Sul); Uruguay (Rocha)

84 BRA 85 URU

Synonyms: hidden

= Isoetes weberi Herter; Beih. Bot. Centralbl. 39(2): 254 (1922), nomen

species :

Isoetes welwitschii A.Braun ex Kuhn; Filic. Afr. [Kuhn] 196 (1868)

Distribution: Ghana; Nigeria; Central African Republic; Sudan; Ethiopia; Eritrea; Kenya; Tanzania; Angola; Zambia; Botswana; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal); Madagascar

22 GHA NGA 23 CAF 24 ERI ETH SUD 25 KEN TAN 26 ANG ZAM 27 BOT SAF 29 MDG

Synonyms: hidden

= Isoetes natalensis Baker; Handb. Fern-Allies 132 (1887)

= Calamaria natalensis (Baker) Kuntze; Revis. Gen. Pl. 2: 828 (1891)

= Calamaria welwitschii (A.Braun ex Kuhn) Kunze; Revis. Gen. Pl. 2: 828 (1883)

species :

Isoetes wormaldii R.Sim; Trans. S. African Philos. Soc. 16: 299, t. 5 (1906)

Distribution: South Africa (E-Cape: Grahamstown to East London)

27 SAF

Conservation Status: IUCN E

species :

Isoetes xiangfei Y.H.Yan, Y.F.Gu & J.P.Shu; Guihaia 42(10): 1626 (2022)

Distribution: China (Hunan)

36 CHI

species :

Isoetes yuhangensis Yu C.Chen & Xing Liu; Phytotaxa 642(1): 42 (2024)

Distribution: China (Zhejiang)

36 CHI

species :

Isoetes yunguiensis C.F.Wang & W.C.Taylor; Novon 12(4): 587-591 (2002)

Distribution: China (Guizhou, Yunnan)

36 CHI

species :

Isoetes ×altonharvillii Musselman & R.D.Bray; Castanea 60(3): 246 (1995)

Distribution: USA (Delaware, Virginia)

78 USA

Remarks: Isoetes caroliniana x engelmannii

species :

Isoetes ×angeloi D.F.Brunt. & K.L.McIntosh; Bot. Lett. 170(2): 281 (2022)

Distribution: Sicily

13 SIC

Remarks: Isoetes gymnocarpa x longissima

species :

Isoetes ×blondeaui D.F.Brunt. & P.C.Sokoloff; Botany (Ottawa) 98(10): 558 (2020)

Distribution: Canada (Quebec)

72 CDN

Remarks: Isoetes echinospora x laurentiana

species :

Isoetes ×brittonii D.F.Brunt. & W.C.Taylor; Amer. Fern J. 80: 82-89 (1990)

Distribution: USA (Connecticut, New Jersey, New York, Pennsylvania, South Carolina, Virginia)

75 USA 78 USA

Remarks: Isoetes engelmanii x riparia

species :

Isoetes ×brochonii Motelay; Actes Soc. Linn. Bordeaux 45: 45 (1892)

Distribution: Andorra; SW-France

12 ADO FRA

Remarks: Isoetes echinospora x creussensis

Synonyms: hidden

= Isoetes lacustris subsp. brochonii (Motelay) R.C.V.Douin; Fl. Ill. France 12: 117 (1934)

species :

Isoetes ×bruntonii Knepper & Musselman; Amer. Fern J. 86(1): 9 (1996)

Distribution: USA (Virginia)

78 USA

Remarks: Isoetes engelmannii x hyemalis

species :

Isoetes ×carltaylori Musselman., R.D.Bray & D.A.Knepper; Canad. J. Bot. 75(2): 301-309 (1997)

Distribution: USA (E-USA)

78 USA

Remarks: Isoetes acadiensis x engelmannii

species :

Isoetes ×dodgei A.A.Eaton; Fern Bull. 6: 6 (1898)

Distribution: Canada (Ontario, Québec); USA (New Hampshire, New Jersey, New York, Pennsylvania, Vermont)

72 CDN 75 USA

Remarks: Isoetes echinospora x septentrionalis

Synonyms: hidden

= Isoetes canadensis A.A.Eaton; Proc. U.S. Natl. Mus. 23 (1226): 650 (1901)

= Isoetes riparia f. canadensis (Engelm.) Proctor; Amer. Fern J. 39: 119 (1949)

= Isoetes riparia var. canadensis Engelm.; Trans. Acad. Sci. St. Louis 4: 383 (1882)

species :

Isoetes ×eatonii R.Dodge; Bot. Gaz. 23: 16, t. 4-5 (1897), pro sp.

Distribution: Canada (Ontario); USA (Connecticut, Massachusetts, New Hampshire, New Jersey, New York, Pennsylvania, Vermont)

72 CDN 75 USA

Remarks: Isoetes echinospora x engelmannii

Synonyms: hidden

= Isoetes eatoni R.Dodge; Ferns & Fern Allies New England 39 (1896)

= Isoetes ×gravesii A.A.Eaton; Fernwort Pap. 2: 14 (1900)

= Isoetes eatoni f. gravesii (A.A.Eaton) C.F.Reed; Amer. Fern J. 35: 82 (1945)

= Isoetes eatoni var. gravesii (A.A.Eaton) Clute; Fern Bull. 16: 55 (1908)

= Isoetes valida var. gravesii (A.A.Eaton) Clute; Fern Allies 243 (1905)

species :

Isoetes ×echtuckeri D.F.Brunt. & D.M.Britton; Canad. J. Bot. 77(11): 1662-1668 (1999)

Distribution: Canada (New Brunswick, Nova Scotia, Ontario, Québec); USA (Maine)

72 CDN 75 USA

Remarks: Isoetes tenella x tuckermanii

species :

Isoetes ×fernaldii D.F.Brunt. & P.C.Sokoloff; Webbia 75(1): 44 (2020)

Distribution: USA (Virginia)

78 USA

Remarks: Isoetes engelmannii x hyemalis

species :

Isoetes ×foveolata A.A.Eaton ex R.Dodge; Ferns & Fern Allies New England 38 (1896)

Distribution: Canada (Ontario); USA (Connecticut, Massachusetts, New Hampshire)

72 CDN 75 USA

Remarks: Isoetes engelmannii x tuckermanii

species :

Isoetes ×gopalkrishnae Sarv.K.Singh, P.K.Shukla & N.K.Dubey; Phytotaxa 362(1): 71 (2018)

Distribution: India (Uttar Pradesh)

40 IND

Remarks: Isoetes coromandelina x sampathkumarnii

species :

Isoetes ×harveyi A.A.Eaton; Fernwort Pap. 11 (1900)

Distribution: Canada (New Brunswick, Newfoundland, Nova Scotia, Ontario); USA (Maine)

72 CDN 75 USA

Remarks: Isoetes lacustris x tuckermanii

Synonyms: hidden

= Isoetes tuckermanii var. harveyi (A.A.Eaton) Clute; Fern Allies 260 (1905)

species :

Isoetes ×herb-wagneri W.C.Taylor; Amer. Fern J. 92(2): 162 (2002)

Distribution: USA (Montana)

73 USA

Remarks: Isoetes echinospora x bolanderi

species :

Isoetes ×heterospora A.A.Eaton; Fernwort Pap. 2: 8 (1900), pro sp.

Distribution: USA (New York)

75 USA

Remarks: Isoetes lacustris x tuckermannii

Synonyms: hidden

= Isoetes tuckermani var. heterospora (A.A.Eaton) Clute; Fern Allies 226, 260 (1905)

= Isoetes macrospora var. heterospora A.A.Eaton; Rhodora 10: 42 (1908)

= Isoetes macrospora f. heterospora (A.A.Eaton) Clute; Fern Bull. 16: 56 (1908)

species :

Isoetes ×hickeyi W.C.Taylor & Luebke; Amer. Fern J. 78(1): 6 (1988)

Distribution: England (Scotland); Canada (Ontario, Saskatchewan); USA (Wisconsin)

10 GRB 71 CDN 72 CDN 74 USA

Remarks: Isoetes echinospora x lacustris

species :

Isoetes ×jeffreyi D.M.Britton & D.F.Brunt.; Canad. J. Bot. 70: 447-452 (1992)

Distribution: Canada (Ontario, Québec)

72 CDN

Remarks: Isoetes lacustris x riparia

species :

Isoetes ×jermyi D.F.Brunt. & Rumsey; Bot. Lett. 168(4): 507 (2021)

Distribution: Great Britain; France

10 GRB 12 FRA

Remarks: Isoetes echinospora x lacustris

species :

Isoetes ×kareniae D.F.Brunt. & P.C.Sokoloff; Webbia 75(1): 38 (2020)

Distribution: USA (Pennsylvania, Virginia)

75 USA 78 USA

Remarks: Isoetes appalachiana x engelmannii

species :

Isoetes ×marensis D.M.Britton & D.F.Brunt.; Canad. J. Bot. 73(9): 1352 (1995)

Distribution: Canada (British Columbia)

71 CDN

Remarks: Isoetes howellii x maritima

species :

Isoetes ×michinokuana M.Takamiya, M.Watanaba & K.Ono; Acta Phytotax. Geobot. 48(2): 109 (1997 publ. 1998)

Distribution: Japan

38 JAP

Remarks: Isoetes japonica x pseudojaponica

species :

Isoetes ×novae-angliae D.F.Brunt. & D.M.Britton; Rhodora 108(935): 238 (2006)

Distribution: USA (Massachusetts)

75 USA

Remarks: Isoetes septentrionalis x tuckermannii

Synonyms: hidden

= Isoetes canadensis var. robbinsii A.A.Eaton; Rhodora 5: 279 (1903)

= Isoetes chapmanii var. robbinsii (A.A.Eaton) M.Broun; Index No. Amer. Ferns: 199 (1938)

= Isoetes dodgei var. robbinsii A.A.Eaton; Rhodora 10: 42 (1908)

= Isoetes riparia var. robbinsii (A.A.Eaton) Proctor; Amer. Fern J. 39: 119 (1949)

species :

Isoetes ×pseudotruncata D.M.Britton & D.F.Brunt.; Canad. J. Bot. 74(1): 58 (1996)

Distribution: Russian Far East (Kamchatka); Alaska; Canada (Alberta, British Columbia)

31 RFE 70 ASK 71 CDN

Remarks: Isoetes echinospora x maritima

Synonyms: hidden

= Isoetes ×paratunica D.F.Brunt., Mochalova & A.A.Bobrov; Amer. Fern J. 105(2): 109, figs. 5-7 (2015)

species :

Isoetes ×robusta (Engelm.) D.F.Brunt.; Rhodora 120(984): 307 (2019)

Distribution: Canada; USA

Error

Synonyms: hidden

= Isoetes braunii f. robusta (Engelm.) N.Pfeiff.; Ann. Missouri Bot. Gard. 9: 173 (1922)

= Isoetes echinospora f. robusta (Engelm.) Clute; Fern Allies 258 (1905)

= Isoetes echinospora var. robusta Engelm.; Trans. Acad. Sci. St. Louis 4: 380 (1882)

= Isoetes muricata f. robusta (Engelm.) C.F.Reed; Amer. Fern J. 35: 82 (1945)

= Isoetes tenella f. robusta (Engelm.) Rothm.; Feddes Repert. Spec. Nov. Regni Veg. 54: 73 (1944)

species :

Isoetes ×truncata (A.A.Eaton) Clute; Fern Allies 260 (1905)

Name: Vancouver Quillwort (EN)

Distribution: SE-Alaska; Canada (Alberta)

70 ASK 71 CDN

Remarks: Isoetes maritima x occidentalis

Synonyms: hidden

= Isoetes echinospora var. truncata A.A.Eaton; Working List N. Amer. Pterid. 27 (1901)

= Isoetes muricata subsp. truncata (A.A.Eaton) H.P.Fuchs; Proc. Kon. Ned. Akad. Wetensch. C 85(2): 239 (1982)

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A complete, synonymic checklist of the Higher Plants of the World

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A complete, synonymic checklist of the Higher Plants of the World

Version 25.10 (Oct. 12th, 2025)

Covering 357,450 accepted species,
8,806 hybrids, 48,075 accepted infraspecific taxa and 1,004,944 synonyms,
**in total 1,419,285* names**

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Contact

How to Cite:

Realization

Search

+ Family

Filter

Order

Family (1)

Genus (1)

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A complete, synonymic checklist of the Higher Plants of the World

Version 25.10 (Oct. 12th, 2025)

Covering 357,450 accepted species,8,806 hybrids, 48,075 accepted infraspecific taxa and 1,004,944 synonyms,in total 1,419,285* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life

Covering 357,450 accepted species,
8,806 hybrids, 48,075 accepted infraspecific taxa and 1,004,944 synonyms,
**in total 1,419,285* names**

[about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus*" are deliberately omitted]