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Ver. 19.2
April 17th, 2024

016.0000 Family Marsileaceae Mirb.; Hist. Nat. Veg. (Lam. & Mirb.) 5: 126 (1802)
3 genera, ca. 55 spp.
Species listed in database: 57; hybrids: 1
= Marsileeae Duby; Bot. Gall. 1: 542 (1828)
= Marsileoideae Eaton; Man. Bot., ed. 6: 81 (1833)
= Pilulariaceae Bercht. & J.Presl; Prir. Rostlin 1: 272 (1820)
= Pilulariaceae Mirb. ex DC.; Essai Propr. Méd. Pl. 48 (1804) (Pilulariae)
016.0001 Genus Marsilea L.; Sp. Pl. 2: 1097 (1753)
Distribution: Worldwide, mostly tropics and subtropics, many in Africa
Species listed in database: 49; hybrids: 1
= Lemma B.Juss.; Juss., Gen. Pl. lxiii (1789)
= Spheroidia Dulac; Fl. Hautes-Pyrénées: 39 (1867)
= Zaluzianskia Neck.; Hist. & Commentat. Acad. Elect. Sci. Theod.-Palat. 3: 303 (1775), nom. inval.
= Zalusianskya Kuntze; Revis. Gen. Pl. 2: 822 (1891), nom. illeg., non Zaluzianskya F. W. Schmidt 1793 (Scrophulariaceae)
species :
Marsilea aegyptiaca Willd.; Sp. Pl. 5(1): 540 (1810)]
Distribution: Greece (S-Peloponnisos); European Russia (E-European Russia); Algeria; Tunisia; Libya; Egypt; Mauritania; Mali; Sudan; Ethiopia; Namibia; Botswana; South Africa (Limpopo, Gauteng, North West, N-Cape Prov.); Madagascar; Kazakhstan; Turkmenistan; Sinai peninsula; Saudi Arabia; N-Yemen; China (N-Xinjiang); India (Bihar, Gujarat, Madhya Pradesh, Odisha, Rajasthan, Uttar Pradesh)
13 GRC 14 RUS 20 ALG EGY LBY TUN 22 MLI MTN 24 ETH SUD 27 BOT NAM SAF 29 MDG 32 KAZ TKM 34 SIN 35 SAU YEM 36 CHI 40 IND
Synonyms: hidden
= Marsilea ballardii K.M.Gupta; J. Bombay Nat. Hist. Soc. 53: 291, t. 1-3 (1956) (fide Fraser-Jenkins 2008)
= Marsilea ballardii var. rajasthanensis (K.M.Gupta) S.M.Almeida & M.R.Almeida; J. Bombay Nat. Hist. Soc. 82(2): 445 (1985) (fide Fraser-Jenkins 2008)
= Marsilea condensata Baker; J. Bot. 24: 281 (1886)
= Marsilea rajasthanensis K.M.Gupta; Marsilea Council Sci. Indus. Res. New Delhi Bot. Monogr. no. 2: 29 (1962) (fide Fraser-Jenkins 2008)
= Marsilea rajasthanensis var. ballardii (K.M.Gupta) K.M.Gupta; Marsilea Bot. Monogr. Council Sci. Industr. Res. India 2: 29 (1962), comb. inval.
= Zaluzianskia aegyptiaca (Willd.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia condensata (Baker) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea aethiopica Launert; Garcia de Orta, Ser. Bot. 6(1-2): 128 (1984)
Distribution: Egypt; Ethiopia; Eritrea
20 EGY 24 ERI ETH
species :
Marsilea ancylopoda A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 434 (1864)
Distribution: +USA (+Florida); Mexico (Aguascalientes, Chihuahua, Colima, Durango, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Nayarit, Oaxaca, Queretaro, Tamaulipas); Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Bonaire; Colombia (Chocó, Cundinamarca, Tolima); Venezuela (Carabobo, Venezuela Distrito Federal, Falcon, Lara, Portuguesa, Sucre, Zulia); Ecuador; Peru; Brazil (Bahia, Paraná, Rio Grande do Sul); Paraguay (Alto Paraguay, Central); Uruguay (Artigas, Canelones, Cerro Largo, Colonia, Maldonado, Montevideo, Paysandu, Rocha, San Jose, Soriano, Tacuarembo); Argentina (Buenos Aires, Chaco, Chubut, Cordoba, Corrientes, Entre Rios, Formosa, Jujuy, La Pampa, Mendoza, Misiones, Rio Negro, Salta, Santa Fe, San Luis)
78 USA 79 MEX 80 COS ELS GUA HON NIC PAN 81 BON HIS JAM PUE 82 CLM VEN 83 ECU PER 84 BRA 85 ARG PAR URU
Synonyms: hidden
= Marsilea berteroi A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 747 (1870)
= Marsilea concinna Baker; J. Bot. 24: 279 (1886)
= Marsilea ernestii A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 746 (1870)
= Marsilea hickenii Herter; Anales Mus. Nac. Montevideo II, 1: 376, t. 29 (1925)
= Marsilea mexicana A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 747 (1870)
= Marsilea striata Ernst; Vargasia (Bol. Soc. Cienc. Caracas) no. 7: 181 (1868)
= Zaluzianskia ancylopoda (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia berteroi (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia concinna (Baker) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia ernestii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia mexicana (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea angustifolia R.Br.; Prodr. Fl. Nov. Holland. 167 (1810)
Distribution: Australia (N-Western Australia, N-Northern Territory)
50 AUS
Synonyms: hidden
= Zaluzianskia angustifolia (R.Br.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea apposita Launert; Senckenberg. Biol. 49: 306, f. 75 (1968)
Distribution: W-Zimbabwe; E-Botswana; South Africa (Mpulamanga, KwaZulu-Natal); Eswatini [Swaziland]
26 ZIM 27 BOT SAF SWZ
species :
Marsilea batardae Launert; Bol. Soc. Broter. Ser. 2, 56: 101 (1983)
Distribution: Portugal; Spain
12 POR SPA
species :
Marsilea berhautii Tardieu; Notul. Syst. (Paris) 16: 85 (1955)
Distribution: Senegal; Mali; Burkina Faso; Ghana; Niger; Nigeria
22 BKN GHA MLI NGA NGR SEN
species :
Marsilea botryocarpa F.Ballard; Hooker´s Icon. Pl. 36, t. 3600 (1962)
Distribution: Kenya
25 KEN
Conservation Status: IUCN V
species :
Marsilea burchellii (Kunze) A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 429 (1864)
Distribution: Namibia; Botswana; South Africa (Limpopo, North West, Free State, W-Cape Prov., E-Cape Prov., N-Cape Prov.)
27 BOT NAM SAF
Synonyms: hidden
= Marsilea biloba Willd.; Sp. Pl. ed. 4. 5: 540 (1810)
= Marsilea glomerata C.Presl; Abh. Königl. Böhm. Ges. Wiss., ser. 5, 3: 580 (1845)
= Marsilea macrocarpa var. biloba (Willd.) Sim; Ferns S. Afr. ed. 2: 315, t. 172, fig. 3 (1915)
= Marsilea macrocarpa var. burchellii (Kunze) Sim; Ferns S. Afr. ed. 2: 315, t. 172, fig. 2 (1915) [as (A. Braun) Sim]
= Marsilea pumila Drège; Zwei Pflanzengeogr. Dokum.: 58 (1843), nom. nud.
= Marsilea quadrifolia var. burchellii Kunze; Linnaea 10: 556 (1836)
= Zaluzianskia biloba (Willd.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia burchellii (Kunze) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea capensis A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 428 (1864)
Distribution: Egypt [I]; Zambia; Namibia; South Africa (E-Cape Prov., W-Cape Prov., N-Cape Prov., Free State, North West, Gauteng, Limpopo, Mpulamanga, KwaZulu-Natal); S-Iraq [I]
20 egy 26 ZAM 27 NAM SAF
Synonyms: hidden
= Zaluzianskia capensis (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Marsilea macrocarpa var. capensis (A.Braun) Sim; The ferns of South Africa, edn. 2: 316 (1915)
species :
Marsilea coromandelina Willd.; Sp. Pl. ed. 4, 5: 539 (1810)
Distribution: Morocco; Cape Verde Isl. (Boavista); Mauritania; Senegal; Gambia; Guinea-Bissau; Burkina Faso; Nigeria; Burundi; Chad; Sudan; Eritrea; Soqotra; Kenya; Tanzania; Angola; Zimbabwe; Mozambique; Namibia; Botswana; South Africa (Limpopo); Madagascar; Iraq [I]; India (Andhra Pradesh, Karnataka, Kerala, Maharashtra, Rajasthan, Tamil Nadu, Uttar Pradesh); Sri Lanka
20 MOR 21 CVI 22 BKN GAM GNB MTN NGA SEN 23 BUR 24 CHA ERI SOC SUD 25 KEN TAN 26 ANG MOZ ZIM 27 BOT NAM SAF 29 MDG 34 irq 40 IND SRL
Synonyms: hidden
= Marsilea microphylla C.Chr.; Dansk Bot. Ark. 7: 180, t. 73, f. 8-9 (1932)
= Marsilea minuta var. coromandelina L.; Mant. Pl. 308 (1771)
= Marsilea muscoides Lepr. ex A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 422 (1864)
= Marsilea muscosa Lepr. ex A.Braun; Flora 300 (1839), nomen
= Marsilea pygmaea Brongn.; Dict. Class d´Hist. Nat. 10: 199 (1826) (pygmea)
= Marsilea trichocarpa Bremek.; Ann. Transvaal Mus. 15: 234 (1933)
= Marsilea trichopoda Lepr. ex A.Braun; Flora 300 (1839) (nomen), et Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 422 (1864)
= Marsilea trichopus Sadeb.; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 420 (1900)
= Zaluzianskia coromandelina (Willd.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia muscoides (Lepr. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia trichopoda (Lepr. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea costulifera D.L.Jones; Muelleria 8(1): 65 (1993)
Distribution: Australia (Australian Capital Territory, New South Wales, Victoria, South Australia, Queensland, S-Northern Territory)
50 AUS
species :
Marsilea crenata C.Presl; Reliq. Haenk. 1: 84, t. 12, f. 13 (1825)
Distribution: ?S-China; Thailand; Laos; Vietnam; peninsular Malaysia; Borneo (Sabah); Philippines; Java; Lesser Sunda Isl.; New Guinea (Papua New Guinea); Australia
41 LAO THA VIE 42 BOR JAW LSI MLY NWG PHI 50 AUS
Remarks: bona sp. fide Fraser-Jenkins et al. (2017), not a synonym of M. minuta
Synonyms: hidden
= Marsilea elata var. crenata (C.Presl) Sadeb.; Engl. & Prantl, Nat. Pflanzenfam. 1(4): 419 (1900)
species :
Marsilea crotophora D.M.Johnson; Syst. Bot. Monogr. 11: 46 (1986)
Distribution: Mexico (Campeche, Guerrero, Tabasco, Veracruz); Nicaragua; Venezuela (Barinas); N-Bolivia (Beni, Santa Cruz); Brazil (Amazonas, Mato Grosso do Sul, Mato Grosso)
79 MEX 80 NIC 82 VEN 83 BOL 84 BRA
species :
Marsilea cryptocarpa Albr. & Chinnock; J. Adelaide Bot. Gard. 22: 1 (2008)
Distribution: Australia (Northern Territory, South Australia)
50 AUS
species :
Marsilea deflexa A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 421 (1864)
Distribution: Mexico (Nayarit, Jalisco, Oaxaca, Puebla, Veracruz); Guatemala; Honduras; Nicaragua; Costa Rica; Colombia (Cundinamarca, Tolima); Venezuela (Amazonas, Anzoategui, Apure, Aragua, Barinas, Bolívar, Carabobo, Cojedes, Falcon, Guarico, Portuguesa, Trujillo, Zulia); Guyana; Ecuador; Peru; Brazil (Amapá, Goiás, Piauí, Roraima); Paraguay (Alto Paraguay, Concepcion, Itapua); Argentina (Corrientes)
79 MEX 80 COS GUA HON NIC 82 CLM GUY VEN 83 ECU PER 84 BRA 85 ARG PAR
Synonyms: hidden
= Marsilea striata Mett.; Ann. Sci. Nat., Bot., sér. V, 3: 310 (1865)
= Zaluzianskia deflexa (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea distorta A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 433 (1864)
Distribution: Mauritania; Senegal; Liberia; Mali; Burkina Faso; Sudan; Tanzania; Zimbabwe; Mozambique; N-Namibia; N-Botswana
22 BKN LBR MLI MTN SEN 24 SUD 25 TAN 26 MOZ ZIM 27 BOT NAM
Synonyms: hidden
= Zaluzianskia distorta (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea drummondii A.Braun; Linnaea 25. 721 (1852)
Name: Common Nardoo
Distribution: Australia (Australian Capital Territory, W-Western Australia, Northern Territory, South Australia, Queensland, Victoria, New South Wales)
50 AUS
Synonyms: hidden
= Marsilea drummondii f. macra (A.Braun) Domin; Biblioth. Bot. 20: 217 (1915)
= Marsilea drummondii var. elata (A.Braun) A.Braun; Index Seminum (B, Berolinensis) 1867: 3 (1867)
= Marsilea drummondii var. hirsutissima (A.Braun) Domin; Biblioth. Bot. 20: 216 (1915)
= Marsilea drummondii var. howittiana (A.Braun) Domin; Biblioth. Bot. 20: 216 (1915)
= Marsilea drummondii var. macra A.Braun; Index Seminum (B, Berolinensis) 1867: 3 (1867)
= Marsilea drummondii var. megalophylla A.Braun; Linnaea 25: 721 (1853)
= Marsilea drummondii var. minor A.Braun; Linnaea 25: 721 (1853)
= Marsilea drummondii var. muelleri (A.Braun) Baker; J. Bot. 24: 282 (1886)
= Marsilea drummondii var. nardu (A.Braun) Domin; Biblioth. Bot. 20: 217 (1915)
= Marsilea drummondii var. orientalis A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 662 (1870)
= Marsilea drummondii var. oxaloides (A.Braun) Domin; Biblioth. Bot. 20: 217 (1915)
= Marsilea drummondii var. salvatrix (Hanst.) A.Braun; Index Seminum (B, Berolinensis) 1867: 3 (1867)
= Marsilea drummondii var. sericea (A.Braun) Domin; Biblioth. Bot. 20: 217 (1915)
= Marsilea elata (A.Braun) A.Braun; App. Ind. Sem. Hort. Berol. 3 (1867)
= Marsilea hirsutissima A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 736 (1870)
= Marsilea howittiana A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 734 (1870)
= Marsilea macra (A.Braun) A.Braun; App. Ind. Sem. Berol. 3 (1867), et Monatsber. Königl. Akad. Berl. 1870: 735 (1870), et Annales des Sciences Naturelles, Botanique ser. 5, 9 (1868)
= Marsilea macropus Hook.; Hooker´s Icon. Pl. 6: t. 909 (1854) (non Engelm. (1848))
= Marsilea muelleri A.Braun; Linnaea 25. 721 (1852)
= Marsilea nardu A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 736 (1870)
= Marsilea nardu var. megalophylla (A.Braun) A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 737 (1870)
= Marsilea oxaloides A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 736 (1870)
= Marsilea salvatrix Hanst.; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1862: 105 (1862)
= Marsilea sericea A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 734 (1870)
= Zaluzianskia drummondii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia macropus (Hook.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea ephippiocarpa Alston; J. Bot. 118, fig. (1930)
Distribution: Angola; Zambia; Zimbabwe; Mozambique; Namibia; Botswana; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, Cape Prov.); Eswatini [Swaziland]
26 ANG MOZ ZAM ZIM 27 BOT NAM SAF SWZ
species :
Marsilea exarata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 732 (1870)
Distribution: Australia (Western Australia, Northern Territory, South Australia, Queensland, New South Wales)
50 AUS
Synonyms: hidden
= Marsilea hirsuta var. exarata (A.Braun) Domin; Biblioth. Bot. 20: 216 (1914)
= Marsilea paradoxa Diels; Repert. Spec. Nov. Regni Veg. 8: 86 (1906)
species :
Marsilea fadeniana Launert; Garcia de Orta, Ser. Bot. 6(1-2): 126 (1984), nom. nov.
Distribution: Kenya
25 KEN
Conservation Status: IUCN E
Synonyms: hidden
= Marsilea glomerata Launert; Senckenberg. Biol. 52: 449 (1971)
species :
Marsilea farinosa Launert; Senckenberg. Biol. 49: 298, f. 41-46 (1968)
subspecies :
subsp. farinosa
Distribution: Ethiopia; Kenya; Tanzania; Angola; Zimbabwe; Mozambique; Namibia; Botswana; South Africa (North West, Limpopo, Gauteng, KwaZulu-Natal)
24 ETH 25 KEN TAN 26 ANG MOZ ZIM 27 BOT NAM SAF
subspecies :
subsp. arrecta J.E.Burrows; Bothalia 11(2): 169 (1989)
Distribution: SE-Botswana; South Africa (Limpopo)
27 BOT SAF
species :
Marsilea fenestrata Launert; Mitt. Bot. Staatssamml. München 3: 507, f. 2 (1960)
Distribution: S-Mozambique; South Africa (KwaZulu-Natal); Eswatini [Swaziland]
26 MOZ 27 SAF SWZ
species :
Marsilea gibba A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 745 (1870)
Distribution: Mali; Mali; Burkina Faso; Congo [Brazzaville]; Central African Republic; D.R.Congo [Zaire]; Rwanda; Chad; Sudan; Eritrea; Uganda; Kenya; Tanzania; Angola
22 BKN MLI 23 CAF CON RWA ZAI 24 CHA ERI SUD 25 KEN TAN UGA 26 ANG
Synonyms: hidden
= Zaluzianskia gibba (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea globulosa Bouchart; Adansonia 41(1): 8 (2019)
Distribution: Madagascar
29 MDG
species :
Marsilea hirsuta R.Br.; Prodr. Fl. Nov. Holland. 167 (1810)
Name: Rough waterclover
Distribution: Azores [I] (Terceira [I]); Australia (Australian Capital Territory, Western Australia, Northern Territory, South Australia, Queensland, Victoria, Tasmania); USA [I] (Florida [I])
21 azo 50 AUS 78 usa
Conservation Status: IUCN E
Synonyms: hidden
= Marsilea azorica Launert & Paiva; Iconogr. Selecta Fl. Azor. 2: 159 (1983)
= Zaluzianskia hirsuta (R.Br.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea latzii D.L.Jones; Fl. Australia 48: 707 (1998)
Distribution: Australia (Northern Territory, Tanami Desert)
50 AUS
species :
Marsilea macrocarpa C.Presl; Abh. Königl. Böhm. Ges. Wiss., Math.-Naturw. Cl. V, 5: 580 (1848)
Distribution: D.R.Congo [Zaire]; Uganda; Kenya; Tanzania; Angola; Zambia; Zimbabwe; Namibia; Botswana; South Africa (Limpopo, Gauteng, Mpulamanga, North West, Free State, KwaZulu-Natal, E-Cape Prov., N-Cape Prov.); Lesotho
23 ZAI 25 KEN TAN UGA 26 ANG ZAM ZIM 27 BOT LES NAM SAF
Synonyms: hidden
= Marsilea dregeana A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 428 (1864)
= Marsilea fischeri Hieron.; Engl., Pflanzenw. Ost-Afrikas C: 90 (1895)
= Marsilea macrocarpa var. dissecta Sim; The ferns of South Africa, edn. 2: 316, pl. 172, fig. 5 (1915)
= Marsilea macrocarpa var. lobata Sim; The ferns of South Africa, edn. 2: 316, pl. 172, fig. 6 (1915)
= Marsilea rotundata A.Br.; Kuhn, Filic. Afr.: 200 (1868)
= Zaluzianskia macrocarpa (C.Presl) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia rotundata (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea macropoda Engelm. ex A.Braun; Amer. J. Sci. Arts, ser. 2, 3: 56 (1847) (nota), ex Kunze, Amer. J. Sci. Arts, ser. 2, 6: 88 (1848)
Name: Bigfoot Water Clover
Distribution: USA (C-Texas, S-Texas, S-Louisiana [I], S-Alabama [I], New Mexico); Mexico (Coahuila, Hidalgo, Nuevo Leon, Tamaulipas)
77 USA 78 USA 79 MEX
Synonyms: hidden
= Marsilea macropus A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 418 (1864) (non Hook. (1854))
= Zaluzianskia macropus (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea megalomanica Launert; Senckenberg. Biol. 51: 435, f. 1-4 (1970)
Distribution: Tanzania
25 TAN
Conservation Status: IUCN I
species :
Marsilea minuta L.; Mant. Pl. 308 (1771)
Name: Dwarf waterclover
Distribution: Morocco; Algeria; Egypt; Mauritania; Senegal; Gambia; Guinea-Bissau; Mali; Burkina Faso; Ghana; Togo; Benin; Niger; Nigeria; Equatorial Guinea (Rio Muni); Congo [Brazzaville]; Central African Republic; D.R.Congo [Zaire]; Rwanda; Burundi; Chad; Sudan; Ethiopia; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; Botswana; South Africa (KwaZulu-Natal); Eswatini [Swaziland]; Madagascar; Comores (Anjouan, Mayotte); Mauritius; La Réunion; Afghanistan; Israel; Palestine; China (Anhui, Fujian, Guangdong, Guizhou, Hainan, Hubei, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang, ?Henan, ?Hunan, ?Shanxi); Japan; Ryukyu Isl.; Taiwan; Pakistan; Nepal; Bhutan; India (Jammu & Kashmir, Andhra Pradesh, Arunachal Pradesh, Assam State, Bihar, Chhattisgarh, Delhi, Gujarat, Himachal Pradesh, Jharkand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, ?Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, ?Sikkim, Tamil Nadu, Tripura, Uttarakhand, Uttar Pradesh, West Bengal); Sri Lanka; Bangladesh; Andaman Isl.; Myanmar; Thailand (widespread); Laos; Cambodia; Vietnam; peninsular Malaysia; Borneo (Sabah); Sulawesi; Philippines; Java; Lesser Sunda Isl. (Bali, Flores, Timor); New Guinea (Irian Jaya, Papua New Guinea); Australia (Northern Territory, Queensland); USA [I] (Florida [I], Georgia [I]); Trinidad [I]; Brazil [I] (Pernambuco [I], Rio de Janeiro [I])
20 ALG EGY MOR 22 BEN BKN GAM GHA GNB MLI MTN NGA NGR SEN TOG 23 BUR CAF CON EQG RWA ZAI 24 CHA ETH SUD 25 KEN TAN UGA 26 ANG MLW MOZ ZAM ZIM 27 BOT SAF SWZ 29 COM MAU MDG REU 34 AFG ISR PAL 36 CHI 38 JAP NNS TAI 40 BAN BHU IND NEP PAK SRL 41 AND CBD LAO MYA THA VIE 42 BOR JAW LSI MLY NWG PHI SUL 50 AUS 78 usa 81 trt 84 bra
Remarks: Marsilea brachycarpa is frequently kept separate
Synonyms: hidden
= Lemma minuta (L.) Desr.; Lam., Encycl. 3: 721 (1792)
= Marsilea aegyptiaca Wall.; Numer. List no. 7096 (1832), nomen
= Marsilea brachycarpa A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 420 (1864) (fide Fraser-Jenkins 2008)
= Marsilea brachypus A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 421 (1864) (fide Fraser-Jenkins 2008)
= Marsilea cornuta (A.Braun) A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 728 (1870)
= Marsilea coromandelica Burm.f.; Fl. Ind., t. 62, f. 3 (1768)
= Marsilea crenulata Desv.; Mem. Soc. Linn. Paris 6, 2: 179 (1827)
= Marsilea diffusa f. algeriensis A.Braun; Monatsber. Kgl. Preuss. Akad. Wiss. Berlin 1870: 727 (1871)
= Marsilea diffusa f. canariensis A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 727 (1871)
= Marsilea diffusa f. gracilipes A.Braun; Monatsber. Kgl. Preuss. Akad. Wiss. Berlin 1870: 728 (1871)
= Marsilea diffusa f. madagascariensis A.Braun; Monatsber. Kgl. Preuss. Akad. Wiss. Berlin 1870: 727 (1871)
= Marsilea diffusa f. microphylla A.Braun; Monatsber. Kgl. Preuss. Akad. Wiss. Berlin 1870: 727 (1871)
= Marsilea diffusa f. nilotica A.Braun; Monatsber. Kgl. Preuss. Akad. Wiss. Berlin 1870: 728 (1871)
= Marsilea diffusa Lepr. ex A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 419 (1864)
= Marsilea diffusa var. approximata A.Braun; Monatsber. Kgl. Preuss. Akad. Wiss. Berlin 1872: 661 (1873)
= Marsilea elata var. cornuta A.Braun; Kuhn, Filic. Afr.: 199 (1868)
= Marsilea erosa Willd.; Sp. Pl. ed. 4, 5: 540 (1810) [possibly a separate species of S-India fide Fraser-Jenkins et al. (2017)]
= Marsilea fimbriata Schumach.; Beskr. Guin. Pl. 461 (1827)
= Marsilea gracilenta A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 421 (1864) (fide Fraser-Jenkins 2008)
= Marsilea kedarmalii Bhardwaja, Gena & D´Souza; Indian Fern J. 11(1-2): 50 (1994 publ. 1995) (fide Fraser-Jenkins et al. 2017)
= Marsilea maheshwarii Gopal; Amer. Fern J. 58: 70 (1968) (fide Fraser-Jenkins 1997)
= Marsilea major (Haines) Chowdhury; Res. Living Pterid. 49 (1971) (fide Fraser-Jenkins 1997)
= Marsilea mearnsii Christ; Philipp. J. Sci., Bot. 8: 276 (1908)
= Marsilea microcarpa A.Braun; Flora 22: 300 (1839)
= Marsilea minuta subsp. brachycarpa (A.Braun) Bhardwaja
= Marsilea minuta var. indica K.M.Gupta; Marsilea Bot. Monogr. Council Sci. Industr. Res. India 2: 25 (1962)
= Marsilea minuta var. major Haines; Botany Bihar & Orissa 3: 1272 (1924) (fide Fraser-Jenkins 1997)
= Marsilea perrieriana C.Chr.; Dansk Bot. Ark. 7: 179 (1932)
= Marsilea poonensis Kolh.; Kew Bull. 1957: 293, f. 1 (1957) (fide Fraser-Jenkins 2008)
= Marsilea sinensis Hand.-Mazz.; Symb. Sin. 6: 47 (1929)
= Marsilea tenax Peter; Abh. Königl. Ges. Wiss. Göttingen, Math.-Phys. Kl., n.f., 13: 38 (1928)
= Marsilea tetraphylla Thunb.; Fl. Jap. 340 (1784)
= Marsilea vulgaris Bory; Bél., Voy. Indes Or., Bot. 2(1): 3 (1833)
= Zaluzianskia coromandelica (Burm.f.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia crenulata (Desv.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia diffusa (Lepr. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia fimbriata (Schumach.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia minuta (L.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
variant :
var. incurva (A.Braun) Launert; Garcia de Orta, Ser. Bot. 6(1-2): 126 (1984)
Distribution: Senegal; Gambia; Guinea; Ghana; Kenya; Tanzania
22 GAM GHA GUI SEN 25 KEN TAN
Synonyms: hidden
= Marsilea crenulata var. incurva (A.Braun) A.Braun; Filices Afr. 198 (1868)
= Marsilea crenulata var. senegalensis (A.Braun) C.Chr.; Index Filic. 419 (1906)
= Marsilea diffusa var. incurva A.Braun; Monatsb. K. Preuss. Akad. Wiss. Berlin 1863: 419 (1864)
= Marsilea senegalensis A.Braun; Flora 22: 300 (1839), nom. nud.
= Zaluzianskia senegalensis (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea mollis B.L.Rob. & Fernald; Proc. Amer. Acad. 30: 123 (1895)
Name: Chihuahuan waterclover
Distribution: USA (Arizona, New Mexico, Texas); Mexico (Aguascalientes, Chiapas, Chihuahua, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sonora, Tamaulipas, Veracruz, Zacatecas); Colombia (Cundinamarca); Ecuador; Peru; Bolivia (Cochabamba, La Paz, Tarija); Argentina (Jujuy, Tucuman); Chile (Coquimbo, Valparaíso)
76 USA 77 USA 79 MEX 82 CLM 83 BOL ECU PER 85 ARG CLE
Synonyms: hidden
= Marsilea punae de la Sota; Darwiniana 20(1-2): 226 (1976)
species :
Marsilea mutica Mett.; Ann. Sci. Nat., Bot., sér. IV, 15: 88 (1861)
Distribution: Australia (N-Western Australia, Northern Territory, E-Queensland, SE-New South Wales, Australian Capital Territory, SE-Victoria [I], S-Tasmania); New Zealand [I]; New Caledonia
50 AUS 51 nzl 60 NWC
Synonyms: hidden
= Marsilea brownii A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 418 (1864)
= Marsilea quadrifolia R.Br.; Prodr. Fl. Nov. Holland. 167 (1810) (non L. (1753))
= Zaluzianskia mutica (Mett.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia brownii (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea nashii Underw.; Bull. New York Bot. Gard. 137 (1906)
Distribution: Bahamas; Turks & Caicos Isl.; Cuba
81 BAH CUB TCI
species :
Marsilea nubica A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 432 (1864)
Distribution: Mauritania; Senegal; Mali; Nigeria; Central African Republic; Chad; Sudan; Kenya; Tanzania; Angola; Mozambique; Namibia; Botswana; Madagascar
22 MLI MTN NGA SEN 23 CAF 24 CHA SUD 25 KEN TAN 26 ANG MOZ 27 BOT NAM 29 MDG
Synonyms: hidden
= Marsilea leiocarpa C.Chr.; Cat. Pl. Madag. Pterid. 66 (1932) (nomen), Dansk Bot. Ark. 7: 180, t. 73, f. 10-12 (1932)
= Zaluzianskia nubica (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
variant :
var. gymnocarpa (A.Braun) Launert; Garcia de Orta, Ser. Bot. 6(1-2): 124 (1984)
Distribution: Mauritania; Senegal; Chad; Sudan; Tanzania; Angola; Namibia; South Africa (Limpopo); Madagascar
22 MTN SEN 24 CHA SUD 25 TAN 26 ANG 27 NAM SAF 29 MDG
Synonyms: hidden
= Marsilea gymnocarpa Lepr.; A. Braun, Flora 300 (nomen), ex Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 422 (1864)
= Zaluzianskia gymnocarpa (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea oligospora Goodd.; Bot. Gaz. 33: 66 (1902)
Name: Pacific waterclover
Distribution: USA (California, Idaho, Montana, Nevada, Oregon, Utah, Washington State, Wyoming)
73 USA 76 USA
Synonyms: hidden
= Marsilea vestita var. oligospora (Goodd.) Dorn; Vasc. Pl. Wyoming 295 (1988)
species :
Marsilea owambo Doweld; Amer. Fern J. 106(2): 143 (2016), nom. nov.
Distribution: ?Ethiopia; Zambia; W-Zimbabwe; Namibia; Botswana
26 ZAM ZIM 27 BOT NAM
Synonyms: hidden
= Marsilea vera Launert; Mitt. Bot. Staatssamml. München 3: 505 (1960), nom. illeg.
species :
Marsilea polycarpa Hook. & Grev.; Icon. Filic. t. 160 (1829)
Distribution: Society Isl. (Moorea); Mexico (Guerrero, Jalisco, Nayarit, Tabasco, Veracruz); Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Cuba; Jamaica; Hispaniola (Haiti, Dominican Republic); Puerto Rico; Trinidad; Colombia (Atlántico, Bolívar, Casanare, Córdoba, Huila); Venezuela (Amazonas, Apure, Aragua, Barinas, Bolívar, Carabobo, Cojedes, Delta Amacuro, Guarico, Monagas, Portuguesa, Sucre, Zulia); Guyana; Surinam; French Guiana; Ecuador; Bolivia (Santa Cruz); Brazil (Bahia, Maranhão, Mato Grosso do Sul, Pará, Pernambuco, Roraima, Santa Catarina)
61 SCI 79 MEX 80 COS GUA HON NIC PAN 81 CUB HIS JAM PUE TRT 82 CLM FRG GUY SUR VEN 83 BOL ECU 84 BRA
Synonyms: hidden
= Marsilea brasiliensis Mart.; Icon. Pl. Crypt. 122, t. 73, f. 2 (1834)
= Marsilea polycarpa var. mexicana A.Braun; (1870 publ. 1871)
= Zaluzianskia polycarpa (Hook. & Grev.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea pyriformis Bouchart; Adansonia 41(1): 14 (2019)
Distribution: Madagascar
29 MDG
species :
Marsilea quadrata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 745 (1870)
Distribution: Borneo (Sabah)
42 BOR
Synonyms: hidden
= Zaluzianskia quadrata (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea quadrifolia L.; Sp. Pl. 2: 1097 (1753)
Name: Vierblättriger Kleefarn (DE); European Water Clover, Water shamrock (EN); Trebol de agua (ES)
Distribution: Germany (Baden-Württemberg, Bayern+, Hessen+, Nordrhein-Westfalen [I], Rheinland-Pfalz); Switzerland; Austria; Poland; Slovakia; Hungary; Portugal; Spain; France; Italy; Slovenia; Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (NE-Greece); European Russia (W-European Russia, E-European Russia); Moldova; Ukraine; Canary Isl. (Gran Canaria); Kazakhstan; Uzbekistan; Tajikistan; Northern Caucasus; Georgia [Caucasus] (Adzharia); Azerbaijan (incl. Talysh); +Turkey (Anatolia); Iran; SE-Afghanistan; China (Gansu, Hebei, Heilongjiang, Henan, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi); South Korea; Japan; E-Canada [I] (Ontario [I]); USA [I] (Connecticut [I], Delaware [I], Illinois [I], Indiana [I], Iowa [I], Kentucky [I], Maine [I], Maryland [I], Massachusetts [I], Michigan [I], Missouri [I], New Hampshire [I], New Jersey [I], New York [I], Ohio [I], Pennsylvania [I], Rhode Island [I], Vermont [I])
11 AUT GER HUN POL SLO SWI 12 FRA POR SPA 13 ALB BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB 14 MLD RUS UKR 21 CNY 32 KAZ TZK UZB 33 AZE GEO NCS 34 AFG IRN TUR 36 CHI 38 JAP KOR 72 cdn 74 usa 75 usa 78 usa
Remarks: Distribution records for China and E-Asia might belong to other species
Synonyms: hidden
= Lemma quadrifolia (L.) Desr.; Lam., Encycl. 3: 720 (1792)
= Marsilea europaea Desv.; Mem. Soc. Linn. Paris 6, 2: 178 (1827)
= Marsilea quadrifolia f. aquatica A.Braun ex Glück; (1911)
= Marsilea quadrifolia f. natans Glück; (1923), nom. illeg.
= Marsilea quadrifolia f. submersa Glück; (1923)
= Marsilea quadrifolia f. terrestris Glück; (1923)
= Pteris quadrifolia (L.) L.; Syst. Nat. ed. X, 2: 1321 (1759)
= Spheroidia quadrifolia (L.) Dulac; Fl. Dépt. Hautes-Pyrenees 39 (1867)
= Zaluzianskia marsileoides Neck.; Acta Theod. pal. phys. 8: 303 (1775)
= Zaluzianskia quadrifolia (L.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea scalaripes D.M.Johnson; Amer. Fern J. 78(2): 68 (1988)
Distribution: peninsular Malaysia (only known from type, Kedah)
42 MLY
species :
Marsilea schelpeana Launert; Mitt. Bot. Staatssamml. München 3: 506, f. 1 (1960)
Distribution: South Africa (E-Cape Prov., W-Cape Prov.)
27 SAF
Conservation Status: IUCN V
species :
Marsilea strigosa Willd.; Sp. Pl. 5(1): 539 (1810)
Name: Gestreifter Kleefarn
Distribution: Spain; Baleares; France; Sardinia; Italy; European Russia (E-European Russia); Morocco; Algeria; Egypt; Kazakhstan; Azerbaijan (Talysh); SE-Afghanistan
12 BAL FRA SAR SPA 13 ITA 14 RUS 20 ALG EGY MOR 32 KAZ 33 AZE 34 AFG
Synonyms: hidden
= Marsilea fabri Dunal ex St.-Hil.; Ann. Sci. Nat., Bot., sér. II, 6: 378 (1836)
= Marsilea pubescens Ten.; Cat. Pl. Hort. Neapol. App. 1: 215 (1815)
= Marsilea strigosa var. pubescens (Ten.) Maire & Weiller; Encyclopédie Biologique 33: 83, fig. 45 (1952)
= Zaluzianskia pubescens (Ten.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia strigosa (Willd.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea subterranea Lepr.; A. Braun, Flora 301 (nomen), et Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1863: 433 (1864)
Distribution: Mauritania; Senegal; Nigeria; Chad; Tanzania
22 MTN NGA SEN 24 CHA 25 TAN
Synonyms: hidden
= Zaluzianskia subterranea (Lepr.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea unicornis Launert; Senckenberg. Biol. 49: 303, f. 53-54 (1968)
Distribution: Angola; Namibia; Botswana
26 ANG 27 BOT NAM
species :
Marsilea vestita Hook. & Grev.; Icon. Filic. t. 159 (1829)
Name: Hairy Water Clover, Hairy peppermint
Distribution: Canada (Alberta, British Columbia, Saskatchewan); USA (Alabama, Arkansas, Arizona, California, Colorado, Florida, Iowa, Idaho, Kansas, Louisiana, Minnesota, Mississippi, Montana, North Dakota, Nebraska, New Mexico, Nevada, Oklahoma, Oregon, South Dakota, Texas, Utah, Washington State, Wyoming); Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chihuahua, Coahuila, Ciudad de Mexico, Durango, Guanajuato, Jalisco, Nuevo Leon, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Zacatecas); Bahamas; Cuba; Barbuda; Peru
71 CDN 73 USA 74 USA 75 USA 76 USA 77 USA 78 USA 79 MEX 81 ABA BAH CUB 83 PER
Synonyms: hidden
= Marsilea brevipes Nutt. ex A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 741 (1870)
= Marsilea fournieri C.Chr.; Index Filic. 418 (1906)
= Marsilea longipes Austin; Bull. Torrey Bot. Club 3(4): 23 (1872)
= Marsilea minuta E.Fourn.; Bull. Soc. Bot. France 27: 329 (1880) (non L. (1771))
= Marsilea mucronata A.Braun; Amer. J. Sci. Arts, Bot. II, 3: 55, f. 2 (1847)
= Marsilea picta Fée; Mém. Foug. 9 (Cat. Foug. Mex.): 47 (1857)
= Marsilea tenuifolia Engelm. ex A.Braun; Amer. J. Sci. Arts, ser. 2, 6: 89 (1848)
= Marsilea uncinata A.Braun; Flora 300 (1889) (nomen), et Amer. J. Sci. Arts, Bot. II, 3: 55, f. 1 (1847)
= Marsilea vestita subsp. tenuifolia (Engelm. ex A.Braun) D.M.Johnson; Syst. New World Spp. Marsilea (Marsileac.) (Syst. Bot. Monog. 11): 69 (1986)
= Marsilea vestita var. mucronata (A.Braun) Baker
= Marsilea vestita var. tenuifolia (Engelm. ex A.Braun) Underw.; Bull. Torrey Bot. Club 14: 92 (1887)
= Zaluzianskia tenuifolia (Engelm. ex A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia uncinata (A.Braun) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
= Zaluzianskia vestita (Hook. & Grev.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea villifolia Bremek. & Oberm. ex Alston & Schelpe; J. S. Afr. Bot. 18: 156, 166 (1952)
Distribution: Namibia; Botswana; South Africa (Limpopo, North West)
27 BOT NAM SAF
Synonyms: hidden
= Marsilea villosa Burch. ex Bremek. & Oberm.; Ann. Transvaal Mus. 16: 400 (1935) (non Kaulf. (1824)
species :
Marsilea villosa Kaulf.; Enum. Filic. 272 (1824)]
Distribution: Hawaii (Niihau, Oahu, Molokai)
63 HAW
Conservation Status: IUCN E
Synonyms: hidden
= Zaluzianskia villosa (Kaulf.) Kuntze; Revis. Gen. Pl. 2: 823 (1891)
species :
Marsilea ×subangulata A.Braun; Monatsber. Königl. Preuss. Akad. Wiss. Berlin 1870: 46 (1870)
Distribution: Jamaica; Colombia (Huila); Venezuela (Venezuela Distrito Federal)
81 JAM 82 CLM VEN
Remarks: Marsilea crotophora x M. deflexa
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A complete, synonymic checklist of the Higher Plants of the World

Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life