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452.0000 Family Asteraceae Dumort.; Commentat. Bot. [Dumortier] 55 (1822)
1620 genera, ca. 29.000 spp. (35.000 spp. incl. microspecies). Systematics taken mostly from Funk et al. (2009, 2020) and Zuntini et al. (2024). Barnadesioideae (so far treated as most basal member of Asteraceae) might be a separate family, more related to Calyceraceae.
Species listed in database: 35255; hybrids: 678
= Acarnaceae Link; (1829)
= Ambrosiaceae Bercht. & J.Presl; Prír. Rostlin 254 (1820), nom. cons.
= Anthemidaceae Bercht. & J.Presl; Prír. Rostlin 254 (1820)
= Aposeridaceae Raf.; New Fl. [Rafinesque] 4: 106 (1838)
= Arctotaceae Bessey; Univ. Stud. Nebraska 14: 107 (1914)
= Arctotidaceae Bercht. & J.Presl; Prír. Rostlin 255 (1820)
= Artemisiaceae Martinov; Tekhno-Bot. Slovar 48 (1820)
= Athanasiaceae Martinov; Tekhno-Bot. Slovar 56 (1820)
= Calendulaceae Bercht. & J.Presl; Prír. Rostlin 254 (1820)
= Carduaceae Bercht. & J.Presl; Prír. Rostlin 255 (1820)
= Carlinaceae Bercht. & J.Presl; Prír. Rostlin 255 (1820)
= Centaureaceae Bercht. & J.Presl; Prír. Rostlin 255 (1820)
= Cichoriaceae Juss.; Gen. Pl. [Jussieu] 168 (1789), nom. cons.
= Cnicaceae Vest; Anleit. Stud. Bot. 273, 297 (1818)
= Compositae Giseke; Prael. Ord. Nat. Pl. 538 (1792), nom. cons.
= Coreopsidaceae Link; Handbuch [Link] 1: 768 (1829)
= Cynaraceae Burnett; Outlines Bot. [Burnett] 926, 931, 1094, 1111 (1835)
= Echinopaceae Bercht. & J.Presl; Prír. Rostlin 255 (1820)
= Eupatoriaceae Bercht. & J.Presl; Prír. Rostlin 253 (1820)
= Gnaphaliaceae Link ex F.Rudolphi; Syst. Orb. Veg. 46 (1830)
= Grindeliaceae Rchb. ex A.Heller; Muhlenbergia 2: 330 (1907)
= Heleniaceae Raf.; Cincinnati Lit. Gaz. 2: 28 (1824)
= Helianthaceae Bercht. & J.Presl; Prír. Rostlin 254 (1820)
= Helichrysaceae Link; Handbuch [Link] 1: 712 (1829)
= Inulaceae Bercht. & J.Presl; Prír. Rostlin 254 (1820)
= Ivaceae Raf.; Fl. Tellur. 1: 31 (1837)
= Lactucaceae Drude; Handb. Bot. 3(2): 369 (1886)
= Lapsanaceae Martinov; (1820)
= Madiaceae A.Heller; Muhlenbergia 2: 332 (1907)
= Matricariaceae Voigt; Hort. Suburb. Calcutt. 400 (1845)
= Mutisiaceae Burnett; Outlines Bot. [Burnett] 934, 935, 1094, 1111 (1835)
= Nassauviaceae Burmeist.; Handb. Naturgesch. 290 (1836)
= Partheniaceae Link; Handbuch [Link] 1: 816 (1829)
= Perdiciaceae Link; Handbuch [Link] 1: 728 (1829)
= Picridaceae Martinov; Tekhno-Bot. Slovar 482 (1820)
= Santolinaceae Martinov; Tekhno-Bot. Slovar 560 (1820)
= Senecionaceae Bercht. & J.Presl; Prír. Rostlin 254 (1820)
= Serratulaceae Martinov; Tekhno-Bot. Slovar 577 (1820)
= Tanacetaceae Vest; Anleit. Stud. Bot. 273, 298 (1818)
= Tussilagaceae Bercht. & J.Presl; Prír. Rostlin 254 (1820)
= Vernoniaceae Burmeist.; Handb. Naturgesch. 296 (1836)
= Xanthiaceae Vest; Anleit. Stud. Bot. 273, 298 (1818)
= Xeranthemaceae Döll; Rhein. Fl. [Döll] 498 (1843)
452.6366 Genus Werneria Kunth; Nov. Gen. Sp. [H.B.K.], ed. fol. 4: 189. tt. 368, 369 (1818)
Distribution: Andes. Revision: Calvo & Moreira-Munoz (2020, as Xenophyllum)
Species listed in database: 22
= Oresigonia Willd. ex Less.; Syn. Gen. Compos. 393 (1832), Prodr. [A.P. de Candolle] 6: 323 (1837)
= Xenophyllum Funk; Brittonia 49(1): 110-117 (1997)
species :
Werneria acerosa Cuatrec.; Brittonia 8: 45 (1954)
Distribution: Ecuador (Azuay, ?Zamora-Chinchipe)
83 ECU
Synonyms: hidden
= Xenophyllum acerosum (Cuatrec.) V.A.Funk; Novon 7(3): 238 (1997)
species :
Werneria amblydactyla S.F.Blake; J. Wash. Acad. Sci. 18: 490 (1928)
Distribution: Peru (Junín, Lima)
83 PER
Synonyms: hidden
= Xenophyllum amblydactylum (S.F.Blake) V.A.Funk; Novon 7(3): 238 (1997)
species :
Werneria ciliolata A.Gray; Proc. Amer. Acad. Arts 5: 140 (1861)
Name: Pupusa, poposa, yaretilla [Chile]
Distribution: Peru (widespread in Andes); Bolivia (La Paz, Cochabamba, Oruro, Potosí); Chile (Arica y Parinacota, Tarapacá, Antofagasta)
83 BOL PER 85 CLE
Synonyms: hidden
= Xenophyllum ciliolatum (A.Gray) V.A.Funk; Novon 7(3): 239 (1997)
species :
Werneria crassa S.F.Blake; J. Wash. Acad. Sci. 18: 495 (1928)
subspecies :
subsp. crassa
Distribution: Colombia (Arauca, Boyacá, Caldas, Cauca, Cundinamarca, Norte de Santander, Quindío, Risaralda, Santander, Tolima, Valle); Ecuador
82 CLM 83 ECU
Synonyms: hidden
= Xenophyllum crassum (S.F.Blake) V.A.Funk; Novon 7(3): 239 (1997)
subspecies :
subsp. orientalis Cuatrec.; Phytologia 45(1): 29 (1980)
Distribution: Colombia (Arauca, Boyacá, Norte de Santander, Santander)
82 CLM
Synonyms: hidden
= Xenophyllum crassum subsp. orientale (Cuatrec.) J.Calvo; PhytoKeys 158: 45 (2020)
species :
Werneria dactylophylla Sch.Bip.; Bonplandia (Hannover) 4: 53 (1856), 55, nomen, et ex Wedd. Chlor. Andina 1: 87
Distribution: Peru; Bolivia (Cochabamba, La Paz, Potosí)
83 BOL PER
Synonyms: hidden
= Werneria dactylophylla f. glabriuscula Rockh.; Bot. Jahrb. Syst. 70: 286 (1939)
= Werneria dactylophylla var. glandulosodenticulata Rockh.; Bot. Jahrb. Syst. 70: 286 (1939)
= Xenophyllum dactylophyllum (Sch.Bip.) V.A.Funk; Novon 7(3): 239 (1997)
= Xenophyllum oscartovarii Linares, J.Campos, Nauray, Vicente Orell. & A.Galán; Arnaldoa 17(1): 104 (2010)
species :
Werneria decora S.F.Blake; J. Wash. Acad. Sci. 18: 491 (1928)
Distribution: Peru (Ancash, Ayacucho, Huancavelica, Huánuco, Junín, Lima)
83 PER
Synonyms: hidden
= Xenophyllum decorum (S.F.Blake) V.A.Funk; Novon 7(3): 239 (1997)
species :
Werneria digitata Wedd.; Chlor. Andina 1: 86 (1855)
Distribution: Peru; Bolivia (Potosí, Oruro)
83 BOL PER
Synonyms: hidden
= Xenophyllum digitatum (Wedd.) V.A.Funk; Novon 7(3): 239 (1997)
= Werneria digitata var. lanata Rockh.; Bot. Jahrb. Syst. 70: 287 (1939)
species :
Werneria esquilachensis Cuatrec.; Brittonia 8: 192 (1956)
Distribution: Peru; Bolivia (Oruro); Chile (Arica y Parinacota, Tarapacá, Antofagasta)
83 BOL PER 85 CLE
Synonyms: hidden
= Senecio pfisteri Ricardi & Martic.; Gayana, Bot. 11: 25 (1964)
= Xenophyllum esquilachense (Cuatrec.) V.A.Funk; Novon 7(3): 239 (1997)
species :
Werneria funkiana (J.Calvo) D.J.N.Hind; Kew Bull. 77: 705 (2022)
Distribution: Ecuador (Chimborazo, Bolívar)
83 ECU
Synonyms: hidden
= Xenophyllum funkianum J.Calvo; PhytoKeys 139: 30 (2020)
species :
Werneria humilis Kunth; Nov. Gen. Sp. [H.B.K.], ed. fol. 4: 191 (1818)
Distribution: Colombia (Boyacá, Caldas, Cauca, Cundinamarca, Huila, Meta, Nariño, Risaralda, Tolima, Valle); Ecuador (widespread); Peru (widespread); Bolivia (Cochabamba, La Paz)
82 CLM 83 BOL ECU PER
Synonyms: hidden
= Oresigonia brevifolia Willd. ex Wedd.; Chlor. Andina 1: 82 (1856), pro syn.
= Werneria articulata S.F.Blake; Contr. U.S. Natl. Herb. 22: 651 (1924)
= Werneria fontii (Cuatrec.) Cuatrec.; Trab. Mus. Ci. Nat., Ser. Bot. 29: 42 (1935), cum descr.
= Werneria humilis f. articulata (S.F.Blake) Rockh.; Bot. Jahrb. Syst. 70(3): 294 (1939)
= Werneria humilis var. fontii Cuatrec.; Trab. Mus. Nac. Ci. Nat., Ser. Bot. 27: cuadro 22 (1934)
= Werneria humilis var. lindenii Wedd.; Chlor. Andina 1: 82 (1856)
= Werneria lehmannii Hieron.; Bot. Jahrb. Syst. 28: 647 (1901), nom. illeg.
= Werneria lehmannii Klatt; Ann. K.K. Naturhist. Hofmus. 9: 368 (1894)
= Xenophyllum fontii (Cuatrec.) V.A.Funk; Novon 7(3): 239 (1997)
= Xenophyllum humile (Kunth) V.A.Funk; Novon 7(3): 239 (1997)
species :
Werneria incisa Phil.; Anales Mus. Nac., Santiago de Chile 40 (1891)
Distribution: Peru; Bolivia (Potosí); Argentina (Jujuy, Salta); Chile (Arica y Parinacota, Tarapacá, Antofagasta)
83 BOL PER 85 ARG CLE
Synonyms: hidden
= Xenophyllum incisum (Phil.) V.A.Funk; Novon 7(3): 239 (1997)
species :
Werneria juniperina Hieron.; Bot. Jahrb. Syst. 21: 365 (1895)
Distribution: Peru (Tacna); Bolivia (Oruro, Potosí); Chile (Arica y Parinacota)
83 BOL PER 85 CLE
Synonyms: hidden
= Werneria lycopodioides S.F.Blake; J. Wash. Acad. Sci. 18: 493 (1928)
= Xenophyllum juniperinum (Hieron.) J.Calvo; Phytotaxa 326(3): 228 (2017)
= Xenophyllum lycopodioides (S.F.Blake) V.A.Funk; Novon 7(3): 240 (1997)
species :
Werneria lorochaqui (J.Calvo & V.A.Funk) Kew Bull.77: 705 (2022)
Distribution: Argentina (Catamarca, Salta, Jujuy, Tucumán)
85 ARG
Synonyms: hidden
= Xenophyllum lorochaqui J.Calvo & V.A.Funk; PhytoKeys 139: 34 (2020)
species :
Werneria marcida S.F.Blake; J. Wash. Acad. Sci. 18: 492 (1928)
Distribution: Peru (Ancash, Arequipa, Ayacucho, Huancavelica, Huanuco, Junin, Lima, Moquegua, Pasco, Puno); Bolivia (La Paz)
83 BOL PER
Synonyms: hidden
= Xenophyllum marcidum (S.F.Blake) V.A.Funk; Novon 7(3): 240 (1997)
= Werneria sedoides S.F.Blake; J. Wash. Acad. Sci. 18: 493 (1928)
species :
Werneria poposa Phil.; Anales Mus. Nac., Santiago de Chile 2(Bot. 8): 40 (1891)
Distribution: Peru; Bolivia (Oruro, Potosí); Argentina (Catamarca, La Rioja, Salta, Jujuy, Tucuman); Chile (Arica y Parinacota, Tarapacá, Antofagasta)
83 BOL PER 85 ARG CLE
Synonyms: hidden
= Werneria humilis Griseb. ex Hieron.; Bot. Jahrb. Syst. 21(3): 364 (1895), nom. inval. pro syn.
= Werneria incisa var. pubescens Rockh.; Bot. Jahrb. Syst. 70(3): 290 (1939)
= Werneria lorentziana Hieron.; Bot. Jahrb. Syst. 21(3): 364 (1895)
= Xenophyllum incisum var. pubescens (Rockh.) Cabrera & S.E.Freire; Monogr. Syst. Bot. Missouri Bot. Gard. 74 (Cat. Pl. Vasc. Rep. Argent. 2): 1245 (1999)
= Xenophyllum poposum (Phil.) V.A.Funk; Novon 7(3): 240 (1997) (´poposum´)
species :
Werneria pseudodigitata Rockh.; Bot. Jahrb. Syst. 70: 288 (1939)
Distribution: Argentina (Jujuy, Salta); Chile (Arica y Parinacota, Tarapacá, Antofagasta)
85 ARG CLE
Synonyms: hidden
= Werneria castillensis Hieron.; nom. nud. in sched.
= Xenophyllum pseudodigitatum (Rockh.) V.A.Funk; Novon 7(3): 240 (1997)
species :
Werneria rigida Kunth; Nov. Gen. Sp. [H.B.K.], ed. fol. 4: 191 (1818)
Distribution: Ecuador (Bolivar, Chimborazo, Cotopaxi, Napo, Pichincha, Tungurahua)
83 ECU
Synonyms: hidden
= Oresigonia pycnophylla Bonpl. ex Wedd.; Chlor. Andina 1: 83 (1856), pro syn.
= Xenophyllum rigidum (Kunth) V.A.Funk; Novon 7(3): 240 (1997)
species :
Werneria rosenii R.E.Fr.; Nova Acta Regiae Soc. Sci. Upsal., Ser. 4, 1(1): 90 (1905)
Distribution: Bolivia (Oruro); Argentina (Jujuy, ?Salta)
83 BOL 85 ARG
Synonyms: hidden
= Xenophyllum rosenii (R.E.Fr.) V.A.Funk; Novon 7(3): 240 (1997)
species :
Werneria rosea Hieron.; Bot. Jahrb. Syst. 28: 648 (1901)
Distribution: Ecuador (Azuay, Canar, Loja)
83 ECU
Synonyms: hidden
= Werneria humilis var. rosea (Hieron.) Rockh.; Bot. Jahrb. Syst. 70(3): 295 (1939)
= Werneria purpurea Spruce ex Rockh.; Bot. Jahrb. Syst. 70: 295 (1939)
= Xenophyllum roseum (Hieron.) V.A.Funk; Novon 7(3): 240 (1997)
species :
Werneria sotarensis Hieron.; Bot. Jahrb. Syst. 21: 363 (1895)
Distribution: Colombia (Cauca, Nariño); Ecuador
82 CLM 83 ECU
Synonyms: hidden
= Werneria leucobryoides S.F.Blake; J. Wash. Acad. Sci. 18: 494 (1928)
= Xenophyllum sotarense (Hieron.) V.A.Funk; Novon 7(3): 240 (1997)
species :
Werneria staffordiae Sandwith; Hooker´s Icon. Pl. 35: t. 3424 (1940)
Distribution: C-Peru; S-Peru
83 PER
Synonyms: hidden
= Xenophyllum staffordiae (Sandwith) V.A.Funk; Novon 7(3): 240 (1997)
species :
Werneria weddellii Phil.; Anales Mus. Nac., Santiago de Chile 40 (1891)
Distribution: Peru (Arequipa, Moquegua, Puno, Tacna); Bolivia (Oruro); Chile (Arica y Parinacota, Tarapacá, Antofagasta)
83 BOL PER 85 CLE
Synonyms: hidden
= Werneria decumbens Hieron.; Bot. Jahrb. Syst. 21: 364 (1895)
= Xenophyllum weddellii (Phil.) V.A.Funk; Novon 7(3): 240 (1997)
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A complete, synonymic checklist of the Higher Plants of the World

Version 25.05 (May 23rd, 2025)

Covering 356,547 accepted species,
8,782 hybrids, 48,090 accepted infraspecific taxa and 1,003,107 synonyms,
in total 1,416,616* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]

With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,209 pictures of 15,787 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life