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Ver. 19.2
April 17th, 2024

456.0000 Family Rubiaceae Juss.; Gen. Pl.: 196 (1789), nom. cons.
589 genera, ca. 14.200 spp. mostly tropics, comparably few temperate
Species listed in database: 14207; hybrids: 48
= Aparinaceae Hoffmanns. & Link; Fl. Portug. 2: 3, 38 (1813-1829)
= Asperulaceae Cham. ex Spenn.; Handb. Angew. Bot. 2: 493 (1835)
= Catesbaeaceae Martinov; Tekhno-Bot. Slovar: 111 (1820)
= Cephalanthaceae Raf.; Ann. Gén. Sci. Phys. Bruxelles 6: 86 (1820)
= Cinchonaceae Batsch; Tab. Affin. Regni Veg.: 234 (1802)
= Coffeaceae Batsch; Tab. Affin. Regni Veg.: 233 (1802)
= Coutareaceae Martinov; Tekhno-Bot. Slovar: 168 (1820)
= Dialypetalanthaceae Rizzini & Occhioni; Lilloa 17: 253 (1948), nom. cons.
= Galiaceae Lindl.; Nat. Syst. Bot., ed. 2: 249 (1836)
= Gardeniaceae Dumort.; Anal. Fam. Pl.: 29, 32 (1829)
= Guettardaceae Batsch; Tab. Affin. Regni Veg.: 235 (1802)
= Hameliaceae Mart.; Consp. Regn. Veg.: 31 (1835)
= Hedyotidaceae Dumort.; Comment. Bot.: 57 (1822)
= Henriqueziaceae Bremek.; Acta Bot. Neerl. 6 (Meded. Bot. Mus. Herb. Rijksuniv. Utrecht 141): 371 (1957)
= Houstoniaceae Raf.; Good Book: 21 (1840)
= Hydrophylacaceae Martinov; Tekhno-Bot. Slovar: 318 (1820)
= Lippayaceae Meisn.; Pl. Vasc. Gen.: Tab. Diagn. 156, Comm. 112 (1838)
= Lygodisodeaceae Bartl.; Ord. Nat. Pl.: 123, 207 (1830)
= Naucleaceae Wernham; New Phytol. 11: 225 (1912)
= Nonateliaceae Martinov; Tekhno-Bot. Slovar: 420 (1820)
= Operculariaceae Juss. ex Perleb; Vers. Arzneikr. Pfl.: 207 (1818)
= Pagamaeaceae Martinov; Tekhno-Bot. Slovar: 447 (1820)
= Psychotriaceae F.Rudolphi; Syst. Orb. Veg.: 49 (1830)
= Randiaceae Martinov; Tekhno-Bot. Slovar: 534 (1820)
= Sabiceaceae Martinov; Tekhno-Bot. Slovar: 555 (1820)
= Spermacoceaceae Bercht. & J.Presl; Prir. Rostlin: 256 (1820)
= Theligonaceae Dumort.; Anal. Fam. Pl.: 15, 17 (1829), nom. cons.
456.2739 Genus Thliphthisa (Griseb.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 776 (2020)
Distribution: SE-Europe, E-Mediterranean to Iran
Species listed in database: 23
species :
Thliphthisa antalyensis (Ehrend.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Turkey (SW-Anatolia)
34 TUR
Synonyms: hidden
= Asperula antalyensis Ehrend.; Notes Roy. Bot. Gard. Edinburgh 22: 352, f. 3b (1958)
= Asperula pseudochlorantha var. antalyensis (Ehrend.) Minareci & K.Yildiz; Ann. Bot. Fenn. 47(2): 122 (2010)
species :
Thliphthisa apuana (Fiori) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Italy
13 ITA
Synonyms: hidden
= Asperula apuana (Fiori) Arrigoni; Webbia 67(1): 42 (2012)
= Asperula purpurea subsp. apuana (Fiori) Bechi & Garbari; Flora Medit. 4: 223 (1994)
= Galium purpureum var. apuanum Fiori; Nuov. Fl. Italia 2: 491 (1927)
species :
Thliphthisa baenitzii (Heldr. ex Boiss.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: S-Greece (Attiki)
13 GRC
Synonyms: hidden
= Asperula baenitzii Heldr. ex Boiss.; Fl. Or., Suppl. 280 (1888)
species :
Thliphthisa baldaccii (Halácsy) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Montenegro (S-Crna Gora)
13 MNT
Conservation Status: extremely threatened, confined to one population only
Synonyms: hidden
= Asperula baldaccii (Halácsy) Ehrend.; Bot. Jour. Linn. Soc. 68: 269 (1974)
= Galium baldaccii Halácsy; Österr. Bot. Zeitschr. 40: 165 (1890)
species :
Thliphthisa brevifolia (Vent.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Name: Kurzblättriger Meister (DE); Short-leaved Woodruff (EN)
Distribution: East Aegaean Isl. (incl. Rhodos); Turkey (SSW-Anatolia, SW-Anatolia); Lebanon (C-Lebanon); Syria (coastal W-Syria)
34 EAI LEB SYR TUR
Synonyms: hidden
= Asperula brevifolia Vent.; Descr. Pl. Jard. Cels. 63, t. 63 (1802)
= Asperula brunnea Boiss.; Fl. Orient. 3: 32 (1875)
species :
Thliphthisa chlorantha (Boiss. & Heldr.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Albania; NW-Greece
13 ALB GRC
Synonyms: hidden
= Asperula chlorantha Boiss. & Heldr.; Boiss., Diagn. Pl. Or. Nov. 3(6): 90 (1859)
species :
Thliphthisa crassula (Greuter & Zaffran) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Crete (NW-Crete, NE-Crete)
13 KRI
Synonyms: hidden
= Asperula crassula Greuter & Zaffran; Willdenowia 14(2): 289 (1984 publ. 1985)
species :
Thliphthisa elonea (Iatroú & T.Georgiadis) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Greece
13 GRC
Synonyms: hidden
= Asperula elonea Iatroú & T.Georgiadis; Willdenowia 14(1): 55 (1984)
species :
Thliphthisa gorganica (Schönb.-Tem. & Ehrend.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: N-Iran
34 IRN
Synonyms: hidden
= Asperula gorganica Schönb.-Tem. & Ehrend.; Bot. Jahrb. Syst., 107(1-4): 80 (1985)
species :
Thliphthisa mazanderanica (Ehrend.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Iran (N-Iran)
34 IRN
Synonyms: hidden
= Asperula mazanderanica Ehrend.; Ann. Naturh. Mus. Wien 56: 225, ff. 4-5 (1948)
species :
Thliphthisa microphylla (Boiss.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Iran (NW-Iran)
34 IRN
Synonyms: hidden
= Asperula microphylla Boiss.; Diagn. Pl. Orient. 3: 34 (1843)
species :
Thliphthisa muscosa (Boiss. & Heldr.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: EC-Greece (Olimbos)
13 GRC
Synonyms: hidden
= Asperula muscosa Boiss. & Heldr.; Boiss., Diagn. Pl. Or. Nov. 3(2): 109 (1856)
species :
Thliphthisa pseudochlorantha (Ehrend.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Turkey (SW-Anatolia)
34 TUR
Synonyms: hidden
= Asperula pseudochlorantha Ehrend.; Notes Roy. Bot. Gard. Edinburgh 22: 351, f. 3a (1958)
species :
Thliphthisa purpurea (L.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
subspecies :
subsp. purpurea
Name: Rotes Labkraut
Distribution: Switzerland; Austria; SE-France; Italy; San Marino; Slovenia; Croatia; Bosnia & Hercegovina; Montenegro; Serbia; Kosovo; North Macedonia; Albania; Romania; Bulgaria; Greece
11 AUT SWI 12 FRA 13 ALB BIH BUL CRO GRC ITA KOS MNT NMA ROM SLV SMA SRB
Synonyms: hidden
= Asperula purpurea (L.) Ehrend.; Österr. Bot. Zeitschr. 122: 260 (1973)
= Asperula purpurea var. luteoviride (Rohlena) Ancev; Fl. Nar. Republ. Bulgariya 9: 41 (1989)
= Asperula purpurea var. trichantha (Vand.) Ancev; Fl. Nar. Republ. Bulgariya 9: 41 (1989)
= Crucianella purpurea (L.) Honck.; Verz. Gew. Teutschl.: 431 (1782)
= Galium purpureum L.; Sp. Pl. 107 (1753)
= Galium purpureum var. luteoviride Rohlena; Sitz. Ber. Böhm. Ges. Wiss.: 54 (1911)
= Galium purpureum var. trichanthum Vand.; Reliq. Forman.: 259 (1909)
= Galium rubrum Scop.; Fl. Carniol., ed. 2, 1: 103 (1771), nom. illeg.
subspecies :
subsp. apiculata (Sm.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Croatia; Serbia; Kosovo; Albania; European Turkey; Greece
13 ALB CRO GRC KOS SRB TUE
Synonyms: hidden
= Asperula purpurea subsp. apiculata (Sm.) Ehrend.; Bot. Jour. Linn. Soc. 68: 269 (1974)
= Galium apiculatum Sm.; Sibth. & Sm., Fl. Graec. Prodr. 1: 91 (1806)
= Galium pycnophyllum Heldr. ex Boiss.; Fl. Orient. 3: 64 (1875)
species :
Thliphthisa pusilla (Bory & Chaub.) J.Krieg., Del Guacchio & P.Caputo; Willdenowia 51(1): 158 (2021)
Distribution: S-Greece (Peloponnisos)
13 GRC
Synonyms: hidden
= Asperula boryana (Walp.) Ehrend.; Bot. Jour. Linn. Soc. 68: 269 (1974)
= Galium boryanum Walp.; Repert. Bot. Syst. 2: 454 (1843)
= Sherardia pusilla Bory & Chaub.; Nouv. Fl. Pélop.: 10 (1838)
= Thliphthisa boryana (Walp.) P.Caputo & Del Guacchio; Pl. Biosystems 154: 777 (2020), nom. superfl.
species :
Thliphthisa rigida (Sm.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Crete
13 KRI
Synonyms: hidden
= Asperula kritsensis Coustur. & Gand.; Bull. Soc. Bot. France 63: 9, 14 (1916 publ. 1917)
= Asperula platygona Gand.; Bull. Soc. Bot. France 62: 154 (1915 publ. 1916)
= Asperula rigida Sm.; Fl. Graec. Prodr. 1: 89 (1806)
= Crucianella rigida (Sibth. & Sm.) Tausch; Flora 18: 359 (1835)
species :
Thliphthisa rupestris (Vis.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: Croatia; Bosnia & Hercegovina; Serbia; Kosovo; Albania; ?Romania
13 ALB BIH CRO KOS SRB
Synonyms: hidden
= Asperula rupestris (Vis.) Rchb.; Fl. Germ. Excurs.: 205 (1831), nom. illeg.
= Asperula scutellaris Vis.; Fl. Dalm. 3: 12 (1852)
= Galium rupestre Vis.; Flora 12(1 Erg.): 5 (1829)
species :
Thliphthisa sapphus Gutermann; PhytoKeys 241: 67 (2024)
Distribution: Greece
13 GRC
species :
Thliphthisa saxicola (Ehrend.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 777 (2020)
Distribution: S-Greece (E-Peloponnisos)
13 GRC
Synonyms: hidden
= Asperula saxicola Ehrend.; Bot. Jour. Linn. Soc. 68: 269 (1974)
species :
Thliphthisa semanensis (Schönb.-Tem. & Ehrend.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 778 (2020)
Distribution: Syria
34 SYR
Synonyms: hidden
= Asperula semanensis Schönb.-Tem. & Ehrend.; Bot. Jahrb. Syst., 107(1-4): 81 (1985)
species :
Thliphthisa serotina (Boiss. & Heldr.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 778 (2020)
Distribution: Turkey (SSW-Anatolia, SW-Anatolia)
34 TUR
Synonyms: hidden
= Asperula serotina (Boiss. & Heldr.) Ehrend.; Davis, Fl. Turkey 7: 766 (1982)
= Galium serotinum Boiss. & Heldr.; Boiss., Diagn. Pl. Orient. 10: 71 (1849)
species :
Thliphthisa suberosa (Sm.) P.Caputo & Del Guacchio; Pl. Biosystems 154: 778 (2020)
Distribution: Cyprus (C-Mountains, E-Cyprus, N-Cyprus, S-Cyprus, W-Cyprus)
34 CYP
Synonyms: hidden
= Asperula cypria Ehrend.; Pl. Syst. Evol. 126: 406 (1977)
= Galium suberosum Sm.; Sibth. & Sm., Fl. Graec. Prodr. 1: 91 (1806)
species :
Thliphthisa tournefortii (Sieber ex Spreng.) P.Caputo & Del Guacchio; Pl. Biosystems 154(5): 778 (2020)
Name: Tournefort´s Meister (DE); Tournefort´s Woodruff (EN)
Distribution: Greece (Kiklades); Crete; East Aegaean Isl. (incl. Rhodos)
13 GRC KRI 34 EAI
Synonyms: hidden
= Asperula majori Barbey; Bull. Herb. Boissier 2: 243 (1894)
= Asperula tournefortii Sieber ex Spreng.; Syst. Veg. 1. 395 (1824)
= Crucianella tournefortii (Sieber ex Spreng.) Tausch; Flora 18: 359 (1835)
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A complete, synonymic checklist of the Higher Plants of the World


Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life