518.0000
Family
Lamiaceae Martinov; Tekhno-Bot. Slovar 355 (1820), nom. cons., nom. alt.
Latest phylogeny: Zhao et al. 2021
Species listed in database: 7814; hybrids: 382
= Aegiphilaceae Raf.; Sylva Tellur. 161 (1838)
= Ajugaceae Döll; Rhein. Fl. [Döll] 375 (1843)
= Chloanthaceae Hutch.; Fam. Fl. Pl., Dicot., ed. 2: 396 (1959)
= Glechomaceae Martinov; Tekhno-Bot. Slovar 288 (1820)
= Labiatae Juss.; Gen. Pl. [Jussieu] 110 (1789), nom. cons.
= Melissaceae Bercht. & J.Presl; Přir. Rostlin 245 (1820)
= Melittidaceae Martinov; Tekhno-Bot. Slovar 390 (1820)
= Menthaceae Burnett; Outlines Bot. [Burnett] 969, 1095, 1106 (1835)
= Monardaceae Döll; Fl. Baden [Döll] 2: 661 (1858)
= Nepetaceae Bercht. & J.Presl; Přir. Rostlin 245 (1820)
= Salazariaceae F.A.Barkley; Phytologia 32: 304 (1975)
= Salviaceae Bercht. & J.Presl; Přir. Rostlin 245 (1820)
= Saturejaceae Döll; Fl. Baden [Döll] 2: 664. med (1858)
= Scutellariaceae Döll; Rhein. Fl. [Döll] 373 (1843)
= Siphonanthaceae Raf.; Fl. Tellur. 4: 87 (1838)
= Stachydaceae Döll; Rhein. Fl. [Döll] 366 (1843)
= Symphoremataceae Wight; Icon. Pl. Ind. Orient. [Wight] 4(3): 13 (1849)
= Viticaceae Juss.; Gen. Pl. [Jussieu] 106 (1789)
518.8702
Genus
Stachys L.; Sp. Pl. [Linnaeus] 2: 580 (1753)
Distribution: temp., subtropical and trop. regions, few in trop. Africa (22), except Malesia, Australia, and New Zealand. The genus is hopelessly polyphyletic and needs splitting in 9 genera (Salmaki et al. 2013, 2019); alternatively all of Sideritis and Prasium would have to be included.
Species listed in database: 371; hybrids: 10
= Aspasia E.Mey. ex Pfeiff.; Nomencl. Bot. [Pfeiffer] 1: 295 (1873)
= Bonamya Neck.; Elem. Bot. [Necker] 1: 316 (1790), opus utique oppr.
= Chamaesphacos Schrenk ex Fisch. & C.A.Mey.; Enum. Pl. Nov. [Fischer & Meyer] 1: 27 (1841)
= Eriostemum Steud.; Nomencl. Bot. [Steudel], ed. 2, 1: 588 (1841), orth. var.
= Eriostomum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)
= Galeopsis Hill; Brit. Herb. [Hill] 359 (1756), nom. illeg.
= Galeopsis Moench; Methodus [Moench] 397 (1794), nom. illeg.
= Koevia Krestovsk.; Novosti Sist. Vyssh. Rast. 45: 94 (2014)
= Lamiostachys Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 91: 1258 (2006)
= Olisia Spach; Hist. Nat. Vég. [Spach] 9: 166 (1840)
= Ortostachys Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 136 (1869)
= Phlomidoschema (Benth.) Vved.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 9: 54 (1941)
= Stachyus St.-Lag.; Ann. Soc. Bot. Lyon 7: 109 (1880)
= Suzukia Kudô; J. Soc. Trop. Agric. 2: 145 (1930)
= Tapeinanthus Boiss.; Pl. Pers. Bor. [Kotschy] 18 (1846), nom. illeg.
= Tapeinanthus Boiss. ex Benth.; Prodr. [A.P. de Candolle] 12: 436 (1848)
= Tetrahitum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 103 (1809)
= Thuspeinanta T.Durand; Index Gen. Phan. [Durand] 78 (1888)
= Trixago Haller; Hist. Stirp. Helv. 1: 101 (1768)
= Trixella Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 135 (1869)
= Zietenia Gled.; Syst. Pl. [Gleditsch] 185 (1764)
Search for: Stachys pilifera-subsp.-ixodes
Nothing found!
species :
Stachys acerosa
Boiss.; Diagn. Pl. Orient. 7: 57 (1846)
Distribution: Iran (SW-Iran)
34 IRN
Remarks: clade 12 Olisia
species :
Stachys aculeolata
Hook.f.; J. Proc. Linn. Soc., Bot. 6: 18 (1861)
Distribution: Nigeria; Cameroon; Bioko Isl. [Fernando Poo]; D.R.Congo [Zaire]; Rwanda; Burundi; Ethiopia; Uganda; Kenya; Tanzania
22 NGA 23 BIO BUR CMN RWA ZAI 24 ETH 25 KEN TAN UGA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys aberdarica T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 639 (1923)
= Stachys aculeolata var. afromontana T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 633 (1923)
= Stachys aspericaulis Pax; Bot. Jahrb. Syst. 39: 645 (1907)
= Stachys bambuseti T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 635 (1923)
= Stachys lindblomiana T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 641 (1923)
= Stachys sidamoensis f. neumannii T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 637 (1923)
= Stachys sidamoensis Gürke; Bot. Jahrb. Syst. 36: 126 (1905)
= Stachys subrenifolia T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 644 (1923)
variant :
var. camerunensis
T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 629 (1921)
Distribution: Cameroon
23 CMN
variant :
var. inermis
Y.B.Harv.; Kew Bull. 51(3): 440 (1996)
Distribution: D.R.Congo [Zaire]; Burundi; Uganda; Tanzania; Malawi
23 BUR ZAI 25 TAN UGA 26 MLW
species :
Stachys adulterina
Hemsl.; J. Linn. Soc., Bot. 26: 300 (1890)
Distribution: China (Hubei, Sichuan)
36 CHI
species :
Stachys aegyptiaca
Pers.; Syn. Pl. [Persoon] 2: 124 (1806)
Distribution: Egypt (Eastern Desert, NE-Egypt); Israel (S-Negev Desert); Sinai peninsula (C-Sinai, S-Sinai); Jordania (S-Jordania)
20 EGY 34 ISR JOR SIN
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys affinis Fresen.; Mus. Senckenberg. 1: 91 (1833), nom. illeg.
= Stachys boveana Bois; Bull. Mus. Natl. Hist. Nat. 24: 149 (1918)
= Stachys palaestina Vahl; Symb. Bot. [Vahl] 2: 64 (1791), nom. illeg.
= Stachys pauciflora Benth.; Labiat. Gen. Spec. [Bentham] 560 (1834)
species :
Stachys aethiopica
L.; Mant. Pl. 1: 82 (1767)
Distribution: Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, N-Cape Prov., W-Cape Prov., E-Cape Prov.)
26 MOZ ZIM 27 SAF
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Betonica capensis Burm.f.; Fl. Ind. [N.L.Burman] Prodr. Fl. Cap. 16 (1768)
= Stachys attenuata Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 351 (1910)
= Stachys capensis C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 530 (1845)
= Stachys fruticetorum Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1083 (1903)
= Stachys harveyi Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 350 (1910)
= Stachys hispidula Hochst.; Flora 28: 66 (1845)
= Stachys pulchella Salisb.; Prodr. Stirp. Chap. Allerton 83 (1796), nom. illeg.
= Stachys serrulata Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 549 (1833)
species :
Stachys affinis
Bunge; Enum. Pl. China Bor. 51 (1833)
Name: Artichoke betony (EN), Knollen-Ziest, Japanknolle (DE)
Distribution: Germany [I]; China (Gansu, Hebei, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Xinjiang); Myanmar (Kachin, Shan); USA [I] (New York [I])
11 ger 36 CHI 41 MYA 75 usa
Synonyms: hidden
= Stachys geobombycis var. alba C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 223 (1965)
= Stachys sieboldii Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 112 (1865)
= Stachys sieboldii var. alba (C.Y.Wu & H.W.Li) H.B.Chen; Fl. Fujianica 4: 604 (1989 publ. 1990)
= Stachys sieboldii var. glabrescens C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 222 (1965)
= Stachys sieboldii var. malacotricha Hand.-Mazz.; Acta Horti Gothob. 9: 83 (1934)
= Stachys tubifera Naudin; Rev. Hort. (Paris) 59: 290 (1887), nom. inval.
species :
Stachys agraria
Schltdl. & Cham.; Linnaea 5: 100 (1830)
Name: Mousesear
Distribution: Mexico (Aguascalientes, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala; Honduras; Nicaragua; Galapagos [I]
79 MEX 80 GUA HON NIC 83 gal
Synonyms: hidden
= Stachys agraria f. albiflora Benke; Rhodora 35: 45 (1933)
= Stachys camporum M.E.Jones; Contr. W. Bot. 12: 70 (1908)
= Stachys confusa Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 122 (1898)
= Stachys polysegia Briq.; Annuaire Conserv. Jard. Bot. Genève 13/14: 395 (1911)
= Stachys umbrosa Scheele; Linnaea 22: 595 (1849)
species :
Stachys ahmetsavranii
Doğu & Bağcı; Bangladesh J. Bot. 50(2): 320, figs. 1-2 (2021)
Distribution: Turkey
34 TUR
species :
Stachys ajugoides
Benth.; Linnaea 6: 80 (1831)
Name: Bugle hedgenettle
Distribution: USA (California)
76 USA
Synonyms: hidden
= Stachys ajugoides var. cymosa Jeps.; Fl. Calif. [Jepson] 3: 427 (1943), nom. inval.
= Stachys ajugoides var. stricta Jeps.; Fl. W. Calif. [Jepson] 457 (1901)
= Stachys ajugoides var. velutina Jeps.; Fl. W. Calif. [Jepson] 457 (1901)
= Stachys cymosa A.Heller ex Jeps.; Fl. Calif. [Jepson] 3: 427 (1943), pro syn.
= Stachys ramosa A.Heller; Muhlenbergia 1: 116 (1905)
species :
Stachys alabamica
B.R.Keener & L.J.Davenp.; J. Bot. Res. Inst. Texas 10(2): 318, figs, 2-4 (2016)
Distribution: USA (Alabama)
78 USA
species :
Stachys albens
A.Gray; Proc. Amer. Acad. Arts 7: 387 (1868)
Name: Whitestem hedgenettle
Distribution: USA (California, Nevada [I], Utah [I])
76 USA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys albens var. juliensis Jeps.; Man. Fl. Pl. Calif. [Jepson] 877 (1925)
= Stachys lanuginosa Greene; Pittonia 3: 342 (1898)
= Stachys malacophylla Greene; Pittonia 3: 343 (1898)
= Stachys pycnostachya Torr.; U.S. Expl. Exped., Phan. Pacific N. Amer. 17: 408 (1874)
= Stachys velutina Greene; Erythea 2: 121 (1894), nom. illeg.
species :
Stachys albicaulis
Lindl.; Edwards´s Bot. Reg., t. 1558 (1833)
Name: Hierba de Santa María [Chile]
Distribution: Argentina (Mendoza); Chile (Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía)
85 ARG CLE
Synonyms: hidden
= Stachys albicaulis var. candidissima Reiche; Anales Univ. Chile 122: 284 (1908)
= Stachys eriophylla Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 113 (1898)
species :
Stachys albiflora
N.E.Br.; Bull. Misc. Inform. Kew 1901: 131 (1901)
Distribution: South Africa (Free State, KwaZulu-Natal)
27 SAF
species :
Stachys albotomentosa
Ramamoorthy; Anales Inst. Biol. Univ. Nac. Autón. México, Bot. 54: 158 (1983 publ. 1987)
Distribution: Mexico (Guanajuato, Hidalgo, Queretaro, San Luis Potosi, Tamaulipas)
79 MEX
Remarks: coccinea agg.
variant :
var. potosina
B.L.Turner; Phytologia 76(5): 393 (1994)
Distribution: Mexico (San Luis Potosi)
79 MEX
species :
Stachys aleurites
Boiss. & Heldr. ex Benth.; Prodr. [A.P. de Candolle] 12: 472 (1848)
Distribution: Turkey (SW-Anatolia)
34 TUR
Synonyms: hidden
= Stachys tengwalliana Rech.f.; Oesterr. Bot. Z. 99: 46 (1952)
species :
Stachys alpigena
T.C.E.Fr.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 641 (1923)
subspecies :
subsp. alpigena
Distribution: Rwanda; Kenya
23 RWA 25 KEN
Remarks: clade 2 Stachys s. str.
subspecies :
subsp. longipetala
Sebsebe; Kew Bull. 48(2): 330 (1993)
Distribution: Ethiopia
24 ETH
species :
Stachys alpina
L.; Sp. Pl. [Linnaeus] 2: 581 (1753)
subspecies :
subsp. alpina
Name: Alpen-Ziest (DE), Limestone Woundwort (EN)
Distribution: England (Wales, W-England); Belgium; Luxembourg; Germany (Baden-Württemberg, Bayern, Hessen, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Saarland, Sachsen, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; N- & C-Spain; Andorra; France; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece (mountains of N-Greece); Ukraine
10 GRB 11 AUT BGM CZE GER HUN LIE LUX POL SLO SWI 12 ADO FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SRB 14 UKR
Remarks: clade 1 Eriostomum, germanica agg.; doubtfully native in England
Synonyms: hidden
= Eriostomum alpinum (L.) Hoffmanns. & Link ex Steud.; Nomencl. Bot. [Steudel], ed. 1, 1: 806 (1821), nom. inval.
= Galeopsis fusca Moench; Methodus [Moench] 397 (1794), nom. illeg.
= Stachys curviflora Tausch; Flora 14: 216 (1831)
= Stachys dinarica De Marb.; Acta Univ. Lund. 27(5): 61 (1892)
= Stachys fusca (Moench) Moench; Methodus [Moench], Suppl. 136 (1802)
= Stachys orientalis L.; Sp. Pl. [Linnaeus] 2: 582 (1753)
= Stachys urticifolia Tausch; Flora 14: 216 (1831)
subspecies :
subsp. dinarica
Murb.; Beitr. Fl. Südbosnien [Acta Univ. Lund. 27(5)]: 61 (1891)
Distribution: Croatia; Montenegro; ?Serbia; Kosovo; Bulgaria
13 BUL CRO KOS MNT
subspecies :
subsp. macrophylla
(Albov) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 277 (1974)
Distribution: Georgia [Caucasus]; Turkey (N-Anatolia); Iran
33 GEO 34 IRN TUR
Synonyms: hidden
= Stachys macrophylla Albov; Prodr. Fl. Colchic. 202 (1895)
species :
Stachys amanica
P.H.Davis; Kew Bull. [6]: 108 (1951)
Distribution: Turkey (S-Anatolia)
34 TUR
Synonyms: hidden
= Stachys debilis Bornm.; Notizbl. Bot. Gart. Berlin-Dahlem 7: 35 (1917), nom. illeg.
species :
Stachys anamurensis
Sümbül; Turk. J. Bot. 14(1): 57 (1990)
Distribution: Turkey
34 TUR
species :
Stachys andongensis
Hiern; Cat. Afr. Pl. [Hiern] 1: 874 (1900)
Distribution: Angola
26 ANG
species :
Stachys angustifolia
M.Bieb.; Fl. Taur.-Caucas. 2: 52 (1808)
Distribution: Croatia; Romania; Bulgaria; European Turkey; Greece (scattered N-Greece, south to Mt. Olimbos, Mt. Athos, Thasos); ?C-European Russia; Ukraine; Crimea
13 BUL CRO GRC ROM TUE 14 KRY UKR
Synonyms: hidden
= Stachys abrotanifolia Boiss. & Heldr.; Fl. Orient. [Boissier] 4(2): 732 (1879)
= Stachys angustifolia subsp. tenuifolia Nyman; Consp. Fl. Eur. 580 (1881)
= Stachys angustifolia var. haynaldii Janka ex Nyman; Consp. Fl. Eur. 580 (1881), nomen
= Stachys tenuifolia Janka; Oesterr. Bot. Z. 22: 180 (1872), nom. illeg.
= Stachys tenuifolia Pall. ex Link; Enum. Hort. Berol. Alt. 2: 109 (1822), nom. illeg.
species :
Stachys anisochila
Vis. & Pančić; Mem. Reale Ist. Veneto Sci. 15: 13 (1870)
Distribution: Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; ?North Macedonia; S- & E-Bulgaria
13 BIH BUL KOS MNT SRB
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys anisochila var. sendteri (Beck) Nyman; Consp. Fl. Eur. Suppl. 2: 252 (1890)
= Stachys recta subsp. anisochila (Vis. & Pančić) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)
= Stachys sendteri Beck; Ann. K.K. Naturhist. Hofmus. 2: 144 (1887)
= Stachys sendteri f. versipila H.Lindb.; Öfvers. Finska Vetensk.-Soc. Förh. 48(13): 92 (1906)
= Stachys sendteri var. adenocalyx H.Lindb.; Öfvers. Finska Vetensk.-Soc. Förh. 48(13): 92 (1906)
species :
Stachys annua
(L.) L.; Sp. Pl. [Linnaeus], ed. 2: 813 (1763)
subspecies :
subsp. annua
Name: Einjähriger Ziest (DE), Annual Yellow-woundwort (EN)
Distribution: England [I] (England [I], Wales [I]); Denmark [I]; Norway [I]; Sweden [I]; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin [I], Baden-Württemberg, Bayern, Hessen, +Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein [I], Saarland, +Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; N- & NW-Spain; France; Italy; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (scattered mountains of NE-, NW-, C-Greece, rare N-Peloponnisos); S-European Russia; Estonia [I]; Latvia [I]; Lithuania [I]; Belarus; C-European Russia; E-European Russia; N-European Russia; NW-European Russia; Ukraine; Crimea; Siberia (Altai, Krasnoyarsk, Tuva, W-Siberia); Russian Far East [I] (Amur [I], Primorye [I]); Kazakhstan; Uzbekistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia, S-Anatolia, SE-Anatolia, SSW-Anatolia, W-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, W-Iran); Lebanon (C-Lebanon, coastal W-Lebanon); Syria (coastal W-Syria, NW-Syria, W-Syrian Mountains); Canada [I] (New Brunswick [I]); USA [I] (Kansas [I], Massachusetts [I], Maine [I], New Jersey [I], New York [I], Pennsylvania [I], Rhode Island [I], Virginia [I])
10 den grb nor swe 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 FRA SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SMA SRB TUE 14 BLR est KRY lat lit RUS UKR 30 SIB 31 rfe 32 KAZ UZB 33 ARM AZE GEO NCS 34 IRN IRQ LEB SYR TUR 72 cdn 74 usa 75 usa 78 usa
Remarks: clade 12 Olisia
Synonyms: hidden
= Betonica annua L.; Sp. Pl. [Linnaeus] 2: 578 (1753)
= Betonica annua var. glabra L.; Sp. Pl. [Linnaeus] 2: 573 (1753)
= Betonica annua var. hirsuta L.; Sp. Pl. [Linnaeus] 2: 574 (1753)
= Olisia annua (L.) Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 136 (1869)
= Olisia delphinensis Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 136 (1869)
= Prasium stachydium E.H.L.Krause; Deutschl. Fl. [Sturm], ed. 2, Abbild., 11: 115 (1903), nom. illeg.
= Sideritis trinervia Gilib.; Fl. Lit. Inch. 1: 81 (1782), opus utique oppr.
= Stachys adenocalyx K.Koch; Linnaea 21: 691 (1848)
= Stachys annua var. maxima K.Koch; Linnaea 17: 299 (1844)
= Stachys balbisii Link; Enum. Hort. Berol. Alt. 2: 110 (1822)
= Stachys betonica Crantz; Stirp. Austr. Fasc., ed. 2, 2: 264 (1769), nom. illeg.
= Stachys decumbens Willd. ex Benth.; Linnaea 11: 336 (1837), nom. illeg.
= Stachys lazica Boiss.; Fl. Orient. [Boissier] 4(2): 745 (1879)
= Stachys micrantha K.Koch; Linnaea 21: 690 (1849)
= Stachys nervosa Gaterau; Descr. Pl. Montauban 107 (1789)
= Stachys procumbens Willd. ex Benth.; Linnaea 11: 336 (1837)
= Stachys pubescens Ten.; Fl. Napol. 1(Prodr.): XXXIV (1812)
= Stachys pubescens var. grandiflora Regel; Index Sem. (St. Petersburg [Petropolitanus]) 1867: 32 (1868)
subspecies :
subsp. ammophila
(Boiss. & C.I.Blanche) Sam.; Ark. Bot., a. s. 5: 376 (1960)
Distribution: Turkey; Lebanon; Syria
34 LEB SYR TUR
Synonyms: hidden
= Stachys annua var. ammophila Boiss. & C.I.Blanche; Diagn. Pl. Orient., Sér. 2, 4: 43 (1859)
subspecies :
subsp. cilicica
(Boiss.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 289 (1974)
Distribution: Turkey; ?Lebanon; Syria
34 SYR TUR
Synonyms: hidden
= Stachys cilicica Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 42 (1859)
variant :
var. lycaonica
R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 289 (1974)
Distribution: Turkey
34 TUR
species :
Stachys antalyensis
Ayaşlıgil & Davis; Notes Roy. Bot. Gard. Edinburgh 42: 72, f. ID-F (1984)
Distribution: Turkey (SW-Anatolia)
34 TUR
species :
Stachys aperta
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 37 (1934)
Distribution: Peru; Bolivia (Chuquisaca, Cochabamba, La Paz)
83 BOL PER
species :
Stachys appalachiana
D.B.Poind. & J.B.Nelson; J. Bot. Res. Inst. Texas 5(2): 405 (2011)
Distribution: USA (North Carolina)
78 USA
species :
Stachys arabica
Hornem.; Hort. Bot. Hafn. [Hornemann] 2: 554 (1815)
Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon); Syria (NW-Syria, Jbel Druze); Israel (coastal W-Israel, Rift Valley, N-Israel); Jordania (S-Jordania, W-Jordania)
34 ISR JOR LEB SYR TUR
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys macrosperma Boiss.; Diagn. Pl. Orient. 12: 84 (1853)
species :
Stachys arachnoidea
Codd; Bothalia 12(2): 182 (1977)
Distribution: South Africa (Limpopo, Mpulamanga); Eswatini [Swaziland]
27 SAF SWZ
species :
Stachys araucana
Phil.; Anales Univ. Chile 90: 563 (1895)
Distribution: Chile
85 CLE
Remarks: provisionally accepted name
species :
Stachys arenaria
Vahl; Symb. Bot. [Vahl] 2: 64 (1791)
Name: Sand-Ziest (DE)
Distribution: Italy; Sicily; Croatia; Morocco; Algeria; Tunisia
13 CRO ITA SIC 20 ALG MOR TUN
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys arenaria subsp. divaricatidens H.Lindb.; Acta Soc. Sci. Fenn., Ser. B, Opera Biol. 1(2) (Itin. Medit.): 133 (1932)
= Stachys arenaria subsp. mollis (Willd. ex Benth.) F.Gómez; Anales Jard. Bot. Madrid 58(1): 199 (2000)
= Stachys arenaria var. angustifolia (Sennen) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 19 (1979), nom. inval.
= Stachys arenaria var. divaricatidens (H.Lindb.) Maire; Bull. Soc. Hist. Nat. Afrique N. 24: 227 (1933)
= Stachys arenaria var. latifolia Maire; Mém. Soc. Sci. Nat. Maroc 21-22: 15 (1929)
= Stachys arenaria var. maroccana Batt.; Contr. Fl. Atl. 73 (1919)
= Stachys arenaria var. peduncularis (Pau) Maire; Cat. Pl. Maroc 4: 1112 (1941)
= Stachys arenaria var. villosissima Andr.; Ind. Horti Bot. Univ. Budapest 5: 22 (1941)
= Stachys diffusa Schweigg.; Enum. Pl. Hort. Regiom. 69 (1812)
= Stachys hirta Georgi; Beschr. Nation. Russ. Reich. 3(5): 1086 (1800), nom. illeg.
= Stachys macedonica D.Don ex Griseb.; Spic. Fl. Rumel. 2: 143 (1844)
= Stachys mollis Willd. ex Benth.; Linnaea 11: 335 (1837)
= Stachys peduncularis Pau; Cavanillesia 8: 111 (1937)
= Stachys prostrata Lag. ex DC.; Mém. Soc. Phys. Genève 1(2): 455 (1823)
= Stachys pseudoarenaria Sennen; Diagn. Nouv. 199 (1936)
= Stachys pseudoarenaria var. angustifolia Sennen; Diagn. Nouv. 199 (1936)
= Stachys purpurea Poir.; Encycl. [J.Lamarck et al.], Suppl. 5: 227 (1817)
= Stachys rubra Donn.Sm.; Hort. Cantabrig., ed. 4: 136 (1807)
= Stachys scordifolia Willd.; Enum. Pl. Hort. Berol. [Willdenow] 616 (1809)
= Stachys scordioides Desf.; Tabl. École Bot. 219 (1804)
species :
Stachys arenariiformis
Rouy; Bull. Soc. Bot. France 37: 166 (1890)
Distribution: Romania; Bulgaria
13 BUL ROM
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys recta subsp. arenariiformis (Rouy) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)
species :
Stachys argillicola
Sebsebe; Kew Bull. 48(2): 333 (1993)
subspecies :
subsp. argillicola
Distribution: Ethiopia; Kenya
24 ETH 25 KEN
subspecies :
subsp. barbisina
Sebsebe; Kew Bull. 48(2): 334 (1993)
Distribution: Ethiopia; Uganda; Kenya; Tanzania
24 ETH 25 KEN TAN UGA
species :
Stachys aristata
Greenm.; Proc. Amer. Acad. Arts 39: 87 (1904)
Distribution: Mexico (Jalisco, Sinaloa)
79 MEX
species :
Stachys arrecta
L.H.Bailey; Gentes Herbarum 1: 43 (1920)
Distribution: China (Anhui, Henan, Hubei, Hunan, Jiangsu, Shaanxi, Shanxi, Zhejiang)
36 CHI
species :
Stachys arriagana
B.L.Turner; Phytologia 77(4): 350 (1994 publ. 1995)
Distribution: Mexico (San Luis Potosi)
79 MEX
species :
Stachys arvensis
(L.) L.; Sp. Pl. [Linnaeus], ed. 2: 814 (1763)
Name: Acker-Ziest (DE); Field Woundwort (EN); Hierba de gato (ES)
Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Sweden; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, +Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Austria; Poland; Slovakia; Portugal; Spain; Gibraltar; Baleares; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Sardinia; Italy; Sicily; Pantelleria; Slovenia; Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Greece (rare lowlands near coast, Kiklades, Kerkyra, Ionian Isl.); Crete; European Russia; Morocco; Algeria; Tunisia; Azores (Santa Maria Isl., Sao Miguel Isl., Terceira, Graciosa, Sao Jorge, Pico, Faial, Flores Isl., Corvo Isl.); Madeira (Madeira Isl., Ilhas Desertas, Porto Santo Isl.); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma); Cape Verde Isl. (Santo Antao Isl., Fogo Isl., Ilha Brava); São Tomé [I]; South Africa [I] (E-Cape Prov. [I]); Ascension Isl. [I]; St. Helena [I]; Mauritius [I]; La Réunion [I]; East Aegaean Isl. (Lesvos, Psara, Ikaria, Samos, Karpathos, Rhodos); Turkey (NW-Anatolia: Bithynia, SSW-Anatolia, SW-Anatolia); Afghanistan (Kandahar); Lebanon (coastal W-Lebanon); Syria (coastal W-Syria); Israel (coastal W-Israel); China (Fujian, Guangdong, Guangxi); Taiwan; India [c]; Sri Lanka [I]; Australia [I] (Western Australia [I], South Australia [I], Queensland [I], New South Wales [I], Victoria [I], Tasmania [I]); Lord Howe Isl. [I]; Norfolk Isl. [I]; New Zealand [I]; New Caledonia [I]; Easter Isl. [I]; Hawaii [I] (Kauai [I], Oahu [I], Molokai [I], Lanai [I], Maui [I], Hawaii Isl. [I]); Canada [I] (British Columbia [I], Nova Scotia [I]); USA [I] (California [I], District of Columbia [I], Louisiana [I], Massachusetts [I], Maine [I], New Jersey [I], New York [I], Oregon [I], Pennsylvania [I], Texas [I], Virginia [I]); Puerto Rico [I]; Guadeloupe [I]; Dominica [I]; Martinique [I]; Colombia [I]; Venezuela [I]; Ecuador [I]; Peru [I]; Bolivia [I]; Brazil [I]; Paraguay [I]; Uruguay [I]; Argentina [I]; Chile [I] (Valparaíso [I], Los Lagos [I]); Juan Fdz. Isl.
10 DEN GRB IRE SWE 11 AUT BGM GER LUX NET POL SLO SWI 12 BAL CIS COR FRA GIB POR SAR SPA 13 ALB CRO GRC ITA KOS KRI MNT NMA PTL SIC SLV SRB 14 RUS 20 ALG MOR TUN 21 AZO CNY CVI MDR 23 sao 27 saf 28 asc sth 29 mau reu 34 AFG EAI ISR LEB SYR TUR 36 CHI 38 TAI 40 ind srl 50 aus lhi nfk 51 nzl 60 nwc 61 eas 63 haw 71 cdn 72 cdn 73 usa 75 usa 76 usa 77 usa 78 usa 81 dom gdl mar pue 82 clm ven 83 bol ecu per 84 bra 85 arg cle JNF par uru
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Cardiaca arvensis (L.) Lam.; Fl. Franç. [Lamarck] 2: 383 (1779)
= Glechoma arvensis L.; Sp. Pl. [Linnaeus] 2: 578 (1753)
= Glechoma belgica L.; Sp. Pl. [Linnaeus] 2: 578 (1753)
= Glechoma marrubiastrum Vill.; Hist. Pl. Dauphiné [Villars] 2: 371 (1787)
= Leonurus arvensis (L.) Durande; Fl. Bourgogne 1: 83 (1782)
= Stachys arvensis f. albiflora H.Lindb.; Acta Soc. Sci. Fenn., Ser. B, Opera Biol. 1(2) (Itin. Medit.): 133 (1932), nom. illeg.
= Stachys arvensis f. stricta (G.Mey.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2416 (1927)
= Stachys arvensis var. albiflora Bolzon; Bull. Soc. Bot. Ital. 1893: 309 (1893)
= Stachys arvensis var. bermudiana Millsp.; Publ. Field Columbian Mus., Bot. Ser. 2: 94 (1900)
= Stachys arvensis var. lamiiformis Domin; Biblioth. Bot. 22(89): 568 (1928)
= Stachys brasiliensis Benth.; Labiat. Gen. Spec. [Bentham] 550 (1834)
= Trixago arvensis (L.) Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 102 (1809)
= Trixago arvensis (L.) Moritzi; Pfl. Schweiz [Moritzi] 312 (1832), isonym
= Trixago colorata C.Presl; Fl. Sicul. [Presl] xxxvii (1826)
= Trixago cordifolia Moench; Methodus [Moench] 398 (1794), nom. illeg.
= Trixago punctata Gilib.; Fl. Lit. Inch. 1: 73 (1782), opus utique oppr.
= Trixella arvensis (L.) Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 135 (1869)
species :
Stachys aspera
Michx.; Fl. Bor.-Amer. [Michaux] 2: 5 (1803)
Name: Hyssopleaf hedgenettle
Distribution: USA (Alabama, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kentucky, Maryland, Maine, Missouri, Mississippi, North Carolina, New Jersey, New York, Ohio, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)
74 USA 75 USA 78 USA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys ambigua (A.Gray) Britton; Mem. Torrey Bot. Club 5: 288 (1894), nom. illeg.
= Stachys aspera var. glabrata Benth.; Labiat. Gen. Spec. [Bentham] 543 (1834)
= Stachys aspera var. tenuiflora C.L.Hitchc.; Trans. Acad. Sci. St. Louis 5: 513 (1891)
= Stachys grayana House; Bull. New York State Mus. Nat. Hist., Nos. 233-4: 60 (1920 publ. 1921)
= Stachys hyssopifolia var. ambigua A.Gray; Syn. Fl. N. Amer. 2: 387 (1878)
= Stachys palustris subsp. aspera (Michx.) Derv.-Sok.; Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 80(3): 89 (1975)
= Stachys palustris var. aspera (Michx.) A.Gray; Manual [A.Gray], ed. 2: 317 (1856)
= Stachys tenuifolia var. aspera (Michx.) Fernald
species :
Stachys asperata
Hedge; Fl. Madagasc., Fam. 175: 28 (1998)
Distribution: Madagascar
29 MDG
species :
Stachys atherocalyx
K.Koch; Linnaea 21: 691 (1849)
Distribution: Germany [I]; Serbia; Kosovo; North Macedonia; Romania; Bulgaria; NE-Greece (S-Thrakia); European Russia; Crimea; ?Northern Caucasus; Georgia [Caucasus]; Armenia; Turkey (NE-Anatolia)
11 ger 13 BUL GRC KOS NMA ROM SRB 14 KRY RUS 33 ARM GEO 34 TUR
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys acanthodonta Klokov; Fl. URSR 9: 648 (1960)
= Stachys linearifolia K.Koch; Linnaea 21: 692 (1849)
= Stachys recta subsp. atherocalyx (K.Koch) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)
= Stachys recta var. sideritidoides Boiss.; Fl. Orient. [Boissier] 4(2): 730 (1879)
= Stachys sideritidoides K.Koch; Linnaea 21: 692 (1849), nom. illeg.
species :
Stachys aucheri
Benth.; Prodr. [A.P. de Candolle] 12: 487 (1848)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Stachys koelzii Rech.f.; Fl. Iranica [Rechinger] 150: 383 (1982)
species :
Stachys aurea
Benth.; Prodr. [A.P. de Candolle] 12: 492 (1848)
Distribution: South Africa (N-Cape Prov., W-Cape Prov.)
27 SAF
Remarks: clade 12 Olisia
Synonyms: hidden
= Betonica heraclea L.; Mant. Pl. 1: 83 (1767)
= Phlomis parvifolia Burch.; Trav. S. Africa 1: 225 (1822), nom. illeg.
= Sideritis plumosa Thunb.; Prodr. Pl. Cap. 2: 95 (1800)
= Stachys hantamensis Vatke; Bot. Zeitung (Berlin) 33: 462 (1875)
= Stachys integrifolia Vahl ex Benth.; Labiat. Gen. Spec. [Bentham] 562 (1834), nom. illeg.
= Stachys teres Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 364 (1910)
species :
Stachys babunensis
Micevski; Prilozi Oddel. Biol. Med. Nauki, Makedonska Akad. Nauk Umet. 13: 30 (1992)
Distribution: North Macedonia
13 NMA
species :
Stachys bakeri
Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 266 (1896)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Stachys debilis Baker; J. Linn. Soc., Bot. 20: 234 (1884) (non Kunth 1817 nec Bornm. 1917)
species :
Stachys balansae
Boiss. & Kotschy; Fl. Orient. [Boissier] 4(2): 722 (1879)
Distribution: Crimea; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, SE-Anatolia); Iraq (NE-Iraq); Iran (N-Iran, Iranian Aserbaijan)
14 KRY 33 ARM GEO 34 IRN IRQ TUR
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Stachys balansae f. latifolia Kusn.; Uchen. Zap. Imp. Yur´evsk. Univ. 6(3): 70 (1898)
= Stachys terekensis Knorring; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 15: 341 (1953)
species :
Stachys balensis
Sebsebe; Kew Bull. 48(2): 331 (1993)
Distribution: Ethiopia
24 ETH
species :
Stachys bayburtensis
R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 279 (1974)
Distribution: Turkey (NE-Anatolia)
34 TUR
species :
Stachys baytopiorum
Kit Tan & Yıldız; Notes Roy. Bot. Gard. Edinburgh 45(3): 450 (1988 publ. 1989)
Distribution: Turkey
34 TUR
species :
Stachys beckeana
Dörfl. & Hayek; Denkschr. Kaiserl. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 94: 186 (1918)
Distribution: Bosnia & Hercegovina
13 BIH
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys hayekii K.Malý; Glasn. Zemaljsk. Muz. Bosni Hercegovini 40: 160 (1920)
= Stachys recta subsp. hayekii (K.Malý) Bjelčić; Posebna Izd. Zemaljsk. Muz. Bosne Hercegovine 3: 18 (1974)
species :
Stachys benthamiana
Boiss.; Fl. Orient. [Boissier] 4(2): 734 (1879) emend. Rech.f., Bot. Jahrb. Syst. 71: 532 (1941)
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
Remarks: clade 12 Olisia
species :
Stachys bergii
G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116(1): 40 (1989 publ. 1990), nom. nov.
Name: Berg´s hedgenettle
Distribution: USA (California)
76 USA
Synonyms: hidden
= Stachys rigida subsp. lanata Epling; Madroño 4: 270 (1938)
species :
Stachys bhattacharjeeae
Ö.Güner; Feddes Repert. 133(4): 258 (2022)
Distribution: Iraq
34 IRQ
species :
Stachys biflora
Hook. & Arn.; Bot. Beechey Voy. 155 (1833)
Distribution: Mexico (Chihuahua, Coahuila, Jalisco, Michoacán, Nayarit, Nuevo León, San Luis Potosi, Veracruz)
79 MEX
Synonyms: hidden
= Stachys agraria var. glabrior Benth.; Prodr. [A.P. de Candolle] 12: 479 (1848)
= Stachys grahamii Benth.; Labiat. Gen. Spec. [Bentham] 551 (1834)
= Stachys latipes Greenm.; Proc. Amer. Acad. Arts 39: 87 (1904)
species :
Stachys bigelovii
A.Gray; Proc. Amer. Acad. Arts 8: 371 (1872)
Name: Rock hedgenettle
Distribution: USA (Texas); Mexico (Baja California Sur, Chihuahua, Coahuila, Colima, Durango, Jalisco, Michoacán, Nuevo León, San Luis Potosi, Sinaloa, Sonora, Zacatecas)
77 USA 79 MEX
species :
Stachys bithynica
Boiss.; Diagn. Pl. Orient. 5: 28 (1844)
Distribution: European Turkey; Crimea; Transcaucasus; Turkey
13 TUE 14 KRY 34 TUR
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Stachys germanica subsp. bithynica (Boiss.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 276 (1974)
= Stachys germanica var. bithynica (Boiss.) Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)
= Stachys germanica var. procera Nábělek; Spisy Přír. Fak. Masarykovy Univ. 70: 61 (1926)
= Stachys pisidica Boiss. & Heldr.; Diagn. Pl. Orient. 12: 75 (1853)
species :
Stachys bizensis
Schweinf. ex Penz.; Atti Congr. Bot. Int. Genova [1892] 356 (1893), nomen
Distribution: Eritrea
24 ERI
species :
Stachys bogotensis
Kunth; Nov. Gen. Sp. [H.B.K.] 2: 309 (1818)
Distribution: Colombia (Antioquia, Boyacá, Cauca, Cesar, Cundinamarca, Meta, Nariño, Norte de Santander, Santander)
82 CLM
Synonyms: hidden
= Stachys hians Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 116 (1898)
species :
Stachys bolusii
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 343 (1910)
Distribution: South Africa (W-Cape Prov.)
27 SAF
species :
Stachys bombycina
Boiss.; Diagn. Pl. Orient. 12: 79 (1853)
Distribution: Turkey (SSW-Anatolia, SW-Anatolia)
34 TUR
Remarks: clade 3 Distans
Synonyms: hidden
= Stachys pseudobombycina Kaynak, Daskin & Yılmaz; Nordic J. Bot. 28: 341 (2010)
species :
Stachys boraginoides
Schltdl. & Cham.; Linnaea 5: 100 (1830)
Distribution: Mexico (Chiapas, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacán, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Veracruz)
79 MEX
Synonyms: hidden
= Stachys boraginoides var. glandulosa Greenm.; Proc. Amer. Acad. Arts 41: 245 (1905)
species :
Stachys brachiata
Bojer ex Benth.; Labiat. Gen. Spec. [Bentham] 547 (1834)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Stachys madagascariensis Briq.; Bull. Herb. Boissier 2: 137 (1894)
= Stachys madagascariensis var. goudotii Briq.; Bull. Herb. Boissier 2: 140 (1894)
= Stachys rupicola Bojer ex Benth.; Prodr. [A.P. de Candolle] 12: 477 (1848)
species :
Stachys brachyclada
de Noé ex Coss.; Ann. Sci. Nat., Bot., Sér. 4, 1: 226 (1854), et in Bull. Soc. Bot. France 2: 583 (1855)
Distribution: E-Spain; Baleares; France; N-Morocco (Rif); Algeria
12 BAL FRA SPA 20 ALG MOR
Synonyms: hidden
= Stachys brachyclada var. immaculata Maire & Wilczek; Bull. Soc. Hist. Nat. Afrique N. 32: 214 (1941)
= Stachys brachyclada var. punctata Maire; Bull. Soc. Hist. Nat. Afrique N. 26: 74 (1936)
= Stachys hirta var. parviflora de Noé ex Bory & Durieu; Expl. Sci. Algérie 1: t. 65 (1848)
species :
Stachys brahuicus
(Boiss.) comb.ined.
Distribution: Iran (EC-Iran, E-Iran, NE-Iran: Mts., SW-Iran); Afghanistan (Balkh, Farah, Herat, Kabul, Kandahar, Orozgan / Daykundi, Samangan); Pakistan (Baluchistan, Kalat)
34 AFG IRN 40 PAK
Remarks: clade 12 Olisia
Synonyms: hidden
= Chamaesphacos afghanicus Aitch. & Hemsl.; Trans. Linn. Soc. London, Bot., Ser. 2, 3: 97 (1886)
= Chamaesphacos brahuicus (Boiss.) Aitch. & Hemsl.; Trans. Linn. Soc. London, Bot., Ser. 2, 3: 97 (1886)
= Thuspeinanta brahuica (Boiss.) Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 229 (1896)
= Tapeinanthus brahuicus Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 29 (1859)
species :
Stachys bridgesii
Benth; Labiat. Gen. Spec. [Bentham] 546 (1834)
Name: Hierba de Santa Rosa [Chile]
Distribution: Chile (Coquimbo, Metropolitana de Santiago, O´Higgins, Maule, Los Rios, Los Lagos)
85 CLE
species :
Stachys bullata
Benth.; Labiat. Gen. Spec. [Bentham] 547 (1833)
Name: California hedgenettle
Distribution: USA (California)
76 USA
Synonyms: hidden
= Stachys californica Benth.; Prodr. [A.P. de Candolle] 12: 469 (1848)
= Stachys acuminata Greene; Bull. Calif. Acad. Sci. 2(7): 410 (1887)
species :
Stachys burchelliana
Launert; Mitt. Bot. Staatssamml. München 7: 301 (1968)
Distribution: Namibia; South Africa (N-Cape Prov.)
27 NAM SAF
Remarks: clade 12 Olisia
Synonyms: hidden
= Phlomis micrantha Burch.; Trav. S. Africa 1: 340 (1822)
= Sideritis rugosa Thunb.; Prodr. Pl. Cap. 2: 95 (1800)
= Stachys burchellii Benth.; Labiat. Gen. Spec. [Bentham] 561 (1834), nom. illeg.
species :
Stachys burgsdorffioides
(Benth.) Boiss.; Diagn. Pl. Orient. 12: 85 (1853)
subspecies :
subsp. burgsdorffioides
Distribution: Turkey (E-Anatolia, Inner Anatolia, S-Anatolia, SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); Iraq (NW-Iraq); Syria (NW-Syria)
34 IRQ SYR TUR
Synonyms: hidden
= Stachys satureoides var. burgsdorffioides Benth.; Prodr. [A.P. de Candolle] 12: 481 (1848)
subspecies :
subsp. ladanoides
Hand.-Mazz.; Ann. K.K. Naturhist. Hofmus. 27: 417 (1913)
Distribution: Turkey
34 TUR
species :
Stachys buttleri
R.R.Mill; Notes Roy. Bot. Gard. Edinburgh 38: 57 (1980)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Stachys byzantina
K.Koch; Linnaea 21: 686 (1849)
Name: Wolliger Ziest (DE), Lamb´s-ear, Woolly Hedgenettle (EN)
Distribution: England [I] (England [I], Wales [I], Isle of Man [I], Scotland [I], Isles of Scilly [I]); Ireland [I] (Ireland [I], Northern Ireland [I]); Germany [I]; Slovakia [I]; Hungary [I]; Spain [I]; France [I]; Monaco [c]; Channel Isl. [I]; Slovenia [I]; Croatia [I]; European Turkey; European Russia; Siberia [I] (Altai [I]); ?Northern Caucasus; Georgia [Caucasus]; Turkey (Inner Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan); China [I]; Australia [I] (South Australia [I], Queensland [I]); Lord Howe Isl. [I]; New Zealand [I]; Canada [I] (British Columbia [I], Ontario [I], Québec [I]); USA [I] (California [I], Connecticut [I], Illinois [I], New York [I], Utah [I], Virginia [I], Vermont [I]); Peru [I]
10 grb ire 11 ger hun slo 12 cis fra mco spa 13 cro slv TUE 14 RUS 30 sib 33 GEO 34 IRN TUR 36 chi 50 aus lhi 51 nzl 71 cdn 72 cdn 74 usa 75 usa 76 usa 78 usa 83 per
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Eriostomum lanatum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)
= Stachys germanica subsp. lanata (Jacq.) Bonnier; Fl. Ill. France 16 (1927)
= Stachys lanata Jacq.; Icon. Pl. Rar. [Jacquin] 1: 11 (1781) non Crantz (1769)
= Stachys lanata var. parvifolia K.Koch; Linnaea 21: 686 (1849)
= Stachys olympica Poir.; Encycl. [J.Lamarck et al.], Suppl. 5: 226 (1817)
= Stachys taurica Zefir.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 14: 348 (1951)
species :
Stachys afra
E.Mey. ex Benth.; Prodr. [A.P. de Candolle] 12: 495 (1848)
Distribution: South Africa (Limpopo, Gauteng, Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.)
27 SAF
species :
Stachys calcicola
Epling; Bull. Torrey Bot. Club 71: 484 (1944)
Distribution: Guatemala; Colombia (Boyacá)
80 GUA 82 CLM
species :
Stachys candida
Bory & Chaub.; Expéd. Sci. Morée, Bot. 3(2): 167 (1832)
Distribution: S-Greece (Mt. Taygetos and surroundings)
13 GRC
Remarks: clade 11 Swainsoniana, candida agg.
species :
Stachys canescens
Bory & Chaub.; Expéd. Sci. Morée, Bot. 3(2): 167 (1832)
Distribution: Albania; S-Greece (S-Peloponnisos: SW-Messinia, Mani peninsula)
13 ALB GRC
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys messeniaca Boiss.; Prodr. [A.P. de Candolle] 12: 475 (1848)
= Stachys messeniaca var. canescens (Bory & Chaub.) Nyman; Consp. Fl. Eur. 578 (1881)
species :
Stachys carduchorum
(R.Bhattacharjee) Rech.f.; Fl. Iranica [Rechinger] 150: 363 (1982)
Distribution: Turkey; Iran
34 IRN TUR
Synonyms: hidden
= Stachys balansae subsp. carduchorum R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 277 (1974)
species :
Stachys caroliniana
J.B.Nelson & D.A.Rayner; J. Bot. Res. Inst. Texas 8(2): 431 (2014)
Distribution: USA (South Carolina)
78 USA
species :
Stachys cataonica
R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 280 (1974)
Distribution: Turkey (Inner Anatolia)
34 TUR
species :
Stachys chamissonis
Benth.; Linnaea 6: 80 (1831)
Name: Coastal hedgenettle
Distribution: USA (California)
76 USA
Synonyms: hidden
= Stachys flaccida Eastw.; Bull. Torrey Bot. Club 29: 80 (1902)
variant :
var. cooleyae
(A.Heller) G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116: 41 (1989 publ. 1990)
Name: Cooleys Hedgenettle
Distribution: Alaska; Canada (British Columbia); USA (California, Oregon, Washington State)
70 ASK 71 CDN 73 USA 76 USA
Synonyms: hidden
= Stachys caurina Piper; Proc. Biol. Soc. Washington 32: 42 (1919)
= Stachys confertiflora Piper; Proc. Biol. Soc. Washington 32: 42 (1919)
= Stachys cooleyae A.Heller; Bull. Torrey Bot. Club 26: 590 (1899)
species :
Stachys chasmosericea
Ayaşlıgil & Davis; Notes Roy. Bot. Gard. Edinburgh 42: 72, f. 1A-C (1984)
Distribution: Turkey (SW-Anatolia)
34 TUR
species :
Stachys chinensis
Bunge ex Benth.; Labiat. Gen. Spec. [Bentham] 544 (1834)
Distribution: Russian Far East (Primorye); China (Gansu, Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi)
31 RFE 36 CHI
Synonyms: hidden
= Stachys aspera subsp. chinensis (Bunge ex Benth.) Krestovsk.; Turczaninowia 7(4): 20 (2004)
= Stachys aspera var. chinensis (Bunge ex Benth.) Maxim.; Bull. Soc. Imp. Naturalistes Moscou 54(1): 45 (1879)
= Stachys baicalensis var. chinensis (Bunge ex Benth.) Kom.; Trudy Imp. S.-Peterburgsk. Bot. Sada 25: 370 (1907)
= Stachys chanetii H.Lév.; Repert. Spec. Nov. Regni Veg. 9: 222 (1911)
= Stachys chinensis var. albiflora C.Y.Li; Fl. Pl. Herb. Chin. Bor.-Or. 7: 256 (1981)
species :
Stachys choruhensis
Kit Tan & Sorger; Pl. Syst. Evol. 155(1-4): 99 (1987)
Distribution: NE-Turkey
34 TUR
species :
Stachys chrysantha
Boiss. & Heldr.; Diagn. Pl. Orient. 1(7): 56 (1846)
Distribution: S-Greece (NC- to SE-Peloponnisos)
13 GRC
Remarks: clade 11 Swainsoniana, candida agg.
Synonyms: hidden
= Stachys candida subsp. chrysantha (Boiss. & Heldr.) Nyman; Consp. Fl. Eur. 580 (1881)
species :
Stachys circinata
L´Hér.; Stirp. Nov. 51 (1786)
Distribution: S-Spain; Gibraltar; Morocco; Algeria; Tunisia
12 GIB SPA 20 ALG MOR TUN
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys circinata f. hozmariensis Pau; Mem. Real Soc. Esp. Hist. Nat. 12: 376 (1924)
= Stachys circinata subsp. numidica (Pomel) Batt.; Fl. Algérie [Battandier et al.] Dicot. 704 (1890)
= Stachys circinata var. zaiana (Emb. & Maire) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 102(11): 1532 (2017)
= Stachys citrina subsp. zaiana Emb. & Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 208 (1932)
= Stachys erioleuca Pomel; Nouv. Mat. Fl. Atl. 305 (1875)
= Stachys latifolia Aiton; Hort. Kew. [W.Aiton] 2: 303 (1789)
= Stachys numidica Pomel; Nouv. Mat. Fl. Atl. 117 (1874)
= Stachys pallida Salisb.; Prodr. Stirp. Chap. Allerton 63 (1796)
= Stachys velutina Willd.; Enum. Pl. [Willdenow], Suppl. 41 (1814)
species :
Stachys citrina
Boiss. & Heldr. ex Benth.; Prodr. [A.P. de Candolle] 12: 490 (1848)
subspecies :
subsp. citrina
Distribution: Turkey (Inner Anatolia, S-Anatolia, SSW-Anatolia)
34 TUR
subspecies :
subsp. chamaesideritis
(Boiss. & Balansa) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 281 (1974)
Distribution: Turkey
34 TUR
Synonyms: hidden
= Stachys chamaesideritis Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 41 (1859)
species :
Stachys clingmanii
Small; Fl. S.E. U.S. [Small] 1032 (1903)
Name: Clingman´s hedgenettle
Distribution: USA (Alabama, Arkansas, District of Columbia, Illinois, Indiana, Kentucky, Maryland, North Carolina, South Carolina, Tennessee, Virginia, Vermont, West Virginia)
74 USA 75 USA 78 USA
species :
Stachys coccinea
Ortega; Nov. Pl. Descr. Dec. 20 (1797)
Name: Scarlet hedgenettle (EN)
Distribution: USA (Arizona, New Mexico, Texas); Mexico (Aguascalientes, Baja California Sur, Chiapas, Chihuahua, Coahuila, Colima, Ciudad de Mexico, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas); Guatemala; Nicaragua
76 USA 77 USA 79 MEX 80 GUA NIC
Remarks: coccinea agg.
Synonyms: hidden
= Stachys cardinalis Kunze; Bot. Zeitung (Berlin) 2: 645 (1844)
= Stachys coccinea Jacq.; Pl. Rar. Hort. Schoenbr. 3: 18 (1798), nom. illeg.
= Stachys jaimehintonii B.L.Turner; Phytologia 76(5): 395 (1994) [fide Cornejo et al. 2017]
= Stachys limitanea A.Nelson; Amer. J. Bot. 25: 115 (1938)
= Stachys manantlanensis B.L.Turner; Phytologia 78(3): 209 (1995) [fide Cornejo et al. 2017]
= Stachys oaxacana Fernald; Proc. Amer. Acad. Arts 35: 564 (1900)
= Stachys pacifica B.L.Turner; Phytologia 76(5): 396 (1994) [fide Cornejo et al. 2017]
= Stachys torresii B.L.Turner; Phytologia 76(5): 397 (1994) [fide Cornejo et al. 2017]
species :
Stachys collina
Brandegee; Univ. Calif. Publ. Bot. 3: 391 (1909)
Distribution: Mexico (Guerrero, Oaxaca, Puebla)
79 MEX
species :
Stachys comosa
Codd; Bothalia 16(1): 51 (1986)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)
27 SAF
species :
Stachys cordata
Riddell; W. J. Med. Phys. Sci. 9(36): 582 (1836)
Name: Heartleaf hedgenettle
Distribution: USA (Alabama, Arkansas, Georgia, Illinois, Indiana, Kentucky, Maryland, North Carolina, New Jersey, New York, Ohio, Pennsylvania, South Carolina, Tennessee, Virginia, West Virginia)
74 USA 75 USA 78 USA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys nuttallii Shuttlew. ex Benth.; Prodr. [A.P. de Candolle] 12: 469 (1848)
= Stachys nuttallii var. occidentalis Benth.; Prodr. [A.P. de Candolle] 12: 469 (1848)
= Stachys palustris var. cordata A.Gray; Manual [A.Gray], ed. 2: 317 (1856)
= Stachys riddellii House; Bull. New York State Mus. Nat. Hist. 233-234: 60 (1920 publ. 1921), nom. illeg.
= Stachys salvioides Small; Fl. S.E. U.S. [Small] 1032 (1903)
= Stachys subcordata Rydb.; Torreya 27: 88 (1927)
species :
Stachys cordifolia
Prain; J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 59: 310 (1890 publ. 1891)
Distribution: Myanmar
41 MYA
species :
Stachys corsica
Pers.; Syn. Pl. [Persoon] 2: 124 (1806)
Distribution: Corsica; Sardinia
12 COR SAR
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Glechoma grandiflora DC.; Fl. Franç. [de Candolle & Lamarck], éd. 3, 3: 538 (1805)
species :
Stachys costaricensis
Briq. ex T.Durand & Pittier; Bull. Soc. Roy. Bot. Belgique 30: 240 (1891 publ. 1892)
Distribution: Guatemala; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Colombia (Antioquia)
80 COS ELS GUA HON NIC PAN 82 CLM
Synonyms: hidden
= Stachys guatemalensis Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 34 (1934)
species :
Stachys crenata
Raf.; Fl. Ludov. 42 (1817)
Name: Mousesear
Distribution: USA (Alabama, Arizona, Florida, Kentucky, Louisiana, Mississippi, Texas); Mexico (Coahuila)
76 USA 77 USA 78 USA 79 MEX
species :
Stachys cretica
L.; Sp. Pl. [Linnaeus] 2: 581 (1753)
subspecies :
subsp. cretica
Distribution: Germany [I]; Greece (C- & S-Greece); Crete (widespread)
11 ger 13 GRC KRI
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Eriostomum creticum (L.) Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)
= Stachys albereana Neyraut & Debeaux; Rev. Bot. Bull. Mens. 10: 263 (1891)
= Stachys altissima Desf.; Tabl. École Bot., ed. 3 (Cat. Pl. Horti Paris.) 101 (1829)
= Stachys pungens Banks & Sol.; Nat. Hist. Aleppo, ed. 2 [Russell]: 2: 255 (1794)
subspecies :
subsp. anatolica
Rech.f.; Ann. Naturhist. Mus. Wien 48: 175 (1937)
Distribution: Turkey
34 TUR
subspecies :
subsp. bulgarica
Rech.f.; Ann. Naturhist. Mus. Wien 48: 171 (1937)
Distribution: Bulgaria; European Turkey; Greece (scattered N-Greece); Turkey
13 BUL GRC TUE 34 TUR
Synonyms: hidden
= Stachys thracica Davidov; Spis. Bulg. Akad. Nauk 12: 109 (1915)
subspecies :
subsp. cassia
(Boiss.) Rech.f.; Bot. Jahrb. Syst. 71: 534 (1941)
Distribution: Croatia; ?Montenegro; North Macedonia; Albania; Bulgaria; Greece (N-Greece, Thasos, Samothraki, Chalkidiki); Turkey
13 ALB BUL CRO GRC NMA 34 TUR
Synonyms: hidden
= Sideritis cassia Boiss.; Diagn. Pl. Orient. 12: 76 (1853)
= Stachys cassia (Boiss.) Boiss.; Diagn. Pl. Orient. 13: 74 (1854)
= Stachys germanica var. cassia (Boiss.) Briq.; Lab. Alp. Mar. 227 (1893)
subspecies :
subsp. garana
(Boiss.) Rech.f.; Ann. Naturhist. Mus. Wien 48: 176 (1937)
Distribution: Turkey; Iraq; Iran
34 IRN IRQ TUR
Synonyms: hidden
= Stachys garana Boiss.; Diagn. Pl. Orient. 12: 76 (1853)
= Stachys cretica var. garana (Boiss.) Boiss.; Fl. Orient. [Boissier] 4(2): 749 (1879)
= Stachys germanica var. garana (Boiss.) Briq.; Lab. Alp. Mar. 223 (1893)
subspecies :
subsp. gulendamiana
Akçiçek; Phytotaxa 539(3): 258 (2022)
Distribution: Turkey (W-Anatolia)
34 TUR
subspecies :
subsp. kutahyensis
Akçiçek; Turk. J. Bot. 34(2): 132 (2010)
Distribution: Turkey
34 TUR
subspecies :
subsp. lesbiaca
Rech.f.; Ann. Naturhist. Mus. Wien 48: 175 (1937)
Distribution: NE-Greece (Samothraki, Limnos); East Aegaean Isl. (Lesvos, Chios, Ikaria); Turkey (W-Anatolia)
13 GRC 34 EAI TUR
subspecies :
subsp. mersinaea
(Boiss.) Rech.f.; Ann. Naturhist. Mus. Wien 48: 175 (1937)
Distribution: Turkey
34 TUR
Synonyms: hidden
= Stachys mersinaea Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 37 (1859)
= Stachys germanica var. mersinaea (Boiss.) Briq.; Lab. Alp. Mar. 223 (1893)
= Stachys cretica var. paniculata Boiss.; Fl. Orient. [Boissier] 4(2): 719 (1879)
subspecies :
subsp. salviifolia
(Ten.) Rech.f.; Ann. Naturhist. Mus. Wien 48: 170 (1937)
Name: Salbeiblättriger Kretischer Ziest (DE)
Distribution: Germany [I]; France; Italy; Sicily; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (Kerkyra, Lefkada, Epirus, N-Peloponnisos, Chalkidiki, Mt. Falakron)
11 ger 12 FRA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA SIC SLV SMA SRB
Synonyms: hidden
= Eriostomum salviifolium (Ten.) C.Presl; Fl. Sicul. [Presl] 37 (1826)
= Stachys germanica subsp. italica (Briq.) P.Fourn.; Quatre Fl. France 832 (1937)
= Stachys germanica subsp. salviifolia (Ten.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2427 (1927)
= Stachys germanica var. boissieri Briq.; Lab. Alp. Mar. 222 (1893)
= Stachys germanica var. italica Briq.; Lab. Alp. Mar. 224 (1893)
= Stachys salviifolia Ten.; Fl. Napol. 34 (1811)
subspecies :
subsp. smyrnaea
Rech.f.; Ann. Naturhist. Mus. Wien 48: 174 (1937)
Name: Smyrna-Ziest (DE); Smyrna Woundwort (EN)
Distribution: European Turkey; East Aegaean Isl. (Chios, Ikaria, Samos, Lipsi, Leros, Kalimnos, Kos, Tilos, Simi, Rhodos); Turkey (widespread); Cyprus
13 TUE 34 CYP EAI TUR
subspecies :
subsp. trapezuntica
Rech.f.; Ann. Naturhist. Mus. Wien 48: 172 (1937)
Distribution: NE-Turkey
34 TUR
subspecies :
subsp. vacillans
Rech.f.; Bot. Jahrb. Syst. 71: 536 (1941)
Distribution: Turkey; Lebanon; Syria; Israel; Jordania
34 ISR JOR LEB SYR TUR
subspecies :
subsp. velata
(Klokov) Greuter & Burdet; Willdenowia 15(1): 78 (1985)
Distribution: Crimea; Georgia [Caucasus]; Armenia
14 KRY 33 ARM GEO
Synonyms: hidden
= Stachys velata Klokov; Fl. URSR 9: 646 (1960)
species :
Stachys cudiensis
Fırat & Ö.Güner; Phytotaxa 570(1): 45 (2022)
Distribution: Turkey
34 TUR
species :
Stachys cuhacioglui
Yıldırım & Ö.Güner; Türl. Hab. 4(2): 100 (2023)
Distribution: Turkey
34 TUR
species :
Stachys cuneata
Banks ex Benth.; Labiat. Gen. Spec. [Bentham] 560 (1833)
Distribution: South Africa (N-Cape Prov., W-Cape Prov.)
27 SAF
Synonyms: hidden
= Stachys denticulata Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 560 (1834)
species :
Stachys cydni
Kotschy ex Gemici & Leblebici; Turk. J. Bot. 22(5): 359 (1998)
Distribution: Turkey
34 TUR
species :
Stachys cymbalaria
Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1088 (1903)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)
27 SAF
species :
Stachys darcyana
A.Pool; Novon 17(1): 61 (2007)
Distribution: Costa Rica; Panama
80 COS PAN
species :
Stachys debilis
Kunth; Nov. Gen. Sp. [H.B.K.] 2: 309 (1818)
Distribution: Colombia (Boyacá, Nariño); Ecuador
82 CLM 83 ECU
Remarks: clade 2 Stachys s. str.
species :
Stachys didymantha
Brenan; Mem. New York Bot. Gard. 9: 53 (1954)
Distribution: Malawi
26 MLW
species :
Stachys dinteri
Launert; Mitt. Bot. Staatssamml. München 2: 313 (1957)
Distribution: Namibia
27 NAM
species :
Stachys distans
Benth.; Prodr. [A.P. de Candolle] 12: 472 (1848)
Distribution: Turkey (SSW-Anatolia); Lebanon (Antilebanon, C-Lebanon, coastal W-Lebanon); Israel (coastal W-Israel, N-Israel)
34 ISR LEB TUR
Remarks: clade 3 Distans
Synonyms: hidden
= Stachys teucriifolia Boiss. & Kotschy; Diagn. Pl. Orient., Sér. 2, 4: 40 (1859)
variant :
var. cilicica
R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33(2): 287 (1974)
Distribution: Turkey; Lebanon; Syria
34 LEB SYR TUR
species :
Stachys diversifolia
Boiss.; Diagn. Pl. Orient. 12: 80 (1853)
Distribution: Turkey (S-Anatolia); Lebanon (coastal W-Lebanon); Syria (W-Syrian Mountains)
34 LEB SYR TUR
species :
Stachys dregeana
Benth.; Comm. Pl. Afr. Austr. [Meyer] 240 (1838)
Distribution: South Africa (Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho
27 LES SAF
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys dregeana var. lasiocalyx (Schltr.) Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 362 (1910)
= Stachys dregeana var. tenuior Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 362 (1910)
= Stachys foliosa Benth.; Comm. Pl. Afr. Austr. [Meyer] 241 (1838)
= Stachys lasiocalyx Schltr.; J. Bot. 36: 317 (1898)
= Stachys rugosa var. foliosa (Benth.) Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 359 (1910)
species :
Stachys drummondii
Benth.; Labiat. Gen. Spec. [Bentham] 551 (1833)
Name: Drummond´s hedgenettle
Distribution: USA (Texas); Mexico (Morelos, Nuevo León, Puebla, San Luis Potosi, Tamaulipas)
77 USA 79 MEX
species :
Stachys durandiana
Coss.; Bull. Soc. Bot. France 20: 256 (1873)
Distribution: Morocco
20 MOR
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Sideritis grantii Batt.; Contr. Fl. Atl. 67 (1919)
= Stachys durandiana var. grantii (Batt.) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 42 (1979)
= Stachys grantii (Batt.) Batt.; Bull. Soc. Hist. Nat. Afrique N. 13: 33 (1922)
species :
Stachys duriaei
de Noé; Bull. Soc. Bot. France 2: 583 (1855)
Distribution: Algeria; Tunisia
20 ALG TUN
Synonyms: hidden
= Stachys argilacea Pomel; Nouv. Mat. Fl. Atl. 118 (1874)
= Stachys duriaei var. argilacea (Pomel) Batt.; Fl. Algérie [Battandier et al.] Dicot. 707 (1890)
= Stachys duriaei var. ochroleuca de Noé; Bull. Soc. Bot. France 2: 583 (1855)
= Stachys duriaei var. purpurea Batt.; Bull. Soc. Bot. France 33: 356 (1886)
= Stachys ochroleuca (de Noé) Pomel; Nouv. Mat. Fl. Atl. 119 (1874), nom. illeg.
= Stachys pulchra Pomel; Nouv. Mat. Fl. Atl. 119 (1874)
species :
Stachys ehrenbergii
Boiss.; Fl. Orient. [Boissier] 4(2): 721 (1879)
Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon)
34 LEB TUR
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Leonurus mollis Ehrenb. ex Boiss.; Fl. Orient. [Boissier] 4(2): 721 (1879)
= Stachys germanica var. ehrenbergii (Boiss.) Briq.; Lab. Alp. Mar. 233 (1893)
species :
Stachys elliptica
Kunth; Nov. Gen. Sp. [H.B.K.] 2: 308 (1818)
Distribution: Colombia (Arauca, Boyacá, Cundinamarca, Meta, Norte de Santander, Santander); Ecuador
82 CLM 83 ECU
Synonyms: hidden
= Stachys elliptica var. aristata Wedd.; Chlor. Andina 2: 153 (1860)
= Stachys elliptica var. grandiflora Wedd.; Chlor. Andina 2: 153 (1860)
species :
Stachys eplingii
J.B.Nelson; Brittonia 31(4): 492 (1979)
Name: Epling´s hedgenettle
Distribution: USA (Alabama, Arkansas, District of Columbia, Georgia, Kentucky, North Carolina, Oklahoma, South Carolina, Tennessee, Virginia, West Virginia)
74 USA 75 USA 78 USA
species :
Stachys erectiuscula
Gürke; Bot. Jahrb. Syst. 28: 315 (1900)
Distribution: South Africa (Mpulamanga, KwaZulu-Natal)
27 SAF
Synonyms: hidden
= Stachys erectiuscula var. natalensis Skan
species :
Stachys eremicola
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 30 (1934), in Repert. Spec. Nov. Regni Veg., Beih. 85: 10 (1935)
Distribution: Chile (Antofagasta, Atacama, Coquimbo)
85 CLE
Synonyms: hidden
= Stachys crenata Phil.; Anales Univ. Chile 90: 562 (1895)
species :
Stachys eriantha
Benth.; Labiat. Gen. Spec. [Bentham] 549 (1833)
Distribution: Mexico (Ciudad de Mexico, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tlaxcala, Veracruz); Colombia (Boyacá, Cauca, Cundinamarca, Nariño, Norte de Santander, Santander); Venezuela (Tachira); Ecuador
79 MEX 82 CLM VEN 83 ECU
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys elliptica var. repens Wedd.; Chlor. Andina 2: 153 (1860)
= Stachys repens M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 194 (1844), nom. illeg.
species :
Stachys euadenia
P.H.Davis; Kew Bull. [6]: 110 (1951)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Stachys euboica
Rech.f.; Bot. Jahrb. Syst. 80: 391 (1961)
Distribution: Greece (S-Evvoia)
13 GRC
Remarks: provisionally accepted name, of uncertain status
species :
Stachys fendleri
Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 115 (1898)
Distribution: Colombia (Cesar, Norte de Santander); Venezuela (Aragua, Venezuela Distrito Federal, Merida, Miranda, Tachira, Zulia)
82 CLM VEN
species :
Stachys filifolia
Hedge; Fl. Madagasc., Fam. 175: 37 (1998)
Distribution: Madagascar
29 MDG
species :
Stachys flavescens
Benth.; Comm. Pl. Afr. Austr. [Meyer] 241 (1838)
Distribution: South Africa (N-Cape Prov., W-Cape Prov.)
27 SAF
Synonyms: hidden
= Stachys gariepina Benth.; Prodr. [A.P. de Candolle] 12: 493 (1848)
species :
Stachys flexuosa
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 352 (1910)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)
27 SAF
species :
Stachys floccosa
Benth.; Labiat. Gen. Spec. [Bentham] 739 (1835)
Distribution: Pakistan (Kurram, Chitral, Swat, Hazara); NW-India (Assam, Himachal Pradesh, Jammu & Kashmir, Uttarakhand)
40 IND PAK
Synonyms: hidden
= Stachys sericea Wall. ex Benth.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830), nom. illeg., non Cav. 1801
= Stachys vestita Benth.; Labiat. Gen. Spec. [Bentham] 538 (1833)
species :
Stachys floridana
Shuttlew. ex Benth.; Prodr. [A.P. de Candolle] 12: 478 (1848)
Name: Florida hedgenettle
Distribution: USA (Alabama, Arkansas, California, Florida, Georgia, Louisiana, Mississippi, North Carolina, South Carolina, Tennessee, Texas, Virginia)
76 USA 77 USA 78 USA
Remarks: clade 2 Stachys s. str.
species :
Stachys fominii
Sosn. ex Grossh.; Fl. Kavkaza [Grossheim] 3: 313 (1932)
Distribution: Azerbaijan
33 AZE
species :
Stachys fontqueri
Pau; Mem. Real Soc. Esp. Hist. Nat. 12: 377 (1924)
Distribution: Morocco
20 MOR
species :
Stachys forsythii
Hedge; Fl. Madagasc., Fam. 175: 30 (1998)
Distribution: Madagascar
29 MDG
species :
Stachys fragillima
Bornm.; Bull. Herb. Boissier 7: 118 (1899)
Distribution: Iraq (NE-Iraq)
34 IRQ
species :
Stachys fruticulosa
M.Bieb.; Tabl. Prov. Mer Casp. Append. 1: 15 (1798)
Distribution: Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (NE-Anatolia); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, W-Iran)
33 ARM AZE GEO 34 IRN TUR
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys araxina Kapeller; Zametki Sist. Geogr. Rast. 16: 22 (1951)
= Stachys fruticulosa subsp. grossheimii (Kapeller) Menitsky; Bot. Zhurn. (Moscow & Leningrad) 77(6): 70 (1992)
= Stachys fruticulosa var. leptophylla K.Koch; Linnaea 21: 694 (1849)
= Stachys fruticulosa var. macrocheilos (Boiss.) Boiss.; Fl. Orient. [Boissier] 3: 737 (1879)
= Stachys grossheimii Kapeller; Trudy Tbilissk. Bot. Inst. 12: 223 (1948)
= Stachys macrocheilos Boiss.; Diagn. Pl. Orient. 5: 30 (1844)
= Stachys sosnowskyi Kapeller; Zametki Sist. Geogr. Rast. 16: 20 (1951)
species :
Stachys gaziantepensis
Dinc & Dogru; Proc. Natl. Acad. Sci. India, B 86(3): 631-635 (2016)
Distribution: Turkey
34 TUR
species :
Stachys geobombycis
C.Y.Wu; Acta Phytotax. Sin. 10: 222 (1965)
Distribution: China (Fujian, Guangdong, Guangxi, Hubei, Hunan, Jiangxi, Zhejiang)
36 CHI
species :
Stachys germanica
L.; Sp. Pl. [Linnaeus] 2: 581 (1753)
subspecies :
subsp. germanica
Name: Deutscher Ziest (DE), Downy Woundwort (EN)
Distribution: England (England); Belgium; Germany (Brandenburg, Baden-Württemberg, Bayern, Hessen, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein [I], Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; ?Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Baleares; France; Corsica; Sardinia; Italy; Sicily; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (scattered N- & C-Greece, Kerkyra, Epirus, south to S-Evvoia); C-European Russia; Ukraine; Crimea; Morocco; Canary Isl. (Tenerife); Northern Caucasus; Georgia [Caucasus]; Armenia; Turkey (N-Anatolia, NW-Anatolia: Bithynia, SW-Anatolia); Canada [I] (Ontario [I]); USA [I] (Florida [I], New Jersey [I], Ohio [I], Pennsylvania [I], Rhode Island [I], Tennessee [I], Virginia [I], West Virginia [I])
10 GRB 11 AUT BGM CZE GER HUN POL SLO SWI 12 BAL COR FRA POR SAR SPA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SIC SLV SMA SRB TUE 14 KRY RUS UKR 20 MOR 21 CNY 33 ARM GEO NCS 34 TUR 72 cdn 75 usa 78 usa
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Betonica germanica (L.) Walther; Fl. Giessen 453 (1802)
= Eriostomum germanicum (L.) Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)
= Eriostomum lusitanicum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 105 (1809)
= Eriostomum polystachyum (Ten.) C.Presl; Fl. Sicul. [Presl] 37 (1826)
= Leonurus germanica (L.) Gueldenst.; Reise Russland [Gueldenstaedt] 2: 20 (1791)
= Stachys acutifolia Bory & Chaub.; Expéd. Sci. Morée, Bot. 168 (1832), nom. superfl.
= Stachys alba Mill.; Gard. Dict., ed. 8: 4 (1768)
= Stachys argentea Tausch; Flora 14: 217 (1831)
= Stachys biennis Roth; Ann. Bot. [Usteri] 14: 27 (1795)
= Stachys castellana Willk.; Suppl. Prodr. Fl. Hispan. [Willkomm & Lange] 153 (1893)
= Stachys cinerea Benth.; Labiat. Gen. Spec. [Bentham] 536 (1833)
= Stachys cordata Klokov; Fl. URSR 9: 647 (1960), nom. illeg.
= Stachys dasyanthes var. alpina Heldr. & Sart.; Exsicc. (Herb. Norm.) 1851: 347 (1851)
= Stachys elongata Benth.; Labiat. Gen. Spec. [Bentham] 536 (1833)
= Stachys excelsa Benth.; Labiat. Gen. Spec. [Bentham] 536 (1833)
= Stachys fuchsii Bubani; Fl. Pyren. [Bubani] 1: 440 (1897)
= Stachys germanica subsp. cordata Greuter & Burdet; Willdenowia 15: 78 (1985)
= Stachys germanica subsp. cordigera Briq.; Lab. Alp. Mar. 232 (1893)
= Stachys germanica subsp. lusitanica (Hoffmanns. & Link) Cout.; Fl. Portugal [Coutinho] 520 (1913)
= Stachys germanica subsp. polystachya (Ten.) Arcang.; Comp. Fl. Ital. [Arcangeli] 552 (1882)
= Stachys germanica var. alpigena K.Koch; Linnaea 21: 687 (1849)
= Stachys germanica var. alpina (Heldr. & Sart.) Briq.; Lab. Alp. Mar. 228 (1893)
= Stachys germanica var. banatica K.Koch; Linnaea 21: 687 (1849)
= Stachys germanica var. canariensis Font Quer & Svent.; Collect. Bot. (Barcelona) 2: 201 (1949)
= Stachys germanica var. lusitanica (Hoffmanns. & Link) Nyman; Consp. Fl. Eur. Suppl. 2: 251 (1889)
= Stachys germanica var. polystachya (Ten.) Nyman; Consp. Fl. Eur. 577 (1881)
= Stachys germanica var. pubescens Benth.; Edwards´s Bot. Reg. 15: t. 1289 (1829)
= Stachys germanica var. serrata Briq.; Lab. Alp. Mar. 231 (1893)
= Stachys germanica var. sublanata Fleischm. ex Nyman; Consp. Fl. Eur. 577 (1881), nomen
= Stachys germanica var. taurica K.Koch; Linnaea 21: 687 (1849)
= Stachys heraclea Webb ex Ball; J. Linn. Soc., Bot. 16: 624 (1878), nom. inval.
= Stachys heterodonta Zefir.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 14: 349 (1951)
= Stachys heterophylla Moench; Methodus [Moench], Suppl. 135 (1802), nom. illeg.
= Stachys hispanica Mill.; Gard. Dict., ed. 8: 6 (1768)
= Stachys lanata Crantz; Stirp. Austr. Fasc., ed. 2, 2: 257 (1769)
= Stachys lusitanica (Hoffmanns. & Link) Steud.; Nomencl. Bot. [Steudel], ed. 1, 1: 806 (1821)
= Stachys major Garsault; Fig. Pl. Med. t. 569 (1764), Descr. Pl. Anim., 33 (1767), Thell. in Bull. Herb. Boissier, Sér. 2, 8: 906 (1908)
= Stachys nova Sadler ex Rchb.; Fl. Germ. Excurs. 319 (1831)
= Stachys obtusata Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)
= Stachys pannonica Láng ex Ten.; Syll. Pl. Fl. Neapol. 291 (1831)
= Stachys polystachya Ten.; Fl. Napol. 1(Prodr.): XXXIV (1812)
= Stachys tomentosa Gaterau; Descr. Pl. Montauban 107 (1789), nom. illeg.
= Stachys verbascifolia Benth.; Labiat. Gen. Spec. [Bentham] 536 (1834)
subspecies :
subsp. dasyanthes
(Raf.) Arcang.; Comp. Fl. Ital., ed. 2 [Arcangeli]: 437 (1894)
Distribution: Italy; Sicily
13 ITA SIC
Synonyms: hidden
= Stachys dasyanthes Raf.; Précis Découv. Somiol. 39 (1814)
= Stachys germanica var. dasyanthes (Raf.) Briq.; Lab. Alp. Mar. 231 (1893)
= Stachys germanica var. orsiniana Briq.; Lab. Alp. Mar. 227 (1893)
subspecies :
subsp. heldreichii
(Boiss.) Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 285 (1929)
Distribution: Albania; Greece (NW-, NC-, C-Greece, Peloponnisos); Transcaucasus; Turkey
13 ALB GRC 34 TUR
Synonyms: hidden
= Stachys heldreichii Boiss.; Fl. Orient. [Boissier] 4(2): 721 (1879)
subspecies :
subsp. penicillata
(Heldr. & Sart. ex Boiss.) Nyman; Consp. Fl. Eur. 578 (1881)
Distribution: C-Greece (rare Evvoia, Peloponnisos)
13 GRC
Synonyms: hidden
= Stachys germanica var. penicillata (Heldr. & Sart. ex Boiss.) Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)
= Stachys penicillata Heldr. & Sartori ex Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 37 (1859)
subspecies :
subsp. velezensis
(Sagorski) Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 286 (1929)
Distribution: Croatia; Bosnia & Hercegovina
13 BIH CRO
Synonyms: hidden
= Stachys reinertii subsp. velezensis Sagorski; Allg. Bot. Z. Syst. 18: 15 (1912)
species :
Stachys gilliesii
Benth.; Labiat. Gen. Spec. [Bentham] 740 (1833)
Distribution: Ecuador; Peru; Bolivia (Chuquisaca, Santa Cruz, Tarija); S-Brazil (Paraná, Rio Grande do Sul, Santa Catarina); Paraguay (Pres. Hayes); Uruguay (Artigas, Cerro Largo, Flores Prov., Florida Prov., Lavalleja, Paysandu, Rivera, Salto, Soriano, Treinta y Tres); Argentina (Chaco, Cordoba, Corrientes, Formosa, Jujuy, Mendoza, Tucuman); Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío)
83 BOL ECU PER 84 BRA 85 ARG CLE PAR URU
Synonyms: hidden
= Stachys boliviana Briq.; Bull. Herb. Boissier 4: 871 (1896)
= Stachys galanderi Briq.; Bull. Herb. Boissier 4: 870 (1896)
= Stachys lorentzii Briq.; Bull. Herb. Boissier 4: 869 (1896)
= Stachys micrantha Griseb.; Abh. Königl. Ges. Wiss. Göttingen 24: 275 (1879), nom. illeg.
= Stachys petiolosa Briq.; Bull. Herb. Boissier 4: 868 (1896)
= Stachys tucumanensis Briq.; Bull. Herb. Boissier 4: 872 (1896)
species :
Stachys glandulibracteata
Y.B.Harv.; Kew Bull. 51(3): 452 (1996)
Distribution: Tanzania
25 TAN
species :
Stachys glandulosa
Hutch. & E.A.Bruce; Bull. Misc. Inform. Kew 1941: 178 (1941)
Distribution: Somalia
24 SOM
species :
Stachys glandulosissima
Floden; Phytoneuron 2016-53: 2 (2016)
Distribution: USA (Tennessee)
78 USA
species :
Stachys glechomifolia
Nábělek; Spisy Přír. Fak. Masarykovy Univ. 70: 61 (1926)
Distribution: Turkey (SE-Anatolia)
34 TUR
species :
Stachys globosa
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 25 (1934)
Distribution: Mexico (Ciudad de Mexico, Hidalgo, Mexico State, Nayarit, San Luis Potosi, Veracruz)
79 MEX
species :
Stachys glutinosa
L.; Sp. Pl. [Linnaeus] 2: 581 (1753)
Distribution: Corsica; Sardinia; Italy (Capraia Isl.)
12 COR SAR 13 ITA
Remarks: clade 11 Swainsoniana
species :
Stachys gossweileri
G.Taylor; J. Bot. 69(Suppl. 2): 167 (1931)
Distribution: Angola
26 ANG
species :
Stachys goulimyi
Rech.f.; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 94: 25 (1957)
Distribution: NC-Greece (Mt. Askio)
13 GRC
species :
Stachys graciliflora
C.Presl; Abh. Königl. Böhm. Ges. Wiss., Ser. 5, 3: 530 (1845)
Distribution: South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Eswatini [Swaziland]
27 SAF SWZ
Synonyms: hidden
= Stachys cooperi Skan; Bull. Misc. Inform. Kew 1909: 420 (1910)
species :
Stachys graeca
Boiss. & Heldr.; Diagn. Pl. Orient. 12: 77 (1853)
Distribution: Greece (scattered NC-, C-Greece, Peloponnisos, S-Evvoia, Attiki)
13 GRC
Remarks: clade 1 Eriostomum
Synonyms: hidden
= Stachys acutifolia Link; Linnaea 9: 575 (1835), sensu auct.
= Stachys lusitanica Bory & Chaub.; Nouv. Fl. Pélop. 37 (1838), sensu auct.
species :
Stachys grandidentata
Lindl.; Bot. Reg. 13: t. 1080 (1827)
Name: Hierba santa [Chile]
Distribution: Chile (Antofagasta, Atacama, Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Lagos)
85 CLE
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys albicaulis Bertero ex Benth.; Prodr. [A.P. de Candolle] 12: 473 (1848), nom. illeg.
= Stachys closii Phil.; Anales Univ. Chile 90: 561 (1895)
= Stachys elliptica Poepp. ex Benth.; Prodr. [A.P. de Candolle] 12: 473 (1848), nom. illeg.
= Stachys serrata Clos; Fl. Chil. [Gay] 4: 500 (1849)
= Stachys toronjilcillo Phil.; Linnaea 33: 193 (1864)
species :
Stachys grandifolia
E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 239 (1838)
Distribution: South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Lesotho; Eswatini [Swaziland]
27 LES SAF SWZ
Remarks: clade 2 Stachys s. str.
species :
Stachys graveolens
Nábělek; Spisy Přír. Fak. Masarykovy Univ. 70: 62, t. 7, f. 3 et p. 63, f. 16 (1926)
Distribution: Iraq (NW-Iraq)
34 IRQ
species :
Stachys guyoniana
de Noé ex Batt.; Fl. Algérie [Battandier et al.] Dicot. 704 (1890)
Distribution: Algeria
20 ALG
Synonyms: hidden
= Stachys guyoniana de Noé ex Munby; Cat. Pl. Algeria 24 (1890)
species :
Stachys hakkariensis
Akçiçek & Fırat; Phytotaxa 257(2): 168 (2016)
Distribution: Turkey
34 TUR
species :
Stachys hamata
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 49 (1934)
Distribution: Colombia (Nariño, Quindío, Tolima); Venezuela (Sucre); Ecuador
82 CLM VEN 83 ECU
Synonyms: hidden
= Stachys urticoides Epling; Repert. Spec. Nov. Regni Veg., Beih. 85: 12 (1935)
species :
Stachys harkerae
J.G.González; Ann. Bot. Fenn. 52(5-6): 345-351 (2015)
Distribution: Mexico (Jalisco, Zacatecas)
79 MEX
species :
Stachys harleyana
A.Pool; Novon 17(1): 63 (2007)
Distribution: Mexico (Chiapas)
79 MEX
species :
Stachys hebens
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 35 (1934)
Distribution: Colombia (Cundinamarca, Magdalena, Norte de Santander, Santander)
82 CLM
species :
Stachys heraclea
All.; Fl. Pedem. 1: 31 (1785)
Distribution: Spain; France; Corsica; Italy; Sicily; +San Marino
12 COR FRA SPA 13 ITA SIC SMA
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Sideritis heraclea Colebr. ex Benth.; Prodr. [A.P. de Candolle] 12: 447 (1848)
= Stachys barbata Lapeyr.; Hist. Pl. Pyrénées 336 (1813)
= Stachys barbigera Viv.; Fl. Cors. Prodr., App. 2: 4 (1830)
= Stachys betonicifolia Pers.; Syn. Pl. [Persoon] 2: 124 (1806)
= Stachys heraclea subsp. malessana Degen & Hervier; Bull. Acad. Int. Géogr. Bot., Sér. 3, 17: 195 (1907)
= Stachys heraclea var. barbigera (Viv.) Nyman; Consp. Fl. Eur. 578 (1881)
= Stachys phlomoides Willd.; Enum. Pl. Hort. Berol. Suppl. [Willdenow] 41 (1814)
= Stachys valentina Lag.; Varied. Ci. 2(4): 39 (1805)
species :
Stachys herrerae
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 18 (1935), in Repert. Spec. Nov. Regni Veg., Beih. 85: 8 (1935)
Distribution: Peru
83 PER
species :
Stachys herrerana
Rzed. & Calderón; Acta Bot. Mex. 3: 1 (1988)
Distribution: Mexico (Hidalgo, Veracruz)
79 MEX
Conservation Status: known only from type
species :
Stachys hildebrandtii
Vatke; Linnaea 43: 95 (1880 publ. 1882)
Distribution: Somalia; Kenya; Tanzania
24 SOM 25 KEN TAN
Remarks: clade 12 Olisia
species :
Stachys hintoniorum
B.L.Turner; Phytologia 77(4): 358 (1994 publ. 1995)
Distribution: Mexico (Coahuila, Nuevo León)
79 MEX
species :
Stachys hispida
Pursh; Fl. Amer. Sept. [Pursh] 2: 407 (1813)
Distribution: Canada (Manitoba, Ontario, Québec); USA (North Dakota, Georgia, Illinois, Indiana, Iowa, Kentucky, Maine, Maryland, Missouri, New York, Ohio, Tennessee, Vermont, Wisconsin)
71 CDN 72 CDN 74 USA 75 USA 78 USA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys palustris var. hispida (Pursh) B.Boivin; Phytologia 22: 381 (1972), nom. illeg.
= Stachys pilosa subsp. hispida (Pursh) Á.Löve & D.Löve; Taxon 31: 354 (1982)
= Stachys tenuifolia var. hispida (Pursh) Fernald; Rhodora 45: 469 (1943)
species :
Stachys hissarica
Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 614 (1889)
Distribution: Kazakhstan; Uzbekistan; Tajikistan; Afghanistan; Pakistan (Chitral, Swat, Hazara, Murree)
32 KAZ TZK UZB 34 AFG 40 PAK
Synonyms: hidden
= Stachys pseudofloccosa Knorring; Fl. URSS [Komarov et al., eds.] 21: 653 (1954)
species :
Stachys huber-morathii
R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 278 (1974)
Distribution: Turkey (N-Anatolia)
34 TUR
species :
Stachys huetii
Boiss.; Fl. Orient. [Boissier] 4(2): 722 (1879)
Distribution: Turkey (E-Anatolia)
34 TUR
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Stachys brantii A.Huet ex Boiss.; Fl. Orient. [Boissier] 4(2): 722 (1879)
= Stachys germanica var. huetii (Boiss.) Briq.; Lab. Alp. Mar. 228 (1893)
species :
Stachys huillensis
Hiern; Cat. Afr. Pl. [Hiern] 1: 875 (1898)
Distribution: Angola
26 ANG
species :
Stachys humbertii
Hedge; Fl. Madagasc., Fam. 175: 34 (1998)
Distribution: Madagascar
29 MDG
species :
Stachys humifusa
Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 547 (1833)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)
27 SAF
Synonyms: hidden
= Stachys subsessilis Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 548 (1834)
= Stachys tenella Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 358 (1910)
species :
Stachys hydrophila
Boiss.; Diagn. Pl. Orient. 12: 81 (1853) et Fl. 4: 726 (1879)
Distribution: +Algeria; Lebanon (C-Lebanon, coastal W-Lebanon)
20 ALG 34 LEB
species :
Stachys hyssopifolia
Michx.; Fl. Bor.-Amer. [Michaux] 2: 4 (1803)
Name: Hyssopleaf hedgenettle (EN), Ysopblättriger Ziest (DE)
Distribution: Germany [I]; USA (Connecticut, Delaware, Florida, Georgia, Illinois, Indiana, Kentucky, Massachusetts, Maryland, Maine, Michigan, Missouri, North Carolina, New Hampshire, New Jersey, New York, Pennsylvania, Rhode Island, South Carolina, Virginia)
11 ger 74 USA 75 USA 78 USA
Synonyms: hidden
= Stachys atlantica Britton; Man. Fl. N. States [Britton] 792 (1901)
= Stachys hyssopifolia f. oswaldiae Oswald; Phytologia 11: 69 (1964)
= Stachys hyssopifolia var. lythroides (Small) J.B.Nelson; Phytologia 46: 94 (1980)
= Stachys lythroides Small; Fl. S.E. U.S. [Small] 1031 (1903)
species :
Stachys hyssopoides
Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 558 (1833)
Distribution: South Africa (Limpopo, North-West Prov., Gauteng, Mpulamanga, Free State, KwaZulu-Natal, N-Cape Prov., E-Cape Prov.); Lesotho
27 LES SAF
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys coerulea Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 558 (1834)
= Stachys macilenta E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 240 (1838)
species :
Stachys iberica
M.Bieb.; Fl. Taur.-Caucas. 2: 51 (1808)
subspecies :
subsp. iberica
Distribution: Crimea; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, S-Anatolia, SE-Anatolia, SSW-Anatolia); Iraq (NE-Iraq); Iran (Iranian Aserbaijan); Syria (W-Syrian Mountains)
14 KRY 33 ARM GEO 34 IRN IRQ SYR TUR
Synonyms: hidden
= Stachys congesta d´Urv.; Mém. Soc. Linn. Paris 1: 324 (1822)
= Stachys iberica var. congesta (d´Urv.) K.Koch; Linnaea 21: 693 (1849)
= Stachys caucasica K.Koch; Linnaea 21: 693 (1849)
variant :
var. densipilosa
R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 284 (1974)
Distribution: Turkey
34 TUR
variant :
var. georgica
(Rech.f.) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 100(6): 585 (2015)
Distribution: Georgia [Caucasus]; Turkey; Iraq; Iran
33 GEO 34 IRN IRQ TUR
Synonyms: hidden
= Stachys iberica subsp. georgica Rech.f.; Feddes Repert. Spec. Nov. Regni Veg. 53: 84 (1944)
subspecies :
subsp. stenostachya
(Boiss.) Rech.f.; Feddes Repert. Spec. Nov. Regni Veg. 53: 84 (1944)
Distribution: Crimea; Turkey
14 KRY 34 TUR
Synonyms: hidden
= Stachys euphratica Hayek; Repert. Spec. Nov. Regni Veg. 21: 258 (1925)
= Stachys memorabilis Klokov; Fl. URSR 9: 648 (1960)
= Stachys stenostachya Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 41 (1859)
species :
Stachys ilicifolius
(Schrenk) Sennikov; Checkl. Vasc. Pl. Tian-Shan 351 (2021)
Distribution: Kazakhstan; Turkmenistan; Uzbekistan; Tajikistan; Iran (EC-Iran, NE-Iran: Mts., W-Iran); Afghanistan (Badghys, Balkh, Herat, Jawzjan / Sar-e-Pol, Samangan); China (Xinjiang)
32 KAZ TKM TZK UZB 34 AFG IRN 36 CHI
Remarks: clade 12 Olisia
Synonyms: hidden
= Chamaesphacos longiflorus Bornm. & Sint.; Mitt. Thüring. Bot. Vereins, N. F. 18: 51 (1905)
= Chamaesphacos ilicifolius Schrenk ex Fisch. & C.A.Mey.; Enum. Pl. Nov. [Fischer & Meyer] 1: 28 (1841)
species :
Stachys iltisii
J.B.Nelson; J. Bot. Res. Inst. Texas 2(2): 761 (2008)
Distribution: USA (Arkansas, Oklahoma)
74 USA 78 USA
species :
Stachys inanis
Hausskn. & Bornm.; Repert. Spec. Nov. Regni Veg. 22: 188 (1925)
Distribution: Turkey (E-Anatolia, Inner Anatolia)
34 TUR
species :
Stachys inflata
Benth.; Labiat. Gen. Spec. [Bentham] 562 (1834)
subspecies :
subsp. inflata
Distribution: Armenia; Azerbaijan; Turkey (NE-Anatolia); Iraq; Iran (EC-Iran, Iranian Aserbaijan, W-Iran, N-Iran)
33 ARM AZE 34 IRN IRQ TUR
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys inflata subsp. albopannosa Rech.f.; Oesterr. Bot. Z. 99: 47 (1952)
subspecies :
subsp. caucasica
(Stschegl.) Takht.; Fl. Erevana 260 (1972)
Distribution: Transcaucasus; Iran
34 IRN
Synonyms: hidden
= Stachys inflata var. caucasica Stschegl.; Bull. Soc. Imp. Naturalistes Moscou 24(2): 474 (1851)
= Stachys schtschegleevii Sosn. ex Grossh.; Fl. Kavkaza [Grossheim] 3: 314 (1932), nom. nov.
species :
Stachys intermedia
Aiton; Hort. Kew. [W.Aiton] 2: 301 (1789)
Distribution: Northern Caucasus; Transcaucasus
33 NCS
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Stachys germanica var. intermedia (Aiton) Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)
= Stachys sibirica Link; Enum. Hort. Berol. Alt. 2: 109 (1822)
species :
Stachys ionica
Halácsy; Consp. Fl. Graec., Suppl. 85 (1908)
Distribution: W-Greece (Ionian Isl.)
13 GRC
Remarks: clade 11 Swainsoniana
species :
Stachys iraqensis
R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 294 (1974)
Distribution: Iraq (NW-Iraq)
34 IRQ
species :
Stachys istanbulensis
Ö.Güner; Turk. J. Bot. 46(6): 625 (2022)
Distribution: Turkey
34 TUR
species :
Stachys iva
Griseb.; Spic. Fl. Rumel. 2: 143 (1844)
Distribution: North Macedonia; NC-Greece
13 GRC NMA
Synonyms: hidden
= Stachys horvaticii Micevski; Acta Bot. Croat. 28: 452 (1969)
= Stachys iva var. ochroleuca Bornm.; Bot. Jahrb. Syst. 61: 89 (1928)
variant :
var. macedonica
(Micevski) Krestovsk.; Novosti Sist. Vyssh. Rast. 48: 121 (2017)
Distribution: North Macedonia
13 NMA
Synonyms: hidden
= Stachys horvaticii var. macedonica Micevski; Acta Bot. Croat. 28: 453 (1969)
species :
Stachys jijigaensis
Sebsebe; Kew Bull. 48(2): 337 (1993)
Distribution: Ethiopia
24 ETH
species :
Stachys keerlii
Benth.; Labiat. Gen. Spec. [Bentham] 551 (1833)
Distribution: Mexico (Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Tamaulipas, Veracruz)
79 MEX
Synonyms: hidden
= Stachys excelsa M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 194 (1844)
= Stachys inclusa Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 12 (1934)
species :
Stachys kermanshabensis
Rech.f.; Pl. SySt. Evol. 134: 290 (1980)
Distribution: Iran (SW-Iran)
34 IRN
Remarks: clade 12 Olisia
species :
Stachys ketenoglui
Kaynak, Daskin & Yılmaz; Nordic J. Bot. 27(3): 238 (2009)
Distribution: Turkey
34 TUR
species :
Stachys komarovii
Knorring; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 15: 343 (1953)
Distribution: Russian Far East (Primorye)
31 RFE
species :
Stachys kotschyi
Boiss. & Hohen.; Diagn. Pl. Orient. 5: 32 (1844)
Distribution: Iraq (NE-Iraq)
34 IRQ
Synonyms: hidden
= Stachys haussknechtii Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 281 (1929)
species :
Stachys kouyangensis
(Vaniot) Dunn; Notes Roy. Bot. Gard. Edinburgh 8: 167 (1913)
Distribution: China (Guizhou, Hubei, Sichuan, Yunnan); SE-Tibet; Myanmar; Thailand; Vietnam
36 CHI TIB 41 MYA THA VIE
Synonyms: hidden
= Lamium kouyangense Vaniot; Bull. Acad. Int. Géogr. Bot. 14: 175 (1904)
= Scutellaria kouyangensis (Vaniot) Dunn; Notes Roy. Bot. Gard. Edinburgh 8: 167 (1913)
= Stachys franchetiana H.Lév.; Repert. Spec. Nov. Regni Veg. 9: 246 (1911)
= Stachys kouyangensis var. franchetiana (H.Lév.) C.Y.Wu; Acta Phytotax. Sin. 10: 228 (1965)
= Stachys kouyangensis var. leptodon (Dunn) C.Y.Wu; Acta Phytotax. Sin. 10: 227 (1965)
= Stachys kouyangensis var. tuberculata (Hand.-Mazz.) C.Y.Wu; Acta Phytotax. Sin. 10: 227 (1965)
= Stachys kouyangensis var. villosissima C.Y.Wu; Acta Phytotax. Sin. 10: 228 (1965)
= Stachys leptodon Dunn; Notes Roy. Bot. Gard. Edinburgh 8: 167 (1913)
= Stachys sieboldii var. tuberculata Hand.-Mazz.; Acta Horti Gothob. 13: 348 (1939)
species :
Stachys kulalensis
Sebsebe; Kew Bull. 48(1): 115 (1993)
Distribution: Kenya
25 KEN
species :
Stachys kuntzei
Gürke; Revis. Gen. Pl. 3(2): 262 (1898)
Distribution: South Africa (Mpulamanga, Free State, KwaZulu-Natal); Lesotho
27 LES SAF
Synonyms: hidden
= Stachys petrogenes Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1085 (1903)
species :
Stachys kurdica
Boiss. & Hohen.; Diagn. Pl. Orient. 5: 31 (1844)
subspecies :
subsp. kurdica
Distribution: Turkey (E-Anatolia, SE-Anatolia); Iraq (NE-Iraq); Iran (Iranian Aserbaijan, W-Iran)
34 IRN IRQ TUR
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys ballotiformis Vatke; Bot. Zeitung (Berlin) 33: 448 (1875)
= Stachys benthamiana var. clinopodioides Boiss.; Fl. Orient. [Boissier] 4(2): 735 (1879)
= Stachys plebeia Vatke; Bot. Zeitung (Berlin) 33: 449 (1875)
= Stachys subnuda var. kurdica (Boiss. & Hohen.) Boiss.; Fl. Orient. [Boissier] 4(2): 734 (1879)
variant :
var. brevidens
Bornm. ex R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 283 (1974)
Distribution: Turkey; Iraq
34 IRQ TUR
subspecies :
subsp. asterocalyx
(Rech.f.) Salmaki; Bot. J. Linn. Soc. 170(4): 597 (2012)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Stachys asterocalyx Rech.f.; Bot. Jahrb. Syst. 71: 533 (1941)
species :
Stachys lallaniana
R.Kr.Singh & Sanjeet Kumar; J. Biodivers. Conservation 7(3)-41: 42 (2023), nom. nov.
Distribution: South Africa
27 SAF
Remarks: provisionally accepted name
Synonyms: hidden
= Stachys malacophylla Skan; Bull. Misc. Inform. Kew 1909(10): 421 (1909), nom. illeg.
species :
Stachys lamarckii
Benth.; Labiat. Gen. Spec. [Bentham] 562 (1833)
Distribution: Namibia; South Africa (N-Cape Prov.)
27 NAM SAF
Remarks: clade 12 Olisia
Synonyms: hidden
= Sideritis decumbens Thunb.; Prodr. Pl. Cap. 2: 95 (1800)
= Stachys nutans Benth.; Labiat. Gen. Spec. [Bentham] 561 (1834)
= Stachys rugosa Lam.; Tabl. Encycl. 3: 66 (1823), nom. illeg.
species :
Stachys lamioides
Benth.; Pl. Hartw. [Bentham] 244 (1846)
Distribution: Colombia (Antioquia, Caldas, Cauca, Huila, Nariño, Putumayo, Quindío, Risaralda, Tolima, Valle); Ecuador
82 CLM 83 ECU
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys lamioides var. aristulata Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 50 (1934)
species :
Stachys langmaniae
Rzed. & Calderón; Acta Bot. Mex. 3: 2 (1988)
Distribution: Mexico (Nuevo León, Tamaulipas)
79 MEX
species :
Stachys lanigera
(Bornm.) Rech.f.; Bot. Jahrb. Syst. 71: 528 (1941)
Distribution: Iraq (NE-Iraq); Iran (W-Iran)
34 IRN IRQ
Synonyms: hidden
= Stachys fragillima Bornm. var. lanigera Bornm.; Beih. Bot. Centralbl. 33(2): 189 (1915)
species :
Stachys latidens
Small; Man. Fl. N. States [Britton] 793 (1901)
Name: Broadtooth hedgenettle
Distribution: USA (Alabama, District of Columbia, Georgia, Kansas, Maryland, North Carolina, South Carolina, Tennessee, Virginia, West Virginia)
74 USA 75 USA 78 USA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys tenuifolia var. latidens (Small) J.B.Nelson; Phytologia 46: 94 (1980)
species :
Stachys lavandulifolia
Vahl; Symb. Bot. [Vahl] 1: 42 (1790)
Distribution: Turkmenistan; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia, SE-Anatolia, SSW-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, NE-Iran: Mts., N-Iran, Iranian Aserbaijan, W-Iran)
32 TKM 33 ARM GEO 34 IRN IRQ TUR
Remarks: clade 12 Olisia
Synonyms: hidden
= Sideritis calycantha M.Bieb.; Fl. Taur.-Caucas. 3: 393 (1819)
= Stachys boissieri Kapeller; Zametki Syst. Geogr. Rast. 16: 17 (1951)
= Stachys calycantha M.Bieb.; Fl. Taur.-Caucas. 3: 393 (1819)
= Stachys lavandulifolia var. glabrescens R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 283 (1974)
= Stachys tomentosa Bunge; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 21(1): 71 (1873), nom. illeg.
= Stachys zuvandica Rzazade; Fl. Azerbaidzhana 7: 611 (1957)
= Zietenia lavandulifolia (Vahl) Link; Enum. Hort. Berol. Alt. 2: 110 (1822)
= Zietenia orientalis Gled.; Mém. Acad. Roy. Sci. Hist. (Berlin) 1766: 3 (1766)
species :
Stachys laxa
Boiss. & Buhse; Nouv. Mém. Soc. Imp. Naturalistes Moscou 12: 179 (1860)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Synonyms: hidden
= Stachys demavendica Bornm.; Russk. Bot. Zhurn. 1: 8 (1911)
species :
Stachys leucoglossa
Griseb.; Spic. Fl. Rumel. 2: 140 (1844)
subspecies :
subsp. leucoglossa
Distribution: ?Serbia; Kosovo; North Macedonia; Bulgaria; NE-Greece (incl. Mt. Athos)
13 BUL GRC KOS NMA
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys recta var. leucoglossa (Griseb.) Boiss.; Fl. Orient. [Boissier] 4(2): 730 (1879)
= Stachys recta subsp. leucoglossa (Griseb.) Stoj. & Stef.; Fl. Bulg. [Stojanov & Stefanoff], ed. 3: 967 (1948)
subspecies :
subsp. samothracica
(Degen) Biel & Kit Tan; Phytol. Balcan. 19(2): 271 (2013)
Distribution: NE-Greece (Samothraki)
13 GRC
Synonyms: hidden
= Stachys patula var. samothracica Degen; Oesterr. Bot. Z. 41: 336 (1891)
= Stachys leucoglossa var. samothracica (Degen) Rech.Fil.; Repert. Spec. Nov. Regni Veg., Beih. 100: 131 (1938)
species :
Stachys libanotica
Benth.; Prodr. [A.P. de Candolle] 12: 462 (1848)
Distribution: Turkey (S-Anatolia); Lebanon (Antilebanon, C-Lebanon); Syria (Jbel Druze)
34 LEB SYR TUR
Remarks: libanotica agg.
Synonyms: hidden
= Stachys ciliaris Boiss.; Diagn. Pl. Orient. 12: 78 (1853)
species :
Stachys lindenii
Benth.; Prodr. [A.P. de Candolle] 12: 467 (1848)
Distribution: Mexico (Chiapas, Guerrero, Jalisco, Mexico State, Michoacán, Oaxaca, Puebla, Queretaro, Veracruz); Guatemala
79 MEX 80 GUA
Remarks: coccinea agg.
Synonyms: hidden
= Stachys boraginoides Benth.; Labiat. Gen. Spec. [Bentham] 540 (1834), nom. illeg.
= Stachys lindenii var. aristulata Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 49 (1934)
species :
Stachys linearis
Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 559 (1833)
Distribution: Namibia; South Africa (Free State, N-Cape Prov., W-Cape Prov., E-Cape Prov.)
27 NAM SAF
Synonyms: hidden
= Stachys recurva Gürke; Bull. Herb. Boissier 6: 549 (1898)
= Stachys rosmarinifolia Benth.; Labiat. Gen. Spec. [Bentham] 559 (1833)
= Stachys rosmarinifolia var. burkei Benth.; Prodr. [A.P. de Candolle] 12: 494 (1848)
= Stachys rugosa var. linearis (Burch. ex Benth.) Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 359 (1910)
species :
Stachys longespicata
Boiss. & Kotschy; Fl. Orient. [Boissier] 4(2): 725 (1879)
Name: Longspike hedgenettle
Distribution: Turkey (Inner Anatolia, S-Anatolia, SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); ?Iraq (?NE-Iraq); Lebanon (Antilebanon, C-Lebanon); Israel (Rift Valley, N-Israel); USA [I] (Maryland [I])
34 ISR LEB TUR 78 usa
species :
Stachys longiflora
Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 39 (1859)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Stachys luchuensis
(Kudô) comb.ined.
Distribution: Ryukyu Isl.; Taiwan
38 NNS TAI
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Suzukia luchuensis Kudô; J. Soc. Trop. Agric. 3: 226 (1931)
= Glechoma luchuensis (Kudô) Masam.; Hokuriku J. Bot. 5: 8 (1956)
species :
Stachys lurestanica
Jamzad; Fl. Iran [ed. Assadi et al.] 76: 230 (2012)
Distribution: Iran
34 IRN
species :
Stachys lyallii
Benth.; Labiat. Gen. Spec. [Bentham] 548 (1834)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Stachys hildebrandtii Briq.; Bull. Herb. Boissier 2: 138 (1894), nom. illeg.
species :
Stachys macraei
Benth.; Linnaea 6: 78 (1831)
Distribution: Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Rios, Los Lagos, Aisén)
85 CLE
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Lamium hirtum Poir.; Encycl. [J.Lamarck et al.], Suppl. 1: 296 (1810)
= Stachys chonotica Hook.f.; Bot. Antarct. Voy. I (Fl. Antarct.) 336 (1846)
= Stachys quadridentata Bertero ex Benth.; Prodr. [A.P. de Candolle] 12: 473 (1848)
species :
Stachys macrotricha
Rech.f. & Goulimy; Anz. Österr. Akad. Wiss., Math.-Naturwiss. Kl. 94: 24 (1957)
Distribution: NC-Greece (around Grevena)
13 GRC
Remarks: very rare
Synonyms: hidden
= Stachys heraclea var. macrotricha (Rech.f. & Goulimy) Krestovsk.; Novosti Sist. Vyssh. Rast. 42: 214 (2011)
species :
Stachys mandoniana
Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 118 (1898)
Distribution: Bolivia (La Paz)
83 BOL
species :
Stachys marashica
Ilçim, Çenet & Dadandý; Ann. Bot. Fenn. 45(2): 151 (2008)
Distribution: Turkey
34 TUR
species :
Stachys mardinensis
(Post) R.R.Mill; Notes Roy. Bot. Gard. Edinburgh 38: 57 (1980)
Distribution: Turkey (E-Anatolia, Inner Anatolia, SE-Anatolia); Iraq (NE-Iraq)
34 IRQ TUR
Synonyms: hidden
= Nepeta mardinensis Post; Bull. Herb. Boissier 7: 159 (1899)
species :
Stachys maritima
Gouan; Fl. Monsp. [Gouan] 91 (1764)
Distribution: NE-Spain; France; Corsica; Italy; Croatia; ?Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; European Turkey; ?Morocco; Algeria; Tunisia; Georgia [Caucasus]; Turkey (N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia)
12 COR FRA SPA 13 ALB BUL CRO ITA KOS NMA SRB TUE 20 ALG TUN 33 GEO 34 TUR
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys annua subsp. maritima Bonnier & Layens; Tabl. Syn. Pl. Vasc. France 253 (1894)
= Stachys maritima var. pubescens K.Koch; Linnaea 17: 299 (1844)
species :
Stachys marrubiifolia
Viv.; Fl. Cors. Prodr., App. 1: 2 (1825)
Distribution: Corsica; W-Italy; Algeria; Tunisia
12 COR 13 ITA 20 ALG TUN
Synonyms: hidden
= Stachys marrubiifoliohirta Batt.; Fl. Algérie Tunisie 268 (1905)
= Stachys poiretii Ten.; Syll. Pl. Fl. Neapol. 538 (1831)
species :
Stachys matthewsii
G.P.Fleming, J.B.Nelson & J.F.Towns.; J. Bot. Res. Inst. Texas 5(1): 9 (2011)
Distribution: USA (North Carolina)
78 USA
species :
Stachys megalodonta
Hausskn. & Bornm. ex P.H.Davis; Notes Roy. Bot. Gard. Edinburgh 21: 46 (1951)
subspecies :
subsp. megalodonta
Distribution: Turkey (SE-Anatolia); Iraq (NE-Iraq); Iran (Iranian Aserbaijan, W-Iran)
34 IRN IRQ TUR
Remarks: clade 12 Olisia
subspecies :
subsp. mardinensis
R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 282 (1974)
Distribution: Turkey
34 TUR
species :
Stachys melampyroides
Hand.-Mazz.; Ann. K.K. Naturhist. Hofmus. 27: 415, t. 16 (1913)
Distribution: Turkey (SE-Anatolia); Iraq (NE-Iraq)
34 IRQ TUR
species :
Stachys menthifolia
Vis.; Flora 12 (Ergänz. 1): 14 (1829)
Distribution: Croatia; ?Montenegro; Serbia; Kosovo; North Macedonia; Albania; NW-Greece (Mt. Grammos, Mt. Timphi to C-Epirus)
13 ALB CRO GRC KOS NMA SRB
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys grandiflora Host; Fl. Austriac. [Host] 2: 169 (1831)
species :
Stachys menthoides
Kotschy & Boiss.; Fl. Orient. [Boissier] 4(2): 727 (1879)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
Remarks: clade 4 Setifolia
species :
Stachys mexicana
Benth.; Labiat. Gen. Spec. [Bentham] 541 (1833)
Name: Mexican hedgenettle
Distribution: Alaska; Canada (British Columbia); USA (California, Oregon, Washington State)
70 ASK 71 CDN 73 USA 76 USA
Synonyms: hidden
= Stachys ciliata Douglas ex Benth.; Labiat. Gen. Spec. [Bentham] 539 (1834)
= Stachys ciliata subsp. macrantha Piper; Proc. Biol. Soc. Washington 32: 43 (1919)
= Stachys ciliata var. pubens A.Gray; Syn. Fl. N. Amer. 2: 388 (1878)
= Stachys emersonii Piper; Erythea 6: 31 (1898)
= Stachys pubens (A.Gray) A.Heller; Bull. Torrey Bot. Club 25: 581 (1898)
species :
Stachys mialhesii
de Noé; Bull. Soc. Bot. France 2: 584 (1855)
Distribution: Algeria
20 ALG
species :
Stachys micheliana
Briq. ex Micheli; Mém. Soc. Phys. Genève 32(10): 7 (1897)
Distribution: Colombia (Antioquia, Arauca, Boyacá, Caldas, Cauca, Chocó, Córdoba, Cundinamarca, Huila, Magdalena, Meta, Nariño, Putumayo, Quindío, Risaralda, Santander, Tolima, Valle); SE-Brazil (São Paulo, Rio de Janeiro); S-Brazil (Paraná, Santa Catarina, Rio Grande do Sul)
82 CLM 84 BRA
Synonyms: hidden
= Stachys mayorii Briq.; Mem. Soc. Neuchâteloise Sci. Nat. 5: 402 (1913)
species :
Stachys milanii
Petrovič; Scrin. Fl. Select. 6: 117 (1887)
Distribution: Serbia; Kosovo; S- & E-Bulgaria
13 BUL KOS SRB
Synonyms: hidden
= Stachys spinulosa subsp. milanii (Petrov ex Magnier) Nyman; Consp. Fl. Eur. Suppl. 2: 252 (1890)
species :
Stachys milasensis
Ö.Güner; Phytotaxa 566(1): 66 (2022)
Distribution: Turkey
34 TUR
species :
Stachys minor
(Boiss.) Akçiçek & Dirmenci; Turk. J. Bot. 36(3): 225 (2012)
Distribution: Turkey
34 TUR
Synonyms: hidden
= Stachys libanotica var. minor Boiss.; Fl. Orient. [Boissier] 4(2): 718 (1879)
species :
Stachys mohinora
B.L.Turner; Phytologia 77(4): 360 (1994 publ. 1995)
Distribution: Mexico (Chihuahua)
79 MEX
species :
Stachys mollissima
Willd.; Hort. Berol. [Willdenow] 60, t. 60 (1805)
Distribution: S-Albania; NW-Greece (Kerkyra, Paxos, Andipaxos, W-Epirus)
13 ALB GRC
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys decumbens Pers.; Syn. Pl. [Persoon] 2: 123 (1806)
species :
Stachys moorei
B.L.Turner; Phytologia 77(4): 360 (1994 publ. 1995)
Distribution: Mexico (Hidalgo, Veracruz)
79 MEX
species :
Stachys mouretii
Batt. & Pit.; Contr. Fl. Maroc (Tours) 32 (1918)
Distribution: Morocco
20 MOR
species :
Stachys mucronata
Sieber ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 2: 733 (1825)
Distribution: E-Crete; East Aegaean Isl. (Kasos, Karpathos)
13 KRI 34 EAI
Remarks: clade 1 Eriostomum, germanica agg.
species :
Stachys multicaulis
Benth.; Prodr. [A.P. de Candolle] 12: 486 (1848)
Distribution: Iraq (NW-Iraq); Iran (W-Iran)
34 IRN IRQ
species :
Stachys munzurdagensis
R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 290 (1974)
Distribution: Turkey (E-Anatolia)
34 TUR
species :
Stachys namazdaghensis
Yıld.; Ot Sist. Bot. Derg. 17(2): 85 (2010)
Distribution: Turkey
34 TUR
species :
Stachys natalensis
Hochst.; Flora 28: 65 (1845)
Distribution: Zimbabwe; Mozambique; South Africa (Limpopo, Gauteng, Mpulamanga, KwaZulu-Natal); Eswatini [Swaziland]
26 MOZ ZIM 27 SAF SWZ
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys leptoclada Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1084 (1903)
= Stachys transvaalensis Gürke; Bot. Jahrb. Syst. 28: 316 (1900)
variant :
var. galpinii
(Briq.) Codd; Bothalia 12(2): 185 (1977)
Distribution: South Africa (Limpopo, Gauteng, Mpulamanga, KwaZulu-Natal); Eswatini [Swaziland]
27 SAF SWZ
Synonyms: hidden
= Stachys galpinii Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1082 (1903)
= Stachys lupulina Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1082 (1903)
= Stachys parilis N.E.Br.; Bull. Misc. Inform. Kew 1901: 131 (1901)
= Stachys villosissima H.M.L.Forbes; Bothalia 4: 38 (1941)
species :
Stachys nelsonii
B.R.Keener & L.J.Davenp.; J. Bot. Res. Inst. Texas 10(2): 315, figs, 1, 3-4 (2016)
Distribution: USA (Alabama)
78 USA
species :
Stachys nemorivaga
Briq.; Bot. Jahrb. Syst. 19: 192 (1894)
Distribution: D.R.Congo [Zaire]; Angola; Zambia
23 ZAI 26 ANG ZAM
species :
Stachys nepetifolia
Desf.; Tabl. École Bot., ed. 3 (Cat. Pl. Horti Paris.) 58 (1829)
Distribution: Mexico (Ciudad de Mexico, Guanajuato, Hidalgo, Jalisco, Mexico State, Michoacán, Morelos, Oaxaca, Queretaro, San Luis Potosi, Tamaulipas, Tlaxcala, Veracruz, Zacatecas)
79 MEX
Synonyms: hidden
= Nepeta circinnata Willd.; Enum. Pl. Hort. Berol. [Willdenow] 602 (1809)
species :
Stachys nephrophylla
Rech.f.; Pl. Syst. Evol. 134: 291 (1980)
Distribution: Iraq (NE-Iraq)
34 IRQ
Remarks: clade 12 Olisia
species :
Stachys neurocalycina
Boiss.; Diagn. Pl. Orient. 12: 85 (1853) et Fl. 4: 748 (1879)
Distribution: Lebanon (Antilebanon, C-Lebanon, coastal W-Lebanon); Israel (coastal W-Israel, Rift Valley, N-Israel); Jordania (W-Jordania)
34 ISR JOR LEB
Remarks: clade 12 Olisia
species :
Stachys nigricans
Benth.; Comm. Pl. Afr. Austr. [Meyer] 238 (1838)
Distribution: South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, E-Cape Prov.); Eswatini [Swaziland]
27 SAF SWZ
Remarks: clade 2 Stachys s. str.
species :
Stachys nivea
Labill.; Icon. Pl. Syr., Dec. 3: 5, t. 3 (1809)
Distribution: Lebanon (Antilebanon); Syria (C-Syrian Desert); Sinai peninsula (S-Sinai)
34 LEB SIN SYR
species :
Stachys nubilorum
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 48 (1934)
Distribution: Guatemala
80 GUA
Synonyms: hidden
= Stachys glechomoides Epling; Bull. Torrey Bot. Club 71: 484 (1944)
species :
Stachys obliqua
Waldst. & Kit.; Descr. Icon. Pl. Hung. 2: 142 (1803 publ. 1804)
Distribution: Croatia; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; European Turkey; NE-Greece (Nomos Epirou, Rodopi, Thasos); East Aegaean Isl. (Lesvos); Turkey (Inner Anatolia, NW-Anatolia: Bithynia, W-Anatolia, WN-Anatolia)
13 ALB BUL CRO GRC KOS MNT NMA SRB TUE 34 EAI TUR
Synonyms: hidden
= Stachys heraclea var. lutea Benth.; Prodr. [A.P. de Candolle] 12: 463 (1848)
= Stachys montbretii Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 48 (1836)
= Stachys pauciflora Vis.; Ill. Piant. Nuov. 1: 10 (1840), nom. illeg.
species :
Stachys oblongifolia
Wall. ex Benth.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830)
Distribution: China (Anhui, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan); North Korea; Taiwan; NE-India (Assam, Manipur, Meghalaya, Mizoram, Nagaland, Tripura, West Bengal); Bangladesh; Vietnam; W-Java
36 CHI 38 NKO TAI 40 BAN IND 41 VIE 42 JAW
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys imaii Nakai; Bot. Mag. (Tokyo) 26: 169 (1912)
= Stachys martini Vaniot; Bull. Acad. Int. Géogr. Bot. 14: 187 (1904)
= Stachys modica Hance; J. Bot. 20: 292 (1882)
= Stachys palustris var. imaii (Nakai) Nakai; Bot. Mag. (Tokyo) 34: 48 (1920)
= Stachys subargentea Hayata; Icon. Pl. Formosan. 8: 94 (1919)
= Stachys tashiroi Hayata; Icon. Pl. Formosan. 8: 95 (1919)
variant :
var. leptopoda
(Hayata) C.Y.Wu; Acta Phytotax. Sin. 10: 222 (1965)
Distribution: China (Fujian, Guangdong, Guangxi, Sichuan, Yunnan); Taiwan
36 CHI 38 TAI
Synonyms: hidden
= Stachys leptopoda Hayata; Icon. Pl. Formosan. 8: 93 (1919)
= Stachys oblongifolia f. leptopoda (Hayata) Kudô; Mem. Fac. Sci. Taihoku Imp. Univ. 2: 188 (1929)
species :
Stachys obscura
Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 43 (1859)
Distribution: Turkey (S-Anatolia, SSW-Anatolia); Lebanon (C-Lebanon, coastal W-Lebanon); Syria (coastal W-Syria, W-Syrian Mountains)
34 LEB SYR TUR
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys arabica var. minor Boiss.; Fl. Orient. [Boissier] 4(2): 747 (1879)
species :
Stachys obtusicrena
Boiss.; Diagn. Pl. Orient. 7: 57 (1846)
Distribution: Iran (EC-Iran, SW-Iran)
34 IRN
species :
Stachys obtusifolia
MacOwan; Bull. Misc. Inform. Kew 1893: 13 (1893)
Distribution: South Africa (E-Cape Prov.)
27 SAF
Synonyms: hidden
= Stachys obtusifolia var. flanaganii Skan
species :
Stachys ochroleuca
Phil.; Linnaea 33: 192 (1864)
Distribution: Chile (Coquimbo, Valparaíso, O´Higgins, Maule, Ñuble, Biobío, Araucanía)
85 CLE
Synonyms: hidden
= Stachys dubia Briq.; Bull. Herb. Boissier 4: 873 (1896)
= Stachys litoralis Phil.; Anales Univ. Chile 90: 560 (1895)
= Stachys meyenii Phil.; Anales Univ. Chile 1873: 520 (1873), nom. illeg.
= Stachys rupestris Phil.; Anales Univ. Chile 90: 562 (1895), nom. illeg.
species :
Stachys ocymastrum
(L.) Briq.; Lab. Alp. Mar. 252 (1893)
Name: Basilikum-Ziest (DE); Italian hedgenettle (EN); Alfabega (ES)
Distribution: Portugal; Spain; Gibraltar; Baleares; France; Corsica; Sardinia; Italy; Sicily; Greece (very rare Kerkyra, SW-Peloponnisos); NW-Crete; Morocco; Algeria; Tunisia; Libya; Madeira (Madeira Isl., Porto Santo Isl.); Canary Isl. (Lanzarote, Fuerteventura, Gran Canaria, Tenerife, La Gomera, Hierro, La Palma); Lebanon (coastal W-Lebanon); USA [I] (New Jersey [I])
12 BAL COR FRA GIB POR SAR SPA 13 GRC ITA SIC 20 ALG LBY MOR TUN 21 CNY MDR 34 LEB 75 usa
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Galeopsis hirsuta L.; Sp. Pl. [Linnaeus] 2: 580 (1753)
= Galeopsis hispanica Mill.; Gard. Dict., ed. 8: no. 6 (1768)
= Sideritis ocymastrum L.; Syst. Nat., ed. 10, 2: 1098 (1759)
= Stachys divaricata Viv.; Fl. Cors. Prodr., App. 3 (1825)
= Stachys foucauldiana Sennen; Diagn. Nouv. 199 (1936)
= Stachys hirta L.; Sp. Pl. [Linnaeus], ed. 2: 813 (1763), nom. illeg.
= Stachys hirta var. hirtula (Pomel) Batt.; Fl. Algérie [Battandier et al.] Dicot. 706 (1890)
= Stachys hirta var. lagascae (Caball.) Sennen & Mauricio; Cat. Fl. Rif Orient. 93 (1933)
= Stachys hirta var. virgata Batt.; Fl. Algérie [Battandier et al.] Dicot. 706 (1888)
= Stachys hirtula Pomel; Nouv. Mat. Fl. Atl. 305 (1874)
= Stachys inscripta Rchb.; Iconogr. Bot. Exot. 1: 28 (1824)
= Stachys lagascae Caball.; Mem. Real Soc. Esp. Hist. Nat. 8: 280 (1915)
= Stachys marrubiastrum Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 266 (1896)
= Stachys ocymastrum f. bicolor Faure & Maire; Bull. Soc. Hist. Nat. Afrique N. 24: 227 (1933)
= Stachys ocymastrum f. lagascae (Caball.) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 35 (1979)
= Stachys ocymastrum subsp. bicolor (Faure & Maire) C.E.Blanco; Actes Simposi Int. Bot. Font Quer 2: 338 (1992)
= Stachys ocymastrum var. bicolor (Faure & Maire) Maire; Cat. Pl. Maroc 3: 639 (1934)
= Stachys ocymastrum var. rifana (Font-Quer & Maire) Guarr.; Doc. Inst. Sci. Univ. Mohamad V. 4: 35 (1979)
= Stachys ocymastrum var. violascens Faure & Maire; Bull. Soc. Hist. Nat. Afrique N. 20: 197 (1929)
= Stachys rifana Font Quer & Maire; Cavanillesia 3: 54 (1930)
= Stachys rifana var. foucauldiana (Sennen) Font Quer & Sennen; Diagn. Nouv. 240 (1936)
= Tetrahitum hirtum Hoffmanns. & Link; Fl. Portug. [Hoffmannsegg] 1: 104 (1809)
species :
Stachys oligantha
Baker; J. Linn. Soc., Bot. 20: 233 (1884)
Distribution: Madagascar
29 MDG
species :
Stachys oreophila
Hedge; Fl. Madagasc., Fam. 175: 31 (1998)
Distribution: Madagascar
29 MDG
species :
Stachys palaestina
L.; Sp. Pl. [Linnaeus], ed. 2: 1674 (1763)
Distribution: Lebanon (Antilebanon); Israel (coastal W-Israel, N-Israel)
34 ISR LEB
Synonyms: hidden
= Stachys lanata Moench; Methodus [Moench] 397 (1794)
species :
Stachys palustris
L.; Sp. Pl. [Linnaeus] 2: 580 (1753)
Name: Sumpf-Ziest (DE), Marsh Woundwort (EN)
Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Italy; Malta; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (scattered N- & C-Greece, Kerkyra); Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Ukraine; Crimea; Siberia (Altai, Krasnoyarsk, Tuva, W-Siberia); Russian Far East [I] (Amur [I], Khabarovsk [I], Primorye [I], Sakhalin [I]); Kazakhstan; Uzbekistan; Kyrgyzstan; Tajikistan; Northern Caucasus; Georgia [Caucasus]; Armenia; Turkey (N-Anatolia, NE-Anatolia); Iran (Iranian Aserbaijan); China (Xinjiang); Mongolia; NW-India (Jammu & Kashmir); Australia [I] (Tasmania [I]); New Zealand [I]; Alaska [I]; Canada [I] (Manitoba [I], New Brunswick [I], Newfoundland [I], Nova Scotia [I], Ontario [I], Prince Edward Isl. [I], Québec [I]); USA [I] (Connecticut [I], District of Columbia [I], Delaware [I], Illinois [I], Massachusetts [I], Maryland [I], Maine [I], Michigan [I], Minnesota [I], New Hampshire [I], New Jersey [I], New York [I], Ohio [I], Pennsylvania [I], Rhode Island [I], Vermont [I], Wisconsin [I])
10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 CIS COR FRA POR SPA 13 ALB BIH BUL CRO GRC ITA KOS MLT MNT NMA ROM SLV SMA SRB TUE 14 BLR EST KRY LAT LIT RUS UKR 30 SIB 31 rfe 32 KAZ KGZ TZK UZB 33 ARM GEO NCS 34 IRN TUR 36 CHI 37 MON 40 IND 50 aus 51 nzl 70 ask 71 cdn 72 cdn 74 usa 75 usa 78 usa
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys aquatica Bubani; Fl. Pyren. [Bubani] 1: 444 (1897)
= Stachys austriaca Heynh.; Alph. Aufz. Gew. 697 (1847), nom. inval.
= Stachys maeotica Postrig.; Bot. Žurn. (Kiev) 7(2): 67 (1950)
= Stachys palustris f. acuminata (Briq.) Ravarut; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 220 (1961)
= Stachys palustris f. angustifolia Serg.; Fl. Sibir. Occid., ed. 2: 2364 (1937)
= Stachys palustris f. canescens Briq. ex Litard.; Candollea 5: 159 (1932)
= Stachys palustris f. cleoniquei B.Boivin; Naturaliste Canad. 94: 648 (1967)
= Stachys palustris f. eglandulosa Ravarut; Fl. Reipubl. Popul. Roman. (Fl. Republ. Socialist. Romania) 688 (1961)
= Stachys palustris f. latifolia Serg.; Fl. Sibir. Occid., ed. 2: 2364 (1937), nom. illeg.
= Stachys palustris f. petiolata (Peterm.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2417 (1927)
= Stachys palustris f. segetum (Hagen) C.W.Christ.; Fl. Kiel [J. A. Christiansen et al.] 199 (1922)
= Stachys palustris f. sessilis (Čelak.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2417 (1927)
= Stachys palustris f. submersa Glück; Biol. Morphol. Untersuch. Wasser- Sumpfgewächse, Teil 3: 62 (1911)
= Stachys palustris f. virescens Briq. ex Litard.; Candollea 5: 159 (1932)
= Stachys palustris subvar. acuminata (Briq.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2418 (1927)
= Stachys palustris subvar. major (Wirtg.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2418 (1927)
= Stachys palustris subvar. oblongifolia (Briq.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2417 (1927)
= Stachys palustris var. acuminata Briq.; Lab. Alp. Mar. 1: 248 (1891)
= Stachys palustris var. agrestis (F.Aresch.) Serg.; Fl. Sibir. Occid., ed. 2: 2365 (1937)
= Stachys palustris var. angustifolia Benth.; Labiat. Gen. Spec. [Bentham] 542 (1834)
= Stachys palustris var. arenicola Farw.; Amer. Midl. Naturalist 11: 82 (1928)
= Stachys palustris var. bracteata Beck; Fl. Nieder-Österreich 2(2): 1013 (1893)
= Stachys palustris var. ciliata Peterm.; Fl. Lips. Excurs. 447 (1838)
= Stachys palustris var. cinerea T.Durand; Bull. Soc. Roy. Bot. Belgique 20(2): 70 (1881)
= Stachys palustris var. hybrida Benth.; Labiat. Gen. Spec. [Bentham] 542 (1834)
= Stachys palustris var. insularis Briq. ex Litard.; Candollea 5: 159 (1932)
= Stachys palustris var. latifolia Peterm.; Fl. Lips. Excurs. 446 (1838)
= Stachys palustris var. major Wirtg.; Fl. Preuss. Rheinprov. 368 (1857)
= Stachys palustris var. oblongifolia Briq.; Lab. Alp. Mar. 1: 248 (1891)
= Stachys palustris var. petiolata Čelak.; Arch. Naturwiss. Landesdurchf. Böhmen 2(2) (Prodr. Fl. Böhmen 2): 358 (1871), nom. illeg.
= Stachys palustris var. petiolata Peterm.; Anal. Pfl.-Schlüss. 349 (1846)
= Stachys palustris var. petiolata T.Durand; Bull. Soc. Roy. Bot. Belgique 18(2): 101 (1879)
= Stachys palustris var. pseudoambigua Erdner; Ber. Naturwiss. Vereins Schwaben Neuburg 39-40: 571 (1911)
= Stachys palustris var. pubescens Schur; Enum. Pl. Transsilv. 539 (1866)
= Stachys palustris var. segetalis Brügger; Naturgesch. Beitr. Umg. Chur [Fl. Curiensis] 75 (1874), nom. inval.
= Stachys palustris var. segetalis Klett & Richt.; Fl. Leipzig 510 (1830)
= Stachys palustris var. segetum (Hagen) Nyman; Consp. Fl. Eur. 578 (1881)
= Stachys palustris var. segetum (Hagen) Rchb.; Fl. Germ. Excurs. 319 (1832)
= Stachys palustris var. sessilis Čelak.; Arch. Naturwiss. Landesdurchf. Böhmen 2(2) (Prodr. Fl. Böhmen 2): 358 (1871)
= Stachys palustris var. subcanescens Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849)
= Stachys palustris var. umbrosa Čelak.; Arch. Naturwiss. Landesdurchf. Böhmen 2(2) (Prodr. Fl. Böhmen 2): 358 (1871)
= Stachys palustris var. viridifolia Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849)
= Stachys palustris var. vulgaris Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849), nom. illeg.
= Stachys palustris var. vulgaris Peterm.; Anal. Pfl.-Schlüss. 349 (1846)
= Stachys segetum Hagen; Chlor. Boruss. [Hagen] 218 (1819)
= Stachys wolgensis Wilensky; Sborn. Statej Mater. Gosplana A.S.S.R. Nemts. Povolzh. 1: 38 (1926)
species :
Stachys paneiana
Mouterde; Saussurea 4: 25 (1973)
Distribution: Israel (N-Israel)
34 ISR
species :
Stachys pannosa
Phil.; Anales Univ. Chile 90: 564 (1895)
Distribution: Chile (Antofagasta, Atacama, Coquimbo, Valparaíso, Ñuble, Biobío)
85 CLE
species :
Stachys parolinii
Vis.; Mem. Reale Ist. Veneto Sci. 1: 46 (1842 publ. 1843)
Distribution: W-Greece (Lefkada, Kefalonia, N-Peloponnisos)
13 GRC
species :
Stachys parviflora
Benth.; Prodr. [A.P. de Candolle] 12: 490 (1848)
Distribution: Tajikistan; Iran (NE-Iran: Mts.); Afghanistan (Badakshan, Badghys, Baghlan, Balkh, Bamyan, Farah, Faryab, Ghazni, Ghorat, Herat, Kabul, Kandahar, Logar, Wardak, Orozgan / Daykundi, Paktia / Khost, Parwan, Samangan, Takhar); Pakistan (Baluchistan, Quetta, Waziristan, Kurram, N. W. Frontier Prov., Peshawar, Swat, Pakistani Punjab); NW-India (Himachal Pradesh, Jammu & Kashmir, Uttarakhand)
32 TZK 34 AFG IRN 40 IND PAK
Remarks: clade 12 Olisia
Synonyms: hidden
= Phlomidoschema parviflorum (Benth.) Vved.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 9 1: 54 (1941)
species :
Stachys parvifolia
M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 193 (1844)
Distribution: Mexico (Chiapas, Chihuahua, Ciudad de Mexico, Durango, Guanajuato, Hidalgo, Mexico State, Michoacán, Queretaro, San Luis Potosi, Sonora, Tamaulipas, Tlaxcala, Veracruz)
79 MEX
species :
Stachys paulii
Grossh.; Trudy Geobot. Obsl. Pastb. S.S.R. Azerbaijdzhana, Ser. B, 1: 77 (1929)
Distribution: Azerbaijan
33 AZE
Synonyms: hidden
= Stachys zappanioides Woronow; Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 1: 221 (1933)
species :
Stachys penanevada
B.L.Turner; Phytologia 77(4): 362 (1994 publ. 1995)
Distribution: Mexico (Tamaulipas, Nuevo León)
79 MEX
species :
Stachys peruviana
Dombey ex Benth.; Labiat. Gen. Spec. [Bentham] 551 (1833)
Distribution: Peru; Bolivia (Chuquisaca, Cochabamba)
83 BOL PER
species :
Stachys petrokosmos
Rech.f.; Svensk Bot. Tidskr. 43: 40 (1949)
Distribution: Turkey (S-Anatolia)
34 TUR
species :
Stachys philippiana
Vatke; Bot. Zeitung (Berlin) 33: 443 (1875)
Distribution: Chile (Coquimbo, Valparaíso, Metropolitana de Santiago, Maule, Ñuble, Biobío, Araucanía)
85 CLE
Synonyms: hidden
= Stachys andina Phil.; Anales Univ. Chile 90: 562 (1895), nom. illeg.
= Stachys elliptica Phil.; Linnaea 33: 193 (1864), nom. illeg.
species :
Stachys pilifera
Benth.; Prodr. [A.P. de Candolle] 12: 487 (1848)
subspecies :
subsp. pilifera
Distribution: Iran (SW-Iran)
34 IRN
subspecies :
subsp. ixodes
(Boiss. & Hausskn. ex Boiss.) Salmaki; Bot. J. Linn. Soc. 170(4): 606 (2012)
Distribution: Iran (SW-Iran)
34 IRN
Synonyms: hidden
= Stachys ixodes Boiss. & Hausskn. ex Boiss.; Fl. Orient. [Boissier] 4(2): 738 (1879)
species :
Stachys pilosa
Nutt.; J. Acad. Nat. Sci. Philadelphia 7: 48 (1834)
Name: Hairy hedgenettle
Distribution: Alaska; Canada (Alberta, British Columbia, Manitoba, New Brunswick, Northern Territories, Ontario, Prince Edward Isl., Québec, Saskatchewan, Yukon); USA (Arizona, California, Colorado, Connecticut, Iowa, Idaho, Illinois, Kansas, Massachusetts, Maine, Minnesota, Missouri, Montana, North Dakota, Nebraska, New Hampshire, New Jersey, New Mexico, Nevada, New York, Oklahoma, Oregon, Pennsylvania, South Dakota, Utah, Vermont, Washington State, Wisconsin, Wyoming)
70 ASK CDN 71 CDN 72 CDN 73 USA 74 USA 75 USA 76 USA 77 USA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys ampla Rydb.; Bull. Torrey Bot. Club 36: 683 (1909)
= Stachys asperrima Rydb.; Bull. Torrey Bot. Club 36: 682 (1909)
= Stachys borealis Rydb.; Brittonia 1: 95 (1931)
= Stachys bracteata Greene; Pittonia 3: 342 (1898)
= Stachys homotricha (Fernald) Rydb.; Brittonia 1: 95 (1931)
= Stachys leibergii Rydb.; Bull. Torrey Bot. Club 36: 682 (1909)
= Stachys palustris f. stevensonis B.Boivin; Phytologia 22: 380 (1972)
= Stachys palustris subsp. homotricha (Fernald) Hultén; Kungl. Svenska Vetenskapsakad. Handl., Ser. 4, 13: 152 (1971)
= Stachys palustris subsp. pilosa (Nutt.) Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 63 (1934)
= Stachys palustris var. homotricha Fernald; Rhodora 10: 85 (1908)
= Stachys palustris var. nipigonensis Jenn.; J. Wash. Acad. Sci. 10: 459 (1920)
= Stachys palustris var. phaneropoda Weath. ex Fernald; Rhodora 45: 475 (1943)
= Stachys palustris var. pilosa (Nutt.) Fernald; Rhodora 45: 474 (1943)
= Stachys palustris var. puberula Jenn.; J. Wash. Acad. Sci. 10: (1920)
= Stachys pilosa var. homotricha (Fernald) Mohlenbr.; Phytoneuron 2015-67: 2 (2015)
= Stachys puberula (Jenn.) Rydb.; Brittonia 1: 96 (1931)
= Stachys pustulosa Rydb.; Brittonia 1: 95 (1931)
= Stachys schweinitzii Rydb.; Brittonia 1: 95 (1931)
= Stachys scopulorum Greene; Pittonia 3: 342 (1898)
= Stachys teucriformis Rydb.; Bull. Torrey Bot. Club 31: 640 (1904)
= Stachys velutina Schwein.; Narrat. Exped. St. Peter´s River [Keating] 2: 390 (1824), nom. illeg.
variant :
var. arenicola
(Britton) G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116: 46 (1989 publ. 1990)
Distribution: Canada (Manitoba, Ontario, Québec); USA (Arkansas, Connecticut, Iowa, Illinois, Indiana, Kansas, Kentucky, Massachusetts, Maine, Michigan, Minnesota, Missouri, Nebraska, New Hampshire, New Jersey, New York, Ohio, Pennsylvania, South Carolina, South Dakota, Virginia, Vermont, Wisconsin, West Virginia)
71 CDN 72 CDN 74 USA 75 USA 78 USA
Synonyms: hidden
= Stachys arenicola Britton; Man. Fl. N. States [Britton] 792 (1901)
= Stachys brevidens Rydb.; Brittonia 1: 95 (1931), nom. illeg.
species :
Stachys pilosissima
M.Martens & Galeotti; Bull. Acad. Roy. Sci. Bruxelles 11(2): 194 (1844)
Distribution: Mexico (Chiapas, Guerrero, Jalisco, Mexico State, Michoacán, Oaxaca, Puebla, Tamaulipas, Veracruz)
79 MEX
Synonyms: hidden
= Stachys exilis Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 25 (1934)
= Stachys flaccida Fernald; Proc. Amer. Acad. Arts 35: 563 (1900)
= Stachys vulcanica Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 33 (1934)
species :
Stachys pinardii
Boiss.; Diagn. Pl. Orient. 5: 29 (1844)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Stachys pinetorum
Boiss. & Balansa; Diagn. Pl. Orient., Sér. 2, 4: 36 (1859)
Distribution: Turkey (S-Anatolia, SSW-Anatolia); Syria (W-Syrian Mountains)
34 SYR TUR
species :
Stachys pittieri
Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 119 (1898)
Distribution: Costa Rica
80 COS
species :
Stachys plumosa
Griseb.; Spic. Fl. Rumel. 2: 139 (1844)
subspecies :
subsp. plumosa
Distribution: ?Serbia; Kosovo; North Macedonia; Albania; W-Bulgaria; Greece (NC-Greece to Thessalia, NE-Greece to Thasos)
13 ALB BUL GRC KOS NMA
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys pangaea Phitos; Oesterr. Bot. Z. 113: 272 (1966)
= Stachys recta subsp. plumosa (Griseb.) Nyman; Consp. Fl. Eur. 579 (1881)
= Stachys viridis Boiss. & Heldr.; Diagn. Pl. Orient., Sér. 2, 4: 39 (1859)
subspecies :
subsp. freynii
(Hausskn.) Hayek; Repert. Spec. Nov. Regni Veg., Beih. 30 (Prodr. Fl. Penins. Balc.): 290 (1929)
Distribution: North Macedonia; NC-Greece
13 GRC NMA
Synonyms: hidden
= Stachys freynii Hausskn.; Mitt. Geogr. Ges. (Thüringen) Jena 5: 70 (1887)
= Stachys dolopica Formánek
species :
Stachys pringlei
Greenm.; Proc. Amer. Acad. Arts 41: 245 (1905)
Distribution: Mexico (Hidalgo, Jalisco, Mexico State, Michoacán, Queretaro, San Luis Potosi, Veracruz)
79 MEX
species :
Stachys pseudohumifusa
Sebsebe; Kew Bull. 48(1): 117 (1993)
subspecies :
subsp. pseudohumifusa
Distribution: Cameroon; Bioko Isl. [Fernando Poo]; Tanzania
23 BIO CMN 25 TAN
Conservation Status: IUCN EN (Cameroon)
subspecies :
subsp. minutiflora
Y.B.Harv.; Kew Bull. 51(3): 451 (1996)
Distribution: Zambia
26 ZAM
subspecies :
subsp. saxeri
Y.B.Harv.; Kew Bull. 51(3): 451 (1996)
Distribution: Nigeria; Cameroon
22 NGA 23 CMN
species :
Stachys pseudonigricans
Gürke; Bot. Jahrb. Syst. 28: 315 (1900)
Distribution: Tanzania; Angola; Zambia; Malawi; Zimbabwe
25 TAN 26 ANG MLW ZAM ZIM
species :
Stachys pseudophlomis
C.Y.Wu; Acta Phytotax. Sin. 10: 226 (1965)
Distribution: China (Hubei, Sichuan)
36 CHI
species :
Stachys pseudopinardii
R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 281 (1974)
Distribution: Turkey (SSW-Anatolia)
34 TUR
species :
Stachys pumila
Banks & Sol.; Nat. Hist. Aleppo, ed. 2 [Russell]: 2: 225, t. 10 (1794)
Distribution: Turkey (S-Anatolia, SE-Anatolia); Syria (NW-Syria, W-Syrian Mountains)
34 SYR TUR
Synonyms: hidden
= Stachys bornmuelleri Hand.-Mazz.; Ann. K.K. Naturhist. Hofmus. 27: 414 (1913)
= Stachys floribunda Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 51 (1836)
species :
Stachys pusilla
(Wedd.) Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 120 (1898)
Distribution: Colombia (Boyacá, Cundinamarca, Tolima); Ecuador; Peru; Bolivia (La Paz)
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Stachys elliptica var. pusilla Wedd.; Chlor. Andina 2: 153 (1860)
species :
Stachys pycnantha
Benth.; Pl. Hartw. [Bentham] 331 (1849)
Name: Shortspike hedgenettle
Distribution: USA (California)
76 USA
species :
Stachys pyramidalis
J.K.Morton; J. Linn. Soc., Bot. 58: 273 (1962)
Distribution: Nigeria
22 NGA
species :
Stachys radicans
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 24 (1934), in Repert. Spec. Nov. Regni Veg., Beih. 85: 9 (1935)
Distribution: Mexico (Chiapas, Ciudad de Mexico, Hidalgo, Jalisco, Mexico State, Michoacán, Tlaxcala, Veracruz); Colombia (Boyacá, Cundinamarca, Nariño)
79 MEX 82 CLM
species :
Stachys ramosissima
Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 50 (1836)
Distribution: Turkey (E-Anatolia, Inner Anatolia)
34 TUR
Synonyms: hidden
= Stachys ramosissima var. elazigensis R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 291 (1974)
species :
Stachys recta
L.; Mant. Pl. 82 (1767)
subspecies :
subsp. recta
Name: Aufrechter Ziest (DE), Stiff hedgenettle (EN)
Distribution: Belgium; Luxembourg; Germany (Brandenburg, +Berlin, Baden-Württemberg, Bayern, Hessen, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein [I], Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; NE-Spain; Andorra; France; Italy; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; ?Greece; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; Ukraine; Northern Caucasus; USA [I] (District of Columbia [I])
11 AUT BGM CZE GER HUN LIE LUX POL SLO SWI 12 ADO FRA SPA 13 ALB BIH BUL CRO ITA KOS MNT NMA ROM SLV SMA SRB 14 BLR LAT LIT RUS UKR 33 NCS 78 usa
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Betonica adscendens Walther; Fl. Giessen 453 (1802), nom. illeg.
= Betonica decumbens Moench; Methodus [Moench] 396 (1794), nom. illeg.
= Betonica hirta L.; Cent. Pl. II: 20 (1756)
= Betonica recta (L.) Baill.; Iconogr. Fl. Franç. t. 208
= Ortostachys recta (L.) Fourr.; Ann. Soc. Linn. Lyon, Sér. 2, 17: 136 (1869)
= Prasium stachys E.H.L.Krause; Deutschl. Fl. [Sturm], ed. 2, Abbild., 11: 116 (1903), nom. illeg.
= Sideritis maculata Gilib.; Fl. Lit. Inch. 1: 81 (1782), opus utique oppr.
= Stachys albanica Markgr.; Ber. Deutsch. Bot. Ges. 44: 428 (1926)
= Stachys betonica Scop.; Fl. Carniol., ed. 2, 1: 411 (1771)
= Stachys bufonia Thuill.; Fl. Env. Paris, éd. 2(1): 295 (1799)
= Stachys czernjaevii Des.-Shost.; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 8: 152 (1940)
= Stachys decumbens Rchb.; Fl. Germ. Excurs. 318 (1831), nom. illeg.
= Stachys delphinensis Jord.; Arch. Fl. France Allem. [F.W.Schultz] 322 (1847), et in Cat. Jard. Bot. Grenoble 1853: 12 (1853)
= Stachys erecta K.Koch; Linnaea 21: 691 (1849)
= Stachys glabrata Simonk.; Enum. Fl. Transsilv. 450 (1886)
= Stachys hirta (L.) Thell.; Repert. Spec. Nov. Regni Veg. 10: 290 (1912), nom. illeg.
= Stachys krynkensis Kotov; Bot. Žurn. (Kiev) 1(2): 297 (1940)
= Stachys nitens Janka; Linnaea 30: 597 (1859 publ. 1860)
= Stachys patula Griseb.; Spic. Fl. Rumel. 2: 142 (1844)
= Stachys procumbens Lam.; Fl. Franç. [Lamarck] 2: 385 (1779)
= Stachys recta f. angusta Borbás; Természettud. Közl. 1884: 158 (1884)
= Stachys recta f. angustifolia (Ten.) Bolzon; Bull. Soc. Bot. Ital. 1903: 37 (1903)
= Stachys recta subsp. delphinensis (Jord.) Nyman; Consp. Fl. Eur. 579 (1881)
= Stachys recta subsp. hirta (Ten.) Briq.; Lab. Alp. Mar. 263 (1893)
= Stachys recta subsp. sarajevensis (K.Maly) Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 181 (1924)
= Stachys recta subvar. alpina (Godr.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2411 (1927)
= Stachys recta subvar. glandulifera (A.Kern.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2411 (1927)
= Stachys recta subvar. midzorica Adamović; Oesterr. Bot. Z. 42: 408 (1899)
= Stachys recta var. alpina Godr.; Fl. France [Grenier] 2: 693 (1853)
= Stachys recta var. angusta (Borbás) Diklić; Fl. Srbije 6: 427 (1974)
= Stachys recta var. angustifolia Godr.; Fl. France [Grenier] 2: 692 (1853), nom. illeg.
= Stachys recta var. angustifolia Ten.; Syll. Pl. Fl. Neapol. 292 (1831)
= Stachys recta var. angustifolia Wender.; Fl. Hassiaca [Wenderoth] 197 (1846)
= Stachys recta var. glabrescens Pančić; Verh. Zool.-Bot. Vereins Wien 6: 543 (1856)
= Stachys recta var. hirta Ten.; Syll. Pl. Fl. Neapol. 292 (1831)
= Stachys recta var. lisae Briq.; Lab. Alp. Mar. 262 (1893)
= Stachys recta var. longiflora Pamp.; Boll. Mus. Republ. San Marino 4: 117 (1920)
= Stachys recta var. luxemburgensis Lefort; Arch. Sect. Sci. Inst. Grand-Ducal Luxemb., n. s. 19: 443 (1950)
= Stachys recta var. major Ten.; Syll. Pl. Fl. Neapol. 292 (1831)
= Stachys recta var. midzorica (Adamović) Diklić; Fl. Srbije 6: 428 (1974)
= Stachys recta var. nitens (Janka) Chrtek; Folia Geobot. Phytotax. 27: 182 (1992)
= Stachys recta var. ramosissima Griseb.; Arch. Naturgesch. (Berlin) 18: 342 (1852)
= Stachys recta var. ramosissima Pančić; Verh. Zool.-Bot. Vereins Wien 6: 543 (1856), nom. illeg.
= Stachys recta var. stenophylla (Spreng.) Nyman; Consp. Fl. Eur. 579 (1881)
= Stachys sarajevensis K.Malý; Glasn. Zemaljsk. Muz. Bosni Hercegovini 40: 160 (1920)
= Stachys sideritis Vill.; Hist. Pl. Dauphiné [Villars] 2: 375 (1787)
= Stachys spinulosa var. glabrata (Simonk.) Nyman; Consp. Fl. Eur. Suppl. 2: 252 (1890)
= Stachys stenophylla Spreng.; Novi Provent. [Sprengel] 40 (1818)
= Stachys sylvestris Forssk.; Fl. Aegypt.-Arab. xxviii (1775)
= Stachys transsilvanica Schur; Enum. Pl. Transsilv. 539 (1866)
= Syderitis maculata Gilib.; Fl. Lit. Inch. 1: 81 (1782), opus utique oppr.
subspecies :
subsp. baldaccii
(Maly) Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 183 (1924)
Distribution: Croatia; ?Montenegro; ?North Macedonia; Albania; Greece
13 ALB CRO GRC
Synonyms: hidden
= Stachys baldaccii (K.Malý) Hand.-Mazz. & Janch.; Oesterr. Bot. Z. 56: 105 (1906)
= Stachys subcrenata f. baldaccii K.Malý; Verh. K.K. Zool.-Bot. Ges. Wien 54: 245 (1904)
subspecies :
subsp. doerfleri
(Hayek) Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 167 (1924)
Distribution: Albania
13 ALB
Synonyms: hidden
= Stachys doerfleri Hayek; Oesterr. Bot. Z. 70: 18 (1921)
subspecies :
subsp. labiosa
(Bertol.) Briq.; Lab. Alp. Mar. 259 (1893)
Distribution: Germany [I]; Austria; France; Italy; +San Marino; Slovenia; Croatia
11 AUT ger 12 FRA 13 CRO ITA SLV SMA
Synonyms: hidden
= Stachys labiosa Bertol.; Fl. Ital. [Bertoloni] 6: 166 (1844)
= Stachys recta subsp. grandiflora (Caruel) Arcang.; Comp. Fl. Ital. [Arcangeli] 551 (1882)
= Stachys recta var. grandiflora Caruel; Prodr. Fl. Tosc. 519 (1860)
= Stachys recta var. labiosa (Bertol.) Briq.; Lab. Alp. Mar. 259 (1893)
= Stachys subcrenata subsp. labiosa (Bertol.) Nyman; Consp. Fl. Eur. 579 (1881)
= Stachys subcrenata var. labiosa (Bertol.) Vis.; Fl. Dalmat. 2: 208 (1847)
subspecies :
subsp. olympica
Stoj. & Jordanov; God. Sofiisk. Univ. Fiz.-Mat. Fak. 34: 205 (1937-1938 publ. 1938)
Distribution: Greece (Mt. Olimbos)
13 GRC
subspecies :
subsp. psammophila
(Fiori) Arrigoni; Fl. Analítica Toscana 7: 163 (2020)
Distribution: Italy
13 ITA
Synonyms: hidden
= Stachys recta f. psammophila Fiori; Sched. Fl. Ital. Exs., Ser. 3, 14: 217, no. 2537 (1921)
subspecies :
subsp. rhodopaea
(Velen.) Chrtek; Folia Geobot. Phytotax. 27(2): 182 (1992)
Distribution: Bulgaria
13 BUL
Synonyms: hidden
= Stachys fragilis var. rhodopaea Velen.; Fl. Bulg. [Velenovský] 456 (1891)
subspecies :
subsp. subcrenata
(Vis.) Briq.; Lab. Alp. Mar. 257 (1893)
Distribution: Germany [I]; Italy; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; European Turkey; Greece (mountains of N-Greece, also C-Greece, Peloponnisos, Thasos); Turkey (NW-Anatolia: Bithynia, SW-Anatolia)
11 ger 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA SLV SRB TUE 34 TUR
Synonyms: hidden
= Stachys fragilis f. serpentini Fiori; Nuovo Giorn. Bot. Ital., n. s. 17(4): 640 (1910)
= Stachys fragilis Vis.; Flora 12(1. Ergänzungsbl.): 14 (1829)
= Stachys glandulifera Dalla Torre & Sarnth.; Fl. Tirol [Dalla Torre & Sarnthein] 6(3): 187 (1912)
= Stachys karstiana (Borbás) Hand.-Mazz.; Oesterr. Bot. Z. 56: 100 (1906)
= Stachys montenegrina K.Malý; Glasn. Zemaljsk. Muz. Bosni Hercegovini 22: (1910)
= Stachys omblae H.Lindb.; Öfvers. Finska Vetensk.-Soc. Förh. 48(13): 93 (1906)
= Stachys petrogena Hand.-Mazz. & Janch.; Oesterr. Bot. Z. 56: 101 (1906)
= Stachys ramosissima Rochel; Bot. Reise Banat 82 (1838), nom. illeg.
= Stachys recta f. eriostachyus (A.Kern.) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2412 (1927)
= Stachys recta subsp. ramosissima Nyman; Consp. Fl. Eur. 579 (1881)
= Stachys recta subsp. serpentini (Fiori) Arrigoni; Inform. Bot. Ital. 12: 138 (1980 publ. 1981)
= Stachys recta var. fragilis (Vis.) Boiss.; Fl. Orient. [Boissier] 4(2): 730 (1879)
= Stachys recta var. hyssopifolia Ten.; Syll. Pl. Fl. Neapol. 292 (1831)
= Stachys recta var. karstiana (Borbás) Chrtek; Folia Geobot. Phytotax. 27: 182 (1992), isonym
= Stachys recta var. karstiana (Borbás) Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2411 (1927)
= Stachys recta var. subcrenata (Vis.) Nyman; Consp. Fl. Eur. 579 (1881)
= Stachys recta var. visianii Briq.; Lab. Alp. Mar. 259 (1893)
= Stachys subcrenata subsp. fragilis (Vis.) Poldini; Stud. Geobot. 21: 8 (2001)
= Stachys subcrenata var. angustifolia Vis.; Fl. Dalmat. 2: 208 (1847)
= Stachys subcrenata var. fragilis (Vis.) Vis.; Fl. Dalmat. 2: 208 (1847)
= Stachys subcrenata var. karstiana Borbás; Math. Term. Közlem. 14: 402 (1877)
= Stachys subcrenata Vis.; Flora 12(1. Ergänzungsbl.): 15 (1829)
subspecies :
subsp. tenoreana
Bornm.; Magyar Bot. Lapok 33: 81 (1934)
Distribution: Italy
13 ITA
Synonyms: hidden
= Stachys tenoreana Bornm.; Magyar Bot. Lapok 33: 81 (1934)
species :
Stachys rehmannii
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 345 (1910)
Distribution: South Africa (Limpopo)
27 SAF
species :
Stachys reptans
Hedge; Fl. Madagasc., Fam. 175: 26 (1998)
Distribution: Madagascar
29 MDG
Remarks: clade 2 Stachys s. str.
species :
Stachys reticulata
Codd; Bothalia 12(2): 183 (1977)
Distribution: South Africa (Mpulamanga)
27 SAF
species :
Stachys richardiana
R.Kr.Singh; Phytotaxa 620(2): 198 (2023)
Distribution: Ethiopia
24 ETH
Synonyms: hidden
= Stachys hypoleuca Hochst. ex A.Rich.; Tent. Fl. Abyss. 2: 197 (1850), nom. illeg.
= Stachys palaestina var. hypoleuca (Hochst. ex A.Rich.) Benth.
species :
Stachys riederi
Cham.; Linnaea 6: 570 (1831)
Distribution: European Russia; Siberia (Buryatia, Chita, Irkutsk, Krasnoyarsk, Yakutia); Russian Far East (Amur, Kamchatka, Khabarovsk, Kuril Isl., Magadan, Primorye, Sakhalin); China (Heilongjiang, Jilin, Liaoning, Nei Mongol, Shaanxi, Shandong, Shanxi); Mongolia; Japan (Hokkaido)
14 RUS 30 SIB 31 RFE 36 CHI 37 MON 38 JAP
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys aspera subsp. baicalensis (Fisch. ex Benth.) Krestovsk.; Turczaninowia 7(4): 20 (2004)
= Stachys aspera var. baicalensis (Fisch. ex Benth.) Maxim.; Bull. Soc. Imp. Naturalistes Moscou 54(1): 44 (1879)
= Stachys baicalensis Fisch. ex Benth.; Labiat. Gen. Spec. [Bentham] 543 (1834)
= Stachys baicalensis var. hispida (Ledeb.) Nakai; Bot. Mag. (Tokyo) 34: 46 (1920)
= Stachys palustris f. baicalensis (Fisch. ex Benth.) Regel; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 4(4): 119 (1861)
= Stachys palustris var. baicalensis (Fisch. ex Benth.) Turcz.; Bull. Soc. Imp. Naturalistes Moscou 7: 156 (1837)
= Stachys palustris var. hispida Ledeb.; Fl. Ross. [Ledebour] 3: 414 (1849)
= Stachys riederi var. hispida (Ledeb.) H.Hara; Bot. Mag. (Tokyo) 41: 144 (1937)
= Stachys rigens Oett.; Trudy Bot. Sada Imp. Yur´evsk Univ. 6: 216 (1906)
variant :
var. hispidula
(Regel) H.Hara; Bot. Mag. (Tokyo) 51: 144 (1937)
Distribution: Russian Far East (Kuril Isl.); China (Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol); Mongolia; South Korea; Japan (Hokkaido, Honshu, Shikoku, Kyushu)
31 RFE 36 CHI 37 MON 38 JAP KOR
Synonyms: hidden
= Stachys aspera var. hispidula (Regel) Krestovsk.; Sosud. Rast. Sovet. Dal´nego Vostoka 7: 344 (1995), isonym
= Stachys aspera var. hispidula (Regel) Vorosch.; Florist. Issl. Razn. Raionakh S.S.S.R. [A.K.Skvortsov] 189 (1985)
= Stachys baicalensis f. intermedia Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43: 32 (1921)
= Stachys baicalensis var. angustifolia Honda; Bot. Mag. (Tokyo) 46: 374 (1932)
= Stachys baicalensis var. hispidula (Regel) Nakai; Bot. Mag. (Tokyo) 34: 46 (1920)
= Stachys japonica f. intermedia Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43(8): 32 (1921)
= Stachys palustris f. hispidula Regel; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 4(4): 119 (1861)
= Stachys riederi f. angustifolia (Honda) H.Hara; Bot. Mag. (Tokyo) 51: 144 (1937)
= Stachys riederi var. intermedia (Kudô) Kitam.; Acta Phytotax. Geobot. 17: 10 (1957)
variant :
var. japonica
(Miq.) H.Hara; Bot. Mag. (Tokyo) 51: 144 (1937)
Distribution: Russian Far East (Amur); China (Anhui, Fujian, Hebei, Henan, Jiangsu, Jiangxi, Liaoning, Nei Mongol, Shandong, Zhejiang); North Korea; South Korea; Japan (Honshu, Kyushu); Ryukyu Isl.
31 RFE 36 CHI 38 JAP KOR NKO NNS
Synonyms: hidden
= Stachys aspera f. glabrata Nakai; Fl. Kor. 2: 147 (1911), nom. illeg.
= Stachys aspera subsp. japonica (Miq.) Krestovsk.; Turczaninowia 7(4): 21 (2004)
= Stachys aspera var. japonica (Miq.) Maxim.; Bull. Soc. Imp. Naturalistes Moscou 54(1): 45 (1879)
= Stachys aspera var. japonica (Miq.) Vorosch.; Florist. Issl. Razn. Raionakh S.S.S.R. [A.K.Skvortsov] 189 (1985)
= Stachys baicalensis var. japonica (Miq.) Kom.; Trudy Imp. S.-Peterburgsk. Bot. Sada 25: 371 (1907)
= Stachys japonica f. angustifolia Miq.; Ann. Mus. Bot. Lugduno-Batavi 3: 197 (1867)
= Stachys japonica f. glabrata Matsum. & Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43(8): 30 (1921)
= Stachys japonica f. villosa Kudô; J. Coll. Sci. Imp. Univ. Tokyo 43: 32 (1921)
= Stachys japonica Miq.; Ann. Mus. Bot. Lugduno-Batavi 2: 111 (1865)
= Stachys japonica var. tomentosa F.Z.Li & Z.Y.Sun; Bull. Bot. Res., Harbin 25(2): 134 (2005)
= Stachys japonica var. villosa (Kudô) Ohwi; Fl. Jap. [Ohwi] 1008 (1953)
= Stachys riederi f. harana H.Ohba & S.Akiyama; Bull. Univ. Mus. Univ. Tokyo 41: 101 (2005)
= Stachys riederi var. villosa (Kudô) Kitam.; Acta Phytotax. Geobot. 17: 10 (1957)
species :
Stachys rigida
Nutt. ex Benth.; Prodr. [A.P. de Candolle] 12: 472 (1848)
subspecies :
subsp. rigida
Name: Rough hedgenettle
Distribution: USA (California, Nevada, Oregon, Washington State); Mexico (Baja California Norte)
73 USA 76 USA 79 MEX
Synonyms: hidden
= Stachys ajugoides var. rigida (Nutt. ex Benth.) Jeps. & Hoover; Fl. Calif. [Jepson] 3: 426 (1943)
= Stachys littoralis Greene; Pittonia 3: 341 (1898), nom. illeg.
= Stachys prattenii Durand; J. Acad. Nat. Sci. Philadelphia, Ser. 2, 3: 100 (1855)
= Stachys rigida subsp. rivularis Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 60 (1934)
= Stachys rivularis A.Heller; Muhlenbergia 1: 33 (1901), nom. illeg.
= Stachys striata Greene; Pittonia 3: 341 (1898)
= Stachys vestita Howell; Fl. N.W. Amer. [Howell] 558 (1901), non Benth.
subspecies :
subsp. quercetorum
(A.Heller) Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 59 (1934)
Distribution: USA (California, Oregon)
73 USA 76 USA
Synonyms: hidden
= Stachys ajugoides var. quercetorum (A.Heller) A.Heller; Fl. Calif. [Jepson] 3(2): 426 (1943)
= Stachys gracilenta A.Heller; Muhlenbergia 2: 319 (1907)
= Stachys nuttallii var. leptostachya Benth.; Pl. Hartw. [Bentham] 331 (1849)
= Stachys quercetorum A.Heller; Muhlenbergia 2: 318 (1907)
= Stachys rigida var. quercetorum (A.Heller) G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116: 47 (1989 publ. 1990)
= Stachys viarum A.Heller; Muhlenbergia 2: 316 (1907)
species :
Stachys riparia
A.Pool; Novon 17(1): 64 (2007)
Distribution: Costa Rica
80 COS
species :
Stachys rivularis
J.M.Wood & M.S.Evans; J. Bot. 35: 489 (1897)
Distribution: South Africa (KwaZulu-Natal)
27 SAF
Synonyms: hidden
= Stachys schlechteri Gürke; Bot. Jahrb. Syst. 26: 74 (1898)
species :
Stachys rizeensis
R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33: 278 (1974)
Distribution: Turkey (NE-Anatolia)
34 TUR
species :
Stachys rosea
(Desf.) Boiss.; Fl. Orient. [Boissier] 4(2): 725 (1879)
Distribution: Algeria
20 ALG
Synonyms: hidden
= Sideritis aegyptiaca Pers.; Syn. Pl. [Persoon] 2: 117 (1806)
= Sideritis rosea Desf.; Ann. Mus. Hist. Nat. 10: 302 (1807)
species :
Stachys rothrockii
A.Gray; Proc. Amer. Acad. Arts 12: 82 (1877)
Name: Rothrock´s hedgenettle
Distribution: USA (Arizona, New Mexico, Utah)
76 USA 77 USA
species :
Stachys rotundifolia
Moc. & Sessé ex Benth.; Labiat. Gen. Spec. [Bentham] 548 (1833)
Distribution: Mexico (Chiapas, Colima, Hidalgo, Jalisco, Mexico State, Morelos, Oaxaca, Puebla, Queretaro, Veracruz)
79 MEX
species :
Stachys rubella
Hedge; Fl. Madagasc., Fam. 175: 33 (1998)
Distribution: Madagascar
29 MDG
species :
Stachys rudatisii
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 347 (1910)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)
27 SAF
species :
Stachys rugosa
Aiton; Hort. Kew. [W.Aiton] 2: 303 (1789)
Distribution: South Africa (N-Cape Prov., W-Cape Prov., E-Cape Prov.); Lesotho
27 LES SAF
Synonyms: hidden
= Sideritis pallida Thunb.; Prodr. Pl. Cap. 2: 95 (1800)
= Stachys crenulata Briq.; Bot. Jahrb. Syst. 19: 192 (1894)
= Stachys deserti Benth.; Prodr. [A.P. de Candolle] 12: 494 (1848)
= Stachys jugalis Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 562 (1834)
= Stachys multiflora Benth.; Prodr. [A.P. de Candolle] 12: 492 (1848)
= Stachys rugosa var. longiflora Benth.; Comm. Pl. Afr. Austr. [Meyer] 1(2): 241 (1838)
species :
Stachys rupestris
Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 49 (1836)
Distribution: Turkey (S-Anatolia, SSW-Anatolia); Syria (NW-Syria, W-Syrian Mountains)
34 SYR TUR
Remarks: clade 12 Olisia
species :
Stachys salisii
Jord. & Fourr.; Brev. Pl. Nov. 2: 92 (1868)
Distribution: Corsica; Sardinia; Italy
12 COR SAR 13 ITA
Synonyms: hidden
= Stachys aimerici Gamisans; Monde Pl. 100: 26 (2005)
= Stachys corsica proles salisii (Jord. & Fourr.) Rouy; Fl. France [Rouy & Foucaud] 11: 313 (1909)
= Stachys corsica var. micrantha Bertol.; Fl. Ital. [Bertoloni] 6(1): 154 (1845)
species :
Stachys sanchezii
Rzed. & A.García; Phytologia 56(5): 321 (1984)
Distribution: Mexico (Ciudad de Mexico, Hidalgo, Jalisco, Mexico State, Michoacán, Puebla, Queretaro, Veracruz)
79 MEX
species :
Stachys sandersii
B.L.Turner; Phytologia 77(4): 372 (1994 publ. 1995)
Distribution: Mexico (San Luis Potosi)
79 MEX
species :
Stachys saturejoides
Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 51 (1836)
Distribution: Turkey (S-Anatolia, SE-Anatolia, SE-Anatolia: Mesopotamian Anatolia); Syria (Jazira)
34 SYR TUR
species :
Stachys saxicola
Coss. & Balansa; Bull. Soc. Bot. France 20: 257 (1873)
subspecies :
subsp. saxicola
Distribution: Morocco; Algeria
20 ALG MOR
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys saxicola subsp. chelifensis Quézel & Simonn.; Bull. Soc. Hist. Nat. Afrique N. 50: 149 (1959)
subspecies :
subsp. laxa
Faure & Maire; Bull. Soc. Hist. Nat. Afrique N. 22: 61 (1931)
Distribution: Morocco
20 MOR
subspecies :
subsp. maweana
(Ball) Maire; Cat. Pl. Maroc 638 (1934)
Distribution: Morocco
20 MOR
Synonyms: hidden
= Stachys maweana Ball; J. Linn. Soc., Bot. 16: 626 (1878)
= Stachys saxicola var. maweana (Ball) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 102(11): 1534 (2017)
subspecies :
subsp. villosissima
(Ball) Maire; Cat. Pl. Maroc 1113 (1941)
Distribution: Morocco
20 MOR
Synonyms: hidden
= Stachys saxicola var. villosissima Ball; J. Linn. Soc., Bot. 16: 626 (1878)
= Stachys saxicola subsp. platyodon Maire; Bull. Soc. Hist. Nat. Afrique N. 23: 208 (1932)
species :
Stachys scaberula
Vatke; Bot. Zeitung (Berlin) 33: 462 (1875)
Distribution: Bhutan; NE-India (Assam)
40 BHU IND
species :
Stachys scabrida
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 349 (1910)
Distribution: South Africa (W-Cape Prov., E-Cape Prov.)
27 SAF
Synonyms: hidden
= Stachys priorii Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 353 (1910)
species :
Stachys schimperi
Vatke; Linnaea 37: 330 (1871 publ. 1873)
Distribution: Sudan; Eritrea; Saudi Arabia
24 ERI SUD 35 SAU
Synonyms: hidden
= Stachys palaestina var. schimperi (Vatke) Benth.; Fl. Trop. Afr. [Oliver et al.] 5: 467 (1900)
species :
Stachys semsurensis
Fırat; Phytotaxa 511(3): 276, figs. 1-3 (2021)
Distribution: Turkey
34 TUR
species :
Stachys serbica
Pančić; Fl. Serbiae 564 (1874)
Distribution: Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (scattered NW-Greece)
13 ALB BIH BUL GRC KOS MNT NMA SRB
Synonyms: hidden
= Koevia serbica (Pančić) Krestovsk.; Novosti Sist. Vyssh. Rast. 45: 94 (2014)
species :
Stachys sericantha
P.H.Davis; Kew Bull. [6]: 111 (1951)
Distribution: Turkey (SW-Anatolia)
34 TUR
Remarks: libanotica agg.
species :
Stachys sericea
Cav.; Descr. Pl. [Cavanilles] 75 (1801)
Distribution: Chile (Valparaíso, Metropolitana de Santiago, O´Higgins, Maule, Ñuble, Biobío, Araucanía, Los Lagos)
85 CLE
Synonyms: hidden
= Stachys candidissima Phil.; Linnaea 29: 19 (1858)
= Stachys glabrata Phil.; Anales Univ. Chile 90: 559 (1895), nom. illeg.
= Stachys longifolia Bertero ex Steud.; Nomencl. Bot. [Steudel], ed. 2, 2: 628 (1841), nom. inval.
= Stachys sideritidoides Gillies ex Benth.; Labiat. Gen. Spec. [Bentham] 740 (1835)
species :
Stachys sericophylla
Halácsy; Consp. Fl. Graec. 2: 519 (1902)
Distribution: S-Albania
13 ALB
Remarks: clade 1 Eriostomum, germanica agg.
species :
Stachys sessilifolia
E.Mey.; Comm. Pl. Afr. Austr. [Meyer] 239 (1838)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.)
27 SAF
Synonyms: hidden
= Stachys bachmannii Gürke; Bot. Jahrb. Syst. 26: 75 (1898)
species :
Stachys sessilis
Gürke; Bot. Jahrb. Syst. 26: 74 (1898)
Distribution: South Africa (Mpulamanga, Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho
27 LES SAF
species :
Stachys setifera
C.A.Mey.; Verz. Pfl. Casp. Meer [C.A. von Meyer] 94 (1831)
subspecies :
subsp. setifera
Distribution: Kazakhstan; Uzbekistan; Tajikistan; Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia, SE-Anatolia: Mesopotamian Anatolia, SSW-Anatolia, SW-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, E-Iran, N-Iran, Iranian Aserbaijan, W-Iran); Afghanistan (Badakshan, Faryab, Kabul); Lebanon (Antilebanon)
32 KAZ TZK UZB 33 ARM GEO 34 AFG IRN IRQ LEB TUR
Remarks: clade 4 Setifolia
Synonyms: hidden
= Stachys bornmuelleri Gand.; Bull. Soc. Bot. France 65: 68 (1918)
= Stachys daenensis Gand.; Bull. Soc. Bot. France 65: 68 (1918)
= Stachys iranica Rech.f.; Oesterr. Bot. Z. 99: 44 (1952)
= Stachys lycopsiformis K.Koch; Linnaea 21: 692 (1848)
= Stachys repens Schur; Enum. Pl. Transsilv. 538 (1866)
= Stachys setifera subsp. daenensis (Gand.) Rech.f.; Fl. Iranica [Rechinger] 150: 366 (1982)
= Stachys setifera subsp. iranica (Rech.f.) Rech.f.; Fl. Iranica [Rechinger] 150: 366 (1982)
= Stachys shirini Parsa; Kew Bull. [3]: 226 (1948)
subspecies :
subsp. lycia
(Gand.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 281 (1974)
Distribution: Turkey
34 TUR
Synonyms: hidden
= Stachys lycia Gand.; Bull. Soc. Bot. France 65: 68, in clavi (1918)
species :
Stachys shikikunensis
(Kudô) comb.ined.
Distribution: Taiwan
38 TAI
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Suzukia shikikunensis Kudô; J. Soc. Trop. Agric. 2: 146 (1930)
= Glechoma shikikunensis (Kudô) Masam.; Hokuriku J. Bot. 4: 88 (1955)
species :
Stachys siirtensis
Ö.Güner & Akçiçek; Phytotaxa 516(3): 253 (2021)
Distribution: Turkey
34 TUR
species :
Stachys simplex
Schltr.; J. Bot. 35: 221 (1897)
Distribution: South Africa (Mpulamanga, Free State, KwaZulu-Natal); Eswatini [Swaziland]
27 SAF SWZ
Synonyms: hidden
= Stachys chrysotrichos Gürke; Bot. Jahrb. Syst. 28: 316 (1900)
= Stachys pascuicola Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1086 (1903)
species :
Stachys sivasica
Kit Tan & Yıldız; Notes Roy. Bot. Gard. Edinburgh 45(3): 449 (1988 publ. 1989)
Distribution: Turkey
34 TUR
species :
Stachys sparsipilosa
R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33: 287 (1974)
Distribution: Turkey (Inner Anatolia, S-Anatolia, SSW-Anatolia)
34 TUR
Synonyms: hidden
= Stachys iberica var. pallidiflora Boiss.; Fl. Orient. [Boissier] 4(2): 731 (1879), nom. illeg.
species :
Stachys spathulata
Burch. ex Benth.; Labiat. Gen. Spec. [Bentham] 559 (1833)
Distribution: Mozambique; Namibia; Botswana; South Africa (Limpopo, North-West Prov., Free State, KwaZulu-Natal, N-Cape Prov.); Madagascar
26 MOZ 27 BOT NAM SAF 29 MDG
Synonyms: hidden
= Stachys karasmontana Dinter; Repert. Spec. Nov. Regni Veg. 17: 203 (1921)
= Stachys minima Gürke; Bull. Herb. Boissier 6: 550 (1898)
= Stachys pachycalymna Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1087 (1903)
species :
Stachys spectabilis
Choisy ex DC.; Mém. Soc. Phys. Genève 1: 457 (1823)
Distribution: Georgia [Caucasus]; Armenia; Turkey (E-Anatolia, NE-Anatolia, S-Anatolia, SE-Anatolia); Iraq (NE-Iraq); Iran (EC-Iran, N-Iran, Iranian Aserbaijan, S-Iran, W-Iran); Israel (Rift Valley, N-Israel)
33 ARM GEO 34 IRN IRQ ISR TUR
Remarks: clade 1 Eriostomum, germanica agg.
Synonyms: hidden
= Stachys elata K.Koch; Linnaea 21: 687 (1848)
= Stachys germanica subsp. spectabilis (Choisy ex DC.) Briq.; Lab. Alp. Mar. 234 (1893)
= Stachys germanica var. elata (K.Koch) Briq.; Lab. Alp. Mar. 235 (1893)
= Stachys germanica var. hypoleuca (K.Koch) Briq.; Lab. Alp. Mar. 235 (1893)
= Stachys germanica var. spectabilis (Choisy ex DC.) Briq.; Lab. Alp. Mar. 234 (1893)
= Stachys hypoleuca K.Koch; Linnaea 21: 688 (1849)
= Stachys rosea Hohen.; Bull. Soc. Imp. Naturalistes Moscou 11: 300 (1838), no diagnostic descr.
= Stachys spectabiliformis Kapeller; Zametki Sist. Geogr. Rast. 16: 14 (1951)
= Stachys spectabilis subsp. elata (K.Koch) Krestovsk.; Novosti Sist. Vyssh. Rast. 33: 203 (2001)
species :
Stachys speluncarum
Contandr. & Quézel; Bull. Soc. Bot. France 123(7-8): 426 (1976)
Distribution: Turkey
34 TUR
species :
Stachys sphaerodonta
Baker; J. Linn. Soc., Bot. 20: 233 (1884)
Distribution: Madagascar
29 MDG
species :
Stachys spinosa
L.; Sp. Pl. [Linnaeus] 2: 581 (1753)
Distribution: Crete (widespread); East Aegaean Isl. (Karpathos)
13 KRI 34 EAI
Remarks: clade 11 Swainsoniana
species :
Stachys spinulosa
Sm.; Fl. Graec. Prodr. [Sibthorp] 1: 410 (1809)
Name: Dorniger Ziest (DE); Spiny Woundwort (EN)
Distribution: Croatia; ?Montenegro; ?Serbia; Kosovo; North Macedonia; Albania; Greece (C- & S-Greece, Kerkyra, Ionian Isl., Kithira, Sporades, Skiros, Evvoia, Andros, Kiklades: Amorgos, Astipaleia); W-Crete; East Aegaean Isl. (Lesvos, Ikaria, Samos, Kalimnos, Kos, Simi, Rhodos, Kastellorhizo Isl.); Turkey (NW-Anatolia: Bithynia, SSW-Anatolia, SW-Anatolia, W-Anatolia)
13 ALB CRO GRC KOS KRI NMA 34 EAI TUR
Synonyms: hidden
= Galeopsis orientalis Mill.; Gard. Dict., ed. 8: no. 5 (1768)
= Stachys betonicifolia Desf.; Ann. Mus. Hist. Nat. 10: 304 (1807), nom. illeg.
= Stachys orientalls (Mill.) Vahl; Symb. Bot. [Vahl] 2: 64 (1791), nom. illeg.
species :
Stachys splendens
Wall. ex Benth.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830)
Distribution: Tibet; Pakistan (Hazara, Murree, Astor); Nepal; Bhutan; N-India (Arunachal Pradesh, Assam, Himachal Pradesh, Jammu & Kashmir, Nagaland, Sikkim, Uttarakhand); Maldives [I]; Bangladesh
36 TIB 40 BAN BHU IND mdv NEP PAK
Synonyms: hidden
= Stachys bikramii Aswal & Mehrotra; J. Econ. Taxon. Bot. 4: 1028 (1983), nom. nov. superfl.
= Stachys emodi Hedge; Fl. Pakistan 192: 185 (1990) (nom. nov.)
= Stachys melissifolia Benth.; Labiat. Gen. Spec. [Bentham] 538 (1834)
= Stachys splendens Wall.; Pl. Asiat. Rar. [Wallich] 1: 64 (1830), nom. inval.
species :
Stachys spreitzenhoferi
Heldr.; Oesterr. Bot. Z. 30. 344 (1880)
subspecies :
subsp. spreitzenhoferi
Distribution: S-Greece (Malea peninsula, Kithira)
13 GRC
Remarks: clade 11 Swainsoniana, candida agg.
subspecies :
subsp. virella
D.Perss.; Biosyst. Stachys 38 (1981)
Distribution: S-Greece (Malea peninsula)
13 GRC
Synonyms: hidden
= Stachys spreitzenhoferi var. virella (D.Perss.) Krestovsk.; Novosti Sist. Vyssh. Rast. 48: 120 (2017)
species :
Stachys sprucei
Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 117 (1898)
Distribution: Ecuador
83 ECU
species :
Stachys spruneri
Boiss.; Prodr. [A.P. de Candolle] 12: 488 (1848)
Distribution: C-Greece (Attiki)
13 GRC
Remarks: clade 11 Swainsoniana, swainsonii agg.
species :
Stachys stebbinsii
G.A.Mulligan & D.B.Munro; Naturaliste Canad. 116(1): 48 (1989 publ. 1990)
Name: Stebbins´ hedgenettle
Distribution: USA (California)
76 USA
species :
Stachys stricta
Greene; Erythea 2: 122 (1894)
Name: Sonoma hedgenettle
Distribution: USA (California)
76 USA
Synonyms: hidden
= Stachys ingrata Greene; Pittonia 3: 341 (1898)
species :
Stachys strictiflora
C.Y.Wu; Acta Phytotax. Sin. 10: 220 (1965)
Distribution: China (Yunnan)
36 CHI
variant :
var. latidens
C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 221 (1965)
Distribution: China (Yunnan)
36 CHI
species :
Stachys subaphylla
Rech.f.; Pl. Syst. Evol. 134: 288 (1980)
Distribution: Iran (N-Iran)
34 IRN
Remarks: clade 12 Olisia
species :
Stachys sublobata
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 354 (1910)
Distribution: South Africa (W-Cape Prov.)
27 SAF
species :
Stachys subnuda
Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 50 (1836)
Distribution: Turkey (E-Anatolia, SE-Anatolia)
34 TUR
species :
Stachys swainsonii
Benth.; Labiat. Gen. Spec. [Bentham] 535 (1834)
subspecies :
subsp. swainsonii
Distribution: C-Greece (Mt. Giona to Mt. Parnassos)
13 GRC
Remarks: clade 11 Swainsoniana, swainsonii agg.
Synonyms: hidden
= Prasium hirsutum Poir.; Encycl. [J.Lamarck et al.] 5: 611 (1804)
= Prasium minus Vahl ex Walp.; Repert. Bot. Syst. [Walpers] 3: 819 (1845)
subspecies :
subsp. argolica
(Boiss.) Phitos & Damboldt; Ber. Deutsch. Bot. Ges. 82: 600 (1969)
Distribution: S-Greece (NE-Peloponnisos)
13 GRC
Synonyms: hidden
= Stachys argolica Boiss.; Diagn. Pl. Orient. 12: 83 (1853)
subspecies :
subsp. scyronica
(Boiss.) Phitos & Damboldt; Ber. Deutsch. Bot. Ges. 82: 600 (1969)
Distribution: C-Greece (around Korinth)
13 GRC
Synonyms: hidden
= Stachys scyronica Boiss.; Diagn. Pl. Orient. 5: 31 (1844)
= Stachys swainsonii var. scyronica (Boiss.) Nyman; Consp. Fl. Eur. 580 (1881)
species :
Stachys sylvatica
L.; Sp. Pl. [Linnaeus] 2: 580 (1753)
Name: Wald-Ziest (DE), Hedge Woundwort (EN)
Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Denmark; Norway; Sweden; Finland; Netherlands; Belgium; Luxembourg; Germany (Brandenburg, Berlin, Baden-Württemberg, Bayern, Hessen, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Saarland, Sachsen, Sachsen-Anhalt, Thüringen); Switzerland; Liechtenstein; Austria; Poland; Czech Republic; Slovakia; Hungary; Portugal; Spain; Andorra; France; Channel Isl. (Alderney, Guernsey, Jersey, Sark); Corsica; Italy; Sicily; Malta; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; European Turkey; Greece (mountains of N-, NE-Greece, south to Mt. Pelion); Estonia; Latvia; Lithuania; Belarus; C-European Russia; E-European Russia; N-European Russia; W-European Russia; Ukraine; Crimea; Madeira (Madeira Isl.); Siberia (Altai, Krasnoyarsk, W-Siberia); Kazakhstan; Turkmenistan; Uzbekistan; Kyrgyzstan; Northern Caucasus; Georgia [Caucasus]; Armenia; Azerbaijan; Turkey (E-Anatolia, N-Anatolia, NE-Anatolia, NW-Anatolia: Bithynia); Iran (N-Iran, Iranian Aserbaijan); China (Xinjiang); China [I]; NW-India (Himachal Pradesh, Jammu & Kashmir); New Zealand [I]; Canada [I] (Ontario [I]); USA [I] (New Jersey [I], New York [I], Ohio [I], Pennsylvania [I])
10 DEN FIN GRB IRE NOR SWE 11 AUT BGM CZE GER HUN LIE LUX NET POL SLO SWI 12 ADO CIS COR FRA POR SPA 13 ALB BIH BUL CRO GRC ITA KOS MLT MNT NMA ROM SIC SLV SMA SRB TUE 14 BLR EST KRY LAT LIT RUS UKR 21 MDR 30 SIB 32 KAZ KGZ TKM UZB 33 ARM AZE GEO NCS 34 IRN TUR 36 CHI 40 IND 51 nzl 72 cdn 75 usa
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys canariensis Jacq.; Icon. Pl. Rar. [Jacquin] 1: 11, t. 108 (1787)
= Stachys cordata Gilib.; Fl. Lit. Inch. 1: 80 (1782), opus utique oppr.
= Stachys cordatifolia Gilib.; Exerc. Phyt. 1: 91 (1792), opus utique oppr.
= Stachys foetida Gueldenst. ex Ledeb.; Fl. Ross. [Ledebour] 3: 413 (1849)
= Stachys glaucescens Muss.-Puschk. ex Spreng.; Syst. Veg., ed. 16 [Sprengel] 2: 736 (1825)
= Stachys sylvatica f. bracteata (Beck) L.Gross; Mitt. Bad. Bot. Vereins 210/211: 82 (1906)
= Stachys sylvatica var. minus Gray; Nat. Arr. Brit. Pl. 2: 372 (1821 publ. 1822)
= Stachys sylvatica unranked mollissima Schultheiss; Abh. Naturhist. Ges. Nürnberg 13, Beil. 1 [Phan.-Fl. Nürnberg-Erlangen]: 633 (1900)
= Stachys sylvatica var. tomentosa Čelak.; Arch. Naturwiss. Landesdurchf. Böhmen 4(3) (Prodr. Fl. Böhmen 4): 840 (1881)
= Stachys trapezuntea Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 38 (1859)
species :
Stachys taliensis
C.Y.Wu; Acta Phytotax. Sin. 10: 225 (1965)
Distribution: China (Yunnan)
36 CHI
species :
Stachys talyschensis
Kapeller; Zametki Syst. Geogr. Rast. 15: 48 (1949)
Distribution: +Azerbaijan
33 AZE
species :
Stachys tamaulipana
B.L.Turner; Phytologia 95(1): 28 (2013)
Distribution: Mexico (Nuevo León, Tamaulipas)
79 MEX
species :
Stachys tenerrima
Epling; Repert. Spec. Nov. Regni Veg., Beih. 80: 32 (1934)
Distribution: Mexico (Baja California Sur)
79 MEX
species :
Stachys tenuifolia
Willd.; Sp. Pl., ed. 4 [Willdenow] 3: 100 (1800)
Name: Smooth hedgenettle
Distribution: Canada (Manitoba, New Brunswick, Ontario, Québec); USA (Alabama, Arkansas, Connecticut, District of Columbia, Delaware, Florida, Georgia, Iowa, Illinois, Indiana, Kansas, Kentucky, Louisiana, Massachusetts, Maryland, Maine, Michigan, Minnesota, Missouri, Mississippi, North Carolina, North Dakota, Nebraska, New Hampshire, New Jersey, New York, Ohio, Oklahoma, Pennsylvania, Rhode Island, South Carolina, South Dakota, Tennessee, Texas, Virginia, Vermont, Wisconsin, West Virginia)
71 CDN 72 CDN 74 USA 75 USA 77 USA 78 USA
Remarks: clade 2 Stachys s. str.
Synonyms: hidden
= Stachys cincinnatensis Kuntze; Revis. Gen. Pl. 2: 531 (1891)
= Stachys glabra Riddell; W. J. Med. Phys. Sci. 9(36): 583 (1836)
= Stachys palustris var. glabra (Riddell) A.Gray; Manual [A.Gray], ed. 2: 317 (1856)
= Stachys palustris var. macrocalyx Jenn.; J. Wash. Acad. Sci. 10: 458 (1920)
= Stachys tenuifolia var. aspera Fernald; Rhodora 10: 85 (1908)
= Stachys tenuifolia var. perlonga Fernald; Rhodora 45: 466 (1943)
= Stachys tenuifolia var. platyphylla Fernald; Rhodora 45: 468 (1943)
species :
Stachys tetragona
Boiss. & Heldr.; Fl. Orient. [Boissier] 4(2): 736 (1879)
Distribution: C-Greece (Sporades, N-Evvoia)
13 GRC
Remarks: clade 11 Swainsoniana
species :
Stachys thirkei
K.Koch; Linnaea 21: 685 (1849)
Name: Thirke-Ziest (DE)
Distribution: Germany [I]; S-Italy; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Bulgaria; European Turkey; Greece (very rare C-, NE-Greece, N-Peloponnisos); East Aegaean Isl. (Chios); Turkey (N-Anatolia, NW-Anatolia: Bithynia, W-Anatolia, WN-Anatolia)
11 ger 13 BIH BUL CRO GRC ITA KOS MNT NMA SRB TUE 34 EAI TUR
Remarks: germanica agg.
Synonyms: hidden
= Stachys janiana Ces., Pass. & Gibelli; Comp. Fl. Ital. [Cesati] 318 (1874)
= Stachys thirkei var. condensata Boiss.; Fl. Orient. [Boissier] 4(2): 720 (1879)
species :
Stachys thunbergii
Benth.; Labiat. Gen. Spec. [Bentham] 540 (1833)
Distribution: South Africa (W-Cape Prov., E-Cape Prov.)
27 SAF
Synonyms: hidden
= Galeopsis hispida Thunb.; Prodr. Pl. Cap. 2: 96 (1800)
= Stachys hispida (Thunb.) Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 263 (1896), nom. illeg.
species :
Stachys thuspeinantha
Sennikov; Checkl. Vasc. Pl. Tian-Shan 351 (2021), nom. nov.
Distribution: Kazakhstan; Turkmenistan; Iraq (W-Iraq: Desert, NW-Iraq); Iran (EC-Iran, NE-Iran: Mts., Iranian Aserbaijan, S-Iran, W-Iran); Afghanistan (Farah, Herat); Syria (C-Syrian Desert); Jordania (S-Jordania, W-Jordania); Saudi Arabia (N-Saudi Arabia)
32 KAZ TKM 34 AFG IRN IRQ JOR SYR 35 SAU
Remarks: clade 12 Olisia
Synonyms: hidden
= Stachys musili Velen.; Sitzungsber. Königl. Böhm. Ges. Wiss. Prag, Math.-Naturwiss. Cl. (Vĕstn. Král. České Společn. Nauk., Tř. Mat.-Přír.) 1911: 5 (1911 publ. 1912)
= Tapeinanthus persicus Boiss.; Prodr. [A.P. de Candolle] 12: 436 (1848)
= Thuspeinanta persica (Boiss.) Briq.; Nat. Pflanzenfam. [Engler & Prantl] IV, 3a: 229 (1896)
species :
Stachys tlaxiacana
B.L.Turner; Phytologia 95(1): 28 (2013)
Distribution: Mexico (Oaxaca)
79 MEX
species :
Stachys tmolea
Boiss.; Diagn. Pl. Orient. 5: 29 (1844)
Distribution: Turkey (Inner Anatolia, NW-Anatolia: Bithynia, W-Anatolia, WN-Anatolia)
34 TUR
Remarks: germanica agg.
species :
Stachys tournefortii
Poir.; Encycl. [J.Lamarck et al.] 13: 227 (1817)
Distribution: NW-Crete; Libya
20 LBY
Remarks: germanica agg.
Synonyms: hidden
= Stachys sieberi K.Koch; Linnaea 21: 685 (1849)
species :
Stachys trichophylla
Baker; J. Linn. Soc., Bot. 25: 342 (1890)
Distribution: Madagascar
29 MDG
species :
Stachys trinervis
Aitch. & Hemsl.; Trans. Linn. Soc. London, Bot., Ser. 2, 3: 9, t. 43 (1888)
Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran); Afghanistan (Badghys, Farah, Faryab, Herat)
32 TKM 34 AFG IRN
Remarks: clade 12 Olisia
species :
Stachys truncata
Kunze ex Benth.; Labiat. Gen. Spec. [Bentham] 550 (1834)
Distribution: Chile (Coquimbo, Valparaíso, Maule, Aisén)
85 CLE
species :
Stachys tubulosa
MacOwan; Bull. Misc. Inform. Kew 1893: 13 (1893)
Distribution: South Africa (KwaZulu-Natal, E-Cape Prov.); Eswatini [Swaziland]
27 SAF SWZ
Synonyms: hidden
= Lamiostachys tubulosa (MacOwan) Krestovsk.; Bot. Zhurn. (Moscow & Leningrad) 91(8): 1258 (2006)
= Stachys dolichodeira Briq.; Bull. Herb. Boissier, Sér. 2, 3: 1081 (1903)
species :
Stachys tundjeliensis
Kit Tan & Sorger; Pl. Syst. Evol. 155(1-4): 99 (1987)
Distribution: Turkey
34 TUR
species :
Stachys turcomanica
Trautv.; Trudy Imp. S.-Peterburgsk. Bot. Sada 9: 463 (1886)
Distribution: Turkmenistan; Iran (EC-Iran, NE-Iran: Mts., N-Iran)
32 TKM 34 IRN
Remarks: clade 12 Olisia
species :
Stachys turkestanica
(Regel) Popov ex Knorring; Fl. URSS [Komarov et al., eds.] 21: 209 (1954)
Distribution: Uzbekistan; Kyrgyzstan; Tajikistan
32 KGZ TZK UZB
Synonyms: hidden
= Stachys alpina var. turkestanica Regel; Trudy Imp. S.-Peterburgsk. Bot. Sada, prepr. 6: 366 (1879)
= Stachys hissarica subsp. turkestanica (Regel) Derv.-Sok.; Novosti Sist. Vyssh. Rast. 14: 205 (1977)
= Stachys hissarica var. turkestanica (Regel) Krestovsk.; Novosti Sist. Vyssh. Rast. 42: 206 (2011)
= Stachys tschatkalensis Knorring; Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 15: 344 (1953)
species :
Stachys turneri
Rzed. & Calderón; Acta Bot. Mex. 32: 4 (1995)
Distribution: Mexico (Guanajuato)
79 MEX
species :
Stachys tymphaea
Hausskn.; Mitt. Geogr. Ges. (Thüringen) Jena 5: 70 (1886)
Distribution: C- & S-Italy; Serbia; Kosovo; North Macedonia; Albania; Bulgaria; Greece (mountains of NE-, NC-, NW- to C-Greece, south to Mt. Parnassos)
13 ALB BUL GRC ITA KOS NMA SRB
Remarks: germanica agg.
Synonyms: hidden
= Stachys alpina var. reinertii (Heldr.) Stoj.; Fl. Reipubl. Popularis Bulg. (Fl. Narodna Republ. Bulg.) 9: 914 (1967)
= Stachys germanica subsp. tymphaea (Hausskn.) R.Bhattacharjee; Notes Roy. Bot. Gard. Edinburgh 33(2): 276 (1974)
= Stachys germanica var. reinertii (Heldr.) Nyman; Consp. Fl. Eur. Suppl. 2: 251 (1889)
= Stachys reinertii Heldr.; Exsicc. Herb. Graecum Normale 1857: 743 (1857)
species :
Stachys tysonii
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 357 (1910)
Distribution: South Africa (Free State, KwaZulu-Natal, E-Cape Prov.); Lesotho
27 LES SAF
Synonyms: hidden
= Stachys obtusifolia var. angustifolia Skan
species :
Stachys uniflora
A.Pool; Novon 17(1): 64 (2007)
Distribution: Costa Rica; Panama
80 COS PAN
species :
Stachys venezuelana
Briq.; Annuaire Conserv. Jard. Bot. Genève 2: 114 (1898)
Distribution: Venezuela (Merida, Trujillo); Ecuador
82 VEN 83 ECU
species :
Stachys venulosa
Greene; Pittonia 1: 157 (1888)
Distribution: Mexico (Durango, etc.)
79 MEX
species :
Stachys veroniciformis
Rech.f.; Pl. Syst. Evol. 134: 289 (1980)
Distribution: Iran (SW-Iran)
34 IRN
species :
Stachys virgata
Bory & Chaub.; Expéd. Sci. Morée, Bot. 3(2): 166 (1832)
Distribution: S-Greece (E-Peloponnisos)
13 GRC
Synonyms: hidden
= Stachys zuccarinii Benth.; Labiat. Gen. Spec. [Bentham] 535 (1833)
species :
Stachys viscosa
Montbret & Aucher ex Benth.; Ann. Sci. Nat., Bot., Sér. 2, 6: 49 (1836)
Distribution: Georgia [Caucasus]; Azerbaijan; Turkey (E-Anatolia, Inner Anatolia, N-Anatolia, NE-Anatolia); Iran (Iranian Aserbaijan)
33 AZE GEO 34 IRN TUR
Synonyms: hidden
= Stachys laetivirens Kotschy & Boiss. ex Rech.f.; Bot. Jahrb. Syst. 71: 529 (1941)
= Stachys odontophylla Freyn; Oesterr. Bot. Z. 41: 59 (1891)
species :
Stachys viticina
Boiss.; Diagn. Pl. Orient. 12: 77 (1853) et Fl. 4: 724 (1879)
Distribution: Turkey (S-Anatolia); Lebanon (C-Lebanon, coastal W-Lebanon); Syria (W-Syrian Mountains); Israel (coastal W-Israel, Rift Valley, N-Israel)
34 ISR LEB SYR TUR
species :
Stachys vulnerabilis
Rzed. & Calderón; Acta Bot. Mex. 3: 3 (1988)
Distribution: Mexico (Coahuila, Nuevo León)
79 MEX
species :
Stachys vuralii
Yıldız, Dirmenci & Akçiçek; Ann. Bot. Fenn. 48(5): 403 (2011)
Distribution: Croatia; Azerbaijan; Turkey
13 CRO 33 AZE 34 TUR
species :
Stachys willemsei
Kit Tan & Hedge; Notes Roy. Bot. Gard. Edinburgh 45(3): 451 (1988 publ. 1989)
Distribution: Turkey
34 TUR
species :
Stachys woronowii
(Schischk. ex Grossh.) R.R.Mill; Fl. Turkey 7: 253 (1982)
Distribution: Armenia; Turkey (E-Anatolia, Inner Anatolia, NE-Anatolia, SSW-Anatolia); Iran (Iranian Aserbaijan); Lebanon (Antilebanon, C-Lebanon); China (Xinjiang)
33 ARM 34 IRN LEB TUR 36 CHI
Synonyms: hidden
= Sideritis balansae Boiss.; Diagn. Pl. Orient., Sér. 2, 4: 35 (1859) et Fl. 4: 707 (1879) (non Stachys balansae Boiss. & Kotschy (1879))
= Sideritis woronowii Schischk. ex Grossh.; Fl. Kavkaza [Grossheim] 3: 290 (1932)
= Stachys pseudosideritis R.Bhattacharjee & Hub.-Mor.; Notes Roy. Bot. Gard. Edinburgh 33(2): 291 (1974) (nom. nov.)
species :
Stachys xanthantha
C.Y.Wu; Acta Phytotax. Sin. 10: 224 (1965)
Distribution: China (Sichuan)
36 CHI
Synonyms: hidden
= Stachys xanthantha var. gracilis C.Y.Wu & H.W.Li; Acta Phytotax. Sin. 10: 225 (1965)
species :
Stachys yemenensis
Hedge; Notes Roy. Bot. Gard. Edinburgh 40: 71, f. 4 (1982)
Distribution: Saudi Arabia (Asir); Yemen (N-Inner Yemen, W-Yemen)
35 SAU YEM
Remarks: clade 12 Olisia
species :
Stachys yildirimlii
Dinç; Ann. Bot. Fenn. 43(2): 143 (2006)
Distribution: Turkey
34 TUR
species :
Stachys yingzuijieensis
L.Wu & Y.P.Chen; PhytoKeys 236: 127 (2023)
Distribution: China (Hunan)
36 CHI
species :
Stachys zepcensis
Formánek; Oesterr. Bot. Z. 38: 383 (1888)
Distribution: Bosnia & Hercegovina; Serbia
13 BIH SRB
Remarks: clade 11 Swainsoniana
Synonyms: hidden
= Stachys anisochila subsp. zepcensis (Formánek) Vandas; Reliq. Formán. [Vandas] 469 (1909)
= Stachys labiosa var. zepcensis (Formánek) K.Malý; Verh. K.K. Zool.-Bot. Ges. Wien 54: 246 (1904)
= Stachys recta var. chrysophaea Hayek; Denkschr. Akad. Wiss. Wien, Math.-Naturwiss. Kl. 99: 183 (1924)
species :
Stachys zeyheri
Skan; Fl. Cap. [Harvey & auct. succ.] 5(1): 363 (1910)
Distribution: South Africa (W-Cape Prov.)
27 SAF
species :
Stachys zoharyana
Eig; Palestine J. Bot., Jerusalem Ser. 4: 172, f. 1 (1948)
Distribution: Israel (coastal W-Israel, N-Israel)
34 ISR
species :
Stachys ×ambigua
Sm.; Engl. Bot. [Smith & Sowerby] 30: t. 2089 (1809)
Name: Hybrid Woundwort (EN)
Distribution: England (England, Wales, Isle of Man, Scotland, Outer Hebrides, Orkney Isl., Shetland Isl., Isles of Scilly); Ireland (Ireland, Northern Ireland); Belgium; Germany (Baden-Württemberg, Bayern, Hamburg, Mecklenburg-Vorpommern, Niedersachsen, Nordrhein-Westfalen, Rheinland-Pfalz, Schleswig-Holstein, Sachsen, Sachsen-Anhalt, Thüringen); France; Czech Republic; Slovakia; Channel Isl. (Alderney, Guernsey, Jersey); Turkey (E-Anatolia); etc.
10 GRB IRE 11 BGM CZE GER SLO 12 CIS FRA 34 TUR
Remarks: clade 2 Stachys s. str., Stachys palustris x sylvatica
Synonyms: hidden
= Stachys palustris subsp. ambigua (Sm.) Arcang.; Comp. Fl. Ital. [Arcangeli] 552 (1882)
= Stachys palustris subsp. ambigua (Sm.) Hartm.; Sv. Norsk Excurs.-Fl. 85 (1846)
= Stachys palustris var. ambigua (Sm.) Bluff & Fingerh.; Comp. Fl. German. [Bluff & Fingerhuth] 1: 31 (1825)
= Stachys palustris var. ambigua (Sm.) Hook.; Fl. Bor.-Amer. [Hooker] 2: 116 (1838)
= Stachys palustris ×sylvatica var. wirtgenii (F.W.Schultz) Bornm.; Beih. Bot. Centralbl. 37(2): 315 (1919)
= Stachys palustri-sylvatica Schiede; De Pl. Hybr. Spont. Nat. 42 (1825)
= Stachys ×ambigua f. ellipticus Clos; Bull. Soc. Bot. France 36: 71 (1889)
= Stachys ×ambigua f. longepetiolatus Clos; Bull. Soc. Bot. France 36: 71 (1889)
= Stachys ×ambigua f. petiolatus Gams; Ill. Fl. Mitt.-Eur. [Hegi] 5(4): 2435 (1927)
= Stachys ×ambigua var. elliptica Wallr.; Scholion zu Hampe´s Prodr. Fl. Hercyn.: 222 (1840), nom. inval.
= Stachys ×ambigua var. lanceolata Wallr.; Scholion zu Hampe´s Prodr. Fl. Hercyn.: 222 (1840)
= Stachys ×ambigua var. ovatifolia Wallr.; Scholion zu Hampe´s Prodr. Fl. Hercyn.: 222 (1840), nom. inval.
= Stachys ×wirtgenii F.W.Schultz; Arch. Fl. 1(2): 117 (1861), nom. inval.
species :
Stachys ×bodeana
Bunge; Mém. Acad. Imp. Sci. St.-Pétersbourg, Sér. 7, 21(1): 71 (1873), nom. nov.
Distribution: Iran (EC-Iran, Iranian Aserbaijan, W-Iran)
34 IRN
Remarks: Stachys inflata x lavandulifolia.
Synonyms: hidden
= Stachys lavandulifolia var. brachyodon Boiss.; Fl. Orient. [Boissier] 4: 743 (1873)
= Stachys ×tomentosa Benth.; Prodr. [A.P. de Candolle] 12: 489 (1848), nom. illeg.
= Stachys ×tomentosa Benth. ex Rech.f.; Fl. Iranica [Rechinger] 150: 395, t. 360 (1982)
species :
Stachys ×burrii
P.H.Davis; Kew Bull. [6]: 109 (1951)
Distribution: Turkey (SW-Anatolia)
34 TUR
Remarks: clade 1 Eriostomum, germanica agg.; Stachys cretica ssp. mersinaea x Stachys sericantha
species :
Stachys ×cryptadenia
Rech.f.; Bot. Jahrb. Syst. 71: 538 (1941)
Distribution: Croatia (Dalmatia)
13 CRO
Remarks: clade 1 Eriostomum, germanica agg.; Stachys cretica ssp. salviifolia x thrikei
species :
Stachys ×digenea
Legué; Bull. Soc. Bot. France 40: 213 (1893)
Distribution: Czech Republic; Slovakia; etc.
11 CZE SLO
Remarks: clade 1 Eriostomum, germanica agg.; Stachys alpina x germanica
species :
Stachys ×leucomalla
Bornm. & Gauba; Repert. Spec. Nov. Regni Veg. 49: 271 (1940)
Distribution: Iran (N-Iran)
34 IRN
Remarks: Stachys inflata x laxa
species :
Stachys ×medebachensis
Feld & O.Koenen; Jahresber. Westfäl. Prov.-Vereins Wiss. 41: 183 (1913)
Distribution: Germany (Bayern, Nordrhein-Westfalen); Switzerland; etc.
11 GER SWI
Remarks: clade 1 Eriostomum, germanica agg.; Stachys alpina x sylvatica
Synonyms: hidden
= Stachys ×hybrida Brügger; Jahresber. Naturforsch. Ges. Graubünden, N. F. 23-24: 102 (1880), nom. inval.
= Stachys ×medebachensis f. superalpina Feld & O.Koenen; Jahresber. Westfäl. Prov.-Vereins Wiss. 41: 183 (1913), nom. inval.
= Stachys ×medebachensis f. supersilatica Feld & O.Koenen; Jahresber. Westfäl. Prov.-Vereins Wiss. 41: 183 (1913), nom. inval.
species :
Stachys ×mirabilis
Rouy; Fl. France [Rouy & Foucaud] 11: 310 (1909)
Distribution: France; etc.
12 FRA
Remarks: clade 1 Eriostomum, germanica agg.; Stachys germanica x palustris
species :
Stachys ×paraplesia
Rech.f.; Fl. Iranica [Rechinger] 150: 395, t. 361 (1982)
Distribution: Iran (EC-Iran, N-Iran)
34 IRN
Remarks: Stachys inflata x turcomanica
species :
Stachys ×sintenisii
Gand.; Bull. Soc. Bot. France 65: 68 (1918)
Distribution: Turkmenistan; NE-Iran (EC-Iran, NE-Iran: Mts.)
32 TKM 34 IRN
Remarks: Stachys lavandulifolia x turcomanica
